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Checklist of Lycidae (Coleoptera: Elateriformia) of Taiwan

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Checklist of Lycidae (Coleoptera: Elateriformia) of Taiwan 研究報告 台灣昆蟲 22: 171-187 (2002) Formosan Entomol. 22: 171-187 (2002) Checklist of Lycidae (Coleoptera: Elateriformia) of Taiwan Ming-Luen Jeng Division of Entomology, Natural History Museum and Biodiversity Research Center, and Department of Ecology and Evolutionary Biology, Snow Hall, 1460 Jayhawk Blvd., University of Kansas, Lawrence, KS66045-7523, U.S.A. Ping-Shih Yang* Department of Entomology, National Taiwan University, Taipei 106, Taiwan, R.O.C. Masataka Satô Laboratory of Natural Conservation, Graduate School of Nagoya Women’s University, Mizuho -ku, Nagoya 467-8610, Japan ABSTRACT The checklist of Lycidae beetles of Taiwan, including five subfamilies, 20 genera and 74 species, is provided. Chinese names, synonyms, original references, type localities, type depositories and distributions are included. Cautires jengi Kasantsev was selected as the valid name according to the first reviser principle and C. yengi is invalid; Cladophorus certus Kleine is replaced by Cautires taiwanensis Jeng et Yang, nom. nov. Key words: Taiwan, Lycidae, checklist, new replacement name. there were 67 species in 22 genera recorded from Taiwan. Since then the The taxonomy of Lycidae of higher classification of Lycidae has Taiwan has been extensively studied undergone more frequent modifications, since the 1920s. Kleine (1926) was first and as a consequence, the taxonomy to review the Taiwanese lycid fauna and scientific names of lycids, from which included 30 species in 19 genera. family to species groups, have changed Miwa (1931) listed 18 genera and 32 considerably during the last two species of Lycidae in his catalogue of decades. Herein we provide the most Taiwanese beetles. Later Kôno (1932) updated checklist of Taiwanese Lycidae, reviewed the Japanese Lycidae in which and a total of 5 subfamilies, 20 genera 30 species in 17 genera were recorded and 74 species are recorded. from Taiwan. Kleine (1933), in Junk’s Coleopterorum Catalogus, listed 28 Material and Methods species in 17 genera from Taiwan. After the World War II, Nakane extensively In the checklist, Chinese names, studied Taiwanese Lycidae in the late original sources, synonyms, type localities, 1960s and early 1970s and raised the type depositories, and distributions are total species number to over 60. Bocák included. Additional remarks, if and Bocáková (1987) mentioned that necessary, were added to account for *Correspondence address e-mail:[email protected] Checklist of Lycidae of Taiwan 171 changes in the scientific names or Tokyo, Japan distributions. The higher classification in NTU Dept. Entomology, National the checklist is a modified system based Taiwan University, Taipei, on Bocák and Bocáková (1990), Lawrence Taiwan and Newton (1995) and Bocáková (2001). OMNH Osaka Museum of Natural Because the tribal and subtribal History, Osaka, Japan classifications have changed frequently, OXUM Hope Department of we did not use these categories but only Entomology, Oxford, U.K. subfamilies. The species list is largely ZMAS Zoological Institute, Russian based on Pic (1912, 1921, 1922a, b, 1925, Academy of Sciences, St. 1926, 1937), Kleine (1924, 1925, 1926, Petersburg, Russia 1933), Kôno (1932), Nakane (1967a, b, 1968, 1969a, b, c, 1971, 1973), Bocák and Checklist of Lycidae of Taiwan Bocáková (1987, 1992), Matsuda (1992), Kasantsev (1993, 2000), Bocáková (1997), LYCINAE 紅螢亞科 Kasantsev and Yang (1998) and Bocák Genus Lycostomus Motschulsky, 1861 (1999); type depositories denote the 吻紅螢屬 depositories of the name bearing types, Motschulsky, 1861: Bull. Moscou XXXIV: and are based on Kleine (1926), 136. Mroczkowski (1959), Nakane (1971, 1973) and Döbler (1982). Type status (syntypes, Lycostomus atrimembris Pic, 1926 軟黑 lectotype or holotype and its sex) is 吻紅螢 specified whenever possible, otherwise we Lycostomus atrimembris Pic, 1926: used “type(s)” to express the unknown L’Échange 42(425): 31 (hors texte). status. The citations of the original Lycostomus jeanvoinei Pic, 1927: Mélange sources are given for each taxon but are exot.-Entomol. 49: 4. not listed in the references; the Type locality: VIETNAM: Chyapa. abbreviations of journal titles are Type depository: Lectotype in according to BIOSIS format. MNHN. The following acronyms were used Distribution: Vietnam and Taiwan. to denote the type depositories: Remarks: Kasantsev (1993) CAS California Academy of Sciences, designated the lectotype for this species San Francisco, CA, U.S.A. and synonymized L. jeavoninei Pic with it. DEI Deutsches Entomologisches This species was recorded from Taiwan Institut, Müncheberg, Germany by Kasantsev and Yang (1998). EUJ Ehime University, Matsuyama, Meanwhile these authors indicated that Japan the record of L. honestus Bourgeois from HUJ Hokkaido University, Sapporo, Taiwan might be a misidentification of L. Japan atrimembris or L. placidus. IZW Instytut Zoologii, PAN, Warszawa, Poland Lycostomus crassus Kleine, 1926 長節吻 MNHN Muséum national d’Histoire 紅螢 naturelle, Pairs, France Lycostomus crassus Kleine, 1926: Stett. MNHU Museum für Nature Künd des Entomol. Z. 87: 97. Humbolt University, Berlin, Type locality: TAIWAN: CHIAI: Germany Dalin (= Taihorin) (嘉義縣大林). NHML Natural History Museum, Type depository: Holotype female London, U.K. in MNHU. NSMT National Science Museum, Distribution: Taiwan. 172 台灣昆蟲第二十二卷第二期 Ann. Mus. Civ. Stor. Nat. Genova Lycostomus formosanus Pic, 1937 蓬萊 XVIII: 626. 吻紅螢 Type locality: INDIA. Lycostomus formosanus Pic, 1937: Type depository: Type(s) in OXUM. L’Échange 53 (469): 147. Distribution: Ceylon, India, Type locality: TAIWAN (= Formose). Philippines, Taiwan and Thailand. Type depository: Holotype male in Remarks: The above synonymy was MNHN. listed by Kleine (1933). Distribution: Japan (Yayeyama Is.) and Taiwan. Genus Macrolycus Waterhouse, 1878 Remarks: No specific locality except 碩紅螢屬 “Formose” was provided by Pic (1937). Waterhouse, 1878: Trans. Entomol. Soc. Nakane (1973) examined the type but did Lond. 1878: 96. not give further information of locality. Satô and Matsuda (1985) recorded this Macrolycus alishanus Nakane, 1967 高 species from Japan. 山碩紅螢 Macrolycus alishanus Nakane, 1967: Lycostomus placidus Waterhouse, 1879 Fragm. Coleopterol. 17: 72. 