Journal of Natural History

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Journal of Natural History Publication details, including instructions for authors and subscription information: http://www.informaworld.com/smpp/title~content=t713192031 A review of the species of Calvia (Coleoptera: Coccinellidae) from the Indian subcontinent, with descriptions of two new species R. G. Booth a a International Institute of Entomology, An Institute of CAB International, London, UK

Online Publication Date: 01 June 1997 To cite this Article: Booth, R. G. (1997) 'A review of the species of Calvia (Coleoptera: Coccinellidae) from the Indian subcontinent, with descriptions of two new species', Journal of Natural History, 31:6, 917 - 934 To link to this article: DOI: 10.1080/00222939700770451 URL: http://dx.doi.org/10.1080/00222939700770451

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The publisher shall not be liable for any loss, actions, claims, proceedings, demand or costs or damages whatsoever or howsoever caused arising directly or indirectly in connection with or arising out of the use of this material. © Taylor and Francis 2007 Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 JOURNAL A nin ucniet with subcontinent, Indian Introduction have America. pale trees beetles, out Entomology almost on to tata preted characters. subcontinent C. Himalayan 1980's. review Iablokoff-Khnzorian various quatuordecimpunctata The For (Fabricius) also the species and entirely many and Iablokoff-Khnzorian Chrysomelidae genus OF been In shrubs Palaearctic International The 56 continent R. Acpe 1 Otbr 1996) October 15 (Accepted biodiversity on of species shown Miyatake K~YWOROS: are Himalaya, C. C. C. external region. NATURAL general, are for which andrewesi trees for years, championorum G. the Queen's trilochana designated recorded. species Calvia on are identification. el with dealt than to BOOTH the based species and characters be characters. are was specimens are They known India, they following of for HISTORY, herbaceous a valid species, valid a Gate, Mulsant shrubs, and (Weise) reviewed. Coleoptera, 0022-2933/97 predominantly synonymized Institute Kapur, (Kanervo, Calvia Some for (1982) submitted (Linnaeus), hs genus. this are are Pakistan, Nepal, Bhutan. Nepal, Pakistan, of and to C. London Oriental below. of predators although descriptions (1982) and keyed Calvia Mulsant Calvia have vishnu One feed This C. reasons. 1997, contains C. the belong The of $12"00 vegetation. flaveola C. pinaki Entomology, 1946). They of for not male on been paper under Coccinellidae, SW7 Many southern sykesii (Crotch), species out 31, provided on occurs regions, other © the (Coleoptera: (Coleoptera: the a identification of to 1997 Eight external synonym are 917-934 Kapur, submitted 20 genitalia, 5JR, below are purposes C. concentrates aphids They this eggs, (Crotch). species prey have prey Taylor & Francis Ltd. Taylor & of and, generally or widely part shiva described C. of the UK two region. a and more An krishna based C. characters are larvae in of of and key Coccinellidae) Kapur. Calvia, and pasupati particular, to of of Institute the C. in new more also predators of nine on ocnlia) from Coccinellidae) psyllids, the species punctata this to s new. as Calvia the They predominantly Himalayas on another (Crotch), and which been various species ava andrewesi Calvia International 15 species the s presented. is paper usually northern taxonomy, Kapur, pupae of can Calvia recorded. distributed species of Calvia are (Mulsant). although ava durgae Calvia CAB from to C. a aphids is all is occasions names 17 endemic associated of buddha to and a single the half now International, species quindecimgut- species region of various describe Two A morphology, on Institute and Indian C. other the Lectotypes unknown key Wie is (Weise) of through- be (Crotch), external species, connexa psyllids species, of Kapur, Indian to North in based based to inter- the with leaf- prey high sub- one the the the of Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 punctata Ambon, China, Caucasus, recognized (Fabricius), 918 required. the Iabtokoff-Khnzorian - from other - been walnut as Mader ry information Prey London. trees to and alni et Apices 2 and C. C. Calvia 1 C. 27 predators shiva al., shirozui margin transverse pronotum closer Anterior and Anterior Apices with (Fig. (Fig. single In Although Unless prey > 5 > quindecimguttata Ghosh Singh Parshad which found specimens species Homoptera addition mm species sculpture 1974). was C. leaves l segments all 2). below) 1). records Indonesia spur than (Mulsant) of muiri of broadly stated C. (1985) long; (Miyatake) Disc were Elytral margin with unknown et margin meso- and both species oeie similar, sometimes meso- of of chinensis present (1980), those prey Phaloura in from Key al. among of two to (Timberlake) ...... the examined of of if four heavily otherwise, Jammu widely black, elytra n Indian on noted as coloration aphidophagous less, elytra the and (Iablokoff-Khnzorian, of on to longer lists of subfamily the has species on are prey. (1982), respectively species however, metatibiae clypeus to elytra. species (Mulsant), clypeus six twelve were metatibiae then and mesotibiae. Indian interrupted been for and infested distributed ava trilochana Calvia him. chestnut n Kashmir, and than C. coccinellid in These pronotum all of Calvia regarded antennae Male alder not this of Singh broadly from recorded of decemguttata there A but Coccinellinae, species material broad. aphid epy concave, deeply subcontinent. recorded aphid on aphid with Calvia revision with as brown study host without genitalia usually C. by Coccinellidae Body in by China and Japan, and Japan, and China R.G. species seems above. (1991) across a on hauseri on Male species truncate, Uttar shorter Aphis three pair as dealt IablokoffoKhnzorian records as is also from with disc India size c. size Quercus dealbata Quercus a very deposited cotton of an a Booth a (Figs of median Kapur 1982). o be to synonym