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88 SYSTEMATIC PART Figs.1-2

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88 SYSTEMATIC PART Figs.1-2 figs.1-2; pl. 2, fig. 1); Geological Survey of Hungary, Budapest and material from the marine Badenian comes from the Locus typicus: Budapest, XIVth district, Hungary nearby Leitha Limestone of Bia. Age: Late Badenian (Early Serravallian), Middle Miocene Badenian (Langhian-Early Serravallian) – Kemence, Hungary NHMW: 1 specimen (NHMW 2004z0001/0054) Material: Late Badenian (Early Serravallian) – Armeniş (= Örményes), Early Badenian (Langhian-Early Serravallian) – Retznei Romania [Weissenegg Fm., Lafarge quarry (formerly Perlmoser)], NHMW: 2 specimens and 1 fragment [MAFI Ech-290-292 Styria, Austria (reference material of VADÁSZ, 1915)] WANZENBÖCK coll.: 2 specimens (no inv. nos.) Late Badenian (Early Serravallian) – Biatorbágy (= Bia), near Badenian (Langhian-Early Serravallian) – Bad Fischau, NÖ, Budapest, Hungary Austria NHMW: 3 specimens [NHMW 1859.XLIII.26; MAFI NHMW: 1 specimen (NHMW 1904.VIII.62) Ech-137 (figured specimen of VADÁSZ, 1915: fig. 12), ? Badenian (Langhian-Early Serravallian) – Brunn am Steinfeld, MAFI Ech-228 (holotype of Scutellina hungarica VADÁSZ, NÖ, Austria 1915: pl. 10 (4), figs. 2-3)] NHMW: 2 specimens (NHMW 2004z0001/0048, Late Badenian (Early Serravallian) – Budatétény (= Tétény), 2004z0001/0051) Hungary Badenian (Langhian-Early Serravallian) – Hainburg NHMW: 1 specimen (NHMW 1997z0178/2090) (Halitherium locality of STACHE, 1867), NÖ, Austria Late Badenian (Early Serravallian) – Buituri (= Bujtur), NHMW: 1 specimen (NHMW 1867.XXII.3) Romania Badenian (Langhian-Early Serravallian) – Haschendorf, W MAFI: 2 specimens [MAFI Ech-240 (figured specimen of Deutschkreutz, Bgld, Austria VADÁSZ, 1915: fig. 14), Ech-264 (reference material of NHMW: 5 specimens (NHMW 1848.III.63-64, 1848.III.66, VADÁSZ, 1915)] 2004z0001/0052, 2004z0001/0060) Late Badenian (Early Serravallian) – Devínska Nová Ves Badenian (Langhian-Early Serravallian) – Hasenberg, near (= Neudorf an der March), Slovak Republic Brunn am Gebirge, NÖ, Austria NHMW: 1 fragment (NHMW 1857.XIX.46), 8 specimens NHMW: 2 specimens (NHMW 1904.VIII.61, 2004z0001/ [NHMW 1851.VI.120, 1851.VI.120.6, 1866.I.1256, 0047) 1929.No.16, 1964/589 (holotype of Scutella Badenian (Langhian-Early Serravallian) – Kalksburg, Vienna, vindobonensis secunda SCHAFFER, 1962: pl. 19, fig. 4), Austria 2004z0001/0046, 2004z0111/0003], 2 partially leached NHMW: 12 specimens [NHMW 1857.38.21, 1857.38.22, specimens (NHMW 1859.XXXVIII.100) 1858.III.13-14, 1858.XXV.40, 1904.VIII.56, 1904.VIII.57 Late Badenian (Early Serravallian) – Sandberg, Devínska Nová (figured specimen of SCHAFFER, 1962: pl. 15, fig. 2) Ves (= Neudorf an der March), Slovak Republic 1904.VIII.58, 1904.VIII.59 (lectotype of Scutella NHMW: 3 specimens (NHMW 2004z0001/0050). 