Atlas of Pennsylvania Butterflies
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ATLAS OF PENNSYLVANIA BUTTERFLIES Compiled & Maintained By: David M. Wright [email protected] 1st Edition 1995 [Dec. 18] 2nd Edition (Revised) 2001 [Feb. 17] 3rd Edition (Revised) 2003 [Jan. 30] 4th Edition (Revised) 2005 [Jan. 5 ] 5th Edition (Revised) 2006 [Jan. 21] 6th Edition (Revised) 2007 [Jan. 3 ] 7th Edition (Revised) 2008 [Jan. 7 ] 8th Edition (Revised) 2009 [Jan. 10] 9th Edition (Revised) 2010 [Jan. 15] 10th Edition (Revised) 2011 [Jan. 10] 11th Edition (Revised) 2012 [Jan. 16] 12th Edition (Revised) 2013 [Jan. 13] 13th Edition (Revised) 2014 [Jan. 15] 2 PAPILIONIDAE Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014 Papilioninae (Swallowtails) 1 2 3 4 Battus philenor Eurytides marcellus Papilio polyxenes Papilio cresphontes Pipevine Swallowtail Zebra Swallowtail Black Swallowtail Giant Swallowtail 5 6 7 8 Papilio glaucus Papilio canadensis Papilio appalachiensis Papilio troilus E. Tiger Swallowtail Canadian T. Swallowtail Appalachian T. Swallowtail Spicebush Swallowtail 9 Papilio palamedes Palamedes Swallowtail 3 PIERIDAE Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014 Pierinae (Whites) 10 11 12 13 Pontia protodice Pieris rapae Pieris virginiensis Euchloe olympia Checkered White Cabbage White West Virginia White Olympia Marble 14 Anthocharis midea Falcate Orangetip Coliadinae (Sulphurs) 15 16 17 18 Colias philodice Colias eurytheme Colias interior Zerene cesonia Clouded Sulphur Orange Sulphur Pink-edged Sulphur Southern Dogface 19 20 21 22 Phoebis sennae Phoebis philea Pyrisitia lisa Abaeis nicippe Cloudless Sulphur Orange-barred Sulphur Little Yellow Sleepy Orange 23 Nathalis iole Dainty Sulphur 4 LYCAENIDAE Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014 Miletinae (Harvesters) Lycaeninae (Coppers) 24 25 26 27 Feniseca tarquinius Lycaena phlaeas Lycaena hyllus Lycaena epixanthe Harvester American Copper Bronze Copper Bog Copper Theclinae (Hairstreaks) 28 29 30 31 Atlides halesus Satyrium titus Satyrium acadica Satyrium edwardsii Great Purple Hairstreak Coral Hairstreak Acadian Hairstreak Edwards’ Hairstreak 32 33 34 35 Satyrium calanus Satyrium caryaevorus Satyrium liparops Satyrium favonius Banded Hairstreak Hickory Hairstreak Striped Hairstreak Southern Hairstreak 36 37 38 39 Callophrys augustinus Callophrys polios Callophrys irus Callophrys henrici Brown Elfin Hoary Elfin Frosted Elfin Henry’s Elfin 40 41 42 43 Callophrys niphon Callophrys gryneus Parrhasius m-album Strymon melinus Eastern Pine Elfin Juniper Hairstreak White M Hairstreak Gray Hairstreak 44 45 Calycopis cecrops Erora laeta Red-banded Hairstreak Early Hairstreak 5 LYCAENIDAE continued Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014 Polyommatinae (Blues) 46 47 48 49 Leptotes marina Cupido comyntas Celastrina ladon Celastrina lucia Marine Blue Eastern Tailed Blue Spring Azure Northern Spring Azure 50 51 52 53 Celastrina neglecta Celastrina serotina Celastrina neglectamajor Celastrina nigra Summer Azure Cherry Gall Azure Appalachian Azure Dusky Azure 54 55 Glaucopsyche lygdamus Plebejus samuelis Silvery Blue Karner Blue RIODINIDAE Riodininae (Metalmarks) 56 Calephelis