Four Highlights from Natura 2000 Forests NGO Recommendations on Management Four Highlights from Natura 2000 Forests: NGO Recommendations on Management

Total Page:16

File Type:pdf, Size:1020Kb

Four Highlights from Natura 2000 Forests NGO Recommendations on Management Four Highlights from Natura 2000 Forests: NGO Recommendations on Management Four Highlights from Natura 2000 Forests NGO recommendations on management Four Highlights from Natura 2000 Forests: NGO recommendations on management Authors and contributors: Wilderness and non-intervention management | Stefan Avramov, Bulgarian Biodiversity Foundation, and Pawel Pawlaczyk, Naturalists Club Poland Management for deadwood retention | Pawel Pawlaczyk and Monika Kotulak, Naturalists Club Poland Light forests | Matthias Dolek, Butterfly Conservation Europe, and Zoltán Ilonczai, Bükki National Park Directorate Ecological coherence | Tamás Papp and Csaba Domokos, Milvus Group, and Andriy-Taras Bashta, Animal Research and Protection Association “Fauna” Four Highlights from Natura 2000 Forests Editor | Ildikó Arany NGO recommendations on management Proofreading | Judit Herman Layout | Kristina Vilimaite Editor responsible | Ágnes Zólyomi Photo on the cover page | Piotr Lewandowski Copyright © CEEweb for Biodiversity, 2013. CEEweb for Biodiversity is an international network ofnon-governmental organizations in Central and Eastern Europe. The mission of the network is the conservation of biodiversity through the promotion of sustainable development. CEEweb for Biodiversity This publication is supported Széher út 40. by the European Commission. H-1021 Budapest, Hungary The donor is not responsible Tel.: (+36 1) 398-0135 for the expressed views Fax: (+36 1) 398-0136 and the use ofthe information www.ceeweb.org made available. 2013 Table of contents Introduction ......................................................................4 Wilderness and non-intervention management ....6 Management for deadwood retention..................22 Light forests .........................................................34 Ecological coherence ...........................................42 Recommendations and conclusions.................................50 References ......................................................................54 2 3 3 support the implementation ofthis target and action along with the Strategy’s other provisions, such as the Biogeographic Process where forestry management is a highlighted issue or the preparation of the Guidance on the management offorests in Natura 2000. Introduction However, it is often unclear for planners and managers alike how the above provisions should be integrated into actual management practice. Therefore, it is crucial to share experience and knowledge In 2010 forests and other wooded land covered 177.8 million on different forest management options and their role in achieving hectares which is more than 40% ofthe EU’s land area. This number favourable conservation status of forests. It is also crucial to enhance has been growing throughout the last twenty years. The majority cooperation between countries and stakeholders. These are some ofwooded land is covered by semi-natural forests or woody ofthe goals expected from the ongoing EU processes. plantations, whereas around 20.4 million hectares (equivalent to 13% ofthe total area) offorests were in protected areas in the EU in With this brochure NGOs wish to contribute to this knowledge by 2010.1 European forests are among the most important ecosystems sharing their field-based experiences. We have decided to highlight providing necessary services for Europe’s citizens, including a four selected issues offorest management and to provide a list number or provisioning, cultural, supporting and regulating services. ofgood examples. These issues are relevant across most biogeographical regions and forest habitat types and are important Despite their growing area European forests are exposed to for achieving favourable conservation status of Natura 2000 forest a number ofpressures such as climate change, habitat fragmentation, habitats and the target species they host. Non-intervention invasive alien species or the intensification offorest management. management is one ofthe management options for all zonal climax At the same time resilience and adaptation capacity offorests to forests with large coverage across Europe, especially those in the these pressures depends on their biodiversity and ecological Boreal and Alpine regions. Ecological coherence is key to ensuring coherence. This was already recognized by the EU’s 2006 Forest forest resilience against climate change and other environmental Action Plan which states maintaining and enhancing forest pressures, and important for emblematic species such as large biodiversity as one ofits aims.2 Maintaining biodiversity, sustaining carnivores. Management for deadwood retention is a rather technical 4 ecological health and functioning in forests