Vol. (~ No.2, June, 1992 79
The Delphacidae of Yukon Territory, Canada (Homoptera: Fulgoroidea)
Stephen W. Wilson Department of Biology, Central Missouri State University, Warrensburg, MO 64093
Abstract Twenty nine species of Delphacidae are recorded from the Yukon Territory, Canada, two additional ones from an adjacent region of Northwest Territories, and one species from coastal Alaska. Two new ienera are described: A8chedelphaxWilsonandYakorwdelphaxWilson. Sixnewspeciesaredescribed:AschedelphaxhJ:Jchne Wilson. DelphacodesanufrieviWilson, D. ernelianoviWilson, Javesella llaWilson, Not1wdelphaxglar:iaWilson, and Yukonodelphax kendalkre Wilson. Aschedelphax colorad.ensis (Beamer), Javesella kilmani (Van Duzee), Yukonodelphaxpediforma (Beamer~ and Y. stramineosa(Beamer) are new combinations. Kusnezoviella matisi Anufriev andEmeljanov isa juniorsynonym ofK. macleaniWilson andDelphacodes hyalinaBeameris a junior synonym ofNot1wdelp/w,x albomrinata (8t8.1). Ofthe 32 species included inthe study, 18 have a Holarctic distribution - 10 ofthese are amphi-Beringian. The remaining 14 species are restricted to the Nearetie, 5 of these are ~rded only from the Yukon and Northwest Territories.
Introduction
~ar northern environments comes from a number of OCCUITe In e ae c. erecen aun surveys aunal and reVISlonary studies. Numerous papers Of MOngOlia and far-eastem RUsSia illCLUded several have been published which include information on speciespreviouslythoughttobelimitedtotheNearetic thePalaearctiefauna(cat81oguedbyNast1972,1979, (Allufrie... andAverkin1982,Anufrie vandEmelj8nO" 1982). Recent summary wOIks on this fauna include 198~~liowingsurvey of the Yukon delphacid thoseforGreatBritain(LeQuesne19(0),Fennoscandia (Ossiannilsson 1978), Estonia (Vilbaste 1971), • 1] ma ;8. G~an .... annmgs, .. uppy aroeas mUSSla evan el]anov . and G. G. Eo SeudE1eP. Infonnation providedfor eash Forthe Nearetie fauna most records are in scattered species includes 1) reference to illustrations of the species descriptions (e. g., Beamer 1948, 1951), revi sions (Scudder 1968, 1964) and the lists of taxa for Quebec and Alberta (Moore 1944, 1950; Strickland specImens eXamlne, In1'onna on on 10 ogy, an ove~liew 1953). Scudder (1979a) noted that 81 specIes of 5} an ofthe species' distribution. New taxa delphacicls are known from Canada and 40 more are are described in detail. undescribedorunrecorded; he didnotprovide a listof Colleetingloealitiespr:'OVicIedinMap1 correspond toloca1i numbers 'venundertheYukondistribution Thedelphacidfaunaofthefarnorth(above &)"N) is virtually unknown. There are a few published records for the NorthwestTemtories(Scudder 1963) were mcorpora m t e y w ere POSSl e; ose and none frOm the Yukon Temtory. Wilson (1988) included in the summary for each species were 80 Insecta Mundi specimens that had identical collecting data as 8(7). Notal median 1000gitudinai pale stripe slighlJy associated males. lighter than rest of notum; aedeagus with elongateslenderapicalprocesses(Figs. 19 21) ...... Nothodelp1uJX tshaunicn Anufrlev Key to Species of Yukon Delphacidae 8'. Notalmedianlongitudinalpalestripemuehlighter thanrestofnotum; aedeaguswith small apical 1. Frons with 2 longttudinal median carinae ...... 2 teeth (e. g., Figs. 32 34) 9 I'. Frons with 1 longitudinal median carina ...... 5 9(8'). Forewmg of brachypter extending to or beyond 2(1). Largepitspresentonfrons. thorax.andabdomen; pygofer,posteriormarginoon.coloro'tls;aedeagus blackwithpartialdorsallongitudinalpalestripe WIth basall/4broadlyroundedondorsalaspect on head and thorax 3 (Fig 25) Notbodelpbax albocarinata (8tlil) 2'. Pitsabsent,lightbrown;posterim: halfofpronotum 9'. Forewmg of brachypter extendmg to fIfth and apicesofforewingscream, abdomenbrown abdominal tergite, posterior margin white; to blaek; male genitalia as in Figs. 1 8 ...... aedeagus with basal V4 sharply pointed on Criemorphus wilhelmi Anufriev and A-leFkin dorsal aspect 3'. Male pygofer with three median posteroventral 11(10'). fdldeagu!l with teeth extending almost one half teethorwith a posteroventralconcavity; styles length of aedeagus (Figs. 38 - 40) .. short, not extendingbeyond dorsal one-haltof ...... NothoGelp.w umbmta Emeljanov pygofer; female mesonotum and forewings 11' Aedeagus with teeth not extending beyond one brown 4 third length ofaedeagus (Figs. 43 40) ...... Nothodelp1uJX guentheri (Dlabola) 4(8). Male py gofer with three median posteroventzal teeth, genitalia as in Figs. 7 - 9A .. 12(6') Distinctpalestripeborderinglongitudinalmedian ...... Achorotile 8Ubm ctica Scudder cannae on notum 13 4'. Male pygofer with a median posteroventral 12' Pale stripe not present (yellow mesonotum may concavity; genitalia as in Figs. 10 12 .. haveblackmarkings lateralto lateralcannae), ...... Achorotile stylata Beamer bodywith brown or pale markings 16 5(1'). Head,includingeyes,muehnarrowerthanthol'ax 13(12). Maleanaltubewithspinescrossing(Figs. 48, 52) (indorsalview, distancebetweenlateraledgeof ...... 14 eyeandtegulasubequaltowidthofeye);pronotal 13'. Male anal tube with spines parallel or diverging lateral carinaestraight, extendingto posterior ...... 15 margin ofpronotum; male pygoferwith lateral fuf18.ted lobes (Figs. 13 - 16) .. 14(13). Frons blackwith yellow carinae; stylewith outer : Mega~lusfla~t8Cr=ord margin strongly concave in middle (Fig. 51); 5'. aedeaguswithnumerous small teeth ondorsal aspect on left side (Fig. 49) . laterally, not extending to posterior margin of ...... Ribautodelp1uJX albostriata (Fieber) pronotum;malepygoferwitboutlateralinftated 14'. Frons mottled brown with yellow ear..nae; style lobes 6 with outer margin broadly convex in middle (Fig. 50); aedeaguswithlargespineinmiddleof 6(5'). Metatibial spur without marginal teeth 7 dorsal aspect (Fig. 54) . 