WTU Herbarium Specimen Label Data Generated from the WTU Herbarium Database October 5, 2021 at 9:54 am http://biology.burke.washington.edu/herbarium/collections/search.php

Specimen records: 672 Images: 86

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Label Query: Genus = "Lonicera" Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera chamissoi Bunge ex P. Kir. U.S.A., WASHINGTON, KING COUNTY: RUSSIAN FEDERATION, SAKHALIN REGION: Cascade Mountains of Western Washington: Cedar River Sakhalin Island, central part of island; along west side of the Orlovka Watershed. Rock Creek wetland system. Head into south end from River circa 25 kilometers west of town of Smirnykh and 4 kilometers rd 40. Drop into system at Binker culvert and head into west side of southeast of mouth of Ognannaya River. the wetland. Elev. 394 ft. Elev. 800 ft. 49° 43' 46.62" N, 142° 31' 27.36" E T22N R7E S16 NE; NAD 27, uncertainty: 500 m., Source: Northeast facing slope above river, with mix of Abies-Betula forest, Georeferenced, Georef'd by WTU Staff open Rubus-Epilobium meadows and small brushy stream gullies. Rock Creek wetland system. Head into south end from rd 40. Drop Fruit bright red; to 1.5 meters tall in open area. Phenology: into system at Binker culvert and head into west side of the wetland. Fruits. Origin: Native. Origin: Native. Ben Legler 913 26 Jul 2003 Clayton J. Antieau 01-33 26 May 2001 with Linda Cooley, Kristie Dinkin WTU-357809 Herbarium: WTU Caprifoliaceae Caprifoliaceae Lonicera caerulea L. Lonicera involucrata (Richardson) Banks ex Spreng. RUSSIAN FEDERATION, SAKHALIN REGION: U.S.A., WASHINGTON, KING COUNTY: Sakhalin Island, central part; along the upper Tym River 18 km due Halmar Gate Wetland; Cascade Mountains of Western Washington. east of Yasnoye and 6 km SW of summit of Mt. Khodovaya. Elev. 1400 ft. Elev. 754 ft. T22N R7E S1; NAD 27, uncertainty: 500 m., Source: 50° 36' 53.94" N, 142° 55' 16.2" E Georeferenced, Georef'd by WTU Staff Mixed forest of Larix, Abies, Betula, Alnus, and Populus; understory Ditches filled with water, drier hummocks with Spiraea sp., Juncus of Rosa, Cornus canadensis, grasses. Fruit blue, glaucous, oblong; effusus, Juncus supinus. Other in vicinity: Salix sp., small shrub in forest. Phenology: Fruits. Origin: Native. Lysichitum americanum. Gaultheria shallon growing from stumps; Menziesia ferruginea common. Upland island with doug fir, western Ben Legler 927 27 Jul 2003 hemlock, red alder. Growing on stumps within wetland. Phenology: Flowers. Origin: Native. WTU-357879 Suzanne Bagshaw 01-14 23 Jun 2001 with Roz Glasser, Mary Clare Schroeder, Neil Dijrani Caprifoliaceae Herbarium: WTU Lonicera chamissoi Bunge ex P. Kir. RUSSIAN FEDERATION, SAKHALIN REGION: Caprifoliaceae Sakhalin Island, on Schmidta Peninsula at northern end of island; along Sukharnyy River ca 7 km SSW of Tri Brata Mountain. Lonicera involucrata (Richardson) Banks ex Spreng. Elev. 394 ft. 54° 2' 2.58" N, 142° 48' 21" E U.S.A., WASHINGTON, WHATCOM COUNTY: Dense Picea-Betula forest, with understory of Cornus canadensis, Ross Lake National Recreation Area. Boundary trail going E along ferns, mosses; also some small logging clearings and buildings. Lightning Creek after suspension bridge. Lightning Creek Trail, Berries bright red; shrub to 1 meter tall in openings. Phenology: flowing stream crossing. Fruits. Origin: Native. 48° 52' 49.78858" N, 121° 31.20124" W; UTM Zone 10, 646000E, 5415864N; Source: Calc. from UTM, UTM from field notes. Ben Legler 1179 9 Aug 2003 Steep rocky slope with exposed bedrock balds. Sparse stand of Pseudotsuga menziesii and Pinus ponderosa. Acer macrophyllum, Herbarium: WTU Acer circinatum, Betula papriferus hugging stream. Growing near stream among Betula papyferis, Acer circinatum, Rosa nootkatensis. Dark purple fruits. Phenology: Fruits. Origin: Native. Caprifoliaceae Kimiora Ward 28 14 Jul 2002 Lonicera glenhii Fr. Schmidt Herbarium: WTU, NPS accession 632, catalog 23680 RUSSIAN FEDERATION, SAKHALIN REGION: Sakhalin Island, east coast; mouth of Tukhaya River circa 17 kilometers north of town of Vzmor'ye and 70 kilometers south of Caprifoliaceae Makarov. Elev. 131 ft. Lonicera glenhii Fr. Schmidt 48.01013°, 142.532024°; WGS 84, uncertainty: 1000 m., Source: Digital Map, Georef'd by John Haskins RUSSIAN FEDERATION, SAKHALIN REGION: Steep windswept slopes and cliffs above beach, with some dwarfed Sakhalin Island; in southern half at Sokol Research Station along Betula forest in depressions, mostly lower growing herbs and talus Belaya River 2 miles east of town of Sokol. on exposed slopes; parent rock partly calcareous. Berries bright red; Elev. 197 ft. shrub under Betula. Phenology: Fruits. Origin: Native. 47° 14' 26.34" N, 142° 46' 47.28" E Forested slope of Betula ermanii and Abies sachalinensis; Ben Legler 1307 20 Aug 2003 overlooking uncut hay fields. Immature fruit green; shrub to 2 meters tall. Phenology: Fruits. Origin: Native. WTU-358088 Ben Legler 744 19 Jul 2003 Herbarium: WTU Caprifoliaceae Caprifoliaceae Lonicera glenhii Fr. Schmidt Lonicera involucrata (Richardson) Banks ex Spreng. RUSSIAN FEDERATION, SAKHALIN REGION: U.S.A., WASHINGTON, OKANOGAN COUNTY: Sakhalin Island, southern part; Sokol Research Station east of town Okanogan National Forest. Road to Hart's Pass (Forest Road 5400). of Sokol; on hillside south of station. Elev. 5812 ft. Elev. 197 ft. 48° 41.482' N, 120° 37.927' W 47° 14' 26.34" N, 142° 46' 47.28" E At seep/small creek above and below road. Berries black; calyces Forested slope dominated by Abies, Picea, Betula, Acer; understory maroon. Phenology: Flowers & Fruits. Origin: Native. with Viburnum furcatum, Sasa, Cacalia, other herbs. Berries bright red; shrub to 1 meter tall on edge of trees. Phenology: Fruits. Origin: Sarah Gage 7454 26 Jul 2003 Native. with Richard G. Olmstead, Sheila Olmstead, Liz Van Volkenburgh, B. Lippett, Robert Goff, Marguerite Goff Ben Legler 1344 22 Aug 2003 WTU-362508 WTU-358117 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ×bella Zabel U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., MICHIGAN, WASHTENAW COUNTY: Near Blewett Pass, Wenatchee Mountains. Ann Arbor, Pontiac Trail at US23 overpass. Elev. 4000 ft. 42.319716°, -83.734115°; WGS 84, uncertainty: 896 m., Source: 47.33556°, -120.57778°; WGS84, uncertainty: 5000 m., Source: Digital Map, Georef'd by Richard Olmstead GeoLocate, Georef'd by WTU Staff Phenology: Fruits. On wooded slopes. Phenology: Flowers. Origin: Native. Richard G. Olmstead 89-003 21 Jun 1989 J. William Thompson 17005 5 Jul 1952 WTU-397757 WTU-171224

Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., OREGON, WALLOWA COUNTY: U.S.A., WASHINGTON, CLALLAM COUNTY: Hell's Canyon National Recreation Area. Forest Road 3965, north of Sequim. Hell's Canyon overlook. 48.07977°, -123.10253°; WGS 84, uncertainty: 1500 m., Source: Elev. 5367 ft. GeoLocate, Georef'd by WTU Staff 45° 7.716' N, 116° 50.052' W In woods. Phenology: Flowers. Origin: Native. Mixed conifer forest with meadow. Red fruit. Phenology: Flowers & Fruits. Origin: Native. J. M. Grant s.n. May 1905 Richard G. Olmstead 02-26 19 Jul 2002 WTU-20228 with Sara Tjessem, Shahid Naeem, Cindy Spurgeon, Don Knoke WTU-360737 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, ISLAND COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Coupe's Woods, near Coupeville, Whidbey Island. 48.22°, -122.685°; WGS 84, uncertainty: 5000 m., Source: U.S.A., WASHINGTON, SKAGIT COUNTY: GeoLocate, Georef'd by WTU Staff Okanogan National Forest. North Cascades Highway 20. Easy Pass Phenology: Flowers. Origin: Native. Trail Boardwalk. Elev. 3756 ft. Lloyd E. McElvain 60 12 May 1950 48° 35.27' N, 120° 48.144' W; T36N R16E S25; uncertainty: 9 m. Bottomland with: Mountain Hemlock, Western Red Cedar, Abies WTU-129213 amabilis, Thuja heterophylla, Tsuga heterophylla, Trisetum spicatum, Viburnum edule, Athyrium filix-femina, Rubus spectabilis, Rubus parviflora, Epilobium latifolium, Vaccinium, Glyceria, Viola, Caprifoliaceae Trillium, Clintonia, and Dryopteris. (GPS 3-4 sat.); Accuracy 26-30'. Phenology: Fruits. Origin: Native. Lonicera ciliosa (Pursh) Poir. ex DC. Ken Davis 03-40 27 Jul 2003 U.S.A., WASHINGTON, KITTITAS COUNTY: with Richard Robohm, Sarah Gage, Robert Goff, Margarite Goff, Southeast Easton, south of Lake Easton. Dale Blum Elev. 2360 ft. 47° 14' 15" N, 121° 11' 30" W; T20N R13E S11 WTU-362416 Phenology: Flowers. Origin: Native. Sarah Gage G&R 259 15 Jun 1992 with Sharon Rodman WTU-344666 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Cougar Mountain. Yellow Island. 47.54056°, -122.09806°; WGS 84, uncertainty: 10000 m., Source: 48.592222°, -123.031944°; WGS 84, uncertainty: 250 m., Source: GeoLocate, Georef'd by WTU Staff Georeferenced, Georef'd by WTU Staff Phenology: Vegetative. Origin: Native. On south side, among in open along trail to beach. Phenology: Flowers. Origin: Native. David M. Chapin s.n. 9 Oct 1982 P. Dunwiddie s.n. 13 May 1991 WTU-314241 WTU-333077 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, KLICKITAT COUNTY: About 2 airmiles northeast of White Salmon, near where powerline U.S.A., WASHINGTON, SKAMANIA COUNTY: crosses road on southwest-facing slope. Columbia River Gorge, 15.6 miles west of Beacon Rock State Park, Elev. 1600 ft. along Washington State Highway 830. T3N R11E; WGS 84, uncertainty: 2187 m., Source: Digital Map, 45.5605°, -122.23761°; WGS 84, uncertainty: 1000 m., Source: Georef'd by Joseph Stormer GeoLocate, Georef'd by WTU Staff Regrowth in powerline swath, and on old road through forest. On old Moist woodland. Phenology: Flowers. Origin: Native. road through forest. Uncommon vine, flowers bright orange. Phenology: Flowers. Origin: Native. Jared J. Davis 151 17 May 1953 Barbara Ertter 7535 14 Jun 1988 WTU-196172 WTU-318851 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, SNOHOMISH COUNTY: Marysville. U.S.A., WASHINGTON, KITSAP COUNTY: 48.05194°, -122.17583°; WGS 84, uncertainty: 3570 m., Source: Bainbridge Island. GeoLocate, Georef'd by Maria Yousoufian Elev. 0 ft. In woods. Phenology: Vegetative. Origin: Native. 47.65528°, -122.51667°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. M. Grant s.n. Jun 1929 At sea level. Phenology: Flowers. Origin: Native. WTU-43879 Mae E. Cornelius 5 23 May 1948 WTU-187658 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, SKAGIT COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. Foot of Mt. Erie, Fidalgo Island. 48.47639°, -122.56944°; WGS 84, uncertainty: 10000 m., Source: U.S.A., WASHINGTON, MASON COUNTY: GeoLocate, Georef'd by WTU Staff Lost Lake Road, 5 miles south of Shelton. In forest. Phenology: Flowers. Origin: Native. 47.142902°, -123.09944°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff Harold W. Smith 1025 12 Jun 1936 In gravelly soil. Phenology: Flowers. Origin: Native. WTU-35761 Phillips B. Freer 157 27 May 1950 WTU-135134 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, SPOKANE COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. 20 miles northeast of Spokane. 47.863187°, -117.121084°; WGS 84, uncertainty: 1000 m., U.S.A., WASHINGTON, LEWIS COUNTY: Source: GeoLocate, Georef'd by WTU Staff Mount Rainier. Canyon Bridge. In woods. Phenology: Flowers. Origin: Native. 46.76613°, -121.63586°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff Sister Mary Milburge 439 8 Jun 1932 Sandy, gravelly, & dry soil. Phenology: Vegetative. Origin: Native. WTU-19369 F. A. Warren 1761 31 Jul 1932 WTU-25195 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, STEVENS COUNTY: U.S.A., WASHINGTON, JEFFERSON COUNTY: (no locality given). Fort Worden State Park, by Port Townsend on the Quimper T35N R41E S4 SW 1/4; NAD 27, uncertainty: 805 m., Source: Peninsula. TRS2LL, Georef'd by Ben Legler Elev. 197 ft. Open woods along edge of road. Phenology: Flowers. Origin: Native. T31N R1W S35; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Monita Engvall 135 15 Jun 1958 Wooded headlands. With Pseudotsuga, Thuja, Rosa, Symphoricarpos, Rubus, Polystichum. Flowers purple-pink and WTU-196748 white; fruit red. a vine climbing over adjacent vegetation. Phenology: Flowers & Fruits. Origin: Native. Caprifoliaceae Richard R. Halse 4094 8 Aug 1990 Lonicera ciliosa (Pursh) Poir. ex DC. WTU-321676 U.S.A., WASHINGTON, THURSTON COUNTY: Road to Lang Lake. Caprifoliaceae Elev. 200 ft. 46.94841°, -122.84591°; WGS 84, uncertainty: 45000 m., Source: Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray GeoLocate, Georef'd by WTU Staff In roadside brush. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, KITSAP COUNTY: Charleston. H. E. Sovereign 449 11 May 1941 47.56201°, -122.65965°; WGS 84, uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff WTU-316125 Side of bluff overlooking sound. Phenology: Flowers & Fruits. Origin: Native. Caprifoliaceae T. C. Frye s.n. 14 Dec 1907 Lonicera ciliosa (Pursh) Poir. ex DC. WTU-20448 U.S.A., WASHINGTON, WHATCOM COUNTY: Bellingham; just south of town. Caprifoliaceae 48.75972°, -122.48694°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Rocky hillside. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, KING COUNTY: Lyman Benson 1509 1 Jun 1929 Clarke Beach Park, southeast end of Mercer Island, Lake Washington. WTU-43569 Elev. 33 ft. T24N R5E S30; NAD 27, uncertainty: 200 m., Source: Georeferenced, Georef'd by WTU Staff Caprifoliaceae Fruit red, transluscent; leaves glaucous below; climbing vine. Phenology: Fruits. Origin: Native. Lonicera conjugialis Kellogg P. F. Zika 14524 6 Oct 1999 U.S.A., WASHINGTON, SKAMANIA COUNTY: Near Flattop Mountain, on N78, 25 miles west of Glenwood. WTU-346839 Elev. 3493 ft. 46.02833°, -121.61278°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff Caprifoliaceae Canadian zone. Phenology: Fruits. Origin: Native. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray E. Hunn 669 2 Aug 1977 U.S.A., WASHINGTON, PIERCE COUNTY: WTU-271247 Near Steilacoom. 47.17°, -122.60139°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff Caprifoliaceae In woods. Phenology: Flowers. Origin: Native. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray J. William Thompson 9961 22 Aug 1933 U.S.A., WASHINGTON, ISLAND COUNTY: WTU-19366JWT Langley. 48.041667°, -122.406667°; NAD 27, uncertainty: 1000 m., Source: Georeferenced, Georef'd by WTU Staff On dry bluffs. Phenology: Flowers. Origin: Native. J. M. Grant 2063 Jun 1923 WTU-16609 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, FERRY COUNTY: Madrona Point Park, Eastsound, Orcas Island. Sanford Ranch, 8 miles south of Republic. Elev. 20-80 ft. 48.532554°, -118.73667°; WGS 84, uncertainty: 1000 m., Source: 48° 41' 30" N, 122° 54' W; T37N R2W S14; Source: Specimen GeoLocate, Georef'd by WTU Staff label Meadow on ranch; deep, wet soil. Phenology: Flowers. Origin: Growing along ground under Arbutus menziesii. Phenology: Flowers. Native. Origin: Native. J. Jorgensen 109 31 May 1936 Sarah Gage G&R 411 3 Jul 1992 with D. Sanford with Sharon Rodman WTU-9478JWT WTU-344641 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: U.S.A., WASHINGTON, SKAGIT COUNTY: Westhaven Beach, Westport. Bowman Bay area. Elev. 20 ft. 48.56194°, -122.39694°; WGS 84, uncertainty: 10000 m., Source: 46° 53' 30" N, 124° 7' 45" W; T16N R12W S1 GeoLocate, Georef'd by WTU Staff Behind dunes along northern shore. Sandy, shallow soil. Flowering Phenology: Flowers. Origin: Native. and fruiting. Phenology: Flowers & Fruits. Origin: Native. Lloyd E. McElvain 198 23 Jul 1950 Sarah Gage G&R 384 22 Jun 1992 with Sharon Rodman WTU-135242 WTU-345246 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, CLALLAM COUNTY: Ozette Island, 1.2 kilometers southwest of Cape Alava. U.S.A., WASHINGTON, ISLAND COUNTY: Elev. 197 ft. Cranberry lake, Deception Pass State Park, Whidbey Island. T31N R16W S27; NAD 27, uncertainty: 805 m., Source: TRS2LL, 48.14389°, -122.49111°; WGS 84, uncertainty: 10000 m., Source: Georef'd by Ben Legler GeoLocate, Georef'd by WTU Staff Occurs commonly on the NE slope of the island. Phenology: Fruits. Phenology: Flowers. Origin: Native. Origin: Native. Harold W. Smith 484 11 May 1936 Keith B. Aubry s.n. 18 Jul 1982 WTU-35762 WTU-315945 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Sphagnum bog 5 miles west of Redmond. Northwest of Leavenworth, Meadow Camp Road (trail), Tumwater 47.67412°, -122.227762°; WGS 84, uncertainty: 1000 m., Source: Mountain. GeoLocate, Georef'd by WTU Staff 47.59639°, -120.66028°; WGS84, uncertainty: 10000 m., Source: On edge of bog. Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. George Neville Jones 6390 20 Apr 1935 Maxine Hagen s.n. 21 May 1950 WTU-28750 WTU-332630 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, KITTITAS COUNTY: Southeast Easton, south of Lake Easton. U.S.A., WASHINGTON, CLARK COUNTY: Elev. 2360 ft. Green Mountain Resort Conservation Easement. 47° 14' 15" N, 121° 11' 30" W; T20N R13E S11 T2N R3E S20; T2N R3E S21; Willamette Meridian; NAD 27, Phenology: Flowers. Origin: Native. uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler In Fraxinus latifolia - Quercus garryana woods. With Symphoricarpos Sarah Gage G&R 253 15 Jun 1992 albus. Phenology: Fruits. Origin: Native. with Sharon Rodman Eliza Habegger 441 11 Jun 1998 WTU-344660 WTU-338755 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, LEWIS COUNTY: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Walupt Lake, Goat Rocks Primitive Area. Mouth of the Snohomish River. Elev. 4000 ft. 47.901249°, -122.088885°; WGS 84, uncertainty: 10000 m., 46.4175°, -121.46139°; WGS 84, uncertainty: 10000 m., Source: Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. On subalpine slopes around lake. Phenology: Fruits. Origin: Native. George Neville Jones 4310 19 Apr 1933 J. William Thompson 15169 4 Aug 1940 WTU-23907 WTU-22290JWT Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SKAMANIA COUNTY: U.S.A., WASHINGTON, MASON COUNTY: Cold Springs Indian Camp, 18 miles north of Trout Lake. Simpson Lake. Elev. 3493 ft. 47.13348°, -123.33929°; WGS 84, uncertainty: 1000 m., Source: 46.12694°, -121.73056°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff In marshy soil at edge of lake. Shrub 4-6 ft. tall. Phenology: Flowers. Canadian zone. Phenology: Flowers. Origin: Native. Origin: Native. E. Hunn 674 2 Aug 1977 Phillips B. Freer 179 27 May 1950 WTU-271213 WTU-135154 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, OKANOGAN COUNTY: 1/2 mile north of Friday Harbor, San Juan Island. Windy Creek trail No. 362, 2 mi. below Windy Peak. 48.548056°, -123.022778°; WGS 84, uncertainty: 800 m., Source: Elev. 6300 ft. Georeferenced, Georef'd by WTU Staff T39N R23E S6 SENE; WGS 84, uncertainty: 1000 m., Source: Fairly common in open fields. Occur singly. Shrub 7-10 feet high. GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Subalpine forest; assoc. Caltha biflora, Kalmia microphylla, Ledum glandulosum, Pinus contorta. Aspect 190 deg., slope 2 deg. Barbara D. Blanchard 42 5 May 1936 Phenology: Flowers. Origin: Native. WTU-45450 George Wooten GW01370 14 Jun 1992 WTU-332873 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, SKAGIT COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Tide Point, Cypress Island (George Fahey homestead). 48.57417°, -122.7075°; WGS 84, uncertainty: 10000 m., Source: U.S.A., WASHINGTON, PACIFIC COUNTY: GeoLocate, Georef'd by WTU Staff Ilwaco. Growing just back of beach. Phenology: Flowers. Origin: Native. 46.30917°, -124.04194°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff A. R. Kruckeberg 5994 14 May 1967 Sphagnum bog. Phenology: Flowers & Fruits. Origin: Native. WTU-320686 L. R. Abrams 11281 9 Jul 1926 WTU-19318JWT Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, SPOKANE COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Summit of Mt. Carleton (Mt. Spokane). 47.65889°, -117.425°; WGS 84, uncertainty: 1000 m., Source: U.S.A., WASHINGTON, PEND OREILLE COUNTY: GeoLocate, Georef'd by WTU Staff Ravine in mountains 4 miles east of Ione. Growing by brook near summit. Phenology: Flowers. Origin: Native. 48.741077°, -117.328331°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff F. O. Kreager 256 19 Jul 1902 Open woods. Shrub 1.5 meters high. Phenology: Flowers. Origin: Native. WTU-68401 H. T. Rogers 545 27 May 1940 WTU-83927 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., WASHINGTON, OKANOGAN COUNTY: Nooksack Mountain. Ptarmigan Creek, Hidden Lakes. 48.92778°, -122.32028°; WGS 84, uncertainty: 10000 m., Source: Elev. 3800 ft. GeoLocate, Georef'd by WTU Staff T39N R19E; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Along a stream. Phenology: Flowers. Origin: Native. Georef'd by WTU Staff Picea-Abies woods. In moist humus. Phenology: Flowers. Origin: W. C. Muenscher 9703 26 Apr 1939 Native. WTU-56047 O. T. Edwards 255 31 May 1936 WTU-3082 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, YAKIMA COUNTY: Lonicera utahensis S. Watson (YIN) At Howard Lake, 55 miles west of Toppenish ACF. Elev. 4887-5248 ft. U.S.A., WASHINGTON, PEND OREILLE COUNTY: 46.371695°, -121.461726°; WGS 84, uncertainty: 500 m., Source: Pass Creek. GeoLocate, Georef'd by WTU Staff 48.836016°, -117.22173°; WGS 84, uncertainty: 10000 m., Hudsonian zone. In forest clearings. Phenology: Flowers. Origin: Source: GeoLocate, Georef'd by WTU Staff Native. In woods along creek. Phenology: Flowers. Origin: Native. E. Hunn 535 30 Jun 1977 Harold St. John 6424 19 Jun 1924 WTU-271269 WTU-112967

Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., WASHINGTON, CLALLAM COUNTY: U.S.A., WASHINGTON, SPOKANE COUNTY: Hurricane Ridge; Olympic Mountains. Mt. Spokane. Elev. 4500 ft. Elev. 3500-4000 ft. 47° 58' 14" N, 123° 29' 29" W; NAD 27 47.65889°, -117.425°; WGS 84, uncertainty: 1000 m., Source: On alpine slopes. Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff In woods. Phenology: Flowers. Origin: Native. J. William Thompson 10606 9 Jun 1934 Sister Mary Milburge 482 20 Jun 1932 WTU-4740JWT WTU-19368 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., WASHINGTON, GARFIELD COUNTY: Umatilla National Forest, intersection of Forest Road 40 and Forest U.S.A., WASHINGTON, WHATCOM COUNTY: Road 4022. Misery Springs. Crest of Skyline Ridge, Mt. Baker. Elev. 6199 ft. Elev. 6000-6500 ft. 46° 7' 25" N, 117° 28' 55" W; T8N R42E S36 SW 1/2 48.83065°, -121.85005°; WGS 84, uncertainty: 750 m., Source: Bush 2-3' high; flowers very pale yellow, twinned with short blunt Nat. Geog. TOPO!, Georef'd by John Bassett spur; fragrant - like mock orange. On woodland borders, not in open First record for the region. Phenology: Flowers. Origin: Native. areas or in dark woods. Phenology: Flowers. Origin: Native. J. William Thompson 10971 11 Jul 1934 Doug Williams 4 27 Jun 1997 with Robert Goff WTU-6086JWT WTU-347085 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., WASHINGTON, KING COUNTY: Kings Lake, 1 mile west of Boyle Lake, northeast of Snoqualmie U.S.A., WASHINGTON, JEFFERSON COUNTY: Falls. Trail to Constance Ridge. Elev. 951 ft. 47.77561°, -123.12014°; WGS 84, uncertainty: 1300 m., Source: T24N R8E; WGS 84, uncertainty: 200 m., Source: Digital Map, GeoLocate, Georef'd by WTU Staff Georef'd by Ibro Polovina On dry rocky mountain side. Phenology: Vegetative. Origin: Native. Shrub 2 meters high; in bud. At lake edge by bridge with rushes and Rhynchospora alba. Phenology: Flowers. Origin: Native. George Neville Jones 5858 8 Aug 1934 G. Lebednik 398 23 Apr 1974 WTU-30350 WTU-260777 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, FERRY COUNTY: U.S.A., WASHINGTON, KING COUNTY: Colville National Forest: Vicinity of Swan Lake. Cascade Mountains of Western Washington. Cedar River Elev. 3600 ft. Watershed. Little Mountain Bog. 48° 31' 4" N, 118° 50' 16" W; T35N R32E S20; WGS 84, Elev. 1600 ft. uncertainty: 532 m., Source: Digital Map, Georef'd by Richard T22N R9E S27; NAD 27, uncertainty: 500 m., Source: Olmstead Georeferenced, Georef'd by WTU Staff Open mixed conifer forest - Douglas fir, Ponderosa pine, Larch. Marshy and boggy area (actually a fen) surrounded by forested bog. Phenology: Flowers. Origin: Native. The fen is a mosaic of sedge marsh, sphagnum moss, and open water. Open bog area. Phenology: Flowers. Origin: Native. Richard G. Olmstead 99-107 8 Jul 1999 Clayton J. Antieau 0125 23 Jun 2001 WTU-349300 with Nancy Job, Sandra Whiting, Jeffery Walker Caprifoliaceae WTU-350896 Lonicera utahensis S. Watson Caprifoliaceae U.S.A., WASHINGTON, FERRY COUNTY: Colville National Forest: Fire Mt./17 Mile Mt. saddle. Lonicera involucrata (Richardson) Banks ex Spreng. Elev. 5430 ft. T35N R34E S29; WGS 84, Source: Digital Map, Georef'd by U.S.A., WASHINGTON, KING COUNTY: Richard Olmstead Halmar Gate Wetland; Cascade Mountains of Western Washington. Mixed conifer woodland; Larix, Abies lasiocarpa, Pseudotsuga, Elev. 1400 ft. Calamogrostis rubescens. Pale yellow/green flowers. Small shrub. T22N R7E S1; NAD 27, uncertainty: 500 m., Source: Phenology: Flowers. Origin: Native. Georeferenced, Georef'd by WTU Staff Ditches filled with water, drier hummocks with Spiraea sp., Juncus Richard G. Olmstead 99-146 10 Jul 1999 effusus, Juncus supinus. Other plants in vicinity: Salix sp., Lysichitum americanum. Gaultheria shallon growing from stumps; WTU-349233 Menziesia ferruginea common. Upland island with doug fir, western hemlock, red alder. Growing on log on hummock neat open water. Phenology: Flowers. Origin: Native. Caprifoliaceae Suzanne Bagshaw 01-31 23 Jun 2001 Lonicera conjugialis Kellogg with Roz Glasser, Mary Clare Schroeder, Neil Dijrani U.S.A., WASHINGTON, SKAMANIA COUNTY: WTU-351122 Gifford Pinchot National Forest: Trailhead at end of spur 040 off Forest Road 8810. Elev. 4899 ft. Caprifoliaceae 46° 5.571' N, 121° 38.986' W; T7N R9E S15 SE 1/4 Solitary shrub by trail below summit. Flowers maroon. Top petal Lonicera involucrata (Richardson) Banks ex Spreng. folds back; bark of thin, linear peels. Shrub to 3 feet, heavily stunted by wind. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Halmar Gates wetland. Alison Colwell 01-18 15 Jun 2001 Elev. 1200 ft. T22N R7E S1 NW 1/4; NAD 27, uncertainty: 500 m., Source: WTU-350097 Georeferenced, Georef'd by WTU Staff Wetlands along Kerriston road and road 33; near tributaries to Williams creek. Other plants in vicinity: Salix sitchensis, Geum Caprifoliaceae macrophyllum, bracken fern, Juncus effusus, evergreen blackberry, Gaultheria sp., Menziesia ferruginea, Typha sp., Eleocharis sp., Lonicera ×bella Zabel Cornus canadensis, Carex sp., Spiraea sp. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, KING COUNTY: South shore of Portage Bay; Montlake, Seattle. Alan C. Yen 01-79 26 May 2001 Elev. 16 ft. T25N R4E S20; NAD 27, uncertainty: 805 m., Source: TRS2LL, WTU-351198 Georef'd by Ben Legler Thickets near shore. Pink flowers, garden relic; 1 shrub. Phenology: Flowers. Origin: Introduced. Caprifoliaceae P. F. Zika 15890 17 Apr 2001 Lonicera involucrata (Richardson) Banks ex Spreng. WTU-350425 U.S.A., WASHINGTON, KING COUNTY: Cascade Mountains of Western Washington: Walsh Lake, south side of lake. Elev. 650 ft. T22N R7E S9 SW 1/4; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler South side of the lake. North-facing second-growth with Thuja plicata, Tsuga heterophylla, Douglas-fir, Polystichum munitum. Lake margin is in shade. Pulustrine; 3 meters tall. Phenology: Fruits. Origin: Native. Eric Beach 01-90 23 Jun 2001 with Alison Colwell, Dick Schaetrel, Tom Lane, Jennifer Thomas WTU-351507 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, FERRY COUNTY: Cascade Mountains of Western Washington: Walsh Lake/Webster Moist forest along trail on north and northwest side of Swan Lake. Creek - at outlet of creek from lake. Elev. 3740 ft. Elev. 650 ft. T35N R31E S20, 29; NAD 27, uncertainty: 500 m., Source: T22N R7E S9 NW 1/4; NAD 27, uncertainty: 200 m., Source: Georeferenced, Georef'd by WTU Staff Georeferenced, Georef'd by WTU Staff Forested wetland adjacent to Swan Lake, 70% canopy closure by At outlet of creek from the lake. The delta is an open site with Pseudotsuga menziesii; associated species included Cornus several channels separating a few areas of higher ground adjacent stolonifera, Aconitum columbianum, Platanthera dilatatum, and to the lake. Collected fom an area along bank west of main channel. Viburnum edule. Phenology: Fruits. Origin: Native. Typha latifolia along lake margin, higher areas dominated by mixed shrubs and herbaceous plants. Yellow flowers; mostly in fruit. F. Weinmann 158 11 Jul 1999 Phenology: Flowers & Fruits. Origin: Native. with Ann Weinmann David M. Chapin 01-910 26 May 2001 WTU-351795 with Suzanne Bagshaw, Marin Bjork WTU-351429 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, CHELAN COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. North Cascades National Park; Near junction of Horseshoe Basin Trail and Stehekin Trail. U.S.A., WASHINGTON, SKAMANIA COUNTY: Elev. 3608 ft. Gifford Pinchot National Forest: Hwy. 141, 3.3 mi. W of Trout Lake 48° 27' 57" N, 121° 1' 36" W; UTM Zone 10, 646657.705E, Ranger Station. 5367998.843N; Source: Field notes, UTM calculated from Lat-Lon. Elev. 2010 ft. Steep, open rockslide area with Penstemon rupicola, Abies 45.96227°, -121.6157°; WGS 84, uncertainty: 1000 m., Source: lasiocarpa (small), Artemisia douglasii, Acer circinatum, Epilobium GeoLocate, Georef'd by WTU Staff angustifolium, Sorbus scopulina, Pachistima mrysinites, Achillea Orange/red flowers. Climbing a Ponderosa pine. Phenology: millefolium, Rubus parviflora, Taxus brevifolia, and Cryptogramma Flowers. Origin: Native. crispa. Spreading shrub to 2 meters tall; flowers yellow, drying, and dehiscing; bracts red; fruits black. Phenology: Flowers & Fruits. David Tank 01-20 16 Jun 2001 Origin: Native. WTU-351931 Alison Colwell 02-36 10 Aug 2002 WTU-353110, NPS accession 632, catalog 22262 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, SKAMANIA COUNTY: Gifford Pinchot National Forest: Dry meadow in South Prairie - U.S.A., WASHINGTON, WHATCOM COUNTY: southwest side of Forest Service Road 6615. Ross Lake National Recreation Area: Little Beaver Creek Trail near Elev. 2998 ft. small, unnamed stream. 45° 55.042' N, 121° 42.819' W Elev. 1886 ft. Seasonally inundated sedge meadow surrounded by Populus 48° 55' N, 121° 5.5' W; UTM Zone 10, 639814.017E, balsamifera, Tsuga heterophylla, and Pseudotsuga mertensiana. 5419728.395N; Source: Field notes, UTM calculated from Lat-Lon. Growing in full sun; heavily browsed; in open area near southwest Douglas-fir forest. Flowers bright orange. Phenology: Flowers. edge of meadow; maroon outer and yellow inner involucral bracts. Origin: Native. Phenology: Flowers. Origin: Native. Ben Legler 42 13 Jul 2002 Sharon Rodman 568 16 Jun 2001 WTU-353709, NPS accession 632, catalog 22869 WTU-352177 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., WASHINGTON, KING COUNTY: North Cascades National Park; Thunder Creek drainage; the first Cascade Mountains of Western Washington: Walsh Lake. North lake large rocky bald encountered on the trail to 4th of July Pass starting margin. at the junction with the thunder Creek Trail. A small way trail goes Elev. 650 ft. north onto the bald. T22N R7E S9 NW 1/4; NAD 27, uncertainty: 200 m., Source: Elev. 1950 ft. Georeferenced, Georef'd by WTU Staff 48° 40' 22.7599" N, 121° 3' 44.95907" W; UTM Zone 10, 3-meter tall shrubs. Phenology: Flowers & Fruits. Origin: Native. 642641E, 5392697N; Source: Calc. from UTM, UTM from field notes. Eric Beach 01-75 23 Jun 2001 Very dry, exposed rock outcrops covered with moss. Uncommon; with Alison Colwell, Dick Schaetrel, Tom Lane, Jennifer Thomas clambering up onto trees and shrubs like a vine; in partial sun; concentrated on the edges of the rocky bald. Phenology: Flowers. WTU-351708 Origin: Native. Steve Hahn 541 15 Jun 2002 WTU-354050, NPS accession 632, catalog 23051 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera glenhii Fr. Schmidt U.S.A., OREGON COUNTY: RUSSIAN FEDERATION, SAKHALIN REGION: Santiam Forest. Sakhalin Island; on west side of island along Hwy 495 4 km SSW of Elev. 4200 ft. Lesogorsk and just north of Tel'novskiy, at mouth of unnamed Wet meadow. Soil character: Black sandy loam. Associated plants: stream emptying into Tatary Straight. Willow, huckleberry. Density in tenths: .2. Slope: North. No. 20. Elev. 10 ft. Ingram. Phenology: Flowers. Origin: Native. Shrubby gully opening to beach, with Alnus, Salix, Petasites, Senecio, grasses. Immature fruit green; small shrub. Phenology: H.G. White 39074 14 Jul 1922 Fruits. Origin: Native. WTU-355262 Ben Legler 848 23 Jul 2003 WTU-357136 Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Caprifoliaceae U.S.A., WASHINGTON, ISLAND COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Whidbey Island, The Nature Conservancy's Ebey's Landing Preserve; along top of bluff above Perego's Lagoon. U.S.A., WASHINGTON, PACIFIC COUNTY: Elev. 200 ft. The Nature Conservancy's Ellsworth Creek Preserve. Entrance to 48° 12' 7.6" N, 122° 43' 54.1" W Yellow Gate Road. Open, grassy, SW-facing bluff backed by Pseudotsuga forest. Elev. 3 ft. Flowers light purple; shrub-like vines forming extensive patches. 46° 25.6' N, 123° 53.675' W Phenology: Flowers. Origin: Native. Estuary with gravel road through it; dominant species include Carex obnupta, Carex lyngbei, and Holcus lanatus. Phenology: Flowers & Ben Legler 1709 4 Jun 2004 Fruits. Origin: Native. WTU-355986 Marianne Spahr 03-22 9 Jul 2003 WTU-357515 Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Caprifoliaceae U.S.A., WASHINGTON, ISLAND COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Whidbey Island, Ebey's Landing, field edge. 48° 11.921' N, 122° 42.902' W U.S.A., WASHINGTON, PACIFIC COUNTY: Full sun; dry. Frequent; pink flowers. Phenology: Flowers. Origin: The Nature Conservancy's Ellsworth Creek Preserve at lower end of Native. creek in salt marsh. Elev. 0 ft. Peter W. Dunwiddie PWD 107 4 Jun 2004 46° 24' 59" N, 123° 53' 14" W with Jeff Walker, Sharon Rodman, Susan Zwinger Salt marsh with Juncus balticus, Triglochia maritima, Potentilla pacifica, and Carex lyngbyei. Flowers yellow; shrub on edge of WTU-356089 marsh. Phenology: Flowers & Fruits. Origin: Native. Ben Legler 501 31 May 2003 Caprifoliaceae WTU-357577 Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, ISLAND COUNTY: Caprifoliaceae Whidbey Island, Ebey's Landing Nature Conservancy Preserve; adjacent to bluff trail along top of bluff. Lonicera involucrata (Richardson) Banks ex Spreng. Elev. 190 ft. 48.195°, -122.7125°; WGS 84, Source: Georeferenced, Georef'd U.S.A., WASHINGTON, OKANOGAN COUNTY: by WTU Staff Okanogan National Forest. Roger's Lake. East trail and down to 3 meters tall; small opening in woods. Phenology: Vegetative. Origin: lake. Native. Elev. 5898 ft. 48° 39.03' N, 119° 58.016' W Jeff Walker 282 10 Sep 2004 Pine woods with little understory to grassy lake edge; dominant trees: Pinus contorta and Picea engelmanii. Growing in partial shade WTU-357077 under Pinus contorta; ripe fruits black. Phenology: Flowers & Fruits. Origin: Native. Sharon Rodman 932 26 Jul 2003 with Jessie Johanson, Joe Johanson Herbarium: WTU Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, COLUMBIA COUNTY: Okanogan National Forest. Rainy Lake Outlet. Ridgeline 0.4 miles southwest of Blakely Springs along Forest Elev. 5010 ft. Service Road 64, southwest of Bluewood Ski Area. UTM Zone 10, 667387E, 5374601N; NAD 27, uncertainty: 50 m., Elev. 6020 ft. Source: Georeferenced, Georef'd by WTU Staff 46° 3' 46" N, 117° 55' 10" W; NAD 83 Sunny, wet meadow by Rainy Lake stream outlet. Dominant trees: Mix of dry, rocky exposed areas and open Picea-Abies forest. Silver fir, Mountain hemlock, Subalpine fir, Engelman. Growing along Flowers white to cream; shrubs to 1 meter tall under conifers. lake margin. Phenology: Flowers & Fruits. Origin: Native. Phenology: Flowers. Origin: Native. Alan C. Yen 03-028 25 Jul 2003 Ben Legler 1983 11 Jul 2004 with Richard Robohm, Suzanne Bagshaw, Liz Van Volkenburgh, Therese Ohlson WTU-362183 Herbarium: WTU Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. U.S.A., WASHINGTON, OKANOGAN COUNTY: Castle Island. Okanogan National Forest. Tiffany Lake. Elev. 0-98 ft. Elev. 6500 ft. 48.42104° N, 122.82176° W; T34N R1W S21; WGS 84, 48° 41.521' N, 119° 56.46' W uncertainty: 100 m., Source: Digital map Lakeshore meadows; Picea englemannii, Pinus contorta, Abies Steep, cliffy island with many rock ledges, cracks and beaches. lasiocarpa. Phenology: Flowers. Origin: Native. Purple flowers, sprawling vine; local on north side; partial shade. Phenology: Flowers. Origin: Native. Alan C. Yen 03-154 26 Jul 2003 with Sharon Rodman, Jessie Johanson, Joe Johanson, Suzanne Ben Legler 2595 28 Apr 2005 Bagshaw, Richard Robohm, Ann Weinmann, Fred Weinmann WTU-383624 WTU-362572 Caprifoliaceae Caprifoliaceae Lonicera ×bella Zabel Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata U.S.A., WASHINGTON, WHITMAN COUNTY: By South Fork Palouse River, near Reaney Park and Lentil Lane, U.S.A., WASHINGTON, SKAGIT COUNTY: Pullman. 2.5 miles southwest of Conway, at Skagit Wildlife Area at end of Elev. 2328 ft. Mann Road, near boat ramp. 46° 43.8' N, 117° 10.3' W; T14N R45E S5 Elev. 5 ft. Pink flowers, adventive shrubs spread from cultivation, thickets. 48° 19' 28" N, 122° 22' 24" W; NAD 27 Phenology: Flowers & Fruits. Mix of muddy farm fields, dikes, and riparian Alnus rubra forest. Fruit deep red-black; shrub 3 meters tall on edge of Alnus woods. P. F. Zika 19719 30 May 2004 Phenology: Fruits. Origin: Native. with R. Old Ben Legler 1724 9 Jun 2004 WTU-359365 WTU-363476 Caprifoliaceae Caprifoliaceae Lonicera tatarica L. Lonicera involucrata (Richardson) Banks ex Spreng. var. U.S.A., WASHINGTON, WHITMAN COUNTY: involucrata Above bike trail, south of Route 270, Pullman. Elev. 2361 ft. U.S.A., WASHINGTON, KING COUNTY: T14N R45E S4; WGS 84, uncertainty: 2500 m., Source: Cedar River Watershed. Wetlands east of 200 and 210 Roads GeoLocate, Georef'd by WTU Staff junction (along 200.12 Road). Base of slope, north aspect; with Dipsacus, Rosa, Clematis. Flowers Elev. 2300 ft. whitish or pink, fruit red, uncommmon adventive shrubs 2.5 meters T22N R9E S29 SW, SW; NAD 27, uncertainty: 805 m., Source: tall, palouse remnant. Phenology: Fruits. Origin: Introduced. TRS2LL, Georef'd by Ben Legler Wetlands. Phenology: Flowers & Fruits. Origin: Native. P. F. Zika 19713 30 May 2004 with R. Old Carla Culver 4-06 13 Jul 2002 WTU-359366 WTU-364507 Caprifoliaceae Caprifoliaceae Lonicera xylosteum L. Lonicera cauriana Fernald U.S.A., WASHINGTON, KING COUNTY: U.S.A., OREGON, LAKE COUNTY: Near west shore of Lake Washington, Washington Park Arboretum. Fremont-Winema National Forest. North Fork Sprague River at Elev. 50 ft. Forest Service Road 3411 crossing near Sandhill Campground. 47° 38' N, 122° 18' W; T25N R4E S38 Elev. 6087 ft. Bank of small creek, partial shade, with Rubus spectabilis, 42° 35.619' N, 120° 52.626' W; NAD 27 Ranunculus repens, Scirpus microcarpus. Adventive shrub 1 meter Damp grassy bench along river. Other plants; Pinus contorta, tall, fruit dark red, shiny, some paired, depressed-globose, individual Artemesia arbuscula, Salix exigua, Juncus sp. Green fruit. berries 7-8 x 8-12 millimeters wide; 1 - 3 seeds per fruit. Phenology: Uncommon. Phenology: Fruits. Origin: Native. Fruits. Don Knoke 498 1 Jul 2004 Peter F. Zika 19883 8 Jul 2004 WTU-365956 WTU-359965 Caprifoliaceae Caprifoliaceae Lonicera conjugialis Kellogg Lonicera ×bella Zabel U.S.A., WASHINGTON, YAKIMA COUNTY: U.S.A., WASHINGTON, SPOKANE COUNTY: Big Klickitat River. South shore of Little Spokane River, north of Dartford Drive, west of 46.044714°, -121.237172°; WGS 84, uncertainty: 10000 m., Route 395 bridge, Dartford. Source: GeoLocate, Georef'd by WTU Staff Elev. 1574 ft. Subalpine slopes, Cascade Mountains. Phenology: Fruits. Origin: 47° 47' N, 117° 24.7' W Native. Thickets with Spiraea douglasii, Poppulus balsamifera, floodplain. Bright red fruit; bracteoles < 1/2 ovary length; 2 meter adventive Louis F. Henderson s.n. 4 Aug 1892 shrubs. Phenology: Fruits. WTU-16624 Peter F. Zika 20134 31 Aug 2004 WTU-360923 Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Caprifoliaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: Lonicera morrowii A. Gray ? Arbutus Point; Orcas Island; near the town of East Sound. 48.690833°, -122.901389°; WGS 84, uncertainty: 200 m., Source: U.S.A., WASHINGTON, OKANOGAN COUNTY: Georeferenced, Georef'd by WTU Staff Washburn Island, Columbia River, 7 air kilometers east of Brewster. Not uncommon along the Sound. Dry ground. A remarkably large Elev. 787 ft. specimen, the stem near ground being 1 inch in diameter. 48° 5' N, 119° 41' W; T30N R25E S15 Phenology: Flowers. Origin: Native. Under Salix exigua, Cornus sericea, willow thickets, occasional escape from wildlife plantings along road. Red fruit, 1.5 meters Louis F. Henderson s.n. 3 Jul 1892 adventive shrubs. Phenology: Fruits. WTU-16620 Peter F. Zika 20331 22 Sep 2004 WTU-360954 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, THURSTON COUNTY: Lonicera periclymenum L. Near Olympia. 47.03806°, -122.89944°; WGS 84, uncertainty: 5000 m., Source: U.S.A., OREGON, CLATSOP COUNTY: GeoLocate, Georef'd by WTU Staff Route 202, 100 m south of Cadonau Road, 1 air kilometer southeast Common from tide waters to high mountains, along marshes and of Mouth of Wallooskee River. creeks. Phenology: Flowers & Fruits. Origin: Native. Elev. 16 ft. 46° 8.4' N, 123° 48.3' W; T8N R9W S33 Louis F. Henderson s.n. 24 May 1892 Partial shade, with Alnus rubra, Rhamnus purshianus, Tsuga, Vaccinium parvifolium, Gaultheria, Rubus spectabilis, near WTU-16494 freshwater marsh. Pink flowers rades yellow-brown, leaves glaucous below and not fused; occasional as an adventive climbing vine. Phenology: Flowers. Peter F. Zika 20042 25 Aug 2004 WTU-361142 Caprifoliaceae Caprifoliaceae Lonicera periclymenum L. Lonicera caerulea L. U.S.A., WASHINGTON, KING COUNTY: U.S.A., OREGON, LINN COUNTY: Foster Island, Lake Washington, north end of Cascade Range, Willamette National Forest, Bruno Mire, along arboretum, Washington Park, Seattle. USFS Road 2234 about 5 air miles southeast of Idanha. Elev. 16 ft. Elev. 1224 ft. 47° 38' N, 122° 18' W; T25N R4E S21 T11S R07E S6; NAD 27, uncertainty: 805 m., Source: TRS2LL, With Alnus rubra, Populus balsamifera. Georef'd by Ben Legler Adventive evergreen vine climbing 3 meters in In mire. Flowers yellow-green, shrub to 0.46 meters tall, associated forest, flowers white, 5.5 cm; upper lip 4-lobed, genera: Kalmia, Vaccinium, Caltha, Dodecatheon, Salix, Picea, 24 mm; lower lip narrow, 29 mm; tube 27 mm; 5 Lysichiton. Phenology: Flowers. ; filaments white, anthers yellow; stigma green. Reference specimen for illustration in Richard R. Halse 5785 14 Jun 2000 Revision of Flora of the Pacific Northwest, WTU Herbarium, 2016. Phenology: Flowers. WTU-347911 Peter F. Zika 19683 Caprifoliaceae 24 May 2004 Lonicera cauriana Fernald WTU-361879, WTU-V-031920 U.S.A., OREGON, DOUGLAS COUNTY: Between Union Creek and Diamond Lake. Meadows in lodge-pole pine forest. Phenology: Flowers. Caprifoliaceae A. R. Kruckeberg 2030 1 Jul 1950 Lonicera maackii (Rupr.) Herder WTU-181069 U.S.A., OREGON, LANE COUNTY: Irving Road near Shirley Street. Eugene. Elev. 393 ft. Caprifoliaceae 44° 6.2' N, 123° 8.4' W Dense blackberry thicket, partial shade, with Rubus armeniacus, Lonicera caerulea L. Symphoricarpos albus. 2 meters tall adventive shrub, fruits bright red, globose, solitary .5-7.0 x 4.7-8.2 mm wide; 1-5 seeded U.S.A., MONTANA, RAVALLI COUNTY: berry. Phenology: Flowers & Fruits. Bitterroot Mountains, along Lost Horse Creek Road. Elev. 5800 ft. P. F. Zika 20407 7 Oct 2004 46.113216°, -114.346589°; uncertainty: 14000 m. with E. R. Alverson Below powerty slide, on wet meadow. Phenology: Fruits. WTU-362640 Klaus H. Lackschewitz 1208 29 Jun 1969 WTU-263903 Caprifoliaceae Lonicera caerulea L. Caprifoliaceae U.S.A., MONTANA, GRANITE COUNTY: Lonicera caerulea L. South of the Skalkaho Road (Highway 38), above the Richt-meyer Ranch and just below the Sapphire Mine. U.S.A., MONTANA, DEER LODGE COUNTY: In a willow thicket on seep. Phenology: Flowers. Anacondda-Pintlar Range, La Marche Lake Meadows. Elev. 8250 ft. Klaus H. Lackschewitz 7189 29 May 1977 On Sphagnum hummocks, inside the cover of Vaccinium Occidentale. And never taller than the former species, up to 12 WTU-288604 inches high, sometimes longer, but then growing prostrate. Frequently found here. Phenology: Flowers. Caprifoliaceae Klaus H. Lackschewitz 3874 17 Jul 1972 Lonicera ×bella Zabel WTU-262744 U.S.A., WASHINGTON, SNOHOMISH COUNTY: King (205th and Alaska Way)/Snohomish Counties. Undeveloped Caprifoliaceae property beginning on north border of LFP. 47.91306°, -122.09694°; WGS 84, uncertainty: 10000 m., Source: Lonicera caerulea L. GeoLocate, Georef'd by WTU Staff Growing in disturbed margin along 205th with other invasives. U.S.A., MONTANA, RAVALLI COUNTY: Phenology: Flowers & Fruits. Bitterroot Mountains, along Bass Creek, just below the big south-slope draw ("Avalanche Run") on St. Joseph. Bass Creek Christie S. Isler 31-5-98 3 May 1998 Trail. Elev. 6200 ft. WTU-335290 With Berula glandulosa. Phenology: Fertile. Klaus H. Lackschewitz 2960 24 Jul 1971 WTU-262757 Caprifoliaceae Caprifoliaceae Lonicera caerulea L. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, IDAHO COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Along Packer Meadows road, 3/4 miles east of Lolo Pass. Tronson Ridge; Wenatchee Mountains. Elev. 5700 ft. Elev. 2000 ft. 46.63778°, -114.55222°; WGS 84, uncertainty: 10000 m., Source: 47.37864°, -120.56581°; WGS84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff Georeferenced, Georef'd by WTU Staff In bog. Phenology: Fruits. Lower wooded slopes. Phenology: Fruits. Origin: Native. B. O. Mulligan s.n. 11 Jul 1974 J. William Thompson 9294 4 Jul 1933 WTU-259658 WTU-19364JWT