depends largely on their modification of current practices with significant benefits for 5 5 management. This was acknowledged by the EU Biodiversity Strategy structural and biological diversity ofthe forest. In the context of this to 2020 by adoptingits Target 3: Increase the contribution of brochure light forests are created and sustained by management agriculture and forestry to maintaining and enhancing biodiversity and practice which opens the canopy and allows light to penetrate to the its Action 12: Integrate biodiversity measures in forest management understorey thus enabling a diverse mosaic offorest and grassland plans.3 These plans should include important measures such as patches and favouring traditional land use practices at the same time. maintaining optimal levels of deadwood, preserving wilderness areas, applying ecosystem-based measures to increase the resilience We especially invite forest planners and managers, land use offorests against fires and adopting specific measures developed for planners, conservation professionals and other interested readers Natura 2000 forest sites. There are several ongoing EU processes to to explore, learn and disseminate our findings. Wilderness and non-intervention management According to the definition given in the EU Guidelines on Wilderness in Natura 2000,4 a wilderness is an area governed by natural processes. It is composed ofnative habitats and species, and large enough for the effective ecological functioning ofnatural processes. It is unmodified or only slightly modified and without intrusive or extractive human activity, settlements, infrastructure or visual disturbance. Wilderness areas are typically national parks or reserves Above: The Three-toed Woodpecker ofIUCN category I covering thousands ofhectares and with a history occurs only in old-growth forests of strict protection. The term ‘wild area’ is used for sites (…) where Right: The highest spruce tree only some ofthe wilderness qualities are found, where the ofBulgaria is in the non-intervention conservation objectives aim at achieving only part ofthe wilderness forest of Parangalitsa reserve, qualities, or where the objective is to fully restore natural processes Rila Mountains and features with the aim to extend the wilderness core zone. Wild areas can have various statuses, for example core zones ofnationally protected or Natura 2000 sites or entire smaller nature reserves under strict protection. Wild areas do not necessarily have a long between minimum intervention and non-intervention along which history ofbeing undisturbed but they need to have the potential the level ofintervention decreases even in case ofnaturally of restoring and sustaining natural processes. occurring disturbances (e.g. bark beetles or forest fires). 6 7 7 Wilderness and wild areas are generally subject to non-intervention Natural dynamics enabled by non-intervention management: all kinds ofhuman intervention are avoided that could Natural structural and functional diversity is a precondition have negative effects. (Exceptions are scientific investigation and in for maintenance ofhigh biodiversity in forests. There are different some cases guided tourism.) In other cases a minimum intervention5 levels and scales ofnatural ecosystem dynamics. In case offorests approach is applied which limits active management to non- it varies from small-scale gap dynamics (as in lowland beech forests) extractive and preventive measures such as eradication ofinvasive through “stand development phases” dynamics (as typical for alien species, maintaining touristic paths, etc. While in each case the montane mixed forests) to the large-scale disturbance dynamics guiding principle is to enable natural processes, there is a continuum (as in typical montane spruce forests). All manifestations of these dynamics should be accepted in wilderness and wild areas as Applicability manifestations of the natural processes maintaining the ecosystem. Even if natural processes are seemingly destructive for the forest Non-intervention management areas significantly contribute stands (as massive wind throws or bark beetle outbreaks), non- to the favourable conservation status ofa large set ofhabitat types, intervention is often a good strategy and may bring along especially primary habitats and dynamic complexes of habitats as conservation benefits. Non-intervention is often less destructive well as the species they host. Examples ofhabitats that benefit from for forest-related biodiversity than active intervention against non-intervention management can be found in all habitat groups the outbreaks; and forest recovery by natural succession is usually under Annex I to the Habitats Directive from aquatic to terrestrial and faster and cheaper than by artificial planting. A network of large non- from closed canopy forests to open grasslands, dunes, peatlands intervention areas contributes
Recommended publications
  • Insect Cold Tolerance: How Many Kinds of Frozen?
