CONTRIBUTIONS
FROM THE CUSHMAN FOUNDATION
FOR FORAMINIFERAL RESEARCH
VOLUME XIX, Part 4 October, 1968
Contents
No. 352. Acceleration of the Evolutionary Rate in the Orbulina Lineage I D. Graham Jenkins ...... _. ___ 133 'No. 353. A New Genus of the Haplophragmoidinae from Malaysia D. S. Dhillon ,- 140 No. 354. Late Eocene and Early Oligocene Planktonic Foraminifera from Port Eliz , abeth and Cape Foulwind, New Zealand M. S. Srinivasan .:, ...... _.... _...... __ 142 No. 355. The Genus Sigmoilopsis Finlay 1947 from Cardigan Bay, Wales. Keith Atkinson ...... _...... ___ 160 No. 356. Preliminary Report on Some Littoral Foraminifera from Tomales Bay, California Don Maurer ...... _.. ___ 163 No. 357. A Taxonomic Note on Massilina carinata (Fornasini, 1905) Keith Atkinson 165 No. 358. A New Species of Pseudoguembelina from the Upper Cretaceous ~ Texas G. C. Esker, ill ...... __ 168 No. 359. Designation of a Lectotype of Globotruncana rosetta (Carsey) G. C. Esker, ill ...... _ ... _. 170 Recent Literature on the Foraminifera Ruth Todd ...... 172 Index to Volume XIX, 1968 ...... _ \83
1968 CONTRIBUTIONS FROM THE CUSHMAN FOU NDATION FOR FORftMINIFERAL RESEARCH In
CONTRIBUTIONS FROM THE CUSHMAN FOUNDA lION FOR FORAMINIFERAL RESEARCH VOLUME XIX, PART 4, OCTOBER, 1968 352. ACCELERATION OF THE EVOLUTIONARY RATE IN THE ORBULINA LINEAGE D. GRAHAM J ENKINS Geology Department, University of Canterbury, Christchurch, New Zealand
ABSTRACT a concommittant increase in the number of aper The stratigraphic ranges of 9 taxa of the Orbulina evo· tures correlated with a decrease in the apertural size. lulionary lineage from 4 Australasian Mi ocene sequences have been plotted against sediment thicknesses and Indl· The first recognized taxon in the Orbulina lin c~\te an evolutionary a cceleration from Globhterillohles eage is Globigerina woodi woodi Jenkins (text fig. trllobus bhUlhericus Todd through to Orbulina. unlversa 2) which has 3-4 chambers in the final whorl and d'Orblgny. A correlation w ith radiometric data indicates that the first 3 taxa of the lineage appeared within about a single umbilical aperture, and the end form is 6 m.y. a nd the last 6 taxa appeared within only 2 m .Y. Orbulina universa d'Orbigny (text fig. 2) which has The possible biological s ignificance of the results and tax~ a spherical test with a large number of small aper onomle rates are discussed. tures scattered over the test surface. All the inter mediate morphologies exist within the lineage, but INTRODUCTION only 7 other named taxa are here recognized. Rep The Orbulina evolutionary lineage has been re resentatives of the 9 taxa have been illustrated from corded by many workers in Miocene sequences the Miocene Muddy Creek section, New Zealand from many parts of the world (see Jenkins, 1965). (text fig. 2). Stratigraphically the lineage can be traced through The taxa within the continually evolving Orbul from the Lower Miocene Globigerina woodi Jenkins ina lineage have been arbitrarily defined, and from to the end form Orbulina universa d'Orbigny in the the writer's experience some are more easily identi Middle Miocene which ranges through to the pres fi able than others. Taxonomically, the difficulties ent. This paper is an attempt to compare the meas of accurate identification and delineation of the urement of the phylogenetic rate of evolution with taxa lie in the region between ancestor and immedi in the O. universa lineage. ate descendant and the problems are especially From a comparison of the stratigraphic initial true of the following taxa in table 1. appearance of 9 named taxa within the Orbulina lineage in 4 stratigraphic sections from New Zea IMMEDIATE ANCESTOR DESCENDANT land and Australia (text fig. 1), it is suggested that there was an increase in the evolutionary tempo in Praeorbulina glomerosa Praeorbulina glomerosa the upper part of the lineage towards the Middle glomerosa (Blow) circularis (Blow) Miocene. There is slight evidence that there was a Praeorbulina glomerosa Praeorbulina glomerosa decrease in the tempo after the appearance of Orb cur va (Blow) glomerosa (Blow) ulina sUlllralis Bronnimann. Globigerinoides trilobus Praeorbulina glomerosa bisphericus (Todd) curva (Blow) ACKNOWLEDGEMENTS Globigerilloides tri/obus Globigerinoides trilobus The writer is indebted to many people, including trilobus (Reuss) bisphericus Todd Professor Alan Wood who provided the academic Globigerina woodi Globigerilla woodi climate at Aberystwyth, Wales during 1956-59 woodi Jenkins connecta Jenkins when this paper was first formulated as part of a Ph.D. thesis, and Dr. W. A. Berggren for providing TABLE 1 the quoted unpublished and radiometric data. Dur Pairs of ancestors and immediate descendants ing the ensuing period I have gained from innumer which are difficult to differentiate where the strati able discussions with many paleontologists. The graphic ranges and morphologies overlap. writer is further indebted to both Mr. R. C. Brazier The separating lines between the above ancestors of the New Zealand Geological Survey who made and descendants have been fixed arbitrarily by vari the illustrations of the specimens and to Miss L. ous micropaleontologists, and one method of pro Fiddes of the Geology Department, University of ducing consistently accurate identifications is by Canterbury, who assembled the figures. relating each taxonomic identification to each holo ORBULINA EVOLUTIONARY LINEAGE type, which is -regarded as the central morphologi The chief morphological changes within the Orb cal type. The writer has discussed the differences ulina lineage are the progressive envelopment of between Globigerilla woodi woodi and Globigerilla the earlier part of the test by the final chamber and woodi COllllecta (Jenkins, 1964), and Blow (1956) 134 JENKINS-EVOL U TION IN ORBULINA LINEAGE
r------r------~r------_,------~IO·S
AUSTRALIA
~------~~~------~------4_------~20·
30·
~------~------~------~~~~~~--~40·
NEW ZEALAND
L------L-______L- ______L- ______~50· 150· 160· 170· 180· TEXT FIGURE 1 Location map discussed the differences between the other taxa A similar clear distinction occurs between Or recorded in table 1. bulina su/uralis and Praeorbulina g/omerosa circu The morphological differences between the other laris where the latter has small sutural openings taxa within the lineage are more definite. Thus the limited to the outline of the Globigerina stage, and ancestor Globigerina woodi cOli/recta differs from the former has the minute apertural openings also its immediate descendant Globigerinoides /ri/obus outside the sutures (text fig. 2). /ri/obus by having a single umbilical aperture as The taxonomic distinction between the two Or opposed to the umbilical aperture plus an additional bulilla species is distinct if 0. ulliversa is defined as aperture or apertures on the spiral side of G. /ri having a scatter of small apertures all over the sur lobus tri/obus (text fig. 2). face of the test (text fig. 2). CONTRIB L' TJONS FROM T H E CUSHMAN FOL'NDATION FOR FORA1\IrNIF I ~RAL R E S E ARCH 135
ORBULINA LINEAGE LAKES NEW ZEALAND ENTRANCE MIDHIRST MUDDY CLlFDEN ,...... -.--.... OIL SHAFT N01 WELL CREEK AUSTRALIA SECTION TARANAKI SECTION
.' ... ' .. ' . ". ': :: . ~feet feet feet feet C0-": . . ORBULINA UNIVERSA / 20 180 320 500. © ~ ORBULINA SUTURALIS 0 90 100 / 0 (@ Q ~ P. GlOMEROSA CIRCULARIS V 4 50 0 100 QCMJ ~ P. GlOME ROSA GlOMEROSA 8 60 235 0 @Q V P. GLOMEROSA CURVA ~ 32 60 825 150 V
988G. TRILOBUS BISPHERICUS ~ 172 no data 1774 950 Q Q Q V G. TRILOBUS TRILOBUS ~ 80 no data no data no data / fitJ8@ ~ G.~ CONNECTA no data no data no data / 152
IRREGULAR IRREGULAR ~ 4ft SAMDl.JW! f100ft SArtRlNG 8 SAMPLING G. WOODI WOOOICD SAMPLING TEXT FIGURE 2 Nine taxa of the Orbulilla lineage illustrated from specimens of the Muddy Creek section. All speci mens approximately X 32 and deposited in the collections of the New Zealand Geological Survey, Low er Hut!, under the numbers FP 1322-1335 inclusive. Stratigraphic separation of individual taxa re corded in feet in 4 named Miocene sequences. 136 JENKINS-EVOLUTION IN ORBULINA LINEAGE
RESULTS Berggren (1968 in press : Table 38) has pub I. Evolutionary rate plotted against sediment lished a chronology of the Cenozoic. A correlation thicknesses. with the Australasian phylogeny of the Orbulilla The stratigraphic ranges of the 9 taxa of the ulliversa lineage indicates that Globigerilla woodi Orbulilla ulliversa lineage have been plotted against woodi appeared at about 28 m.y. B.P., Globigerill thicknesses of sediment of Lakes Entrance oil shaft, oides trilobus bisphericus at about 22 m.y. B.P. and Victoria, Australia, and in three New Zealand sec O. universa at about 20 m.y. B.P. Consequently tions: Midhirst No.1 Well , Taranaki, Muddy Creek the fir st 3 taxa of the O. ulliversa lineage appeared section and Clifden section, (text figs. 1-3) . By within approximately 6 m.y. and the last 6 taxa ap comparison of sediment thicknesses in the fir st 3 peared within only approximately 2 m.y. This pos sections, it can be demonstrated that the evolution sibly crude chronology tends to support the postu of Globigeril1a woodi woodi to Globigerilloides tri lated acceleration in the tempo of evolution within lobus bisphericus took place over relatively thicker the O. ulliversa lineage. sequences of rock as compared with the evolution of G. trilobus bisphericus to O. ulliversa (table 2). DISCUSSION OF RESULTS Rensch (1959) quoted both Darwin (1859) and G. woodi woodl G. trilobus Gaudry (1896) as concluding that in the lowe r to G. trilobus bi sllheric us to bisphericus O. unil'ersa animals the tempo of evolution is generally slow, Lakes Entrallce oil shaft 364 ft. 64 ft. but Rensch considered these generalisations to be Midhirst No. J 2673 + ft. 1470 ft. unsafe. He cited the evidence of the fossil record Muddy Creek 2950+ ft. 850 ft. of the rapid splitting of the Foraminiferida Num mulites and Orbitoides in the Eocene and also nu TABLE 2 merous lines of ammonites. Rensch (1959) further Stratigraphic separation of taxa. It should be stated "that the speed of evolution depends on the noted that in both Midhirst No. 1 and Muddy rates of selection, because in more mobile animals Creek, the sequences do not extend stratigraphi the evolutionary tempo is increased since such types cally low enough to record the initial appearance of are able to adapt more easily." It is therefore pos Globigerilla woodi woodi; this is indicated by the sible to interpret the Orbulilla ulliversa lineage as + sign. demonstrating an increase in the rate of selection from Globigerilloides trilobus bisphericus through The morphological change in the evolutionary to O. ufliversa. lineage from G lobigerilla woodi woodi through to According to Simpson (1953) the ideal measure Globigerinoides trilobus bisphericus involved radi ment of evolutionary rate would be the amount of cal changes in test morphology. The main changes genetic change in continuous populations per year were a reduction in the height of the aperture and or other unit of absolute time. Simpson concluded the test becoming more compact, these resulting in that "there is little hope that fossils can ever be used G lobigerina woodi connecta; the next stage was the for any extensive study of genetic rate of evolution." appearance of an aperture on the spiral side of the Nevertheless fossils can be used to study the rates test at the base of the final chamber, resulting in of evolution and Simpson (1 953) distinguished two Globigerilloides trilobus trilobus. The final cham main methods. The phylogenetic rate of evolution ber showed a gradual envelopment of the rest of measures the changes involved within a lineage, the test; envelopment proceeded to about 15-30% and the taxonomic frequency rate records the of the chambers of the earlier part of the test, and changes in the numbers or frequencies of the phyla was accompanied by an increase in apertures to or larger taxonomic units existing at a given time. about 2-4, resulting in G. trilobus bisphericus (text Evolutionary rates are expressed against various fig. 2). temporal scales. A frequently used scale is based The morphological test changes from Globiger on the thickness of strata, but the validity of the illoides trilobus bisphericus to Orbulina ulliversa method demands a constant relationship between involved a complete envelopment of the earlier not only succession but also thickness of strata and test by the final chamber; the lineage has been di time. Simpson (1953) in a further discussion stated vided into 5 further taxa (text fi g. 2). that the method demands the rate of deposition to A possible interpretation of the evolution in have been approximately constant, but this is im volves the postulate that once the final chamber en possible to prove and deposition is normally not veloped the earlier part of the test, beyond about constant. Simpson concluded "that this method 30% , then the tempo of evolution increased (see can give some information about fluctuations in text fig 2.). rates of evolution and it is a valid way of compar 2. Evolutionary rate plotted against radio ing rates of different characters or of different phyla metric data. during the same span of time .. . ." The present CONTRIBUTIONS FROM THfo':; CUSHMA N FOUNDATION FOR FORAl\'Il N IFERAL RESEARCH 137
i ; ;: g t • !¥ < G. WOODI WOODI • n r- " -i :to G. WOODI CONNECTA !ilo~ G. TRILOBUS TRILOBUS :tor---VI G. TRILOBUS BISPHERICUS :toVl'" c::I: Z P. GLOMEROSA CURVA VI:to-i -i"'~ P. GLOMEROSA GLOMEROSA ~-i:to :to Z P. GLOMEROSA CIRCULAR IS r- n O. SUTURAlIS -:to '" O. UNIVERSA
I Ii i. . if K • •. ~[;',. G. WOODI WOODI n !:.., G. TRILOBUS TRILOBUS 0 G. TRILOBUS BISPHERICUS '"Z P. GLOMEROSA CURVA VI P. GLOMEROSA GLOMEROSA n '"-i P. GLOMEROSA CIRCULARIS 0 O. SUTURALIS Z O. UNIVERSA
i i I I ~, I.t 0' G. WOODI WOODI " 3: G. WOODI CONNECTA a Z :I: '" G. TRILOBUS TRILOBUS ~$ ~ ~VI G. TRILOBUS BISPHERICUS :to-i N P. GLOMEROSA CURVA Zz :to - :too '"r- P. GLOMEROSA GLOMEROSA :to - "..... Z P. GLOMEROSA CIRCULARIS ~ 0 O. SUTURALIS '"r- r- O. UNIVERSA
n i ~, i a ,~;, G. WOODI WOODI ,. 3: G. WOODI CONNECTA c: VlO G. TRILOBUS TRILOBUS ",0 n-< G. TRILOBUS BISPHERICUS -i P. GLOME ROSA CURVA -n o~ P. GLOMEROSA GLOMEROSA z'" P. GLOMEROSA CIRCULARIS '" O. SUTURALIS " O. UNIVERSA
TEXT FIGURE 3 Stratigraphic ranges of 9 named taxa of the O. universa lineage from 4 Australasian Miocene sequences. 138 JENK1NS-EVOL UTtON IN ORUULINA LINEAGE paper is presented to "give some information about function (Huxley, 1957) : a change from the multi fluctuations in the rates of evolution" of the Or chambered botryoidal G lobigerilla to the spherical bulilla ulliversa lineage. test of Orbulilla was the result of selective survival. It is assumed in the present work that the rocks It is possible that functionally the spherical test has a containing the Orbulilla LIIliversa lineage at Lakes buoyancy advantage in the planktonic environment. Entrance, Australia, and in the New Zealand local ities of Clifden, Midhirst No. I, Taranaki and TAXONOMIC RATES Muddy Creek were deposited over the sa me span of Measurements of taxonomic rates are at present time in the Lower to Middle Miocene. As can be crude, but such studies have given approximate es seen from text figs. 2 and 3, the thicknesses of timates of the rate of speciation in certain animal strata are different, indicating the expected different groups. Thus Rhodes (1966) recorded that the rate rates of sedimentation in the various areas. of speciation of Cenozoic mammals has been cal An argument which could be levelled against culated as 500,000 years. the postulated increase in the evolutionary tempo The duration of the Miocene Orbulilla lineage within the Orbulilla Ulliv ersa lineage concerns its from the initial appearance of Globigerilla woodi taxonomic subdivision. For example, it could he through to the initial appearance of Orbulilla wli argued that the G lobigerinoides Irilobus bispirericus versa is approximately 8 m.y. (Berggren, 1968, in to O. ulliversa part of the lineage has been split up press). The number of species recorded in the lin into more taxa than the earlier part of the lineage. eage is 5 and therefore the average speciation rate is 1.6 m.y. A similar objection to studies of evolutionary Published records of evolutionary acceleration rate was recently presented by Rhodes (1966), but within well documented lineages are rare. The pres fortunately there is some support for the writer's ent paper attempts to demonstrate a possible inter postulated acceleration in evolutionary tempo from pretation of the Orbulilla LIIli versa lineage as anoth the statistical work of Scott (1966, 67). In the er example of changes in evolutionary tempo with Clifden section the rate of change of certain char in a well-known lineage. acters plotted against sediment thickness is consid erably more rapid in Praeorbulilla glomerosa than either before or after this species (see Scott 1966, REFERENCES figs. 13,39; 1967, fig. 9). BERGGREN, W. A., 1968 (in press) , Tertiary Boun daries ill Micropaleontology of Marine Bot If a taxonomist lumped the whole of the lineage tom Sediments: Symposium of SCOR (Work taxa from Globigerilla woodi to Orbulilla LIIliversa ing Group 19, September 1967, Cambridge, into only one species, then it could be argued that England.) its mono-taxonomic nature would not demonstrate the evolutionary acceleration. On the other hand, BLOW, W. H ., 1956, Origin and Evolution of the the observable (and measureable - see Scott 1966, Foraminiferal genus Orbulilla d'Orbigny: Mi 67) morphological changes of the lineage plotted cropaleontology, vol. 2, pp. 57-70. against a temporal scale such as rock thicknesses DARWIN, C., 1859, On the Origin of Species by demonstrate a more rapid change in morphology Means of Natural Selection. 2 vols., London. in the upper part of the lineage. GAUDRY, A., 1896, Essai de Paleontologie Philo Huxley (1957) used the word allagenesis to de sophique. Paris. scribe "all types or degrees of biological improve HUXLEY, J., 1957, 'The Three Types of Evolution ment, from detailed adaptation to general organiza ary Processes," Nature (Lond.) vol. 180, pp. tional advance" and slasigell esis "to cover all proc 454-455. esses leading to stabilization and persistence of JENKINS, D . G., 1964, A New Planktonic Foram types and of patterns of organization, from spe iniferal sub-species from the Australasian Low cies up to phyla." Thus the Orbulilla lilli versa lin er Miocene: Micropaleontology, vol. 10, No. eage could be interpreted as demonstrating ana 1, p. 72, text-fig. 1. genesis followed by stasi genesis, with the appear ---, 1965, Planktonic Foraminifera and Terti ance of O. LIIliversa in the Middle Miocene, a form ary Intercontinental correlations: Micropale which has persisted for about 20 m.y. to the present ontology, vol. 11, No.3, pp. 265-277, pis. 1-2. day. This interpretation agrees with the general statement by Sylvester-Bradley (1967) that "peri RENSCH, B., 1959, Evolution above the Species ods of anagenesis alternate with periods of stasi Level. Methuen & Co. Ltd., London. genesis during which modification is slow and the RHODES, F. H . T., 1966, The Course of Evolution: species remain in the same adaptational grade." Proc. Geol. Assoc., vol. 77, pt. 1, pp. 1-53. The Orbulilla ulliversa lineage could be further SCOTT, G. H., 1966, Description of an Experimen interpreted as an advance in the efficiency of a major tal Class within the Globigerinidae (Forami- CONTRIBUT IONS F R OM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEAitC H 139
nifera) - 1: N. Z. Jl Geol. Geophys., vol. 9, SIMPSON, G. G., 1953 , "The Major Features of pp. 513-40. Evolution": New York. ---., 1967, Description of an Experimental Class within the Globigerinidae (Foraminife SYLVESTER-BRADLEY, P. C., 1967, Evolution versus ra) - 2: N. Z. Jl Geol. Geophys., vol. 10, pp. Entropy: Proc. Geol. Assoc., vol. 78, pt. 1, 55-73. pp. 137-147. 140 DHILLON- :MALA YS IAN H APL OPHRAGMOIDINAE
CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH VOLUME XIX, PART 4, O CTOBER, 1968 353. A NEW GENUS OF THE HAPLOPHRAGMOlDINAE FROM MALAYSIA D. S. DHILLON Department of Geology, University of Malaya, Kuala Lumpur, West Malaysia
ABSTRACT DESCRIPTION OF THE NEW GENUS T he new genu s and species, desc ribed below, a loe believed AND SPECIES at present to be r estricted to the brackis h-waler environ ments of two estua ries in M a laysi a. Their mOI"phoJogy i s Family LlTUOLlDAE de Blainville, 1825 described in detail and their taxonomic position s h own to Subfamily HAPLOPHRAGMOIDINAE be among the Haplophrasmoidinae. Maync, 1952 Genus Gobbettia Dhillon, n. gen. INTRODUCTION Type Species Gobbettia wi]fordi Dhillon, n. sp. In the course of study of the Recent brackish Test free, medium size, pl anispiral; wall chitin water foraminifera of three estuaries in Malaysia ous or pseudochitinous, glossy, smooth, translucent undertaken for the purpose of preparing a thesis to transparent, flexible; chambers inflated near pe for the degree of Master of Science, species of fo riphery, increasing in length gradually but notice raminifera were found in the Recent sediments of ably in width as added, chambers not reaching axis two estuaries, the Labuk Estuary, Sabah, East Ma of coiling, leaving an open umbilicus which is great laysia, and the Kuala Sungei Selangor, West Ma ly depressed relative to the peripheral margin; pe laysia, which did not fall within existing generic riphery broadly rounded, not lobulate; typically 8 and specific categories. It was therefore considered chambers in final whorl; sutures slightly or irregu desirable to erect a new genus for these forms. larly curved, depressed; primary aperture a nar The collections that form the basis of the present row, arched, interiomarginal slit , supplementary paper were made by the author during the months aperture a comparatively wide, arched opening sur of May, 1967, and November, 1967. The majority rounded by an everted lip placed low in the aper of the specimens come from Labuk Estuary, while tural face of the last chamber. only one specimen was from the Kuala Sungei Se Remarks.-The genus Gobbellia is closest to langor, and all were found in soft, brown muds. Trochammillila Cushman and Bronnimann, which The salinities of the waters correspond to the plio in the early stages is pl anispiral with a single or haline (9-16 parts per thousand) and brachyhaline multiple aperture. However, Trochammillila later (16.5-30 parts per thousand) brackish-marine water develops rather irregular chambers, and numerous facies of Hiltermann (1949). Temperatures cor apertures that are no longer in an equatorial posi respond to those of shallow, warm tropical waters. tion. According to Helen Tappan (personal com A study of the available literature indicated the munication) they could not be regarded as con new genus to be most closely related to the Haplo generic. phragmoidinae described by Maync (1952), but the The genus is named in honour of Derek J. Gob difference between it and the other members of the bett, Sedgewick Museum, Cambridge, England. Haplophragmoidinae is sufficient to establish its generic distinction. Gobbettia wilfordi Dhillon, n. sp. T ext fi gures la, b DEPOSITORY OF TYPES Test free, medium size, planispirally coiled; wall are deposited in the Re The figured specimens chitinous or pseudochitinous, glossy and smooth, Department of Geology, search Collection of the translucent to transparent; chambers inflated near Kuala Lumpur, West University of Malaya, periphery, increasing gradually in length but rapid Malaysia. ly in width as added; umbilicus open, depressed rel ACKNOWLEDGEMENTS ative to the periphery; characteristically 8 chambers This work was made possible by the research in fin al whorl; sutures slightly or irregularly curved, facilities granted by the University of Malaya and slightl y depressed; periphery broadly rounded, not by a grant from the Geological Survey, Borneo lobulate; primary aperture a low, arched, interio Region, Kota Kinabalu, Sabah, East Malaysia. Sin marginal slit, supplementary aperture a wide, arched cere thanks are due to Helen Tappan, Ruth Todd, opening placed low in the apertural face of the last K. Asano, S. B. Bhatia, and D. J. Gobbett for chamber. Specimen sli ghtly deflated due to flexi examining and commenting on the new genus bility of the test. and species. R emarks.-This species is named in honour of CONTRIBU TIONS }<.... ROM THE CUSHl\lAN FOUNDATION F OR FORAAnNIF ERAL RESEARCH 141
1a 1b 2a 2b
TEXT FIGURE 1 Gobbettia wilfordi Dhillon, n. gen., n. sp.; a. side view of Recent holotype; b. edge view. X63. TEXT FIGURE 2 Gobbettia klompei Dhillon, n. sp.; a. side view of Recent holotype; b. edge view. X63 .