Caprifoliaceae Caprifoliaceae Lonicera villosa (Michx.) Schult. var. villosa Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., IDAHO, VALLEY COUNTY: U.S.A., WASHINGTON, KING COUNTY: Along Big Creek at Green Tables Forest Camp, Payette National Alki Point. Forest. 47.57639°, -122.40861°; WGS 84, uncertainty: 10000 m., Source: Elev. 6000 ft. GeoLocate, Georef'd by WTU Staff 44.583659°, -115.881559°; WGS 84, uncertainty: 4800 m., Phenology: Flowers & Fruits. Origin: Native. Source: GeoLocate, Georef'd by WTU Staff Phenology: Flowers & Fruits. G. L. Jenks s.n. 6 Jun 1908 J. William Thompson 13880 1/2 16 Jul 1937 WTU-16610 WTU-10621JWT Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera villosa (Michx.) Schult. var. villosa U.S.A., WASHINGTON, KING COUNTY: West Seattle. U.S.A., OREGON COUNTY: 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: Bank of Big Springs Creek, along Fort Klamath-Bend road. GeoLocate, Georef'd by WTU Staff Phenology: Fruits. Phenology: Fruits. Origin: Native. Morton E. Peck 9574 19 Jul 1920 W. V. Lovitt s.n. 19 Nov 1909 WTU-12782JWT WTU-16611

Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, KING COUNTY: U.S.A., MONTANA, FLATHEAD COUNTY: Alki Point. One mile south of Columbia Falls. 47.57639°, -122.40861°; WGS 84, uncertainty: 10000 m., Source: Open woods. Twining vine. Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff Phenology: Flowers & Fruits. Origin: Native. H. T. Rogers 966 21 Jun 1942 with J. M. Rogers G. L. Jenks 697 6 Jun 1908 WTU-241178 WTU-272465 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera cauriana Fernald U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., OREGON, DOUGLAS COUNTY: Mount Ingalls; Mount Stuart District. Diamond Lake. Elev. 3936 ft. Damp ground. Phenology: Fruits. 47.47263°, -120.94637°; WGS84, uncertainty: 2000 m., Source: Georeferenced, Georef'd by WTU Staff Morton E. Peck 19245 8 Jul 1936 Along stream. Zone = transition. Range = Alaska to Canada southward to California and Colorado. Common. Black Twinberry. WTU-8444JWT Phenology: Flowers & Fruits. Origin: Native. W. J. Eyerdam 1082 4 Jul 1937 WTU-292944 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., IDAHO COUNTY: U.S.A., OREGON COUNTY: Palouse Country and about Lake Coeur d' Alene. Along the Umatilla River and tributaries, western Blue Mountains, -116.747304°, 0°; uncertainty: 24000 m. eastern Oregon. Phenology: Flowers. Origin: Native. Elev. 1640-3936 ft. Extending to near the summit. New to Eastern Oregon. Phenology: E. A. Shumway s.n. Jun 1892 Flowers. Origin: Native. WTU-188498 Wm. C. Cusick 3247 12 Jun 1908 WTU-139247 Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Caprifoliaceae U.S.A., OREGON COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Along Cow Creek. Rocky places. Phenology: Flowers & Fruits. Origin: Native. U.S.A., WASHINGTON, SAN JUAN COUNTY: Kanaka Bay, San Juan Island. Thomas Howell s.n. 30 Jun 1887 48.485556°, -123.085°; WGS 84, uncertainty: 500 m., Source: Georeferenced, Georef'd by WTU Staff WTU-19321JWT Phenology: Fruits. Origin: Native. A. S. Foster s.n. Jul 1904 Caprifoliaceae WTU-16481 Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., OREGON COUNTY: Caprifoliaceae Granite Pass. Phenology: Flowers & Fruits. Lonicera ciliosa (Pursh) Poir. ex DC. C. V. Piper 6078 17 Jun 1904 U.S.A., OREGON, WHEELER COUNTY: Near tributary to Service Creek, about 13 miles southeast of Fossil. WTU-19393JWT Elev. 3000 ft. T8S R22E S3; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Caprifoliaceae Climbing vine in shrubbery. Flowers bright orange. Phenology: Flowers. Origin: Native. Lonicera involucrata (Richardson) Banks ex Spreng. Arthur Cronquist 6650 22 Jun 1950 U.S.A., WASHINGTON, CHELAN COUNTY: Wenatchee Forest. WTU-144266 Elev. 4800 ft. T26N R16E S20 SE 1/4; WGS84, uncertainty: 32000 m., Source: Georeferenced, Georef'd by WTU Staff Caprifoliaceae Slope: 0. Deep moist humus soil. Meadow. Density: .1 Willow-Ledum-spiree. Limited distribution. Scattering abundance. Lonicera involucrata (Richardson) Banks ex Spreng. Flowering period: July. Forage value: Browsed by sheep with other shrubs. Phenology: Flowers & Fruits. Origin: Native. U.S.A., WASHINGTON, YAKIMA COUNTY: Upper Wenas Creek. Frank B. Lenzie 451 12 Jul 1918 46.869342°, -120.775868°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-322659 Wet soil. Phenology: Flowers & Fruits. Origin: Native. F. A. Warren 2032 28 May 1933 Caprifoliaceae WTU-213475 Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., OREGON, LANE COUNTY: Caprifoliaceae Cottage Grove. Forests. Abnormal - procumbent; white flowers. Phenology: Flowers. Lonicera ciliosa (Pursh) Poir. ex DC. Origin: Native. U.S.A., IDAHO, IDAHO COUNTY: Wm. C. Cusick 4115 13 Jun 1914 Lowell-Kooskia Road, 3 miles west of Lowell. T32N R6E S12; WGS 84, uncertainty: 1000 m., Source: WTU-139250 GeoLocate, Georef'd by WTU Staff In old burn area on sandy-loam. Flowers orange. Phenology: Flowers. Origin: Native. Quentin Jones 50 17 Jun 1950 WTU-141848 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, WHATCOM COUNTY: U.S.A., OREGON, DOUGLAS COUNTY: Bellingham; about 1 mile west of town. [No locality given on label] 48.75972°, -122.48694°; WGS 84, uncertainty: 10000 m., Source: Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff In woods. Bush . Phenology: Flowers & Fruits. Origin: F. D. Bailey s.n. Jun 1934 Native. WTU-73343 T. C. Frye s.n. 10 Aug 1909 WTU-16484 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. Near Lake Quinault. 47.46722°, -123.84417°; WGS 84, uncertainty: 10000 m., Source: CANADA, BRITISH COLUMBIA: GeoLocate, Georef'd by WTU Staff Slocan Lake. Open woods. Phenology: Flowers. Origin: Native. Hillside in aspenbirch association. Phenology: Flowers. Origin: Native. J. William Thompson 6262 9 May 1931 James Hatter 33 28 Jun 1949 WTU-19374JWT WTU-143026 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., IDAHO, LATAH COUNTY: U.S.A., MONTANA, LINCOLN COUNTY: Along Elk River, 1 mile east of Elk River. Above Kootenai River about 60 miles west of Kalispell; near 46.783274°, -116.160368°; WGS 84, uncertainty: 1600 m., Highway 2. Source: Digital Map, Georef'd by Joseph Stormer, Invalid Granitic cliffs. Phenology: Flowers. Origin: Native. coordinates (located outside county bounds). Thickets. Phenology: Flowers. Origin: Native. C. L. Hitchcock 17678 25 Jun 1948 William H. Baker 15597 28 Jun 1959 WTU-125452 WTU-297784 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., IDAHO, IDAHO COUNTY: Along Meadow Creek above junction of Meadow Creek and Selway U.S.A., OREGON, JOSEPHINE COUNTY: River. Siskiyou National Forest, 20-23 miles west and north of Selma on 45.497311°, -115.937845°; WGS 84, uncertainty: 10000 m., Illinois River Road (3504) or 8-11 miles north of intersection of 3504 Source: GeoLocate, Georef'd by WTU Staff and 374 on 3504. Common. Phenology: Flowers. Origin: Native. Elev. 1180 ft. 42.35201°, -123.779622°; WGS 84, uncertainty: 4000 m., Source: J. L. McMullen 1550 3 Jun 1954 Digital Map, Georef'd by Joseph Stormer Lowland deciduous woods mostly of Quercus, also Holodiscus, WTU-297788 Pseudotsuga, Calocedrus. Only one plant seen; vine; flowers with pinkish corolla lobes and white tubes, but tubes pinkish abaxially toward base. Phenology: Flowers & Fruits. Origin: Native. Caprifoliaceae Melinda F. Denton 2476 21 Jun 1973 Lonicera involucrata (Richardson) Banks ex Spreng. var. ledebourii (Eschsch.) Jeps. WTU-259228 U.S.A., WASHINGTON, CLALLAM COUNTY: Lake Ozette. Caprifoliaceae Elev. 50 ft. 48.155°, -124.66583°; WGS 84, uncertainty: 10000 m., Source: Lonicera involucrata (Richardson) Banks ex Spreng. GeoLocate, Georef'd by WTU Staff Phenology: Flowers. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Ocean City. Ruby L. C. Sovereign s.n. 26 Apr 1941 47.07111°, -124.16472°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-316087 Swampy woods. Phenology: Flowers & Fruits. Origin: Native. G. N. Jones 3929 27 Jun 1933 WTU-37174 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., IDAHO, LATAH COUNTY: U.S.A., OREGON, CURRY COUNTY: Palouse Range. Crumarine Creek Drainage at base Moscow Along the Rogue River, 2 miles south of Illahe. Mountain. T34S R11W S30; NAD 27, uncertainty: 805 m., Source: TRS2LL, 46.760766°, -116.908462°; WGS 84, uncertainty: 10000 m., Georef'd by Ben Legler Source: GeoLocate, Georef'd by WTU Staff Thickets. Phenology: Flowers & Fruits. Origin: Native. Phenology: Flowers. Origin: Native. William H. Baker 11525 13 Jun 1954 Ben W. Chichester 1093 17 Jun 1954 WTU-297990 WTU-299455 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, BONNER COUNTY: U.S.A., WASHINGTON, YAKIMA COUNTY: Along Trestle Creek, 2 miles west of Hope, Lake Pend Orielle. Snoqualmie Forest, Tieton R.S. 48.279605°, -116.347341°; WGS 84, uncertainty: 500 m., Source: Elev. 2700 ft. GeoLocate, Georef'd by WTU Staff T14N R14E S21; WGS 84, uncertainty: 10000 m., Source: Hillside, open woods. Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff Deep sandy loam; timber, browse. Principal associated plants in William H. Baker 14938 22 Jun 1957 order of abundance: Afo. Bsn. Phenology: Fruits. Origin: Native. WTU-296245 John A. Reggem 330 15 Jun 1937 WTU-324050 Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Caprifoliaceae U.S.A., OREGON, CURRY COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. Mouth of Pistol River. 42.282898°, -124.404958°; WGS 84, uncertainty: 1835 m., U.S.A., IDAHO, LATAH COUNTY: Source: Digital Map, Georef'd by Joseph Stormer South slope of West Twin, Moscow Mountains, five miles northeast Wooded slopes near ocean. Phenology: Flowers & Fruits. Origin: of Moscow. Native. 46.7325°, -116.99917°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. William Thompson 12831 21 Jun 1936 Phenology: Flowers. Origin: Native. WTU-2828JWT Ben Chichester 319 28 Jun 1950 WTU-302157 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., IDAHO, BENEWAH COUNTY: Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray var. Junction of Skyline Drive and Highway 95. hispidula T43N R4W S20; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler U.S.A., OREGON, CURRY COUNTY: Woods. Phenology: Flowers. Origin: Native. Dry bank above Illinois River, 2 miles south of Agness. Dry bank above river. Clambering over Rhus diversiloba; corollas William H. Baker 15855 9 Jun 1960 deep pink. Phenology: Flowers & Fruits. Origin: Native. WTU-296248 C. L. Hitchcock 19920 1 Jul 1953 WTU-151214 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. Ocean City. 47.07111°, -124.16472°; WGS 84, uncertainty: 10000 m., Source: U.S.A., IDAHO, LATAH COUNTY: GeoLocate, Georef'd by WTU Staff East Twin, Moscow Mountains, five miles northeast of Moscow. Meadow along seabeach. Phenology: Flowers. Origin: Native. 46.81194°, -116.90417°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff G. N. Jones s.n. 23 Jun 1933 South slope, open woods. Phenology: Flowers. Origin: Native. WTU-37173 Ben Chichester 266 30 Jun 1950 WTU-302156 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., OREGON, JOSEPHINE COUNTY: Big Boulder Creek trail, above Cle Elum River. Oregon Caves National Monument and vicinity. 47.44242°, -121.057772°; WGS 84, uncertainty: 10000 m., 42.0833°, -123.4°; WGS 84, uncertainty: 5000 m., Source: Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by Joseph Stormer On serpentine. Phenology: Flowers & Fruits. Origin: Native. Cliff 100 feet above the opening of the caves. Phenology: Flowers. Origin: Native. A. R. Kruckeberg 5774 15 Jun 1964 William H. Baker 770 25 Jun 1950 WTU-313383 with George C. Ruhle WTU-298305 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., IDAHO, BONNER COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. Along Priest River, 3 miles south of outlet to Priest Lake. 48.446432°, -116.905153°; WGS 84, uncertainty: 5000 m., U.S.A., OREGON, JOSEPHINE COUNTY: Source: GeoLocate, Georef'd by WTU Staff Oregon Caves National Monument and vicinity. Woods. Phenology: Flowers. Origin: Native. 42.0833°, -123.4°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by Joseph Stormer William H. Baker 15632 5 Jul 1959 On a cliff 100 feet above cave opening. Phenology: Flowers. Origin: Native. WTU-296055 William H. Baker 756 18 Jun 1949 with George C. Ruhle Caprifoliaceae WTU-298306 Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., OREGON, JACKSON COUNTY: Caprifoliaceae East of Brownsboro. Rocky hillsides. Phenology: Vegetative. Origin: Native. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray var. hispidula J. William Thompson 2143 8 Apr 1927 U.S.A., OREGON, JOSEPHINE COUNTY: WTU-19372JWT Illinois River Valley on 4-wheel drive road off 8 Dollar Mountain Road, downstream from Illinois River bridge 2.1 miles. 42° 16' 2" N, 123° 41' 15" W; T38S R8W S19 Caprifoliaceae Roadside under Douglas Fir. Pink-flowered vine. Phenology: Flowers & Fruits. Origin: Native. Lonicera ciliosa (Pursh) Poir. ex DC. Hillary S. Hayden 108 26 Jun 1998 U.S.A., IDAHO, IDAHO COUNTY: Along Middle Fork of Clearwater River, 2 miles west of Lowell. WTU-342644 46.144162°, -115.636514°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff Edges of woods. Phenology: Flowers. Origin: Native. Caprifoliaceae William H. Baker 14813 30 May 1957 Lonicera ciliosa (Pursh) Poir. ex DC. WTU-296057 U.S.A., WASHINGTON, KING COUNTY: Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Caprifoliaceae Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray E. S. Meany 119 Jun U.S.A., OREGON, DOUGLAS COUNTY: North Umpqua River at Winchester. WTU-16468 Bank of river. Phenology: Flowers & Fruits. Origin: Native. W. C. Cusick 4350 21 Jul 1914 Caprifoliaceae WTU-149315 Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., IDAHO, IDAHO COUNTY: Above Kamiah. 46.22722°, -116.02833°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff, Invalid coordinates (located outside county bounds). In coniferous forest, grade, Canadian-Arid Transition Zones. Twining liane, corolla orange, old burn. Phenology: Flowers. Origin: Native. L. Constance 1690 31 May 1936 with R. C. Rollins WTU-9477JWT Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., OREGON, CLACKAMAS COUNTY: U.S.A., OREGON, COLUMBIA COUNTY: Near Estacada. Saint Helens, Vernonia, 11 miles. Phenology: Flowers. Phenology: Flowers. Origin: Native. J. William Thompson 2606 11 Jun 1927 Lilla Leach 3164 29 May 1931 WTU-19376JWT WTU-255318

Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, LEWIS COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: Mission Creek. Eight-Mile Canyon, ca. [circa] 8 miles south of Leavenworth. 46.23988°, -116.68151°; WGS 84, uncertainty: 10000 m., Source: 47.480593°, -120.66028°; WGS84, uncertainty: 1000 m., Source: Nat. Geog. TOPO!, Georef'd by John Bassett GeoLocate, Georef'd by WTU Staff Woods on upper slope in ravine. Phenology: Flowers. Origin: Native. Along stream in yellow pine forest. Phenology: Flowers. Origin: Native. H. St. John 6347 30 May 1924 with J. A. Cary, P. Putnam, F. A. Warren C. L. Hitchcock 11409 26 May 1945 WTU-110719 WTU-100665

Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. CANADA, BRITISH COLUMBIA: U.S.A., MONTANA, LIBERTY COUNTY: About 4.5 miles north of Milnes Landing on east side of Sooke River Near Kootenai Falls. west of Victoria. Roadside thickets. Phenology: Flowers. Origin: Native. Trailing over shrubs in logged-over area by river. Very common, flowers orange-red. Phenology: Flowers. Origin: Native. W. B. Cooke 17327 6 Jun 1946 with V. G. Cooke J. A. Calder 29514 24 May 1961 with K. T. MacKay WTU-134848 WTU-215728 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray var. hispidula Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., OREGON, JOSEPHINE COUNTY: U.S.A., WASHINGTON, OKANOGAN COUNTY: Along Forest Service Road 4105; 2.8 miles from junction with Forest Near summit of Twisp Cut-off road. Service Road 4103. Siskiyou National Forest. 48.36361°, -120.12111°; WGS 84, uncertainty: 10000 m., Source: Elev. 2400 ft. GeoLocate, Georef'd by WTU Staff 42° 18' 55" N, 123° 42' 58" W; T37S R9W S35; T37S R9W S36 Phenology: Flowers. Origin: Native. Growing on rise to left of road. Purple buds, pinkish-white flowers. One stem with evidence of insect gall. Phenology: Flowers & Fruits. Chas. B. Fiker 912 26 Jun 1932 Origin: Native. WTU-19370JWT Roger N. Fuller 98-2 27 Jun 1998 WTU-349147 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, KITTITAS COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. Along Swauk Creek near Liberty, Wenatchee Mountains. Elev. 2000 ft. CANADA, BRITISH COLUMBIA: 47.251245°, -120.69124°; WGS 84, uncertainty: 5000 m., Source: U.B.C. Forest, Marine Drive, Vancouver. GeoLocate, Georef'd by WTU Staff Elev. 229 ft. Phenology: Flowers & Fruits. Origin: Native. 49° 15' N, 123° 14' W Phenology: Fruits. Origin: Native. J. William Thompson 8312 21 May 1932 V. J. Krajina 781 8 Oct 1949 WTU-19371JWT WTU-214335 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, CLALLAM COUNTY: U.S.A., IDAHO, SHOSHONE COUNTY: Olympic Peninsula, Quillayute Prairie. Mouth of Uranus Creek. 47.94634°, -124.52951°; WGS 84, uncertainty: 1500 m., Source: T50N R4E S5 SW 1/4; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Phenology: Fruits. Origin: Native. Phenology: Flowers. Origin: Native. Geo. N. Jones 3786 Aug 1931 Jean G. Witt 1007 16 Jun 1946 WTU-21516 WTU-116185

