ATLAS OF PENNSYLVANIA

Compiled & Maintained By:

David M. Wright [email protected]

1st Edition 1995 [Dec. 18] 2nd Edition (Revised) 2001 [Feb. 17] 3rd Edition (Revised) 2003 [Jan. 30] 4th Edition (Revised) 2005 [Jan. 5 ] 5th Edition (Revised) 2006 [Jan. 21] 6th Edition (Revised) 2007 [Jan. 3 ] 7th Edition (Revised) 2008 [Jan. 7 ] 8th Edition (Revised) 2009 [Jan. 10] 9th Edition (Revised) 2010 [Jan. 15] 10th Edition (Revised) 2011 [Jan. 10] 11th Edition (Revised) 2012 [Jan. 16] 12th Edition (Revised) 2013 [Jan. 13] 13th Edition (Revised) 2014 [Jan. 15]

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PAPILIONIDAE Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

Papilioninae (Swallowtails) 1 2 3 4

Battus philenor Eurytides marcellus polyxenes Papilio cresphontes Pipevine Swallowtail Zebra Swallowtail Black Swallowtail Giant Swallowtail 5 6 7 8

Papilio glaucus Papilio appalachiensis Papilio troilus E. Tiger Swallowtail Canadian T. Swallowtail Appalachian T. Swallowtail Spicebush Swallowtail 9

Papilio palamedes Palamedes Swallowtail

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PIERIDAE Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

Pierinae (Whites) 10 11 12 13

Pontia protodice Pieris rapae Pieris virginiensis Euchloe olympia Checkered White Cabbage White West Virginia White Olympia Marble 14

Anthocharis midea Falcate Orangetip

Coliadinae (Sulphurs) 15 16 17 18

Colias philodice Colias eurytheme Colias interior Zerene cesonia Clouded Sulphur Orange Sulphur Pink-edged Sulphur Southern Dogface 19 20 21 22

Phoebis sennae Phoebis philea Pyrisitia lisa Abaeis nicippe Cloudless Sulphur Orange-barred Sulphur Little Yellow Sleepy Orange 23

Nathalis iole Dainty Sulphur

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LYCAENIDAE Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

Miletinae (Harvesters) Lycaeninae (Coppers) 24 25 26 27

Feniseca tarquinius Lycaena phlaeas Lycaena hyllus Lycaena epixanthe Harvester American Copper Bronze Copper Bog Copper

Theclinae (Hairstreaks) 28 29 30 31

Atlides halesus titus Satyrium acadica Great Purple Hairstreak Coral Hairstreak Acadian Hairstreak Edwards’ Hairstreak 32 33 34 35

Satyrium calanus Satyrium liparops Satyrium favonius Banded Hairstreak Hickory Hairstreak Striped Hairstreak Southern Hairstreak 36 37 38 39

Callophrys augustinus Callophrys polios Callophrys irus Callophrys henrici Brown Elfin Hoary Elfin Frosted Elfin Henry’s Elfin 40 41 42 43

Callophrys niphon Callophrys gryneus Parrhasius m-album Strymon melinus Eastern Pine Elfin Juniper Hairstreak White M Hairstreak Gray Hairstreak 44 45

Calycopis cecrops Erora laeta Red-banded Hairstreak Early Hairstreak 5

LYCAENIDAE continued Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

Polyommatinae (Blues) 46 47 48 49

Leptotes marina Cupido comyntas ladon Celastrina lucia Marine Blue Eastern Tailed Blue Spring Azure Northern Spring Azure 50 51 52 53

Celastrina neglecta Celastrina serotina Celastrina neglectamajor Celastrina nigra Summer Azure Cherry Gall Azure Appalachian Azure Dusky Azure 54 55

Glaucopsyche lygdamus Plebejus samuelis Silvery Blue Karner Blue

RIODINIDAE

Riodininae (Metalmarks) 56

Calephelis borealis Northern Metalmark

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NYMPHALIDAE Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

Libytheinae (Snouts) 57

Libytheana carinenta American Snout

Danainae (Milkweed Butterflies) 58 59

Danaus plexippus Danaus gilippus Monarch Queen

Heliconiinae (Longwings & Fritillaries) 60 61

Agraulis vanillae Euptoieta claudia Gulf Fritillary Variegated Fritillary 62 63 64 65