G. E. Wilford, Geological Survey, Borneo Region, Remarks.-This species is similar to Gobbettia Kota Kinabalu, Sabah, East Malaysia. wilfordi in almost all respects, the only difference Dimensions.-Maximum diameter of Recent hol being in the number of the supplementary apertures. otype 0.34 mm., maximum thickness 0.15 mm. This species is named in honour of the late Pro Material.-One hundred specimens. fessor T. H. F. Klompe, formerly Professor and Geographic Range.-At present this species is Head, Department of Geology, University of Ma known only from Kuala Sungei Selangor, and laya, Kuala Lumpur, West Malaysia. Labuk Estuary. Dimensions.-Maximum diameter of Recent hol Types and Repository.-Holotype (Research Col otype 0.33 mm., maximum thickness 0.14 mm. lection No. 5788) and one paratype deposited in Material.-Two specimens. the Department of Geology, University of Malaya, Kuala Lumpur, West Malaysia. Geographic Range.-At present this species IS known only in Labuk Estuary. Types and Repository.-Holotype (Research Col Gobbettia klompei Dhillon, 11 . sp. lection No. 5789) and paratype deposited in the Text fi gures 2a, b Department of Geology, University of Malaya, Test medium size, free, planispiral; chambers not Kuala Lumpur, West Malaysia. reaching to the axis of coiling, leaving an open um bilicus that is greatly depressed relative to the pe riphery; wall chitinous or pseudochitinous, trans REFERENCES lucent to transparent; surface glossy and smooth; HILTERMANN, H ., 1949, Klassifikation der natur chambers greatly inflated near periphery, increasing lischen Brackwasser : Erdiil und Kohle, v. 2, gradually in length but rapidly. in width as added; p.4-8 . typically 8 chambers in last whorl; sutures slightly MAYNe, WOLF, 1952, Critical taxonomic study and curved, depressed; periphery broadly rounded but nomenclatural revision of the Lituolidae based not lobulate; primary aperture a low, arched, inte upon the prototype of the family, Lituola nau riomarginal slit, 2 supplementary apertures with tiloidea Lamarck, 1804: Cushman Found. Fo everted lips around them placed low in the aper ram. Research, Contrib., v. 3, pI. 2, p. 35-56, tural face of the last chamber. pI. 9-12. 142 SRIt'\ lVASAN-EOCENE:-OLIGOCEL\'1!: ~EW ZEALAND FORAMINIFERA
CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH VOLUM E XIX, PART 4, OCTOBER, 1968 354. LATE EOCENE AND EARLY OLIGOCENE PLANKTONIC FORAMINIFERA FROM PORT ELIZABETH AND CAPE FOULWIND, NEW ZEALAND M. S. SRINIVASAN Geology Department, Banaras Hindu University, India
ABSTRACT regarded it as a junior synonym of Slaill/orlhia Twent y-eight species and subspecies of planktonic Fe Hofker, wbich bas a similar aperture and tooth l'aminifera are recorded from Upper E ocene (Kaialan and Runangan) and Lower Oligocene (Whaingaroan) strata of pl ate, but tbe writer agrees witb Loeblicb and Tap the west coast of Sou th ] s land. New Zealand. Five bie pan (1956), witb Beckmann (1957), and witb Reiss s ll'aligraphic zones are defined and correlated with zones (1963) tbat Chilogllembelina bas biserial cbambers el sewhere in t h e wodd. Comparison with the sequence of tbroughout and no triseri al initial cbambers. planklonic Foraminifera in the Caribbean r egion a nd in A comparison of topotypes of C. ololara (Fin East Afri ca is reasonably close. lay) with topotypes of C. cubellsis (Palmer) sbows tbese forms to be identical. INTRODUCTION The Tertiary of New Zealand is remarkably well Genus Zeauvigerina Finlay, 1939 developed in a variety of facies. Much of it con Type Speci tains abundant fossils; it has been divided into 21 es Zeauvigerina zelandica Finlay, 1939 Zeanvigerina zelaudica Finlay stages grouped into 7 series. This paper describes fossil planktonic Foraminifera of the Kaiatan, Run Plate 13, figure 2 angan and Whaingaroan stages (early Upper Eo 1939 Z eallvigerilla zelalldica FINLAY, Trans. Roy. cene to Lower Oligocene) at Port Elizabeth 5 miles Soc. N. Z. 68:541 , pI. 69, fig . 4a-b. north of Greymouth (text fig. I) and at Cape Foul 1964 Zeauvigerilla zelalldica LOEBLI CH and TAP wind near Westport (text fig. 2), on tbe west coast PAN, In Moore; Treat. Inv. Paleont. (C) of South Island, New Zealand. Protista 2(2) :556, fig. 440(4). The figured specimens are deposited in the Geol Remarks.-Finl ay ( 1939) separated tbis genus ogy Department, Victoria University of Welling from EOllvigerilla Cushman because tbe early cbam ton. Sample numbers are the New Zealand Fossil bers show no spiroplectine coiling and tbe final Record numbers based on sbeet districts of the New chambers sbow no tendency to become irregularly Zealand Map Series. The illustrations on plates triserial. Loeblich (1951) and Loeblich and Tap 13-17 were drawn by tbe writer, with tbe aid of a pan (1964) pointed out, bowever, tbat Eouvigerilla camera lucida, at twice their present scale. Tbe has neitber coiled early cbambers nor triserial adult systematic order closely follows tbat of Loeblicb chambers, and tberefore considered Zeauvigerilla a and Tappan (1964). junior synonym. Gallitelli ( 1957) also snggested tbatZeallvigerilla may be a synonym of Eouvigerilla. Toothplates were sbown to be present in EOllvig SYSTEMATIC DESCRIPTIONS erilla by Hofker (1957) and by Gallitelli (1957) Family HETEROHELICIDAE Cushman, 1927 but are not present in Z eauvigerilla; therefore Zea Subfamily HETEROHELICINAE Cusbman, 1927 uvigerina is regarded as a valid genus. Genus Chiloguembelina Loeblich and Tappan, 1956 Because Zeal/vigerilla bas significant similarities Type Species Gumbelina midwayensis to Chiloguembelilla, Reiss (1963) included both Cushman, 1940 genera in a new family, Chiloguembelinidae. Z ea Chiloguembelina cubensis (Palmer) uvigerilla is very close to Chilogl/embelilla, differ Plate 13, fi gure 1 ing only in having an apertural neck ; therefore, 1934 Gumbelilla ClIb ellsis PALM ER, Mem. Soc. both of tbese genera are here included in the fam Cubana Hist. Nat. 8:74, text-figs. 1-6. ily Heterobelicidae. 1940 GlImbelilla OIOlara FINLAY, Trans. Roy. Soc. Finlay (1939) noted it from many localities in N. Z. 69(4) :453, pI. 63, figs. 50-52. New Zealand but considered it to be a good index 1961 Cililoguembelilla orolara HORNIBROOK, N. Z. of Upper Bortonian (Upper Middle Eocene) age . Geol. Surv. Pal. Bull. 34( I): 84. Possibly tbe specimens found at Port Elizabeth are Remarks.-Hofker (1956, 1963) considered that derived ones. This is the first record of Zeauviger Chiloguembelilla has triserial initial chambers and ilia in post-Bortonian Stage Arnold rocks. C"l COLUMNAR SECTtON t 0 ? ~ UNTROON'.6.N Z lim lf.~ t o~ '" ..,., G'auconi t ~ "nJY 1im~a t ()lW, f ~ 1%2 -1% 7
I~,\ F-=--=1 "l I I 0 "'" :0 _ _ Dork brownicih )Jr"'y :> . __ _ E~-=-~,;i l1y n 'll. ... I:!I..x'I.,: II is! "'" Z :;; VERTICAL ",",,0 UORIZONTAL SCA.LE 01' FEET ."" I l'l LOCA.LITY MAP ,.. , F~~:j z :0 500 250" 500 1000 1500 :.:JOO 2500 3000 3so.;, ,oU T :> F-=--"l ~ t" SOU'f .. ,au.wo. "'" .. :0 N[W lU'L ..... O '" l'l rn F--"l I I l'l ''''' :> :ol PORT ELIZABETH SECTION C"l "" I I I :I: f'l4S 1--=-'-= 1 0 1-)47 -=-~~ -- TEXT FIGURE 1 Sketch map from pace and compass traverse and air photographs, cross-section, and columnar section, showing microsample positions (S44 F947 to S44 F967) at Port Elizabeth_ ...,-A COL UMNA I2 S~C11ON L I T~OlOGY .,. OI=-T1ol I ~ .,. II'UT 4SO SAH\.' .t sf-t>. P.5$5 .;;~:}":~:?:.~ QWlrtert'l4ry sa.nd. , t f55' _-=-.::-:.::: '=.-: \ 4co f 55~ =-::- ::~c , tz \\ ~~ f·551 _ ___ _ ,, f.551 --~ :-- t , ~ ~~~ F.55O -- - - M ed. ~UJT\. brown.- qre1.4 ~ , f 5'~ ------modualel.4 ca.tca.reo!,l.s (.!) ,I >50 , ..& ::::. ~~~ sUbtone wUh OCCClsiora.l Z , h
Family HANTKENINIDAE Cushman, 1927 1958 Halltkellilla allstralis HORNIBROOK, (in part) , Subfamily HASTIGERININAE Bolli, Micropaleontology, A(1) :27, 28,29 (table). Loeblich and Tappan, 1957 1961 Halltkenilla alabamellsis HORNIBROOK, N. Z. Genus Glohanomalina Haque, 1956 Geol. Surv. Pal. Bull. 34(1):142, pI. 20, fig. Type Species Glohanomalina ovalis Haque, 1956 424, 429 (in part). Globanomalina micra (Cole) R emarks.-The writer agrees with R a m say Plate 13, fi gures 3, 4 ( 1962 ), who recogni sed that H. australis is a dis 1927 NOllioll micrus COLE, Bull. Amer. Paleont. tinct species from H. alabamellsis. H. australis 14(51):22, pI. 5, fig. 12. differs from H . alabamellsis in side view, having a 1940 Nonioll iota FINLAY, Trans. Roy. Soc. N. Z. more or less angul ar outline, with triangular cham 69(4):456, pI. 65, figs . 108-110. bers and slightly sigmoidal sutures. Well preserved 1947 Globigerina iota FINLAY and MARWICK, N. specimens of H. alabamellsis are rare because of Z. J. Sci. Tech. B 28(4):232. the fragile test. 1958 Globigerilla iota HORNIBROOK, N. Z. J. Geol. Thalmann (1942) and Banner and Blow (1959) Geophys. 1(4) :664-5, fig. 20, 24. gave the range of the genus Halltkellilla (S.S.) as 1958 Globigerille/la iota HORNIBROOK, Micropa Middle and Upper Eocene. In New Zealand Hant leontology, 4(1) : 27-29 , 34, pI. I, figs. 22-24. kellina has been reported by Finlay (1939) and 1961 Pseudo/wstigerilla iota HORNIBROOK, N. Z. Hornibrook (1961) from many localities. Geol. Surv. Pal. Bull. 34(1): 148 . Finlay (1939) described H. australis from a num 1964 Globallomalilla micra (Cole) , LO EB LI CH and ber of Upper Bortonian localities and considered it TAPPAN, In Moore; Treat. Inv. Paleont. (C) to be different from H. alabamellsis. Hornibrook Protista 2(2) :665, fig . 531(6-8). (1958) considered all the occurrences of Halltkell Variatioll.-Runangan specimens show consider ilia from Bortonian to Runangan to represent H . able variation in the inflation of chambers and the australis. But later (1961) he reported H. alabam acuteness of the periphery. A few Runangan speci ellsis from Kaiatan and Runangan at Port Elizabeth mens from Port Elizabeth have a well-developed and Runangan at Oamaru, considering that all bulla covering the aperture. Whaingaroan speci New Zealand Halltkellilla belonged to the alabam mens are smaller than the earlier ones. ellsis group. R emarks.-From a comparison of topotypes of All records of allstralis (at Hampden by Finlay, P. iota (Finlay) with the to po types of P. micra and at Te Uri stream, Pahi, and Kaipara Harbour (Cole) , Berggren (1960) considered these forms by Hornibrook) are from Upper Bortonian strata. to be identical. At Port Elizabeth alabamellsis is restricted to the The writer here agrees with Reiss (1963) that topmost Kaiatan and lower Runangan. At Cape Pseudo/wstigerilla is related to Halltkellina and not Foulwind it occurs only at the base of the section a descendant of Planomalina as thought by Banner which is considered to be lower Runangan in age. and Blow (1959). At Oamaru it occurs at the base of the Totara lime G. micra has been reported by Finlay (1939) and stone [basal Runangan, Hornibrook (pers. comm.)]. by Hornibrook (1961) from many New Zealand Consequently, it appears that the two species dis localities, ranging from Porangan (Lower Eocene) tinguish two rather narrow zones. The H. australis to Runangan (Upper Eocene) in age. At Port zone corresponds to the Upper Bortonian (Upper Elizabeth it ranges up to the Whaingaroan. The Middle Eocene) and the H. alabamellsis zone cor range in New Zealand is thus similar to that re responds to the uppermost Kaiatan and the lower ported by Blow and Banner (1962) in the Lindi Runangan (middle Upper Eocene). area of Tanganyika. G. micra attained world-wide Famil y GLOBOROTALIIDAE Cushman, 1927 distribution during the Upper Eocene and is useful Subfamily GLOBOROTALIINAE Cushman, 1927 for correlation. Genus Turborotalia Cushman and Bermudez, 1949 T ype Species Globorotalia centraJis Cushman Subfamily HANTKENININAE Cushman, 1927 and Bermudez, 1937 Genus Hantkenina Cushman, 192 5 Turborotalia centralis (Cushman and Bermudez) Type Species Hantkenina aJabamensis Plate 14, fi gures 8, 9 Cushman, 1925 1937 Globorotalia celltralis CUSHMAN and BER Hantkenina alahamensis Cushman MUDEZ, Contr. Cushman Lab. Foram. Res. Plate 13, fi gures 5, 6, 9 13 : 26, pI. 2, figs. 62-65. 1925 Halltkenilla alabamensis CUSHMAN, U. S. 1964 Turborotalia celltralis LOEBLICH and TAP Nat. Mus. Proc. 66 (2567):3, pI. I, figs. 1- PAN, In Moore; Treat. Inv. Paleont. (C) 6, pI. 2, fig . 5, p. 3, text-fig. 1. Protista 2(2) :668, fig. 533(6). 146 SRIN[VASAN- EOCENg-OLIGOCgr-;g ~E;W ZEALAND FORAl\l1NIFERA
Remarks.-The high-spired form of T . celltralis T. opima lIalla (Bolli) was first described from and the forms intermediate between T . centralis the opima lIalla zone (Oligocene) of Trinidad. and G. ampliapertllra noted by Bolli (1957) in the Blow and Banner (1962) recorded it from Middle upper part of the Navet formation and from the Eocene to the top of the Oligocene in the Lindi G. cocoaellsis zone of Trinidad probably should be area and also from the Oligocene of Germany. The referred to G. ampliaperlllra pselldoampliapertllra form referred to as Globigerilla sp. by Batjes Blow and Banner, as should be the si mil ar New ( 1958) from the Oligocene of Belgium is probably Zealand forms frequent in the Runangan. this species. In Trinidad T. celltralis makes its first appear ance in tbe upper part of tbe Globigerapsis kllgleri Tnrborotalia aculeata (Jenkins) zone (Middle Eocene), and is replaced by G. ampli Plate 13, figures 7, 8 apertllra pselldoampliapertllra (high-spired inter 1965 Globorotalia illcollspicua aClileata J EN KINS, mediate forms of T. celltralis alld G. ampliapertllra) N . Z. Jour. Geol. Geophy., vol. 8(6):1118, in the upper part of tbe Navet Formation. fig. 13 , nos. 119-125. In the Lindi area of Tanganyika T. celltralis Remarks.-Tn its spinose surface ornamentation ranges from the Middle Eocene within the Cribro- this species closely resembles Trllllcorotaloides col 1/{/lItkellilla dallvillellsis z 0 n e (Upper Eocene) , lactea (Finlay) and T. rohri Bronnimann. Tbe above the last appearance of the genera Halltkell specimens examined by the writer lack the carinate ilia and Globigerapsis. peripbery on the dorsal side and the spines on tbe In New Zealand T. celltralis makes its last ap extremities of the chambers, characteristic of G. pearance above tbat of Halltkellilla and below that illcollspicu(I Howe. of Globigerapsis. From many records of tbis species from various Turborotalia gemma (Jenkins) countries, it appears tbat T . celltralis first enters in Plate 14, figures 3,4, 11 -13 tbe Middle Eocene and makes its last appearance before tbe end of the Eocene. It is replaced by its 1965 Globorotalia gemma J ENKINS, N . Z. Jour. direct descendant G. ampliapertllra pselldoampli Geol. Geophy., vol. 8(6): 1115, fig. 11, nos. apertllra. Study of the transitional stages of T. cell 97-103. tralis to G. ampliapertllra pseudoampliapertllra in Remarks.-In New Zealan:! the Kaiatan forms different parts of the world may prove of value in of T. gemma show a more trochoidal shell than the tbe stratigrapbic correlation of Upper Eocene strata. Upper Runangan and Whaingaroan ones and have a distinctive aperture. Turborotalia increbescens (Bandy) Subbotina ( 1953) recorded the form from tbe uppermost Eocene to the lower part of the Oligo Plate 14, figures 5-7 cene of tbe Caucasus. Blow and Banner (1962) 1949 Globigerilla illcrebescells BANDY, Bull. Amer. recorded it from the Oligocene of Lindi, East Af Paleont. 32(131):120, pI. 23, figs. 3a-c. rica, and from tbe Rupelian of Elmsheim, Germany. Remarks.-Tbe type of T. illcrebescells is from the Jackson (Upper Eocene) of Alabama. It bas Subfamily TRUNCOROTALOTDINAE also been reported from tbe Claiborne Group (Mid Loeblich and Tappan, 1961 dle Eocene), Red Bluff Formation (Lower Oligo Genus Truncorotaloides Bronnimann and cene) and Vicksburg Group (Middle Oligocene). Bermudez, 1953 Blow and Banner (1962) reported tbe species to Type Species Truncorotaloides rohri Bronnimann range from the Upper Eocene to tbe Oligocene in and Bermudez, 1953 the Lindi area of East Africa. In New Zealand T. Truncorotaloides collactea (Finlay) illcrebescells makes its first appearance just above 1939 Globorotalia collactea FINLAY, Trans. Roy. tbe top of tbe basal Kaiatan and makes its last ap Soc., N. Z. 69(3):327, pI. 29, fi gs. 164-165. pearance just below tbe top of tbe Kaiatan Stage. 1940 Globorotalia collactea FINLAY and MARWICK, Its range witbin the type Kaiatan section at Port Trans. Roy. Soc. N. Z. 70(1): 108, 109. Elizabeth is taken to mark a distinct planktonic zone. 1961 Globigerilla col/actea HORNIBROOK, N. Z. Geol. Surv. Pal. Bull. 34(1): 149. Turborotalia opima nana (Bolli) 1965 Trllllcorotaloides col/acrea JENKINS, N . Z. Plate 14, fi gures 1, 2 Jour. Geol. Geophy., vol. 8(5) :843, figs. 1957 Globorotalia opima IU/lUI BOLLI, U. S. Nat. 1-27. Mus. Bull. 215: 118, 169, pI. 28, fig. 3. Remarks.-The record of this species in tbe Remarks.-This form differs from T . illcrebescells Kaiatan at Port Elizabeth is additional . to tbree (Bandy) in its tigbter coiling and much lower existing records from the Kaiatan by Hornibrook aperture. (1961) . CONTR1BUTI ONS FROl\1 THE CUSHMAN FOUNDATION FOR FOR AMINIF E R AL RESEARCH 147
T. col/aclea has been reported from the upper row-rimmed slit, and in the adult appears to be Middle Eocene at Hahajima, Japan (Saito, 1962) . completely sealed. In the Caribbean region, however, this species is In Victoria, Australia, Carter ( 1958) reported restricted to the Lower Eocene .. S. lillaperla having typical flattened chambers from the Brown Creek clays (Upper Eocene). He re Family GLOBIGERINIDAE Carpenter, ported a variety with a tendency to develop swol Parker and Jones, 1862 len and more evenly rounded chambers, in the Subfamily GLOBIGERININAE Carpenter, lower Glen Aire clays (Lower Oligocene). This va Parker and Jones, 1862 riety, referred to by Carter as "G . Iinaperla (round Genus Globigerina d'Orbigny, 1826 chambered form ) ," is probably G. allgiporoides. T ype Species Globigerina bnlloides Parker, A few specimens with smoother wall ornamenta Jones and Brady, 1865 tion and much smaller than the typical G. allgipor Globigerin3 3mpliapertura Bolli oides were observed from the Kaiatan samples. Plate 16, figures 5, 6 The lowest typical G. allgiporoides was found at the base of the Runangan Stage. This species is 1957 Globigeril1a ampliaperlura BOLLI , U. S. Nal. abundant and one of the dominant pelagic species Mus. Bull. 215:108, pI. 22, figs. 5-7 ( not in the Whaingaroan at Port Elizabeth, but not at fig. 4) . Cape Foulwind. 1962 Globigerilla ampliaperlura BLOW and BAN NER, In Eames el.al. Fund. Mid. Tert. Stral. Globigerina brevis Jenkins Correl., pI. 2: 83, pI. XIa-d, XVlIC, fig. 12b. R emarks.-The writer (Srinivasan, 1965) has Plate 13, fi gures 10, II stated that Globigerilla ampliaperlura appears at 1965 Globigerilla brevis JENKINS, N. Z. Jour. the base of the Whaingaroan Stage (base of Oligo Geol. Geophy. , vol. 8(6) : 1100, fi g. 7, nos. cene) , but it since has been found near the top of 58-63. the Runangan Stage (uppermost Eocene). G. am R emarks.- The deeply incised sutures of G . pliaperlura is a direct descendant of G. ampliaper brevis suggest affinities to Globigerapsis illdex. A lura pseudoampliaperlura Blow and Banner, pres few specimens having a small bulla-like structure ent in the lower part of the G. illdex zone. partly covering the umbilical aperture were ob served in the upper part of the G. illdex zone. Globigerina ampliapertura pselldoampliapertura Globigerina euapertura Jenkins Blow and Banner Plate 13, fi gures 12, 15 Plate 17, fi gures 4-6 1960 Globigerilla euaperlura JENKINS, Micropa 1962 Globigerilla pseudoampliaperlura BLOW and leontology, 6( 4) : 350, pI. 1, fig. 8. BANN ER, In Eames el.al. Fund. Mid. Terl. 1961 Globigerilla reliculala HORNIBROOK, N. Z. Stral. Correl., pI. 2:95, pI. XII, A-C XVlIA Geol. Surv. Pal. Bull. 34( 1): 146 (in part), E, fig . 12C. text-fig. b, c (not a, d, e). R emarks.-The s ub s p e c i e s G. ampliaperlura R emarks.-Young specimens of G. euaperlura, pseudoampliaperlura is here restricted to inter common in the lower part of the Whaingaroan mediate forms between T. celliralis and G. ampli Stage, closely resemble G. ampliaperlura Bolli; they aperlura present in the Runangan. It is distin are similar in size and have the same type of aper guished from G. ampliaperlura by its asymmetrical ture. The two species were probably derived from test, which resembles that of T. cell Ira lis, and from an Upper Eocene common ancestor. T. celliralis by its relatively high-spired test and The first appearance of G. euaperlura marks the intraumbilical aperture. beginning of the Landon Series. At Port Elizabeth (Whaingaroan) forms of this species having the Globigerina angiporoides Hornibrook umbilicus covered by a bulla with one opening are Plate 15, fi gure 9 frequent; they resemble forms referred to as C. 1965 Globigerilla allgiporoides HORNIBROOK, N. ullicavus by Jenkins (1960). Z. Jour. Geol. Geophy., vol. 8, no. 5:835, figs. 1a -i, 2. Globigerina ouachitaensis Howe and \ Vallace Remarks.- G. allgiporoides, which seems to be Plate 15, fi gures 4-8 the direct descendant of S. Iillaperla (Hornibrook, 1932 Globigerilla ouachilaellsis HOWE and WAL 1961), is characterised by spherical chambers of LACE, Bull. La. Conserv. Geol., No. 2:74, nearly the same size in the last whorl, and with pI. 10, fig. 7. these chambers more embracing than in S. Iill R emarks.-The species differs from G. o/ficillalis aperla. The aperture in the young is a very nar- in its less tightly and more trochospirally coiled 148 SRl i'\IVASAN-EOCE1'\E-OLICOC E:\l<:; ~E W ZE A L AND F OR A MIN IFERA shell and in its distinctly broader aperture and more Globigerina d . trilocularis d'Orhigny open umbilicus. Plate 17, figures 1, 2 1826 Globigerilla triloeularis O'ORBIGNY, Ann. Sci. Globigerina oflicinalis Subbotina Nat. 7:277 (Unpublished figures of d'Or Plate IS, figures 1-3 bigoy in Fornasini, Rend. Acad. Sci. Inst. 1953 Globigerilla officillalis SUBBOTINA, (Part), Bologna. n.s., vol. 2, fasc. 1, pI. 1, figs. 6, Trudi, UNIGRI, N. S. 76:78, pI. 11, fig. 1 7, p. 12, text fig .). (Holotype) and fig. 2 and 6. Remarks.-The species is characterised by a low Remarks.-The form described by Jenkins ( 1960) spire and rapidly enlarging chambers in the final as G. parva Bolli, from the Globoquadrilla dehiseell s whorl. G. ouachitaellsis is similar but has a more zone, southeast Australia, is probably G. officillalis . trochospiral shell and chambers of more uniform size in the final whorl. Globigerina angustiumbilicata Bolli Forms similar to G. triloeularis were recorded Plate 15, fi gures 10, II from the Eocene San Fernando Formation of Trini 1957 Globigerilla eiperoellsis all gustiumbilicata dad (Bolli, 1957), and from the Eocene Kyoragi BOLLI, U. S. Nat. Mus. Bull. 215:109, pI. and Sakasegawa groups of Japan (Asano, 1962). 22, figs. 12-13. The present specimens do not match G. praebul Remarks.-This species is distinguished from G . loides occ/usa Blow and Banner. officillalis Subbotina by having five chambers in the final whorl instead of four. A few Runangan Genus Globorotaloides Bolli, 19 57 samples contain intermediate forms, and G. offic T ype Species G loborotaloides variabilis Bolli, 1957 illalis is probably ancestral to G. allgustiumbilieata. Globorotaloides suteri Bolli This species makes its initial appearance at the top Plate 14, fi gures 10, 14 of the C. verrueosus zone (Whaingaroan) at Port 1957 Globorotaloides suteri BOLLI, U. S. Nat. Elizabeth, where it is comparatively rare. Mus. Bull. 215 : 117, pI. 27,9-13 (not fig. 14). R emarks.-Frequent to infrequent specimens oc Globigerina labiacrassata Jenkins cur both at Cape Foulwind and at Port Elizabeth. Plate 13, fi gures 13, 14 1961 Globigerina retieulata Stache, HORNIBROOK Globorotaloides turgida (Finlay) (in part), N. Z. Geol. Surv. Pal. Bull 34(1): Plate 17, fi gures 3, 7, 8 146, Text-fig. 4, a, d, e. 1939 Globigerilla lillaperta Finlay, var. IlIrgida 1965 Globigerilla labiaerassata J ENKINS, N. Z. FINLAY, Trans. Roy. Soc. N. Z. 69:125. Jour. Geol. Geophy., vol. 8(6): 1102, fig. 1964 Globorotaloides lurgida J ENK INS, Contr. 8, nos. 64-71. Cushman Found. Foram. Res. 15(3):117, Remarks.-Hornibrook (1965) found G . reticul pI. 8, l3a-c. ata to be a lIomell dubium . G. labiaerassata is one of the more distinctive pelagic species of the R emarks.-From the figure given by Asano, Whaingaroan Stage. It is characterized by a hi gh Globigerina ariakellsis (1962, pI. 20, figs. 5-7) arched aperture, a thick apertural rim, and a rel seems to be closely related to G. IlIrgida (Finlay). atively low spire with four chambers in the last As already pointed out by Jenkins (1964), the il whorl, giving the test an almost square shape. lustration given by Bolli (1957) of G. cf. suteri Specimens from Port Elizabeth are smaller and (pI. 27, fig. 14) shows a flattened bulla with a dis have a less distinct apertural rim than specimens tinct lip, strongly resembling the bulla of G. tur examined by the writer from southern Hawkes Bay. gida (Finlay).
