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8. Archosaur Phylogeny and the Relationships of the Crocodylia
8. Archosaur phylogeny and the relationships of the Crocodylia MICHAEL J. BENTON Department of Geology, The Queen's University of Belfast, Belfast, UK JAMES M. CLARK* Department of Anatomy, University of Chicago, Chicago, Illinois, USA Abstract The Archosauria include the living crocodilians and birds, as well as the fossil dinosaurs, pterosaurs, and basal 'thecodontians'. Cladograms of the basal archosaurs and of the crocodylomorphs are given in this paper. There are three primitive archosaur groups, the Proterosuchidae, the Erythrosuchidae, and the Proterochampsidae, which fall outside the crown-group (crocodilian line plus bird line), and these have been defined as plesions to a restricted Archosauria by Gauthier. The Early Triassic Euparkeria may also fall outside this crown-group, or it may lie on the bird line. The crown-group of archosaurs divides into the Ornithosuchia (the 'bird line': Orn- ithosuchidae, Lagosuchidae, Pterosauria, Dinosauria) and the Croco- dylotarsi nov. (the 'crocodilian line': Phytosauridae, Crocodylo- morpha, Stagonolepididae, Rauisuchidae, and Poposauridae). The latter three families may form a clade (Pseudosuchia s.str.), or the Poposauridae may pair off with Crocodylomorpha. The Crocodylomorpha includes all crocodilians, as well as crocodi- lian-like Triassic and Jurassic terrestrial forms. The Crocodyliformes include the traditional 'Protosuchia', 'Mesosuchia', and Eusuchia, and they are defined by a large number of synapomorphies, particularly of the braincase and occipital regions. The 'protosuchians' (mainly Early *Present address: Department of Zoology, Storer Hall, University of California, Davis, Cali- fornia, USA. The Phylogeny and Classification of the Tetrapods, Volume 1: Amphibians, Reptiles, Birds (ed. M.J. Benton), Systematics Association Special Volume 35A . pp. 295-338. Clarendon Press, Oxford, 1988. -
Re-Evaluation of the Genus Rubidgea in the Lower Permian of the Paraná Basin, Brazil, and Its Biostratigraphic Consequence
Versão online: http://www.lneg.pt/iedt/unidades/16/paginas/26/30/185 Comunicações Geológicas (2014) 101, Especial I, 455-457 IX CNG/2º CoGePLiP, Porto 2014 ISSN: 0873-948X; e-ISSN: 1647-581X Re-evaluation of the genus Rubidgea in the Lower Permian of the Paraná Basin, Brazil, and its biostratigraphic consequence Reavaliação do género Rubidgea, no Pérmico Inferior da Bacia do Paraná, Brasil, e suas consequências bioestratigráficas R. Iannuzzi1*, G. P. Tybusch1 Artigo Curto Short Article © 2014 LNEG – Laboratório Nacional de Geologia e Energia IP Abstract: The plant megafossil specimens revised in this study came *Corresponding author / Autor correspondente: [email protected] from the outcrops located in the São Paulo and Rio Grande do Sul states and positioned stratigraphically on top of the Itararé Group 1. Introduction and/or base of the Rio Bonito (= Tietê) Formation, Lower Permian of the Paraná Basin. The study material comprises leaves fragments The record of the genus Rubidgea in Brazil was first preserved as impressions, previously included in the morphogenus mentioned by Cazzulo-Klepzig et al. (1980) for the Itararé Rubidgea because of their supposed gangamopterid venation pattern, Group of the Rio Grande do Sul state, and it is represented lacking anastomoses. However, re-evaluation of this material showed by a specimen classified as Rubidgea sp. Later, Rubidgea that these leaves bear at some degree, rare anastomoses along the lamina. Based on this observation, we proposed that these leaves obovata Maithy (1965) and Rubidgea lanceolatus Maithy should be transferred to the morphogenus Gangamopteris, which (1965) were illustrated and reported for the outcrops displays this type of venation. -
O Esqueleto Pós-Craniano De Exaeretodon Riograndensis Abdala Et Al
