Phasmid Study Group Newsletter, 28
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Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea
Brigham Young University BYU ScholarsArchive Theses and Dissertations 2018-07-01 Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco Brigham Young University Follow this and additional works at: https://scholarsarchive.byu.edu/etd Part of the Life Sciences Commons BYU ScholarsArchive Citation Pacheco, Yelena Marlese, "Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea" (2018). Theses and Dissertations. 7444. https://scholarsarchive.byu.edu/etd/7444 This Thesis is brought to you for free and open access by BYU ScholarsArchive. It has been accepted for inclusion in Theses and Dissertations by an authorized administrator of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco A thesis submitted to the faculty of Brigham Young University in partial fulfillment of the requirements for the degree of Master of Science Michael F. Whiting, Chair Sven Bradler Seth M. Bybee Steven D. Leavitt Department of Biology Brigham Young University Copyright © 2018 Yelena Marlese Pacheco All Rights Reserved ABSTRACT Ecomorph Convergence in Stick Insects (Phasmatodea) with Emphasis on the Lonchodinae of Papua New Guinea Yelena Marlese Pacheco Department of Biology, BYU Master of Science Phasmatodea exhibit a variety of cryptic ecomorphs associated with various microhabitats. Multiple ecomorphs are present in the stick insect fauna from Papua New Guinea, including the tree lobster, spiny, and long slender forms. While ecomorphs have long been recognized in phasmids, there has yet to be an attempt to objectively define and study the evolution of these ecomorphs. -
Insecta: Phasmatodea) and Their Phylogeny
insects Article Three Complete Mitochondrial Genomes of Orestes guangxiensis, Peruphasma schultei, and Phryganistria guangxiensis (Insecta: Phasmatodea) and Their Phylogeny Ke-Ke Xu 1, Qing-Ping Chen 1, Sam Pedro Galilee Ayivi 1 , Jia-Yin Guan 1, Kenneth B. Storey 2, Dan-Na Yu 1,3 and Jia-Yong Zhang 1,3,* 1 College of Chemistry and Life Science, Zhejiang Normal University, Jinhua 321004, China; [email protected] (K.-K.X.); [email protected] (Q.-P.C.); [email protected] (S.P.G.A.); [email protected] (J.-Y.G.); [email protected] (D.-N.Y.) 2 Department of Biology, Carleton University, Ottawa, ON K1S 5B6, Canada; [email protected] 3 Key Lab of Wildlife Biotechnology, Conservation and Utilization of Zhejiang Province, Zhejiang Normal University, Jinhua 321004, China * Correspondence: [email protected] or [email protected] Simple Summary: Twenty-seven complete mitochondrial genomes of Phasmatodea have been published in the NCBI. To shed light on the intra-ordinal and inter-ordinal relationships among Phas- matodea, more mitochondrial genomes of stick insects are used to explore mitogenome structures and clarify the disputes regarding the phylogenetic relationships among Phasmatodea. We sequence and annotate the first acquired complete mitochondrial genome from the family Pseudophasmati- dae (Peruphasma schultei), the first reported mitochondrial genome from the genus Phryganistria Citation: Xu, K.-K.; Chen, Q.-P.; Ayivi, of Phasmatidae (P. guangxiensis), and the complete mitochondrial genome of Orestes guangxiensis S.P.G.; Guan, J.-Y.; Storey, K.B.; Yu, belonging to the family Heteropterygidae. We analyze the gene composition and the structure D.-N.; Zhang, J.-Y. -
PROCEEDINGS of the WORKSHOP on TRADE and CONSERVATION of PANGOLINS NATIVE to SOUTH and SOUTHEAST ASIA 30 June – 2 July 2008, Singapore Zoo Edited by S
PROCEEDINGS OF THE WORKSHOP ON TRADE AND CONSERVATION OF PANGOLINS NATIVE TO SOUTH AND SOUTHEAST ASIA 30 June – 2 July 2008, Singapore Zoo Edited by S. Pantel and S.Y. Chin Wildlife Reserves Singapore Group PROCEEDINGS OF THE WORKSHOP ON TRADE AND CONSERVATION OF PANGOLINS NATIVE TO SOUTH AND SOUTHEAST ASIA 30 JUNE –2JULY 2008, SINGAPORE ZOO EDITED BY S. PANTEL AND S. Y. CHIN 1 Published by TRAFFIC Southeast Asia, Petaling Jaya, Selangor, Malaysia © 2009 TRAFFIC Southeast Asia All rights reserved. All material appearing in these proceedings is copyrighted and may be reproduced with permission. Any reproduction, in full or in part, of this publication must credit TRAFFIC Southeast Asia as the copyright owner. The views of the