EXPLANATION OF PLATE 13 FIGS. PAGE 1. Chiloguembelilla cubell sis (Palmer). Hypotype, Ff. 699; X 70, (S23f534). 142 2. Zeauvigerilla zelalldiea Finlay. Hypotype, Ff. 702; X70, (S44f951)...... 142 3,4. Globall omalilla micra (Cole) . Hypotype, Ff. 698; X 70, (S44f954) ...... 145 5,9. Halltkellilla alabamell sis Cushman. H ypotype, Ff. 700; X 70, (S23f521). 145 6. Halltkellilla alabamellsis Cushman. Hypotype Ff. 701; X70, (S44f954)...... 145 7,8. Turborotalia awleata (Jenkins) . Hypotype, Ff. 729 ; X 70, (44f947) ...... 146 10, 11. Globigerilla brevis Jenkins. H ypotype, Ff. 730; X 70, (S44f960)...... 147 12,15. Globigerilla euapertura Jenkins. Hypotype, Ff. 731 ; X 44, (S44f966) ...... 147 13,14. Globigerilla labiacrassata Jenkins. Hypotype, Ff. 732; X44, (S44f967) ...... 148 CONTRIB. CUSHMAN FOUND. FORAM. RESEARCH, VOL. 19 PLATE 13
7 8 6
11
14
Srinivasan: Eocene-Oligocene New Zealand Foraminifera CONTRIB. CUSHMAN FOUND. FORAM. RESEARCH, VOL. 19 PLATE 14
7
11 13
Srinivasan: Eocene-Oligocene New Zealand Foraminifera CONTRIB UTIONS FROM THE CUSHMAN FOUN DATION FOR FORA~nN IF ER A L RESEARCH 149
Genus Subbotina Brotzen and Pozaryska, 1961 1960 Globigerilla yeguaensis BERGGREN, Stock T ype Species Globigerina trilocuJinoidcs holm Cont. Geol. 5 (3) : 73-83, pI. 2, figs . Plummer, 1927 1-4; pI. 3, figs . 1-3 ; pI. 4, figs. 1-2; pI. 8, figs. Subbotina Jinaperta (Finlay) 1-5; text-fig. Ii. Plate 16, figures 7,10, 11 Remarks.-Berggren (1960) described the large 1939 Globigerina lillaperla FINLAY, Trans. Roy. degree of variation shown by individuals of yegua Soc. N. Z. 69( I): 125, pI. 13, figs. 54-57. ensis and gave a complete synonymy. 1948 Globigerilla lillaperla DORREEN, J. Paleont. The figured specimen from Port Elizabeth is very 22(3) :299. similar to one of the specimens illustrated by Berg 1958 Globigerina lillaperla HORNIBROOK, Micro gren (1960, pI. 2, fig. la), but has a more prom paleontology, 4(1) :26, 29, 33, pI. I, figs. inent apertural lip. 19,21. This species closely resembles S. Iriloculinoides 1961 Globigerina lillaperla HORNIBROOK, N. Z. (Plummer), the type species of Subbolina, partic Geol. Surv. Pal. Bull. 34( I): 145. ularly in having a distinct apertural lip. The close Remarks.-Brotzen and Pozaryska (1961) pro phylogenetic relationship of yeguaellsis to Iriloc posed tbe genus Subbolilla for Globigerina-like ulilloides was pointed out by Berggren, and yegua forms having a distinct reticulate or pitted surface ellsis is here incluued in the genus Subbolilla. ornamentation and an aperture with a distinct lip and cited G. Iriloculilloides Plummer as its type Subfamily ORBULININAE Schultze, 1854 species. Genus Candeina d'Orbigny, 1839 The species G. lillaperla closely resembles G. T ype Species Candeina nitida d'Orbigny, 1839 Iriloculilloides, the type species of Subbolilla, and Candeina zeocenica Hornibrook and Jenkins has been shown by several workers, including Bolli Plate 15, fi gures 12, 13 (1957), Loeblich and Tappan (1957), and Horni 1965 Calldeina zeocellica HORNIBROOK and JENK brook (1961), to be a direct descendant of G. Iri INS, N. Z. Jour. Geol. Geophy., vol. 8 (5) : loculinoides. Therefore it is here included in the 839, figs . 1-5. genus Subbolilla. It differs from S. Iriloculilloides (Plummer) in being larger and in having later Remarks.-Hornibrook and Jenkins' description ally compressed chambers and a less distinct aper appears to be the first record of Candeina earlier turallip. than Miocene. S. linaperla has been reported from Danian to Its record in Cape Foulwind marks an additional Paleocene in Egypt (Nakkady, 1950), Lower Eo stratigraphic occurrence. cene in Trinidad (Bronnimann, 1952), Eocene on mid-Pacific sea monnts (Hamilton, 1953), Lower Genus Globigerapsis Bolli, Loeblich and Eocene in the Salt Range, Pakistan (Haque, 1956), Tappan, 1957 Lower to Upper Eocene in Trinidad (Bolli, 1957), T ype Species Globigerapsis kugleri Bolli, Middle to Upper Eocene in Australia (Carter, Loeblich and Tappan, 19 57 1958), Middle to Upper Eocene in Japan (Asano, Globigerapsis ind~ (Finlay) Saito, 1962), and Upper Paleocene to Upper Eo Plate 16, fi gures 8, 9, 12 cene in East Africa (Blow and Banner, 1962). 1939 Globigerinoides index FINLAY, Trans. Roy. Judging from its world-wide distribution, as shown Soc. N. Z. 69(1) : 125, pI. 14, figs. 85-88. by these records, it is probably a valuable index 1948 Globigerilloides index DORREEN, J. Paleont. fossil for intercontinental correlation. 22(3) :298, pI. 41 , fig. 2. 1958 Globigerinoides index HORNIBROOK, Micro Subbotina yegnaensis (W einzerl and Applin) paleontology, 4(1) :27-29, 34, pI. I, figs. Plate 16, figures 1-4 11-14. 1929 Globigerina yeguaensis WEINZERL and Ap 1961 Globigerinoides illdex HORNIBROOK, N . Z. PLIN, J. Paleont. 3 :408, pI. 43, fig. 2. Geol. Surv. Pal. Bull. 34( I): 152.
EXPLANATION OF PLATE 14 FIG,. PAGE 1,2. Turborolalia opi",a Ilalla (Bolli). Hypotype, Ft. 709; X70, (S44f952) ...... 146 3, 4. Turborolalia gemma (Jenkins). Hypotype Ft. 710; X 87, (S44f954) ...... 146 5-7. Turborolalia illcrebescens (Bandy) .. Hypotype, Ft. 712; X70, (S44f949)...... 146 8, 9. Turborolalia cenlralis (Cushman and Bermudez). Hypotype, Ft. 713; X70, (S44f956) ...... 145 10, 14. Globorolaloides suler; Bolli. Hypotype, Ft. 714; X87, (S44f959) ...... 148 11-13. Turborolalia gemma (Jenkins). Hypotype, Ft. 711 ; X70, (S44f963) ...... 146 150 SRI N IVASAN- E OCE N E-O LIGOCE N E ~ E'V ZEALAN D FOR AMIN lFERA
Remarks.-This species has deeply cleft sutures Blow and Banner ( 1962) considered that in the and a sub globular shape, as stated by Finlay ( 1939) bullate globigerinids there appears to be no funda and Hornibrook (1958) , and the typical form has mental difference in structure of the bullae, and , as never more than three apertures (one large final the morphology of the primary test is in every case opening, and two others, somewhat smaller, above generically the same, they regarded Catapsydrax and the sutures of previous chambers). This makes it Tillophode/la as junior synonyms of Globigerillita. distinct from G. tropieatis Blow and Banner. The writer agrees with Loeblich and Tappan Blow and Banner (1962) state that G. illdex (1964) in regarding Catapsydrax as distinct from does not extend into the Upper Eocene strata any C lobigerillita and Tillophode/la. Catapsydrax dif where in the world. Their contention may be va li d fers from Clobigerillita in that it has a distinct um for the East African area but may not be applic bilical bu ll a, with one or more accessory infralam able to other parts. Furthermore their suggestion ina I apertures, whereas Clobigerillita has a modi that G . tropiealis has been incorrectly reported as fied final chamber with supplementary apertures. G. illdex from other countries such as Cuba, Israel, Tillophode/la differs from Catapsydrax in having Syria and South-east Australia is not warranted. an umbilical bulla expanding along the earlier su The form illustrated by Carter (1958) shows the tures with numerous small accessory apertures typical globular shape and cleft sutures and also opening beneath its margin. the presence of the three openings, features char acteristic of G. illdex. Catapsydrax echinatus Bani The writer (1965) has found that in New Zea Plate 17, fi gures 10·12 land the genus Globigerapsis makes its last appear ance later than H. alabamellsis and T. celltralis. 1957 Catapsydrax eehillatus BOLLI, U. S. Nat. According to Blow and Banner, in East Africa it Mus. Bull. 215 : 165, 166, pI. 37, figs. 2-5. makes its last appearance earlier than H. alaham R emarks.-It is characterised by distinctly spin ellsis and T. eelltratis. This clearly points out the ose surface ornamentation, which distinguishes it inconsistency in the range of the genus Globiger from C. dissimilis. In New Zealand all the known apsis in these two localities. pelagic species wit h spi nose ornamentation (T. It is certain that G. illdex sti ll existed in New aeuleata, T. eo/la etea, C . primitiva, and C. eehin Zealand and Australia during the Upper Eocene, atus) range up to the Upper Eocene. In the Carib making its last appearance at the top of the Eo bean region, however, the planktonic species with cene. For some reason it evidently does not range spinose ornamentation range from Upper Pale quite to the top of the Eocene in East Africa. ocene to Middle Eocene only. The top of the Runangan Stage at Port Elizabeth (Type locality) has been defined by Finlay and Genus Globigerinatheca Briinnimann, 1952 Marwick (1947) as coinciding with the highest oc T ype Species Globigerinatbeca barri currence of G. illdex. Brii nnimann, 1952 Globigerinatheca d. barri Briinnimann Subfamily CATAPSYDRACINAE Bolli, Plate 17, fi gure 9 Loebli ch and Tappan, 1957 1952 Clobigerinatheea barri BRONNIMANN (part), Genus Catapsydrax Bolli , Locblich and Cushman Found. Foram. Res. Contr. 3 (1): Tappan, 195 7 27, 28, text-fig. 3a-c, g, h. T ype Species Globigerina dissimilis Cushman 1964 C lobigerillatheea barri LOEBLICH and TAP and Berm udez, 1937 PAN, In Moore, Treat. Inv. Paleon!. (C) The genus Catapsydrax was erected for the Clo Protista 2(2) :676, fig. 543(5). bigerilla-like forms having a single umbilical bulla R emarks.-Rare specimens resembling G. barri over the aperture in the ad ult, with one or more were found only at Port Elizabeth. accessory infralaminal apertures. Opinions differ as to whether the umbilical bulla is of use for taxonomic differentiation or merely BIOSTRATIGRAPHY represents variation. Hofker (1961), for instance, Planktonic Foraminiferal Zones interpreted the bulla to represent a closed cyst of Check lists of actual occurrences of planktonic the reproductive phase in the life history. Takay foraminifera at Port E li zabeth and Cape Foulwind anagi and Saito ( 1962) considered that mere de are given in text fig. 3a and text fig. 3b respectively. velopment of a transformal chamber or bulla is not On the basis of the stratigraphic ranges of plank sufficient for specific distinction, and suggested that tonic foraminifera five zones are distinguished (text these modified final chambers may develop in the fig. 4). These biostratigraphic zones, which are reproductive phase or when individual growth is here regarded as being informal units, are defined interrupted by some environmental causes. in terms of American and International codes on CONTRTBUTIONS FROM THE Ct!SHMAN F OUNDATION FOR FORAMINIFERAL RESEARCH 151
STAGE KAIATAN RUNANGAN WHAINGAROAN
Planktonic T. acu/~ala T. incr~b~sc~ns H . ala- I G. a",pliap~rtura ZONE Ibom~n.ri~ Benthonic G. ulIssi B. cf. moodys~ nsis L. po flli~ C. verrll COSilS T. lealal/diea
Foraminifera Fossil No. . " . . . . ." . ." .- .- . · Amm odi~cu~ archimedis (Stache) __ 2 2 I Vuh'u/ina gra1luloSll Finlay ._._ .... ____..... _... _._. __ .____ ._ ... 2 4 Semivulvulil/a eapitata (Stache) _. __. __ 2 Textularia eusp;s Finlay ..... _._ .... _.... _.. 2 Textularia waleotli Srinivasan _... _..... __ ...... _.. _... _...... Gaudryil/a proreussi x reussi ... __ ... __ ... _.. _ Goudryilla reu~si Stacbe ___ .__ .__ .. __ .__ ... I KarrtruUfla obscura Srinivasan _. _____._ .. _ ... _ ... _.. _ ) Art nodosario antipoda (Stacbe) _. _____._ ... _._._. __ ... __ 2 4 4 4 4 I 4 A. cr. kaiala~ nsis Dorreen . ) 2 Tritaxi/il/a ualandiea Finlay ..... _...... _.. 2 ) Ali/ioUl/tlla vel/a; Srinivasan _... _ 2 2 2 Seutulor;~ eli1.llb~ thiae Srinivasan _. __ .__ .____ . ___ ._ ... I 2 ) Dentalil/a obliquesuturala Stacbe ... ____ ._._. ______. Robulus dorOlhiae Finlay _._. __ .______.. _ .. ) 2 ) ) 2 2 Proxi/rotls whaingaroica (Stacbe) .... _ .. _.. __ .__ ._. __ I I ) 4 2 I I I I 4 Sphatroidilla I'ariabifis Reuss _... _...... __ ..... _._ ... _.... _...... _...... ) 2 ) ) 2 4 4 2 S. blllloidts d'Orbigny ..... _.. _... _...... 2 ) ) ) 4 4 ) 4 4 ) 2 Bril a/illa cr. moodysttls;s (Cushman &: Todd) _._ ...... 2 2 2 Lalibolil'ina pofllis (Finlay) ' ___ '_"_" __' ___ '_' __ ' ___ '" ) 2 4 ) L. anostomoso (Finlay) ..... ___. ______.. _._. __ I I Tappanina olson; (Hornibrook) • ______. ___ ._. 2 ) 2 2 2 2 I 2 2 Bulimina II'Ullcont lla Finlay _. __ 4 4 4 2 4 4 4 4 4 4 4 2 Nov;ul'a borlolara (Finlay) 4 4 I 4 4 4 4 4 2 4 Recluvigtrina poslprandia Finlay 2 I R . slriatiss;nla (Stache) ..... _... _.. __ .... _. ___ 2 2 DiscopuMnulina apposila (Finlay) ._____ .. __ ._ .. __ 2 ) 2 D . seopos ( Finlay) ______._._._. ______. __ Sip/lOlIina australis Cushman .__ . _____ . __ _ I Chiloguembelina eubensis ( Palmer) ..... _ .. _.. _. ___ .__ .. _._._ 2 2 2 2 4 4 4 4 4 4 4 4 2 ) 2 ) 2 2 Zelllll'igeri/ia ltlandica Finlay ..... _ .. _...... _...... __ ... _... .. I I Gioballomalilla micra (Cole) . .. .. _...... _...... _ ...... _ I 4 4 4 4 4 2 2 Halllke/lina alabamensis Cushman ._ .. ____... __ ._._.__ ._. ) 2 TlITborolalia cenlrolis (Cushman &: Bermudez) __ I I I )
T. illcrebescens (Bandy) ___ no .. _ ... __._._. __ 2 ) ) 2 T. opimanana (Bolli) ._ .. _. ____. ___ . ____ _ I I 2 2 2 T . acufeato (Jenkins) _... _ ... _._.~ .... _.. ___ .. _... _._ .... __ ._ .. _... __._._ 2 2
T. g~mllla (Jenkins) ...... _._. __ .M .. __ • ___._ I I I I I 1 I G/obigerilla ampliaperlura Bolli ..... _..... ___ .... _ .. ___. _ I 2 ) 2 2 2 2 2 G. ampfiaperluro pseudoampliaperlura Blow &: Banner 2 2 I G. allgiporoides Hornibrook ___ ~" ___' _____ "_'''_''M'_'' __ 2 2 2 2 4 4 4 ) 4 2 G . brr\'is Jenkins _____. ______I I I G. eauap~rtura Jenkins ""'_"_'_' ___' __"_'_' __ "'_ 2
G. angusliumbiliealo Bolli ... M __ ..... _ ..... _._. __ .... _._ .. _ .. _.M __.... _ G. oOkillolis Subbotina .. __ ...... _...... _..... _.. G. ollochilat!nsis Howe & Wallace _... _ ..... _...... _.M. __ .__ .. _... G. labiacrassata Jenkins _. __ ...... _... _.. _.. _... ___. __ ._._ ... _.. _._ G. d . Irilocufaris d'Orbigny ____ .______. __ Tmncorolafoides colloctea ( Finlay) ______I G loborololoides sutui Bolli .__ .. ____~ ___._._ .. __.. 1 I I G. lurgida (Finlay) .. __ ...... __ ._. __ .... _._. __ ._ ..... _._. __ ...... _._ .. _. 1 I I I I I Sflbbo/il/a Iillopula (Finlay) ..... _._ ... _...... _ 2 2 2 2 2 ) 2 4
S. yeg"aellsis Wienzierl &: Applin ..... _.. _... _ ..... __ .M .. _ ... __._. I I I Ca"d~ina zeocenica Hornibrook &: Jenkins .___ . ___ ._ 1 2 I I I 2 )
Globigerapsis index (Finlay) _. ______M_ 4 4 4 4 4 4 4 4 4 4 4 4 4 Calopsydrax echinalus Bolli ______I I I Globigerillathtca d. barr; Bronnimann __ ._._._.__ .__ 1 C;bicid~s ps~udocanvexus Parr ..... _._ .... ____ .. __ ... __ .... ___ 1 I I I 2 C. parki Finlay ...... _._ ... _._ .... _..... _..... 2 3 2 2 ) 4 ) 2 2 4 2 ) C. semip~rloralus Hornibrook ..... _... _. __ .. __ ...... _._. __ ._ 2 2 2 ) 2 ) ) 2 C. per/oralus (Karrer) ..... _.... _. __ .__ .. ______.. _. ) 2 2 ) 4 4 4 2 C. hampd~n~fUis Hornibrook ______2 C. \'errucosus Finlay _ .. ______._. __ 2 2 I C. collinsi Finlay ___. ___ ._._._.. __ ... _._ ... _ .. _.. _.... ___ .... _. I I 2 C. kafuri/ormis Hornibrook I £lfipsoglandulina lobialo (Schwager) ..... _.M .... _...... __ ...... I I 2 I R Olaliatina sulcigua (Stacbe) _... _...... _ ...... _._ ... _. __ .. __ ... 4 4 4 ) 2 ) ) 2 2 Anomalinoides semiterts (Finlay) ___ .. _.. _.. _._. ___ .__ _ ) 2 Melonis dorretni (Hornibrook) _. ______.____ 4 4 ) ) 4· ) ) ) Vel/aellO l~alandico Srinivasan ____ .__ .___ .. __. __ _ I Cerob~rlina kakahoica Finlay ___.... __ ..... ___ .. _...... _ ..... _ .. __ 2 2 ) 2 2 )
Note: Only stratigraphically important benthonic s~ies are listed. Key: ( .. ) = nil specimens; 1 = less than 5 specimens; 2 = 5 to 10 specimens; 3 = 10 to 20 specimens; 4 = more than 20 specimens
TEXT FIGURE 3a Check L ist of the Late Eocene a nd E arly Oligocene foraminifer a from Port Elizabeth section. New Zealand 152 SRINIVASAN-EOCENE-OLIGOC E N E N EW ZEALAND FORAMINlFERA
STAGE R UN A NGA N W HAI NGA ROAN Planktonic ::"~~~: I G. i"d~x G. a",p/iapuwra I ~f/Si.f z 0 N E ------~~~------~------B~ntho n ic L. pomjs C. I·t.'rrueosus T. ua/al/diea
0 _ .. to " .., "PO .. ~: :; ::~ ~, ., . . . .~ ~, ~, ~, ~, Fossil No , . ~ ., '" Fo ram i n i f~ra . '" '" -... "'" '" .~ " ...... , Trip/asia marll'icki Loeblich & Tappan ...... ~. 2 2 2 TU lIIlaria subrltombica (Stac b~ ) ...... _ 2 2 I I Galldryina r(' ussi Stache ...... _ ...... _ 2 2 I Arellodosaria antipodo ( Stacb~) ...... _...... 2 2 2 2 2 2 2 3 2 2 2 3 222212 233222 Qllillqucfoculilla zealandica Srinivasan ~ ... _...... 2 2 2 2 2 2 Mifiofineifa I'elloi Srinivasan . 3 222 222 D ellialina obliquullturalo Stacbe I Robulus gyroscalprlls (Stach~ ) .. I I I 1 I I I _ 2 I I Palmufu bil'ium Finlay _... _.. _... _ ProxiJrons wl/aingo roico (Stache) ..... _... __ .._ ... _ Bolidlldla ('/('go lls Parr M~'." .. ~._~ ... ~._'~""_" " "~ ..... ~ .. ~ ...... _ Spho('roidilla \'aritlbifis R~u ss ...... _ .. ~ .... _.. _ S. bulloidu d'Orbigny ..... ___..... ___.. _. __... _.~ ... _.. _... _._ ...... 22323 2 2 2 2 22 33332 222222322 2 2 2 2 L atiboli villa pontis (Finl ay) .~~~" ____"._"._"._,,. 2222222 2 I I I LtJlibolil'illtl all/stomostl (Finlay) ..._ .. _._._ ... _.. _... _ 1 I I I I 1 1 1 TClppallilla olsolli ( Horni brook ) ._.___ ...... _...... _ .. _.... _ 2 1 I I 2 2222333 2 3 2222333 23 2 3 3 2 2 Bufi",intl trtll/condla Finlay .__ ... _~.~.M._._M .... ~ ... * ...... _ 3 2 222 3 2 2 2 3 2 2 2 2 2 2 Non'm'o bortorara (Finlay) .. M_ ...... _~.M. __ ..... 4 4 4 4 3 3 3 2 2 2 1 I I R(,Cllll"iguillu poslprandio ( Finlay) ___ .. _. __ ..... ___ .__ _ I 4 4 4 1 4233324 1 R . slTialiss;ma (S t ach~) ..... ~ ...... ~ ...... _...... _...... _ 14 4 4 4 4 4 4 443443 Pijp('rsia gracilis Srinivasan ..... ~ .... _...... _._...... 1 I D iscopuil"imtiirru apposila ( Finlay) 2 3 2 2 2 2 3 3 3 444 3 3 3 3 223 2 333222222322 2 D . scopos (Finlay) ...... _ ...... _ 1 Glabrtltdlu erassa Dorr~~n I G. {illfayi Srinivasan ...... I Porosororafia tail/Ilia (Dorr~~ n ) 2 I 2 2 222222 I,,: : 1::: 2 Porosorolnlia knIT; (Hornibrook) I 1 1 1 P. okokotllSis (Hornibrook) ...... 1 2 2 Chilogll('mbt!lil1a ClIb~ns is (Palmer ) ...... _...... I I I 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 1 1 1 Globanomalil/a micra (Cole) ...... __ ... _.. __ ._.__ ._._ ...... _...... ~_._._ 2 2 I H UlI/k(',,;lIa flfabamtnsis C ushman .....~ .... _...... _...... 2 Tllrb~"oUtlio c~lII ralis (Cushman & B ~rm ud~z) ..... _ ...... _... 2 T . opima " mlG (Bolli) ..... _..... __ ...... ~ ..... __~~ ..... ~ ...... ~_ ....._~ .. _._
T . g('IIIIIIO ( J~nk i n s) ..... M ...... _ ...... _ ...... _ .. _. __~ __.... __~ ...... _ Gfobigtrinfl ampliaptrtllro Bolli _._._._._... ___.... _... _ . 1 1 2 2 ... I G. lIlIIpiiaptrlllra pSt.'udOtllllpliaptrtura Blow & Bann~r ... _ I .. I
G . (mgiporoidn Hornibrook M ... M .. M ... ~ .. _ ..... __• ___.... __... _. 2 2 2 34423 22 321 G. eaupt'rlllra J~nk i n s ..._M ... _.M~ .. __ ... _~~ .. _ ... _ ...... G. ollacililt/(,IISis Howe & Wa ll ac~ ...... _ .. _~ ...... _~ .... _~ ...... 2 2 - I I I 2 1 ... G. ol/icillalis Subbotina ... ~ ..... _..... 4 3 2 2 2 GloborOlaloides Sllltri Bolli ._ .. ~ ... _.. _...... __..... _...... 2 G. IIIrgid// (Finlay) 2 S"bbotilla lil1t1p('rla (Finlay) ...... ~ ...... _...... _ 2 4 Cmrd('ill(l Ztoceniea Hornibrook and Jenkins I 4 2 Globigu apsis illdex (Finlay) ...... _...... ~ .. .. 3 2 3 2 2 4 4 4 4 4 3 3 2 4 4 4 444 Cibicid('s pSt'lIdocol/I·txIlS Parr ...... ~ 2 1 1 1 1 1 1 I 1 I I C. parki Finlay . • 2 2 2 I 222 1 22 1 22 3 222 2 2 2 2 3 2 1 C. puJora/lIs Karrer . 2 3 33 3 3 2 2 2 2 2 2 I I 1 3 3 3 3 3 2222 2 2 3 3 2 2 C. r('rrIlCOSIIS Finlay ...... _.. .. 1 I H fllkyardia bart rum; Parr ..... ~ ... 1 Ellipsogflllldlifina labiato Sc b wag~r I Rotalial illo suleigua (Stac h~) ..... ~ .. _._ .. _. __...... _.. .. 2 2 3 2 2 2 2 2 2 3 2 Allomalilloides St.'mittrt.'S (Finlay) I I I I I I M ('/ollis dorrulli ( Hornibrook) ...... ~ .... __ ... _ ... ~ ...... ~ 2 2 23 33333 3 3 4 3 3 3 3 2 3 333 22 33 2 2 3 2 2 M . pocimaoricllm Srini vasan ... __ ...... _... __ ...... _...... __ . 2 1 1 ... Ct r/llolamarckina clarki Srinivasan .. _..... __ ...... __. __ ~ .. _ .. . Vl!llu('na ualandico Srinivasan .. __ .~ ...... ~ __._~ .. _.. _.. _ ... _...... 3 3 I 2 I 23 1 : 3 Ctroberlina knknlroica Finlay ___~_'M 2 11 2222 1 2233 222 1 3 2 ... NOI~ : Only stratlgrapblcally important benthonic spcci~s are listed. Key: C... ) = nil s pecim~n s ; I = I~ ss tban S specimens; 2 = 5 to 10 specimens; 3 = 10 to 20 specim~ns ; 4 -= more tban 20 s pecim~ns TEXT FIGURE 3b Check L isl of the Late E ocene and E a rly Oli gocene foraminltel'a f l'om Cape Foulwind section, New Zealand stratigraphic nomenclature and also in terms of the pearance of Turborotalia illcrebescells (Bandy). biostratigraphic zones proposed by Vella (1964). The zone is not represented at Cape Foulwind. At They are here treated in ascending order. Port Elizabeth it is represented by the lowest ex posed 120 ft. of the Kaiata mudstone. The base of (i) Tlirborotalia aculeata Zone the zone is not exposed; the top is immediately be The Tlirborotalia aculeata zone has its boundary low the lowest occurrence of T. iflcrebescefls. The defined by the initial appearance of the zone fossil, planktonic assemblage of T. aculeata zone at Port and its upper boundary is defined by the initial ap- Elizabeth is the same as that in the upper part of CO ~TRIB U TION S FROM THE CL"SHMA N F OL"NDA TION F OR 1" O RA~nNJFERAL RESEARCH 153
New Zealand Part Elizabeth Ra nges of Zone name Age Stages Planktonic Foraminiferal Zones fossils at Port Elizabeth
,~ c ~ <; " Globigerina am pliapertura •"- "g Whaingaroon .2 •0 ~ --' (5'" E Zone • G
Globigerapsis index