Punctation the large were pair listed very predator with craccivora Pradesh the indicate preying three (Linnaeus) apical Mader unidentified with they the Agarwala different (Pawar 4-5 lateral are Palaearctic, lateral 16-21) Asiatic of similar, yellow in very in published Indian Calvia parvinotata in oe etc. lobe, (see off-white mm had emns 9 segments ava albida Calvia summarized spurs apical northern India in and this on of aphids, and projections of The projections and little long n Parry, and C. no Koch spots at C. the and (Fig. Calvia Chromaphis juglandicola ...... subcontinent C. and, paper, spurs, shiva and Sikkim, most Calvia records Natural C. in sculpture muiri (Fig. to and C. aphid, septenaria information by Ghosh Afghanistan (Fig. (1982) at Manipur, psyllids anomala 3). and yellow although rosti pronotal antennae species C. Kapur in Ghara, Gordon although 22) relatively Body and the and species relatively tricolor 12). 10 for Table Mollitrichosiphum India. 1989). History (Weise) quindecimguttata of (1988) spots, (Miyatake) other and quadrate a and C. size (Crotch) Calvia. Punctation Mulsant elytra below) adults below) is subspecies punctures Nepal. India. they elongate, on 1. small, a (1985) from durgae Korschefsky shiva clearly Label narrow It usually (Stolyarov sometimes C. lateral broad Museum, currently from reviewed prey has referred and Wadhi sicardi Calvia to Kaput Singh Alnus from from data also and and (see still the for on of 3 2 Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 Table Species C. C. C. C. C. C C F[GS flaveola punctata shiva breiti tricolor o albida vulnerata Og 1-11. Unless 1. punctation; clypeus; outline (7) O Host C. punetata, Characters indicated (dorsal); Walnut Adelges Aphis Undetermined Undetermined Eggs Undetermined Mollitrichosiphum Mollitriehosiphum Maerosiphoniella Chromaphis Tuberculatus (3) Vesiculaphis records persicae Acyrthosiphon (Pentatomoidea: Cervaphis (5) C. of gossypii Calvia 0 quatuordecimguttata, ° C. (10) aphid elytral otherwise, (Adelgidae) for Urostylis :~0. of ~., ~J~o~ ° (Sulzer) monosha, a~o °~-'~ juglandicola Calvia Calvia C. quercus indicus o0.0 :~ species Glover, aphids, aphids aphids punctation; andrewesi, Prey °'~J o gossypii pseudoartemisiae punctigera alni alni species all spp. Urostylidae) Ghosh, Takahashi association elytral species Ghosh Ghosh from psyllids, Aphis : ( (1) Mordvilko, Kaltenbach) outline from (8) punctation; Westwood the craccivora tibial C 6 are et et C. scale the 0 tricolor, al. aL Indian Aphididae (lateral); "Ooo punctata, ° o Shinji o spurs; Indian o'a insects o,, ° Myzus o o Koch, (6) clypeus; a subcontinent "o (4) subcontinent (11 outline (sensu C. C. ) C vulnerata, quatuordecimguttata, (2) Phaloura andrewesi, Phaloura Pawar Data Data Singh Korschefsky, Data Data Data Stolyarov Data Data lato). (lateral); C. 7 and labels labels label labels labels label label quatuordecimguttata, and and S elytral . • o o o. adjacent and Source and o et • . outline Singh, (9) • Parry, ... o al., , :;:..'oo o; 1940 Qo Singh, Singh, C. o punctation; 1974 ""- punctata, "° o 1985 (dorsal). 1989 regions. elytral o 1991 1991 919 Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 920 4 FIGSl2-15. - - 5 3 lobe, pink eyes Pronotum and Larger and Elytral bands. (Fig. arrangement either Elytral seven obvious Pronotum Coloration (Fig. interstices with is with etc. 5). 5), well-defined with microreticulate Male punctation punctation elytral contrast or Elytral five (Fig. seven Habitus duller, shining larger very larger median different) large rather 23) punctures black coloration variable, between punctures punctures interstices, drawings pale on between on yellow lobe, convex ...... disc spots disc yellow ...... the close uneven, usually etc. spots clearly (arranged of black punctures, close more (Fig. at two Calvia spots (Fig. and least arranged with and with widely punctures (15) 4). double, R.G. 24) (arranged spp. deep, 2-2-2-1). Elytral on deep, basal C. contrasting six ...... : separated disc, flaveola. Booth 2-2-1, (12) or interstices a ...... variable interstices mixture coloration and fewer 1-3-2-l) microreticulate Body C. or tricolor; external and with spots spots length in of convex or convex size interstices either large head quatuordecimguttata partly (13) reddish-pink (if margins c. but 4-6 with with yellowish-brown C. where and as with rather joined vulnerata; mm. on flatter seven small narrowly small small frons, frons monosha close Male laterally to (Figs spots punctures, yellowish- punctures punctures (14) together between without median yellow (Linnaeus) 6, Bielawski with then into C. 7). breiti; 4 8 5 Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 - FIGS - 6 7 18 entirely Fig. large more Elytral Larger yellow Elytra orangey spots always Large lobe Species transverse punctures two Large except or 16-21. (ventral); (17) with diameters, 13, of than punctures absent elytral less elytral laterally colouration yellow elytral male same currently a spots for black, brown. variable 16 (Fig. Male one strongly subbasal j (21) then genitalia fine l punctures (lateral); either punctures diameter, punctures to usually 7). except (Fig. Male genitalia t( same known yellowish-brown, ...... elytra Calvia number rufous convex Elytral variable, band. yellow 6). median with (lateral). with for (18) on i usually with on from Elytral more of species Black (Fig. coloration of outer or very disc fine a disc but C. parameres narrow l spots, double lobe, black, the widely with 8), more tricolor: broad, rufous sublateral Scale coloration not generally from more margin, 17 generally rarely etc. but a with as very black row widely marker single separated explanate above (Fig. usually outer (dorsal); the northern (16) variable, or of black, line closer, suture variable, then row longer Indian without small spaced, 25) =0.5 Median --4-- margin, never ...... six of usually and parts sides (19) and pronotum separated mm, punctures from ...... (arranged in small subcontinent present. separated but a proportion interstices black lobe, (Fig. paler sipho; then of Figs entirely punctures with background the parameres, 29) sublateral or 16-19; pronotum on between If 3-2-1), (20) subcontinent. black six on elytra darker pale average to flatter spots average between sipho, breadth 0.25 front yellow punctata medially entirely colour adjacent line. outer vulnerata (Figs trabes arranged mm, unicolorous by detail three by adjacent (Fig. If to Median slightly ring. Figs almost black, about 6, elytra (Mulsant) black (ventral); large in and of 7). 