Rev. bras. paleontol. 10(2):79-94, Maio/Agosto 2007 © 2007 by the Sociedade Brasileira de Paleontologia O ESQUELETO PÓS-CRANIANO DE EXAERETODON RIOGRANDENSIS ABDALA ET AL. (CYNODONTIA, TRAVERSODONTIDAE), TRIÁSSICO DO BRASIL TÉO VEIGA DE OLIVEIRA, CESAR LEANDRO SCHULTZ & MARINA BENTO SOARES Instituto de Geociências, UFRGS, Avenida Bento Gonçalves, 9500, 91501-970, Porto Alegre, RS, Brasil. [email protected], [email protected], [email protected] RESUMO – Pela primeira vez, elementos pós-cranianos de Exaeretodon riograndensis, um cinodonte traversodontídeo da Cenozona de Rhynchosauria da Formação Santa Maria, Neotriássico do sul do Brasil, são descritos e comparados com E. frenguellii e outros cinodontes não-mamalianos. O material inclui parte da coluna vertebral, radio, ulna e elementos da cintura pélvica. As diferenças mais significativas em relação a E. frenguellii estão na morfologia e nas dimensões do arco neural do atlas, na presença de intercentros em vértebras cervicais posteriores ao áxis e na ausência da dilatação do ápice do espinho neural das vértebras truncais. A morfologia da região sacral da coluna, do rádio, da ulna e da cintura pélvica são concordantes com o observado em E. frenguellii. A comparação com outros táxons de cinodontes não-mamalianos permite a observação da nítida evolução em mosaico do esqueleto pós-craniano destes animais: E. riograndensis mostra alguns caracteres similares a outros Cynognathia (grupo ao qual pertence), enquanto outros são mais próximos aos observados em alguns Probainognathia (incluindo mamíferos), como a natureza mais avançada do complexo atlas-áxis. Palavras-chave: Triássico, Brasil, Formação Santa Maria, Traversodontidae, Exaeretodon riograndensis, pós-crânio. ABSTRACT – THE POSTCRANIAL SKELETON OF EXAERETODON RIOGRANDENSIS ABDALA ET AL. -
Micheli Stefanello
UNIVERSIDADE FEDERAL DE SANTA MARIA CENTRO DE CIÊNCIAS NATURAIS E EXATAS PROGRAMA DE PÓS-GRADUAÇÃO EM BIODIVERSIDADE ANIMAL Micheli Stefanello DESCRIÇÃO E FILOGENIA DE UM NOVO ESPÉCIME DE CINODONTE PROBAINOGNÁTIO DO TRIÁSSICO SUL-BRASILEIRO Santa Maria, RS 2018 Micheli Stefanello DESCRIÇÃO E FILOGENIA DE UM NOVO ESPÉCIME DE CINODONTE PROBAINOGNÁTIO DO TRIÁSSICO SUL-BRASILEIRO Dissertação apresentada ao Curso de Mestrado do Programa de Pós-Graduação em Biodiversidade Animal, Área de Concentração em Sistemática e Biologia Evolutiva, da Universidade Federal de Santa Maria (UFSM, RS), como requisito parcial para obtenção do grau de Mestre em Ciências Biológicas – Área Biodiversidade Animal. Orientador: Prof. Dr. Sérgio Dias da Silva Santa Maria, RS 2018 Micheli Stefanello DESCRIÇÃO E FILOGENIA DE UM NOVO ESPÉCIME DE CINODONTE PROBAINOGNÁTIO DO TRIÁSSICO SUL-BRASILEIRO Dissertação apresentada ao Curso de Mestrado do Programa de Pós-Graduação em Biodiversidade Animal, Área de Concentração em Sistemática e Biologia Evolutiva, da Universidade Federal de Santa Maria (UFSM, RS), como requisito parcial para obtenção do grau de Mestre em Ciências Biológicas – Área Biodiversidade Animal. Aprovada em 28 de fevereiro de 2018: Santa Maria, RS 2018 AGRADECIMENTOS Ao meu orientador, Dr. Sérgio Dias da Silva, pela orientação, por todo o tempo despendido ao longo desse mestrado e por possibilitar meu aprimoramento na área a qual tenho apreço. Aos colegas do Centro de Apoio à Pesquisa Paleontológica da Quarta Colônia da Universidade Federal de Santa Maria (CAPPA/UFSM) e do Laboratório de Paleobiodiversidade Triássica, dessa mesma instituição, pela convivência e por terem ajudado-me de diferentes formas ao longo do mestrado. Em especial, ao colega Rodrigo Temp Müller, pela coleta do espécime (objeto de estudo dessa dissertação), por toda a ajuda com o TNT e com as figuras, e por auxiliar-me de inúmeras formas. -
Studies on Continental Late Triassic Tetrapod Biochronology. I. the Type Locality of Saturnalia Tupiniquim and the Faunal Succession in South Brazil
Journal of South American Earth Sciences 19 (2005) 205–218 www.elsevier.com/locate/jsames Studies on continental Late Triassic tetrapod biochronology. I. The type locality of Saturnalia tupiniquim and the faunal succession in south Brazil Max Cardoso Langer* Departamento de Biologia, FFCLRP, Universidade de Sa˜o Paulo (USP), Av. Bandeirantes 3900, 14040-901 Ribeira˜o Preto, SP, Brazil Received 1 November 2003; accepted 1 January 2005 Abstract Late Triassic deposits of the Parana´ Basin, Rio Grande do Sul, Brazil, encompass a single third-order, tetrapod-bearing sedimentary sequence that includes parts of the Alemoa Member (Santa Maria Formation) and the Caturrita Formation. A rich, diverse succession of terrestrial tetrapod communities is recorded in these sediments, which can be divided into at least three faunal associations. The stem- sauropodomorph Saturnalia tupiniquim was collected in the locality known as ‘Waldsanga’ near the city of Santa Maria. In