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T.C. BARTIN ÜNİVERSİTESİ FEN BİLİMLERİ ENSTİTÜSÜ ORMAN MÜHENDİSLİĞİ ANABİLİM DALI BARTIN İLİ CARABIDAE (COLEOPTERA) TÜRLERİ YÜKSEK LİSANS TEZİ HAZIRLAYAN SEMA KARA DANIŞMAN PROF. DR. AZİZE TOPER KAYGIN BARTIN-2016 T.C BARTIN ÜNĠVERSĠTESĠ FEN BĠLĠMLERĠ ENSTĠTÜSÜ ORMAN MÜHENDĠSLĠĞĠ ANABĠLĠM DALI BARTIN ĠLĠ CARABIDAE (COLEOPTERA) TÜRLERĠ YÜKSEK LĠSANS TEZĠ HAZIRLAYAN Sema KARA DANIġMAN Prof. Dr. Azize TOPER KAYGIN JÜRĠ ÜYELERĠ Danışman : Prof. Dr. Azize TOPER KAYGIN - Bartın Üniversitesi Üye : Prof. Dr. Erol AKKUZU - Kastamonu Üniversitesi Üye : Yrd. Doç. Dr. Nuri Kaan ÖZKAZANÇ - Bartın Üniversitesi BARTIN-2016 KABUL VE ONAY Sema KARA tarafından hazırlanan “BARTIN İLİ CARABIDAE (COLEOPTERA) TÜRLERİ” başlıklı bu çalışma, 23.05.2016 tarihinde yapılan savunma sınavı sonucunda oy birliği ile başarılı bulunarak jürimiz tarafından Yüksek Lisans Tezi olarak kabul edilmiştir. Başkan : Prof. Dr. Azize TOPER KAYGIN (Danışman) Üye : Prof. Dr. Erol AKKUZU Üye : Yrd. Doç. Dr. Nuri Kaan ÖZKAZANÇ Bu tezin kabulü Fen Bilimleri Enstitüsü Yönetim Kurulu‟nun ..…/..…/20… tarih ve 20…../…..-….. sayılı kararıyla onaylanmıştır. Doç. Dr. H. Selma ÇELİKYAY Fen Bilimleri Enstitüsü Müdürü ii BEYANNAME Bartın Üniversitesi Fen Bilimleri Enstitüsü tez yazım kılavuzuna göre, Prof. Dr. Azize TOPER KAYGIN danışmanlığında hazırlamış olduğum "BARTIN İLİ CARABIDAE (COLEOPTERA) TÜRLERİ" başlıklı yüksek lisans tezimin bilimsel etik değerlere ve kurallara uygun, özgün bir çalışma olduğunu, aksinin tespit edilmesi halinde her türlü yasal yaptırımı kabul edeceğimi beyan ederim. 23/05/2016 Sema KARA iii ÖNSÖZ Yüksek lisans öğrenimim boyunca desteğini ve güvenini esirgemeyen, her aşamada bilgi birikimi ile ufkumu açan danışman hocam Prof. Dr. Sayın Azize TOPER KAYGIN'a teşekkür ederim. Tez çalışmalarım sırasında destek ve yardımlarını gördüğüm, tezin fotoğraf çekimlerinde bana yardımcı olan değerli hocam Yrd. Doç. Dr. Sayın Nuri Kaan ÖZKAZANÇ'a; bilgi birikimlerinden yararlandığım değerli hocalarım; Prof. Dr. Sayın İsmet DAŞDEMİR‟e, Yrd. Doç. Dr. Sayın Yafes YILDIZ‟a, Yrd. Doç. Dr. Sayın Ersin GÜNGÖR‟e teşekkür ederim. Carabus (Lamprostus) spinolai, Carabus (Procrustes) chevrolati, Carabus (Oxycarabus) saphyrinus, Carabus (Archicarabus) wiedemanni wiedemanni, Poecilus (Poecilus) cupreus, Harpalus (Harpalus) caspius, Parophonus (Parophonus) dia; türlerinin teşhislerini ve teşhisi yapılan Carabus (Procerus) scabrosus, Harpalus (Harpalus) serripes serripes, Ophonus (Hesperophonus) azureus, Anchomenus (Anchomenus) dorsalis, Psedoophonus (Psedoophonus) griseus, Psedoophonus (Psedoophonus) rufipes türlerinin kontrolunu yapan Doç. Dr. Sayın Sakine Serap AVGIN‟a (Kahramanmaraş Sütçü İmam Üniversitesi Eğitim Fakültesi) teşekkür ederim. Tür teşhis kaynakları konusunda yardımcı olan ve tezimdeki bütün türlerin teşhislerinde ve kontrolünde hoşgörüyle yardımcı olan, engin bilgi ve deneyimlerinden faydalandığım Yrd. Doç. Dr. Sayın Memiş KESDEK‟e (Muğla Sıtkı Koçman Üniversitesi, Fethiye Ali Sıtkı Mefharet Koçman Meslek Yüksekokulu) teşekkür ederim. Carabus (Procerus) scabrosus, Anchomenus (Anchomenus) dorsalis, Psedoophonus (Psedoophonus) griseus, Psedoophonus (Psedoophonus) rufipes, Paradromius (Manodromius) linearis türlerinin teşhislerinin yapılmasında ve teşhis kaynaklarının temin edilmesinde yardımcı olan Arş. Gör. Sayın Ebru Ceren FİDAN‟a (Eskişehir Osmangazi Üniversitesi Fen Edebiyat Fakültesi) teşekkür ederim. Arazi çalışmalarımda araç desteği sağlayan Bartın Orman İşletme Müdürlüğüne, arazi çalışmalarımın düzenli şekilde yapılmasını sağlayan eski Yenihan Şefi Sayın Oğuz Benkul BOLLUKÇU‟ya, Arıt İşletme Şefi Sayın Ali DUYMUŞ‟a, Kozcağız İşletme Şefi Sayın Ramazan ONUR‟a, Amasra İşletme Şefi Sayın Erkan Sade‟ye, Kurucaşile İşletme Şefi Sayın İsmail ŞEN‟e ve Merkez İşletme Şefi Sayın Mehmet ÖZDEMİR‟e teşekkür ederim. Arazi