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From Australian Palaeozoic Rocks AUSTRALIAN MUSEUM SCIENTIFIC PUBLICATIONS Fletcher, Harold O., 1943. The genus Conocardium from Australian Palaeozoic rocks. Records of the Australian Museum 21(4): 231–243, plates xiii–xiv. [22 December 1943]. doi:10.3853/j.0067-1975.21.1943.537 ISSN 0067-1975 Published by the Australian Museum, Sydney nature culture discover Australian Museum science is freely accessible online at http://publications.australianmuseum.net.au 6 College Street, Sydney NSW 2010, Australia THE GENUS CONOCARDIUM FROM AUSTRALIAN PALAEOZOIC ROCKS. By H. O. FLETCH~JR, The Australian :rvrUseU1TIo (PlateR xiii-xiv.) Introduction. This" paper deals with the known species of ConocaTdi1t1n from Palaeozoic rocks in Australia. Reference is made to the rather limited records of the genus in literature, and new species are described from the Lower Middle Devonian (Murrumbidgee Series) of New South Wales, from the Upper Middle Devonian (Atrypa limestone) of Western Australia, and from the Permian of New South Wales and Queensland. The material examined is in the collection of the Australian Museum, with the exception of three specimens, described as Conocardium gogoense, sp. nov" from the collection of the University of Western Australia. I am indebted to Dr, Curt Teichert for the opportunity of describing this interesting species. The known geological range of the genus in Australia extends from the SHurian to the Upper Permian. The genus Oonocardiu1n embraces one of the early highly specialized types of shelI and was first recorded from Permian rocks in Australia by McCoy (1847, p. 300). Opinion is divided regarding the position of the animal within the shell and the correct orientation is still a subject of discussion. Some workers in this group consider the short truncated end of the shell to be anterior, while others are certain it should be considered the posterior end. Tryon (1884, p. 195), comparing the shell with Adacna, concluded that the conical and gaping end was anterior, the aperture being for the passage of the foot and the function of the long tube or rostrum being siphonal. This view, held by a number of authors, was elaborated by Hind (1900, pp. 452~453) who, after a close examination of the internal characters of a perfect specimen of Oonocardium hercule1tm, brought forward several reasons in" support of it. He considered the external ligament of Conocardium to be restricted, occupying a depression between the umbones and the long rostrum. He remarks: "This surely cannot be placed anterior to the umbones." Other authors, however, are of the opinion that the ligament is amphidetic or paravincular­ types one would rather expect in a shell of this type. Hind also states that in bivalve!> the anterior adductor scar is usually the deeper and better marked, and in many genera, as in Oonocardium, is bounded behind by a ridge of shelL In Cucullaea and other genera, however, we find the exact opposite, the posterior adductor scar being well defined and with a strongly developed ridge. The pallial line in Conocardiu1n is simple and the assumption that it is a siphonate form appears to be attributed to the gape, the presence of the elongated rostrum or analogies with other siphonate genera which have no pallial sinus. These are exceptional, and as a general rule siphonate forms are always recognized by the presence of a pallial sinus. This fact raises the question: what definite evidence have we that Conocardi1trn is a siphonate genus? Hind is of the opinion that the gaping aperture was used for the extrusion of the foot and with this I agree. He states, however, that the foot would be unlikely to protrude over the prominent or sharp interlocking teeth of the valve. In many genera this is exactly what occurs, a case in point being OardiuJ1!, in which the foot protrudes over much heavier teeth than are present in Oonocardiul1t. After a study of the evidence in favour of the trullcated end of the shell being posterior, and after several interesting discussions with Mr. Tom Iredale, Conchologist G K8COHDS or,' THE AFWrRALIAN :\lrS~:UM. at the Australian Museum, on the behaviour of recent shells, I conclude this view should be accepted with a great deal of reservation. Any suggestion that the tubular process or rostrum was used for the passage of siphons appears untenable, the extreme length and small diameter of the tube being strong evidence against it. XO reasonable suggestion can be put forward as to the function of the rostrum, unless it was for some respiratory purpose. Certainly it was not for a byssus, as the shell was obviously a mobile sand-burrower and not a sedentary byssifel'ous type. The general appearance of Conoca1'dit~m affords the best evidence regarding the orientation of its shell. The winged, conical and gaping end, through which the foot protruded, is obviously the burrowing end and when the shell was at rest would remain buried in the