that area, the deposits of the Alemoa Member yield the ‘Alemoa local fauna,’ which typifies the first association; includes the rhynchosaur Hyperodapedon, aetosaurs, and basal dinosaurs; and is coeval with the lower fauna of the Ischigualasto Formation, Bermejo Basin, NW Argentina. The second association is recorded in deposits of both the Alemoa Member and the Caturrita Formation, characterized by the rhynchosaur ‘Scaphonyx’ sulcognathus and the cynodont Exaeretodon, and correlated with the upper fauna of the Ischigualasto Formation. Various isolated outcrops of the Caturrita Formation yield tetrapod fossils that correspond to post-Ischigualastian faunas but might not belong to a single faunal association. The record of the dicynodont Jachaleria suggests correlations with the lower part of the Los Colorados Formation, NW Argentina, whereas remains of derived tritheledontid cynodonts indicate younger ages. -
The Anatomy of Asilisaurus Kongwe, a Dinosauriform from the Lifua
THE ANATOMICAL RECORD (2019) The Anatomy of Asilisaurus kongwe,a Dinosauriform from the Lifua Member of the Manda Beds (~Middle Triassic) of Africa 1 2 3 STERLING J. NESBITT , * MAX C. LANGER, AND MARTIN D. EZCURRA 1Department of Geosciences, Virginia Tech, Blacksburg, Virginia 2Departamento de Biologia, Universidade de Sao~ Paulo, Ribeirao~ Preto, Brazil 3Sección Paleontología de Vertebrados CONICET—Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Buenos Aires, Argentina ABSTRACT The diagnosis of Dinosauria and interrelationships of the earliest dino- saurs relies on careful documentation of the anatomy of their closest rela- tives. These close relatives, or dinosaur “precursors,” are typically only documented by a handful of fossils from across Pangea and nearly all speci- mens are typically missing important regions (e.g., forelimbs, pelves, skulls) that appear to be important to help resolving the relationships of dinosaurs. Here, we fully describe the known skeletal elements of Asilisaurus kongwe, a dinosauriform from the Middle Triassic Manda Beds of the Ruhuhu Basin of Tanzania. The taxon is known from many disarticulated and partially articulated remains and, most importantly, from a spectacularly preserved associated skeleton of an individual containing much of the skull, pectoral and pelvic girdles, forelimb and hindlimb, and parts of the vertebral column including much of the tail. The unprecedented detail of the anatomy indi- cates that Asilisaurus kongwe had a unique skull that was short and had both a premaxillary and dentary edentulous margin, but retained a number of character states plesiomorphic for Archosauria, including a crocodylian- like ankle configuration and a rather short foot with well-developed meta- tarsals I and V. -
University of Birmingham a New Dinosaur With
University of Birmingham A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil Marsola, Julio; Bittencourt, Jonathas; Butler, Richard; Da Rosa, Atila; Sayao, Juliana; Langer, Max DOI: 10.1080/02724634.2018.1531878 License: None: All rights reserved Document Version Peer reviewed version Citation for published version (Harvard): Marsola, J, Bittencourt, J, Butler, R, Da Rosa, A, Sayao, J & Langer, M 2019, 'A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil', Journal of Vertebrate Paleontology, vol. 38, no. 5, e1531878. https://doi.org/10.1080/02724634.2018.1531878 Link to publication on Research at Birmingham portal Publisher Rights Statement: Checked for eligibility: 25/07/2018 This is the accepted manuscript for a forthcoming publication in Journal of Vertebrate Paleontology. General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. -
Plant Fossils and Gondwana Flora
UNIT 12 PLANT FOSSILS AND GONDWANA FLORA Structure_____________________________________________________ 12.1 Introduction Vertebraria Expected Learning Outcomes Thinnfeldia 12.2 Plant Fossils Sigillaria Definition Nilssonia Classification Williamsonia Modes of Preservation Ptilophyllum Significance 12.5 Activity 12.3 Gondwana Flora of India 12.6 Summary 12.4 Descriptions of some Plant 12.7 Terminal Questions Fossils 12.8 References Glossopteris 12.9 Further/Suggested Readings Gangamopteris 12.10 Answers 12.1 INTRODUCTION The animals, plants and micro-organisms are the three main life forms surviving today. Even their fossilised remains are found in rocks that tell us about their past history. The animals comprise invertebrates and vertebrates. In Block 4, you will read about the invertebrates and their geological history that began in the latest Precambrian time. You also read about the microfossils in Unit 10 that too have a long geological record beginning from Precambrian onwards. In Unit 11, you read the evolutionary history of one of the vertebrate groups i.e., horse. In this unit, you will read the plant fossils and the Gondwana flora of India. Introduction to Palaeontology Block……………………………………………………………………………………………….