. ~ .","• •c .S Runongon Zone •E c G " •c ~• u -2 "0 u• W • • Hantkenina alabamensis ~• <; ,• ::: c. Zone u c. ·s :::> • ;; ...: Turborotalia increbescens ~, u Zone • Koioton .• ...: T urborota lia aculeata Zone TEXT FIGURE 4 Stratigraphic zones distinguished by planktonic Foraminifera In the Upper Eocene and Lower Oligo cene at Port Elizabeth. the upper greensands (Basal Kaiatan) at McCul (iii) Halltkellina alabamellsis Zone lough's Bridge, south Canterbury, except that Glo The Halltkellina alabamellsis zone is defined by boquadrilla primitiva (Finlay) does not occur at the total range of the zone fossil. The zone is rep Port Elizabeth. This similarity plus the presence in resented ' both at Cape Foulwind and Port Eliza both areas of a form intermediate between Gaudry beth. At Cape Foulwind it is about 60 ft. thick and illa prorel/ssi Finlay and G. rellssi Stache indicates rests unconformably on granite; the oldest part is that the T. aculeata zone at Port Elizabeth is sim missing. At Port Elizabeth it is about 220 ft. thick, ilar in age to the upper greensands at McCullough's conformably overlies the T. illcrebescells zone, and Bridge. It appears that G. primitiva makes its last contains an extremely rich planktonic fauna. appearance within the Gal/dryina reussi zone (Bas The following become extinct within the H. ala al Kaiatan) and does not range quite to the top of bamellsis zone: Catapsydrax echinatus Bolli, Zea that zone. Therefore, the T. aculeata zone at Port uvigerina zelandica Finlay. Globigerilla angipor Elizabeth is considered to be slightly younger than oides Hornibrook makes its initial appearance in the the upper greensands at McCullough's Bridge and middle part of the zone. Halltkenilla itself is rare, to represent the upper part of the G. reussi zone and may be missed if not searched for carefully. (Basal Kaiatan). The H . alabamellsis zone represents a short pe riod at the close of the Kaiatan and the beginning (ii) Turborotalia illcrebescell s Zone of the Runangan during which several faunal The Turborotalia illcrebescells zone is defined by changes took place. the presence of the zone fossil and is present only at Port Elizabeth, being represented by a thickness (iv) Globigerapsis illdex Zone of about 510 feet of strata in the lower part of the The G lobigerapsis index zone has its lower boun section and conformably overlying the T. acul dary defined by the extinction of H. alabamensis eata zone. and its upper boundary by the extinction of G. The following species become extinct within the illdex. The zone is represented at Cape Foulwind T . illcrebescells zone: T. aculeata (Jenkins) and by 200 ft. of shallow-water micaceous mudstone Trullcorotaloides collactea (Finlay). and at Port Elizabeth by about 590 ft . of relatively The top of the T. illcrebescells zone marks the deep-water calcareous mudstone. At both places base of the upper Kaiatan Stage. it rests conformably on the H. alabamellsis zone. 154 SRINIVASAN-EOCENE-OLlGOCENE NEW ZEALAND FQRAloDNIFERA
~ ~ m A R N 0 l 0 LANDON ;Ji '" '<> N ~ ~ '"m 0 c 0 ~ ~ > 5 ' ... ~ ,... ~ ~ Q ~ 0 m > ~ -g ~ z - ~ 0 ~ PLANKTONIC FORAMINIFERA
cnilofvU16~/iIl8 ClIDehJ/6 ( Palmer )
Z~auv'l~ri"" Je/8nak8 F i n l ~y -- G/(JDan0I118/illtJ micu ( Co le) Hantkeninll alaMmensis CLlshman
Turbofo/6lia centre/Is (C ushma n & Bermudt~) TurbortJMlia inudescens (Ban dy )
Tu,#(JI'oI8//a opima h8/Ja (Bolli) . - Tur6oroltl/ia 8clJl~l1t4 (Jenkins) - TurJorol8/i6 femm8 (Jenkins) - . 7hll1C'orofl1/oides Coleetea (Finlay) - , Glo6iferil18 11111/li4perlur4 Bolli Glo6iferint1 4l11pll4pertvrr1 pseud06mpl16plrlllN Blow 6: Banner - , Glopreril1t1 tlnf,p CJI'o,aes Hornibrook GIOPi(~riI14 j;1"~ris Jenkin'
Glopif~ril16 ~(JQpert(/l"t1 Jenk ins
Glopiferil1t1 oV4c1I1Y4~I1$i$ Howe & Wa llace - 610P1erl114 officll14/is Subbotina GloPiferin4 6nfQslivmpilicQ/" Bolli Glopijeril14 14Pi6C'/'t1s$611 Jen kins - Gloporol6lo,aes silferi Bo lli fl/oPOI"ot4/oities 1111"1'"'' ( Finlay ) Suph/il1t1 liMpedt1 ( Finlay)
SuMofin8 "P"ns/s (W, inJ,,1 & Applin)
GloP'f~ft1pSI$ ind~x (Finlay )
Cal1t1eil14 Jeocenic~ Hornibrook 6. Jenk ins C,t"lsytlrtJx eC'hint1tll$ Bolli - 6/oPlferll1llhect1 c/ . hrri Bronniman n -
~ ~ a~if! , ~ S· ~ ... ,... ,... ,.. ,.. , ~ . ~ i Ii- ~ . ~ ~ , f ~ g a f '" ~ a ". ~-t"!'f , ~ t "'n ;P ~ ,.. i: ,.. ,.. ., ~~l=D '. ~ ~ ! ~ v- S"'5- ., 11- ~ ..., 3 iC :::0 ..... ~ ....--:;:: ;- -r~...,~~~ '"' ~ ,... I: ~ ,.., ~., ,.. ~ oDpoa.:F~ ,.. ~ ,.. ~ ~ a .,~ ~ . ~: '" ~"r-. frg-
~ 2:' = = ~ ~[':E" n ~ c~, _. , ~ ~ eg' ~ ~ !I.JI ' 0 3=-~ ~ n ~ ~ ~ '" :::o. a"'O ~ 100"1- ~ ~ ;:; ,:::0 t-~ ·
TEXT FIGURE 5 Correlation chart showing some significant planktonic Foraminifera in the Upper Eocene - Lower Oligocene of New Zealand. Arrows indicate taxa also found in younger New Zealand strata. Cl N. West and West North o New East Assam Middle South Austra lia Western Indonesia Burma Pakistan Boluch - Age Z Zealand Corribeon Africa India East >-l New Australia N. W. India istan :- til Zealand Blow and Glaessner, Von der c: Present Bolli, Carter, Wade, Eames, Grimsdole, Bonner 1942 Vlerk ::! Chapman and Nogappa, 1959 o Stages paper 1957 1962 1958, 1959 1964 1955 1951 1951 z Crespin, 1935 rn
o~ G. G. G. :<: 'linoperta' Schwezettaw Baroi! Lower Lower Lower Lower >-l ampliaperturo ompliapertura Se llii :I: Whoingoroan Unit - 3 ? 'c' Stage l'J Zone Stage 'Series' Nori Nod Oligocene Oligocene Cl Zone Zone Zone c: ? ? rn :I: '">: G. z G. 6or14nii o'J coc06ensis Zone with c: G. 'inde,,' z Zone Pellalispirll o ------Unit - 2 Giralllo- tb' Stage Yaw Kapili 'Tapti Upper > indu Zone >-l limestone with Stage Stage S.rie.· Chocolate (3 Runangan with Pelloti.s pira with with with Cloys Z Zone G. Pellatispiro Pellatispiro Pellotispiro Pello'ispiro 'J JonvilleMu Pello'ispiro o ------aod Dod :0 Zone 'olabomensis Upper Upper G. H. c. 6 H. Unit - 1 compresso· : ------alabamensis danvillensis Eocene Eocene alabamensis semiinvalut4 Zone > Zone ------aod 5 Zone z G. II. 'J alobomensis l'J semi-involuto : T. > t' Zone increbescens : l'J Kaiatan ------? ? rn lone l"l ____ 1-___ f- ----L _____ ~ Cl :I: T. T. T. ? ? Pondaung Upper aculeata ronri ronri Zone Stage Prong Zone Zone f-----1------1.---- Upper 10' Stage Lime.tane Middle Middle Bartonian Eocene Eocene
TEXT FIGURE 6 .... \A New Zealand Upper Eocene to Lower Oligocene Foraminiferal zones and some overseas correlations. \A 156 SRINIVASAN-EOCE N E ·OLIGOCENE N E"\V ZEALAN D F OR AMIN JFERA
The following become extinct in the lower part zones suggested by Vella (1964), the zones fall of the G. illdex zone: Turborotalia celltralis (Cush into the following categories: man and Bermudez) ; G. officillalis Subbotina, and Rallge ZOll e: T. illcrebescells zone, H. alabam G. turgida (Finlay) ; G. brevis Jenkins. G. ampli ellsis zone. apertura Bolli makes its initial appearance towards Successive appearallce Wile: T. aculeata zone. the upper part of the zone. Successive disappearance zone: G. illdex zone, The top of the G. index zone is, by definition, the G. ampliapertura zone. top of the Runangan Stage and of the Arnold Series. Age and Correlation (v) G lobigerina ampliapertura Zone In the Tertiary strata of the Tropical-Tethyan and The Globigerilla ampliapertura zone is marked Indo-Pacific belts, the larger foraminifera are dom by the presence of G. ampliapertura. The lower inant; they are the main basis for time-stratigraphic boundary is defined by the extinction of G. illdex divisions and for intercontinental correlation. Dur and S. lillaperta, both of which range commonly ing the Upper Eocene to Lower Oligocene epoch, throughout the underlying zone. The G. ampli New Zealand was situated well outside the Tropi apertura zone conformably overlies the G. index cal-Tethyan and Indo-Pacific belts, and to date zone both at Cape Foulwind and at Port Elizabeth. Asterocyciilla is the only genus of larger foraminif At Cape Foulwind the zone is represented by 150 era known to have invaded the New Zealand area feet of shallow-water micaceous mudstone and the during the Eocene epoch. The earliest attempt at upper part of the zone is missing. At Port Eliza extensive long-distance correlation by means of beth the G. ampliapertura zone is about 180 feet planktonic foraminifera was Grimsdale's (1951) thick and contains a rich pelagic fauna. correlation of the Tertiary rocks of western Europe Sampling was not high enough at Port Elizabeth with those of the middle East. The works of to determine the upper boundary of the G. ampli Glaessner (Indo-Pacific, 1942), Finlay (New Zea apertura zone. Consequently, it was not possible to land, Australia, Indo-Pacific, 1947), Crespin (Aus determine those species that are restricted to the tralia, 1950) , Hamilton (Pacific, 1953), Carter (S. zone or that make their last appearance at the top Australia, 1958), Hornibrook (New Zealand, of the zone. 1958), and Nagappa (Pakistan, India, Burma, It may be possible to distinguish an upper sub 1959) have enriched the knowledge of foraminif zone of the ampliapertura zone itself, based on the era in dating and correlating the Tertiary strata first appearance of G. labiacrassata Jenkins. As throughout the world . Eames et al. (1962) gave this species is rare at Port Elizabeth more work is extremely valuable data for correlation of zones required elsewhere regarding its first appearance based on planktonic foraminifera with those based before it can be formally proposed; however, a on large foraminifera. Probably the most useful well defined zone based on benthonic foraminifera, standard for middle Tertiary intercontinental cor the Tritaxililla zealalldica zone, corresponds to this rel ation is the sequence of zones based on plank interval of time. tonic foraminifera established in the Caribbean region by Bolli (1957). Nature 0/ the biostratigraphic ZOll es Comparison of the ranges of New Zealand Low The planktonic foraminiferal zones fall into the er Tertiary planktonic species with those of the following categories in the American Code and ac Caribbean and East Africa is not entirely straight cording to the International Subcommission on forward. This is probably due to latitudinal differ Stratigraphic Terminology (Hedberg, ed., 1961): ence. The Upper Eocene - Lower Oligocene foram Rallge ZOll e: T. increbescells zone, H. alabam inifera of New Zealand are different in a number ellsis zone. of respects from those described from Trinidad by COli current rallge Zone: T. aculeata zone, G. Bolli (1957) and from East Africa by Blow and illdex zone, G. ampliapertura zone. Banner (1962). An instance of such differences is In relation to the usage of the biostratigraphic the complete absence of the common tropical index
EXPLANATION OF PLATE 15 FIGS. PAGE 1-3 . Globigerilla officillalis Subbotina. Hypotype, Ff. 715 ; X 87, (S44f954) ...... 148 4-6. Globigerina ouachitaellsis Howe and Wallace. Hypotype, Ff. 716; X70, (S44f950) . .... 147 7,8. Globigerilla ouachitaellsis Howe and Wallace. Hypotype, Ff. 717; X70, (S44f950) . ... . 147 9. Globigerilla allgiporoides Hornibrook. Hypotype, Ff. 718 ; X44, (S44f964) ...... 147 10, 11. Globigerina angustiumbilicata Bolli. Hypotype, Ff. 719; X70, (S44f962) ...... 148 12,13. Candeina zeocellica Hornibrook and Jenkins. Hypotype, Ff. 720; X 70, (S23f555) ...... 149 CONTRIB. CUSHMAN FOUND. FORAM. RESEARCH, VOL. 19 PLATE 15
5 6
7
9
Srinivasan: Eocene-Oligocene New Zealand Foraminifera CONTRIB. CUSHMAN FOUND. FORAM. RESEARCH, VOL. 19 PLATE 16
Srinivasan: Eocene-Oligocene New Zealand Foraminifera CONTRTBUTIONS FROM THE CUSHMAN 1.... OUNDATION FOR FORAMIN IFERAL RESEARCH 157 forms Truncorotalia, Cribrohalltkellilla and several ACKNOWLEDGEMENTS species of Hantkenina. On the other hand common Sincere thanks are expressed to Dr. Paul Vella New Zealand forms such as G. allgiporoids Horni for his helpful advice and criticism during the writ brook, G. brevis Jenkins, and G. labiacrassata ing of this paper. Thanks are also extended to Mr. Jenkins are not known in the tropical regions. N. de B. Hornibrook for allowing full access to lit Finlay and Marwick (1947) and Hornibrook erature and collections held at the New Zealand (1958, 1959) considered the Kaiatan and Runang Geological Survey, and to Dr. D . G . Jenkins for an Stages [marked by the latest occurrence of rare many interesting discussions. Discocyclilla (now A sterocyclilla) , Hantkellilla al This research was carried out during the tenure abamellsis and Globigerapsis illdex] to be Upper of a New Zealand Commonwealth Scholarship at Eocene in age. They considered the Whaingaroan Victoria University of Wellington, New Zealand. Stage (with no Halltkellilla and Globigerapsis) , con REFERENCES formably overlying the Runangan, to be Lower ALLAN, R. S., 1948, Geological correlation and pa Oligocene. leoecology: Geol. Soc. Amer. Bull., vol. 59, The writer (Srinivasan, 1965) has previously p. 1-10. correlated the boundary between the Arnold and ASANO, K. , 1962, Tertiary Globigerinids from Ky Landon Series of New Zealand with the Eocene ushu, Japan: Prof. Enzo Kon'no memorial Oligocene boundary as described by Blow and volume. Sci. Reports of the Tohoku Univer Banner in the Lindi area, East Africa. sity, second series (Geol.) special volume no. The New Zealand upper Eocene - Lower Oligo 5, 1962. cene planktonic foraminiferal zones (text fi g. 5) BATJ ES, D. A . J ., 1958, Foraminifera of the Oligo are correlated with a si mil ar set of zones estab cene of Belgium: Inst. Roy. des Sci. Nat. de lished in the Caribbean region by Bolli (1957) and Belgique, Mem. 143, pp. 1-188, plates 1-13. in the Lindi area of East Africa by Blow and Ban ner (1962) . The correlation is mainly based on the BECKMANN , J. P., 1957, Chilogllembelina Loeblich stratigraphic occurrences of the following plank and Tappan and related Foraminifera from the tonic species present in all three areas: Lower Tertiary of Trinidad, B.W.I.: U. S. Nat. Mus. Bull. 215, pp. 83-95, pI. 21, text-figs. Globallomalilla micra (Cole) , Turborotalia 14-16. opima lIalla (Bolli), Turborotalia illcrebescells (Bandy), Turborotalia centralis (Cushman BERGGREN, W. A ., 1960, Some planktonic Foram and Bermudez), Hantkellilla alabamensis inifera from the Lower Eocene (Ypresian) of Cushman, Subbotilla linaperta (Finlay), Glo Denmark and Northwestern Germany: Stock bigerapsis illdex (Finl ay). Globigerilla ampli holm Contributions in Geology, vol. v:3, pp. apertura pseudoampliapertura, G. ampliaper 41-100, pIs. 1-13. tura and G. euapertllra. BOLL!, H . M., LOEBLlCH, A. R., TAPPAN, H., 1957, The accompanying chart (text fi g. 6) is an at Planktonic foraminiferal families, Hantkenin tempt to correlate the Upper Eocene-Lower Oligo idae, Orbulinidae, Globorotaliidae and Globo cene strata of New Zealand with the strata else truncaniidae: U. S. Nat. Mus. Bull. 215, pp. where, primarily on the basis of planktonic foram 3-50. inifera. Where data on planktonic foraminifera are BOLLI, H. M., 1957, Planktonic Foraminifera from inadequate, as in the cases of Baluchistan, Pakistan, the Oligocene-Miocene Cipero and Lengua for India, Burma, Indonesia, and northwest and west mations of Trinidad, B.W.I.: Ibid, pp. 97-123, ern Australia (Tethyan and Indo-Pacific belt), pis. 22-29, text-figs. 17-21. larger foraminifera have been utilized in compila ---., 1957, Planktonic Foraminifera from the tion of this chart. Eocene Navet and San Fernando formations
EXPLANATION OF PLATE 16 FIGS. PAGE 1-3. SlIbbotina yegllaellsis (Weinzierl and Applin). Hypotype, Ff. 703; X70, (S44f958) . 149 4. SlIbbotina yegllaellsis (Weinzierl and Applin). H ypotype, Ff. 704; X70, (S44f953) ...... 149 5, 6. Globigerina ampliapertllra Bolli. Hypotype, Ff. 705; X70, (S44f961) ...... 147 7, 10,11. Subbotina lillaperta (Finlay). H ypotype, Ff. 706; X70, (S44f956) ...... 149 8,9. Globigerapsis index (Finlay). Hypotype, Ff. 707; X44, (S44f959) ...... 149 12. Globigerapsis index (Finlay). Hypotype, Ff. 708; X35, (S44f959) ...... 149 158 SRINIVASAN-EOCENE-OLlGOCENE NEW ZEALAND FORAMINIFERA
of Trinidad, B.w.!.: Ibid, pp. 155-172, pIs. HORNIBROOK, N. DE B., 1958, New Zealand upper 35-39, text-figs. 25-26. Cretaceous and Tertiary foraminiferal zones BRON NIMANN, P., 1960, The genus Halltkellilla and some overseas correlations: Micropaleon Cushman in Trinidad and Barbados, B.W.!.: tology, vol. 4, No. I, p. 25. Jour. Paleontology, vol. 24(4), pp. 397-420. ---, 196 1, Tertiary foraminifera from Oamaru CARTER, A. N., 1958, Pelagic Foraminifera in the district (NZ.): part I, Systematics and distri Tertiary of Victoria: Geol. Mag., vol. 95(4), bution: New Zealand Geol. Surv. Pal. Bull. pp. 297-304, table I. 34(1), pp. 1-192, pI s. 1-28 . - --, 1958, Tertiary Foraminifera from the Aire ---, 1965, Globigerilla allgiporoides n. sp. from district, Victoria : Geol. Surv. Vict. Bull. No. the Upper Eocene and Lower Oligocene of 55, pp. 1-76, pI. 1-10. New Zealand and the status of Globigerilla CR ES PIN, I., 1950, Australian Tertiary microfaunas allgipora Stache, 1865, New Zealand Jour. and their relationships to assemblages else Geol. & Geophys., vol. 8, No. 5, pp. 834-83 8. where in the Pacific region: Jour. Paleontol ---, and JENKINS, D. G., 1965, Calldeilla zeo ogy, vol. 24(4), pp. 421-429. cenica Hornibrook and Jenkins, a new species EAMES, F. E., 1951 a, A contribution to the study of of Foraminifera from the New Zealand Eo the Eocene in Western Pakistan and Western cene and Oligocene, Ibid., 839-842. India; the geology of the standard sections in the Western Punjab and in the Kohat district; JENKINS, D. G., 1960, Planktonic Foraminifera Geol. Soc. London Quar. Jour., vol. 107, pI. from the Lakes Entrance oil shaft, Victoria, Australia: Micropaleontology, vol. 6, No.2, 2, pp. 159-171. pp. 345-371. ---, 1951 b, Discussion of the faunas of certain standard sections and their bearing on the clas ---, 1963 , The Eocene-Oligocene boundary in sification and correlation of the Eocene in New Zealand : New Zealand Jour. Geol. & Western Pakistan and Western India: Geol. Geophys., vol. 6, No. 5, pp. 707. Soc. London Quar. Jour., vol. 107, pI. 2, pp. ---, 1964, A history of the holotype, ontogeny 173-196, 5 pIs., 3 text-figs. and dimorphism of Globorotaloides turgida EAM ES, F. E., BANNE R, F. T., BLOW, W. H., CLARK, (Finlay): Contr. Cushman Found. Foram. W. J., 1962, F undamentals of mid-Tertiary Res., vol. 15(3 ), pp. 117-121. stratigraphical correlation (Cambridge Univer ---, 1965, Planktonic foraminiferal Zones and sity Press, London). New Taxa from the Danian to Lower Miocene FINLAY, H. J., 1939c, New Zealand Foraminifera: of New Zealand. New Zealand Jour. Geol. & the occurrence of R zelwkilla, Halltkellina, R o Geophys., vol. 8, No.6, pp. 1088- 1126. taliatina and Zeauvigerilla: Roy. Soc. New LOEBLICH, A. R. , JR. , and TAPPAN, H., 1964, In Zealand Trans., vol. 68 , pp. 534-543. Moore, R. C., ed .: Treatise on invertebrate FINLAY, H. J., and MARWICK, J ., 1940, The divisions Paleontology (C), Protista 2(1) & (2),900 p. of the Upper Cretaceous and Tertiary in New NAGAPPA, Y., 1959, Foraminiferal biostratigraphy Zealand: Roy. Soc. New Zealand Trans., vol. of the Cretaceous-Eocene succession in the 70(1), pp. 77-135. India-Pakistan-Burma region : Micropaleontol ---., and , 1947, New divisions of the ogy, vol. 5, No.2, pp. 145-192. New Zealand Upper Cretaceous and Terti ary: PARR, W. J., 1947, An Australian record of the fo New Zealand Jour. Sci. Tech. B 28(4), pp. raminiferal genus Halltkenina: Roy. Soc. Vic 228-236. toria Proc. 58(N.S.), pts. I-II, pp. 45-47. GLA ESSNER, M. F., 1943, Problems of stratigraphic RAMSEY, W. R. , 1962, Hantkenininae in the Terti correlation in the Indo-Pacific region: Roy. ary rocks of Tanganyika: Contr. Cushman Soc. Victoria, Proc., new ser., vol. 55, pt. I, Found. Foram. Res., vol. XIII, pI. 3, pp. 79-89. pp. 41-80. SAITO, T ., 1962, Eocene planktonic Foraminifera GRIMSDALE, T. F., 1951, Correlation, age determi from Hahajima (Hillsborough Island): Pale nation, and the Tertiary pelagic Foraminifera: ontology. Soc. Japan Trans. Proc. n.s., No. World Petr. Congr., III, Proc. Sec. 1, pp. 463- 45, pp. 209-225, pIs. 32-34. 475. SAlTO, T., and BE, A. W. H., 1964, Planktonic Fo HAMILTON, E. L., 1953 , Upper Cretaceous, Terti raminifera fro m the American Oligocene: ary and Recent Planktonic Foraminifera from Lamont Geol. Observ. Contr. No. 718, Sci. mid-Pacific fl at-topped sea mounts: Jour. Pa vol. 145, pp. 702-705. leontology, vol. 27, No.2, pp. 204-237. SRINIVASAN, M. S., 1965, The Eocene-Oligocene CONTR IBU T IONS FROM THE CUSHMAN FOUNDAT ION F OR FO RA~n N I F E RAL RESEARCH 159
boundary in New Zealand. Nature, vol. 207, THALMANN, H. E., 1942, Foraminiferal genus Hant No. 4996, pp. 514-575. kenina and its subgenera: Amer. Jour. Sci., ---, 1966, Descriptions of New Species and vol. 240, pp. 809-820. Notes on Taxonomy of Foraminifera from the VLERK, I. M. VAN DER, 1955, Correlation of the Upper Eocene and Lower Oligocene of New Tertiary of the Far East and Europe: Micro Zealand : Trans. Roy. Soc. N. Z., Geol. vol. 3, paleontology, vol. 1, No.1, pp. 72-75. No. 17, pp. 231-256, 6 pis. --- , 1966, Foraminifera and Age of the Type WADE, M., 1964, Applicatfon of the lineage concept Section of the Tahuian Stage: New Zealand to biostratigraphic zoning based on planktonic Jour. Geol. & Geophys., vol. 9, No.4, pp. 504- Foraminifera: Micropaleontology, vol. 10, No. 512. 3, pp. 273-290. 160 ATKINSON- SlonOILOPSlS FROM \VALES
CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH VOLUME XIX, PART 4, OCTOBER, 1968 355. THE GENUS SIGMOILOPSIS FINLAY 1947 FROM CARDIGAN BAY, WALES KEITH ATKINSON Department of Geology, University College of Wales, Aberystwyth, Wales, U.K.