珠海吻紅螢 Type locality: TAIWAN: CHIAI: Lycostomus placidus Waterhouse, 1879: Alishan (嘉義縣阿里山). Illustrations of Typical Specimens of Type depository: Holotype male in Coleoptera in the Collection of the NSMT. British Museum I: 7. Distribution: Taiwan. Type locality: CHINA: Hong Kong? Type depository: Holotype in NHML. Macrolycus crassicornis Nakane, 1967 Distribution: China and Taiwan. 粗角碩紅螢 Remarks: Kasantsev and Yang (1998) Macrolycus crassicornis Nakane, 1967: first recorded this species from Taiwan. Fragm. Coleopterol. 17: 72. Type locality: TAIWAN: NANTOU: Lycostomus similis (Hope, 1831) 相似吻 Sung-gang (= Sung-kang) (南投縣松崗). 紅螢 Type depository: Holotype male in Lycus similis Hope, 1831: in Gary (ed.) NSMT. Zool. Misc.: 26. Distribution: Taiwan. Lycostomus similis (Hope): Waterhouse, 1879: Illustrations of Typical Macrolycus dominator Kleine, 1925 碩 Specimens of Coleoptera in the 紅螢 Collection of the British Museum I: Macrolycus dominator Kleine, 1925: 10. Dtsche. Entomol. Z. 1925: 327. Lycus triangularis Hope, 1831: in Gray Type locality: TAIWAN: (ed.) Zool. Misc.: 26. KAOHSIUNG: Fengshan (= Hoozan), Lycus suturalis Redtenbacher, 1848: Jiashian (= Kosempo); CHIAI: Dalin (= Hugel’s Kaschmir Insecta IV: 508. Taihorin) (高雄縣鳳山、甲仙與嘉義縣大林). Lycus geminus Walker 1858: Ann. Mag. Type depository: Syntypes in Nat. Hist. 3(II): 281. MNHU and IZW (type No. 623 and 624). Lycus cinnabarinus Cand., 1861: Mem. Distribution: Japan (Yayeyama Is.) Liege: 355. and Taiwan. Lycus coccineus Motschulsky, 1861: Bull. Remarks: Kleine (1925) described Moscou XXXIV: 136. this species based on 14 syntypes. The Lycostomus patruelis Bourgeois, 1883: Japanese population was another Checklist of Lycidae of Taiwan 173 subspecies named ishigakianus Satô and NSMT. Ohbayashi. Distribution: Taiwan. Macrolycus erythropterus Nakane, 1967 Macrolycus taiwanus Nakane, 1967 臺 赤翅碩紅螢 灣碩紅螢 Macrolycus erythropterus Nakane, 1967: Macrolycus taiwanus Nakane, 1967: Fragm. Coleopterol. 17: 72. Fragm. Coleopterol. 17: 72. Type locality: TAIWAN: NANTOU: Type locality: TAIWAN: NANTOU: Sung-gang (= Sung-kang) (南投縣松崗). Sung-gang (= Sung-kang) (南投縣松崗). Type depository: Holotype male in Type depository: Holotype male in NSMT. NSMT. Distribution: Taiwan. Distribution: Taiwan. Remarks: Nakane (1967) spectulated Macrolycus flabellatus (Motschulsky, that the record of M. flabellatus from 1860) 扇角碩紅螢 Taiwan might be a misidentification of M. Lygistopterus flabellatus Motschulsky, taiwanus 1860: Schrenck’s Reisen Amurlande II(2): 114. Genus Dilophotes Waterhouse, 1879 細 Macrolycus flabellatus (Motschulsky): 紅螢屬 Bourgeois, 1902: Bull. Mus. Fr. VIII: Waterhouse, 1879: Illustrations of 89. Typical Specimens of Coleoptera in Celetes pectinifer Kiesenwetter, 1874: the Collection of the British Berl. Entomol. Z. XVIII: 251. Museum I: 75. Macrolycus pectinifer (Kiesenwetter): Gorham, 1883: Trans. Entomol. Soc. Dilophotes atricollis Pic, 1926 黑胸細紅 Lond. 1883: 391. 螢 Cerceros pectinicornis Kraatz, 1879: Dilophotes atricollis Pic, 1926: Entomol. Dtsche. Entomol. Z. XXIII: 127. Mitt. XV: 68. Type locality: RUSSIA: E. Type locality: TAIWAN: KAOH- SIBERIA. SIUNG: Jiashian (= Kosempo) (高雄縣甲 Type depository: Type(s) in ZMAS. 仙) Distribution: China, Japan, Korea, Type depository: Holotype in DEI. Mongolia, Russia and Taiwan. Distribution: India and Taiwan. Remarks: The above synonymy
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