….............….…........ 3 Like the kingdom Animalia, plants also form a separate kingdom known as the Plantae. It is thought that plants appeared first in the Precambrian, but their fossil record is poor. It is also proposed that earliest plants were aquatic and during the Ordovician period a transition from water to land took place that gave rise to non-vascular land plants. However, it was during the Silurian period, that the vascular plants appeared first on the land. The flowering plants emerged rather recently, during the Cretaceous period. -
Comparative Bone Microstructure of Three Archosauromorphs from the Carnian, Late Triassic Chañares Formation of Argentina
Comparative bone microstructure of three archosauromorphs from the Carnian, Late Triassic Chañares Formation of Argentina JORDI ALEXIS GARCIA MARSÀ, FEDERICO L. AGNOLÍN, and FERNANDO E. NOVAS Marsà, J.A.G., Agnolín, F.L., and Novas, F.E. 2020. Comparative bone microstructure of three archosauromorphs from the Carnian, Late Triassic Chañares Formation of Argentina. Acta Palaeontologica Polonica 65 (2): 387–398. The Chañares Formation exhibits one of the most important archosauriform records of early Carnian ecosystems. Here we present new data on the palaeohistology of Chañares archosauriforms and provide new insights into their paleobiology, as well as possible phylogenetically informative traits. Bone microstructure of Lagerpeton chanarensis and Tropidosuchus romeri is dominated by fibro-lamellar tissue and dense vascularization. On the other hand, Chanaresuchus bonapartei is more densely vascularized, but with cyclical growth characterized by alternate fibro-lamellar, parallel-fibered and lamellar-zonal tissues. Dense vascularization and fibro-lamellar tissue imply fast growth and high metabolic rates for all these taxa. These histological traits may be tentatively interpreted as a possible adaptative advantage in front of Chañares Formation environmental conditions. Key words: Archosauromorpha, Lagerpeton, Tropidosuchus, paleobiology, paleohistology, Mesozoic, South America. Jordi Alexis Garcia Marsà [[email protected]] and Fernando E. Novas [[email protected]], Labora- torio de Anatomía Comparada y Evolución de los Vertebrados, -
A Re-Evaluation of the Enigmatic Dinosauriform Caseosaurus Crosbyensis from the Late Triassic of Texas, USA and Its Implications for Early Dinosaur Evolution
A re-evaluation of the enigmatic dinosauriform Caseosaurus crosbyensis from the Late Triassic of Texas, USA and its implications for early dinosaur evolution MATTHEW G. BARON and MEGAN E. WILLIAMS Baron, M.G. and Williams, M.E. 2018. A re-evaluation of the enigmatic dinosauriform Caseosaurus crosbyensis from the Late Triassic of Texas, USA and its implications for early dinosaur evolution. Acta Palaeontologica Polonica 63 (1): 129–145. The holotype specimen of the Late Triassic dinosauriform Caseosaurus crosbyensis is redescribed and evaluated phylogenetically for the first time, providing new anatomical information and data on the earliest dinosaurs and their evolution within the dinosauromorph lineage. Historically, Caseosaurus crosbyensis has been considered to represent an early saurischian dinosaur, and often a herrerasaur. More recent work on Triassic dinosaurs has cast doubt over its supposed dinosaurian affinities and uncertainty about particular features in the holotype and only known specimen has led to the species being regarded as a dinosauriform of indeterminate position. Here, we present a new diagnosis for Caseosaurus crosbyensis and refer additional material to the taxon—a partial right ilium from Snyder Quarry. Our com- parisons and phylogenetic analyses suggest that Caseosaurus crosbyensis belongs in a clade with herrerasaurs and that this clade is the sister taxon of Dinosauria, rather than positioned within it. This result, along with other recent analyses of early dinosaurs, pulls apart what remains of the “traditional” group of dinosaurs collectively termed saurischians into a polyphyletic assemblage and implies that Dinosauria should be regarded as composed exclusively of Ornithoscelida (Ornithischia + Theropoda) and Sauropodomorpha. In addition, our analysis recovers the enigmatic European taxon Saltopus elginensis among herrerasaurs for the first time. -