    POINT OF VIEW Eur. J. Entomol. 96:157—164, 1999 ISSN 1210-5759 Insect cold tolerance: How many kinds of frozen? B rent J. SINCLAIR Department o f Zoology, University o f Otago, PO Box 56, Dunedin, New Zealand; e-mail: [email protected] Key words. Insect, cold hardiness, strategies, Freezing tolerance, Freeze intolerance Abstract. Insect cold tolerance mechanisms are often divided into freezing tolerance and freeze intolerance. This division has been criticised in recent years; Bale (1996) established five categories of cold tolerance. In Bale’s view, freezing tolerance is at the ex­ treme end of the spectrum o f cold tolerance, and represents insects which are most able to survive low temperatures. Data in the lit­ erature from 53 species o f freezing tolerant insects suggest that the freezing tolerance strategies o f these species are divisible into four groups according to supercooling point (SCP) and lower lethal temperature (LLT): (1) Partially Freezing Tolerant-species that survive a small proportion o f their body water converted into ice, (2) Moderately Freezing Tolerant-species die less than ten degrees below their SCP, (3) Strongly Freezing Tolerant-insects with LLTs 20 degrees or more below their SCP, and (4) Freezing Tolerant Species with Low Supercooling Points which freeze at very low temperatures, and can survive a few degrees below their SCP. The last 3 groups can survive the conversion of body water into ice to an equilibrium at sub-lethal environmental temperatures. Statistical analyses o f these groups are presented in this paper. However, the data set is small and biased, and there are many other aspects o f freezing tolerance, for example proportion o f body water frozen, and site o f ice nucleation, so these categories may have to be re­ vised in the future.
    [Show full text]
  • Potential European Bison (Bison Bonasus) Habitat in Germany
    PROJECT REPORT Potential European bison (Bison bonasus) habitat in Germany Tobias Kuemmerle Humboldt-University Berlin Benjamin Bleyhl Humboldt-University Berlin Wanda Olech University of Warsaw & European Bison Friends Society Kajetan Perzanowski Carpathian Wildlife Research Station MIZ, PAS & European Bison Friends Society 1 PROJECT REPORT CONTENTS CONTENTS ...............................................................................................................................................................2 INTRODUCTION .......................................................................................................................................................3 HISTORIC DISTRIBUTION OF EUROPEAN BISON IN EUROPE .....................................................................................4 EUROPEAN BISON HABITAT PREFERENCES ..............................................................................................................6 HABITAT USE OF CONTEMPORARY EUROPEAN BISON POPULATIONS .......................................................................................... 6 PALEO-ECOLOGICAL DATA ON EUROPEAN BISON HABITAT USE ............................................................................................... 10 MAPPING EUROPEAN BISON HABITAT IN GERMANY ............................................................................................ 10 APPROACH .................................................................................................................................................................
    [Show full text]
  • Green-Tree Retention and Controlled Burning in Restoration and Conservation of Beetle Diversity in Boreal Forests
    Dissertationes Forestales 21 Green-tree retention and controlled burning in restoration and conservation of beetle diversity in boreal forests Esko Hyvärinen Faculty of Forestry University of Joensuu Academic dissertation To be presented, with the permission of the Faculty of Forestry of the University of Joensuu, for public criticism in auditorium C2 of the University of Joensuu, Yliopistonkatu 4, Joensuu, on 9th June 2006, at 12 o’clock noon. 2 Title: Green-tree retention and controlled burning in restoration and conservation of beetle diversity in boreal forests Author: Esko Hyvärinen Dissertationes Forestales 21 Supervisors: Prof. Jari Kouki, Faculty of Forestry, University of Joensuu, Finland Docent Petri Martikainen, Faculty of Forestry, University of Joensuu, Finland Pre-examiners: Docent Jyrki Muona, Finnish Museum of Natural History, Zoological Museum, University of Helsinki, Helsinki, Finland Docent Tomas Roslin, Department of Biological and Environmental Sciences, Division of Population Biology, University of Helsinki, Helsinki, Finland Opponent: Prof. Bengt Gunnar Jonsson, Department of Natural Sciences, Mid Sweden University, Sundsvall, Sweden ISSN 1795-7389 ISBN-13: 978-951-651-130-9 (PDF) ISBN-10: 951-651-130-9 (PDF) Paper copy printed: Joensuun yliopistopaino, 2006 Publishers: The Finnish Society of Forest Science Finnish Forest Research Institute Faculty of Agriculture and Forestry of the University of Helsinki Faculty of Forestry of the University of Joensuu Editorial Office: The Finnish Society of Forest Science Unioninkatu 40A, 00170 Helsinki, Finland http://www.metla.fi/dissertationes 3 Hyvärinen, Esko 2006. Green-tree retention and controlled burning in restoration and conservation of beetle diversity in boreal forests. University of Joensuu, Faculty of Forestry. ABSTRACT The main aim of this thesis was to demonstrate the effects of green-tree retention and controlled burning on beetles (Coleoptera) in order to provide information applicable to the restoration and conservation of beetle species diversity in boreal forests.