sand or mud. It is impossible to visualize the blunt, truncated end being used for this purpose. Making analogies with other burrowing genera, it is found in almost every case that the posterior end of the shell is used for burrowing and is developed structurally towards that purpose. For the present, therefore, I am following the authors who consider the trullcate end, provided "dth a tubular prolongation, as the anterior portion. Systematic Position of the Genus. The family Conocardiidae has been adopted almost universally to receive the single genus Conoca1'diu1n. The systematic pOSition of the family is still very much in doubt. In dealing with specialized or degenerate genera it is difficult and often misleading to arrive at definite conclusions by analogies with other bivalve groups. They are usually a law unto themselves. Hind rightly stated that it is difficult, if not impossible, to assign the to a correct systematic position. Historical Notes on the Genus in Australia. MeCoy (1847, p. 300) described and figured a shell as PleU1'01'hynchus austmlis from Permian rocks, stating the species was "not uncommon of larger size in the sandy schists of Wollongong, New South Wales". Dana (1849, p. 701) recorded the same species from Glendon, but referred it to the genus Ca1'diu1n. He wrongly suspected that McCoy's specimen did not come from Wollongong but was collected at Glendon. Dana also described and figured one end of a shell as Ca1'diu1n ferox. Etheridge (1881, p. 250) thought this may be another species of Gonoca1'diurn, but an analysis of Dana's description reveals that the large size of the shell, the thickness of the valves, and the character of the muscle scars preclude any reference to Conocardi1lm. De Koninck (1877, p. 109) recorded Gonocardium sowerbyi from "the black argillaceous limestone of the Yass District" which he rightly considered to be Devonian in age. He also (1877, p. 146) recorded Conoca1'diu1n australe? McCoy, from a railway cutting between Maitland and Stony Creek, an horizon almost certain to be the Branxton Stage of the Upper Marine Series. De Koninck was not confident his shell was identical with McCoy's species, though his description reveals few points of dissimilarity. Thp Rpe(;imfm was not figured, and as De Koninck'R type matprial was destroyed in the Garden Palace fire of 1882, the true identity of this shell must remain doubtful. Etheridge (1881, p. 250) described a single specimen as Conoca1'dium, sp. indet., from the :VJ:urrumbidgee Series, Lower Middle Devonian, at Bungaralaby, Lake Bathurst, near Goulburn, New South Wales. This shell is considered to be identical with Gonocardi7t1n soweTbyi De Koninck. Two species, Conoca1'diurrlo bellulu1n and C. costaturn, were described by Creswell (1893, p. 43) from the Lilydale limestone in Victoria. The descriptions are very limited, but fortunately the figures are good. The Lilydale limestone, previously considered Upper Silurian in age, is now known to be Middle Devonian (Hill, 1939, p. 219), Cbapman (1908, p. 45) recorded both the Victorian species with additional localities. The only Queensland record of the genus is that of Etheridge (1892, p. 278), who described a shell from the Gympie Series as Conoca1'diu1n aust1'ale McCoy. Later (1902, page 68) he stated: "The shell I formerly figured from the Permo-Carboniferous of ,Queensland as C. aust1'ale I now know is 110t that species; it is separated therefrom by the characters of its anterior end." The Queensland shell is distinct from the typical O. australe and has been named GonocaTdiu1n gyntlJiense, sp. novo Etheridge (1902, p. 68) described Gonocardiu1n australe from the Upper Marine Series at Hartley, New South Wales. This shell is an inflated type now renamed OonocoTdium Tobustu1n, sp. novo Dun (1907, p. 265) described two specimens from the Upper Silurian rocks at Oakey Creek, near Molong, New South Wales, as Gonocanliu1n davidis. Another specimen from the Upper Silurian Lower Limestone bed of the Bowning Series at Limestone Creek near Silverdale, was exhibited by Mitchell (1906, p. 262) at a meeting of the Linnean Society of New South Wales. The shell was of small size, not more than half an inch along the hinge-line; unfortunately the shell has been mislaid and its occurrence must remain solely as a record. Walkom (1913, p. 137) recorded Gonocarcliu1n australe in a list of fossils from the tipper part of the Branxton Stage, and also listed it as occurring in the Muree Stage. The genus Gonocardiuln has been recorded from several localities in Western Australia. Etheridge (1907, p. 33) recorded a small shell, differing from C. australe McCoy, from Permian beds of the Irwin River district. Teichert (1941, p. 377) records it from the Fossil Cliff limestone. Other records have been listed by Glauert (1910, p. 91) including GonocaTdiu1n f hibernicurn Sowerby, an unlikely species in Australian rocks, from the Kimberley district.
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