6'. Metatibial spur wiLlI marginal Leeth 12 ...... R~1X1UtodiilphaXpustlla Eirieljanov 7(6). DistincLpalestlipebortleting lougitudinalllledian 15(13'). Apex ofstyle acute (Fig. 68) .. carinae on notum 8 Chilodelphax magnifrons (Crawford) 7'. Stripenotpresent;notumpalewithdarkmarkings 15'. Apex of style WIth Irregular teeth (Flg. 61) ...... to solid black 10 ...... Unkanodes excisa (Melichar) J Vol. 6, No. 2, June, 1992 81 f\ () (l r -..... '" ~ --~ ~ r-l. ~ l:f: ( ~ ~ I ..' V ., . I 0\ ~\ U \ Q "" '" ~ ('J"--- 2 I:' \. - / ~\ ~ ~ A \\ If I Vi ./ ,- -- / ~~~/- \'\ // / l,. 1\/1 ~ /7 ,.., \. / / ./ Y,J'-,'( ..:> ). \f /- 0 / / II 0 ./ ./ ~ I "( I \ A n 1')1 Y' \ \ I ~ \ I ,r- ~ 1Ll. 10 / 11\ 11 1 \ / \ J(\ -<..~ • #--- /\~ J \ /..-- r - ...... (j _t-- I \ II \ ) \ 11 \ \ ~ ," ...... I -,,, ...... r"" A J .... / ,',- -...... r ~ " ~ '--", \~~J \ / t, -- \ V f / ...... --: °::°.'::-: '-.:.~ :'.: ::-Z·.: ---- " ./ " / ------12 "'" \' ------!Y ...... / 1'2 "" "- -- Figures 1-15. 1 • 3. Criomorphus wilhelmi. 4 •6. Achorotile aeuta: 7. 9. A subaretiai. 10· 12. A 8tjLata. 13· 15. Megamelii8 floous. 1,4, 10, 14·Analtube,lateralview.2, 5,8, 11,15.Aedeaglls, lateralview 3,6,9, 12·Styles,csudalview 7·Malegenitalia, lateralview. 130 Pygofer and anal tube, caudalVlew. Figure 4 • 6, 10 - 12 afterSCUdder (1963). 16(12') Male pygofer, in caudal View, as long or longer 17(16) FronspaleyeIlowtobIack;carinaeCOllCOIorous(at than wide, not appearing to flare outwards least anteriorly) 18 postenorly(e. g., FIgs. 17, 63); Ifstyles dIverge 17'. Frons mottled brown to black, cannae strongly then not appearingto layalongventral margin contrastingyellow orwhite 23 of pygofer (e. g., Figs. 66, 81) 11 16'. Male pygofer, in caudal view, wider than long, 18(17). Male mesonotum yellow with black markings ovaLe, appearing to flare ouLwards posteriorly lateral to lateral carinae 19 (e. g., Figs. 116, 118); styles stronglydivergent, 18'. Male mesonotum not as above 20 appearing tolay along venLralmarginofpygofer (e. g., Figs. 106, 115) 26 82 Insecta Mundi 19(18). Male pygofel with dorsolateral aspect strongly 29'. Aedeagus withblunt venual projection and deep producedcaudally(Figs.62, 63); analtubewith indentation on dorsal aspect (Fig. 114) .. spines AsChedelphax hoehne 'Nilson . Javesella simillima (LiImav tlOri) 19'. Malepygoferwithdorsolateralaspectnotproduced eaudally; anal tube without spines . 80(28'). Aedeagus with apex sharply angled ...... Delphacodes campestris (Van Duzee) anteroventrally, lacking numerous teeth (Fig. 119) JavB8ellii llii Wilson 2~18') Male pygofer with caudally directed median 30'. Aedeagus with apex broadly rounded, with proJection ventral to base ofstyles (Fig. 70) ... numerous teeth (FIgS. 122, 126) 31 ACXlnthodelphax analis (Crawford) 20'. Male pygofer Without median proJection ...... 21 31(30'). AedeagusWith teeth alongmostofventral aspect and teeth in basalonehalfofdorsalaspect(Fig 21(20'). Male head and notum black; styles each With 122) Javesella beringiaca Emeljanov dorsally projecting tooth near base on medial 31'. Aedeagus with teeth only in apical one half(Fig. aspect (Fig. 76) .. 126) Javesella kilmani (Van Duzee) ...... Detp1uJ.codes dentipennis Beamer 21'. Male with head and pronotum brown or black marked with yellow; styles each without dorsally projecting tooth 22 Criomorphus wilhelmi Anufrie." and A."erkin (Figures 1 - 3, Map 2) 22(21'). Apex ofstyle acute (Fig. 81) . .. Yukonodelphax kendallae Vlilson CnomorplUis bOrealts(S8.hIberg), Ariufriev1972:613, 22'. Apex ofstyle broadly rounded (Fig. 87) . 1977:864 neeSahlberg ...... Delp1ureedes amt{rievi Wilson Criomorphus wilhelmi Anufriev and Averkin 23(17'). styles '/lith apices OOll'lerging (Fig. 91) . 1982a:181. .. Kusnemviella macleani Wilson 23' stYles with apices dP~ng (Figs. 94, 98, 103) • DistribUtiOn reooi'CIs fOr the specimens used iii ...... 24 this study are: YUKON TERRITORY: 28, 54, 55; 3 - 20July;ALASKA: Gobbler'sKnob,66°45'N 15O"40W; 24(23') 9 male braclIYpl:ers, 1 male macropl:er, 6 female brachypters. Otherrecordsarefrom Anumev(1972), Anufriev and Averkin (1982), and Wilson (1988). 24'. 25(24'). Aedeagus curved dorsally (Fig. 96) . mSTRlBUTION NEARCTIC: Canada: Yukon ...... Paradelp1uJ.codes titoratis (Reuter) Territory; USA. Alaska, PALAEARCTIC: Mongolia, 25'. Aedeagus curved ventrally (Fig. 101) . Russia: Amur Region, Kamchatka Region, ...... DelpllllWl1es emeliarwvi Wilson Khabarovsk Territory, Kuril Islands, Magadan Region, Primor'yeTerritory, SakhalinRegion, Yakut 26(16'). Aedeagus strongly recurved (Fig. 105) .. AutOnomous Republic...... Javesella pellucioo (Fabricius) 26'. Aedeagus decuned or stIaight (e. g., Figs. 108, 111) 27 Achorotife scuta Scudder (Figures 4 6, Map 2) 27(26'). Aedeagus forked (Fig. 108) ...... JavB8ella obscurella (Boheman) Achrotde (sic) acuta SCudder 1963:167. 27'. Aedeagus not forked 28 28(27'). Aedeagus much broader in basal one third than Distribution records for specimens used in this near apex (F'Igs. 111, 114) 29 stud.yare:'TOKONTERRITORY~26,80,48, 50, 51; 10 28' Aedeagus approximately equal in width males, 21 females, all brachypters; 1 .June - 20 .July throughout most of length (FIgs. 119, 122) .... Other recordS are from Scudder (1963). 30 DISTRIBUTION· NEARCTIC: Canada: Alberta, 29(28). Quebec, Saskatchewan, Yukon Terrilmy...... Javesella discolor (Baheman) Vol. 6, No. 2, June, 1992 83 / ~ "\ n / \ -I ... / --- ~ .LV / \ V , /J /J \(' ( J .( '\ \ V \\ ~, /~ ~, lY 21 \"V V -,-- I ...... ~~ :{~-- I ')() ------..., "'~ -..., '7___ -. .. J / ~ ./ /" ./ J ~ ~ 28~ ...... ::,:..... ~ ------: ....