9). (Hope) as 20, If a apex 921 21. 6 7 Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 922 - Flos 8 . Elytra rently Median Elytral 22-28. (27) (23) j/ known relatively C. punctation C. lobe sykesii; Median 26 quatuordecimguttata; of from 2" male shorter (28) lobe, southern clearly genitalia albida. C (Fig. parameres, double, India; 11) narrower Scale (24) and Nilgiri R.G. a trabes 27 mixture more marker=0'5mm. C. 24 (Fig. Booth and monosha; (ventral strongly ' " 30) of Anaimalai large view) (25) convex and Hills, of C. small (Fig. Calvia vulnerata; Kerala/Tamil 2: punctures, 10). spp.: . andrewesi Species (26) (22) Nadu. larger C. cur- C. (Weise) breiti; shiva; Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 - - 10 - 9 - FIGS 11 elytra. irregular Prosternal and Median often Prosternal median Elytra ized carinae. Larger apex carinae. slender Median stripes Coloration pronotum General to without Elytral Smaller obvious markings punctures Fig. 29-32. same (ventral); brownish-yellow, sperm 7). area of with rather on Sixth species, and punctation lobe, Ground sipho such (lateral); Narrowed species, lobe Narrowed dark i lobe shape darker (Fig. or Male broadly process more elytra. process duct yellow numerous sperm (30) with etc. obvious visible of of markings broader oblong as 43) 6.5-8.0mm genitalia relatively colour markings male 5-7-6.6mm male widely C. for (Fig. Sixth two (32) to convex, Calvia transverse. duct apical convex, apical even andrewesi, sometimes brownish-yellow, sternite other ...... contrast in genitalia slightly (Fig. or genitalia dull 27) C. nor relatively visible spaced, 30 of or shape, three portion portion longer andrewesi, species (Fig. but brownish-yellow Calvia without long. 40). with Calvia uneven, ...... long. of Elytra with paler median II between small sternite shorter mottled long 14). pronotum Base on male (Fig. small, Prosternal shorter of of species, from spp.: Prosternal carinae carinae. average or Male median with median sipho. dark if and of lobe, either 41 subtruncate large of and uneven dark darker or (29) bursa the ) (Figs two 31 slender median brown elytra male either with only parameres, ...... Scale broader process to lobe uniform Indian and brown Coloration separated lobe C. process or copulatrix greenish-yellow spots 42, then punctata, with paler weakly oval, three well small marker=0.5mm. spots of of (Figs lobe, 43) or spots male generally subcontinent male (Fig. punctures or forming deeper developed yellow generally 1 strongly small trabes weakly punctures. by etc. with 34, yellow transverse, with median genitalia genitalia 28)...... about (Fig. 38). dark emargination three (ventral); stripes, with larger, a with emarginate rounded Spermatheca variable with to or mottled Spermatheca brown 26) lobe, two Coloration paler shorter weak brownish-yellow, weak bold longer both weakly but well-developed diameters (31) parameres, longitudinal at flaveola dark spots. albida to in sykesii to pattern never with (Fig. (Fig. sides medially. medially. obsolete. size C. obsolete breiti sclerot- bulkier brown yellow andrewesi, more Male Bielawski bold with 38), 34), (Crotch) and and sp. (as on Mader trabes nov. 923 l 10 11 9 Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 924 FIGS 37 33-40. trabes genitalia 33-35, (38) same 33 (lateral); 37-39; Male of (ventral); J / / C. genitalia 0"25 flaveola, (34) 38 ram, (39) same of ; Figs sipho; paratype. C. 34 (ventral); 36, championorum, (40) R.G. 40. details (37) (35) Booth median of sipho; holotype: sipho 39 35 lobe, (36) apex. (33) details parameres, 40 Scale median 3 marker of sipho lobe, trabes =0"5 apex. parameres, mm, (lateral); Male Figs Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 that FIGS mens unsexed, for leave foveae. any (Fig. apex shown (Fig. this Dieke Collection Michelia Material Calvia the unknown Coelophora Comments < 0.66 sclerotized apex This Two Korschefsky study. original. other species this and 41-43. the 1) 12). from (Anisocalvia) Cotl'n in the As of also species specimens, species examined species the detail champaca 'Bagdogra The to Oriental width sipho it 1952'. appears three Mulsant support region lack Iablokoff-Khnzorian has Spermatheca 1965'. very is in (1940) does in more of not distinct localities of One Figs tricolor genus. at Calvia Calvia the both distinctive to a / Range, this been end not (including median slender 704 be (43) 3, 16-21. head described from notion. from of Korschefsky, properly restricted 'Assam, for For of possible species / in C. Calvia sperm TYPUS Kurseong, and flaveola. (Fig. the swelling Calvia The Bengal. ,/ example, male all the (Figs However, the Lemnia time this other from duct (1982) 36). tricolor United Shillong structure belong to to genitalia, deeply spp.: [printed 1, examine Scale species 1940: With being, or northeast (Fig. Base the Indian it Bengal, 12, lateral who Mulsant) States shares in Korschefsky marker Indian the (41) 2, 42) vi.1945 concave 16 of of the until Calvia. from Syntype illustrated Coccinellidae the on the species gave bursa 21) wings record N. India National C. characters = more subcontinent orange-red female sipho an 0.25 beating C. a but albida; anterior The copulatrix [examined] unrecorded does (cf. brief and ram. Chatterjee from material (Fig. here lacks relatively genitalia, Figs 43 Museum. is not description bushes with (42) in margin poorly Shillong, for championorum card] 19), its 32, with fit distinct becomes Synona the