Speyeria cybele Speyeria aphrodite Speyeria idalia Speyeria atlantis Great Spangled Fritillary Aphrodite Fritillary Regal Fritillary Atlantis Fritillary 66 67 68

Speyeria diana Boloria selene Boloria bellona Diana Fritillary Silver-bordered Fritillary Meadow Fritillary [unverified old record]

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NYMPHALIDAE continued Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

Nymphalinae (True Brushfoots) 69 70 71 72

Chlosyne gorgone Chlosyne nycteis Chlosyne harrisii Phyciodes tharos Gorgone Checkerspot Silvery Checkerspot Harris’ Checkerspot Pearl Crescent 73 single-brooded (▪) 74 75 76

Phyciodes cocyta (2 sp. ?) Phyciodes batesii Euphydryas phaeton Junonia coenia Northern Crescent Tawny Crescent Baltimore Common Buckeye 77 78 79 80

Polygonia interrogationis Polygonia comma Polygonia faunus Polygonia progne Question Mark E. Comma/Hop Merchant Green Comma Gray Comma 81 82 83 84

Nymphalis l-album j-album Nymphalis californica Nymphalis antiopa Aglais milberti Compton’s Tortoiseshell California Tortoiseshell Mourning Cloak Milbert’s Tortoiseshell 85 86 87

Vanessa virginiensis Vanessa cardui Vanessa atalanta American Lady Painted Lady Red Admiral

Limenitidinae (Admirals & Relatives) ssp. arthemis 88 89 90 91

Limenitis a. arthemis-like L. arthemis ssp. intergrades Limenitis a. astyanax Limenitis archippus White Admiral (full bands) Hybrid Purple (partial bands) Red-spotted Purple Viceroy

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NYMPHALIDAE continued Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

Apaturinae (Emperors) 92 93

Asterocampa celtis Asterocampa clyton Hackberry Emperor Tawny Emperor

Satyrinae (Satyrs & Browns) 94 95 96 97

Lethe anthedon Lethe eurydice Lethe appalachia Hermeuptychia sosybius Northern Pearly Eye Eyed Brown Appalachian Brown Carolina Satyr 98 99 100

Megisto cymela Coenonympha tullia Cercyonis pegala Little Wood Satyr Common Ringlet Common Wood Nymph

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HESPERIIDAE (Skippers) Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

Eudaminae (Dicot Skippers) 101 102 103 104

Epargyreus clarus Urbanus proteus Autochton cellus Achalarus lyciades Silver-spotted Long-tailed Skipper Golden-banded Skipper Hoary Edge 105 106 107

Thorybes bathyllus Thorybes pylades Thorybes confusis Southern Cloudywing Northern Cloudywing Confused Cloudywing

Pyrginae (Spread-winged Skippers) 108 109 110 111

Staphylus hayhurstii Erynnis icelus Hayhurst’s Scallopwing Dreamy Sleepy Duskywing Juvenal’s Duskywing 112 113 114 115

Erynnis horatius Erynnis martialis Erynnis zarucco Erynnis funeralis Horace’s Duskywing Mottled Duskywing Zarucco Duskywing Funereal Duskywing 116 117 118 119

Erynnis lucilius Pyrgus centaureae Columbine Duskywing Wild Indigo Duskywing Persius Duskywing Grizzled Skipper 120 121

Pyrgus communis Pholisora catullus Common Checkered Skipper Common Sootywing

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HESPERIIDAE continued Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

Heteropterinae (Skipperlings) 122

Carterocephalus palaemon Arctic Skipper

Hesperiinae () 123 124 125 126

Nastra lherminier accius numitor lineola Swarthy Skipper Clouded Skipper Least Skipper European Skipper 127 128 129 130

Hylephila phyleus Hesperia metea Fiery Skipper Sachem Leonard’s Skipper Cobweb Skipper 131 132 133 134

Hesperia attalus Hesperia sassacus peckius Polites themistocles Dotted Skipper Indian Skipper Peck’s Skipper Tawny-edged Skipper 135 136 137 138

Polites origenes Polites mystic Wallengrenia egeremet Crossline Skipper Long Dash Whirlabout Northern Broken Dash 139 140 141