authors expressed in these proceedings do not necessarily reflect those of the TRAFFIC Network, WWF or IUCN. The designations of geographical entities in this publication, and the presentation of the material, do not imply the expression of any opinion whatsoever on the part of TRAFFIC or its supporting organizations concerning the legal status of any country, territory, or area, or its authorities, or concerning the delimitation of its frontiers or boundaries. The TRAFFIC symbol copyright and Registered Trademark ownership is held by WWF. TRAFFIC is a joint programme of WWF and IUCN. Layout by Sandrine Pantel, TRAFFIC Southeast Asia Suggested citation: Sandrine Pantel and Chin Sing Yun (ed.). 2009. Proceedings of the Workshop on Trade and Conservation of Pangolins Native to South and Southeast Asia, 30 June-2 July -
The Culture of Bornean Phasmids. P.E
The culture of Bornean phasmids. P.E. Bragg, 51 Longfield Lane, Dkeston, Derbyshire, DE7 4DX, UK. Key words Phasmida, Culture potential, Borneo, Taxonomic assessment, Polyphagy, Monophagy Introduction At the PSG meeting at the Natural History Museum, London, on 22Dd 1995, I gave an illustrated lecture on the species of Bornean phasmids which have been reared in captivity. The talk stressed that the "culture potential" i.e. the likelihood of establishing a culture, of a given species appears to depend upon the flight capability of the species. There is a close correlation between culture potential and taxonomic position; this is not unexpected as flight capability is generally closely linked to the taxonomic position of phasmids. This article summarises the lecture. Since December 1987 I have visited Borneo on 7 occasions, spending a total of about six months on the island. Borneo is the third largest island in the world and is composed of four political units: Brunei (an independent country), Kalimantan (part of Indonesia), Sarawak (a State of Malaysia) and Sabah (also part of Malaysia). Most of my collecting has been done in Sarawak, although I spent five weeks in Kalimantan, three in Brunei, and two in Sabah. My first visit to Borneb was a two week holiday with my wife and Lee Yong Tsui in December 1987 and January 1988, during this time we stayed with Dr Lee's family in Sarawak. Although not a collecting trip, seven species were collected over two nights and five of these were bred in the UK. All subsequent trips have been made specifically to collect phasmids and, whenever possible, to attempt to rear them in the UK. -
Insect Egg Size and Shape Evolve with Ecology but Not Developmental Rate Samuel H
ARTICLE https://doi.org/10.1038/s41586-019-1302-4 Insect egg size and shape evolve with ecology but not developmental rate Samuel H. Church1,4*, Seth Donoughe1,3,4, Bruno A. S. de Medeiros1 & Cassandra G. Extavour1,2* Over the course of evolution, organism size has diversified markedly. Changes in size are thought to have occurred because of developmental, morphological and/or ecological pressures. To perform phylogenetic tests of the potential effects of these pressures, here we generated a dataset of more than ten thousand descriptions of insect eggs, and combined these with genetic and life-history datasets. We show that, across eight orders of magnitude of variation in egg volume, the relationship between size and shape itself evolves, such that previously predicted global patterns of scaling do not adequately explain the diversity in egg shapes. We show that egg size is not correlated with developmental rate and that, for many insects, egg size is not correlated with adult body size. Instead, we find that the evolution of parasitoidism and aquatic oviposition help to explain the diversification in the size and shape of insect eggs. Our study suggests that where eggs are laid, rather than universal allometric constants, underlies the evolution of insect egg size and shape. Size is a fundamental factor in many biological processes. The size of an 526 families and every currently described extant hexapod order24 organism may affect interactions both with other organisms and with (Fig. 1a and Supplementary Fig. 1). We combined this dataset with the environment1,2, it scales with features of morphology and physi- backbone hexapod phylogenies25,26 that we enriched to include taxa ology3, and larger animals often have higher fitness4. -