- ..- /~ ~..., ---- Ll - c...::. .. .->: c=-. -. - ./ /' - ~ -- /1 ...... ~ / "'"\\ \L.L.J J ~ ?h // // I \. ./ \\ \( I \~~{ ( / , I '\. J " I L.';) / V / ,.,.,.,. 1"')0 - ~ ""'- ~JU \ - /- ..::J£. - " / / ~~./~ .... , ,.,.,.,. ~,~ c.-~~:~·_?) .;:)..::J c~~-:::'.../ r--.... f \ - J7 \ , " -I """ \ ) / v.L ~ 34 ~~ 40 \ '0 ....- - ...... "- / /" " \ ~ A1 / \ \" / / / ) I // --- ~ 1" / I "'" ~ "'" / ( / / f /,," 37 c-c- / I ./ ....1 '- / / \..~ ~ .jo L / Figures 16·41. 16 22. M:>tJwdelpJw.x tshatmioo. 23 26. N. alOOmMata 27 29. N. eW~ 00 37. No gkwia. 38 40. N. umbmta.41. N.guenth21i. 16, 00, 41- pYgofer, Iateralview.17-Pygofer,caudalview.18,23,27,21-Analtube, lateralview. 19, 21, 24, 25, 28, 32, 34, 35, 36, 38, 40 -Aedeagus, lateral'..jew 20, 33, 39 -Aedeagus, ventral·Jiew. 22, 26 Styles, GaUdal·Jiew. 29, 37 - Right style, caudalview. Figures 23 - 26 after Anufriev and Emeljanov (1988). 84 Insecta Mundi Achorotite subarctica Swdder 51 ,r-,_ .... ,...... ,..~ Map 1 I' • '" Iy,g"" '"' :~<~\ -_.... lor useain uris :~ ~)/~r~~~~::;::::::::,::::=_==:;'H;:::====== , v .....~... Territory; USA: Alaska; PALAEARCTIC: Mongolia; 0.' n~' ~, • ..... , '-'&&'&'-'-, AutonomousDistrict Khabal'OvskTenitorv.Tavmvr , omous nepuollc. Achoroti/e sty/ata Beamer \r _Q IV - I~, Map ~J 15 14 13 12 11 10 9 8 7 6 5 4 3 2 1 Achorotile stylata Beamer 1954:147. Map1. Yukm 'Ifuiitmy 001 kYiting localities 1. Rosa RiVer, 2.Quiet Distribution records for specimens used in this Lake, 3 - Takhini Hot Springs, 4 - Atlin Rd, 15mi. S. AIaska Hwy, Ii. 15 5 lUi. S Jake'sComer, 6 Hwy7 B C. atYukmBonIer, 7 Tagisb, studyare: YUKONTERRITORY: 4, 5, 22,64;5males, 6 females, all brachypters; 8 June - 31 July. Other records are from Beamer (1954) and Scudder (1963) who recorded this species fmm Pnn. proum.siR I. Hwy., mi Iffi4, Kluane1..., 3)- Cmmacks, 18km S, 21- Koidem and mSTRIBUTlON NEARCl'IC: Canada: British AlBBkaHwy.,kml881, WbiteRiver,22-LewesLa1re,23-TatclnmCr., 62"17'N 13&17W, 2( • ven Wilczek L, 62"44N 13&42'W, 25 • Pelly Columbia, Yukon Territory, USA. Wyoming. crossing. 26 - MtQJesten, 10kin E, 71 • MtQJesten 33kin NW, 28 - Klcndike R. Dempster Comer, 1km W, 29 • Dawson Aillxrt, 30 • Boondmy77kmE and2DkmE, 31-Daw_,10kmE,37kmE,and KIn 690, 32-Dawson,33-BensenCro88ing, MoUN138" 33'W, 34-N. KlendikeRiver,Km42,DempsterHwy.,35-N.KlondikeRiver,Km64, Highway, 37 • N. F ~ ~ ...... t' /' " "- :.L .... I( \ \ "- ... :-:--.,. ~ ':US ~ ~ ~ ~- ~ ~~A 44 , t::,.,\ --- ~..., <-- - .J \ I , ... / ~~~ ... 4~/-----" , "'- / /~ ... AI::; ~ ~.~ ...... / !..L/ ...... ~ ...- ~ t::A ~ '" 1"\ , / ....----. ./-:1:: /' ...... \I--_ ,JV ( ..;/ / ... -- ..... "" /I '" .;:-- ...... [ // <"\. /"'\ ~ /J ..... / .....\ \"" I' I" ~ I \J \// 1::;1l )( I \\ j ) v 46 \Y /47 55 ...... -.Jl .-/ ~ ~ ~ r-<~ 0';1 - ~~ / (' / \ \ /n ~ / / nl J ~ )J ';)1 I ~- -L'i I / ~., 561 r.,.. r /I / '"' .f " r / I ~ ly\ 1q3 / ....., "- /' ".... / / \ ,( ) \\/ / / ~? / r-n \ '\../ / ~1 ( /' ~~ .... ., ...... " / ~ -I Figure42- 62; 42· 47. Not1lotklphaxgu.entheri, 48· 61. Ribautodelphaxalbostriatao 62 . 66. Ro pusitla. 66· 68. Chi1l>delpbo.x magnifrom. 59 . 61. Unkanodes excisa. 62. Aachedelphax 1ux:1uJe. 42 . Pygofer, caudalview. 43, 45, 49, 50, 63, 54, 67, 59, 60· Aedeagus.lateral view. 44· Aedeagus. ventral view. 46· Anal tube. lateral view. 47. 61, 66, 68; 61· Styles, caudal view. Nothodefphax tshaunics Anufriev rnSTRlBUTION NEARCllC: Canada: Yukon (Figures 16 - 22, Map 3) TeIrit. 7yrphOdelp1UiX tshiiunwus Ariufriev 1979:297. Nothodelphax albocarinata (Stal) .\Tothodelphax t&hcmnioa (Anumev), ..A.nufriev and (Figures 23 26, Map 3) Averkin 1982:137. '!hedistributionrecordforthe specimenusedinthis DelplUIi albOCCirinatus St811858:357. study is: YUKON TERRITORY: 55; 1male brach.ypter; Liburnia albocarinata (St8l), J. Sahlberg 1871.426. ~)July.OtherrecordsarefromAnufrievandEmeIjanov Delphaoodes albocarinata (St8l), China 1938:197. (1988). 86 Insecta Mundi Kaszab, z. 1958. DIe MelOJden Afghamstsns Kirby, w. 1818b. MelLttophagus. In W. Kirby and (Coleoptera). Acta Zool. Acad. Sci. Hung. 3: W. Spence, An introduction to entomology: or 245-312. elements ofthe natural history ofinsects: with plates. Vol. 1. Ed 3, p. 164 (footnote). Longman, Kaszab, Z. 1959. Phylogenetische Beziehungen des Hurst, Rees, Orme, & Brown, London. fljjgelgeaders der Meloiden (Coleoptera), nebst Bescbreibnng neuer ('T8ttnngen Hnd Arten Kirsch, T. 1866 Beittiige zHr Kaferfauna von Acta Zoot. Acad. SCI. Hung. 5: 67-114. Bogota. (Zweltes Stuck). Berhn. Entomol. Z. 10: 173216. Kaszab, Z. 1960. Die Arten und die systematische StellWlg der Meloidell-Gattung Calydus Reitter Kiug, F. 1825. Entomologiae Brasilianae specimen (Coleoptera). Acta Zoot Acad. Sci. Hung. 6: alterum, sistens Insectorum Coleopterorum 125-134 nondum descriptonlrn centuriam Nova Acta Leopoldina [Verb. Kaiser}. Leopold. Carolin. :Kaszab, Z. 1962. Uber das System der asiatischen Akad. NatuTf.] 12(2): 421-476, pis. 40-44. Lytta Arten, nehst Besehreibung drei neuer Arten (Coleoptera, Meloidae). Ann. Hist.-Natur. Kolbe, H. J. 1894. Coleopteren aus Africa. Stet Mus. Nat. Hung., Zoot 54. 289-298. tiner Entomot Zeitung 55. 178-190. Kaszab, z. 1964. Ergebmsse der Mongohsch-Deut Kolbe, H. J. 1897. Kiifer und Netzfluger Ost-Mn schen biologischen Expedition 1962. nr. 3. kas. In K Mobius, Die Thienvelt Ost Afrikas Meloidae (Coleoptera). Folia Entomol. Hung. und der Nachbargebiete. Wirbellose Thiere. In (n.s.)17. 317-324. Deutsche-Ost-Mtika. Vol. 4, pp. 1-367, pIs. 1-4. Dietrich Reimer, Berlin. Kaszab, Z. 1966a Meloetyphllls fllscatllS horni ssp nov. sowie die systematische Stellung der Gat- Kraatz, G. 1863. Revision der Cerocomiden-Grup- tung Meloetyphlus Waterhouse (Coleoptera: pe. Berlin. EntomoL Z. 7: 109-115, pI. 4., figs. Meloidae). Beitt'. Enoomol. 