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray var. hispidula U.S.A., WASHINGTON, CHELAN COUNTY: South of Navarre Peak. U.S.A., OREGON, JOSEPHINE COUNTY: Elev. 6000 ft. Along Forest Service Road 4105; 2.8 miles from junction with Forest T30N R20E; WGS84, uncertainty: 4800 m., Source: Service Road 4103. Siskiyou National Forest. Georeferenced, Georef'd by WTU Staff Elev. 2400 ft. Timbered slope. Phenology: Fruits. Origin: Native. 42° 18' 55" N, 123° 42' 58" W; T37S R9W S35; T37S R9W S36 Growing on rise to left of road. Purple buds, pinkish-white flowers. George H. Ward 42 29 Jul 1942 One stem with evidence of insect gall. Phenology: Fruits. Origin: Native. WTU-134683 Roger N. Fuller 98-2 27 Jun 1998 Caprifoliaceae WTU-345717 Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., IDAHO, BONNER COUNTY: Priest River Experimental Forest. Lonicera ciliosa (Pursh) Poir. ex DC. 48.355678°, -116.787586°; WGS 84, uncertainty: 4000 m., Source: Digital Map, Georef'd by Melissa Bowers U.S.A., WASHINGTON, CHELAN COUNTY: Seral community, arborvitae-hemlock vegetation zone. Phenology: Slopes of Mount Stuart. Fruits. Origin: Native. Elev. 3500 ft. 47.475118°, -120.903144°; WGS 84, uncertainty: 1500 m., R. F. Daubenmire 44497 17 Sep 1944 Source: GeoLocate, Georef'd by Dan Post Phenology: Flowers. Origin: Native. WTU-271594 J. H. Sandberg 554 24 Jul 1893 with J. B. Leiberg Caprifoliaceae WTU-116037 Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray var. hispidula Caprifoliaceae U.S.A., OREGON, JOSEPHINE COUNTY: Along Forest Service Road 4105; 2.8 miles from junction with Forest Lonicera involucrata (Richardson) Banks ex Spreng. Service Road 4103. Siskiyou National Forest. Elev. 2400 ft. U.S.A., WASHINGTON, OKANOGAN COUNTY: 42° 18' 55" N, 123° 42' 58" W; T37S R9W S35; T37S R9W S36 Ashnola River Camp, Winthrop Ranger Distrist, Okanogan National Growing on rise to left of road. Purple buds, pinkish-white flowers. Forest. One stem with evidence of insect gall. Phenology: Flowers & Fruits. Elev. 4800 ft. Origin: Native. T40N R20E S14; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Roger N. Fuller 98-2 27 Jun 1998 Slope: River bottom; Soil: sandy loam; Vegetation type: Lodgepole Pine; Principal associated plants: Carex sp., Senecio sp.; WTU-345717 28-year-old timber burn. Bearberry honeysuckle. Phenology: Fruits. Origin: Native. Caprifoliaceae R. Gibbons 131-56 8 Aug 1956 Lonicera ciliosa (Pursh) Poir. ex DC. WTU-322658 CANADA, BRITISH COLUMBIA: Inonoaklin River Valley. In woods. Common. Phenology: Flowers. Origin: Native. M. Ridewood 39 20 Jun 1960 WTU-196967 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, CLEARWATER COUNTY: U.S.A., WASHINGTON, WHATCOM COUNTY: Above junction of Elk River and North Fork clearwater River. Gooseberry Point. 46.817408°, -115.466508°; WGS 84, uncertainty: 1000 m., 48.73222°, -122.67083°; WGS 84, uncertainty: 10000 m., Source: Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Climbing overe Holodiscus and Symphoricarpos, under open stand Edge of woods. Phenology: Flowers & Fruits. Origin: Native. of Pinus ponderosa, Pseudotsuga taxifolia and Abies grandis. On hillside. Corolla orange. Phenology: Flowers. Origin: Native. C. L. Hitchcock 3491 22 May 1938 C. W. Sharsmith 3582 5 Jun 1938 WTU-45074 WTU-48813 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, KITSAP COUNTY: Port Orchard. U.S.A., WASHINGTON, CLALLAM COUNTY: 47.645917°, -122.609352°; WGS 84, uncertainty: 10000 m., Beaver Creek. Source: GeoLocate, Georef'd by WTU Staff 48.071272°, -124.291322°; WGS 84, uncertainty: 10000 m., Phenology: Flowers & Fruits. Origin: Native. Source: GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. C. V. Piper 771 20 Jun 1889 G. N. Jones 4550 8 Apr 1934 WTU-16618 WTU-23583 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, SNOHOMISH COUNTY: Marysville. CANADA, BRITISH COLUMBIA: 48.05194°, -122.17583°; WGS 84, uncertainty: 3570 m., Source: Reta Lake, along R. R. tracks. GeoLocate, Georef'd by Maria Yousoufian Phenology: Flowers. Origin: Native. Moist land. Phenology: Flowers. Origin: Native. Garman 5485 31 Jul 1967 J. M. Grant s.n. May 1930 with Henry WTU-43311 WTU-318938 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., MONTANA, FLATHEAD COUNTY: U.S.A., WASHINGTON, MASON COUNTY: 4 miles northwest of Whitefish along U.S. highway 93. Near Shelton. Elev. 3100 ft. 47.21528°, -123.09944°; WGS 84, uncertainty: 5000 m., Source: T31N R22W; WGS 84, uncertainty: 3000 m., Source: Digital Map, GeoLocate, Georef'd by WTU Staff Georef'd by Joseph Stormer At edge of meadow. Phenology: Flowers. Origin: Native. Climbing vine, flowers 6-8 feet off the ground, corolla deep orange outside (?), paler inside. Phenology: Fruits. Origin: Native. George Neville Jones 6530 11 May 1935 Fred Zwickel 12 22 Jun 1949 WTU-29188 WTU-182192 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, KING COUNTY: Near margin of Lake Washington, on Mercer Island. U.S.A., WASHINGTON, YAKIMA COUNTY: 47.57083°, -122.22083°; WGS 84, uncertainty: 10000 m., Source: Cascade mountains, east of Chinook Pass. GeoLocate, Georef'd by WTU Staff Elev. 5000 ft. Near margin of lake. Climbing over brush. Phenology: Flowers & 46.87222°, -121.51444°; WGS 84, uncertainty: 10000 m., Source: Fruits. Origin: Native. GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. J. William Thompson 7588 24 Jul 1931 John Broadbent s.n. 16 Jun 1935 WTU-19373JWT WTU-35483 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., IDAHO COUNTY: U.S.A., OREGON, JOSEPHINE COUNTY: Moscow Mountains. Siskiyou National Forest; on highway 3897, 3.3 miles north & east of 46.815368°, -116.830321°; WGS 84, uncertainty: 15000 m., Grayback Road (3941) or [circa] 20 miles east of Cave Junction. Source: Digital Map, Georef'd by Joseph Stormer Elev. 4390 ft. In woods. Climbing up to 12 feet. Phenology: Flowers. Origin: Cut-over woods of Pseudotsuga and Ceanothus. Flowers Native. reddish-orange, infrequent vine. Phenology: Flowers. Origin: Native. F. L. Pickett 1528 20 May 1934 Melinda F. Denton 2552 23 Jun 1973 WTU-65940 WTU-259229 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SNOHOMISH COUNTY: Stanwood. U.S.A., WASHINGTON, CHELAN COUNTY: 48.24139°, -122.36944°; WGS 84, uncertainty: 10000 m., Source: Near Cottonwood, Entiat Valley. GeoLocate, Georef'd by WTU Staff Elev. 3000 ft. South slope thicket. Phenology: Flowers. Origin: Native. 47.67611°, -120.20722°; WGS84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Lyman Benson 1168 20 Apr 1929 (Common at this and higher elevations). Phenology: Fruits. Origin: Native. WTU-43575 G. E. Morrill 414 23 Aug 1933 Caprifoliaceae WTU-27751 Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., IDAHO, LATAH COUNTY: Cedar Mountain. Lonicera involucrata (Richardson) Banks ex Spreng. 46.80343°, -116.86878°; WGS 84, uncertainty: 1000 m., Source: Nat. Geog. TOPO!, Georef'd by John Bassett U.S.A., MONTANA, BEAVERHEAD COUNTY: Over bush in more open woods. Common, straggling over bush. Wise River near Lodgepole Forest Camp. Phenology: Flowers. Origin: Native. 45.64334°, -113.07749°; uncertainty: 5000 m. Phenology: Flowers. Origin: Native. F. L. Pickett 1116 7 Jul 1917 C. L. Hitchcock 15274 29 Jul 1946 WTU-66498 with C. V. Muhlick WTU-105102 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., IDAHO, KOOTENAI COUNTY: Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray East end of Hayden Lake. 47.76611°, -116.78556°; WGS 84, uncertainty: 10000 m., Source: U.S.A., WASHINGTON, JEFFERSON COUNTY: GeoLocate, Georef'd by WTU Staff Point Hadlock, Olympic Peninsula. Shady woods. Phenology: Flowers. Origin: Native. 47.75028°, -123.74972°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff William H. Baker 14876 15 Jun 1957 Bluff along beach. Phenology: Flowers & Fruits. Origin: Native. WTU-296249 Geo. N. Jones 3141 Jun 1931 WTU-21890 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., IDAHO, BOISE COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Along Forest Service Road 304, North of Idaho City. Elev. 5248 ft. U.S.A., OREGON, HOOD RIVER COUNTY: 43° 48' N, 115° 48' W; T5N R6E S7 To Bluegrass Ridge, Mount Hood. Shrub to 40 centimeters tall, corolla yellow. Phenology: Flowers. Elev. 3000 ft. Origin: Native. 45.364284°, -121.596463°; uncertainty: 6000 m., Georef'd by Joseph Stormer J. F. Smith 4225 8 Jun 2002 Underbrush by trail. Phenology: Flowers. Origin: Native. WTU-351881 J. William Thompson 3288 31 Jul 1927 WTU-19385JWT Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., OREGON, JEFFERSON COUNTY: CANADA, BRITISH COLUMBIA: Lake Suttle. Three miles north of Takla Landing. 44.421253°, -121.741412°; uncertainty: 3200 m. Forest edge. Abundant; fruit ripe; purple. Phenology: Flowers. Phenology: Flowers. Origin: Native. Origin: Native. C. L. Hitchcock 4891 23 Jun 1939 T. T. McCabe 7903 18 Jul 1940 WTU-53620 WTU-78209

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SNOHOMISH COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: 3 miles north of Sylvanus [Silvana]. Near Taholah. 48.23587°, -122.25106°; WGS 84, uncertainty: 500 m., Source: 47.3475°, -124.29194°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Sunshine, edge of woods. Phenology: Flowers. Origin: Native. On bluffs along the ocean. Abundant. Phenology: Flowers & Fruits. Origin: Native. Lyman Benson 1396 11 May 1929 George Neville Jones 6491 11 May 1935 WTU-43576 WTU-29193 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. CANADA, BRITISH COLUMBIA: Sunup Cabin, southwest of Mount Baldy, 8 1/2 miles northwest of CANADA, BRITISH COLUMBIA: Ross Spur. At west edge of town, Arrowhead. Elev. 4200-4500 ft. Along rocky bluffs. Occasional, flowers orange-red. Phenology: Open coniferous forest, decomposed granite soil, moist level place. Flowers. Origin: Native. Phenology: Flowers. Origin: Native. J. A. Calder 8936 10 Jun 1953 George G. Hedgcock s.n. 14 Jun 1939 with D. B. O. Savile WTU-102268 WTU-176889

Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, PIERCE COUNTY: CANADA, BRITISH COLUMBIA: Tacoma. Twenty five miles south of Fort Saint James. 47.25306°, -122.44306°; WGS 84, uncertainty: 10000 m., Source: Under alder and willows in bog. Abundant; shrub to 4 feet; flowers GeoLocate, Georef'd by WTU Staff yellow, fruit dark purplish-black. Phenology: Flowers. Origin: Native. Woods. Phenology: Fruits. Origin: Native. T. T. McCabe 7604 26 Jun 1940 Geo. B. Rigg s.n. 7 Dec 1907 WTU-78210 WTU-16473 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, ISLAND COUNTY: U.S.A., OREGON, JACKSON COUNTY: Edge of Cranberry Lake, Whidbey Island. Ragsdale Butte Lookout, circa 12 miles northwest of Trail. 48.14389°, -122.49111°; WGS 84, uncertainty: 10000 m., Source: Elev. 4500 ft. GeoLocate, Georef'd by WTU Staff Outcrop of blue schistose shale. Phenology: Flowers. Origin: Native. Phenology: Fruits. Origin: Native. C. L. Hitchcock 5019 29 Jun 1939 G. N. Jones 4936 2 Jun 1934 with J. S. Martin WTU-25101 WTU-54298 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON COUNTY: U.S.A., WASHINGTON, ISLAND COUNTY: [No locality given on label] One mile west of Oak Harbor. 47.55564°, -120.536966°; WGS 84, uncertainty: 321158 m., 48.29333°, -122.64194°; WGS 84, uncertainty: 10000 m., Source: Source: Digital Map, Georef'd by Joseph Stormer GeoLocate, Georef'd by WTU Staff Woods near seashore. Indefinite loan by Tacoma High School to Phenology: Fruits. Origin: Native. University of Washington. Phenology: Flowers & Fruits. Origin: Native. C. H. Harrison s.n. 22 Sep 1935 J. B. Flett s.n. 29 Jun 1896 WTU-36943 WTU-37511 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SKAMANIA COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Moosbach (Moss Creek). Along US Hwy 101, near Prairie River. 45.793118°, -121.640474°; WGS 84, uncertainty: 10000 m., T23N R10W S15; NAD 27, uncertainty: 805 m., Source: TRS2LL, Source: GeoLocate, Georef'd by WTU Staff Georef'd by Ben Legler Phenology: Fruits. Origin: Native. In young woodland (about 5-10 years old). Phenology: Fruits. Origin: Native. Wilhelm N. Suksdorf 9434 8 Jul 1896 D. R. Simpson 129 23 Jun 1964 WTU-229655 WTU-287655 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. CANADA, BRITISH COLUMBIA: Sarita River, drainage Sarita River, Victoria. CANADA, BRITISH COLUMBIA: Elev. 30 ft. Queen Charlotte Islands; just south of Skidegate Mission, Graham In gravel bank on seacoast; exposure: west; slope: 30. Flowers Island. yellow. Distributed by the Herbarium, University of Victoria, Victoria, Occasional 5 inches shrubs in thickets at edge of coniferous woods; Canada. Phenology: Flowers & Fruits. Origin: Native. flowers yellow. Phenology: Flowers. Origin: Native. A. Young 312 12 Jun 1963 J. A. Calder 21431 11 Jun 1957 with W. Hubbard with D. B. O. Savile, R. L. Taylor WTU-219315 WTU-241259 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Friday Harbor. Fish Lake; 12 miles northeast of Salmon La Sac. 48.53528°, -123.00528°; WGS 84, uncertainty: 10000 m., Source: 47.51513°, -121.06729°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Phenology: Flowers & Fruits. Origin: Native. Near stream. Shrubs 3 feet tall; corolla yellow, glandular. Phenology: Flowers. Origin: Native. A. S. Pope s.n. 26 Jun 1904 C. L. Hitchcock 7946 21 Jun 1942 WTU-16623 WTU-76204 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. CANADA, BRITISH COLUMBIA: Just south of Lawnhill between Tlell and Skidgate, Graham Island. U.S.A., OREGON, LINCOLN COUNTY: On steep slope of rocky bluffs back from beach. Common; a couple U. S. 101, just south of Lost Creek State Park, 5 miles south of of 6-8 feet shrubs; flowers orange-yellow. Phenology: Flowers. Newport. Origin: Native. T12S R12W S7 Shrub. Phenology: Flowers. Origin: Native. J. A. Calder 20899 25 May 1957 with D. B. O. Savile, R. L. Taylor Noel H. Holmgren 3156 13 May 1968 WTU-241258 WTU-242493 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., IDAHO, BOISE COUNTY: Friday Harbor, San Juan Island. South side of Pilot Peak about 10 miles southwest of Lowman. 48.53528°, -123.00528°; WGS 84, uncertainty: 10000 m., Source: Elev. 7800 ft. GeoLocate, Georef'd by WTU Staff 43.981107°, -115.762294°; WGS 84, uncertainty: 1000 m., Roadsides. Common. Phenology: Flowers & Fruits. Source: GeoLocate, Georef'd by WTU Staff Open grassy slope; near seepage. Phenology: Flowers & Fruits. Barbara D. Blanchard 133 5 Jul 1936 Origin: Native. WTU-45445 C. L. Hitchcock 9917 13 Jul 1944 with C. V. Muhlick Caprifoliaceae WTU-93737 Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, KING COUNTY: Seattle. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: Digital Map, Georef'd by Joseph Stormer U.S.A., WASHINGTON, KITSAP COUNTY: Phenology: Flowers. Origin: Native. Seabeck. 47.63972°, -122.82722°; WGS 84, uncertainty: 10000 m., Source: E. S. Meany s.n. 1898 GeoLocate, Georef'd by WTU Staff On banks and cliffs. Phenology: Flowers & Fruits. Origin: Native. WTU-37180 F. S. Hall s.n. 8 Jul 1928 Caprifoliaceae WTU-44504 Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., MONTANA, GRANITE COUNTY: Skalkaho Pass Road, 32 miles east of Hamilton. Lonicera involucrata (Richardson) Banks ex Spreng. Phenology: Flowers. Origin: Native. CANADA, BRITISH COLUMBIA: C. L. Hitchcock 23941 10 Jul 1965 Between Cascade and Rossland. Moist woods in mountains. Phenology: Flowers. Origin: Native. WTU-223697 J. William Thompson 14426 19 Jul 1939 Caprifoliaceae WTU-21282JWT Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., MONTANA, PARK COUNTY: Near Cooke City. Lonicera ciliosa (Pursh) Poir. ex DC. On sandy bank. Phenology: Fruits. Origin: Native. U.S.A., WASHINGTON, KING COUNTY: C. L. Hitchcock 13587 13 Aug 1945 Alki Point, Seattle. with C. V. Muhlick 47.57639°, -122.40861°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-102149 Phenology: Flowers. Emma A. Shumway s.n. 17 May 1892 Caprifoliaceae WTU-183010 Lonicera involucrata (Richardson) Banks ex Spreng. CANADA, BRITISH COLUMBIA: Caprifoliaceae Huntingdon. Elev. 150 ft. Lonicera involucrata (Richardson) Banks ex Spreng. Along roadside. Flowers yellow, exceptionally early. Phenology: Flowers. Origin: Native. U.S.A., OREGON COUNTY: McKenzie Pass. T. T. McCabe 3309 26 Apr 1936 Bush Honeysuckle, Ink-berry, Twinberry. Phenology: Flowers. Origin: Native. WTU-45282 L. Javete s.n. 29 Jun 1936 WTU-73318 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., IDAHO COUNTY: U.S.A. COUNTY: St. Marie's Highway 6 miles north of Harvard. On campus. Low meadow. Phenology: Fruits. Origin: Native. Yellow flowered honey. [No collection date given on label]. Phenology: Flowers. F. L. Pickett 1516 27 May 1934 E. C. s.n. WTU-65947 WTU-16464 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SAN JUAN COUNTY: Stuart Island. CANADA, BRITISH COLUMBIA: 48.674722°, -123.210556°; WGS 84, uncertainty: 3700 m., Near Guichon Creek, 13 miles south of Savona. Source: Georeferenced, Georef'd by WTU Staff Elev. 3500 ft. Moist cliffs along sound. Not very common. Phenology: Flowers. 50° 32' N, 120° 52' W Origin: Native. Among Salix in thicket small swale. Shrubs 1-2 meters tall. Phenology: Flowers. Origin: Native. W. H. Lawrence 100 20 Jun 1904 C. L. Hitchcock 7413 23 Jun 1941 WTU-95440 with J. S. Martin WTU-68256 Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Caprifoliaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. Yellow Island. Slopes above cabin at west spit. 48.54361°, -123.16333°; WGS 84, uncertainty: 10000 m., Source: U.S.A., WASHINGTON, KING COUNTY: GeoLocate, Georef'd by WTU Staff Seattle, campus, State University. Dry woodland. Phenology: Flowers & Fruits. Origin: Native. 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff E. Habegger s.n. 5 Jul 1996 In woods. Phenology: Flowers. WTU-333076 T. C. Frye S-297 15 May 1904 WTU-20315 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., IDAHO, VALLEY COUNTY: Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Along Big Creek at Green Tables Forest Camp, Payette National Forest, Sawtooth Mountains. U.S.A., WASHINGTON, KING COUNTY: Elev. 6500 ft. Seattle. 44.583659°, -115.881559°; WGS 84, uncertainty: 4800 m., 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: GeoLocate, Georef'd by WTU Staff Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers & Fruits. Origin: Native. Phenology: Fruits. Origin: Native. J. William Thompson 13739 8 Jul 1937 C. V. Piper 408 15 Jun 1888 WTU-38733 WTU-16615