borealis Northern Metalmark 6 NYMPHALIDAE Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014 Libytheinae (Snouts) 57 Libytheana carinenta American Snout Danainae (Milkweed Butterflies) 58 59 Danaus plexippus Danaus gilippus Monarch Queen Heliconiinae (Longwings & Fritillaries) 60 61 Agraulis vanillae Euptoieta claudia Gulf Fritillary Variegated Fritillary 62 63 64 65 Speyeria cybele Speyeria aphrodite Speyeria idalia Speyeria atlantis Great Spangled Fritillary Aphrodite Fritillary Regal Fritillary Atlantis Fritillary 66 67 68 Speyeria diana Boloria selene Boloria bellona Diana Fritillary Silver-bordered Fritillary Meadow Fritillary [unverified old record] 7 NYMPHALIDAE continued Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014 Nymphalinae (True Brushfoots) 69 70 71 72 Chlosyne gorgone Chlosyne nycteis Chlosyne harrisii Phyciodes tharos Gorgone Checkerspot Silvery Checkerspot Harris’ Checkerspot Pearl Crescent 73 single-brooded (▪) 74 75 76 Phyciodes cocyta (2 sp. ?) Phyciodes batesii Euphydryas phaeton Junonia coenia Northern Crescent Tawny Crescent Baltimore Common Buckeye 77 78 79 80 Polygonia interrogationis Polygonia comma Polygonia faunus Polygonia progne Question Mark E. Comma/Hop Merchant Green Comma Gray Comma 81 82 83 84 Nymphalis l-album j-album Nymphalis californica Nymphalis antiopa Aglais milberti Compton’s Tortoiseshell California Tortoiseshell Mourning Cloak Milbert’s Tortoiseshell 85 86 87 Vanessa virginiensis Vanessa cardui Vanessa atalanta American Lady Painted Lady Red Admiral Limenitidinae (Admirals & Relatives) ssp. arthemis 88 89 90 91 Limenitis a. arthemis-like L. arthemis ssp. intergrades Limenitis a. astyanax Limenitis archippus White Admiral (full bands) Hybrid Purple (partial bands) Red-spotted Purple Viceroy 8 NYMPHALIDAE continued Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014 Apaturinae (Emperors) 92 93 Asterocampa celtis Asterocampa clyton Hackberry Emperor Tawny Emperor Satyrinae (Satyrs & Browns) 94 95 96 97 Lethe anthedon Lethe eurydice Lethe appalachia Hermeuptychia sosybius Northern Pearly Eye Eyed Brown Appalachian Brown Carolina Satyr 98 99 100 Megisto cymela Coenonympha tullia Cercyonis pegala Little Wood Satyr Common Ringlet Common Wood Nymph 9 HESPERIIDAE (Skippers) Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014 Eudaminae (Dicot Skippers) 101 102 103 104 Epargyreus clarus Urbanus proteus Autochton cellus Achalarus lyciades Silver-spotted Skipper Long-tailed Skipper Golden-banded Skipper Hoary Edge 105 106 107 Thorybes bathyllus Thorybes pylades Thorybes confusis Southern Cloudywing Northern Cloudywing Confused Cloudywing Pyrginae (Spread-winged Skippers) 108 109 110 111 Staphylus hayhurstii Erynnis icelus Erynnis brizo Erynnis juvenalis Hayhurst’s Scallopwing Dreamy Duskywing Sleepy Duskywing Juvenal’s Duskywing 112 113 114 115 Erynnis horatius Erynnis martialis Erynnis zarucco Erynnis funeralis Horace’s Duskywing Mottled Duskywing Zarucco Duskywing Funereal Duskywing 116 117 118 119 Erynnis lucilius Erynnis baptisiae Erynnis persius Pyrgus centaureae Columbine Duskywing Wild Indigo Duskywing Persius Duskywing Grizzled Skipper 120 121 Pyrgus communis Pholisora catullus Common Checkered