2 vindobonensis LAUBE, 1869), 2004z0001/0059, partially leached and fragmented specimens (NHMW 2004z0072/0002, 2004z0111/0001] 2004z0001/0049) Badenian (Langhian-Early Serravallian) – Neckenmarkt, Bgld, ? Late Badenian (Early Serravallian) – Gârbova de Sus Austria (= Felsö-Orbó), Romania NHMW: 6 specimens [NHMW 291/1961, 390/1961 NHMW: 2 specimens [MAFI Ech-130 (figured specimen of (holotype of Scutella multiconcava SCHAFFER, 1962: pl. 18, VADÁSZ, 1915: fig. 16), Ech-234 (figured specimen of figs. 1-2), 391/1961 (figured specimen of SCHAFFER, 1962: VADÁSZ, 1915: fig. 15)] pl. 15, fig.3), 1964/590 (figured specimen of SCHAFFER, Late Badenian (Early Serravallian) – Globu Craiovei 1962: pl. 17, fig. 4), 2004z0001/0055-56, 2004z0001/ (= Globukrajova), Romania 0058] NHMW: 1 specimen and 1 fragment [MAFI Ech-286, 289 Badenian (Langhian-Early Serravallian) – Oslip, Bgld, Austria (reference material of VADÁSZ, 1915)] NHMW: 1 specimen (NHMW 1976/1843/18) and Late Badenian (Early Serravallian) – Harka, Györ-Moson- numerous test fragments (NHMW 2003z0083/0006) Sopron, Hungary Badenian (Langhian-Early Serravallian) – Pötzleinsdorf, NHMW: 1 specimen [MAFI Ech-284 (reference material of Vienna, Austria VADÁSZ, 1915)] NHMW: 1 specimen (NHMW 2004z0072/0001) Late Badenian (Early Serravallian) – Iablanitja (= Bélajablánc), Badenian (Langhian-Early Serravallian) – Romania Rauchstallbrunngraben, near Baden, NÖ, Austria NHMW: 1 specimen [MAFI Ech-282 (reference material of NHMW: 1 specimen (NHMW 2004z0001/0062) VADÁSZ, 1915)] Badenian (Langhian-Early Serravallian) – Siegenfeld, Gaaden Late Badenian (Early Serravallian) – Minişu des Sus Bay, NÖ, Austria (= Felménes), Romania NHMW: 1 specimen (NHMW 2004z0111/0002) NHMW: 1 specimen and 1 fragment [MAFI Ech-285, 288 Late Badenian (Early Serravallian) – Müllendorf (Mühlen- (reference material of VADÁSZ, 1915)] dorfer Kreide AG quarry), Bgld, Austria Late Badenian (Early Serravallian) – Paptelek, Hungary NHMW: 19 specimens (NHMW 1981/55/20, 1997z0178/ NHMW: 1 specimen [MAFI Ech-263 (reference material of 1754, ../1756-1757, 2002z0181/0023-34, 2004z0112/ VADÁSZ, 1915)] 0009-11), Late Badenian (Early Serravallian) – Sopron (= Ödenburg), Late Badenian (Early Serravallian) – Unterpetersdorf (= Hungary Alsópéterfa), Bgld, Austria NHMW: 1 specimen (NHMW 2004z0001/0053) NHMW: 2 specimens [MAFI Ech-283 (reference material of Late Badenian (Early Serravallian) – Tusa (= Tusza), Romania VADÁSZ, 1915)] NHMW: 1 specimen and 2 fragments [MAFI Ech-287 (reference material of VADÁSZ, 1915), Ech-158 (figured Foreign material for comparison: specimen of VADÁSZ, 1915: fig. 17)] Badenian (Langhian-Early Serravallian) – “Bia gravel”, Soskút, Hungary Dimensions (in mm): NHMW: 4 specimens (NHMW 1865.XXX.78-80) Inv. No. TL TW TH Remark: According to VADÁSZ (1915: 105, footnote) there NHMW 1997z0178/1756 158.6 172.5 28.9 are only strata of Sarmatian age outcropping at Soskút NHMW 1997z0178/1757 125.5 137.8 20.1 88 SYSTEMATIC PART SYSTEMATIC PART 89 NHMW 1964/589 133.3 ~140 21.5 Differential diagnosis: NHMW 390/1961 95.8 99.4 20.0 The type species of Parascutella, Ps. leognanensis (LAMBERT, NHMW 1964/590 92.6 102.3 ~17 1903) from the Early Miocene of Western France, differs from NHMW 1904.VIII.59 150.3 163.8 30.6 Ps. gibbercula by its flatter profile, without conspicuous eleva- tion in adapical interambulacrum 5, its broader interporiferous Description: zones and more closed petals [compare AGASSIZ, 1841a (under Size and shape: Test large to very large, ranging from 15 (juve- the name Scutella subrotunda) and DURHAM, 1953]. nile specimen with gonopores still closed) to 159 mm TL. In a