    [Show full text]
  • 11701-19-A0558 RVH Landmarke 4 Engl
    Landmark 4 Brocken ® On the 17th of November, 2015, during the 38th UNESCO General Assembly, the 195 member states of the United Nations resolved to introduce a new title. As a result, Geoparks can be distinguished as UNESCO Global Geoparks. As early as 2004, 25 European and Chinese Geoparks had founded the Global Geoparks Network (GGN). In autumn of that year Geopark Harz · Braunschweiger Land · Ostfalen became part of the network. In addition, there are various regional networks, among them the European Geoparks Network (EGN). These coordinate international cooperation. 22 Königslutter 28 ® 1 cm = 26 km 20 Oschersleben 27 18 14 Goslar Halberstadt 3 2 1 8 Quedlinburg 4 OsterodeOsterodee a.H.a.Ha H.. 9 11 5 13 15 161 6 10 17 19 7 Sangerhausen Nordhausen 12 21 In the above overview map you can see the locations of all UNESCO Global Geoparks in Europe, including UNESCO Global Geopark Harz · Braunschweiger Land · Ostfalen and the borders of its parts. UNESCO-Geoparks are clearly defi ned, unique areas, in which geosites and landscapes of international geological importance are found. The purpose of every UNESCO-Geopark is to protect the geological heritage and to promote environmental education and sustainable regional development. Actions which can infl ict considerable damage on geosites are forbidden by law. A Highlight of a Harz Visit 1 The Brocken A walk up the Brocken can begin at many of the Landmark’s Geopoints, or one can take the Brockenbahn from Wernigerode or Drei Annen-Hohne via Schierke up to the highest mountain of the Geopark (1,141 meters a.s.l.).
    [Show full text]
  • Cephalcia Lariciphila
    Prepared by CABI and EPPO for the EU under Contract 90/399003 Data Sheets on Quarantine Pests Cephalcia lariciphila IDENTITY Name: Cephalcia lariciphila Wachtl Synonyms: Cephalcia alpina Klug Cephalcia annulata Hartig Lyda alpina Klug Taxonomic position: Insecta: Hymenoptera: Pamphiliidae Common names: European web-spinning larch sawfly (English) Lärchen-Gespinstblattwespe (German) Bayer computer code: CEPCAL EU Annex designation: II/B HOSTS This insect attacks Larix spp., e.g. L. decidua and L. leptolepis. Interspecific hybrids are also attacked. GEOGRAPHICAL DISTRIBUTION EPPO region: Austria, Belgium, Czech Republic, Denmark, Finland, France, Germany, Italy, Netherlands, Russia (European, Siberia), Slovakia, Sweden, Switzerland, UK. EU: Present. BIOLOGY Adults fly in May-June, males often being more common. Eggs are deposited singly on short-shoot needles. Larvae spin a web-tube at the bases of the short shoots and remain in the web while consuming the needles. Most feeding takes place in July. The old larvae drop to the ground where they remain free as eonymphs without a cocoon 5-20 cm below the surface. In September they develop into pronymphs which overwinter. Pupation occurs in spring. Larvae may however remain in the soil as eonymphs for as many as four winters before they pupate (Röhrig, 1953; Pschorn-Walcher, 1982). A development cycle of 1-2 years is normal but, in some places, e.g. the Netherlands, up to 44% of the population was reported to have at least a 3-year generation cycle (Luitjes & Minderman, 1959). During outbreaks, population densities of 57-372 nymphs/m2 of soil were found. DETECTION AND IDENTIFICATION Symptoms C. lariciphila cuts the needle at the bases and pulls it into the web where it is eaten.