16: 183 188. 8-10. Kaszab, z. 1966b. Revision der Triben Ertliini und Kraatz, G. 1882. In L. von Heyden and G. Kraatz, Derideini (Coleoptera, Meloidae). Ann. Hist. Kafer urn Samarkand, gesammelt von Haber Natur Mus Nat Hung, Zoot 58' 343-360. hauer. Deut. Entomol. Z. 26: 334 335. Kaszab, Z. 1969. The system of the Meloidae Kuzin, B. S. 1953. The blister beetles of Kazakh (Coleoptera). Mem. Soc. Entomol. Ital. 48. 241- stan. [In Russian.] Tr. Respubl. Stantsyi Zas- 248. chity Rast., Kazakhstansky Filial Vashnil (Alma-Ata) I' 72-154 Kaszab, Z. 1983. Insects of Saudi Arabia. Coleop- tera: Faro. Meloidae. i .... synopsis of the Arabian KuziD, B. S. 1964. Contribution to the knowledge Meloidae. Fauna Saudi Arabia 5. 144-204 of the systematics of Coleoptera, Meloidae, Mylabrini. [In Russian.] Tr. Vses. Entomol. Kirby, W. 1802 Monograpbia Apllrn AngHae; or, Obshcbest 44' 336-377 an attempt to divide into the natural genera and fimiilies such species ofthe Linnean genus LacOI'daire, J. T. 1859. Histoire naturelle des Apis as have been discovered in England: with insectes. Genera des coIeopteres ou expose descriptions and observations. Vol. 2. J. Raw, methodique et critique de tous les genres Ipswich, 388 pp., pIs. 15-18. proposes jusqu'ici dans cet ordre d'Insectes. Vol. 5. Librarie Encyclopedique de Roret, Paris, Kirby, W. 1818a A century Of insects, including 75/1 pp several new genera described from his cabinet. Trans. Linn. Soc. London 12: 375 454, pis. 21 22. Vol. 6, No.2, June, 1992 87 A /\ r ~ ~ f'\ oJ '>---- /' ...... "- ~ '\ (\ ~h7 ~~ ~ /w ~ I 1"71 L (\ "" l j ( ~ /"-- \ " ----v \ / -I ~ v '-- '---" /// \. \ r / A OLf 00 If) ~ UJ \ "'---/'"/ <.~ \r- - / (\ I ... '" "- ./ v "- / \ -- I n [I r-... /' 7 \ "\/' ( / ...., r / \/ ( \ / 7(1 "- / - ~ "- ...... IJ ,;~~ / //\ \\/1 ~ \/ nn A v ~ \..-/ '--'" I, Figures 63 • 8L 63 . 66. Aschedelphax hochae. 67 . 69. DelpJuuxx1es mmpestris. 70 . 73. Acanthodelphax analis. 74 . 76. D. dentipermis.77 81. YukmvxJelphaxkeruialku!. 63 Pygefer, eaudah"iew. 64 Analtube, oonnective, andaedeagtlS, 1ateJ:alview. 66, 67, 68, 72, 715, 79, 80 Aedeagus, laieralview. 66, 69, 78, 76, 81 Styles, caudalview. 71, 74, 78 Anal tube, lateral riew. 70, 77· Pygofer, lateral view. (Figures 67 . 00 after Muir and Giffard (1924),74·76 after Beamer(1948). Distribution records for specimens usedin this Nothodelphax guentheri (Dlabola) study are' YUKONTERRITORY: 12, 15, 16,21; 22 (Figures 41- 47, Map 3) male brachypters; 31 May· 20 June. Otherrecords are from Anufrie-.. and Emeljano... (1988). Koswigianella (sic) gilntheri Dlabala 1966:444. Nothodelphax guentheri (Dlabola) Emeljanov 1982.90. DISTRIBUTION· NEARCTIC: Canada: Yukon Territory; PALAEARCTIC: Mongolia, Russia: The distribution record for the speClDlen used Kamchatka Region, Magadan Region, Yakut in this study is: YUKON TERRITORY: 52; 1 male Autonomous Republic. brachypter, 29 June. Other records are from Anufriev and Emeljanov (1988). 88 Insecta Mundi DISTRIBUTION - NEARePIC. Canada. Yukon Chilodelphax magnifions (Crawford), VInson 1988. Territory;PALAEARCTIC:Mongolia,Russia:Altay 337. TelTltory, Taymyr Autonomous District. Distributionrecordsfarspecimensusedinthisstudy are: YUKON IERRlTORY: 5; 14 male brachypters, 12 Riballtodelpbax albostriata (Fieber) =e~~~==~4JUne-27July. (Figures 48 - 51, Map 4) DISTRIBUTION • NEARGrIC: Canada: British Delpha:Je albostriata Fieber 1866:525. ColWlibia, Northwest TerritOry, Yukon T~l'ritory; Libumia albostriata (Fieber), Fieber 1872:5. USA: Alaska, Colorado, Montana, Wyoming. Delphae0de8 albostriata (Fieber), Muir and Giffard 1924:25. Unkanodes excisa (Melichar) Rib:=~.hax albostl'iatus (Fieber), Wagner (Figures 59 - 61) Distributionreoo:rdsfm specimenswoodinthisstudy Libumia excisa Melichar 1898:67 are:YUKONTERRlTORY:27,33,39;7malebrachypte:rs; Elymodelphax excisa (Melichar); Wagner 1963:167. ::wilstd~~reeaIds are RemVllbaste (1971) Unkanodes (- Elymodelphu;i), Diahala 1965.86. This species has been reported from coastal DISTRIBUTION - NEARCTIC: Canada: Yukon western Alaska ~ilson 1988¥and may3lCCur alon: Territory; V5;.-\: Alaska; PALAEARCTIC: Austria, the north coast 0 he 'Yukonerritory~Distributio Belgium, Cypros, Czechoslovakia, DeIlD18lk, Fstonia, records are from Anumevand Emeljanov(1988) and FiDl8Ud.,F.l'anre,E8StGermany,WestGermany,Hungary, Wilson (1988). Italy,Mongolia,Netherlands,Nmway,Poland,Portugal, Rom8iii8, Spain, SWeden, SWitzerland, Tunisia,Russia, DISTRIBUTION - NEARCTIC. USA. Alaska, Yugoslavia. PALAEARCTIC: Denmark; Finland; Germany; Poland; Russia: Kuril Islands, Yakut Autonomous Republic; Sweden. Ribautodelphax pusilla Emeljanov (Figures 52 - 55, Map 4) 62"- Eurysa magni{rons (Crawford), Muir and Giffard Vol. 6, No.2, June, 1992 89 Q') '\ "":..~. "::"""'" R~ (\ I ...... - ./ /' ------'-=VV-tz:lt;.'I£,/ ~\ J } l/J \ / -v;.....~ ~ ~ ~ I ~~, I / ~v v '\ { ...... L ,V 07 /~ 0:/"'-.---- ~ .., ~U I ~'l ---J :J - / / OJ J I/ /1 '" .----... "...... / / " /\ J? ) ( ( C)1 "") ') ./ ./ ..... / \ / \\ 87 / J V ) \( I ~ f'.--J) (\ / uu )\ ) \\/1 \ ,- \ \I I """1 r------.I /r./ \ ~ r-...... , 1 G - ~ ~ ~5 V \ ~ (" I .... 1"\1"\ ...... " '\ III \ f\) L--..J ~ \ I / ~ -" ':J'L. J"< ~ ~ /1... ()1"":'\... '- / ...., '- " " '-J --- ~b ~. 1"\'" ,,-;;-- .-- II V ...... c - 07 \ /./ ...... '.' .". . . ./ .... 'I' ~ --\f ~ ... / \./ ~v.. ~ ( / ') \\ / / /00 \\ .... I"\~ r/ "'-v J \ \ ";10 I / \\ j) "- J 0Ll. \ ''y'l I \V/" -' ~ \...... Figures 82 -103. 82 . 87. Delpha00de8 aJ11.l[rievi. 88 - 91. Kusnezoviella macleani. 92 . 94. D. deans 95- 98 Pamdelphomdes litmulis. 