Kataloge Der Wissenschaftlichen Sammlungen Des Naturhistorischen Museums in Wien Band 13
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Kataloge der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien Jahr/Year: 1998 Band/Volume: 13 Autor(en)/Author(s): Brock Paul D. Artikel/Article: Catalogue of type specimens of Stick- and Leaf-Insects in the Naturhistorisches Museum Wien (Insecta: Phasmida). 3-72 ©NaturhistorischesKataloge Museum Wien, download unter www.biologiezentrum.at der wissenschaftlichen Sammlungen des Naturhistorischen Museums in Wien Band 13 Entomologie, Heft 5 Paul D. BROCK Catalogue of type specimens of Stick- and Leaf-Insects in the Naturhistorisches Museum Wien (Insecta: Phasmida) Selbstverlag Naturhistorisches Museum Wien Juli 1998 ISBN 3-900 275-67-X ©Naturhistorisches Museum Wien, download unter www.biologiezentrum.at 5 ©Naturhistorisches Museum Wien, download unter www.biologiezentrum.at Catalogue of type specimens of Stick- and Leaf-Insects in the Naturhistorisches Museum Wien (Insecta: Phasmida) P. D. Brock* Abstract Type specimens of784 taxa of Phasmida have been located in the Naturhistorisches Museum Wien (NHMW), which is the most important collection in the world for phasmid taxonomy. The species are listed alphabetically, with the number of specimens, sex and locality data, which, excepting very few instances, have never been recorded before. The most important material relates to species described by Brunner von Wattenwyl and Redtenbacher (mainly published in their monograph, between1906-1908) and the majority of Stäl's types. There are a number of discrepancies in the literature, relating to the where abouts of type specimens, which are commented on; in particular, a number of specimens recorded from other museums are only present in the NHMW and data labels invariably refer to the other museum(s) and, in some instances, are known to have been 'loaned' especially for the monograph. -
The Pregenital Abdominal Musculature in Phasmids and Its Implications for the Basal Phylogeny of Phasmatodea (Insecta: Polyneoptera) Rebecca Klugã, Sven Bradler
ARTICLE IN PRESS Organisms, Diversity & Evolution 6 (2006) 171–184 www.elsevier.de/ode The pregenital abdominal musculature in phasmids and its implications for the basal phylogeny of Phasmatodea (Insecta: Polyneoptera) Rebecca KlugÃ, Sven Bradler Zoologisches Institut und Museum, Georg-August-Universita¨tGo¨ttingen, Berliner Str. 28, 37073 Go¨ttingen, Germany Received 7 June 2005; accepted 25 August 2005 Abstract Recently several conflicting hypotheses concerning the basal phylogenetic relationships within the Phasmatodea (stick and leaf insects) have emerged. In previous studies, musculature of the abdomen proved to be quite informative for identifying basal taxa among Phasmatodea and led to conclusions regarding the basal splitting events within the group. However, this character complex was not studied thoroughly for a representative number of species, and usually muscle innervation was omitted. In the present study the musculature and nerve topography of mid-abdominal segments in both sexes of seven phasmid species are described and compared in detail for the first time including all putative basal taxa, e.g. members of Timema, Agathemera, Phylliinae, Aschiphasmatinae and Heteropteryginae. The ground pattern of the muscle and nerve arrangement of mid-abdominal segments, i.e. of those not modified due to association with the thorax or genitalia, is reconstructed. In Timema, the inner ventral longitudinal muscles are present, whereas they are lost in all remaining Phasmatodea (Euphasmatodea). The ventral longitudinal muscles in the abdomen of Agathemera, which span the whole length of each segment, do not represent the plesiomorphic condition as previously assumed, but might be a result of secondary elongation of the external ventral longitudinal muscles. -
Fam: Bacillidae, Suborden: Areolatae, Orden: Phasmida