9 Paleontological Conference Th
Polish Academy of Sciences Institute of Paleobiology 9th Paleontological Conference Warszawa, 10–11 October 2008 Abstracts Warszawa Praha Bratislava Edited by Andrzej Pisera, Maria Aleksandra Bitner and Adam T. Halamski Honorary Committee Prof. Oldrich Fatka, Charles University of Prague, Prague Prof. Josef Michalík, Slovak Academy of Sciences, Bratislava Assoc. Prof. Jerzy Nawrocki, Polish Geological Institute, Warszawa Prof. Tadeusz Peryt, Polish Geological Institute, Warszawa Prof. Grzegorz Racki, Institute of Paleobiology, Warszawa Prof. Jerzy Trammer, University of Warsaw, Warszawa Prof. Alfred Uchman, Jagiellonian University, Kraków Martyna Wojciechowska, National Geographic Polska, Warszawa Organizing Committee Dr Maria Aleksandra Bitner (Secretary), Błażej Błażejewski, MSc, Prof. Andrzej Gaździcki, Dr Adam T. Halamski, Assoc. Prof. Anna Kozłowska, Assoc. Prof. Andrzej Pisera Sponsors Institute of Paleobiology, Warszawa Polish Geological Institute, Warszawa National Geographic Polska, Warszawa Precoptic Co., Warszawa Cover picture: Quenstedtoceras henrici Douvillé, 1912 Cover designed by Aleksandra Hołda−Michalska Copyright © Instytut Paleobiologii PAN Nakład 150 egz. Typesetting and Layout: Aleksandra Szmielew Warszawska Drukarnia Naukowa PAN ABSTRACTS Paleotemperature and paleodiet reconstruction on the base of oxygen and carbon isotopes from mammoth tusk dentine and horse teeth enamel during Late Paleolith and Mesolith MARTINA ÁBELOVÁ State Geological Institute of Dionýz Štúr, Mlynská dolina 1, SK−817 04 Bratislava 11, Slovak Republic; [email protected] The use of stable isotopes has proven to be one of the most effective methods in re− constructing paleoenvironments and paleodiet through the upper Pleistocene period (e.g. Fricke et al. 1998; Genoni et al. 1998; Bocherens 2003). This study demonstrates how isotopic data can be employed alongside other forms of evidence to inform on past at great time depths, making it especially relevant to the Palaeolithic where there is a wealth of material potentially available for study. -
Macropredatory Ichthyosaur from the Middle Triassic and the Origin of Modern Trophic Networks
Macropredatory ichthyosaur from the Middle Triassic and the origin of modern trophic networks Nadia B. Fröbischa,1, Jörg Fröbischa,1, P. Martin Sanderb,1,2, Lars Schmitzc,1,2,3, and Olivier Rieppeld aMuseum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung an der Humboldt-Universität zu Berlin, 10115 Berlin, Germany; bSteinmann Institute of Geology, Mineralogy, and Paleontology, Division of Paleontology, University of Bonn, 53115 Bonn, Germany; cDepartment of Evolution and Ecology, University of California, Davis, CA 95616; and dDepartment of Geology, The Field Museum of Natural History, Chicago, IL 60605 Edited by Neil H. Shubin, The University of Chicago, Chicago, IL, and approved December 5, 2012 (received for review October 8, 2012) The biotic recovery from Earth’s most severe extinction event at the Holotype and Only Specimen. The Field Museum of Natural His- Permian-Triassic boundary largely reestablished the preextinction tory (FMNH) contains specimen PR 3032, a partial skeleton structure of marine trophic networks, with marine reptiles assuming including most of the skull (Fig. 1) and axial skeleton, parts of the predator roles. However, the highest trophic level of today’s the pelvic girdle, and parts of the hind fins. marine ecosystems, i.e., macropredatory tetrapods that forage on prey of similar size to their own, was thus far lacking in the Paleozoic Horizon and Locality. FMNH PR 3032 was collected in 2008 from the and early Mesozoic. Here we report a top-tier tetrapod predator, middle Anisian Taylori Zone of the Fossil Hill Member of the Favret a very large (>8.6 m) ichthyosaur from the early Middle Triassic Formation at Favret Canyon, Augusta Mountains, Pershing County, (244 Ma), of Nevada.