çalışmalarında bana yardımcı olan, bu bölümde yüksek lisans yapmamı öneren ve iv destekleyen babam Yılmaz KARA‟ya, eniştem Sadık ARSLAN‟a, ablam Esra ARSLAN‟a; örnek toplamamda yardımcı olan dünyalar tatlısı yeğenim Yusuf Efe ARSLAN‟a; hayatımın her aşamasında yanımda olan biricik annem Zülbiye KARA‟ya; canım kardeşim Hüseyin KARA‟ya, teşekkür ederim. Manevi desteğini benden esirgemeyen değerli arkadaşlarıma teşekkür ederim. Sema KARA v ÖZET Yüksek Lisans Tezi BARTIN ĠLĠ CARABIDAE (COLEOPTERA) TÜRLERĠ Sema KARA Bartın Üniversitesi Fen Bilimleri Enstitüsü Orman Mühendisliği Anabilim Dalı Orman Entomolojisi ve Koruma Bilim Dalı Tez DanıĢmanı: Prof. Dr. Azize TOPER KAYGIN Bartın - 2016, sayfa: xviii + 165 2014-2016 yılları arasında gerçekleştirilen bu çalışma ile Bartın ilinde yayılış gösteren Carabidae familyasına ait türlerin belirlenmesi amaçlanmıştır. Arazi çalışmalarında çukur tuzak yöntemi kullanılmış serbest dolaşan türler ise elle toplanmıştır. Bartın ilinin her bir ilçesinde (Merkez, Amasra, Ulus, Kurucaşile) farklı habitatlar tercih edilerek, çukur tuzaklar kurulmuştur. Çukur tuzak yöntemine ek olarak kış döneminde de Carabidae faunasına ait veriler elde etmek için kışın böceklerin saklanabileceği kışlak tuzaklar kurulmuştur. Fakat bu yöntemden başarılı sonuç alınamamıştır. Çukur tuzaklarda yakalanan ve serbest dolaşan böcekler toplanıp laboratuvar ortamında preparasyonları yapılmıştır. Preparasyonu yapılan türler toplandıkları tarih, mevkii gibi arazi bilgilerinin bulunduğu etiketler ile koleksiyon kutularına yerleştirilmiştir. Tür teşhisleri yapılarak, her bir tür fotoğraflanmıştır. Ayrıca Carabus türlerinin erginlerinin beslenmesi, çiftleşmesi ve yumurta bırakma döneminin bir kısmı gözlenmiştir. Bu çalışmada Carabidae familyasından 419 adet ergin ve 41 adet larva incelenmiştir Çalışmanın sonucunda Bartın ilinde 10 alt familyaya ait 26 cins, 51 tür ve alttür tespit vi edilmiş olup 2 tür Türkiye faunası için yeni kayıt özelliğindedir. Bartın ilinde Carabidae türlerinin sistematiği çıkarılmıştır. Ayrıca bu çalışma Bartın ilinde Carabidae familyasına ait olan türlerin araştırılmasına yönelik ilk çalışma niteliğinde olup, tespit edilen türler de Bartın için ilk kayıt olma özelliğini taşımaktadır. Anahtar Kelimeler Bartın; Carabidae; Çukur Tuzak; Kışlak Tuzak; Predatör. Bilim Kodu 502.02.01 vii ABSTRACT M. Sc. Thesis CARABIDAE (COLEOPTERA) SPECIES IN BARTIN PROVINCE Sema KARA Bartın University Graudate School of Natural and Applied Sciences Forest Engineering Science of Forest Entomology and Protection Thesis Advisor: Prof. Dr. Azize TOPER KAYGIN Bartın - 2016, pp: xviii + 165 The purpose of this study, which was carried out in 2014-2016, was to determine the species belonging to Carabidae family and expanding in Bartın province. Pitfall trap method was used in the field work and those free species were collected by hand. Pitfall traps were set in different habitats in each district of Bartın province (Center, Amasra, Ulus, Kurucaşile). In addition to pitfall traps, winter traps were set to collect data on Carabidae fauna during the winter time. However, this method wasn‟t successful. The insects that fell into the pitfall trap or move freely were collected, and then the samples were prepared in the laboratory. With stickers showing the collection date and place, they were put into collection boxes. The species were identified and each species was photographed. In addition, the eating, copulating and a part of egg laying periods of Carabus species adults were observed. In this study, totally 419 adults and 41 larvae of Carabidae family were examined. At the and of the study, 26 genera, 51 species and subspecies of 10 subfamilies belonging to Carabidae family in the province of Bartın were identified; 2 species are new record for Turkey‟s fauna. Carabidae species of Bartın province with this systematic detection has viii been