    [Show full text]
  • Small-Mammal Assemblages Inhabiting Sphagnum Peat Bogs in Various Regions of Poland
    BIOLOGICAL LETT. 2012, 49(2): 115–133 Available online at: http:/www.versita.com/science/lifesciences/bl/ DOI: 10.2478/v10120-012-0013-4 Small-mammal assemblages inhabiting Sphagnum peat bogs in various regions of Poland MATEUSZ CIECHANOWSKI1, JAN CICHOCKI2, AGNIESZKA WAŻNA2 and BARBARA PIŁACIŃSKA3 1 Department of Vertebrate Ecology and Zoology, University of Gdańsk, al. Legionów 9, 80‑441 Gdańsk, Poland 2 Department of Zoology, Faculty of Biological Sciences, University of Zielona Góra, ul. prof. Z. Szafrana 1, 65‑516 Zielona Góra, Poland 3 Department of Systematic Zoology, Adam Mickiewicz University, Umultowska 89, 61‑614 Poznań, Poland Corresponding author: Mateusz Ciechanowski, [email protected] (Received on 19 May 2011; Accepted on 1 March 2012) Abstract: We studied species composition of assemblages of small mammals (rodents and shrews) inhab­ iting Polish 25 ombrotrophic mires and quaking bogs in several regions in order to reveal characteristic features of their quantitative structure and compare them between regions, internal zones of the bog habitats, and different levels of anthropogenic degradation. We reviewed also all published results of small-mammal trapping in such habitats. Mammals were captured in pitfalls, snap traps and live traps on 12 bogs of the Pomerania region, 4 bogs of the Orawa-Nowy Targ Basin (Kotlina Orawsko-Nowotarska), 3 bogs in the Świętokrzyskie Mts, and 6 bogs in Wielkopolska and the Lubusz Land. Additionally, we included materials collected from Barber traps (pitfalls) used during studies of epigeic invertebrates on 4 bogs. In total, 598 individuals of 12 species were collected. The number of pitfall captures per 100 trap- nights was very low (7.0–7.8), suggesting low population density.
    [Show full text]
  • 00A-Okladka.Vp:Corelventura
    SOCIETY OF ECOLOGICAL CHEMISTRY AND ENGINEERING ECOLOGICAL CHEMISTRY AND ENGINEERING A CHEMIA I IN¯YNIERIA EKOLOGICZNA A Vol. 17 No. 12 OPOLE 2010 EDITORIAL COMMITTEE Witold Wac³awek (University, Opole, PL) – Editor-in-Chief Milan Kraitr (Western Bohemian University, Plzen, CZ) Jerzy Skrzypski (University of Technology, £ódŸ, PL) Maria Wac³awek (University, Opole, PL) Tadeusz Majcherczyk (University, Opole, PL) – Secretary PROGRAMMING BOARD Witold Wac³awek (University, Opole, PL) – Chairman Jerzy Bartnicki (Meteorological Institute – DNMI, Oslo-Blindern, NO) Mykhaylo Bratychak (National University of Technology, Lviv, UA) Bogus³aw Buszewski (Nicolaus Copernicus University, Toruñ, PL) Eugenija Kupcinskiene (University of Agriculture, Kaunas, LT) Bernd Markert (International Graduate School [IHI], Zittau, DE) Nelson Marmiroli (University, Parma, IT) Jacek Namieœnik (University of Technology, Gdañsk, PL) Lucjan Paw³owski (University of Technology, Lublin, PL) Krzysztof J. Rudziñski (Institute of Physical Chemistry PAS, Warszawa, PL) Manfred Sager (Agency for Health and Food Safety, Vienna, AT) Mark R.D. Seaward (University of Bradford, UK) Jíøi Ševèik (Charles University, Prague, CZ) Piotr Tomasik (University of Agriculture, Kraków, PL) Roman Zarzycki (University of Technology, £ódŸ, PL) Tadeusz Majcherczyk (University, Opole, PL) – Secretary EDITORIAL OFFICE Opole University ul. kard. B. Kominka 4, 45–032 OPOLE, PL phone +48 77 455 91 49 email: [email protected] http://tchie.uni.opole.pl SECRETARIES Agnieszka Do³hañczuk-Œródka, phone
    [Show full text]
  • Guidance Document on the Strict Protection of Animal Species of Community Interest Under the Habitats Directive 92/43/EEC
    Guidance document on the strict protection of animal species of Community interest under the Habitats Directive 92/43/EEC Final version, February 2007 1 TABLE OF CONTENTS FOREWORD 4 I. CONTEXT 6 I.1 Species conservation within a wider legal and political context 6 I.1.1 Political context 6 I.1.2 Legal context 7 I.2 Species conservation within the overall scheme of Directive 92/43/EEC 8 I.2.1 Primary aim of the Directive: the role of Article 2 8 I.2.2 Favourable conservation status 9 I.2.3 Species conservation instruments 11 I.2.3.a) The Annexes 13 I.2.3.b) The protection of animal species listed under both Annexes II and IV in Natura 2000 sites 15 I.2.4 Basic principles of species conservation 17 I.2.4.a) Good knowledge and surveillance of conservation status 17 I.2.4.b) Appropriate and effective character of measures taken 19 II. ARTICLE 12 23 II.1 General legal considerations 23 II.2 Requisite measures for a system of strict protection 26 II.2.1 Measures to establish and effectively implement a system of strict protection 26 II.2.2 Measures to ensure favourable conservation status 27 II.2.3 Measures regarding the situations described in Article 12 28 II.2.4 Provisions of Article 12(1)(a)-(d) in relation to ongoing activities 30 II.3 The specific protection provisions under Article 12 35 II.3.1 Deliberate capture or killing of specimens of Annex IV(a) species 35 II.3.2 Deliberate disturbance of Annex IV(a) species, particularly during periods of breeding, rearing, hibernation and migration 37 II.3.2.a) Disturbance 37 II.3.2.b) Periods