ABSTRACT Sigmoililla schilimbergeri SILVESTRI, 1904, Mem. Four species of Sigmoilullsis Finlay are desc l'ibed (rom Accad. Pont. Rom. Nuovi. Lincei., vol. 22, the Recent depo s its of Card igan Bay. Two of the sl)ecies. pp. 267, 269. (figs.-Schlumberger, 1888, pI. Sil;lJIoiloJlSiS lIloyi and S igmoilollsis woodi, are new. 7, figs. 12-14; p. 481, text fig. 6; p. 482, text INTRODUCTION fig. 7). In 148 surface sediment samples from Cardigan Descriplioll .-Test free, elongate oval in outline, Bay, Wales, four species of Sigmoilopsis Finlay over twice as long as broad, posterior end sub 1947 were found . The sediments were collected by rounded, apertural end slightly truncate; test not the Department of Geology, Aberystwyth, between greatly compressed, rounded tri angular in cross sec 1963 and 1965 during numerous cruises of the R.Y. tion, unequally biconvex in apertural view and pe AlIllIr. These samples form part of the Cardigan riphery subrounded. Sigmoidal chamber arrange Bay - Irish Sea Research Project at present being ment, chambers more distinct on the side with carried out in the Department. greater convexity than on the other, up to six visible Immediately after collection, forma ld ehyde solu externall y, final chamber extending only slightly be tion, buffered with borax, was added to the sedi yond the penultimate one at the posterior end. Su ment to preserve the living organisms. In the lab tures indistinct, only roughly marked, slightly de oratory the samples were washed in clean water and pressed. Inner layer of wall calcareous; exterior stained using rose Bengal (Walton, 1952). No spe arenaceous, composed of fairly large grains for the cimens of Sigmoilopsis were found with the aper size of the test which do not protrude beyond the tural plug of dark, red-stained protoplasm which is outline so that the finish is relatively smooth, most indicative of living foraminifera. of the grains are colourless, markedly few are dark The holotypes of the new species have been pre pyroxenes and amphiboles. Aperture terminal, oval sented to the British Museum (Nat. Hist.). Para to semi-circular on a short neck with a phialine lip types are deposi ted in the author's collection at the and a small, broad, bar-like tooth parallel to the Museum of the Department of Geology, Aberyst base of the aperture. All Cardigan Bay specimens wyth. The type localities given in the systematics are sectioned were microspheric. Cardigan Bay - Irish Sea Project sample locations. Dimell siolls of th e figured specimell.-Length SYSTEMATIC PALAEONTOLOGY 0.67 mm.; breadth 0.30 mm.; thickness 0.23 mm. Superfamily MlLIOLACEA Ehrenberg, 1839 Locality.-C.B. 555; 52 ' 13'34" N; 4' 20'48" W. Family MILIOLIDAE Ehrenberg, 1839 Depth .-7 fathoms. Subfamily QUINQUELOCULININAE Observed rallge ill dimell siolls.-Length 0.35- Cushman, 1917 0.70 mm.; breadth 0.20-0.40 mm.; thickness 0.17- Genus Sigmoilopsis Finlay, 1947 0.25 mm. Sigmoilopsis schlumbergeri (Silves tri), 1904 Occlirrellce.-14 stations, depth range 0 to 13 Plate 18, figur es la, I b, Ie fathoms. Plwlispirilla celata (Costa) BRADY, 1884, Chall. R emarks.-Before the recognition of this as a Repts. Zool., vol. 9, pp. 197, 198, pI. 8, figs. distinct species (Silvestri, 1904), there can be little 1-4. doubt that many authors grouped S. schilimbergeri
EXPLANATION OF PLATE 17 FIGS. PAGE 1,2. G lobigerilla ct. trilocularis d'Orbigny. Hypotype, Ft. 721; X 70, (S44f951) ...... 148 3. Globorotaloides tllrgida (Finlay). Hypotype, Ft. 723; X44, (S44f965) ...... 148 4-6. Globigerilla ampliapertllra pselldoampliapertura Blow and Banner. Hypotype, Ff. 722; X70, (S44f956)...... 147 7,8. Globorolaloides tllrgida (Finlay). Hypotype, Ft. 724; X70, (S44f965)...... 148 9. Globigerillatheca cf. barri Bronnimann. Hypotype, Ft. 725; X44, (S44f949)...... 150 10-12. Catapsydrax echillatlls Bolli. Hypotype, Ft. 726; X 70, (S44f949)...... 150 CONTRIB. CUSHMAN FOUND. FORAM. RESEARCH, VOL. 19 PLATE 17
Srinivasan: Eocene-Oligocene New Zealand Foraminifera CONTRIB. CUSHMAN FOUND. FORAM. RESEARCH, VOL. 19 PLATE 18
( . / ~: .le.-_e
Atkinson : Sigmoilopsis From Wales CONTRIB UTIONS FRO:\:I THE CUSHMAN FOUNDATION FOR FORAMIN IFERAL RESEARCH 161
(Plallispirilla celalo sensu Brady 1884) under P. periphery. It differs from S. schlumbergeri (Sil celala Costa 1855 .. Costa's type has a shorter test vestri) in being more compressed, concavo-convex and more conspicuous sutures. and having less well defined sutures. The subacute A lectotype for this species was designated by periphery distinguisbes it from both Silvestri's spe Loeb1icb and Tappan ( 1964, p. 466) as the speci cies and S. wa llgolluiensis Vella. Compared with men figured by Brady 1884 from Porcupine Station S. filliayi Vella it has a much more irregular, 23, west of Ireland at 630 fatboms. The Cardigan coarsel y arenaceous wall and sub-acute periphery Bay specimens have a ge nerally more roughened as well as possessing a phialine lip. appearance than Brady's type; this may be some di rect result of the shallower depth from which they Sigmoilopsis wanganuiensis Vella, 1957 were coll ected . Brady recorded the world-wide Plate 18, fi gures 2a, 2b, 2c depth range as 28 to 2435 fathoms. Sigl11 oilopsis wallgalluiellsis VELLA , 1957, N. Z. Ge- 01. Surv. Pal. Bull. no. 28, p. 20, pI. 4, figs . Sigmoilopsis moyi, sp. nov. 67-70. Plate 18, figures 3a, 3b, 3c, 3d Descriptioll .-Test free, sub-circul ar to ovate in Descriplioll .-Test free, ovate in outline, basal outline, compressed, sides onl y gently convex in end broadly rounded with the last chamber extend apertural view ; peri phery rounded. Chambers sig ing beyond the penultimate one, apertural end moidally arranged, indistinct, fin al chamber extend gently attenuated , compressed concavo-convex in ing posteriorly beyond the penultimate one, cham cross section; periphery sometimes keeled, more bers not greatly inflated; sutures very indistinct. In often subacute to subrounded. Chambers fairly ner layer of wall calcareous and exterior arena distinct, sigmoidall y arranged, four visible external ceous composed of medium sized grains with an ly, gentl y inflated. Sutures onl y roughly visible, de even finish, including occasional dark minerals. pressed. Wall arenaceous, fai rly coarse witb many Aperture terminal, oval to crescentic, on a short angular grains scattered unevenl y over the test. neck with a rudimentary pl ate-like tooth and phi Aperture terminal on the attenuated fin al cbamber, aline lip. witb short neck, phialine lip and short bar-like tootb. Dimensiolls of Ih e fig ured specil11 ell.-Length Dimell siolls of Ih e hololype.- Lengtb 0.73 mm.; 0.6 mm.; breadth 0.52 mm.; thickness 0.27 mm. breadth 0.57 mm.; thickness 0.30 mm. LocaIiIY.-C.B. 552; 52°13'27" N ; 4°22'3 2" W. LocalilY of Ih e holotype.-C.B. 561; 52° 16' 16" Depth .-9 fathoms. N; 4 ° 16'00" W. Observed rallge ill d il11 ell siolls.-Length 0.27- Deplh.-9 fathoms. 0.65 mm.; breadth 0.23 -0.55 mm.; thickness, 0.14- Type level.- Recent. 0.29 mm. Types.-Holotype presented to the British Mu Occurrell ce.-9 stations, depth range 0 to 12 seum (Nat. Hist.); paratypes deposited in the au fathoms. thor's coll ection at the Department of Geology, Remarks.-This species is si milar to S. fli lltii Aberystwyth. (Cushman ) but is broader, has a more irregul ar Observed rallge ill dimell siolls.-Length 0.25-0.75 grain size and a shorter neck. The Cardigan Bay mm.; breadth 0.20-0.59 mm.; thickness 0.12-0.31 specimens are smaller than the type and have a mm. shorter neck and a less pronounced tooth structure. Localities of paralypes.-C.B. 502 : 52°17'21" N , Vella (1957) recorded this species as very per 4°25'54" W, depth 13.5 fathoms; C.B. 561: 52"16' sistent from 43 to 63 fathoms in Cook Strait. 16" N , 4° 16'00" W, depth 9 fathoms. Sigmoilopsis woodi, sp. nov. Occurrellce.-ll stations, depth range 2 to 13.5 fathoms. Plate 18, fi gures 4a, 4b, 4c, 4d R el11arks.- This species varies in the amount of Descriptioll.- Test free, sub-circular to ovate in compression, the concavity of the side having few outline, unequall y biconvex, rounded triangular in er chambers visible, and in the acuteness of the cross section, not compressed; periphery rounded.
EXPLANATION OF PLATE 18 FIGS. PAGE l a, b, c. Sigm oilopsis schlul11bergeri (Silvestri), 1904, length 0.67 mm...... 160 2a, b, c. Sigm oilopsis wallgalluiell sis Vella, 1957, length 0.60 mm...... 161 3a, b, c. Sigmoilopsis m oyi sp .. nov., holotype, length 0.73 mm. 161 3d. Sigl11oilopsis moyi sp. nov., paratype, thin section, X 60...... 161 43, b, c. Sigl11 oilopsis woodi sp. nov., holotype, length 0.48 mm...... 161 4d. Sigl11 oilopsis woodi sp. nov., paratype, thin section, X 47 ...... 161 162 ATKINSON-SIGlUOILOrSIS FROM WALES
Chambers sigmoidally arranged, very indistinct, our' along the Continental Shelf of the south gently inflated, fin al chamber not extending posteri east coast of Australia: Trans. Roy. Soc. S. orly beyond the penultimate one, in transverse sec Aust., Vol. 65(2), pp. 145-211. tion the chambers are oval to circular; sutures ex CORYELL, H. N., and RIVERO, F. C., 1940, A Mio ceedingly indistinct, slightly if at all depressed. cene microfauna of Haiti: Journ. Paleont., Wall coarsely arenaceous, with many dark coloured Vol. 14, pp. 324-344, pis. 41-44. grains, exterior rough. Aperture terminal, oval to CUSHMAN, J. A., 1929, The Foraminifera of the semi-circular, a little raised above the penultimate Atlantic Ocean. Pt. 6: U. S. Nat. Mus. Bull. chamber on a short neck with a phialine lip and a 104. narrow, bar-like tooth structure. ___, 1946, The Genus Sigmoilina and its spe Deri vato nominis.-In honour of Professor Alan cies: Contr. Cush. Lab. Foram. Res., Vol. 22, Wood. pt. 2, pp. 29-45. Dimensions of the holotype.-Length 0.48 mm.; breadth 0.37 mm.; thickness 0.23 mm. CUSHMAN, J. A., and RENZ, H. H ., 1947, The Fo Locality of the holotype.-C.B. 561 ; 52°16'16" raminiferal Fauna of the Oligocene Ste. Croix N; 4 ° 16'00" W. Formation of Trinidad, British West Indies: Depth.-9 fathoms. Cush. Lab. Foram. Res. Sp. Pub. no. 22., Type level.-Recent. ELLIS, B. F., and MESSINA, A., 1940, Catalogue of Types.-Ho!otype presented to the British Mu Foraminifera: Amer. Mus. Nat. Hist. seum (Nat. Hist.) ; paratypes deposited in the au FINLAY, H. J., 1947, New Zealand foraminifera: thor's collection at the Department of Geology, Key species in stratigraphy - No.5: N. Z. Jour. Aberystwyth. Sci. Technol., sec. B, vol. 28, no. 5, p. 270. Observed range of dimellsions.-Length 0.29- FLINT, J. M., 1899, Recent foraminifera. A des 0.58 mm.; breadth 0.23-0.42 mm.; thickness 0.21- criptive catalogue of specimens dredged by the 0.25 mm. U. S. Fish Commission Steamer Albatross: U. Localities of paratypes.-C.B. 646: 52°19' 14" N, S. Nat. Mus. Ann. Rept., pt. 1, pp. 249-349. 4° 13'00" W, depth 10 fathoms; C.B. 421 : 52°25' HERON-ALL EN, E., and EARLAND, A., 1932, Foram 05 " N, 4°10'44" W, depth 8 fathoms; C.B. 736: inifera; Part 1 - The ice-free area of the Falk G.R. 556 748 (SN 60U.K. Nat.Grid), beach sample. land Islands and adjacent seas: Discovery Occurrellce.-IO stations, depth range 0 to 12 Repts., Vol. 4, pp. 291-459. fathoms. R emarks.-This species is very variable in cross LERoY, L. W. , 1941, Small foraminifera from the section and some thin sections show a gradual re Late Tertiary of the Netherlands East Indies; duction in the degree of triangularity with growth Pt. 1: Colorado Sch. Mines, Quart., Vol. 36, (plate 18, fig. 4d). Most large specimens are still no. 1, pp. 12-62. very triangular. The amount of neck development LOEBLICH, A. R., JR., and TAPPAN, H., 1964, Trea also varies. Externally this species resembles Flillt tise on Invertebrate Paleontology. Part C, ilia crassatina (Brady), but they differ in chamber Protista 2 (1): Geol. Soc. Amer. arrangement. It is similar to S. flintii (Cushman) SCHLUMBERGER, C., 1888, Note sur Ie genre Plani but is less compressed and has a shorter neck. It spirilla: Soc. Zoo!.. France, Bull., Paris, vol. 12, differs from S. wallgalllliensis Vella in being pp. 475-488, pI. 7, text figs. 1-8. rounded triangular in cross section and having a SILVESTRI, A., 1904, Ricerche strutturali su alcune coarser arenaceous wall. forme dei Trubi di Bonfornello (Palermo): Accad. Pont. Romano Nuovi Lincei, Mem., ACKNOWLEDGMENTS Roma, vol. 22, pp. 235-276. This and associated work was supported by a THALMANN, H. E., 1960, An index to the genera grant from the Natural Environment Research and species of the Foraminifera, 1890-1950: Council. The author wishes to thank Dr. J. R. George Vanderbilt Found., Stanford Univ., Haynes for critically reading the manuscript. Calif., pp. 1-393. VELLA, P., 1957, Studies in New Zealand Foram REFERENCES inifera - Part 1 - Foraminifera from Cook BRADY , H . B., 1884, Report on the Foraminifera Strait: New Zealand Geol. Surv., Paleont. dredged by H.M.S. 'Challenger' during the Bull., no. 28. years 1873-1876 : Chall. Repts. Zool., Vol. 9. WALTON, W. R., 1952, Techniques for recognition CHAPMAN, F., 1941, Report on Foraminiferal of living foraminifera: Contr. Cush. Found. soundings and dredgings of the F.I.S. 'Endeav- Foram. Res., vol. 3, pt. 2, pp. 56-60. CONTRIB UTI ONS F ROM THE CUSH i\IAN F OUN DATI ON FOR F ORAMI N IFERAL RES E AR CH 163
CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH VOLUME XIX, PART 4, OCTOBER, 1968 356. PRELIMINARY REPORT ON SOME LITTORAL FORAMINIFERA FROM TOMALES BAY, CALIFORNIA DON MAURER University of Delaware, Marine Laboratory, Lewes, Delaware
ABSTR ACT mentation, test abrasion, faunal composllIon (eco T went y-one species a r e listed f r om T om a les Bay, Cal logic homogenity), density, size-frequency distri iforni a, and a p r ol)osed paleoecologic study i s outlined, bution, and sediment association. These criteria Results of the preli minar Y work showed t h at the fau na of T omal es Bay is vel'Y simi l a r t o tha t in Sun se t Bay, Or egon, would be evaluatec! by checking the results of the fossil analysis with the distribution of modern fo INTRODUCTION raminifera in the bay. The purpose of the study is to report the prelimi Tn addition to possible paleoecologic implica nary results of an investigation of littoral foramin tions, the present research is of interest for several ifera in Tomales Bay, California, the first phase of reasons. Considerably richer in species than orig a proposed paleoecologic study. The macrofauna inally reported by Bush (1935), the fauna contains and flora of the Pleistocene Millerton formation at least twenty-one species rather than the nine she surrounding Tomales Bay, 40 miles north of San reported. On the basis of studies in other bays and Francisco at 38° 15'N. Lat., have been described by estuaries (Phleger 1960) , it would be expected that Dickerson (1922) and Weaver (1949). Although the marsh at the southern end of the bay would foraminifera have been reported from these depos yield a characteristic assemblage of arenaceous its, best developed as cliffs and headlands jutting in species. to Tomales Bay (Santa Rosa, California Sheet The present research extends the distribution of 1963), none have been listed or described. How nearshore foraminifera north from San Francisco. ever, Professor Don Steinker has recently examined Bandy (1953) reported on some collections taken samples of Millerton material for foraminifera near San Francisco Bay and Detling (1958) de (personal communications). scribed and figured some intertidal foraminifers Johnson (1962) was concerned with developing from Sunset Bay, Oregon. More recently Grivetti valid criteria for distinguishing between a bioco (1960) studied intertidal species of the Farallon enosis and a thanatocoenosis. By comparing fo ss il Tslands and Lankford (1962) the recent foraminif mollusks with modern ones from Tomales Bay he era from the nearshore turbulent zone of the entire attempted to reconstruct the mode of formation of western United States and northwest Mexico. Sev fossil assemblages in the Millerton. Since foram eral of Lankford's collecting sites were in Bodega inifera are within the sediment size range of silt, Bay, the northern extension of the fault zone that the effects of reworking and post-mortem transpor Tomales Bay occupies (personal communication). tation are prohably more critical on such micro fossils than the macrofossils. For example, M ur MATERIALS AND METHODS ray (1965) described how benthic foraminifera While working on the distribution and abundance were raised in suspension in surface waters and re of the tube-building polychaete OlVellia fusiform is distributed over relatively wide areas by storms. (Della Chiaje) during the summer of 1964, the He indicated that small specimens were preferen writer noticed that its tube was composed of sand tially removed from source sediments and settled grains and tests of foraminifers. Although it is elsewhere. As a result the death assemblages were common for O. fusiform is to utilize material from altered in composition: by removal and by addi the substratum, the abundance and variety of fo tion. Failure to develop and apply objective dis raminifera thus used was so striking as to prompt criminatory criteria would make it difficult to dis further investigation. A pilot study was undertaken tinguish a biocoenosis from a thanatocoenosis in and twenty quantitative samples were collected with this case. a Van Veen grab from White Gulch, a small in Foraminifera can be used to explore further the cline about three miles south of the mouth of validity of criteria suggested by Johnson (1962) for Tomales Bay. In addition, thirty-six samples were recognition of exotic elements in a fossil assem taken intertidally at various sites around the bay. blage. Following the present investigation of the Moreover, Mr. Ali Imam and Mr. George Seddon, modern foraminifers, the fossil situation would be graduate students attending the summer session at examined with special attention to criteria like frag- the University of the Pacific, kindly allowed the Contribu tion No. 56, Uni v . of Delaware M a rine L aborator y author to examine material which they had col- 164 l\:rAURER-TOl\IALES BAY (CALIF.) FORAl\flNIFERA
lected for class projects. Their samples included been made, as an additional 120 quantitative sam intertidal stations slightly north of the bay and sub ples have been collected from Tomales Bay, but tidal sites in the southern portion. the analysis of these is still to be accomplished. The writer would like to acknowledge the aid of RESULTS Professor Don Steinker, who kindly examined the From all these sources the following preliminary species and brought the writer's attention to some list of calcareous benthic foraminifera of Tomales references. Bay has been compiled. The asterisk following the specific name indicates that the species was listed REFERENCES and figured by Delling. BANDY, O. L., 1953, Ecology and paleoecology of Oolilla coslala (Williamson) some California Foraminifera. Part I. The Elphidium illcerlum (Williamson) * frequency distribution of Recent Foraminifera NOllioll graleloupii (d'Orbigny) * off California: Jour. Paleontology, v. 27, no. Bulimillella e/egalllissima (d'Orbigny)' 2, p. 161-182. Fissurilla lucida (Williamson) " BUSH, J. B., 1930, Foraminifera of Tomales Bay, Quillque/oculilla semillula (Linne) ' California: Stanford Micropaleontology Bull., Quinqueloculilla sp.* v. 2, no. 2, p. 38-42. Quillqueloculilla rhodiellsis Parker' DETLlNG, M. R., 1958, Some littoral Foraminifera Cibicides lobalulus (Walker and Jacob) " from Sunset Bay, Coos County, Oregon: Cush Bolivilla vaughalli Nalland man Found. Foram. Research, Contr., v. 9, Comuspira invo[vells (Reuss) " part 2, p. 25-31. Discorbis columbiellsis (Cushman) ' DICKERSON, R. E., 1922, Tertiary and quaternary Elphidiella lIilida Cushman' history of the Petaluma, Point Reyes and Santa Bucella inusilala Anderson" Rose Quadrangles: Calif. Acad. Sci. Pro., 4th NOllionella miocenica Cushman var. slella ser., v. 11, p. 527-601. Cushman and Moyer' Discorbis ornalissima Cushman" GRIVETTI, L. E., 1960, Intertidal Foraminifera of Epollides columbiensis Cushman* the Farallon Islands: Unpublished M.A. The Cassidulilla Iimbala (Cushman and Hughes) " sis, University of California at Berkeley, 307 p. Bulimilla affillis d'Orbigny JOHNSON, R. G., 1962, Mode of formation of ma Elphidium crispwn (Linne) rine fossil assemblages of the Pleistocene Mill Trochammilla illflata (Montagu) erton Formation of California: Geol. Soc. Globigerilla bulloides d'Orbigny Amer. Bull., v. 72, p. 113-120. ROlalia beccari (Linne) has not as yet been iden LANKFORD, R. R., 1962, Recent Foraminifera from tified, but its presence is suspected. The planktonic the nearshore turbulent zone, western United species G. bulloides was washed in. The fauna re States and northwest Mexico: Unpublished sembles to a marked degree the one described from Ph.D. dissertation, University of California at Oregon by Detling (1958). It is suggested that San Diego, 234 p., illus. Quillqueloculilla semillula, Quillqueloculilla sp. and MURRAY, J. W., 1965, Significance of benthic fo Quillqueloculina rhodiensis are Quinqueloculilla raminiferids in plankton samples: Jour. Pale species A, species B and species C respectively of ontology, v. 39, no. I, p .. 156-157. Delling's paper. WEAVER, C. E., 1949, Geology of the Coast Ranges This paper is to be considered as a preliminary immediately north of the San Francisco Bay report; specimens have not as yet been deposited in region, California: Geol. Soc. Amer. Mem. a repository. Efforts to continue the study have 35, p. 1-245. CONTRIBUTIONS FRO?'I THE C USHi\l AN l"O L"NDATION F OR F OR A1\1I N Jl"ERAL RES EARCH 165
CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH VOLUME XIX, PART 4, OCTOBER, 1968 357. A TAXONOMIC NOTE ON MASSlUNA CARlNATA (FORNASINI, 1905) KEITH ATKINSON Department of Geology, University College of Wales, Aberystwyth, Wales, U.K.