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, IDAHO COUNTY: U.S.A., OREGON, DESCHUTES COUNTY: Headwaters of Sheep Creek, Seven Devils Mountains. Just below Lava Lake, 1/4 mile east of McKenzie Pass. 45.456717°, -116.535407°; WGS 84, uncertainty: 10000 m., In Pinus Albicaulis-Abies lasiocarpa-Tsuga mertensiana forest. Source: GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Open, wet meadow. Phenology: Flowers. Origin: Native. A. R. Kruckeberg 2128 5 Jul 1950 Marion Ownbey 2078 26 Jun 1940 with Fred G. Meyer WTU-183620 WTU-73617 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. CANADA, BRITISH COLUMBIA: U.S.A., IDAHO, VALLEY COUNTY: At south end of Bolean Lake, 4 3/4 miles northeast of Falkland. Along Big Creek at Green Tables Forest Camp, Payette National Elev. 4715 ft. Forest, Sawtooth Mountains. In marsh. Shrubs 1 meter tall. Phenology: Flowers. Origin: Native. Elev. 6500 ft. 44.583659°, -115.881559°; WGS 84, uncertainty: 4800 m., C. L. Hitchcock 7522 1 Jul 1941 Source: GeoLocate, Georef'd by WTU Staff with J. S. Martin Phenology: Flowers. Origin: Native. WTU-68006 J. William Thompson 13739 8 Jul 1937 WTU-10121JWT Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae CANADA, BRITISH COLUMBIA: Lonicera ciliosa (Pursh) Poir. ex DC. Vicinity of Upper Hat Creek, base of Marble Mountains. Elev. 3000 ft. U.S.A., WASHINGTON, KING COUNTY: 50.817167°, -121.574099°; uncertainty: 8000 m. State University, Seattle. Phenology: Flowers. Origin: Native. 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. William Thompson 355 12 Jul 1938 Open woods. Phenology: Flowers. Origin: Native. with Emily M. Thompson Lyman Benson 1487 29 May 1929 WTU-17425JWT WTU-43572 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, KING COUNTY: Seattle. U.S.A., MONTANA COUNTY: 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Flathead National Forest; Echo Lake. Source: Digital Map, Georef'd by Joseph Stormer Elev. 3400 ft. Phenology: Flowers. Origin: Native. 48.17196°, -113.891917°; uncertainty: 15000 m. Phenology: Flowers. Origin: Native. E. S. Meany s.n. 1898 C. H. Kauffman 104 27 Jun 1928 WTU-37179 with G. B. Cummins WTU-252537 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., IDAHO, IDAHO COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. Bimerick Meadows. T34N R8E S22; WGS 84, uncertainty: 10000 m., Source: U.S.A., WASHINGTON, KING COUNTY: GeoLocate, Georef'd by WTU Staff In Seattle. In moist soil at the margins of wet sedge meadow. Shrub to 1 meter 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: tall. Berries black. Phenology: Flowers & Fruits. Origin: Native. GeoLocate, Georef'd by WTU Staff Moist woods. Phenology: Flowers. Quentin Jones 254 17 Jul 1950 J. William Thompson 10392 6 May 1934 WTU-141849 WTU-6177JWT Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, KING COUNTY: Revenna Park. U.S.A., WASHINGTON, KING COUNTY: Phenology: Flowers. Origin: Native. Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Gavett 693 25 May 1908 Source: Digital Map, Georef'd by Joseph Stormer Open woods. Phenology: Flowers. WTU-272196 J. William Thompson 6542 23 May 1931 WTU-19386JWT Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. CANADA, BRITISH COLUMBIA: U.S.A., WASHINGTON, KING COUNTY: Mount Revelstoke National Park; West Woolsey Creek (Silver 1/4 mile north east of Seattle. Creek) along Trail no. 6, West Fork Woolsey Trail. 47.687541°, -122.293619°; WGS 84, uncertainty: 10000 m., 51° 9' N, 117° 55' W Source: Digital Map, Georef'd by Joseph Stormer In damp soil on rocky slope of ravine. 9 Repls. Phenology: Flowers. In bushes. Phenology: Flowers. Origin: Native. Origin: Native. W. J. Eyerdam 6519 20 May 1949 J. H. Soper 12791 3 Aug 1970 with M. J. Shchepanek WTU-290961 WTU-257556 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON COUNTY: Wenatchee National Forest. Near Chiwaukum Ranger Station. U.S.A., MONTANA COUNTY: Deep clay soil; associated species: Amelanchier, Acer, Salix. Lac. specimen cer. Unknown species is plentiful in vicinity of Chiwauku. Flowering in Elev. 7000 ft. May/June; seed dissemination July. Unknown, probably eaten by Moist woods. Phenology: Flowers. Origin: Native. sheep; browse, density: 0.3. Phenology: Flowers & Fruits. Origin: Native. D. B. Swingle s.n. 29 Jun 1933 Frank B. Lenzie 242 17 Jun 1917 WTU-37096 WTU-322657 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, CHELAN COUNTY: Slope of Stuart Ridge. U.S.A., OREGON, BAKER COUNTY: Elev. 3500 ft. 4 miles below Anthony Lake, Elkhorn Range; Blue Mountains. 47.47024°, -120.78503°; WGS84, uncertainty: 13000 m., Source: Moist soil along stream in spruce-fir woodland. Frequent. Phenology: Georeferenced, Georef'd by WTU Staff Fruits. Origin: Native. Yellow pine slopes. Phenology: Flowers. Origin: Native. Bassett Maguire 26951 10 Aug 1946 J. William Thompson 11726 19 Jun 1935 with Arthur H. Holmgren WTU-6239JWT WTU-111616

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, PIERCE COUNTY: U.S.A., WASHINGTON, STEVENS COUNTY: Upper Valley of the Nesqually [Nisqually]. Addy, 3 miles northwest. Gardiner road. 46.79101°, -121.748729°; WGS 84, uncertainty: 2766 m., Source: 48.35611°, -117.83639°; WGS 84, uncertainty: 10000 m., Source: Digital map, Georef'd by Maria Yousoufian GeoLocate, Georef'd by WTU Staff Open ground. Phenology: Flowers & Fruits. Origin: Native. Phenology: Flowers. Origin: Native. O. D. Allen 215 16 Jun 1896 Carl H. Spiegelberg 254 5 Jun 1923 WTU-37175 WTU-174533

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., MONTANA, GRANITE COUNTY: CANADA, BRITISH COLUMBIA: Off the Dry Creek Road. Lake Wapta. Elev. 6700 ft. 51.439549°, -116.350259°; uncertainty: 1000 m. Several flow bushes seen on the extensive clearcuts. Phenology: Fly-honey suckele. Phenology: Flowers. Origin: Native. Flowers. Origin: Native. unknown B-11 Jul 1927 Klaus H. Lackschewitz 7299 14 Jun 1977 WTU-318946 WTU-288582 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., IDAHO, BONNER COUNTY: U.S.A., WASHINGTON, SPOKANE COUNTY: Along Lambs Creek, Priest Lake. Mount Carleton (Mount Spokane). 48.49056°, -116.9025°; WGS 84, uncertainty: 10000 m., Source: 47.9167°, -117.1°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by Dan Post Moist ground. Phenology: Flowers & Fruits. Origin: Native. Sandy wood. Phenology: Flowers. Origin: Native. William H. Baker 15981 25 Jun 1960 F. O. Kreager 200 17 Jul 1902 WTU-261419 WTU-68518

Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., MONTANA, MISSOULA COUNTY: San Juan Island, road to False Bay, circa 1.5 miles west of Cattle Lolo Pass, Montana-Idaho border. Point Road, on south end of island. 46.635261°, -114.579964°; uncertainty: 3000 m., Georef'd by 48.450555°, -122.9944°; WGS 84, uncertainty: 500 m., Source: Joseph Stormer GeoLocate, Georef'd by WTU Staff Phenology: Flowers & Fruits. Origin: Native. Along the road. Vine common, flowers mostly reddish-orange, the extreme distal end of lobes orange. Phenology: Fertile. Origin: C. L. Hitchcock 14654 12 Jul 1946 Native. with C. V. Muhlick Melinda F. Denton 3490 2 Aug 1974 WTU-104415 WTU-259588 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, CHELAN COUNTY: CANADA, BRITISH COLUMBIA: Wenatchee Mountains. Mission Creek Canyon ca. [circa] 8 miles Lockside Drive, Victoria. south of Cashmere. Woods. Black train ferry. Phenology: Flowers. Origin: Native. 47.394627°, -120.500256°; WGS84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff unknown 5596 5 May 1970 Sandy soil from sandstone cliffs. Flowers orange. Phenology: Flowers. Origin: Native. WTU-318945 C. L. Hitchcock 17229 14 Jun 1948 Caprifoliaceae WTU-125237 Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae CANADA, BRITISH COLUMBIA: Yoho Valley. Lonicera involucrata (Richardson) Banks ex Spreng. Field. Involuchsed fly-honeysuckle. Phenology: Flowers. Origin: Native. U.S.A., OREGON, WALLOWA COUNTY: At Wallowa Lake. unknown 1728 8 Aug 1957 Moist thickets. Phenology: Flowers. Origin: Native. WTU-318944 William H. Baker 6796 2 Jun 1950 WTU-302094 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., IDAHO, VALLEY COUNTY: Lonicera utahensis S. Watson At summit of divide between Warm Lake and Cascade. 44.51639°, -116.04083°; WGS 84, uncertainty: 10000 m., Source: CANADA, BRITISH COLUMBIA: GeoLocate, Georef'd by WTU Staff Road 3 miles from Tulameen River. Phenology: Flowers. Origin: Native. In open spruce-subalpine fir forest, along Grasshopper Mountain. Large red fruits in pairs. Phenology: Vegetative. Origin: Native. C. L. Hitchcock 14017 26 Jun 1946 with C. V. Muhlick A. R. Kruckeberg 5707 7 Sep 1963 WTU-105105 WTU-218719 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, OKANOGAN COUNTY: U.S.A., IDAHO, CUSTER COUNTY: Below Big Craggy. Redfish Lake, Stanley Basin, Sawtooth Mountains. Elev. 3000 ft. 44.4425°, -114.92722°; WGS 84, uncertainty: 10000 m., Source: 48.76328°, -120.32716°; WGS 84, uncertainty: 1800 m., Source: GeoLocate, Georef'd by WTU Staff Georeferenced, Georef'd by WTU Staff Woods. Phenology: Flowers. Origin: Native. By mountain stream. Phenology: Fruits. Origin: Native. William H. Baker 10601 22 Jul 1953 J. William Thompson 10873 30 Jun 1934 WTU-296051 WTU-6180JWT Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, BONNER COUNTY: U.S.A., MONTANA, PARK COUNTY: Priest River Exptl. Forest. Along the old flume between the dam and the power plant at the falls 48.349577°, -116.807015°; WGS 84, uncertainty: 4000 m., of the Clarks Fork of the Yellowstone River, about 4 miles east of Source: Digital Map, Georef'd by Melissa Bowers Cooke City. Roadside. Community: Status: seral. Type: dense, scrub. Phenology: Fruits. Origin: Native. Phenology: Flowers & Fruits. Origin: Native. Jean G. Witt 1418 6 Sep 1948 R. F. Daubenmire 44276 12 Jun 1944 WTU-129063 WTU-271604

Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, CHELAN COUNTY: U.S.A., WASHINGTON, CHELAN COUNTY: North end of ridge on east side of Tumwater Canyon. Slope of Stuart Ridge. T25N R17E; WGS84, uncertainty: 10000 m., Source: GeoLocate, Elev. 3500 ft. Georef'd by WTU Staff 47.47024°, -120.78503°; WGS84, uncertainty: 13000 m., Source: Forest slope. Phenology: Flowers. Origin: Native. Georeferenced, Georef'd by WTU Staff Yellow pine slopes. Phenology: Flowers. Origin: Native. George H. Ward 523 21 Jun 1946 J. William Thompson 11726 19 Jun 1935 WTU-134685 WTU-33461 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., MONTANA, JEFFERSON COUNTY: About 10 miles south of Boulder. U.S.A., IDAHO, VALLEY COUNTY: 46.091814°, -112.120834°; uncertainty: 12800 m. North end of Payette Lake, 8 miles north of McCall. Moist Douglas fir forest. Phenology: Flowers. Origin: Native. 45.026962°, -116.09778°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff W. E. Booth 54471 27 Jun 1954 Edge of meadow. Phenology: Flowers. Origin: Native. WTU-302158 William H. Baker 10286 8 Jul 1953 WTU-296049 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, LEWIS COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Near Centralia. Elev. 50 ft. U.S.A., IDAHO COUNTY: 46.71639°, -122.95306°; WGS 84, uncertainty: 5000 m., Source: Troy Road. GeoLocate, Georef'd by WTU Staff Along the edge of woods forming thickets, streamside. Phenology: Roadside ditches. Phenology: Fruits. Origin: Native. Flowers. Origin: Native. J. William Thompson 10105 7 Apr 1934 William H. Baker 12831 27 May 1955 WTU-6181JWT WTU-296118 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, ISLAND COUNTY: U.S.A., IDAHO, CLEARWATER COUNTY: West side of Camano Island. 4 miles northeast of Elk River. Elev. 10-20 ft. 46.824541°, -116.119093°; WGS 84, uncertainty: 1000 m., 48.17417°, -122.52694°; WGS 84, uncertainty: 10000 m., Source: Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Roadside. Phenology: Flowers. Origin: Native. Shade and sun. Vine habit of growth. Phenology: Flowers. Origin: Native. Mary Hafercamp 132 21 Jul 1948 with Xerpha M. Gaines Ora P. Quense 95 19 May 1940 WTU-132722 WTU-316065 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, IDAHO COUNTY: U.S.A., IDAHO, SHOSHONE COUNTY: Meadow Creek. above Selway Falls. Mouth of Bullion Creek at junction with North Fork of the Saint Joe 45.481973°, -115.931284°; WGS 84, uncertainty: 10000 m., River. Source: GeoLocate, Georef'd by WTU Staff 47.377453°, -115.764825°; WGS 84, uncertainty: 10000 m., In shade under coniferous climax, slopes overhanging creek. Small Source: GeoLocate, Georef'd by WTU Staff shrub, 2-4 feet high. Phenology: Fruits. Origin: Native. Phenology: Flowers. Origin: Native. L. Constance 1651 30 May 1936 William H. Baker 13482 15 Jul 1955 with R. C. Rollins WTU-296119 WTU-5731JWT

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., MONTANA, GLACIER COUNTY: Swauk Creek, near Liberty. Glacier National Park, foot of Mount Wilbur; Swiftcurrent Valley. 47.260278°, -120.694722°; NAD 27, uncertainty: 10000 m., Elev. 5200 ft. Source: Georeferenced, Georef'd by WTU Staff Area from which snow had recently melted at the base of a small Along the banks of creek. Phenology: Flowers & Fruits. Origin: cliff. Phenology: Flowers. Origin: Native. Native. Daniel H. Brant 114 28 Jul 1948 J. William Thompson 11819 27 Jun 1935 WTU-129561 WTU-6211JWT Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., IDAHO, BLAINE COUNTY: U.S.A., WASHINGTON, THURSTON COUNTY: Pettie Lake, Sawtooth Range. Near Deschutes River. T8N R14E S31; NAD 27, uncertainty: 805 m., Source: TRS2LL, 47.017618°, -122.90455°; WGS 84, uncertainty: 5000 m., Source: Georef'd by Ben Legler GeoLocate, Georef'd by WTU Staff Wet marshy banks. Phenology: Flowers. Origin: Native. In open woods. Phenology: Flowers. Origin: Native. William H. Baker 10849 6 Aug 1953 I. C. Otis 2058 24 May 1936 WTU-296120 WTU-13009JWT