    [Show full text]
  • 25Th U.S. Department of Agriculture Interagency Research Forum On
    US Department of Agriculture Forest FHTET- 2014-01 Service December 2014 On the cover Vincent D’Amico for providing the cover artwork, “…and uphill both ways” CAUTION: PESTICIDES Pesticide Precautionary Statement This publication reports research involving pesticides. It does not contain recommendations for their use, nor does it imply that the uses discussed here have been registered. All uses of pesticides must be registered by appropriate State and/or Federal agencies before they can be recommended. CAUTION: Pesticides can be injurious to humans, domestic animals, desirable plants, and fish or other wildlife--if they are not handled or applied properly. Use all pesticides selectively and carefully. Follow recommended practices for the disposal of surplus pesticides and pesticide containers. Product Disclaimer Reference herein to any specific commercial products, processes, or service by trade name, trademark, manufacturer, or otherwise does not constitute or imply its endorsement, recom- mendation, or favoring by the United States government. The views and opinions of wuthors expressed herein do not necessarily reflect those of the United States government, and shall not be used for advertising or product endorsement purposes. The U.S. Department of Agriculture (USDA) prohibits discrimination in all its programs and activities on the basis of race, color, national origin, sex, religion, age, disability, political beliefs, sexual orientation, or marital or family status. (Not all prohibited bases apply to all programs.) Persons with disabilities who require alternative means for communication of program information (Braille, large print, audiotape, etc.) should contact USDA’s TARGET Center at 202-720-2600 (voice and TDD). To file a complaint of discrimination, write USDA, Director, Office of Civil Rights, Room 326-W, Whitten Building, 1400 Independence Avenue, SW, Washington, D.C.
    [Show full text]
  • A Comparison of Three Trapping Methods Used to Survey Forest-Dwelling Coleoptera
    Eur. J. Entomol. 103: 397–407, 2006 ISSN 1210-5759 A comparison of three trapping methods used to survey forest-dwelling Coleoptera ESKO HYVÄRINEN, JARI KOUKI and PETRI MARTIKAINEN Faculty of Forest Sciences, University of Joensuu, P.O. Box 111, FIN – 80101 Joensuu, Finland; e-mail: [email protected] Keywords. Abundance distribution, beetles, biodiversity inventories, boreal forest, Coleoptera, interception traps, pitfall traps, sampling, species richness, window traps Abstract. Sampling of insect communities is very challenging and for reliable interpretation of results the effects of different sam- pling protocols and data processing on the results need to be fully understood. We compared three different commonly used methods for sampling forest beetles, freely hanging flight-intercept (window) traps (FWT), flight-intercept traps attached to trunks (TWT) and pitfall traps placed in the ground (PFT), in Scots pine dominated boreal forests in eastern Finland. Using altogether 960 traps, forming 576 sub-samples, at 24 study sites, 59760 beetles belonging to 814 species were collected over a period of a month. All of the material was identified to species, with the exception of a few species pairs, to obtain representative data for analyses. Four partly overlapping groups were used in the analyses: (1) all, (2) saproxylic, (3) rare and (4) red-listed species. In terms of the number of species collected TWTs were the most effective for all species groups and the rarer species the species group composed of (groups 1-2-3-4) the larger were the differences between the trap types. In particular, the TWTs caught most red-listed species. However, when sample sizes were standardized FWTs and TWTs caught similar number of species of all species groups.