ABSTRACT Millett (1898) attributed specimens from the i\[U~l!IiliIlU, nitidn cl"rinatn (Fol'nasini, 1905) and M.1ll!lsll Malay Archipelago to S. lIitida d'Orbigny. Al ina. mllldti (W iesnel', 1912) are shown to be synonymous. though illustrations of rather wild-growing individ The vadeta! na m e curinata is elevated to speciflc r ank and the differences from Slli_roloculina nitida d'Orblgny al'e dis uals, his figures 9 to 12 on Plate 5 have cylindrical cussed, The types of ~r. carinatn nre r ei llustmted togeth chambers and fig. 12b shows well the rounded pe er with a specimen [rom Cludigan Bay. "\-Val es, The spe riphery. Discussi ng Brady's carinate form he com cies is described and i ts geographical occurrence Is outlined. mented "A similar form occurs at several of the Malay stations but the test is thinner and the cham INTRODUCTION bers have not the inflation characteristic of S. The author has studied the Recent foraminifera IIi/ida." No indication is given in the text as to in 148 bottom sediment samples from Cardigan whether this carinate type is illustrated, but hi s fig Bay, Wales. Two samples contained specimens ure 13 , plate 5, is very si milar to Brady's figures. which were identical to SpirO/OClI/illa lIitida d'Or In the plate explanation for plate 5 of Millett's bigny as figured by Brady ( 1884) . This rarely re work, figure 13 is described as "S. lIitida var." It is corded miliolid has been referred to Spir%clI/illa here concluded that Millett's figures 9 to 12 on milletli Wiesner, 1912, by subsequent aut h 0 r S Plate 5 are S. IIi/ida d'Orbigny and figure 13 is S. (Boltovskoy, 1954; Barker, 1960). The present IIi/ida sensu Brady. work has shown that this name is invalid (Thal In 1905 Fornasini erected S. lIitida d'Orbigny mann, 1960) and the prior designation, Spir%c- var. carilla/a, designating as the type figures 9 and 1Ilina lIitida var. carillata Fornasini, 1905, should 10 of Brady 1884. Wiesner (1912) erected S. mil be used for this form. /elli, which was described as differing from S. IIi/ida d'Orbigny by its slightly roughened surface as com DISCUSSION pared with the shining surface of d'Orbigny's spe The first mention of this species as a discre:e cies. He attributed the wild growth patterns of form was by Brady in 1884. In a discussion of S. Millett's specimens to the habitat and suggested nitida d'Orbigny, 1826, he remarked, "Soldani 's that with growth the species altered from a smooth, drawing .... named by d'Orbigny Spir%clI/illa round-chambered form to one with clear, narrow lIitida appears to be intended for one of the vari chambers with keels or ribs. Whilst it is probably eties of Spir%clI/ina p/allll/a/a which differs from correct that environment resulted in the wild the typical form in its discrete and slightly inflated growth, tbere has been no further evidence avail segments and the somewhat produced superior ex able to support the idea that the smooth round tremity of the final chamber. The complanate shal peripberied forms are juveniles of the forms il low water Spiroloculinae of the tropics are very lustrated by Brady. variable in minor characters but a considerable sec Wiesner did not give a type figure or designate tion of them, well typified by the figures PI. 9, figs. a type specimen, but his synonymy included Brady 9, 10, may be assigned to this particular form." (1884, p. 149, pI. 9, figs. 9,10) and Millett (1898, The figures 9 and 10 of Brady are here reproduced p. 265, pI. 5, figs. 9-12). Significantly he did not as text figures 3 and 2 respectively. quote Millett's figure 13. Wiesner was apparently Cushman and Todd ( 1944; 40) described the in ignorance of Fornasini's prior designation var. first apertural view of S. lIitida d'Orbigny, given by carilla/a for Brady's figures. Neglect of tbis fact, Terquem in 1878, as having a "distinctly rounded" togetber witb lack of a designated type, must render periphery. Fornasini's ( 1900) line drawing of this Spir%clI/illa millelli Wiesner, 1912 invalid. species, traced from d'Orbigny's 1826 unpublishe:l In 1944 C ushman and Todd (p. 76) described plates, also showed a slightly rounded periphery. Wiesner's species as belonging to tbe genus Massil Brady's illustrations are of a species with an acute ilia. Although these authors stated that it was syn or carinate periphery on the final chamber and a onymous witb S. IIi/ida Brady (not d'Orbigny) double keel or excavated periphery on the penul tbey retained Millett's figures 9 to 12 and ignored timate chamber. It is apparent therefore that S. bis figure 13. ni/ida sensu Brady is not S. lIitida d'Orbigny. Tbis species bas also been illustrated as M. mi/- 166 ATKINSON-NOTE ON l\IASSILINA CARINATA
/elli by Boltovskoy (1954). The name was also into one keel at the apertural end. Test in early used by Barker (1960) for Brady's original fi gures. stages quinqueloculine, later chambers added in one Thalmann ( 1960 ) noted "s. millelli : invalid name : pl ane; chambers distinct, at least six visible from see S. Ilitida carillata Fornasi oi." each side. Sutures distinct, depressed. Wall cal careous, imperforate, porcellaneous, surface rough CONCLUSION ened, not polished, some suggestion of faint lon Although Brady (1884) was the first to illustrate gitudinal stri ae near apertural end, double keel ir this form, it was Fornasini ( 1905 ) who gave it an regu larly developed. Aperture terminal at the end original and undisputed name. According to Ar of a short neck, large, circular, with a conspicuous ticle 17 (9) of the International Code for Zoolog lip and small, often poorly developed bifid tooth ical Nomenclature, the variety earinata Fornasini which does not project beyond the end of the ul is available because it was proposed before 1961. timate chamber. Many authors [Wiesner, Bol!ovskoy, McKenzie Type specimens designated by Fornasini as (1962) and Hulme ( 1964)] have regarded this va Brady, 1884, pI. 9, fi gs. 9, 10. riety as specificall y distinct from S. nitida by its Dimensions oj th e type specimens.-Text Fig. 9: carinate or keeled periphery and roughened surface. Length 0.74 mm.; breadth 0.60 mm. ; thickness 0.16 This foraminifer, therefore, should be called Mas mm. Text Fig. 10: Length 1.10 mm.; breadth 0.85 si/ina earinata (Fornasini , 1905). mm.; thickness 0.24 mm. Depository.-The type specimens are deposited SYSTEMATIC PALAEONTOLOGY in the British Museum (Natural History). The Superfamily MILIOLACEA Ehrenberg, 1839 registered numbers are ZF 2411 (text fig. 9) and Family MILIOLIDAE Ehrenberg, 1839 ZF 2412 (text fig. 10). Subfamily QUINQUELOCULININAE Dimensions of the fi gured Cardigan Bay speci Cushman, 19 17 men (text fig. 1}.-Le ngth 1.00 mm.; breadth 0.73 Genus MassiJina Schlumberge r, 1893 mm.; thickness 0.22 mm. Massilina carinata (Fornasini, 1905) This specimen is in the author's collection at the Text fi gures I, 2, 3 Museum of the Department of Geology, Aberyst 1884 Spir%eulina ni/ida BRADY, Chall. Rep. wyth. Zool. , vol. 9, p. 149, pI. 9, fi gs. 9, 10. Observed range of dimensions of the Cardigan pars 1898 Spir%Clllinll nitida d'Orbigny MILLETT, Bay specimens.-L eng t h 0.50 mm.-1.00 mm.; JI. R. microsc. Soc., p. 265, pI. 5, fig. breadth 0.22 mm.-0.73 mm.; thickness 0.10 mm.- 13 (S. nitida d'Orbigny WIT. in plate 0.22mm. explanation) . Distribution.-This species has been recorded 1905 Spir%Clllina nitida d'Orbigny var. ea from 15 fathoms off Japan and at 8 fathoms in the rinata FORNASINI, Boll. Soc. geol. ital., Torres Strait (Brady, 1884). Cushman ( 1917 ) vol. 24, p. 389. listed specimens from Honolulu, Tokyo and Ha 1954 Massilina millelli (Wiesner) BOLTOV waii. Boltovskoy (1954, 1957) described it from SKOY, Revta lnst. naco Invest. Cienc. off the coast of central east Argentina and in the nat. Mus. argent, vol. 3, p. 261, pI. 21, estuary of the river Platte. Occurrences in Austra fi g. 6. lia and New Zealand are noted by McKenzie 1960 Massilina millelli (Wiesner) BARKER, (1962) and Hulme (1964). Spec. Pubis. Soc. econ. Paleont. Miner., Massi/ina carillata has not previously been re vol. 9, p. 18, pI. 9, fi gs. 9, 10. corded from British waters. The author retrieved non 1826 Spir%eulina nitida D'ORBIGNY, Annis specimens from Cardigan Bay Stations C.B. 113 Sci. nat., ser. I, vo l. 7, p. 298. and C.B. 426. At the time of recovery the water non 1923 Spir%Clllina carina/a WI ESNER, Die temperature was 7"C and the salinity was 32.4){,. Miliolid. der ost li ch., p. 28, pI. 5, fi gs. These stations are in 9 and 10 fathoms respectively. 42-43. non 1947 Spir%eulina earinata LE CALVEZ, Mem. Servo Carte geol. det. Fr., p. 26, pI. 2, ACKNOWLEDGMENTS figs. 32, 33. The author wishes to thank Dr. J. R. Haynes and Description.-Test free, elongate, roughly oval Dr. R. L. Moy for critically reading the manu in outline, compressed, greatest breadth at the cen script. Thanks are also due to Dr. C. G . Adams ter, almost parallel sided in edge view, sides con for making ava ilable the collections of the British cave. Periphery subacute, penultimate chamber Museum (Nat. Hist. ) and to Dr. R. Hedley and with rectangular periphery and double keel, final Dr. R. C. Whatley for many helpfu l discussions chamber has double keel at the aboral end merging and communications. The photographs of Brady's CONTRIBUTIONS FROM THE CUSHMAN ~'OUN DATION FOR FORAMINU'ERAL RESEARCH 167 original drawings are reproduced by kind permis di foraminiferi adriatici. Memorie R. Accad. sion of the Publications Division of Her Majesty's Sci. 1st. Bologna Sci. nat. Ser. 5, 8 : 360, text Stationery Office. fig. 4. ---, 1905, Sulle Spiroloculine italiane fossili e REFERENCES recente. Bol!. Soc. geo!. ita!. 24: 387-399. BARKER, R. W., 1960, Taxonomic Notes on the spe H ULME, S. G., 1964, Recent Foraminifera from cies figured by H. B. Brady in his Report on Manukau H arbour, Auckland, New Zealand. the Foraminifera Dredged by H.M.S. 'Chal N. Z. Jl Sci. 7 : 305-340. lenger' during the years 1873-1876. Spec. LOEBLICH, A. R., JR., and TAPPAN, H., 1964, Trea PubIs. Soc. econ. Paleont. Miner., Tulsa, 9 : tise on Invertebrate Paleontology. Part C. 2-240. Protista 2. Geo!. Soc. Am., I & 2 : 1-900. BRADY, H . B., 1884, Report on tbe Foraminifera McKENZIE, K. G., 1962, A record of foraminifera dredged by H.M.S. 'Challenger' during the from Oyster Harbour, near Albany, Western years 1873-1876. 'Challenger' Rep. Zoo!., 9 : Australia. J. Proc. R. Soc. West. Aust., 45 : 1-800. 117-132. BOLTOVSKOY, E., 1954, Foraminiferos de la bahia MILLETT, F. W., 1898, Report on the Recent Fo San BIas. Revta. Inst. naco Invest. Cienc. nat. raminifera of the Malay Archipelago collected Mus. Argent. Cienc. nat. Bernardino Riva by Mr. A. Durrand, F.R.M.S. Jl R. microsc. davia, 3 : 245-300. Soc. : 258-269. ---, 1957, Los Foraminiferos del Estuario de D'ORBIGNY, A., 1826, Tableau methodique de la Rio de La Plata y su zona de inftuencia . Revta. classe des Cephalopodes. AnnIs Sci. nat., ser. lnst. naco Invest. Cienc. nat. Mus. Argent. 1,7:96-3 14. Cienc. nat. Bernardino Rivadavia, 6 : 3-77. TERQUEM, 0., 1878, Les foraminiferes et les en to CALVEZ, Y. LE, 1947, Revision des foraminiferes mostraces ostracodes du Pliocene superieur de lutetiens du Bassin de Paris: 1 - Miliolidae. l'Ile de Rhodes. Mem. Soc. geo!. Fr. ser. 3, 1 : Mem. Servo Carte geo!. det. Fr. : 1-45. 1-135. CUSHMAN, J. A., 1917, A monograph of the foram THALMANN, H . E., 1960, An Index to the Genera inifera of the North Pacific Ocean: Part 6 - and Species of the Foraminifera 1890-1950. Miliolidae. Bul!. U. S. natn. Mus., 71 : 1-104. George Vanderbilt Found., Stanford Univ., ---, and TODD, R., 1944, The Genus Spir%c Calif.: 1-393. u/illQ and its species. Spec. PubIs. Cushman WIESNER, H ., 1912, Zur Systematik adriatischer Lab. 11 : 1-82. Nubecularien, Spiroloculinen, Miliolinen und ELLIS, B., and MESSINA , A ., 1940, A Catalogue of Biloculinen. Arch. Protistenk. 25 : 201-239. Foraminifera. Am. Mus. Nat. Hist., N .Y. --- , 1923, Die Miliolideen der iistlichen Adria. FORNASINI, C., 1900, Intorno ad alcuni esemplari Prague, the author. 168 E SKER- TEXAN CRETACEOUS P SEU DOGU _ El\IDELI~'-\
CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH YOLUM E XIX, PART 4, OCTOBER, 1968 358. A NEW SPECIES OF PSEUDOGUEMBELlNA FROM THE UPPER CRETACEOUS OF TEXAS GEORGE C. ESKER, III Louisiana State University, Geology Department
ABSTRACT ture is reduced to a narrow slit which tends strongly A new species of I)S6Udoguembelina appears to be re to become or actuall y forms three separate aper s tricted to the Globotruncnna gRnsseri zone. Though ra re in occurrence. It may prove to be a useful Index silecies tures (text fig. 2). In addition to the primary aper fOl' this zone. ture or apertures, there are usuall y at least three INTRODUCTION pairs of narrow slit-like accessory apertures with Pseudoguembelil/a kempel/ sis n. sp. has been prominent fl aps. found in surface and subsurface sa mples from Tex P. kempel/sis n. sp. can be distingui shed from P. as. These samples contain abundant planktonic Fo cOTllllla Seiglie, 1959; P. coslll/ala (Cushman, raminifera belonging to the G/obo/rtlll ca l/a gal/sseri 193 8); P. exco/ala (Cushman, 1926) of which P. zone. P. kempel/sis is rare in these samples. It may coslala (Carsey, 1926) is a junior synonym; P. prove to be a useful indicator of the G/obolrlll/cal/a palpebra Bronnimann and Brown, 1953 , and P. gal/sseri zone, since at present it is known to occur slriala (Ehrenberg, 1838) in lacking we ll developed only in this zone. The species, therefore, appears costae or striae. P. kempel/ sis differs from P. pUI/C to have at least local stratigraphic significance. Ill/ala (Cusbman, 1938) and P. slriala (Ehrenberg, 183 8) in having more elongate arcuate later cham ACKNOWLEDGEMENT bers and more circular early cbambers. Also, P. The writer wishes to thank Mr. D. Nugent for kem pel/sis appears to differ from all other species photographing the type specimens. of Pselldoguembe/il/ a in possessing a thick prom Superfamily GLOBIGERINACEA Carpenter, inent carina. Parker and Jones, 1862 Dimel/siol/ s.-Holotype (text figs. 1, 2): 0.407 Pseudoguembelina kempensis n . sp. mm. length; 0.259 mm. width; 0.148 mm. thickness, H .Y.H. no. 8082 .. Para type (text fi gs. 4, 5): 0.383 T ext figures 1-5 mm. length; 0.284 mm. width; 0.136 mm. thick Test plano-biserial or biserial, consisting of 7-12 ness, H.Y.H. no. 8083. Paratype (text fig . 3): 0.420 large chambers, increasing rapidly in size, and may mm. lengtb; 0.252 mm. width; 0.144 mm. thickness, have a small pointed early part consisting of about H.Y.H. no. 8084. 7 additional chambers. The small early chambers Deposilory.-H. Y. Howe Collection, School of may be coiled initially (text fig. 4). The small early Geology, Louisiana State University. Holotype, chambers are not inflated and the sutures are nearly H.Y.H. no. 8082; paratype, H.Y.H. no. 8083; para flu sh with the chamber surface. The surface is type, H.Y.H. no. 8084; and 7 paratypes, H.Y.H. punctate with the alignment of the pores, particu no. 8085. larly towards the outer margin, sometimes suggest Type /oca/ily.-Kemp Clay, 2-4' below the con ing faint striae (text fig. 3). The microspheric tact between tbe Littig Conglomerate and the Kemp forms tend to lack spines and are not as thick. The Clay, on Walker's Creek about 5Y2 miles NE of third pair of chambers is the most strongly inflated, Cameron, Texas. after which later chambers tend to be less inflated and, finally, somewhat compressed. The last cham REFERENCES ber may be very elongate and may extend com BRONNIMANN, P., and BROWN, N. K., JR. , 1953 , Ob pletely over the preceding two chambers. On the servations of some pl anktonic Heterohelicidae megalospheric forms, the first few large chambers usually have short thick spines that may tend to from tbe Upper Cretaceous of Cuba: Cush man Found. Foram. Res., Contr., v. 4, pt. 4, merge to form a sort of ridge. These tend to im part a knobby appearance to the chamber (text fig. pp. 150-155. 2). Text figs. 3 and 4 show the extremes in the CUSHMAN, J. A., 1926, Some foraminifera from the degree to which the test is pointed. The sutures are Mendez shale of Eastern Mexico: Contr. Cush deeply depressed between the large chambers. A man Lab. Foram. Res., v. 2, pt. I, no. 26, rather thick conspicuous carina is present. The p. 20. primary aperture is a large lunate arch in all cham ---, 1938, Cretaceous species of G iimbelil/a bers except the last one (text fi g. 5). With the and rel ated genera: Contr. Cusbman Lab. Fo completion of the final chamber, the primary aper- ram. Res., v. 14, pt. I, pp. 13, 16. CONTRIB. CUSHMAN FOUND. FORAM. R ESEARCH, VOL. 19
Text Figures 1-3
Atkinson: Massi/ina carillata (Fornasini, 1905)
EXPLANATION OF TEXT FIGURES F IGs. P AGE 1. Massi/ilia carill a/a ( Fornasini, 1905) . X 40. Cardigan Bay Station C. B. 426. la, side view; Ib, apertural view...... 166 2. Massi/ilia carilla/a (Fornasini , 1905). X 43. Challenger Stati on 187A.. ( Brady 1884, pI. 9, fig. 10)...... 166 3. M aSl'ililla carilla/a (Fornasi ni , 1905). X 50. C hallenger Station 233B. ( Brady 1884, pI. 9, fig . 9) . .3a, side view; 3b, apertural view...... 166 CONTRIB. CUSHMAN FOUND. FORAM. RESEARCH, VOL. 19 Text Figures 1-5
Esker: Pselldogltembelina from Texas
Text Figures 1-3
Esker: Lectotype of Globotnmcana 1'osetta (Carsey) CONTRIBUTIONS FROM THE CUSHMAN FOU NDATION FOR FORAlIlINIFERAL R ESEARCH 169
EHRENBERG, C. G., 1838, Ober die Bildung der SEIGLIE, G. A., 1959, Notas sobre algunas especies Kreidefelsen und des Kreidemergels durch un de Heterohelicidae del Cretacico superior de sichtbare Organismen: K. Akad. Wiss. Berlin, Cuba: Asoc. Mexic. Geol. Petr., Bol., v. II, Physik. Abh., p. 135. no. 1-2, p. 60.