Skipper Common Sootywing 10 HESPERIIDAE continued Atlas of Pennsylvania Butterflies 13th Edition (Rev.) – 2014 Heteropterinae (Skipperlings) 122 Carterocephalus palaemon Arctic Skipper Hesperiinae (Grass Skippers) 123 124 125 126 Nastra lherminier Lerema accius Ancyloxypha numitor Thymelicus lineola Swarthy Skipper Clouded Skipper Least Skipper European Skipper 127 128 129 130 Hylephila phyleus Atalopedes campestris Hesperia leonardus Hesperia metea Fiery Skipper Sachem Leonard’s Skipper Cobweb Skipper 131 132 133 134 Hesperia attalus Hesperia sassacus Polites peckius Polites themistocles Dotted Skipper Indian Skipper Peck’s Skipper Tawny-edged Skipper 135 136 137 138 Polites origenes Polites mystic Polites vibex Wallengrenia egeremet Crossline Skipper Long Dash Whirlabout Northern Broken Dash 139 140 141 Pompeius verna Atrytone arogos Anatrytone logan Little Glassywing Arogos Skipper Delaware Skipper 11 HESPERIIDAE continued Atlas of Pennsylvania Butterflies th 13 Edition (Rev.) – 2014 142 143 144 145 Poanes massasoit Poanes hobomok Poanes zabulon Poanes viator Mulberry Wing Hobomok Skipper Zabulon Skipper Broad-winged Skipper 146 147 148 149 Euphyes dion Euphyes conspicua Euphyes bimacula Euphyes vestris Dion Skipper Black Dash Two-spotted Skipper Dun Skipper 150 151 152 153 Atrytonopsis hianna Amblyscirtes hegon Amblyscirtes vialis Oligoria maculata Dusted Skipper Pepper & Salt Skipper Common Roadside Skipper Twin-spot Skipper 154 155 156 Calpodes ethlius Panoquina panoquin Panoquina ocola Brazilian Skipper Salt Marsh Skipper Ocola Skipper 12 INDEX - SCIENTIFIC NAMES cardui (Vanessa), 86 Euchloe olympia, 13 Carterocephalus palaemon, 122 Euphydryas phaeton, 75 Abaeis nicippe, 22 caryaevorus (Satyrium), 33 Euphyes bimacula, 148 acadica (Satyrium), 30 carinenta (Libytheana), 57 conspicua, 147 accius (Lerema), 124 catullus (Pholisora), 121 dion, 146 Achalarus lyciades, 104 cecrops (Calycopis), 44 vestris, 149 Aglais milberti, 84 Celastrina ladon, 48 Euptoieta claudia, 61 Agraulis vanillae, 60 lucia, 49 eurydice (Lethe), 95 Amblyscirtes hegon, 151 neglecta, 50 eurytheme (Colias), 16 vialis, 152 neglectamajor, 52 Eurytides marcellus, 2 Anatrytone logan, 141 nigra, 53 Feniseca tarquinius, 24 Ancyloxypha numitor, 125 serotina, 51 faunus (Polygonia), 79 anthedon (Lethe), 94 cellus (Autochton), 103 favonius (Satyrium), 35 antiopa (Nymphalis), 83 celtis (Asterocampa), 92 funeralis (Erynnis), 115 Anthocharis midea, 14 centaureae (Pyrgus), 119 gilippus (Danaus), 59 aphrodite (Speyeria), 63 Cercyonis pegala, 100 Glaucopsyche lygdamus, 54 appalachia (Lethe), 96 cesonia (Zerene), 18 glaucus (Papilio), 5 appalachiensis (Papilio), 7 clarus (Epargyreus), 101 gorgone (Chlosyne), 69 archippus (Limenitis), 91 claudia (Euptoieta), 61 gryneus (Callophrys), 41 arogos (Atrytone), 140 clyton (Asterocampa), 93 halesus (Atlides), 28 arthemis-like (Limenitis), 88 cocyta (Phyciodes), 73 harrisii (Chlosyne), 71 arthemis intergrades (Limenitis), 89 coenia (Junonia), 76 hayhurstii (Staphylus), 108 arthemis astyanax (Limenitis), 90 Coenonympha tullia, 99 hegon (Amblyscirtes), 151 Asterocampa celtis, 92 Colias eurytheme, 16 henrici (Callophrys), 39 clyton, 93 interior, 17 Hermeuptychia sosybius,