99-108.D.emeljanovi.82,99-Pygofer,lateralview.88,85,90,98,96,97, 101, 102-Aedeagus,lateralVlew.84-Aedeagus, ventralview. 86-Analtubeandinnerviewofpro Aschedelpl7ax Wi1501 I, I lew gelIUS DESCRIPTION.Vertexslightly longerthanwide, lateral margins almost parallel; lateral, median, and oblique carinae weak, with a concavity between eaah oblique Type species: Aschedelphax hochae WIlson carma and posf.enor margm. Frons WIth lateral margins carinate, almost straight and parallel, slightlyconvergent 90 Insecta Mundi point offorking on juncture with vertex. Antennal seape MALE GENITALIA: PygofeI obliquelysubcyHndrieal, in length subequal to width at apex; pedicel ca. 2 X length of lateral view, posterodorsal area strongly produced on each scape Beiikextendingtomesotl"OChanters Pi'OnotallateraI side, height ca 2X width; in caudal view, strongly prOOueed carinae slightly posterolaterallycurved, extending across posterodorsalareacurvedinwards,bilobed;ventralmarginof &'4 of prulwtUnI before beooming obsolete. Mesonotal diaphragmopeningslightlyeonea....e; diaphragmannature stronglyproduced andsubtriangularin lateralview. Anal tUbe subcjrIiiidrlCal; two elOngate, parallel acute spiIies transverse row of 7 teeth (5 + 2) on plantar surface. Pygofer obliquely subcytindrical, posterodorsal area stronglyproducedoneach side; posterodorsal areacurved in-...-aros, bilobed; ';.-entroeaudal margin broadly ooneave. Diaphragm armature subtriangular in lateral view. Anal This speCIes IS named In recognItion of Dr. tubewithpairofelongate,slenderventrooaudally direeted Hannalore Hoeh's research on planthopper acutespines.StylesCliverging. Aedeagusdirectedcaudally, systematics. strai(Jlit. Distribution records fOI the specimens used in This genus includes two species, A. coloradensis this study are: YUKON TERRITORY: 26, 29, 31, 32, (Beamer), newcombinationand. A. hocJw.e\Vilson, 33, 46; 7 male brachypters; 6 June - 16July. new species. The genus name is in honor of Dr. MallfcedAsche'ssigllmcantwnLribuLionsludeiphacid DISTRIBUTION . NEARCTIC: Canada: YukOn systematics. Territory. Wilson, new species ASChedelj)hiiX l70ChBe Delphacodes campestris (Van Duzee) (Fig!Ires 62 - 66, Map 5) (Figures 67 - 69, Map 5) HOLOTYPE. male brachypter with label: "YT. Dawson; 10kmE;16vii1983;G.G.E. Scudder"inthe L~bumza oampestns Van Duzee 1894:191. CanadianNationalCollection, Ottawa;PARA.TYPES: Delphaoodesoampestris(VanDuzee),MWrandGift'ard sixmale brachypterswiththefollowing data' Yukon, 1924:35. Dawson 37 kDi E, 10 Krii E, 1980, R: J. Cannings; Benson Cr., 64°11'N 138°33'W, 12July 1983, G. G. E. '5 6 Scudder; Mile 150, DempsterHwy., 22June 1980, R. urneo. • Nothodelphax guenth~ri l , lateralto lateralcarinae.Mesonotum ale , forewing hyaline, costal veinwhite. Legs yellow, apices of terminal tarsomeres black. Vol. 6, No. 2, June, 1992 91 ~ J /J ... ",..... /1 ---'- ./ / II 1/ / ~~ ~N ~ ../' Iv / r .~~.J:.. ,.- '\ ./ ...... ~Lt~\ C /'\"" / \ \ LjY "(~, ~~~ ~ ...... ( r , , 10J:\ IVO '-... J ~ .-----:::::=> -- i) -----,,-? 7 /~ / - ../ 1(\~ ~ ~ ...., ~~ J 112 ...... " .... '"''"' -- ./ 11'-1 -'/ .... 'J-' / ~~r--~ .// ...... ", ...... / ./ II - \ / "- ...... ---- \\ ------~ y~ 113 / ~ ~ ... ~ \ } I \ \ ~ "1 "11"'7 / '\...-J Ll/ ~ 114 / ~ .A..A.V .:::..... "'\/ / I ./'" -" ...----...... / ./ //r~~~" ~ "1"1r:" f? \ ~ J.J.J~) ---"/ \ / \ ...... / '-- " / '\ )( ( ---116 I \/ -/ /\.. \ " / \ , I / "- / -c . "'- / .... ,. " ...... ~ -. / 110 -- V .A. .... ( ~ 11"\1"'"., ----- I r-- .. \ ...... -~ .- \'.::." ". 120 \) '" ...... ---- - lL~ - Figuresl04-127.104-100.Javesellapellucida. 107 -100.J. obsatrello. 110-112.J. disrolor. 113-1l6.J. simiUima. 116-121. J. 114 122 124. J. beringiam 125 127. J. kilmMi. 104, 107, 113, 117, 125 Anal tube, lateral'liew. 105, 108, 111, 114, 119, 120, 122, 123, 126 -.AEldeagus, lateralVIew. 106, 116, 121, 124, 127·Styles, caudal. VIew. 100, 112·Right style, caudal. view. 110 .Malegenita1ia,Jateralview 116-~fer,lateralview118-Pygofer,C8ndalview Fignres122-124afterAuuroevaudEmeljaDDv (1988), 125 • 127 afterDuBose (1900). 92 Insecta Mundi DisLribution records for the specimens used in Yukonodelphax Wilson, new genus thisstudyare: YUKONTERRITORY: 15, 19; 15maJe brachypters, 8 females, 30 May • 21 JUly. Other TYPE SPECIES: Yukonodelphax kendallaeWilson records arefromDu&se(1960), Omet al. (1985) and the S.W. Wilson insect collection. DESCRIPTION: Vertex slightly longer than wide to subequal in length and width, lateral mfll'gins almost DISTRmUTION· NEARCTIC: Canada: Alberta, parallel; lateral, median, and oblique carinae weak, with Yukon Territory; USA: Connecticut, Illinois, Iowa, a concavity between each oblique carina and posterior Kentucky, Michigan, Missouri, Minnesota, margin Frons with lateral margins carinate, convex, MiSSlSSlppl, NebraSka,Nevada, NewHampShire,New convergmg posteriorly; median canna distmct, almost York, North Carolina, North Dakota, Oklahoma, obsolete atpointofforkingonjuncturewithvertex Antennal Pennsylvania, South Dakota, Tennessee, Virginia. scape length subequal to WIdth at apex; pedicel 2 • 3 X length of scape Beak extending to metacoxae Pronotal carinae poorly defIned; lateral carInae curving posterolaterally ft>lIOWing curvature of eye, becoming Acanthode/phax ana/is (Crawford) obsolete between eye and posterior margin ofpronotum. (Figures 70 - 73, Map 5) Mesonotal carinae weak; lateral carinae diverging and extending to posterior margin. Metatibia with apical transverserowof6 teethonplantarsurface:spurfoliaceouS; MegamelUS anal7-8 CraWford 1914:620. with 10 . 