exerted. Moreover, it is the first study on identifying the species of Carabidae family in the city. The identified species have been the first records for the area. Key Words Bartın; Carabidae; pitfall trap; Predator; winter trap. Science Code 502.02.01 ix ĠÇĠNDEKĠLER Sayfa KABUL VE ONAY .............................................................................................................. ii BEYANNAME ..................................................................................................................... iii ÖN SÖZ ................................................................................................................................ iv ÖZET .................................................................................................................................... vi ABSTRACT ..................................................................................................................... viii İÇİNDEKİLER ...................................................................................................................... x ŞEKİLLER DİZİNİ ............................................................................................................ xiii TABLOLAR DİZİNİ ......................................................................................................... xvii SİMGELER VE KISALTMALAR DİZİNİ ..................................................................... xviii BÖLÜM I GİRİŞ ................................................................................................................... 1 1.1 Genel Bilgiler .............................................................................................................. 1 1.2 Carabidae Familyasının Önemi ................................................................................... 2 1.3 Carabidae Familyasının Morfolojisi
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    © Biologiezentrum Linz/Austria; download unter www.biologiezentrum.at Linzer biol. Beitr. 41/1 901-935 30.8.2009 New or interesting records of Carabid beetles from Europe, Madeira, northern Africa, Turkey, from the Near East, Iran, Iraq, Kuwait, and Pakistan, with nomenclatorial and taxonomic notes (Coleoptera, Carabidae, Bembidiini, Brachinini, Cyclosomini, Elaphrini, Harpalini, Lebiini, Nebriini, Platynini, Pterostichini, Scaritini, Sphodrini, Zabrini) D.W. WRASE A b s t r a c t : A new assignment is proposed: Harpalus honestus ssp. creticus MAŘAN 1934 = Harpalus (s.str.) sulphuripes ssp. creticus MAŘAN 1934. Change of rank: Acupalpus (s.str.) schnitteri JAEGER 1999, nov. status is upgraded from: Acupalpus (s.str.) suturalis ssp. schnitteri JAEGER 1999. Following species are recorded for the first time: Leistus (s.str.) terminatus (PANZER 1793): Iceland (Austurland; Suðurland). Brachinus (s.str.) alexandri F. BATTONI 1984: Israel (Golan; Central District; North District). Elaphrus (s.str.) weissi DOSTAL 1996: Turkey (Ankara). Israel (Golan). Asaphidion austriacum SCHWEIGER 1975: Iran (Golestan). A. curtum ssp. curtum HEYDEN 1870: Tunisia (Béja; Jendouba). A. festivum (JACQUELIN DU VAL 1851): Greece (Macedonía: Flórina/Péla). A. transcaspicum (SEMENOV) 1889: Iran (Golestan; Khorāsān). Bembidion (Trepanes) articulatum (PANZER 1796): Turkey (Samsun). B. (Trepanes) maculatum spp. maculatum DEJEAN 1831: Israel (Golan). Sphaerotachys fumicatus (MOTSCHULSKY 1851): Israel (North District). Paratachys fasciatus spp. fasciatus (MOTSCHULSKY 1851): Israel (North District). Tachyura (s.str.) euphratica (REITTER 1885): Israel (North District). T. (s.str.) poeciloptera (BATES 1873): Turkey (Antalya). T. (s.str.) walkeriana (SHARP 1913): Greece (Fthiótida; Samos). Israel (Golan; North District). Pogonus (s.str.) orientalis ssp. orientalis DEJEAN 1828: Romania (Tulcea). Trechus (Epaphius) secalis ssp. secalis (PAYKULL 1790): Iceland (Austurland; Suðurland).