Fásmidos espinosos. La Familia Heteropterygidae ( orden: Phasmatodea, suborden: Areolatae, Zompro 2005) Por Sergi Romeu 1- Introducción: En esta familia Heteropterygidae encontramos los insectos más peculiares que podemos imaginarnos, llenos de espinas por todo el cuerpo y con un camuflaje de formas y colores típico del hábitat de sotobosque de las selvas húmedas. Hojas secas, líquenes, musgos, cortezas, pequeñas ramas, brotes, astillas...toman vida al intentar leerlos en este artículo. Principalmente estamos hablando de especies de distribución Asiática presentes en Malaysia, Sumatra, Borneo y muchas otras islas de Indonesia. 2- Clasificación: Durante los últimos años, varios autores han estudiado la sistemática del orden phasmatodea. Principalmente se trata de revisiones teóricas, basadas en descripciones de los ejemplares tipo depositados en los museos de todo el mundo. Paul Brock trata el grupo que nos interesa dentro la familia Bacillidae, como una sub-familia llamada Heteropteryginae, dividiéndola a su vez en cuatro tribus: Datamini, Anisacanthini, Obrimini y Heteropterygini. La mayoría de especies de esta familia Bacillidae no tienen alas, exceptuando algunas especies con rudimentos alares o alas reducidas dentro de nuestra sub-familia Heteropteryginae. Desde la familia Bacillidae, la clave taxonómica para llegar a la sub-familia Heteropteryginae es según P. Brock (1999): - 1) Antena mas larga que el fémur delantero. Alados o sin alas, pero nunca presentes en África y Europa........................................................................................................................................................2 -
Phasmid Studies, 2(1&2)
ISSN 0966-0011 PHASMID STUDIES. volume 2, numbers 1 & 2. June & December 1993. Editor: P.E. Bragg. Published by the Phasmid Study Group. Phasmid Studies ISSN 0966-0011 volume 2, numbers 1 & 2. Contents The World of Stick and Leaf-Insects in Books, some general remarks Paul D . Brock . A very pretty phasmid : Parectatosoma hystrix J. Roget . 7 Looking at Baculum eggs John Sellick . .. 10 Keeping and breeding Haani ella species successfully Ian Abercrombie . 14 List of stick and leaf-insect (Phasmatod ea = Phasmida) type material in the Natural History Museum, published since Kirby's 1904 Cat alogue Paul D. Brock . .. 17 A survey of the distribution of the unarmed stick insect Acanthoxyla inermis in Port Gaverne and Port Isaac, North Cornwall in 1992 Malcolm Lee. .. 25 The Phasmid Database: changes to version 1 P.E . Bragg 33 Reviews and Abstracts Phasmid Abstracts . .. 35 PSG 121 , Phenac ephorus spinulosus (Haus1eithner) P .E . Bragg . .. 41 Pharnacia serratipes (Gray) Frank Hennemann . ... 45 Phena cocephalus coronatus Werner P.E. Bragg . 51 The leaf-piercing eggs of Asceles John Sellick . 54 Defensive and flying behaviour in Sipyloidea sp. (PSG 103) R .P. Bradburne 56 A new Libethra from Ecuador Wim Potvin . 59 Some notes on Din ophasma gutti gera (Westwood) from Borneo P.E. Bragg . 62 Revi ews and Abstracts Pha smid Abstracts . 66 Publications not ed . 67 Cover illustration: Female Phenacephorus spinulosus (Hausleithner) by P.E . Bragg. The World of Stick and Leaf-Insects in Books, some general remarks. Paul D. Brock, "Papillon , 40, Thorndike Road , Slough, SU ISR. UK. Key words Stick and Leaf Insects, Books. -
Standardisation of Bioacoustic Terminology for Insects
Biodiversity Data Journal 8: e54222 doi: 10.3897/BDJ.8.e54222 Research Article Standardisation of bioacoustic terminology for insects Edward Baker‡, David Chesmore‡ ‡ University of York, York, United Kingdom Corresponding author: Edward Baker ([email protected]) Academic editor: Therese Catanach Received: 12 May 2020 | Accepted: 22 Jul 2020 | Published: 04 Aug 2020 Citation: Baker E, Chesmore D (2020) Standardisation of bioacoustic terminology for insects. Biodiversity Data Journal 8: e54222. https://doi.org/10.3897/BDJ.8.e54222 Abstract After reviewing the published literature on sound production in insects, a standardised terminology and controlled vocabularies have been created. This combined terminology has potential for use in automated identification systems, evolutionary studies, and other use cases where the synthesis of bioacoustic traits from the literature is required. An example implementation has been developed for the BioAcoustica platform. It is hoped that future development of controlled vocabularies will become a community effort. Keywords insect, sound production, vocabulary, bioacoustics Introduction "Two dangers face the student seeking to rationalize and codify a terminology that has grown up empirically and that is beginning to differentiate regionally or according to faculty or in other ways - as must always tend to happen. One danger is that of legislating prematurely and clumsily for hypothetical future requirements; the other is a too easy-going and long-sustained attitude of laissez-faire arising from wishing to let the mud settle before trying to penetrate the shadows of often chaotic and obscure usages. If the former danger © Baker E, Chesmore D. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. -