Pompeius verna Little Glassywing Arogos Skipper Delaware Skipper

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HESPERIIDAE continued Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014

142 143 144 145

Poanes massasoit hobomok Poanes zabulon Mulberry Wing Hobomok Skipper Zabulon Skipper Broad-winged Skipper 146 147 148 149

Euphyes conspicua Euphyes bimacula Euphyes vestris Dion Skipper Black Dash Two-spotted Skipper Dun Skipper 150 151 152 153

Atrytonopsis hianna hegon Oligoria maculata Dusted Skipper Pepper & Salt Skipper Common Roadside Skipper Twin-spot Skipper 154 155 156

Calpodes ethlius panoquin Brazilian Skipper Salt Marsh Skipper Ocola Skipper

12 INDEX - SCIENTIFIC NAMES cardui (Vanessa), 86 Euchloe olympia, 13 palaemon, 122 Euphydryas phaeton, 75 Abaeis nicippe, 22 caryaevorus (Satyrium), 33 Euphyes bimacula, 148 acadica (Satyrium), 30 carinenta (Libytheana), 57 conspicua, 147 accius (Lerema), 124 catullus (Pholisora), 121 dion, 146 Achalarus lyciades, 104 cecrops (Calycopis), 44 vestris, 149 Aglais milberti, 84 Celastrina ladon, 48 Euptoieta claudia, 61 Agraulis vanillae, 60 lucia, 49 eurydice (Lethe), 95 , 151 neglecta, 50 eurytheme (Colias), 16 vialis, 152 neglectamajor, 52 Eurytides marcellus, 2 Anatrytone logan, 141 nigra, 53 Feniseca tarquinius, 24 Ancyloxypha numitor, 125 serotina, 51 faunus (Polygonia), 79 anthedon (Lethe), 94 cellus (Autochton), 103 favonius (Satyrium), 35 antiopa (Nymphalis), 83 celtis (Asterocampa), 92 funeralis (Erynnis), 115 Anthocharis midea, 14 centaureae (Pyrgus), 119 gilippus (Danaus), 59 aphrodite (Speyeria), 63 Cercyonis pegala, 100 Glaucopsyche lygdamus, 54 appalachia (Lethe), 96 cesonia (Zerene), 18 glaucus (Papilio), 5 appalachiensis (Papilio), 7 clarus (Epargyreus), 101 gorgone (Chlosyne), 69 archippus (Limenitis), 91 claudia (Euptoieta), 61 gryneus (Callophrys), 41 arogos (Atrytone), 140 clyton (Asterocampa), 93 halesus (Atlides), 28 arthemis-like (Limenitis), 88 cocyta (Phyciodes), 73 harrisii (Chlosyne), 71 arthemis intergrades (Limenitis), 89 coenia (Junonia), 76 hayhurstii (Staphylus), 108 arthemis astyanax (Limenitis), 90 Coenonympha tullia, 99 hegon (Amblyscirtes), 151 Asterocampa celtis, 92 Colias eurytheme, 16 henrici (Callophrys), 39 clyton, 93 interior, 17 Hermeuptychia sosybius, 97 atalanta (Vanessa), 87 philodice, 15 Hesperia attalus, 131 Atalopedes campestris, 128 comma (Polygonia), 78 leonardus, 129 atlantis (Speyeria), 65 communis (Pyrgus), 120 metea, 130 Atlides halesus, 28 comyntas (Cupido), 47 sassacus, 132 Atrytone arogos, 140 confusis (Thorybes), 107 hianna (Atrytonopsis), 150 Atrytonopsis hianna, 150 conspicua (Euphyes), 147 hobomok (Poanes), 143 Attalus (Hesperia), 131 cresphontes (Papilio), 4 horatius (Erynnis), 112 augustinus (Callophrys), 36 Chlosyne gorgone, 69 Hylephila phyleus, 127 Autochton cellus, 103 harrisii, 71 hyllus (Lycaena), 26 baptisiae (Erynnis), 117 nycteis, 70 icelus (Erynnis), 109 batesii (Phyciodes), 74 Cupido comyntas, 47 idalia (Speyeria), 64 bathyllus (Thorybes), 105 cybele (Speyeria), 62 interior (Colias), 17 Battus philenor, 1 cymela (Megisto), 98 interrogationis (Polygonia), 77 bellona (Boloria), 68 Danaus gilippus, 59 iole (Nathalis), 23 bimacula (Euphyes), 148 plexippus, 58 irus (Callophrys), 38 brizo (Erynnis), 110 diana (Speyeria), 66 j-album (Nymphalis), 81 Boloria bellona, 68 dion (Euphyes), 146 Junonia coenia, 76 selene, 67 edwardsii (Satyrium), 31 juvenalis (Erynnis), 111 borealis (Calephelis), 56 egeremet (Wallengrenia), 138 ladon (Celastrina), 48 calanus (Satyrium), 32 Epargyreus clarus, 101 laeta (Erora), 45 Calephelis borealis, 56 epixanthe (Lycaena), 27 l-album (Nymphalis), 81 californica (Nymphalis), 82 Erora laeta, 45 , 124 Callophrys augustinus, 36 Erynnis baptisiae, 117 Lethe anthedon, 94 gryneus, 41 brizo, 110 appalachia, 96 henrici, 39 funeralis, 115 eurydice, 95 irus, 38 horatius, 112 Leptotes marina, 46 niphon, 40 icelus, 109 leonardus (Hesperia), 129 polios, 37 juvenalis, 111 lherminier (), 123 ethlius, 154 lucilius, 116 Libytheana carinenta, 57 campestris (Atalopedes), 128 martialis, 113 Limenitis archippus, 91 canadensis (Papilio), 6 persius, 118 arthemis-like, 88 Calycopis cecrops, 44 zarucco, 114 arthemis intergrades 89 ethlius (Calpodes), 154 a. astyanax, 90