  • Potential Impact of Diabrotica Resistant Bt-Maize Expressing Cry3bb1 on Ground Beetles (Coleoptera: Carabidae)

    Potential Impact of Diabrotica Resistant Bt-Maize Expressing Cry3bb1 on Ground Beetles (Coleoptera: Carabidae)

    Potential impact of Diabrotica resistant Bt-maize expressing Cry3Bb1 on ground beetles (Coleoptera: Carabidae) Von der Fakultät für Mathematik, Informatik und Naturwissenschaften der RWTH Aachen University zur Erlangung des akademischen Grades eines Doktors der Naturwissenschaften genehmigte Dissertation vorgelegt von Diplom-Biologe Kai Uwe Priesnitz aus Iserlohn Berichter: Universitätsprofessor Dr. Alan Slusarenko Universitätsprofessor Dr. Ingolf Schuphan Tag der mündlichen Prüfung: 17.12.2010 Diese Dissertation ist auf den Internetseiten der Hochschulbibliothek online verfügbar. Contents 1 Introduction ....................................................................................1 2 General background ......................................................................4 2.1 Ecology and biology of maize and its role in agriculture .......................... 4 2.1.1 Botanical characteristics and systematics of Zea mays ......................................4 2.1.2 Domestication of Maize .......................................................................................5 2.1.3 Maize as genetically modified crop......................................................................8 2.1.4 Event MON 88017: A herbicide tolerant Bt-maize protected against Diabrotica virgifera virgifera..................................................................................................9 2.2 Diabrotica virgifera virgifera: A major pest in maize............................... 11 2.3 Ground beetles: Widespread epigeal arthropods on arable
  • Biodiversity and Geodiversity Background Paper

    Biodiversity and Geodiversity Background Paper

    Biodiversity and Geodiversity Background Paper CONTENTS 1 INTRODUCTION 5 1.1 Purpose 5 1.2 What Is Biodiversity 5 1.3 What Is Geodiversity 6 2 DESIGNATIONS RELEVANT TO NUNEATON AND BEDWORTH 7 2.1 Natura 2000 Site Network 7 2.2 Special Areas of Conservation 8 2.3 Special Sites of Scientific Interest 8 2.4 Local Nature Reserves 8 2.5 Local Geological Sites 8 2.6 Local Wildlife Sites 8 2.7 Priority Habitats and Species 8 2.8 Ancient Woodlands 9 2.9 Veteran Trees 10 3 INTERNATIONAL LEGISLATION 10 3.1 The Convention on the Conservation of European Wildlife 10 and Natural Habitats (the Bern Convention) 3.2 Conservation (Natural Habitats, etc) Regulations 1994 10 (regulation 38). 3.3 Directive 2009/147/EC (the Birds Directive), as amended 11 3.4 Directive 92/43/EEC (the Habitats Directive) 11 4 NATIONAL LEGISLATION 11 4.1 Natural Environment and Rural Communities (NERC) Act 11 2006 4.2 Wildlife and Countryside Act 1981, as amended 12 4.3 The Hedgerow Regulations 12 4.4 Natural Choice: Securing the Value of Nature 13 4.4.1 Local Nature Partnerships 14 4.4.2 Biodiversity Offsetting 14 4.4.2.1 Mitigation Hierarchy 15 4.5 National Planning Policy Framework 15 4.6 Local Sites: Guidance on their Identification, Selection and 16 Management 4.7 Keepers of Time: A Statement of Policy for England’s 16 Ancient Woodland 4.8 Geological Conservation: A Good Practice Guide 16 5 REGIONAL STRATEGIES / POLICIES 16 5.1 Enhancing Biodiversity Across the West Midlands 16 2 6 SUB-REGIONAL STRATEGIES / POLICIES 17 6.1 Warwickshire Geodiversity Action Plan 17 6.2 Warwickshire,