C. dorsal comfortably number 35, in 24.vi.1935 short it J. small, India: / known. of first particular championorum; seems 39), Unyal Korschefsky Meghalaya, taken the hypomeral coloration antennae, Pope time, available well- of suggests Syntype clypeus sp. best It / speci- / from into nov. GH and was 925 the are On to Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 unusual 926 mens from the being species' detailed pale centre, by extends A18353 do Kapur of showed spurs Uttar and examined Material Pradesh, Calvia shiva Calvia Calvia Calvia triloehana Calvia Bhatkot, (Miyatake, synonymy Calvia durgae Calvia Calvia connexa Calvia Oenopia dorsonotata Oenopia Coccinella 12-maculata Coccinella quatuordecimguttata: Calvia Coccinella 14-guttata Comments the Kaput This This not oe 1989:353 Pope, This The Sixteen onymized above C. elytral Mandal pinaki from pasupati pending n the on Pradesh, genitalia was present was (1963) pasupatL appear genitalia generic into that species is and character and s a is Sukhatal, Ranikhet (1963) synonymic a of specimens. Kapur, Mandal, margins, 1985). Kapur, very variably very by Kapur, accurate the single a very metatibiae, CIE C. Miyatake, Kapur, attributed and a o pl to apply to Kapur, West Belicek, 1976:327 placement, is which pasupati more drawings more all and distinct 1963: widespread Mulsant, in A19155. Kumaon, in Linnaeus, 1963: H. Kumaon, 1963: Uttar Sikkim, Gebler, 1963: similar Bhatkot, the list. the species India: drawings Miyatake Calvia occurred detailed 1963: northerly G. 40, 1985: 43, to 42, Mandal usn, 1846:140 Mulsant, with and from Coccinellidae. for Examination coloured 45, Champion; Holotype it Pradesh, the his 1850: Holotype 42, Holotypes This Holotype 1832: to 1758: is quatuordecimguttata 17, Singhik, Uttar was Calvia Holotype the 8000ft C. Kumaon and study species present Holotype probably other during the Calvia of (1985). Holotype 424, regions duodecimmaculata series. species 76, 367, Syntype [not examined] 367, [not Syntype (Figs single involved. the R.G. variably illustration Pradesh species male trilochana India, (Fig. male [examined] male of Lectotype [examined], synonymized [examined], Lectotype Calv& female [examined] female ytp nteaie] nm rocpe) syn- preoccupied), (name examined] [not Syntype May 5000ft can genitalia Uttar shiva of male slide species. of its Habitus 2-4, male [examined] male 4000 Although [examined] spur best of Booth male C. is 22) Kapur's largely true the and 1920 [examined] shiva, species preparation. the Kapur The matched also coloured Pradesh, 23) ft to 24.iv.1924 present [not examined] [not given atr comprising latter Iablokoff-Khnzorian's of position. I for May Indian retain figures H. differences have recorded (Linnaeus) be C. C. there holotypes in C. (Gebler) by G. Syn. durgae, explained connexa 1920 very lacking on Syn. Holarctic Syn. adl various Mandal, connexa. no it Iablokoff-Khnzorian of subcontinent. Syn. Champion; a. R. Maj. nov. within Freshly is the hesitation the H. nov. closely nov. from some nov. Syn. in C. is and mesotibiae a species G. but by trilochana, the erroneous. pair A Calvia species five nov. dissected Champion; Godavari, W. ob about doubt slight a without with black genitalia Uttar series of in colour were G. The (1979, by for the for apical establishing Miyatake's dates, distortions which form Hingston; Booth s a is C. Pradesh, of included material material the (1982) forms, pinaki Nepal which speci- Uttar 1982) tibial most time with CIE pale just and the Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 was examined Material the Khnzorian they spots BHUTAN: Belicek specimens. colour he larger Comments eoi dorsonotata, Oenopia 6.7-7"0 referred C. Calvia Phodrang (Fig. Material Anisocalvia Anisocalvia Anisocalvia indicated Coccinella Calvia Coccinella Calviaflaccida paratype quatuordecimguttata, gave Recent Five This Three lectotype Pope, This by and Booth by Booth 1989:368 originally synonymized 5) than monosha vulnerata: Booth than pattern, mm the specimens. species to will is a is examined examined 1989:367 (1976) and vulnerata by belonged uniramosa specimens. 15.vi. 1972 vishnu krishna (1982) also (1982) buddha authors, Bielawski the by in long, body Thimphu h above the and variably separate the was Mulsant, Pope, described Polska Bielawski, Iablokoff-Khnzorian typical male Korschefsky, was Pope, Pope, Crotch, list i.e. designated length Crotch, Crotch, Hope, from C. in NEPAL: Calvia 1989:368 to described Hope, for of and (Holotype scarcely described 'Bengal' dorsonotata Akademia Bhutan: coloured this recorded the 1989:368 1853: 1989:368 4.iii.1985 duodecimmaculata the regarding synonyms, and and the and from example Indian 1979: 1874: 1831: as female 1874: 1874: species dorsonotata species 1831: 7.3-7-6mm 151, 1932:529 following Caivia Kathmandu by longer 145, 122, Bengal ava vulnerata Calvia from 31, Holotype (Oxford Himalayan subcontinent the 145, Lectotype here designated [examined], synonymized designated here 145, Lectotype and 145, Booth genitalia, Lectotype [examined], synonymized same 31, Nauk, CIE Gordon Lectotype [examined] with from the from (Figs Lectotype designated here [examined], synonymized Holotype male [examined] male Holotype from is species species monosha C. (Figs near 3 Lectotype here designated [examined], synonymized designated here Lectotype (1982). ettp [xmnd,snnmzdb Booth by synonymized [examined], Lectotype than largely from paratype was A16905. three C. University and colour Warsaw). long, suggesting long, 6, others duodecimmaculata and Wangdi 4500 5, form. quatuordeeimguttata. C. antenna from (1985) implied 13, material in species. Indian Pope 24) due vulnerata. Bielawski pce, I species, Paratypes, detail 25) form (Hope) ft in in the Under [INDIA]: to 20.v.-23.vi. 1983 20.v.-23.vi. Phodrang, (1989). Naturhistorisches and Museum). the by and subcontinent The range The the Himalayas. in and which Booth F. region. Sasaji elytral The colour this measured figured that 1 W. 3, has Lectotype, dense latter name, latter Bhutan, and Pope and Hope's as this (1985) punctation. 