20 smallmarginal teeth, apical tooth verysmall. Delphaeocks analis (Crawford), Muir and GiffMd Metatarsomere 1 with apical transverse row of 7 teeth (5 1924:24. + 2) on plantar surface. Acanthodelphaxana-lis (Crawford),Wilson1988:888. Pygof'et subeylindrical,ventroeaudalmmginexcavated. Ornamenteddiaphragmarmaturepresent.Analtubewith acute Distribution records for SpeCImens used In this large pair of widely separated, elongate, curved, spines.Stylesdiverging, apicesblunttosubaeute.Aedeagus study ue: YUKON TERRITORY: 35, 36, 37, 39; 10 liUile braehypters, 7 female brachypters, 2 female directed caudally, bearing teeth ancVor spines. Y. maeropters, 9 June 12July. Otherrecords arefrom This genus includes three species: kendallae Wilson (1988). Wilson, new species, Y. pediforma (Beamer), new combination, and y: .9tromineosa (Beamer), new combination. nTSTRmTmON . NEARCl'IC CanlLa' d Albel"tra, YukOn TelTitOij; USA: Alaska, Imnois, MiChigan, 134' 128' 122' i • •• , ....T. • v. __1 , ....T ... - , 99•• . I M '(uRonodelp17ax Renda/me Wilson, MALEGENITALIA.Pygofe:t subcylimhical,inlateral new species view, height ca 1.5 X width;ventral mar~ofdi;;hra= openingwith largethumb like diaphra~rma0e,a (Figures 77 - 81, Map 6) ofarmaturegranular.Analtubesubcylindrical,twoelongate spines extending from dorsooaudal margin posteriorly thencurvingventrallynearmiddle. Styles divergingfrom base, curved inwards near apiGes sum that aeute apiees SeuddCr", in the Canadian National Colleetion, recul'Y ; roa at ase, a rupt y narrowmg an curvmg Ottawa; PARATYPES: five male brachypters, with ventroposteriorly in basall,l3; 7 slender spines scattel'Eld the following data. YUkOll, Slims Ro Delta, 21 June ondorsalaspect, a slightlyobliquerowof6smaIlteetli(two 1982, R. D. Wilkie & S. G. Cannings (1 male); Alaska illustrated) on ventrolateral aspect on left side. Hwy. nu. 1054 Kluane Lo, 30 May 1979, G. G. E. This species is named for my daughter, Kendall. Scudder(3males); KluaneN P,SlimsR :flats; 2June Distribution records for the specimens used in 1979; G. G. E. Scudder (1 male); inthe University of this study are: YUKO~ TERRITORY: 14, 15, 19; 6 British. Columbia and S. J,Jf. Wilson eolleetions. male brachypLers, 30 May - 12 June. DESCRIPI'ION. Vertex pale yellow. Frons brown DISTRIBUTION . NEARCTIC: Canada: 'Yukon mottledwithyellowbasally,becomingpaleyellowapically; TeIT.itory carinae paleyellow. Clypeusbrownwith palemedian and lateral carinae. Pronotum pale yellow, brown behind eye lateral to Delphacodes anufnevi Wilson, new speCIes lateJ: alcarinae. Mesonotum brown to dullblack, posterior (Figures 82 - 87) margin yellow. Brachypterous forewing hyaline, costal vein pale ,ellow. Coxae brown to black, margined with yellow, remainder of legs yellow, tarsomeres brown to HOLOTYPE: male brachypter with label: ·Yukon, blackapically.AbdomenbIe \Vllteblaek, pygefermargined AlaskaHwy.;km 1671; 1.VI-1979G, G G Scudder", with yellow apically. m the Canadian National Collection, Ottawa. Abdomen bllWkish, marked with. pale beige. DESCRIPTION:Bodypaleyellow.Lightbrowninfusions on genae, elypeus and on legs. FIontal lateral car inae, slightly convex; median carina forking at juncture with vertex. Braehypterous foIewing hyaline, short, not extendingtopygofer, light brown. Metatibialspurwith 10 ex WI su on oraa as longitudinal axis, nearly straight, club-shaped; gonopore in· si e' in a base,broadlyrounded,withrowof4 visiblestrongteethon n en ra aspect; in right lateral view, with 1 toothondorsal aspect, , .. . apical 1/3 (not illustrated' in ventral view with 1 and a row of 13 visible teeth on 94 Insecta Mundi tspecies (Asche 1985). Delphacodes (8. l.), as DISTRIBUTION-NEARCfIC: Canada:Northwest traditionally recognized by North American Territory; lISA· Texas researchers, IS an obVIOUS paraphyletlc and polyphYleticenr~~fcludes12~NorthAmerican Paradelphacodes Htoralis (Reuter) (Figures 95 - 98, Map 6) De~haoodes~1.;::reviseditcan serve as a taxon (al~t inu::rt s~) for placement of problematic Ubilrnm litoralis Reuter 1880'198 SPeci~ De1phaoodes litom1is (Reuter), Metcalf 1943:463. PamlibUmia (StroelJingianellG) litomlis (Reuter), S SpecIes IS namedIn recognItion of Dr. G. A. bufriev'scontributionstothesystematicsofnorthem LeQuesne 1964:57. Holarctic delphacids. Stl'uebingianella lito1 alis (Reuter), Nast 1972.62. The distribution record for the specimen used in Paro.de1phaoodes 1itom1is (Reuter), Anumev 1980: this study is: YUKON TERRITORY: 16; 1 male 211. brachypter; 1 June. Distribution records for specimens used in this DISTmBUTION· NEARCI'IC: Canada: Yukon stud, are: YUKON TERRITORY. 21, 49, a male Territory. brachypters; 31 May-10July. Otherrecordsarefrom Nast(1972)anda specimenintheCanadianNational Kusnezoviella macleaniWilson Collection, Ottawa This species was recorded from Heleocharis and Phragmites (Ossiannilsson 1978). (Figures 88 91, Map 6) DISTRIBUTION - NEARCTIC: Canada: New- KUBn€ZQviella macleani WilSOIl 1988.339. founcHand, Yukon TeuiLmy, PALAEARCl'IC. Fin Kusnezoviella matisi Anufriev and Emeljanov land; Russia: Buryat Autonomous Republic, Yakut 1988:406, New Synonymy. Autonomous Republic; SCOtland. Distribution rec.mrls for specimens used in this study are: YUKON TERRITORY: 18, 19, 31, 33, 42, 49,58,60; 13 malebraehypters, 1malemaeropter,26 female brachypters, 1 female macropter, 30 May - 27 July. Otherrecordsarefroll1Anufriev andEmeljanov (lQRR\ ..nil Wil..nn (lQAA\ '-r ~ ~ " " I • N"'~4 '~~~,j~. V"I7~~ 1'- - TeITitory; • Delphacodes emeljanovi USA: Alaska; PALAEARCTIC: Russia: he.(11 " T. • ',. _ . - ! 'J, 'g' .l-'aradelphacodes litoralis ~ ~l .... " ,, .. \ OJ..",.,,' ,u..,vues aencis tseamer JI~- ...... '" V -\:"'\ IUUI C;, ~c. - ::7"+1 .....- "V _,.e' I\.