13 INDEX - (continued) phaeton (Euphydryas), 75 Staphylus hayhurstii, 108 philenor (Battus), 1 Strymon melinus, 43 lineola (Thymelicus), 126 philea (Phoebis), 20 tarquinius (Feniseca), 24 liparops (Satyrium), 34 philodice (Colias), 15 tharos (Phyciodes), 72 lisa (Pyrisitia), 21 phlaeas (Lycaena), 25 themistocles (Polites), 134 logan (Anatrytone), 141 Phoebis philea, 20 Thorybes bathyllus, 105 lucia (Celastrina), 49 sennae, 19 confusis, 107 lucilius (Erynnis), 116 Pholisora catullus, 121 pylades, 106 Lycaena epixanthe, 27 Phyciodes batesii, 74 Thymelicus lineola, 126 hyllus, 26 cocyta, 73 titus (Satyrium), 29 phlaeas, 25 tharos, 72 troilus (Papilio), 8 lyciades (Achalarus), 104 phyleus (Hylephila), 127 tullia (Coenonympha), 99 lygdamus (Glaucopsyche), 54 Pieris rapae, 11 Urbanus proteus, 102 maculata (Oligoria), 153 virginiensis, 12 Vanessa atalanta, 87 m-album (Parrhasius), 42 Plebejus samuelis, 55 cardui, 86 marcellus (Eurytides), 2 Poanes hobomok, 143 virginiensis, 85 marina (Leptotes), 46 massasoit, 142 vanillae (Agraulis), 60 massasoit (Poanes), 142 viator, 145 verna (), 139 martialis (Erynnis), 113 zabulon, 144 vestris (Euphyes), 149 Megisto cymela, 98 polios (Callophrys), 37 vialis (Amblyscirtes), 152 melinus (Strymon), 43 Polites mystic, 136 viator (Poanes), 145 melissa (see samuelis), 55 origenes, 135 vibex (Polites), 137 metea (Hesperia), 130 peckius, 133 virginiensis (Pieris), 12 midea (Anthocharis), 14 themistocles, 134 virginiensis (Vanessa), 85 milberti (Aglais), 84 vibex, 137 Wallengrenia egeremet, 138 mystic (Polites), 136 Polygonia comma, 78 zabulon (Poanes), 144 , 123 faunus, 79 zarucco (Erynnis), 114 Nathalis iole, 23 interrogationis, 77 Zerene cesonia, 18 neglecta (Celastrina), 50 progne, 80 L neglectamajor (Celastrina), 52 polyxenes (Papilio), 3 L nicippe (Abaeis), 22 , 139 L nigra (Celastrina), 53 Pontia protodice, 10 L niphon (Callophrys), 40 progne (Polygonia), 80 L nycteis (Chlosyne), 70 proteus (Urbanus), 102 L Nymphalis antiopa, 83 protodice (Pontia), 10 L californica, 82 pylades (Thorybes), 106 L l-album j-album, 81 Pyrgus centaureae, 119 L numitor (Ancyloxypha), 125 communis, 120 L ocola (Panoquina), 156 Pyrisitia lisa, 21 L origenes (Polites), 135 rapae (Pieris), 11 L Oligoria maculata, 153 samuelis (Plebejus), 55 L olympia (Euchloe), 13 sassacus (Hesperia), 132 L palaemon (Carterocephalus), 122 Satyrium acadica, 30 L palamedes (Papilio), 9 calanus, 32 L panoquin (Panoquina), 155 caryaevorus, 33 L Panoquina ocola, 156 edwardsii, 31 L panoquin, 155 favonius, 35 L Papilio appalachiensis, 7 liparops, 34 L canadensis, 6 titus, 29 L cresphontes, 4 selene (Boloria), 67 L glaucus, 5 sennae (Phoebis), 19 L palamedes, 9 serotina (Celastrina), 51 L polyxenes, 3 sosybius (Hermeuptychia), 97 L troilus, 8 Speyeria aphrodite, 63 L Parrhasius m-album, 42 atlantis, 65 peckius (Polites), 133 cybele, 62 pegala (Cercyonis), 100 diana, 66 persius (Erynnis), 118 idalia, 64 14