1 ~ 1 of variation This Oenopia dorsonotata Oenopia much its colour M. by elytral a species the 21 body species Museum, Booth collection, J. based km have synonym smaller specimens (1989) D. unsexed, of varieties, punctation O shape Iablokoff- was Although Brendell. and followed the on was Wangdi Basel; black much when three Pope, spots and and 927 not and of of by at as Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 928 hand ment 39211 / dull black pale and original Crotch. the designated Kurseong (1982), from the Harmoniapunctata form Material Propylea Fry (University A. Dalbergia tion unique Crotch's Hardwicke designated 20.v.-23.vi.1983 Gokarna elytral specimens occupied specimen agree May krishna vishnu, University Calviapunctata: Comments buddha Twenty-five This Bielawski, The preceding same yellow their Coll. of areas. ]', Kurseong, of 1920 with very spots ground syntypes Anisocalvia is above although [Fry A descriptions. is black three examined obversepunctata by fitting 4500ft immediately hereby labels Inde sissoo have 1905-100', another closely all female lectotypes . G. H. a as The Bequest; of to of (arranged each 1963:17 black catalogue the 'India black. species specimens colour spots Cambridge synonymic Korschefsky, Verschraeghen Cambridge); and median specimens. from spot M. West the as leaf. designated 7.vi.1983 Mulsant, lectotype his Champion; with specimen spot very vishnu sublateral J. the Terai, buddha just description Orient'; Pale positions Alexander figures Bengal, for are NEPAL: may The D. is surrounding that 3-2-1), label[/TYPE lobe is preceding variably usn, 1853: Mulsant, hereby discernable represented Hope's Brendell. except and specimens list Bardia constant, 1853: typical have M. Crotch India: of 1932:524 in Fig. in as colour fitting Calvia of o the for Sikkim, Lectotype with was Delhi, Anisocalvia labelled stripe J. 1904; the marked Lectotypes or the 143, Fry's six of designated coloured two 323 / '39223 D. 330 Lectotypes (Figs vulnerata almost lectotype established specimen with punctata Collection Anisocalvia the 25. R.G. male pale borders. forms Lectotype [examined] Lectotype Uttar a Brendell; are median when does in m 23.ii. 1984 m 22.iii.1946, species. collection. Mungphu / feature 'Type description only The India of the 7-9, spots 156, usually genitalia Booth entirely buddha Buddha species, were not of Pradesh, viewed Calvia Natural of form as by number (Mulsant except In [circular, lobe 29) The Lectotype of Coccinella vulnerata Coccinella Orient by (arranged Kathmandu was nscli krishna Anisocalvia also conform buddha the darker all lutae by illustrated spotted, A Anisocalvia E. CIE Booth black flaecida M. Crotch. lectotype is is of with from male the of Cr. designated lectotype noted West 321 / '39221 F. History characteristic shown / G. the two the of ) nscli krishna Anisocalvia 11518, Vishnu red spotted and rings. elytra [Crotch's elytra ranging T. [examined], specimen Allen; below. to and spots 3-2-1) with Bhatkot, by Northern bordered Atkinson; British the with the with of in Museum vishnu, Pope Krishna feeding Specimens had of Crotch former a Cr Iablokoff-Khnzorian Kathmandu Calviaflaccida reduced. Fig. forms, by or above Only basic nscli vishnu Anisocalvia fitting hand]', [Crotch's and a Crotch. with (Fig. (1989) Gordon Embassy Kumaon 13. typical synonymized Museum A. India same the by forms do forms on pattern West (1874) from the C. pattern. Cr A a krishna the 29). oe pale Some with apparently aphids pattern uniramosa uniramosa is who [Crotch's specimen positions A and aes as labels Deyrolle arrange- District, descrip- hand[ Bengal, entirely (1987). hereby 4000 4500ft female in label] in of black with also The and not the his six on by of ft / Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 various feeding and Ranikhet, obversepunctata Material Anisocalvia and Union on 30.vi.1975 8.vi.1977 Champion; considered in but in is the is Comments 1.iii.1962 lar sipho Material Calvia Anamalai Universit~it, pink [Weise's unmarked c~c~, [printed] H. C. Comments currently a punctata, southern dorsal Thirty-seven The This This E. Five Weise J. printed is sides were valid Kashmir, Nilgiri and Andrewes' andrewesi: noticeably regia; card] (Iablokoff-Khnzorian, on dates above species species specimens. examined handwriting]'. examined CIE view / designated CIE of the species, andrewesi Kumaon (1908) CIE Hills, aphids elytra Andrewes by Uttar label] known Hills India, the Berlin). Otrore the H. / median A10431, Iablokoff-Khnzorian two (Fig. Srinagar 18235. A12741. Northern is is Korschefsky, median specimens. Anisocalvia G. S. collection. median more G. and Pradesh, also very / on based whereas separate lectotypes only Calvia India: Weise, 474 India H. Champion; 11). F. by 16.viii.1962 pear. Bequest lobe One Afghanistan: one recorded similar convex pred Hampson. G. lobe Gordon from [red Paralectotype Calvia 30.ix.1939 Pakistan: The his India lobe Lectotype / 1908: species of specimen Joshimath Champion; ~, C. from There Anamalais India: 1932:521 is ink] on the the description are male 1982). punctata Andrewesim B.M. to 'Cotype greatly in are andrewesi (Figs 220, Deyrolle Uttar from aphids (1987). male Nilgiri CIE Calvia in lateral / C. from are Lectotypes Gilgit Anamalai only genitalia A. Babur Lectotype (1979, the ? 1922 punctata, with 10, the 25.iii. two Pradesh, expanded [here genitalia ~, 18897, P. Uttar the is / [green on punctata University and (University view narrowly 11, Andrewes (Weise) Central 25.vi.1962 a of 'Anamalais 221 elytral Kapur; specimens Garden, 1982). Indian Ulmus; northern 1985 designated], Anaimalai 30 (Figs this here [Weise's Pradesh, Hills, (Fig. of on [printed] and bordered, for S. 32) (Fig. CIE Harmonia (Mulsant) Asian markings stat. designated There species Garhwal, scale subcontinent Jammu expanded 30-32) the of not Jammu 10) S. Bequest Kabul CIE of of species. A17174, 29). [printed] Cambridge rev. two India W. Cambridge); and handwriting]' Hills, this a / States on are 'Type 18503, Anisocalvia synonym circular on In also Almora and species. 