Parascutella paulensis (AGASSIZ, 1841), from the Burdigalian 44.6 mm TL specimen the gonopores are still closed. The out- of the Mediterranean and Eggenburgian to Karpatian of the line of the test is subcircular, slightly transversely elongate Central Paratethys, likewise differs from Ps. gibbercula by its (rarely antero-posteriorly elongated individuals are found, less strongly developed marginal indentations, its rounded most of these, however, show a regenerated marginal area and posterior margin without or with faint anal notch, and flat- thus do not show the characteristic outline of this species). The tened profile with central maximum height. anterior margin is rounded with shallow indentations corre- Parmulechinus hoebarthi (KÜHN, 1936) from the Early Egg- sponding to the ad-, inter- and perradial sutures. The posterior enburgian (Early Burdigalian) differs from Parascutella gibber- margin is transversely truncated, with two strong indentations cula by it shorter petals (c. half the length of the corresponding corresponding to the perradial sutures of the ambulacra I and test radius), its antero-posteriorly elongated outline, and the V. The anal notch is shallow but distinct and rather broad. In different position of its peristome (between the 4th or 5th pair of juveniles and specimens with thicker coronal plates (i.e. speci- postbasicoronal plates). mens from high-energy settings, e.g. from Neckenmarkt), For the difference between Ps. gibbercula and Ps. jaquemeti however, it may be deeper. The maximum width lies slightly (DE LORIOL, 1902), Ps. striatula (DE SERRES, 1829), Ps. michaleti posterior of the apical disc, at the line interambulacra 1b-4a. (LAMBERT, 1906), Ps. depereti (LAMBERT, 1912), and Ps. savini In profile the test is distinctly wedge shaped, with a thicker, (LAMBERT, 1915) the reader is referred to PHILIPPE (1998). rounded anterior margin and a thinner, relatively sharp poste- rior margin. The maximum height lies 10 to 13 % of the Discussion: TL posterior of the centre of the apical disc along the central Originally the species was named Scutella gibercula by DE suture of interambulacrum 5. The test width is about 103 SERRES (1829: 156). AGASSIZ (1841a: 86) changed the name to S. to 110 % of the TL and the height about 13-25 % of the gibbercula (there is no indication whether the change was in- TL. tentional or unintentional). According to the ICZN rules this Apical disc: The apical disc lies subcentrally and belongs to the represents an unjustified emendation or incorrect subsequent tetrabasal type. The madreporite is pentagonal and crowded spelling. The name S. gibbercula is in prevailing usage during with many small madreporic pores. Four gonopores are situat- the last 150 years and is attributed to the original author and ed at the apices of the madreporite, except in interambulacrum date of the original spelling. According to the ICZN it is there- 5. The ocular pores are small and rounded. fore deemed to be a justified emendation, respectively a cor- Ambulacra: The ambulacra are petaloid. The petals are straight, rect original spelling (in case of an unintentional change) (ICZN moderately closed distally and are usually flush with the
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