INDEX - COMMON NAMES

Admiral Duskywing Lady Red, 87 Columbine, 116 American, 85 White, 88 Dreamy, 109 Painted, 86 Azure Horace’s, 112 Little Glassywing, 139 Appalachian, 52 Funereal, 115 Little Wood Satyr, 98 Cherry Gall, 51 Juvenal’s, 111 Little Yellow, 21 Dusky, 53 Mottled, 113 Long Dash, 136 Northern Spring, 49 Persius, 118 Marble Spring, 48 Sleepy, 110 Olympia, 13 Summer, 50 Wild Indigo, 117 Metalmark Baltimore, 75 Zarucco, 114 Northern, 56 Black Dash, 147 Eastern Tailed Blue, 47 Monarch, 58 Blue Elfin Mourning Cloak, 83 Eastern Tailed, 47 Brown, 36 Mulberry Wing, 142 Karner, 55 Eastern Pine, 40 Orange Marine, 46 Frosted, 38 Sleepy, 22 Silvery, 54 Henry’s, 39 Orangetip Broken Dash Hoary, 37 Falcate, 14 Northern, 138 Emperor Painted Lady, 86 Brown Hackberry, 92 Pearly Eye Appalachian, 96 Tawny, 93 Northern, 94 Eyed, 95 Eyed Brown, 95 Purple Buckeye Falcate Orangetip, 14 Hybrid, 89 Common, 76 Fritillary Red-spotted, 90 Cabbage White, 11 Aphrodite, 63 Queen, 59 Checkered Skipper Atlantis, 65 Question Mark, 77 Common, 120 Diana, 66 Red Admiral, 87 Checkered White, 10 Great Spangled, 62 Ringlet Checkerspot Gulf, 60 Common, 99 Gorgone, 69 Meadow, 68 Roadside Skipper Harris’, 71 Regal, 64 Common, 152 Silvery, 70 Silver-bordered, 67 Sachem, 128 Cloudywing Variegated, 61 Satyr Confused, 107 Glassywing Carolina, 97 Northern, 106 Little, 139 Little Wood, 98 Southern, 105 Hairstreak Scallopwing Comma Acadian, 30 Hayhurst’s, 108 Eastern, 78 Banded, 32 Skipper Gray, 80 Coral, 29 Arctic, 122 Green, 79 Early, 45 Arogos, 140 Copper Edwards’, 31 Brazilian, 154 American, 25 Gray, 43 Broad-winged, 145 Bog, 25 Great Purple, 28 Checkered, Common, 120 Bronze, 26 Hickory, 33 Clouded, 124 Crescent Juniper, 41 Cobweb, 130 Northern, 73 Red-banded, 44 Common Checkered, 120 Pearl, 72 Southern, 35 Common Roadside, 152 Tawny, 74 Striped, 34 Crossline, 135 Dash (see Broken Dash) White M, 42 Delaware, 141 Black, 147 Harvester, 24 Dion, 146 Long, 136 Hoary Edge, 104 Dotted, 131 Dogface Hop Merchant, 78 Dun, 149 Southern, 18 Dusted, 150 European, 126 15