Evolution of Flight Morphology in Stick Insects
1 2 A tale of winglets: evolution of flight morphology in stick insects 3 4 Yu Zeng1,2,†, Conner O’Malley1, Sonal Singhal1,3, Faszly Rahim4,5, 5 Sehoon Park1, Xin Chen6,7, Robert Dudley1,8 6 7 1Department of Integrative Biology, University of California, Berkeley, CA 92870, 8 USA 9 2Schmid College of Science and Technology, Chapman University, Orange, CA 10 92866, USA 11 3 Department of Biology, CSU Dominguez Hills, Carson, CA 90747 USA 12 4Islamic Science Institute (ISI), Universiti Sains Islam Malaysia, 71800 Bandar Baru 13 Nilai, Negeri Sembilan, Malaysia 14 5Centre for Insect Systematics (CIS), Universiti Kebangsaan Malaysia, 43600 15 Bangi, Selangor, Malaysia 16 6Department of Biology, The College of Staten Island, The City University of New 17 York, NY 10314, USA 18 7Department of Biology, The Graduate School and University Center, The City 19 University of New York, NY 10016, USA 20 8Smithsonian Tropical Research Institute, Balboa, 21 Republic of Panama 22 23 †Corresponding author: [email protected] 24 25 1 26 27 Abstract 28 29 The evolutionary transition between winglessness and a full-winged morphology requires 30 selective advantage for intermediate forms. Conversely, repeated secondary wing 31 reductions among the pterygotes indicates relaxation of such selection. However, 32 evolutionary trajectories of such transitions are not well characterized. The stick insects 33 (Phasmatodea) exhibit diverse wing sizes at both interspecific and intersexual levels, and 34 thus provide a system for examining how selection on flight capability, along with other 35 selective forces, drives the evolution of flight-related morphology. Here, we examine 36 variation in relevant morphology for stick insects using data from 1100+ individuals 37 representing 765 species. -
New Stem-Phasmatodea from the Middle Jurassic of China
Eur. J. Entomol. 108: 677–685, 2011 http://www.eje.cz/scripts/viewabstract.php?abstract=1667 ISSN 1210-5759 (print), 1802-8829 (online) New stem-Phasmatodea from the Middle Jurassic of China LIANGJIE SHANG1, OLIVIER BÉTHOUX2 and DONG REN1* 1College of Life Science, Capital Normal University, 105 Xisanhuanbeilu, Haidian District, Beijing 100048, China; e-mail: [email protected] 240 rue d’Aveillans, 38770 La Motte d’Aveillans, France; e-mail: [email protected] Key words. Adjacivena rasnitsyni gen. n., sp. n., Phasmatodea, Susumaniidae, Heteropteryx dilatata, China, Daohugou, intra-specific variability, sexual dimorphism, fossil, Middle Jurassic Abstract. Adjacivena rasnitsyni gen. n., sp. n., from the Middle Jurassic Jiulongshan Formation (Daohugou Village, Inner Mongolia, China), is described based on a set of well-preserved specimens, interpreted as two females and a male of the same species. The new taxon is assigned to the family Susumaniidae, i.e. is a candidate stem-Phasmatodea. This ordinal assignment is supported by genital elements, chiefly a putative operculum concealing the ovipositor observed in one female specimen and a putative extension of the 10th tergum with a thorn pad in the male specimen. Variation in the venation of the forewing is appreciated based on more or less complete pairs of wings and a comparison with that in the forewings of male Heteropteryx dilatata (Parkinson, 1798). This material offers new perspectives on the primary homologies of the hind wing venation of stick-insects. INTRODUCTION “Cu” is concave). This situation has two interrelated con- Stick-insects (also known as Cheleutoptera, Phasmato- sequences: conjectures about the primary homologies of dea, Phasmida and Phasmatoptera, among others) com- stick-insect wing venation and the identification of sev- pose a rather small order of insects, which currently eral Mesozoic representatives of the order are both uncer- includes over 3,000 extant species (Bragg, 2001; Brock & tain.