~- .l...t".,,., De1phaoodes dentis Beamer 1948:102. Iv[ \\ \< l'\. --" The distribution record for the specimen used in 64"- r~ ~ v~\ -64 r ") -J ; - '-, male brachypter; 9 July. This species was described I~Y~ ~ J/1 from Texas, the only previously recorded locality '\~ n-. • ~ .~, ~ ., •• .".. Ib -r ') .L07"to _• .L 11.. "'UK~" - ~ <-V-, c\., identical in genitalic features to D. dentis specimens ft.~~ f'\ ) ~ '--J~\ ",, 'OJ I include this species in this study but feel that the I '~M~~ rt~t~ ~~\ . . ",",'.. ,. ~._ ~ ,~ -- -.I"' .. .. ~~ ~'~J "~ ,~ ~~.. -----'-;. __ =.>.-.___.1..,- \ mislabeled. Jo. 136' 132' 1k· 124' Vol. 6, No.2, June, 1992 95 Delphacodes erneljanovi V~lsor I, ThisspeciesisnamedforDr.AF.ErneljaIlovwhohas new species ro~=::a~('==f~an::e:am:t;; (Figures 99 - 103) studyare'NORrHWESTTERRITORY'Akla'dk.j Imele bi'8cliypter; 25 JUly. YUKON 1'ERRl1URY: 9j 1 rii8Ie HOLOTYPE: male bracbypter with label: "Yr. braehypter; 21 July. C8l'Cl'08s;21·vii·1987jS.G.Cannings",intheCanaman National ColleetioR, Ottawa.; PARATYPE: male DISTRIBUTION·NEARCfIC: Canada: Northwest brachypter with the following data: Northwest Territory, Yukon Territory. Tetlitory, Aklavik, 25 July 1931, Dlyant, Lot 000, CAS,intheUniversityofBritishColumbiacollection. Javesella peIlucida (Fabricius) (Figures 104 -106, Map 7) DESCRIPl'ION: BOOy mediumbrown. Fronswith area betweenlateralcarinaesUffusedwithdarkbrowntoblaCk. Bl'&Chypterous forewings hYiiline; short, not extending to F'ulgOm pelliicidii Fabricius 1794:7 pygofer.Metatibialspurwithca.20lateralteeth.Abdomen Delphacodespellucida(Fabricius), Muirand Giffard reddish bmWn With black transverse markings 1924:20. MALEGENITALIA:Pygofersubcylindrical;inlateral Javesella pellucida (Fabricius), Fennah 1963:15. View, height ca 2 5 X Width, with strongly produood, subtriangular diaphragm armature. Anal tube DistributionreconJsforspecimensusedinthisstudy are: ALASKA: Skagway; Paxson, 21 mi Nj Mosquito I eke, 59"27'N 136"02'W. NORtHWEST'IERRl'lURY: longitudinal axis at which point shaft of style inflects Aklavik.YUKONTERRITORY: 1,3,12,21,38,40,41,42, lllwar d; remainderofshaftsubequalhi width throughout 43, 44, 45, 46, 49, 56, ID male brachyptem, 17 male length; apex obliquely angled. Aedeagus laterally JIl8Cl'QPters; 15 May - 2 August. Other:records are from campr essed, recur ved, broader atbase nanowing toward Vllbaste (1971)andWiJson(1988): J.pellucidaisa vector apex; gonopore subapical, on right side; left side with 3 ofviral pathogensofseveral rereals(W'Jlson andO'Brien small teeth, right side with 4 small teeth. 1987); studies of the biology of this planthopper were ThissptJGit!tsisplaa:dinlhegelluslMPJauxdes($.I.) SlBllDlflI"ized by Moehida andKisimoto (1971). farthereasonsgivenunderthediscussionofD.Q1W,{rieui. DISTRIBUTION - NEARefIC: Canada: Alberta, NorihwP.St TAmtOl"}', Vllknn TAmtory; TTSA' Ala..k", 134' 128' 122' COnnecticut, IllinOIS, Mam.e, Massachusetts, New Nno*h Nno*h ~-• 68'- • Javesella pellucida -ea lee ana, lTalY, . .. .f. .11. , .... , (Figures 107 - 109, Map 8) 104- Delphax obswrella Baheman 1847:53. Javesella obswrella(Baheman), LeQuesne 1964:57. -62 Distribution records far specimens used in this -J YUKONTERRITORY: 51; 3 malebrachypters;11-22 ~-J' ~_.~. -~ ~~~ (1971) and Wilson (1988). This species is a vector of 132' 1:28' 124' viralpathogensofcereals(Wilson andO'Brien 1987); 96 Insecta Mundi the biology ofit; on cereal crops was summarized by DISTRIBUTION - NEARCTlC: Canada: Yukon Ossiannilsson (1978). Tenitory; USA: Alaska; PALAEARarIC: Estonia, Finland; RUSSIa: Khabarovsk Terntory, Magadan DISTRIBUTION-NEARCl'IC: Canada: Northwest Region, Taymyr Autonomous District. Territory, Yukon Territory; USA: Alaska; PALAEARCTIC. Austria, Belgium, Bulgaria, JaveseHa f1a Wilson, new species Czechoslovakia Denmark Estonia, Finland France, (Figure 116 121, Map 9) East German;, West Germany, Great Britain, HOLOTYPE: m8IEl maGl'OpterWith label: ·YUKON, Svl1itzerland, Turkey, Yugoslavia. KOIDERN: 22-VII-1979; G. G. E. Scudder", in the CanadianNationalCollection, Ottawa., PARATYPB. Javesella discoLOi (Soheman) malemacroJLterwiththesamedata,intheUniversity of British Columbia collection. (Figures 110 112, Map 8) DESCRIPI'ION: Vertex dark brown spot, frons and Delphax discoLOr Boheman 1847:61. slypeusblaek;allwithlightyellowGarinae.Antennal&Gape Javesella discolor (Boheman1l.eQuesne 1964:57. and base of pedicel dark brown, rest of pedicel yellow. Pronotumpaleyellow, arealateraltolateralearinaeblaek. .Distribution records for specimens used in this Mesonotumblack;tegulaeyellow;forewings milkyhyaline, study are: NORTHWEST TERRITORY: Aklavik. oostal vein dark brown. Legs yellow, suffusedwith brown 'iT IKON 1WRlTORY: 21, 35, 37, 40, 41, 52, 53. to~ ALASKA: Paxson, 21 mi; 16malebrachypters, 2male 'DlaeJ:opteIS, 9 June •22July. Othenecords are from VIew, WIdth ca 1.5 X heIght, lateral aspects appeartoflare Vilbaste (1971) and Wilson (1988). This species has outwards; ventral margin of diaphragm opening almost been reported to be a vector of viral pathogens of straIghttooonvex m mlddle, diaphragmarmaturelaCking cereals (Wilson and O'Brien 1987); host plants were but region under diaphragm opening slightJy produced summarized by Ossiannllsson (1978). alongmidline.Analtubesubcylindrical, two acuteslightly curved spines extending frOm dorsocaudal margin and DISTRIBUTION-NEARcrIC'Canada' Northwest meeting along inner margins (in holotype, not paratype - .., ~p , ~ --~ .. J.u....uu J. ~y, UOfi~ , 134" 128' 1 Fig. 117). Styles stlongly divelging, each style widest at (1972, 1979, 1982) or Zoological Record fliJm1972 In base,nalTOWing distally;outerandinnermarginssinuous, 1988. ronstnetedjustbeforefoot shapedapex:. Aedeaguslaterally oompressed, almost straight; gonopore apical on ventral aspect;apexsharpIYangledantel'O'.rentr&lly, apexsubacute ern ry; AH: c at a gIon, dorsally, acute and tooth-like ventrally. Magadan Region, Sakhalin Region. Distribution records fot specimens used in this study ale. YUKON TERRITORY. 