EXPLANATION OF TEXT FIGURES Paper No. 358 FIGS. PAG E 1,2. Pseudoguembelilla kempellsis n. sp., holotype, showing the appearance of the final chamber and the primary aperture, H.V.H . no. 8082. X 125...... 168 3. P. kempensis n. sp., paratype, megalospheric form showing the presence of faint striae towards the outer margins, H.V.H. no. 8084. X 125...... 168 4,5. P. kempellsis n. sp., paratype, microspheric form showing the initial coiled part, well devel oped carina, and the primary aperture as it exists in the chambers formed before the last chamber, H.V.H. no. 8083. X 125...... 168
EXPLANATION OF TEXT FIGURES Paper No. 359 FIGS. PAGE 1-3. Globotruncalla rosetta (Carsey), lectotype, Univ. of Texas Collections no. 1510. ' I, spiral view; 2, side view showing single keel on the final chamber and closely spaced double keel on the preceding chamber; 3, umbilical view. All X87 ...... 170 170 ESKER- LECTOTYPE OF GLOUOTRUNCANA ROSETTA
CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH VOLUME XIX, P ART 4, O CTOBER, 1968 359. DESIGNATION OF A LECTOTYPE OF GLOBOTRUNCANA ROSETTA (CARSEY) G EORGE C. ESKER, III Louisiana State University, Geology Department
ABSTRACT cies had one or two keels, nor did she include a fi g Because of the apparent loss of the holotype of Globo truncalH\ rosetta (Car seY), the author has consider ed it ure by which this could be determined. Plummer necessary to des ignate and descl'ibe a lectotype. After the ( 193 1) reillustrated the holotype, including a side examination of the three specimens labeled as cotypes of view that showed the presence of only a single keel. Globil;erin~l rosetta Carsey in the Carsey coll ection, Later, Bro nnimann and Brown (1955) stated that the s pecime n which most closely fitted the hololype' 8 char acteristics was selected to be the lectotype. upon examining the holotype they found two close ly spaced keels on all of the chambers except the INTRODUCTION final one. Pessagno ( 1960) has shown that G. ro Statements such as those made by Bronnimann sella has a rather wide double keel in the early and Brown (1955), that the holotype of Globo stages at least, and that there is a tendency for the Irull calla rosetta (Carsey, 1926) had two closely ventral keel to disappear in the adult stages. Ols spaced keels except for the final chamber, and by son (1964) and van Hinte ( 1963) consider within Bandy ( 1967 ), that the holotype of Gioboirullca lla the population of G. rosella forms that have two rosetta had only a single keel , are contradictory. keels throughout the test. Douglas and SHter (1966) The author, in an attempt to resolve such contra state that G. rosetta is more commonly biconvex, dictory statements by studying the holotype of Glo has a less acute axial periphery, and that its cham bolruncalla rosella, was unable to locate any speci bers have somewhat curved umbilical sides. They men in the Carsey collection that could be consid say that only a small percentage of typical G. ro ered to be the holotype. It apparently has been sella have two keels in the early stage, yet in their lost. Therefore, in an attempt to avoid consider article (pI. 3, fi g. 6) one specimen shows a very able controversy about the species in the future, the closely spaced double keel and another (fi g. 7) ap author decided to designate and describe a lecto pears to show an imperforate keel band. Bandy type of Globolruncana rosetta (Carsey) from the (1967) has stated that the holotype of Carsey has cotypes in the Carsey collection. only a single keel and that only an occasional spe cimen shows any trace of a second keel on the final ACKNOWLEDGEMENTS whorl. Band y believes that Ol sson and van Hinte The author wishes to thank Dr. Samuel P. El have erroneously considered forms of G. velliricosa lison of the University of Texas for his help in ob to be G. rosella. These references should serve to taining the specimens from the Carsey collection indicate the controversial status of G. rosetta. for this study. Thanks are also due to Mr. Don The author has examined specimens from the Nugent for the photographs of the type specimen. upper Taylor Marl of Texas and has reached the conclusion that the double-keeled specimens therein DISCUSSION may be assigned to two distinct species, G. velllri The Carsey collection includes three specimens cosa and G. rosetta, and to more obviously differ labeled as cotypes of Globolruncalla rosella. Un ent species such as G. Iillfl eiafla. The specimens as fortunately, these do not appear to be conspecific. signed to G. vefllricosa generally have a nearly fl at All three clearly have double keels. One specimen spiral side and two moderately widely spaced keels. belongs to G. aegypliaca Nakkady, of which G. On immature specimens the keels are widely spaced gagnebini· Tilev is a junior synonym. Another spe similar in appearance to those of G. lilllleialla Iri~ cimen should be assigned to G. gansseri Bolli dica carillala. The speci mens assigned to G. rosella rillala Pessagno. The third specimen appears to fit generally have a slightly convex spiral side, usually the concept of the species of G. rosella, as held at more convex than specimens assigned to G. vell lri least by some workers. This specimen has two cosa, and two closely spaced keel s. While the keels keels on all of the chambers except the final one. may become progressively more closely spaced in The keels become more closely spaced on the both species, those of G. rosetta are more closely younger chambers until the ventral keel disappears spaced than those of G. velliricosa. altogether on the fin al chamber. DESCRIPTION In the original description of G. rosetta by Car Low trochospiral test, slightly convex on the sey (1926) , she did not mention whether her spe- spiral side, strongly convex on the umbilical side. CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION F OR F ORAMINIl<'ERAL RESEARCH 171
Moderately to strongly lobate equatorial periphery, Cretaceous of Central Texas: Univ. Texas angular rhomboidal axial periphery ; with two mod Bull., 2612, pp. 1-56, 1-7 pIs. erately spaced keels in the first chamber of the final DOUGLAS, R. , and SLITER, W. V., 1966, Regional whorl, becoming more close ly spaced on later distribution of some Cretaceous Rotaliporidae chambers until the ventral keel disappears on the and Globotruncanidae (Foraminiferida) with final chamber. There are adumbilical and perium in North America: Tulane Studies Geol., v. 4, bilical extensions of the ventral keel. The chamber no. 3, pp. 84-131, pIs . 1-5, text fig. I, tabs. 1,2. surface is spinose on the umbilical side and on all HINTE, J . E. VAN, 1963, Zur Stratigraphie und Mi but the last two chambers on the spiral side. There kropalaontologie der Oberkreide und des Eo are five chambers in the final whorl. The sutures zans des Krappfeldes (Karnten): Geol. Bundes are moderately curved and raised on the spiral side, ansI., Jahrb., Sonderband 8, pp. 1-147, pis. nearly radial and slightly depressed on the umbil 1-22. ical side. Umbilicus wide, primary aperture interio marginal umbilical; umbilicus covered by a tegil NAKKADY , S. E., 1950, A new foraminiferal fauna lum with infralaminal and intralaminal accessory from the Esna shales and Upper Cretaceous apertures. Chalk of Egypt: Jour. Pal. , V. 24, no. 6, pp. 675-692, pIs. 89, 90. Dimellsiolls.-Max. diam. 0.52 mm.; min. diam. 0.42 mm.; thickness 0.30 mm. OLSSON, R. K., 1964, Late Cretaceous planktonic foraminifera from New Jersey and Delaware: D epository.-University of Texas collections no. Micropal., v. 10, no. 2, pp. 157-188, pIs. 1-7. 1510. PESSAGNO , E. A., JR., 1960, Stratigraphy and micro paleontology of the Cretaceous and lower Ter REFERENCES ti ary of Puerto Rico: Micropal., v. 6, no. 1, pp. 87-110, pIs. 1-5. BANDY, O. L., 1967, Cretaceous planktonic foram iniferal zonation : Micropal. , v. 13, no. I, pp. PLU MM ER, H ELEN J ., 1931 , Some Cretaceous Fo 1-31 , Text figs. 1-13. raminifera in Texas: Univ. Texas Bull., no. 3101, pp. 109-202, pIs. 8-15 . BRONNIMANN, P., and BROWN, N. K., JR. , 1955, TILEV, N., 1952, Etude des Rosalines Maestrichti Taxonomy of the Globotruncanidae: Eclogae ennes (genre G /obotrllIlCQIlQ) du Sud-Est de Geol. Helv., v. 48, no. 2, pp. 503-561 , pIs. 20- la Turquie (Sondage de Ramandag): Maden 24, text figs . 1-24. Tetkik ve Arama Enstitlisli, ser. b, no. 16, pp. CARSEY, DOROTHY 0., 1926, Foraminifera of the 1-101, pIs. 1-3, text figs. 1-24. 172 TODD-REC}<~NT LITERATURE ON THE FORAMINIFERA
CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH VOLUME XIX, PART 4, OCTOBER, 1968 RECENT LITERATURE ON THE FORAMINIFERA
Below are given some of the more recent works ennes) et les Basses-Alpes.-Mem. du Bureau on the Foraminifera that have come to hand. de Recherches Geol. et Min., No. 58, Colloque sur I'Eocene, Paris, May 1968, p. 13-26, pIs. 1, BERGGREN, W. A., PHILLIPS, J. D., BERTELS, A., and 2, tables 1-3 , maps 1,2. WALL, D. Late Pliocene-Pleistocene stratig BOLLI, H . M., BOUDREAUX, J. E., EMILIANI, CESARE, raphy in deep sea cores from the south-central North Atlantic.-Nature, v. 216, No. 5112, HAY, W. W., HURLEY, R. J., and JONES, J. I. Oct. 21, 1967, p. 253-254, text figs. 1, 2 Biostratigraphy and paleotemperatures of a (graphs, diagram) .-Preliminary report of pa section cored on the Nicaragua Rise, Carib leomagnetic studies. bean Sea.-Bull. Geol. Soc. America, v. 79, No.4, April 1968, p. 459-470, pI. 1, text figs . BIELLE, MARIETTE, and CARALP, MICHELLE. Les 1-3 (map, profiles, graphs).-Study of 56.4- Miogypsinidae de deux coupes profondes des meter core. Plio-Pleistocene boundary (700, Landes.-Bull. Soc. Geol. France, ser. 7, v. 9, 000 years) recognized by correlation of calca No.1, 1967 (March 1968), p. 36-42, pI. 4, reous nannoplankton. Glacial stages present tables 1-3 (range charts). in the Pliocene. BIGNOT, GERARD. Nouvelle decouverte de Fabiallia BOULANGER, D., and DEBOURLE, A. Precisions sur cassis (Oppenheim) dans Ie Lutetien du Bas A ngolia aqllilallica, Cuvillier, 1963, Foraminif sin de Paris.-Mem. du Bureau de Recherches ere du Lutetien d'Aquitaine Occidentale.-Re Geol. et Min., No. 58, Colloque sur I'Eocene, vue de Micropaleontologie, v. 10, No. 4, Paris, May 1968, p. 79-83, 1 pI., 1 text fig. March 1968, p. 243-249, pIs. 1, 2, text figs. I, (geol. section). 2 (graphs) .- Based on free specimens. Genus BlZON, GERMAINE. Contribution a la connaissance related to Gypsilla. des Foraminiferes planctoniques d'Epire et BRADSHAW, JOHN S. Environmental parameters des Iles Ioniennes (Grece occidentale) depuis and marsh Foraminifera.-Limnology and Ie Palt~ogene superieur jusqu'au Pliocene. Oceanography, v. 13, No.1, Jan. 1968, p. 26- Publ. Instit. Fran~ais du Petrole, 1967 (April 38, text figs. 1-10 (maps, diagram, graphs), 1968), p. 1-142, pIs. 1-29 (fossils), pIs. 1-14 table I.-Continuous recording. (maps, range charts, evolutionary series, vari ation series, correlation table), text figs. 1-22 BRONNIMANN, PAUL, and CONRAD, MARc-A. Cinqu (maps, range charts, geol. sections, drawings, ieme note sur les Foraminiferes du Cretace in columnar sections) .-Illustrated systematic cat f6rieur de la region genevoise. M elalhrokerioll alog includes about 75 species and subspecies, valserinellsis, n. gen., fl. sp., un Foraminifere none new. nouveau du Barremien 11 facies urgonien dans BLEAHU, M., BABUCEA, YVONN E, and PILIUTA, ANA Ie Jura fr a n ~a i s.-Compte Rendu des Seances, MARIA. Contribution to the microbiostratig Soc. Physique Hist. Nat. Geneve, n. ser., v. 1, raphy of the Eocretaceous rocks in the Metal fasc. 3, Sept.-Dec. 1966 (June 1967), p. 129- liferous Mountains (English abstract of Ru 151, pIs. 1-3, text figs. 1-11 (map, graphs, manian text) .-Acad. Repub. Populare Ro drawings) .-Similar to Alveolophragmilll11. mini, Studii si Cercetari de Geol., Geofiz., Geo BRONNIMANN, PAUL, CURRY, DENNIS, POM EROL, graf., ser. Geol., tom. 13 , No.1, 1968, p. 137- CHARLES, and SZOTS, ENDRE. Contribution a 155, pIs. 1-4, table I.-An exclusively arena la connaissance des Foraminiferes plancton ceous fauna of Aptian-Middle Albian age. iques de l'Eocene (incluant Ie Paleocene) du BLONDEAU, ALPHONSE. Revision des Nummulites bassin Anglo-Franco-Belge.-Mem. du Bureau et des Assilines des Alpes-Maritimes Franco de Recherches Geol. et Min., No. 58, Colloque Italiennes.-Mem. du Bureau de Recherches sur I'Eocene, Paris, May 1968, p. 101-108, Geol. et Min., No. 58, Colloque sur I'Eocene, chart.-Planktonic species listed from Than Paris, May 1968, p. 27-55, 1 pI., range chart. etian to Ludian. BLONDEAU, ALPHONSE, BODELLE, JACQUES, COM CHALILOV, D. M. Mikrofauna i Stratigrafija Pale PREDON, ROB ERT, LANT EAUME, MARCAL, and ogenovykh Otlozhenij Azerbajdzhana.-Akad. NEUMANN, MADELE INE. Repartition stratig Nauk Azerbajdzhan. SSR, Instit. Geol., Baku, raphique des grands Foraminiferes de 1'100- 1967, p. 1-244, pIs. 1-42, text figs. 1-9 (map, cene dans les Alpes-Maritimes (Franco-Itali- phylogenetic diagrams) .-IlIustrated system at- CONTRIBU TIONS FRO"M THE CUSHl\'IAN FOU NDATION FOR FORAMINIFERAL RESEARCH 173
ic catalog of 260 species and subspecies (33 pI. 3, June 1968, p. 371-375, text fig. 1 (draw species and 10 subspecies new) from Danian ings) .-Developed from a globorotaloid an to lower Oligocene. Nonionelleta gen. n. cestor. (type species NOlliollella tumida Chalilov 1957) DE ZANCHE, VITTORIO, PAVLOVEC, RAJKO, and is erected. PROTO DECIMA , FRANCA. Microfauna and CIFELLI, RICHARD, BLOw, WALTER H., and MEL Microfacies of the Eocenic Flysch series near SON, WILLIAM G. Paleogene sediment from Ustje in the Vipava valley (Vipavska Dolina, a fracture zone of the Mid-Atlantic ridge. SW Slovenia).-Acad. Sci. Art. Sloveniea, Jour. Marine Res., v. 26, No.2, May 15, 1968, Razprave, Ljubljana, v. 10, 1967, p. 205-263, p. 105-109, text figs . I, 2 (map, diagram). pIs. 1-17, text figs. 1-6 (map, columnar sec Two samples, one Thanetian, the other Ypresi tion, drawings, graphs, diagram), tables 1-6. an, with planktonic Foraminifera. Mostly concerning Nummulites; 2 new species. COLA LONGO, MARIA LUISA. Biostratigrafia del Meso DR EYER, EVA. Mikrofossilien des Rat und Lias zoico nei dintorni del Passo del Diavolo ( Par von SW-Brandenburg.-Jahrbuch fUr Geolo co Nazionale d'Abruzzo) .-Giornale Geol., gie, Berlin, Band 1, 1965 (1967), p. 491-531, Ann. Mus. Geol. Bologna, ser. 2, v. 34, fasc . pIs. 1-9, table I (range chart).-Illustrations I, 1966 (1967) , p. 87-130, pIs. 10-13, text figs. of a few Foraminifera accompanying Ostra 1-3 (map, correl. chart, columnar section), coda. tables I, 2.-Cretaceous Foraminifera in thin EAMES, F. E. Sillduliles, a new genus of the Num section. mulitidae (Foraminiferida) .-Palaeontology, v. II , pt. 3, June 1968, p. 435-438.-Type spe COLA LONGO, MARIA LUISA , and SARTONI, SAMUELE. cies is Operculilla silldellsis L. M. Davies 1927. Globorotalia hirmta aemilialla nuova SOllO species cronologica del Pliocene in Italia. EISENACK, ALFRED. Foraminiferen aus dem Ordo Giornale Geol., Ann. Mus. Geol. Bologna, ser. vizium und Gotlandium des baltischen Ge 2, v. 34, fasc. I, 1966 (1967), p. 265-284, pIs. bietes.-Neues Jahrb. Geol. Palaonl. Abh., 30, 31, text figs. I, 2 (map, drawings), table Stullgart, Band 128, Heft 3, July 1967, p. 244- 1 (phylogenetic diagram).- A fivefold zona 274, pIs. 24-28, text figs. 1-10 (drawings). tion of Pliocene on the basis of species of Forty-seven species, 17 new, from Ordovician Globorotalia. and Silurian. Pachyammilla n. gen. (genero type A mphitremoidea? pachytheca Eisenack) COLE, W. STORRS. More on variation in the genus erected. Lepidocyclilla (Larger Foraminifera) .-Bull. Am. Paleontology, v. 54, No. 243 , June 21, FOLLADOR, UMBERTO. 11 Pliocene ed il Pleistocene 1968, p. 291-327, pIs. 19-24.-Variation in L. dell'Italia centro-meridionale, versante Adri vauglwlli, and several species included as atieo. Biostratigrafia.- Boll. Soc. Geol. Ital., synonyms. v. 86, fa sc. 3, 1967, p. 565-584, text figs. I, 2 (map, diagram) , table I (range chart) .-Defi CONATO, VITTORIO, and FOLLADOR, UMBERTO. nition and description of 5 assemblage zones Globorotalia crotoll ellsis e Globorotalia cras and 4 subzones. sacrotollellsis nuove specie del Pliocene Itali ano.-Boll. Soc. Geol. Ital., v. 86, fasc. 3, FUCHS, WERNER. Ober Ursprung und Phylogenie 1967, p. 555-563, text figs . 1-6 (drawings). der Trias-"Globigerinen" und die Bedeutung Related to G. crassaformis, but with lower dieses Formenkreises fUr das echte Plankton. ranges. -Austria Geol. Bundes., Verhandl., Heft 1-2, 1967, p. 135-176, pIs. 1-8, text figs. 1-3 (maps). CONKIN, JAMES E., CONKIN, BARBARA M., and -Origin of G lobigerilla-like forms traced to CANIS, WAYNE F. Mississippian Foraminifera Diplotremilla in the Middle Triassic and they of the United States. Part 3-The limestones evolved into Gubkillella, the first true plankton of the Chouteau Group in Missouri and Illi of the Jurassic. They are frequent and may nois.-Micropaleontology, v. 14, No.2, April have been planktonic in the Rhaetic. Five new 1968, p. 133-178, pIs. 1-4, text figs. 1-16 (map, genera and 24 species, 21 new, in upper Ladini columnar sections, diagram, check lists, range an to Rhaetic. Kollmallllita n. gen. (generotype chart), tables 1-42.-Illustrated systematic cat G lobigerilla ladillica Oberhauser), Schmidita alog includes 42 species, 5 new and 5 emended. n. gen. (generotype S. hedbergelloides n. sp.), CORDEY, W. G. A new Eocene Cassigerillella from Oberhauserella n. gen. (generotype Globiger Florida.-Palaeontology, v. II, pI. 3, June ilia mesotriassica Oberhauser) , Schlagerilla n. 1968, p. 368-370, text fig. 1 (drawings). gen. (generotype S. allgllslillmbilicata n. sp.), Morphology and phylogeny of Orbulilloides beck and Praegllbkillella n. gen. (generotype P. mallllii (Saito 1962) .-Palaeontology, v. II, kryptllmbilicata n. sp.). 174 TODD-RECENT LiTERATURE ON THE FORAl\IINIFERA
GOFMAN, E. A. Foraminifery Jury Severnogo Bucuresti, 1968, 391 p., 41 pis., 19 text figs ., Kavkaza.-Akad. Nauk SSSR, Minist. Neft. 9 tables, geol. map.-Illustrated systematic Promysh. SSSR, Instit. Geol. Razrab. Gor. catalog (p. 257-286, pis. 10-34) includes 107 Iskop., 1967, p. 1-148, pis. 1-18, text figs . 1-3 species and varieties, none new. (map, columnar sections, range chart), tables KAHLER, FRANZ and GUSTAVA. Zur Systematik der 1-10 (check lists) .-IIIustrated systematic cat Fusuliniden.-Ann. Naturhist. Mus. Wien, alog includes 92 Jurassic species (35 new and Band 71, Nov. 1967, Kiihn-Festschrift, p. 107- 6 indeterminate) and 4 varieties (2 new). 115. GOLEV, B. T. Zur Frage der Morphologie und KOEHN-ZANIN ETTI, LOU ISETTE, and BRONNIMANN, Systematik der Unterfamilie Nummulitinae. PAUL. De la paroi de Triasina IllIntkeni Maj Austria Geol. Bundes., Verhandl., Sonderheft zon, 1954.-Compte Rendu des Seances, Soc. G, 1965, p. 265-287, pis. 1-3 , text figs. 1-5. Physique Hist. Nat. Geneve, n. ser., v. I, fasc. GORBATCHIK, T. N. Javlenie Gomeomorfii u Fo 2, May-Aug. 1966 (Nov. 1966), p. 83-88, 1 pI. raminifer.-Paleont. Zhurnal, 1968, No. I, p. -Various degrees of crystallization of the wall. 3-10, pis. 1,2, text figs. 1-3 (diagrams).-Two LEVEN, E. YA. Stratigraphy and Fusulinides of new species in the Lituolidae, one in Melath the Pamirs Permian deposits (in Russian). rokerian and one in TOllasia gen. nov. (type Acad. Sci. USSR, Geol. Instit., Trans., v. 167, species T. evoluta sp. nov.), both from the 1967, p. 1-224, pis. 1-39, text figs. 1-35 (maps, Berriasian. geol. sections, columnar sections, photos, range GUDlNA, V. I., NUZHDlNA, N. A., and TROITSKII, S. charts), tables 1-5.-IIIustrated systematic cat L. New Foraminifera Forms and Recent Data alog of 126 species and subspecies (40 species on Sea Pleistocene of Taimyr Lowland (in and 7 subspecies new). Russian).-Acad. Sci. USSR, Siberian Branch, LINDENBERG, HANS GEORG. Untersuchungen an Geol. and Geophys., No. 1 (97), Jan. 1968, Iituoliden Foraminiferen aus dem SW-deut p. 40-48, I pI., text figs. 1-3 (map, geol. sec schen Dogger, 2: Die Arten von Haplophrag tions).-Two species (1 new) and a new sub mium und Triplasia. Eine Bearbeitung auf bio species. metrischer und palaokologischer Grundlage. HOFKER, J. Foraminifera of the Upper Cretaceous Abhandl. Senckenberg. Naturforsch. Gesell of South-Limburg, Netherlands. LXXXV. schaft, No. 514, Nov. I, 1967, p. 1-73, pis. 1-5, Siderolites calcitrapoides Lamarck and Sidero text figs. 1-190 (drawings, graphs), table 1. lites laevigata Reuss in the Tuff Chalk of Taxonomic revision resulting in 1 species (4 Maastricht, especially in the type-section, Sec formae) of Haplophragmium and 2 species (1 tion I.-Natuurhist. Maandblad, 55e Jrg., No. new) of Triplasia. 9, Sept. 28, 1966, p. 140-142, text figs. 1, 2 LOURDES DIAZ DE GAMERO, MARIA. Paleontologia (graphs) .--Significance of variation in proloc de la Formacion EI Veral (Mioceno), Estado ulus size. Falcon.-Geos, Escuela Geol., Minas y Metal Hat die feinere Wandstruktur der Foraminiferen urgia, Univ. Central de Venezuela, Caracas, supragenerische Bedeutung? - Palaont. Zeitschr., No. 17, May 1968, p. 7-51, pis. 1-3, map, Stuttgart, Band 41, Heft 3/ 4, Oct. 1967, p. check list, correl. table.-Catalog of 85 species, 194-198, pis. 19-21.-Examplesof different wall a few illustrated. structure in phylogenetically-related genera. MABESOONE, J. M., TINOCO, I. M., and COUTlNHO, Foraminifera from the Bay of Jakarta, Java. P. N. The Mesozoic-Tertiary boundary in Bijdragen tot de Dierkunde, Aft. 37, Amster northeastern BraziL-Palaeogeography, Palae dam, 1968, p. 11-59, pis. 1-12, text figs. 1, 2 oclimatology, Palaeoecology, v. 4, No.3, April (map, check Ji st).-Thirty-five species, 1 new. 1968, p. 161-185, text figs. 1-7 (maps, outcrop HORNtIlROOK, N. DE B. A handbook of New Zea photos, sedimentary log, drawings of forams), land microfossils (Foraminifera and Ostra tables 1-6.-Based on Foraminifera. coda) .-New Zealand Geol. Survey Hand MANTOVANI UGUZZONI, M. P., and PIRINI RADRIZ book, Inform. Ser. 62, 1968, p. 1-136, text figs. ZANI, C. I Foraminiferi delle Marne a Fu 1-29 (plates of fossils), tables 1-11 (range coidi.-Riv. Ital. Paleont. Stratig., v. 73, No. charts, correl. table) .-Beautifully illustrated 4, Dec. 1967, p. 1181-1256, pis. 85-94.-About and usable handbook of about 225 key species 90 species, none new, of Aptian-Albian age giving brief descriptions and local ranges. recorded and illustrated. IONESI, BICA. Stratigrafia Depozitelor Miocene de MARGEREL, J EAN-PIERRE. Les petits Foraminif Platforma dintre Valea Siretului si Valea Mol eres de l'Eocene de l'ouest de la France et leur dovei.-Edit. Acad. Repub. Social Romania, interet stratigraphique.-Mem. du Bureau de CONTRIBUTIONS FROM THE CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH 175