5.ix.1983 and from punctata [examined] species more P. in Kumaon differences B.M. [printed] addition, (Fig. Kerala four with of [red size excelsa; clearly / Kashmir, Kashmir, feeding the Type Crotch 'Anamalais' strongly printed 30), and In 1922-221'. from bordered, Divn, specimens aphids; of Uttar / former and CIE 6500 C. Andrewesi Tamil the / show C. (Humboldt- [printed punctation, in Parachinar whereas Anamalais Andrewes' on andrewesi, collection andrewesi Kumaon Propylea Pradesh, Srinagar Srinagar both A15369, rounded ft latter label] Jammu psyllid that Nadu, Soviet H. circu- from Two with 929 the G. on as in m it / Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 be Wetherall; collection 930 label elytral is (1982) species Pradesh, when Anisocalvia Department Bengal, the H. and The Brendell. Everest excluded ft~r and syntypes Material examined Material Dalhousie Material examined Material Himalayan Phulcoki hrih West Bharmikh, n hemlock; on Champion. Calvia Calvia 1961-62 only an Twenty-two Nine This This Mader Naturknnde, Stevens; strength Shillong, this Natural is adequate alive sykesii: (Anisocalvia) breiti (Anisocalvia) markings single a remark by here and from and is a is Kurseong species has R. specimens. are from Chakrata 8800 in sykesii (1932) its Deccan, H. (Kapur, Pakistan: region L. The of West designated of very been not oshfk, 1932:529 Korschefsky, History rather Sikkim, Meghalaya, ft Coe; E. the the female lectotype about Agriculture, seems specimens. Natural 27-31.v. Humboldt-Universit~it, known. and Crotch, Sikkim described Tiruchchirapalli, characteristic Andrewes; examined Bengal, in original green Inde India Chautara Division India: oblong 1963). India. Murree was Museum the syntype to Lachung as Mader, designation. 1874: coloration History Verschraeghen by 1983 be G. type and referred Lectotype the Darjeeling syntype this India: Kapur shape Bhutan). Wadhi and rarely E. 7000ft Hills, Himachal District, 146, Lectotype here designated here 146, Lectotype Calvia lectotype. M. of Iablokoff-Khnzorian of are 1932: species Himalayan Calvia Bryant. 20.xii.1961, labelled Museum. this C. J. Sikkim, and to Tamil (1963) (Figs Thobba collected, fairly series R.G. remaining and D. vi.1928 6, andrewesi The (Fig. ~ sykesii species in 7000ft Nauling breiti from Manali coloration Dukhun Syntype [not examined] [not Syntype Brendell. Nepal: Parshad Berlin Pradesh, the The 1904; 14, (ICZN constant Nadu. as Booth whereabouts recorded 27) The ahn June Lachung species, H. a original (Crotch) from Weise's 26) Mader . G. H. feeding but second 25-31.iii.1924 paralectotype. after Roberts. (M. West Lekh, specimen being Taplejung The Sykes; Art. Bhutan: (1982). is (Fig. Kadrala, in Uhlig, personal it separated death. apparently Kulu, Champion; (1982) species female, description Bengal, maculation on 9500 in from 72(b) of Assam Nepal: 14). Berlin with Adelges the All Thimphu collection Himachal 1960, District, ft [examined] The recorded Darjeeling, was Bashahr (i)) and labelled 11-20.vi.1983 two Iablokoff-Khnzorian's the Weise's Darjeeling from W. Kathmandu species is not Himachal because feeding also as remaining 1-14.vi.1924 and specimens not on F. communication), Sangu restricted CIE all 'aber. in Pradesh, it recorded as Badgley; determination 9000ft spruce; here differ is from other the West a A17186 on it light green light 29.iv.1918 cotype, c. a'. has Museum Pradesh, taken M. District, original Adelges only seen Mount Bengal 6200 Indian H. to India. There Uttar West J. from Miss dark the (in D. G. to in in ft is Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 red-bordered TYPE specimens History East India East to of A18525 Verschraeghen Vesiculaphis Comments Calvia Material pronotum but suture, more times than slightly M-shaped Pronotum Comments with metatibiae but twice slightly sternite evenly basal sternites the the In This Thirteen Notes Bielawski Length Head Prosternal slightly uneven, metasternum; weak albida ore daet to adjacent coarse two the diameter elytra genitalia, region Anisocalv. punctured, one Museum darker dull and truncate, is entire. examined with on original he following another with and Company surface mark with 6.5-8.0mm, Bielawski, 1972:308 discal, coarser A19776. punctures to with had the printed sp. (1972) specimens. of process 1904; frons elytra brownish-yellow, punctation shining, of leave pair separation process. is seen on sixth sykesii medially; shining description, weak bloom Himalayan smallest, claws one Calvia hereby species, Uttar adjacent described very Museum, Museum bamboo; explanate. of no in convex, clearly separated sublateral; 5.5-6.5mm somewhat the suggestion apical Crotch'. doubt and side with apparent finely India: Sterna species Pradesh, ava championorum Calvia designated and of Bielawski's India underside all to ava albida Calvia apices margins. rather obsolete rather notched species. this Crotch label] Holotype this came to came with sculpture Manipur about spurs. ucue coarser punctures punctured, Upperparts eyes, (Figs mes- from Sikkim, very by Part Museum. (Figs pronotum species and irregular, at Mandal of of weak broad, c. / as higher its intervals 17) gave (1874) Abdomen Although and Duk=hun. of 33-36, the sparsely medially; paler mandibles n mouthparts, and original Elytral the male [not examined] [not male the 1.5-3 28, the similar correct the Doherty; from to nin subcontinent Indian Bielawski Gangtok metepisterna following British ucue separated punctures 14 A 41) only lectotype. dull magnification with longitudinal collections microsculpture. ih very with almost diameters, 42) single and punctation with description, Likhu and sp. to identity. it yellow female finely, weakly convex weakly alone weak 16.vi.1985 Col. is Museum no that laterally nov. West female fine finely similar species obsolete 9.ii.1985 indication It Khola on Sykes weakly darker to with sparsely o obsolete to of is and largest reticulate especially Bengal, stripes, frons, on brownish-yellow, labelled ( x ( the syntype punctured. in in and in are CIE Male fifth epimera Tal / particular disc carinae 80). 