INDEX - (continued) Yellow L Little, 21 L

Skipper (continued) L L Fiery, 127 L L L Golden-banded, 103 L L Grizzled, 119 L L L Hobomok, 142 L Indian, 132 L L L Least, 125 L L Leonard’s, 129 L L L Long-tailed, 102 L L Ocola, 156 L L

Peck’s, 133 L L Pepper & Salt, 151 L L Roadside, Common, 152 L L Salt Marsh, 155 L

L Silver-spotted, 101 L L Swarthy, 123 L L L Tawny-edged, 134 L L Twin-spotted, 153 L L

Two-spot, 148 L L Zabulon, 144 L L L Snout L L American, 57 L L Sootywing L Common, 121 L L L Spring Azure, 48 L

Sulphur L L L Clouded, 15 L L Cloudless, 19 L L L Dainty, 23 L L Orange, 16 L L L Orange-barred, 20 L Pink-edged, 17 L L L Summer Azure, 50 L L Swallowtail L L L Appalachian Tiger, 7 L L Black, 3 L

L Canadian Tiger, 6 L L Eastern Tiger, 5 L L L Giant, 4 L L Palamedes, 9 L L L Pipevine, 1 L Spicebush, 8 L L L Zebra, 2 L L Tortoiseshell L

L California, 82 L L Compton’s, 81 L L

Milbert’s, 84 L L Viceroy, 91 L L L West Virginia White, 12 L L Whirlabout, 137 L L L White L L Cabbage, 11 L L L Checkered, 10 L L West Virginia, 12 L L L White Admiral, 88 L Wood Nymph L Common, 100 L

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NOTES:

County distribution sources include:

Compiler’s extensive bibliography of Pennsylvania butterfly literature (1798-2013), Field Summary of The Lepidopterists’ Society (Lepidopterists’ News, 1947-1953), Season Summary of The Lepidopterists’ Society (“News of the Lepidopterists’ Society”, 1959-2013), specimen vouchers in natural history museums in U.S. & Canada and private collections (50+), personal correspondence (1969-2013), reports to online listserve group at: http://groups.yahoo.com/group/PaLepsOdes/ (2005-2013), reports to NABA Recent Sightings at: http://www.naba.org/sightings/sightings.html (1999-2013), and reports to the Pennsylvania State Coordinator for Butterflies and Moths of North America at: http://npwrc.usgs.gov/resource/distr/lepid/bflyusa/bflusa.htm followed by http://www.butterfliesandmoths.org/ (1995-2005 and 2005-2013 respectively).

Scientific names follow:

Pelham, J. 2008. Catalogue of the butterflies of the United States and Canada. Journal of Research on the 40:1-672 pp. [Catalogue available at: http://butterfliesofamerica.com/US-Can-Cat-1-30-2011.htm. This online version is frequently updated with new taxonomic information. Last revision dated 2/14/2012.]

Common names follow:

North American Butterfly Association (NABA) Checklist & English Names of North American Butterflies, 1st Edition (1995) and 2nd Edition (2001), with modifications from Opler, P.A. 1998. A Field Guide to Eastern Butterflies. Peterson Field Guides. Houghton Mifflin Company, 512 pp. ; J.P. Brock, J. & K. Kaufmann. 2003. Butterflies of North America. Kaufmann Field Guides. Houghton Mifflin Harcourt Publishing Co., 392 pp.; and Opler, P.A., K. Lotts, and T. Naberhaus [coordinators]. 2012. Butterflies and Moths of North America. Bozeman, MT: Big Sky Institute. Data set last accessed on 12/18/2013 at: http://www.butterfliesandmoths.org/.