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    View metadata, citation and similar papers at core.ac.uk brought to you by CORE provided by ZENODO A peer-reviewed open-access journal ZooKeys 100:Recent 475–485 records (2011) of steppe species in Belarus, first indications of a steppe species invasion? 475 doi: 10.3897/zookeys.100.1541 RESEARCH ARTICLE www.zookeys.org Launched to accelerate biodiversity research Recent records of steppe species in Belarus, first indications of a steppe species invasion? Oleg Aleksandrowicz Pomeranian University, Slupsk, Poland Corresponding author: Oleg Aleksandrowicz ([email protected]) Academic editor: R. Vermeulen | Received 1 December 2009 | Accepted 21 January 2011 | Published 20 May 2011 Citation: Aleksandrowicz O (2011) Recent records of steppe species in Belarus, first indications of a steppe species invasion? In: Kotze DJ, Assmann T, Noordijk J, Turin H, Vermeulen R (Eds) Carabid Beetles as Bioindicators: Biogeographical, Ecological and Environmental Studies. ZooKeys 100: 475–485. doi: 10.3897/zookeys.100.1541 Abstract Belarus is situated at a crossroad of natural borders of species distributions: the NE part is situated in a taiga zone, whereas the other part of terrain is in the European forest zone. The distance of Belarus to the steppe zone is about 330 kilometers. This geographical position and the extensive knowledge of its fauna can be used to monitor changes in the distribution of different species. An intensive study of open habitat ground beetles was carried out from 1975–2008 in Belarus, using pitfall traps, quadrate- sampling methods, hand collecting, netting and light traps. In total, more than 130 000 specimens of ground beetles belonging to 169 species were collected from 62 fields and 11 meadows of different types.
  • The Ground Beetles (Coleoptera: Carabidae) of the Bulgarian Black Sea Coast

    The Ground Beetles (Coleoptera: Carabidae) of the Bulgarian Black Sea Coast

    370 Bulgarian Journal of Agricultural Science, 18 (No 3) 2012, 370-386 Agricultural Academy THE GROUND BEETLES (COLEOPTERA: CARABIDAE) OF THE BULGARIAN BLACK SEA COAST T. TEOFILOVA1, E. MARKOVA1 and N. KODZHABASHEV2 1 Sofia University, Faculty of Biology, Department of Ecology and Environmental Protection, BG - 1164 Sofia, Bulgaria 2University of Forestry, Faculty of Forestry, Department of Hunting and Game Management, BG - 1756 Sofia, Bulgaria Abstract TEOFILOVA, T., E. MARKOVA and N. KODZHABASHEV, 2012. The ground beetles (Coleoptera: Carabidae) of the Bulgarian Black Sea coast. Bulg. J. Agric. Sci., 18: 370-386 The publication represents the first complete overview of the established in the area of the Bulgarian Black Sea coast spe- cies from the Carabidae family. Full checklist of the all 465 species is given. Species of ground beetles are characterized and classified by their zoogeographical belonging, degree of endemism, habitat preferences, and life form they refer to Carabid’s subsuming to the subdivisions Northern or Southern Black Sea coast is pointed. Key words: Carabidae, ground beetles, Black Sea coast Introduction occurring in the area of the Bulgarian Black Sea coast 98 species were described, and for 5 more it was consid- Combination of various environmental factors con- ered that they would probably be found there. Data for tributed to the definition of the Black Sea coast as a de- the species of the researched zoogeographical region tached zoogeographical region (Gruev and Kuzmanov, was found also in the revision of Rambousek (1912). 1994) and in terms of wildlife, it could be claimed that The second period of studies included the time until ground beetles are convenient and expedient group for around 1950 and was characterized by more intensive monitoring and bioindication researches (Desender and and detailed faunistic researches.