Recherches de Geol. et Min., No. 58, Colloque pl anctoniques de I'Ypresien de la Belgique et sur l'Eocene, Paris, May 1968, p. 93 -99, text du nord de la France.-Mem. du Bureau de fig. 1 (map). Recherches Geol. et Min., No. 58, Colloque sur l'Eocene, Paris, May 1968, p. 109-129, pI. MARTINIS, BRUNO. Sull'eta delle argi ll e di Gal lipoli (Lecce).-Rend. Atti Accad. Naz. Lincei, I, table 1, map I.-Eight species recorded and ill ustrated. ser. 8, v. 42, fasc. 6, June 1967, p. 824-829, pI. I, text figs. I, 2 (map, columnar section). NEAGU, THEODOR. Biostratigraphy of Upper Cre Foraminifera of Calabrian age. taceous deposits in the southern Eastern Car pathians near Brasov.-Micropaleontology, v. MASOLI , MARIO. Microfaune nel Domeriano di 14, No.2, April 1968, p. 225-241, pis. I, 2, Yerona.--Studi Trentini Sci. Nat., n. ser., sez. text figs. 1-3 (map, zonal charts).-Lists and A, BioI., v. 44, fasc. 2, 1967, p. 131-151 , pis. outline drawi ngs of forams from 14 horizons 1-4, text fig I (map).-Twenty-one species of wit hin a section from Cenomanian to lower Foraminifera, II indeterminate. Maestrichtian. Four new species and 2 new MCGOWRAN, BRIAN. Reclassification of early Ter subspecies are described. tiary Globorolalia.-Micropaleontology, v. 14, NEMKOV, G. Les Nummulites de I'V.R.S.S ., leur No.2, April 1968, p. 179-198, pis. 1-4, lext evolution, systematique et distribution stratig figs. 1-3 (phylogenetic diagrams) .-Trw/co raphique.-Mem. du Bureau de Recherches rOlaloides is subdivided into 3 subgenera : Geol. et Min., No. 58, Colloque sur l'Eocene, Acaril/il/a, Morozol'ella, and Trlll/corolaloides Paris, May 1968, p. 7 1-78, phylogenetic dia s.S., and retained in Truncorotaloidinae. Plal/o gram, range chart. rOlaliles is retained in G loborotaliinae. NEUMANN, MADELEINE. A propos de Gyroidil/ella MEDIZZA , F. La struttura del guscio nel genere magI/ a Le Calvez en Aquitaine et en Mesogee. Aragol/ia Finlay 1939 (Foraminifera).-Boli. -Mem. du Bureau de Recherches Geol. et Soc. Paleont. Ital., v. 5, No.1, 1966, p. 92-98, Min., No. 58, Colloque su r l'Eocene, Paris, pi s. 33, 34, text fig . 1 (diagram).-A lamellar May 1968, p. 85-91, 1 pI., text fig. 1 (drawing). genus. NORLING, ERIK . On Liassic Nodosariid Foraminif MEHES, KALMAN. Two species of the genus Or era and their wall structures.--Sver. Geol. hilolil/opsis from the Yillany Mountains, Hun Unders., ser. C, No. 623, v. 61 , No.8, 1968, gary.- Micropaleontology, v. 14, No.2, April p. 1-75, pis. 1-9, text figs. 1-12 (map, range 1968, p. 22 1-224, pI. I, text fig. 1 (map), table charts, drawings), tables 1-5.-A few species \.- Two Barremian species, neither one new. are described as to their wall texture, perfora MELLO, JAMES F., and Buus, MART tN A. An ap tion, lamination and layering, and ornamenta plication of cluster analysis as a method of de tion. Two new genera are separated off from termining biofacies.-J our. Paleontology, v. Delllalilla-one not laminated, the other par 42, No.3, May 1968, p. 747-758, text figs . 1-6 ti ally laminated, leaving Del/lalil/a completely (graphs, maps), tables 1-4.-Based on Phleger's laminated. Prodelllalilla nov. gen. (type spe (\956) paper on Recent Foraminifera off Tex cies Del/lalil/a rerqll emi d'Orbigny 1849) and as. The method is useful for biofacies analysis. Mesodelllalilla nov. gen . (type species Del/lal ME RIC, ENGIN. Sur quelques Loftusidae et Or iI/a mallllilla d'Orbigny 1849). bitoididae de la Turquie.-Revue Fac. Sci. OKJMURA , YUJI. Carboniferous palaeotextulariid Univ. Istanbul, ser. B: Sci. Nat., tome 32, fasc. foraminifers from the Akiyoshi Limestone 1-2, 1967, p. 1-58, pis. 1-36, text figs. 1-22 Group, Southwest Japan.-Jour. Sci. Hiroshima (map, range charts, stratigraphic profile, graphs, Univ., ser. C (Geol. and Min.), v. 5, No.3, drawings).-Twenty species (5 new) and a May 1967, p. 255-266, pI. 17, text figs. 1, 2 variety from upper Maestrichtian. (map, range chart) .-Nine species, 5 indeter MICHA EL, ERHARD. Variations in the number of minate. species of benthonic Foraminifera in the Bar OLBERTZ-WEHRLI , GERTA . Epislomil/ella a I a t a remian of northwestern Germany.-Palaeo (Marsson), ein Beitrag zur Kliirung des Be geography, Palaeoclimatology, Palaeoecology, griffs Coleiles reliclIloslls (Plummer) [Foram.]. v. 4, No.4, June 1968, p. 287-303, text figs. -Wissenschaft. Zeitschr. Ernst-Moritz-Arndt 1-5 (map, graphs), tables I, 2.-Number of Vniv. Greifswald, Jahrg. 16, Math.-naturwiss. species increases away from shorelines. Max Reihe Nr. 3, 1967, p. 189-198, pis. 1-3.- Pos im a and minima are observed in faunal sec sible relationship between the two species. tions, showing conformity between different PERCONIG, E. Microfacies of the Triassic and Ju sections. rassic sediments of Spain.- Internat. Sedimen MOORK ENS, THIERRY. Quelqu es Foraminiferes tary Petrographical Ser., v. X, 1968, p. 1-63, 176 TODD-RECENT LITERATURE ON THE FORAMIN IFERA
pis. 1-123, text figs. 1-11 (maps).-Book in A. Truncorotalia aragollellsis (Nuttall) iz eludes 341 photomicrographs illustrating mi Nizhnepaleogenovykh Otlozhenij Bassejna r. crofacies, some ineluding Foraminifera. Vedi (Armjanskaja SSR) .-Izvest. Akad. Nauk PHILLIPS, J. D., BERGG REN, W. A., BERTE LS , A., Armjans. SSR, Nauki 0 Zemle, v. 20, No. 1-2, and WALL, D. Paleomagnetic stratigraphy and 1967, p. 3-12, 1 pI., 1 table.-Three unnamed micropaleontology of three deep sea cores varieties in the lower Eocene of Armenia. from the central North Atlantic Ocean.-Earth SA IDOVA, KH. M. The biomass and quantitative and Planetary Science Letters 4, North-Hoi distribution of live Foraminifera in the Kurile land Publ. Co., Amsterdam, 1965, p. liS-130, Kamchatka Trench area (translation) .-Dok text figs. 1-5 (graphs, phylogenetic diagram, lady Acad. Sci. USSR, Earth Sci. sec., v. 174, correl. diagram) , table I.-Absolute dating of May-June 1967, p. 216-217, text fig . 1 (graphs). abundance changes and evolution in Foram SALAJ, JOZ EF, and JEND REJAKOVA , OTILIA. Die inifera. Foraminiferen aus der Oberen Trias der West PICHA, FRANTISEK, HANZLIKOVA, EVA , and CtCHA, karpaten.-Geol. Sbornik, Geol. Carpathica, IVAN. Geology of the Cejc-Zajeci zone (Eng Bratislava, v. IS, No.2, Nov. 1967, p. 311- lish summary of Czech text) .-Sbornik Geol. 313, pis. 19, 20.-Assemblages photographed. Ved, Geologie, rada G, sV. 13 , 1965, p. 37-74, SCHMID, MANFRED E. Das Genus A ustrocolomia pis. 1-9, text fi g. 1 (diagram), tables 1-3.-In Oberhauser, 1966 (Foraminifera, Nodosari eludes illustrations of key fossil s between Mae idae) .-Austria Geol. Bundes., Verhandl., Heft strichtian and Oligocene. 1-2, 1967, p. 189-192, pI. I.-Emendation; the PI ERCE, STANLEY, Kossoy, VICTOR, VALENTI, ROB wall is single-layered and has an elevated ring ERT, and SMETANA, DENNIS G. Cytochemical like portion at the upper end of each chamber. studies on the test of A Ilogromia laticollare. Zwei neue pl anktonische Foraminiferen aus dem Micropaleontology, v. 14, No.2, April 1965, Badener Tegel von Sooss, NO.-Ann. Natur p. 242-246, text fi gs. 1, 2 (photomicrographs) . hist. Mus. Wien, Band 71, Nov. 1967, Kiihn PROSN IAKOVA , L. V. Orbitolina from the Lower Festschrift, p. 347-352, text figs. 1, 2 (draw Cretaceous deposits of the Crimean lowlands ings) .-Globigerilloides kuehlli and Globiger (English summary of Russian text) .-Paleont. illopsis grilli from the Tortonian. Sbornik, No.4, vyp. 1, Izdat. L'vov. Univ., SCHROEDER, ROLF, and CONRAD, MARc-A . Huit 1967, p. 43-49, pis. 1-3 , text fig. 1 (drawing). ieme note sur les Foraminiferes du Cretace in REGUANT, SALVADOR. Une echelle de Nummulites ferieur de la region genevoise. Eopalorbitolilla pour la stratigraphie Sud-Pyreneenne.-Mem. charollaisi, n. gen., n. sp., un Orbitolinide du Bureau de Recherches Geol. et Min., No. nouveau du Barremien a facies urgonien. 5S, Colloque sur 1'J:;ocene, Paris, May 1965, p. Compte Rendu des Seances Soc. Physique Hist. 63-70, text figs. 1-5 (map, graphs).-Correla Nat. Geneve, n. ser., v. 2, fasc. 3, Sept.-Dec. tion by size of specimens. 1967 (March 1968), p. 145-162, pis. 1-4, text REISS, Z. Victoriella (Foraminiferid a) from Israel. fi gs. 1-4 (map, columnar section, drawings) . -Israel Jour. Earth-Sci., v. 16, No.3, Sept. SEIGLIE, GEORGE A. Systematics of the foramin 1967, p. 111-119, pis. 1, 2.-V. cOll oidea in ifers from Araya-Los Testigos shelf and upper the Oligocene. slope, Venezuel a, with special reference to sub ROCHA, A. TAVARES, and GOMES, J. DO NASCIM ENTO. order Rotaliina and its distribution.-Carib Ensaio critico sobre a morfo logia e ocorrencia bean Jour. Sci., v. 7, No. 3-4, Sept.-Dec. 1967, de Iberina lusitanica (Egger) (foraminifero). p. 95-133, 82 figs. on 8 pls.-IIIustrated system -Servicos Geol. de Portugal, Comunicac,:oes, atic catalog ineludes 201 species. Percentage tomo LI, 1967, p. 169-219, pis. 1-23 , text fig. of total benthonic population is given. 1-4 (diagrams, graph, maps), tables 1, 2.-In SERPAGLI, E., and SIROTTI, A. Eta aquitaniana lower Kimmeridgian to lower Valanginian in delle brecciole a lepidocieline e miogipsine Portugal. delle "Arenarie del Monte Cervarola" (Appen RUDDIMAN , WILLIAM F. Historical stability of the nino Sett.) .-Boll. Soc. Paleont. Ital., v. 6, No. Gulf Stream meander belt: foraminiferal evi 1, 1967, p. 18-29, pis. 4, 5, text figs . 1-3 (map, dence.-Deep-Sea Research, v. 15, No.2, April graphs, drawing), tables 1, 2.-Quantitative 1968, p. 137-148, text figs. 1-6 (maps), tables study of Lepidocyclina and Miogypsilla per 1, 2.-High percentages of equatorial species mits the dating to be lower Aquitanian. of planktonics in surface sediments under the SKINNER, JOHN W., and WILDE, GARNER L. Per present-day meander belt. mi an Foraminifera from Tunisia.-Univ. Kan SAAKJ AN-G EZALJAN, N. A., and MARTIROSJAN, Ju. sas Paleont. Contr., Paper 30, Nov. 30, 1967, CONTRIBUTIONS FROM THE CUSHMAN F OUNDATION FOR F ORAMIN IFERAL RESEARCH 177
p. 1-22, pIs. 1-32, text figs. 1-3 (maps, check Kanagawa Prefecture, Japan (in Japanese with list) .-Fourteen species, 10 new. English abstract) .-Contribs. to celebrate Prof. Ichiro Hayasaka's 76th Birthday, Dec. 1967, SMITH, P. B. , and EMILIANI, C . Oxygen-isotope p. 211-224, text figs . 1-4 (distrib. maps, graph, analysis of Recent tropical Pacific benthonic drawing), tables 1-5 .-Based on Foraminifera, Foraminifera.--Science, v. 160, No.3 834, June temperature was similar to present under the 21, 1968, p. 1335-1336, text figs. I, 2 (map, warm Kuroshio Current. graph), table I.-Off western central America, results of analyses for temperature turn out UJIIE, HIROSHI. Distribution of living planktonic Foraminifera in the southeast Indian Ocean. as expected. Bull. Nat. Sci. Mus., v. 11, No. I, March 15, STOERMER, NORBERT, and WI ENHOLZ, EVA . Mi 1968, p. 97-125, pis. 1-10, text figs. 1-23 (dis krobiostratigraphie an der Lias/ Dogger-Grenze trib. maps, graphs), tables I, 2.- lllustrations in Bohrungen nordlich der Mit:eldeutschen and abundance maps for 16 species. Hauptscholle.-J ahrbuch fiir Geologie, Berlin, VENGLINSKI, I. V. A new representative of the Band I, 1965 ( 1967), p. 533-591 , pIs. 1-10, Sigm oi/illa from the Lower Miocene deposits tables 1-9 (range charts) .-A few Foraminif of the Transcarpathians (English summary of era, 2 species new. Russian text) .-Paleont. Sbornik, No.4, vyp. STRANIK, ZDENEK, BEN EsovA, EVA , and PI CHA, I, Izdat. L'vov. Univ., 1967, p. 58-61 , 1 pl. FRANTISEK. Geology of the deep boring Bul Sigmoi/illa abbreviata sp. nov. hary-I (English summary of Czech text) . VENKAC HALAPATHY, V. Some new types of Nodo Sbornik Geol. Ved, Geologie, rada G, sV. 13 , sari ida from Berriasian and Valanginian depos 1968, p. 75-131 , pis. 1-9, text figs . 1-4 (colum its of Crimea (in Russian) .- Bull. Moscow nar section, geol. sections, map), tables 1-4. Soc. Naturalists, Geol. Ser., v. 43, pI. I, 1968, Includes illustrations of Foramini fera from the p. 83-96, I pI., text figs. 1-7 (diagrams, graphs). boring, from MaIm to Tortonian. -Three Frolldieu/aria, 1 Lillgulilla and I Sar TAMBAREAU, Y., and VILLATTE, 1. Les zones de aeellaria, all new, and Pseudosaracellaria gen. grands Foraminiferes du Paleocene (Thaneti nov. (genotype P. truneata sp. nov.) . en-Sparnacien) de I'avant-pays Pyreneen a I'est VIALOV, O. S., and DABAGIAN, N . V. Notes on du plateau de Lannemezan et leur extension. some Paleogene Foraminifera from Grzybow Mem. du Bureau de Recherches Geol. et Min., ski's collection (English summary of Russian No. 58, Colloque sur l'E:ocene, Paris, May text) .-Paleont. Sbornik, No.4, vyp. 1, Izdat. 1968, p. 57-62, tables 1, 2. L'vov. Univ., 1967, p. 24-34, pis. 1-3, text figs. TORlYAMA, Ryuzo, and KANM ERA , KAM ETOSHI. 1-3.- Three species of Delldrophrya. Fusulinacean fossils from Thailand, Part II. VITALlS-ZILAHY, LIDYA. Zones provisoires de Fo Two new Permian genera from Thailand. raminiferes planctoniques dans la serie E:ocene Geol. and Paleo. of Southeast Asia, v. 4, Feb. du bassin de Dorog en Hongrie.-Mem. du 25, 1968, p. 29-44, pIs. 6-8, text figs . I, 2 Bureau de Recherches Geol. et Min., No. 58, (map, stratigraphic section) .-Thai/alldilla n. Coll oque sur l'Eocene, Paris, May 1968, p. gen. (type species T. buravasi n. sp.) and N eo 131-135, table 1 (range chart) .--Six plank thai/alldilla n. gen. (type species N. pitakpaiv tonic zones. ani n. sp.), both in the Neoschwagerinidae, VOLOSHINA, A. M. Eight species of Ataxophrag subfamily Thailandininae n. subfam. miidae (Foraminifera) from the Upper Cre UBALDO, MARIA DE LOURDES. Contribuic;ao para 0 taceous and Paleocene of the east part of the estudo dos Foraminiferos recentes do Arqui Crimea (English summary of Russian text). pelago dos Ac;ores.- Rev. Fac. Ciencias Lis Paleont. Sbornik, No.4, vyp. I , Izdat. L'vov. boa, ser. C, v. 15, fasc. 1, 1967, p. 35-63, pIs. Univ., 1967, p. 50-57, pI. 1 (in 2 pts.), 2, 3. 1-6.-IIlustrated systematic catalog of 48 spe Eight species, 4 new. cies and varieties, none new, from shore WILLE-JANOSCH EK, URSULA. Zur Abgrenzung von samples around the Azores. G /oborota/ia aragollellsis aragoll ellsis Nuttall UCHlO, TAKAYAsu. Foraminiferal assemblages in gegen G. aragoll ellsis eaucasiea Glaessner (Fo the vicinity of the Seto Marine Biological Lab raminifera) aus dem Eozan von Schorn, Salz oratory, Shirahama-cho, Wakayama-ken, Japan burg (Osterreich).-Ann. Naturhist. Mus Wien, (Part I) .-Publ. Seto Marine BioI. Lab., v. 15 , Band 71 , Nov. 1967, Kiihn-Festschrift, p. 395- No.5, March 1967, p. 399-417, text fig . I 400, I pl.-Transitional forms illustrated. (map), tables 1-5.-Percentage analysis of 6 RUTH TODD samples. U. S. Geological Survey Paleo-temperature of the Naganuma Formation, Washington, D. C . 20560 CUSHMAN FOUNDATION FOR FORAMINIFERAL RESEARCH. INC. 1968
PATRONS
ARTHUR N. DUSENBURY, JR., 19 Sherman Ave., White Plains, New York WALDO L. SCHMITT, U. S. National Museum, Washington, D . C. SUN OIL COMPANY, Dallas, Texas
FELLOWS
HAROLD V. ANDERSEN, Louisiana State University, Baton Rouge, Louisiana HARRY W. ANISGARD, Humble Oil & Refining Co., New Orleans, Louisiana ZACH M. ARNOLD, Univ. California, Berkeley, California ORVILLE L. BANDY, Univ. Southern California, Los Angeles, California C. LOTHROP BARTLETT, 842 East Drive, Beaumont, Texas R. STANLEY BECK, 621 Truxtun, Bakersfield, California LEROY E. BECKER, R. D. I, Spencer, Indiana NOEL BROWN, JR., Esso Production Research Co., Houston, Texas JAMES BUSH, Box 2936, Palm Beach, Florida CHEVRON RESEARCH CORP ., La Habra, California RICHARD CiFELLI, U. S. National Museum, Washington, D. C. DANA K. CLARK, Shell Oil Co., Los Angeles, California W. STORRS COLE, 310 Fall Creek Drive, Ithaca, New York G. ARTHUR COOPER, U. S. National Museum, Washington, D. C. RAYMOND C. DOUGLASS, U. S. National Mmeum, Washington, D. C. JOHN B. DUNLAP, 167 O.K. Ave., New Orleans, Louisiana ROBERT L. ELLISON, Univ. Virginia, Charlottesville, Virginia Esso PROnUCTION RESEARCH CO ., Houston, Texas ROBERTA K. SMITH EV ERN DEN, U. S. National Museum, Washington, D. C . M. A. FURRER, Creole Petroleum Corp., Caracas, Venezuela J. B. GARRETT, JR., Pan American Petroleum Corp., Houston, Texas GEOLOGISCH-PALAONTOLOGISCH ES INST., Univ. Kiel , Kiel, Germany GEORGE L. HARRINGTON, 850 Webster St., Palo Alto, California HOLLIS D. HEDBERG, 118 Liberty Place, Princeton, New Jersey LLOYD G. HENBEST, U. S. Geological Survey, Washington, D. C. ISMAIL A. KAFESCIOGLU, Western Reserve University, Cleveland, Ohio MRS. F. H. KJERSTEAD , Smith College, Northampton, Massachusetts ROBERT M. KLEINPELL, 5959 Margarido Drive, Oakland, California H. G . KUGLER, Natural History Museum, Basel, Switzerland DORIS L. Low, U. S. Geological Survey, Washington, D. C. GEORGE W. LYNTS, Duke University, Durham, North Carolina DONALD A . MYERS, Federal Center, Denver, Colorado ENRICO F. DINAPOLI, Via G. A . Guttani 14, Rome, Italy NATUR. VEREIN FUR STEIERMARK, Graz, Austria KA:rHIlRINE V. W. PALMER, Paleontological Research Institution, Ithaca, New York FRANCES L. PARKER, Scripps Institution of Oceanography, La Jolla, California FRED B PHLEGER, Scripps Institution of Oceanography, La Jolla, California MORTON POLUGAR, Standard Oil Co. of California, OildaJe, California C. M. QUIGLEY, 5303 Brae Burn Drive, Bellaire, Texas CHARLES A. Ross, Western Washington State College, Bellingham, Washington K. NORMAN SACHS, U. S. National Museum, Washington, D. C. F. MARlON SETZER, 536 College St., Bellaire, Texas CHARLES R. STELCK, University of Alberta, Edmonton, Canada HANs E. THALMANN, P. O. Box 4407, Stanford, California M. RUTH TODD, U. S. Geological Survey, Washington, D. C. DONALD F. TOOMEY, Pan American Petroleum Corp., Tulsa, Oklahoma WALTER H. TRENCHARD, 309 Theodora Blvd., Lafayette, Louisiana JAMES A. WATERS, 308 Shadywood Lane, Seagoville; Texas R. T. D . WICKENDEN, 2757 Dunlevy St., Victoria, Canada VIRGIL WINKLER, c/ o Creole Petroleum Corp., Caracas, Venezuela GORDON A. YOUNG, c/ o Mene Grande Oil Co., Caracas, Venezuela INDEX TO VOLUME XIX, 1968
Acceleration of the evolutionary rate in the Orbulilla lineage. By D. G. Jenkins ...... _... __ 133 Albani, A. D.: Recent Foraminiferida from Port Hacking, New South Wales ...... 85 A IIdersellia rumalla, n. gen., n. sp., and some taxonomic observations on the subfamily Valvulininae. By Theodor Neagu ...... 120 A new genus of the Haplophragmoidinae. By D. S. Dhillon ...... 140 A new species of Pselldogllembelina from the Upper Cretaceous of Texas. By G . C. Esker, III ...... 168 An intertidal Margillopora colony in Suva Harbor, Fiji. By R. K Smith ...... 12 Annotated bibliography of Paleozoic nonfusulinid Foraminifera, Addendum 5. By D. F. Toomey and B. Mamet ...... 41 Archallispira, Notes on the synonymy of R eichelillella and . By G. A. Seiglie and C. W. Poag ...... 30 A taxonomic note on Massililla carillala (Fornasini , 1905). By K. Atkinson. 165 Atkinson, K: A taxonomic note on Massi/ina carillala (Fornasini, 1905) ...... 165 Atkinson, K: The genus Sigmoi/opsis Finlay, 1947, from Cardigan Bay, Wales ...... 160
Bock, W. D.: Two new species of Foraminifera from the Florida Keys 27 Buzas, M. A.: On the spatial di stribution of Foraminifera ...... I
California, Preliminary report on some littoral foraminifera from Tomal es Bay. By D. Maurer 163 Church, C.: Memorial: Dr. Joseph John Graham .. 81 Correction in Pagination ...... 131 Corrections ...... 32 Cretaceous of Texas, A new species of Pselldogllem belilla from the Upper. By G . C. Esker, Til 16
Designation of a lectotype of Clobolfllllcalla rosella (Carsey). By G. C. Esker, III ...... _.. I 0 Dbillon, D. S.: A new genus of tbe Haplopbragmoidinae from Malaysia ...... _...... __ ...... _... _ .... _ I ~O Discocyciilla from Pondicberry, Soutb India. By B. K Samanta ...... _.... _. 0 Distribution of Foraminifera, On the spatial. By M. A. Buzas ...... _...... _...... _..... _ I
Eocene faunule from the basal San Juan Bautista Formation of California. By J. N. Waters I Esker, G. C., ITT: A new species of Pseudoguembelilla from tbe Upper Cretaceous of Texas 16 Esker, G . C., Ill: Designation of a lectotype of Clobolruncana rosella (Carsey) ...... _ I 0 Evolutionary rate in the Orbulilla lineage, Acceleration of the. By D. G. Jenkins 133
Fiji, An intertidal Margillopora colony in Suva Harbor. By R. K. Smith ...... 12 Florida Keys, Two new species of Foraminifera from the. By W. D. Bock ...... 2 Foraminifera from the Florida Keys, Two new species of. By W. D. Bock ...... 27 Foraminifera from Tomales Bay, California, Preliminary report on some littoral. By D. Maurer 163 Foraminifera, On the spatial distribution of. By M. A. Buzas ...... I Foraminifera, Recent Literature on. By Ruth Todd ...... 33 , 75, 123 , 2 Foraminiferida, Recent from Port Hacking, New South Wales. By A. D . Albani . 5
Clobolfllllcalla rosella (Carsey), Designation of a lectotype of. By G. C. Esker, III o Graham, Dr. Joseph John, Memorial. By C. Church ...... I Gupta, S. C. and M. Mohan: The microfauna and age of the "Gypseous Shales," Western Kut h. India ...... _I
Haplophragmoidinae, A new genus of the, from Malaysia. By D. S. Dhillon 140
India, South, Discocyclilla from Pondicherry. By B. K. Samanta ...... 0 India, The microfauna and age of the "Gypseous Shales," Western Kutch. By M. Mohan and S. C. Gupta ...... _I
Jenkins, D . G.: Acceleration of the evolutionary rate in the Orbulilla lineage .. 133
Literature on the Foraminifera, Recent. By Ruth Todd ...... 33 , 75, 123. I -_ Littoral Foraminifera from Tomales Bay, California, Preliminary report on some. By D. Maurer 1 3
Malaysia, A new genus of the Haplophragmoidinae from. By D. S. Dhillon ...... 1.;0 Mamet, B. and D. F. Toomey: Annotated bibliography of Paleozoic nonfusulinid Foraminifera. Addendum 5 ...... _. ~ 1 Margillopora colony in Suva Harbor, Fiji , An intertidal. By R. K Smith ...... 1_ Massililla carillala (Fornasini, 1905) , A taxonomic note on. By K Atkinson ...... 165 Maurer, D.: Preliminary report on some littoral foraminifera from Tomales Bay, California .. 163 Memorial: Dr. Joseph John Graham. By C. Churcb ...... _... _ I Mohan, M. and S. C. Gupta: The microfauna and age of the "Gypseous Shales," Western Kutch, India ...... _.. 21 Neagu, Theodor: A Ildersellia rumana, n. gen., n. sp., and some taxonomic observations on the sub- family Valvulininae ...... 120 New South Wales, Recent Foraminiferida from Port Hacking. By A. D. Albani ...... 85 New Zealand, Late Eocene and Early Oligocene planktonic Foraminifera from Port Elizabeth and Cape Foulwind. By M. S. Srinivasan ...... 142 Nonfusulinid Foraminifera, Annotated bibliography of Paleozoic, Addendum 5. By D. F. Toomey and B. Mamet ...... 41 Notes on the synonymy of R eichelillella and Arcllnispira. By G. A. Seiglie and C. W. Poag ...... 30
On the spatial distribution of Foraminifera. By M. A. Buzas ...... 1 Orbulina lineage, Acceleration of the evolutionary rate in the. By D. G. Jenkins ...... 133
Pagination, Correction in ...... 131 Paleozoic nonfusulinid Foraminifera, Addendum 5, Annotated bibliography of. By D. F. Toomey and B. Mamet ...... 41 Poag, C. W. and G. A. Seiglie: Notes on the synonymy of R eichelinella and A rcollispira 30 Preliminary report on some littoral foraminifera from Tomales Bay, California. By D. Maurer ...... 163 Pseudoguembelilla from the Upper Cretaceous of Texas, A new species of. By G. C. Esker, III...... 168
Recent Foraminiferida from Port Hacking, New South Wales. By A. D. Albani 85 Recent Literature on Foraminifera. By Ruth Todd ...... 33, 75, 123 , 172 Reichelillella and Arcanispira, Notes on the Synonymy of. By G. A. Seiglie and C. W. Poag ...... 30
Samanta, B. K.: DiscocyC/illa from Pondicherry, South India ...... 70 San 1 uan Bautista Formation of California, Eocene Faunule from the basal. By J. N. Waters ...... 18 Seiglie, G. A. and C. W. Poag: Notes on the synonymy of Reichelillella and Arcanispira ...... 30 Sigmoi/opsis Finlay 1947 from Cardigan Bay, Wales. By K. Atkinson ...... 160 Smith, R. K. : An intertidal Margillopora colony in Suva Harbor, Fiji ...... 12 Spatial distribution of Foraminifera, On the. By M. A. Buzas ...... 1 Srinivasan, M. S.: Late Eocene and Early Oligocene planktonic foraminifera from Port Elizabeth and Cape Foulwind, New Zealand ...... 142
Texas, A new species of Pseudoguembelino from the Upper Cretaceous of. By G. C. Esker, III . 168 The genus Sigmoi/opsis Finlay 1947 from Cardigan Bay, Wales. By K. Atkinson ...... 160 The microfa una and age of the "Gypseous Shales," Western Kutch, India. By M. Mohan and S. C. Gupta ...... 21 Todd, Ruth: Recent Literature on the Foraminifera ...... 33,75, 123 , 172 Toomey, D. F. and B. Mamet: Annotated bibliography of Paleozoic nonfusulinid Foraminifera, Addendum 5 ...... 41 Two new species of Foraminifera from the Florida Keys. By W. D. Bock ...... 27
Valvulininae, Alldersellia rumana, n. gen., n. sp., and some taxonomic observations on the subfam- ily. By Theodor Neagu ...... 120
Wales, The genus Sigmoi/opsis Finlay 1947 from Cardigan Bay. By K. Atkinson ...... 160 Waters, 1. N.: Eocene faunule from the basal San Juan Bautista Formation of California ...... 18