21: 2 male brachypters; 22July; ~EWHAMPSIDRE: Coos Co., JaveseDa kilmani (Van Duzee) Moodse FIIlls Cpgd, NW Second ConnAclicnt flo; 23 new combination July; 2 male macropters; Mt. Washington, Auto Rd., (Figures 125 - 127, Map 8) 2700 ft.; 1 July; 1 male maeroptel'. Libumia kilmani Van Duzee 1894:191. DlSTRlBUPION . NEARaPIC. Canada. Yukon DelpJUJiXXJ.e8 kilmani (VanDuzee), Muirand Giffard Territory; USA: New Hampshire. 1924:85. Jave881la berihgiaca Emeijanov Distribution records for specimens used in this (Figures 122 - 124, Map 7) studyare' YIIKONTERRTTORY: 2, 19, 23, 29,61,62; 9 maIes, 3 females, all bracliypters; 3 June· 19 July. Javesella beringiaca Emeljanov, Anumev and Otherrecords are from DuBose (1960). Emeljanov 1988:421 DlSTRIBUPION . NEARCTIC. Canada: AlLeI ta, The distribution record for the specimen used in Quebec, Manitoba, YukonTerritory;USA: Michigan, this study is: YUKO~ TERRITORY: 36; 1 male New Hampshire, North Carolina, New York, Ohio. brachypter.11June.OtherrecordsarefromAnufriev and Emeljanov (1988). Anufrlev and Emeljanov (1988) list this speCIes as Javesella beringzaca Comments on Diversity and Distl1butlon Emel.janov; no reference is made to itin either Nast The Yukon delphacid diversity of 32 species is roughly comparable to that of other Jar northern areaswhichhavebeensurveyed.Thirtysixdelphacid species have been recoriled fi'Om Norway, 70 fi'Om 134" 1i<1l" 122" £"0_ .:I t!1 L ..",., .:I'~' • • "'...,,, 'L>A ,n~a.... / , , r,'~ .from Estonia (Vilbaste.. .1971). These countries each 68'- 1Q)~\i •- Javesella simillima -68 Finland has no substantial area BOuth of600 ~. The ! UtI; have beencataloguedinclude Magadan Region with -r' 'C 'Y .....:. , u 10, '.\ - --<: •.• 01\ .J "u..;o- ~ .. - / " &" , IJIM1 / ?( "?) with37(AnufrievandEmeljanov1988).Fifteenspecies " W In ,) ~~\\~, lightoftheresultsofthis study,itislikelythatthisis ''''''- ~. -64 ",n .i""" +~ '",,,IF ~..- r ( >L-/~~~ collecting especially in BOutheastern. Alaska. I~ '-" ~ ~ /~ TheYukondelphacidfaunaincludesspecieswith TT . " ...... ,.., ,-J . " ...... e 6Z- If( \ ~~J / \""'\ -62 (see Scudder(1979b)fordiscussionofthecategoriesof ~,,~ i .. I)J (\'\ 1~ "\ f)" A Yukon delphacid species are Holarctic. Fourofthese , ".-~ ,- . , , ~. .. 6ff_'v._r-L7~~ S ~ V X ]1_~6o albocarinata)arewidelydistributedinthePalaearctic 140" 136" 132' 128' 124" and are also found in the northwestern. ~earctic. 98 Insecta Mundi Threespecies(J.simillima,P.litorolis, R.albostriata) Anufriev, G. A. 1979. Notes on the genus are circumpolar, restricted to a far northern Tyrphodelphnx Vilbaste, 1968 with descriptions distributionWlthffiboththePalaearcticandNearctic. of two new ASIatIc specIes (Homoptera, Tenspecies(A. subarctica, C. wilhelmi, J. beri7'lgiaoo, Delphacidae). Reiehenb8£hia 17:295 301. K. macleani, M. flavus, N. eburneocarinata, N. guentheri, N. tshaunica, N. umbmta, R. pusilla)have Anufriev,G.A. 1980.Anewspeciesandnewgenera an amphi-Beringian distribution occurring in the of leafhoppers of the family Delphacidae northeaslern Palaearctic and the westenl Nearcti.c. (Homoptera, Auchenorrhyna) frolli Palaearctic. The remaining species have been recorded only Zool Zh 59·208-216 fromtheNearctic. Fourspec1es(A. analzs, A. acuta, J. kilmani, J. lta) 8l'e boreal andtwo others (A. stylata, .."-nutrie...., Go A. & A. M. Averkin. 1982. CiGadinea C. magnifrOns) are Cordilleran in distribution. One of the family Delphacidae (Homoptera, species (D. campestri8) is found throughout most of Auchenorrhyncha) from the Sokhondo ReBel ve theeasternNearet.ic. Anotherspecies(D.dentipennis) (Transbaicalia). Ins. Mongolia 8:123- 139. hasbeenreporledpreviouslyh'OllltheeastemNearet1c; it.spresenceintheYukonmayrepresenta disjunction Anufriev, G. A. & A. F. Emeljanov 1981 The FIve species (A. hOChiie, D. anuj'rieVt, D. emel}anovi, genusAchOl"OttleFieb. (Homoptera,Delphacidae) N. glacia, Y. kendallae), all ofthemne'wly deseribed, and its systematics in the Palearetic Region. havebeenrecordedonlyfromtheYukonorNorthwest Entomol. Rev. 59:87-97. Tenitories. The record of D. ckntis from Northwesl Territories is probably accidental as it has been Anufriev,G.A. &A.F.Emeljanov.l9B8.Cicadinea recorded previously only from Texas (Beamer 1948). (AuchenOIThyncha)-Leafhoppers,pp.12-495. In: Twelve of the 15 Alaskan delphacids (Wilson ~ra, P. A., ed. [Identificationguidetotheinsects 1988)are alsorecorded fi'Om theYukonTerritory. Of of the far eastern USSR. Vol. 2. Homoptera and those not recorded nom the Yukon, two species, J. Hemiptera.] arcanastyla(Beamer)andJ. atrata(Osborn), appear lo extend only as far north as southeastern Alaska Asche, M. 198fl. Zur Phylogenie der Delphacidae and another, lIn kanodes€XCi sa (Melichar),isbroadly Leach, 1815 (Homoptera Cicadina distributedm the PaIaearctic, was found in western Fulgoromorpha). Marburger EntomoI. Publ. Alaska and might occur in the northern. Yukon 2(1):1910. Territory. Beamer,RH.l946.SomenewspeciesofDelphaa:xks Acknowledgments I thank Dr. G. G. E. Scudder, Department of EntnmnJogy, IIniVersity Of British Columbia, fOr loaning specimens and Dr. K G. A. Hamilton, Beamer,RH.1948. SomenewspeciesofDelphaoxles BiOSYstematiesResearehCentre,AgrieultureCanada, (lIomoptera,Fulgolidae,Delphacinae)ParlIV.J. Ottawa, for offeringhelpful suggestions. I appreciate Kansas Entomol. 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