    [Show full text]
  • Status and Development of Old-Growth Elements and Biodiversity During Secondary Succession of Unmanaged Temperate Forests
    Status and development of old-growth elementsand biodiversity of old-growth and development Status during secondary succession of unmanaged temperate forests temperate unmanaged of succession secondary during Status and development of old-growth elements and biodiversity during secondary succession of unmanaged temperate forests Kris Vandekerkhove RESEARCH INSTITUTE NATURE AND FOREST Herman Teirlinckgebouw Havenlaan 88 bus 73 1000 Brussel RESEARCH INSTITUTE INBO.be NATURE AND FOREST Doctoraat KrisVDK.indd 1 29/08/2019 13:59 Auteurs: Vandekerkhove Kris Promotor: Prof. dr. ir. Kris Verheyen, Universiteit Gent, Faculteit Bio-ingenieurswetenschappen, Vakgroep Omgeving, Labo voor Bos en Natuur (ForNaLab) Uitgever: Instituut voor Natuur- en Bosonderzoek Herman Teirlinckgebouw Havenlaan 88 bus 73 1000 Brussel Het INBO is het onafhankelijk onderzoeksinstituut van de Vlaamse overheid dat via toegepast wetenschappelijk onderzoek, data- en kennisontsluiting het biodiversiteits-beleid en -beheer onderbouwt en evalueert. e-mail: [email protected] Wijze van citeren: Vandekerkhove, K. (2019). Status and development of old-growth elements and biodiversity during secondary succession of unmanaged temperate forests. Doctoraatsscriptie 2019(1). Instituut voor Natuur- en Bosonderzoek, Brussel. D/2019/3241/257 Doctoraatsscriptie 2019(1). ISBN: 978-90-403-0407-1 DOI: doi.org/10.21436/inbot.16854921 Verantwoordelijke uitgever: Maurice Hoffmann Foto cover: Grote hoeveelheden zwaar dood hout en monumentale bomen in het bosreservaat Joseph Zwaenepoel
    [Show full text]
  • Conspicuousness, Phylogenetic Structure, and Origins of Müllerian
    www.nature.com/scientificreports OPEN Conspicuousness, phylogenetic structure, and origins of Müllerian mimicry in 4000 lycid beetles from all zoogeographic regions Michal Motyka1, Dominik Kusy1, Michal Masek1, Matej Bocek1, Yun Li1, R. Bilkova1, Josef Kapitán2, Takashi Yagi3 & Ladislav Bocak1* Biologists have reported on the chemical defences and the phenetic similarity of net-winged beetles (Coleoptera: Lycidae) and their co-mimics. Nevertheless, our knowledge has remained fragmental, and the evolution of mimetic patterns has not been studied in the phylogenetic context. We illustrate the general appearance of ~ 600 lycid species and ~ 200 co-mimics and their distribution. Further, we assemble the phylogeny using the transcriptomic backbone and ~ 570 species. Using phylogenetic information, we closely scrutinise the relationships among aposematically coloured species, the worldwide diversity, and the distribution of aposematic patterns. The emitted visual signals difer in conspicuousness. The uniform coloured dorsum is ancestral and was followed by the evolution of bicoloured forms. The mottled patterns, i.e. fasciate, striate, punctate, and reticulate, originated later in the course of evolution. The highest number of sympatrically occurring patterns was recovered in New Guinea and the Andean mountain ecosystems (the areas of the highest abundance), and in continental South East Asia (an area of moderate abundance but high in phylogenetic diversity). Consequently, a large number of co-existing aposematic patterns in a single region and/or locality is the rule, in contrast with the theoretical prediction, and predators do not face a simple model-like choice but cope with complex mimetic communities. Lycids display an ancestral aposematic signal even though they sympatrically occur with diferently coloured unproftable relatives.
    [Show full text]