1879. India brown. general Ind. suggested CIE punctures by given apically; of and punctures dorsally, and A18126, one and Kurseong with moderately microsculpture. 'Type in 2-4 the reticulation Mus. metasternum, Museum, more Jiri, A16784, The paler below. restricted sixth adjacent appearance Meso- number fifth diameters, his [circular, browner A18353, intervals Natural Nepal. sides 96. / 79.64. at slightly figures or yellow visible visible some- most Inde 931 and fine the less of ex of to or to Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 leading slender 932 Etymology is Champion; Pradesh, Champion Pradesh, at vals metasternum ~, becoming species with ground distinctly from in coarser disc diameters, sternites carinae a punctured. and sublateral, of of sternite Type Comments C. C. 1-3 weak generally Uttar flaveola base small This PARATYPES HOLOTYPE Male This championorum. Head Prosternal Length pronotum the C. with diameters, somewhat three material C. are sclerotized colour. extending to surface truncate, laterally, of apex species Pradesh, species entire. championorum; either Nainital championorum, genitalia with W. fine notched obsolete well-developed intervals / apical the are readily broad Meso- 2 rather 5-7-6-6mm, G. Almora of usually ~, reticulate process frons 4 sutural given ~, In slightly bloom uneven C. is largest the and is ~, sixth intervals U. Uttar third anteriorly Division, C. named brighter (Figs more rods. and India: very Champion separable laterally Upper medially. with sixth finely Gumti albida, under slightly 14 Division, convex elytra and sometimes with metatibiae paler ih pair with punctures with microsculpture. Pradesh, Female even similar 33-36), the reticulate ~, visible 5-0-5.5mm for sclerotized Uttar punctured, body parts generally sublateral yellow Valley, Kumaon the the broad notch broad where to elytra a by browner. India: or father although basally, Notes explanate Calviaflaveola B.M. middle mixture prosternal latter (Figs genitalia to Kumaon, slightly tergite their pale median Pradesh, Gori form visible of with about surfaces W. stripes, of to ava albida Calvia R.G. 4 and 1927 shining, wings Sept. on area species. punctures 15, stripes weakly flatter the yellow genitalia. of ~, Almora broad, Valley, some of pair Pronotal of darker son, prosternum. medially; lobe at proportionally the (Fig. 37-40, twice Booth (Fig. former process 409. the male at same June, 1917 one various S. well are high apically, joined of base specimens without to sp. transverse G. separation slender Kumaon, only Division 42), Garhwal, more 15), apical adjacent diameter spots In separated H. G. H. developed, brownish-yellow, data 43) magnification C. August nov. punctures Bielawski also is of female with C. cornu sized is at and convex, weakly bursa. with shining. but Sterna in only reticulate against as albida, have spurs. Champion; apex. April scarcely microseulpture ventral . G. H. 7000 holotype; to of ucue sprtd by separated punctures 1917, of Kumaon, slender, slightly by well-developed weakly less a fine, externally, apex entirely suture, very convex Underside smallest, C. Male 1919 1-5-3 ft the elytral the weak the Elytral fifth Champion. ( x ( and H. microsculpture, view, differ championorum strongly separated sparsely with pale sperm 2 H. emarginated, sometimes 80). with G. diameters, narrower and 1 mottled ~, lacking one undated 6500fl dorsally, G. punctation 3, sipho in all Champion; yellow two but pale on sixth fifth 7 Champion. discal, duct punctation punctation ~, 1 prosternal punctures and ~, so by disc, pairs carinae, the swollen pattern yellow, H. visible visible H. short, elytra Uttar Uttar back- finely inter- 1-5-3 sides than than two and one not but but G. G. on of c. l Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 portion well-developed pair assoc, H. Champion; Murree walnut Joshimath walnut; on short the Etymology F. H. generally individual not Type characters. Washington), Acknowledgements separation in material, C. S. Comments and Museum the AGARWALA, References BELICEK, to A. E. thank PARATYPES arspoila pseudoartemisiae Macrosiphoniella HOLOTYPE albida. Male This The the Roberts; I E. colour Department always of M. am material with Venning. India. A review and bibliography, review and A India. n zoegahc eainhp btente fauna the between relationships zoogeographic ane Isca oepea Coccinellidae), Coleoptera: (Insecta: Slipiflski with Andrewes; curved aphids; University shorter than shorter Brancucci specific 1 Hills, J., species genitalia grateful respectively. my and to and 9, separate relatively Its of variation of 1976, 4.v.1985 The B. aphids, from 1 India 2 colleague the 9, all K. and K. M. dorsal ~, Thobba, size, 8 ~, and pair presence 1 is name Coccinellidae W. of (~, 1 three India: C. species. ~, to Uhlig CIE (Figs very (Naturhistorisches ~, 1 unsexed, it Entomology, CIE T. of Calvia 4 slightly A. of championorum. CIE large, in R. 1 rods. GHOSH, in 13 rm most from unsexed, Space is Huflejt 9, species. A12459; Cambridge wings, Foster G. Jammu similar C. Uttar undated D. 9~, 37-40), all A17293/Pres. (Museum from of Kashmir A17174, championorum, species J. Female weakly Gordon prosternal Uttar three narrower 18 A. of and du Pradesh, (Polska and apical the western Canada and Alaska with and analyses western Canada to, unsexed: . 1988, records Prey K., India: and 1 Heaume median The or examples facilities ~, from species Pradesh, but Latin G. spermatheca sclerotized for Valley assoc, Zoology vi.1888 The (USDA, Burma: Kashmir, ein with region Natural Tropical Museum, MacGavin body Uttar carinae Akademia generally Naturkunde, by Joshimath on Quaestiones the 4 lobe diminutive male for sipho during precludes with 20.vii.1983 Comm. ~, Artemisia of Indian Joshimath H. form Haka, Pradesh, Pest Museum the slender History area c/o will C. and Anantnag 6 Basel) Roberts; (Fig. aphids; swollen smaller for habitus this 9~, stout championorum Nauk, Management, United 3.iv.1984 and separate Inst. EntomoIogicae, subcontinent at base at Chin female flaveolus Humboldt-Universit~it, of h use the permission 43), vulgaris; 12 in study of CIE Museum, Kumaon, for piohgu Coccinellidae in aphidophagous 17.vi.1990 more and ventral Ent. at ventral unsexed, rts ouba n Alberta and Columbia British 1 than Instytut Hills, ~, 1 drawings. cornu States the vi. 1942 unsexed, base genitalia of A15632, it were N. the B.M. of shiny from (yellow) bursa loan C. Punjab, 34, 2 U. Debnath sclerotized of view, London. to slender, hs as these National ~, W. and kindly championorum Oxford 12, Zoologii, Pakistan: IIE 1-14. Burma . P. A. apical of examine 1985-1. all appearance copulatrix. provide pred N.W. 283-409. of Uttar Almora type and other type and narrow C. A21302, C. and Khyra the transmont- provided / sperm albida, Kapur, third, albida, I University aphids 1.viii.1910 good Sp. Museum, base also Province Pradesh, Warsaw) refers material Punjab, Berlin), reliable H. apical Gully b160, pred with duct wish and and 933 will key but but on G. on by to Downloaded By: [NEICON Consortium] At: 10:19 11 September 2007 WADm, WEISE, WaDm, STOLY~ROV, PAWAR, PnALOURA, SINGH, IABLOKOFF-KHNzORIAN, KORSCrmFSKV, KORSCHEFSKY, MIYATAKE, LINNAEUS, MULSANT, SASAJI, HOPE, MULSANT, MADER, MULSANT, BIELAWSKI, GORDON, IABLOKOFF-KHNzORIAN, KANERVO, GORDON, BOOTH, CROTCH, BIELAWSKI, KAPUR, GEBLER, 934 BIELAWSK1, Himalayas, program Bulletin English Manipur-l, The Indian Annates de Annates arthropod Himalaya, Ladakh Biologie of in Botanicae Fennieae Botanicae (Paris: Maison), (Paris: Janson), Faunistica, General (Coleoptera), Boubre), Nertchinsk, Nouveaux Mkmoires de Mkmoires Nouveaux Physiques (Coleoptera) Collections, CoccineUidae), hyoeia) luontaisista Chrysomelidae), Occasional Universitets F. J., K. H., S. A. . G. R. L., S. Basel the A. F., G. W., R. R. 1908, Coleopteren Coleopterists' R. C. . 1758, C., M., E., E., E., R., D. 1985, V., R. R., M. S. R., 1932, New P., 1832, D., . 1874, R., alder D., and and translation of 1831, Synopsis (J Hardwicke, R., und R., Coleoptera: and P. 1846, 95 ocnlia collected 1985, Coccinellidae for 311 1979, Ergebnisse 1853, and V., 1850, 568 and 1963, Beitr~ige 1963, et 1972, Entomology, 1946, & community 1963, 1985, 1968 la 18, 283-312. 18, Papers aiy Coccinellidae, Family 1987, A 1987, Bulletin York Entomological Society, Entomological York Entomon, Zur flrsskrift S. P~sH~, dans Oxford, StNaH, 1932, Fam. 1940, cotton Alnus nepalensis Alnus Naturettes, d'Agriculture Naturettes, Systematik Notice described SUGONJAEV, pp. PARRY, K), pp. SociOt~ Bulletin PARS~aD, ytm Naturae. Systema POPE, Histoire naturelle des colkoptkres de France. Sulcicolles.--Skcuripalpes. France. colkoptkres de des naturelle Histoire and Entomologische Bliitter Supplrment Calvia pce des Species (Coleoptera), Kenntnis Die 280 pp. 280 The revision A la The Tutkimuksia Indian Journal Indian Vier on S. Association S. in pests). Vanamo, SINOH, Sibrrie T. K., T. of catalogue sur in Entomotogica . 1989, M., Fam. Marienk~ifer (N.F. Linnkenne de Linnkenne M., . D., R. Coccinellidae ytmtc Entomology, Systematic Entomological Review, Entomological 10, of . 1982,Some B., Entomology, aus M., Coccinellidae by of . E. J. in species neue 23, la Coccinellidae, der les zur h rts Msu (aua Hsoy Entomology, History) (Natural Museum British the B., E. 291-295. der the iie Coccinelliden, einiger F. Socibtk Impkriale Socibtk northern 1979, Ostindien, Coccinellidae, 1], 1985, orientale, 5-8. T., 2), Colropt~res Kgifer, 1982, h of S. Gray, 12(3), colropt6res Coccinelliden W. Kenntnis 1980, D. Bhutan-Expedition new 1989, Entomologicheskoe Obozrenie, Entomologicheskoe Insecta Matsumurana Insecta of la vihollisista, of 59(4), and Record h coleopterous the from 1991, lep~in Editio Genera Hope, Don., of Aphidophagous No. the monographie Japan), species (Coleoptera: Lyon 36, Scandinavica, Les The Plant Protection, Plant (Coleoptera) Some 1 144-147. 21, UMagOV, 1-206. review A Afghanistan avec et of Crotch 26 Stettin by the new fiir Coccinellid 1-21. 1-3. Coleopterorum Catalogus, der decima Uttar Pradesh Journal Uttar and lehtikuoriaisen, Zoological Miscellany Zoological of d'Industrie, Lyon d'Industrie, 93, Coccinelles qui (N.S.), the of der the trimrres Entomologica Basiliensia, Entomologica Biologie und of 24 nin subcontinent Indian la description de la new records natural der Coccinelliden Check-list des nae Zooii oitts Zoologicae Societatis Zoologici Annales by 1-912. Hokkaido se Entomologische Zeitung, Entomologische 53(2), Nepaul pp. collection a~akice Coccinellini pal~iarktischen third S. , 1-46. 3, of E. (Holmiae), Alten records trouvent des Naturalistes family Entomologischer Anzeiger, Coccinellidae) A., 1, 20, 343-370. 20, of 1972 (the coccinellids Mulsant (Coleoptera) the of enemies srcuripalpes, 129-333. America Mount colroptrres (New 1-10] 1974, insects ColOoptOres-Coccinellidae 17, Coccinellidae Welt, Systematik des of type case University Coccinellidae of of 291-292. Melasoma at le darts Coleoptera von (2), Naturhistorischen [Dynamics eis, 30, series), Coccinellidae 824 in quelques of Coccinellidae Entomologische for material in de vrs expedition, Everest 53, north the of of (London), , 1-1104. 2, Afghanistan. important pp. the Moscou, aus fauna 245-257. [In 245-257. an trimrres Annales Zoology, North pars 120, pars district Hope der Kiifer, Expedition collection 4, from of integrated Nepal, of esp~ces nouvelles, aenea (London: of 69, 83-125. of 1-33. Mexico, soitd with associated Japan Entomological Eastern 2, a Sikkim (Coleoptera), Coccinellidae e mines des fruit pp. 213-230. srcuripalpes, des (Coleoptera. from 11, cotton 23-78. 225-659. 12, Bliitterffir Fragmenta 75, III, 14, L. Museums 21-32. of to Russian, 79-80. (Tokyo: Sciences pests 5-7. Journal control (Paris: 37-75. Major Nepal . W. E. Nepal India: 1-48. Lunds State, (Col., 1924 field de in