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, IDAHO COUNTY: U.S.A., MONTANA, LAKE COUNTY: Bimrick [Bimerick] Meadows. Several miles southeast of McDonald Lake; Mission Range. 46.272222°, -115.444444°; uncertainty: 2000 m., Georef'd by On cliffs high above. Phenology: Flowers. Origin: Native. Joseph Stormer Marshy deforested area; spruce-fir zone. Phenology: Flowers & C. L. Hitchcock 18305 12 Jul 1948 Fruits. Origin: Native. WTU-125236 R. F. Daubenmire 38172 4 Jun 1938 WTU-271522 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, THURSTON COUNTY: U.S.A., IDAHO, VALLEY COUNTY: Near Deschutes River. Along North Fork of Lake Fork Payette River, 12 miles northeast of 47.017618°, -122.90455°; WGS 84, uncertainty: 5000 m., Source: McCall. GeoLocate, Georef'd by WTU Staff 44.922416°, -115.946308°; WGS 84, uncertainty: 1000 m., In open woods. Phenology: Flowers. Origin: Native. Source: GeoLocate, Georef'd by WTU Staff Open woods. Phenology: Flowers. Origin: Native. I. C. Otis 2058 24 May 1936 William H. Baker 11637 6 Jul 1954 WTU-56616 WTU-297789 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., IDAHO, CARIBOU COUNTY: Forest Service Campground, Gravel Creek. CANADA, BRITISH COLUMBIA: Elev. 6700 ft. Between Cascade and Rossland. 42° 55' N, 111° 20' W; T5S R43E S35 SW; WGS 84, uncertainty: Elev. 4000 ft. 500 m., Source: Nat. Geog. TOPO!, Georef'd by John Bassett 49.040529°, -118.0093°; WGS 84, uncertainty: 14442 m., Source: Shrub to 1.5 meters, berries black, bracts bright red. Phenology: Digital Map, Georef'd by Joseph Stormer Flowers. Origin: Native. Crest of High Mountain. Legit: J. W. Thompson. Phenology: Fruits. James F. Smith 2624 24 Jul 1993 J. W. Thompson 14412 18 Jul 1939 WTU-328802 WTU-21322JWT

Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Klickitat River. Above Elk Lake. Phenology: Flowers. Origin: Native. T23N R2E S32; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff B. Flett 1264 16 Jul 1899 Moist marshy banks. Phenology: Flowers. Origin: Native. WTU-37182 William H. Baker 12250 30 Jul 1954 WTU-298010 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae CANADA, BRITISH COLUMBIA: Lonicera utahensis S. Watson Outlet of Wolverine Lake east of Manson Creek village. 55° 41' N, 124° 26' W U.S.A., IDAHO, VALLEY COUNTY: At edge of deciduous thickets. Common; fruit black. Phenology: On Gold Fork lookout, Payette National Forest, Sawtooth Mountains. Fruits. Origin: Native. 44.67194°, -115.83667°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. A. Calder 13640 28 Jul 1954 Deep canyon. Phenology: Fruits. Origin: Native. with D. B. O. Savile, J. M. Ferguson J. William Thompson 13909 19 Jun 1937 WTU-173118 WTU-10566JWT Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, VALLEY COUNTY: Deep canyon on Gold Fork Lookout, Payette National Forest, U.S.A., IDAHO, SHOSHONE COUNTY: Sawtooth Mountains. 1 mile northeast of summit; between Bovill and Clarkia. Elev. 7000 ft. Elev. 3000 ft. 44.67194°, -115.83667°; WGS 84, uncertainty: 10000 m., Source: T42N R1E S27; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Deep canyon. On gold fork lookout. Phenology: Fruits. Origin: In dry meadow area along draw in mixed coniferous area. Shrub Native. about 1 meter or less tall; bracts red; fruits black. Phenology: Fruits. Origin: Native. J. William Thompson 13906 19 Jul 1937 Arthur Cronquist 6008 30 Jul 1949 WTU-10569JWT with Quentin Jones WTU-181974 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., MONTANA, BEAVERHEAD COUNTY: U.S.A., MONTANA, PARK COUNTY: At Pintlar Falls. Near the X-L-O mine's ball mill, north of Cooke Guard Station, In partial shade. Phenology: Fruits. Origin: Native. vicinity of Cooke City. 45.033593°, -109.872273°; WGS 84, uncertainty: 3036 m., C. L. Hitchcock 12881 27 Jul 1945 Source: Digital Map, Georef'd by Joseph Stormer with C. V. Muhlick Meadow. Phenology: Flowers. Origin: Native. WTU-102018 Jean G. Witt 1222 8 Jul 1948 WTU-129064 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., MONTANA, PONDERA COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. var. Ten miles southwest of Glacier Park, in Marias Pass. involucrata Phenology: Fruits. Origin: Native. U.S.A., OREGON, COOS COUNTY: C. L. Hitchcock 18147 7 Jul 1948 Cape Arago, south of Coos Bay. T26S R14W; WGS 84, uncertainty: 1784 m., Source: Digital Map, WTU-125449 Georef'd by Joseph Stormer Jutting promotory. Shrub 1-2 meters high; flowers yellow. Phenology: Flowers. Origin: Native. Caprifoliaceae Arthur Cronquist 6848 14 Apr 1953 Lonicera japonica Thunb. WTU-152445 U.S.A., WASHINGTON, ASOTIN COUNTY: South shore of reservoir, (Lower Granite "Lake"), Snake River, 1.5 miles east of Silcott. Caprifoliaceae Elev. 721 ft. 46° 25.3' N, 117° 10.1' W Lonicera conjugialis Kellogg South shore of reservoir. Flowers white, fragrant, fade to yellow, 4.8 centimeters long, lip = tube, spreading from garden relic, into U.S.A., OREGON, JOSEPHINE COUNTY: thickets, former homestead site. Phenology: Flowers. Grayback Mountain. Dry slope. Phenology: Fruits. Peter F. Zika 16661 1 Oct 2001 William H. Baker 910 18 Jul 1950 WTU-346390 with George C. Ruhle WTU-298304 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., IDAHO, LATAH COUNTY: Lonicera conjugialis Kellogg Near mouth of Cedar Creek. Elev. 1600 ft. U.S.A., OREGON, JOSEPHINE COUNTY: T38N R2W S11; WGS 84, uncertainty: 10000 m., Source: Siskiyou Mountains. Bolan Lake. GeoLocate, Georef'd by WTU Staff Elev. 5000 ft. Along old road next to creek, noutheast exposure, Abigra/Pacmyr H. 42.022068°, -123.45918°; uncertainty: 750 m., Georef'd by Joseph T., decomposed granite. Several. Phenology: Flowers. Origin: Stormer Native. Mountain meadow. Phenology: Fruits. C. A. Wellner 178 13 Jun 1974 J. William Thompson 12499 5 Aug 1935 WTU-353915 WTU-4336JWT Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. var. U.S.A., IDAHO, LATAH COUNTY: ledebourii (Eschsch.) Jeps. Along the Troy Road, eight miles east of Moscow. 46.7325°, -116.99917°; WGS 84, uncertainty: 10000 m., Source: U.S.A., OREGON, LANE COUNTY: GeoLocate, Georef'd by WTU Staff Along the Siltcoos outlet, near the ocean. Thickets, open banks. Phenology: Flowers. Origin: Native. T19S R12W In swales among partly stabilized dunes. Shrub 2 meters tall; flowers William H. Baker 7543 25 May 1951 yellow. Phenology: Flowers. Origin: Native. WTU-302159 Arthur Cronquist 6876 21 Apr 1953 WTU-152444 Caprifoliaceae Caprifoliaceae Lonicera conjugialis Kellogg Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., OREGON, JOSEPHINE COUNTY: U.S.A., OREGON, JOSEPHINE COUNTY: Cliffs above Bolan Lake, Siskiyou Mountains, near the California Bolan Lake, Siskiyou Mountains. border. Elev. 5500 ft. Elev. 6000 ft. In damp Douglas Fir-White Fir-Tsuga forest. Around rim of lake. Crevices of rocks, clliffs. Phenology: Fruits. Phenology: Flowers. Origin: Native. C. L. Hitchcock 5252 7 Jul 1939 A. R. Kruckeberg 1929 23 Jun 1950 with J. S. Martin WTU-183673 WTU-53628 Caprifoliaceae Caprifoliaceae Lonicera maackii (Rupr.) Herder Lonicera utahensis S. Watson U.S.A., OREGON, LANE COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Near west bank of Willamette River, near east end of Merry Lane, Middle Fork Clearwater River. At 3 Devils Creek, 4 miles below Eugene. Lowell, Nez Perce Forest. Elev. 393 ft. Elev. 1500 ft. 44° 5.1' N, 123° 7.1' W 46.086257°, -115.59472°; WGS 84, uncertainty: 1000 m., Source: Partial shade, with Alnus rubra, Populus balsamifera, Fraxinus GeoLocate, Georef'd by WTU Staff latifolia, Rubus armeniacus, Symphoricarpos albus, riparian forest. Exposed south bank of river. Shrub, 2-3 feet high, berries red. Two wild shrubs, 2-3 meters tall immature. Phenology: Flowers. Phenology: Fruits. Origin: Native. P. F. Zika 20094 27 Aug 2004 L. Constance 1610 31 May 1936 with E. R. Alverson with R. C. Rollins WTU-359307 WTU-9476JWT Caprifoliaceae Caprifoliaceae Lonicera conjugialis Kellogg Lonicera japonica Thunb. U.S.A., OREGON COUNTY: U.S.A., WASHINGTON, KING COUNTY: Top of Siskiyou Mountains, near Waldo. Clarke Beach Park, southeast end of Mercer Island, Lake Phenology: Flowers & Fruits. Washington. Elev. 19 ft. Thomas Howell s.n. 9 Jul 1887 T24N R5E S30; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler WTU-19313JWT Adventive vine. Flowers cream, fade yellow, fruit black. Phenology: Flowers. Caprifoliaceae Zika 14525 6 Oct 1999 with A. Weinmann Lonicera involucrata (Richardson) Banks ex Spreng. var. ledebourii (Eschsch.) Jeps. WTU-340086 U.S.A., OREGON, LINCOLN COUNTY: On the Pacific Coast 4 miles south of Waldport. Caprifoliaceae Along the beach. Shrubs to 4 feet high along the beach. (Scarcely typical of inland plants.) Flowers yellow. Phenology: Fruits. Origin: Lonicera conjugialis Kellogg Native. U.S.A., OREGON, JOSEPHINE COUNTY: C. L. Porter 7712 6 Sep 1958 Siskiyou National Forest; around Bolan Lake. with Marjorie W. Porter Elev. 5300-6000 ft. Woods of Tsuga mertensiana, Abies lasiocarpa, Abies grandis, WTU-186460 Pseudotsuga. Shrub, locally common in open wooded area, flowers maroon. Phenology: Flowers. Caprifoliaceae Melinda F. Denton 2724 25 Jun 1973 Lonicera conjugialis Kellogg WTU-259486 U.S.A., OREGON COUNTY: Ashland Butte. Near sun line. Phenology: Fruits. Drake s.n. Jul 1986 with Dickson WTU-16786 Caprifoliaceae Caprifoliaceae Lonicera conjugialis Kellogg Lonicera utahensis S. Watson U.S.A., OREGON, DESCHUTES COUNTY: U.S.A., IDAHO, FRANKLIN COUNTY: West base of Brokentof Mountain. Cache National Forest, Bear River Range. North of Franklin on Stream bank. Phenology: Fruits. Highway 91, northeast on Cub River Road approximately 17 miles, then 2.4 miles east on the German Dugway. Morton E. Peck 14390 12 Jul 1925 Elev. 9197 ft. 42° 12.11' N, 111° 37.33' W WTU-19392 Under mixed conifers. Moist area with clay loam soil. North exposure. Assocaited with Geranium viscosissimum and Pedicularis racemosa. Phenology: Fruits. Origin: Native. Caprifoliaceae Allyson Davis 141 14 Aug 1997 Lonicera conjugialis Kellogg WTU-336072 U.S.A., OREGON, JEFFERSON COUNTY: East slope of the Cascades, off of United States Highway 20 at Corbett Sno Park, 8.8 miles east of Santiam Junction. Caprifoliaceae Elev. 4080 ft. 44° 25.23' N, 121° 47.49' W Lonicera involucrata (Richardson) Banks ex Spreng. Flowers purple, shrub to 2 meters tall, associated genera: Pseudotsuga, Pteridium, Abies, Acer, Rubus, Ribes, Ceanothus. CANADA, BRITISH COLUMBIA: Phenology: Flowers & Fruits. New Hazelton. 55.25°, -127.583333°; WGS 84, uncertainty: 3000 m., Source: Richard R. Halse 6053 21 Jun 2001 GeoLocate, Georef'd by Joseph Stormer Forming with Lonicera and Alnus dense wet bottom WTU-347893 jungles.Abundant and large, occasionally to 5-6 feet, usually 4-5 feet in height. Now in full fruit, berries purple. Phenology: Fruits. Origin: Native. Caprifoliaceae T. T. McCabe 8102 4 Aug 1940 Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata WTU-78211 U.S.A., OREGON, WALLOWA COUNTY: Caprifoliaceae Wallowa-Whitman National Forest, Eagle Cap Wilderness; from Two Pan to 2 miles south. Lonicera periclymenum L. Elev. 5750-8000 ft. 45.193933°, -117.340333°; WGS 84, uncertainty: 8000 m., U.S.A., WASHINGTON, KING COUNTY: Source: Digital Map, Georef'd by Joseph Stormer Marsh Island, Union Baby, Lake Washington. Exposed loose rocks on east-faceing slope. Shrub circa 3 feet tall; Elev. 13 ft. rare; corollas yellow. Phenology: Flowers & Fruits. Origin: Native. T25N R4E S21; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Melinda F. Denton 3310 15 Jul 1973 In willow thickets. White flowers fade yellow, bird-sown 3 meters vine adventive. Reference WTU-258694 specimen for illustration in Revision of Flora of the Pacific Northwest, WTU Herbarium, 2016. Phenology: Flowers. Caprifoliaceae P. F. Zika 16207 Lonicera involucrata (Richardson) Banks ex Spreng. with A. L. Jacobson U.S.A., WASHINGTON, KITTITAS COUNTY: 8 Jun 2001 Scatter Creek Camp, upper Cle Elum River. 47.176973°, -120.991239°; WGS 84, uncertainty: 10000 m., WTU-358222, WTU-V-031919 Source: GeoLocate, Georef'd by WTU Staff Along canyon bottom in Abies lasiocarpa - Picea engelmanni forest. Phenology: Flowers. Origin: Native. Caprifoliaceae A. R. Kruckeberg 2968 24 Jun 1952 Lonicera involucrata (Richardson) Banks ex Spreng. WTU-156515 U.S.A., WASHINGTON, KITTITAS COUNTY: Swauk Creek, near Liberty. Caprifoliaceae 47.260278°, -120.694722°; NAD 27, uncertainty: 10000 m., Source: Georeferenced, Georef'd by WTU Staff Lonicera periclymenum L. Along the banks of creek. Phenology: Fruits. Origin: Native. U.S.A., WASHINGTON, CLARK COUNTY: J. William Thompson 11819 27 Jun 1935 Floodplain of Shanghai Creek near Northeast 202 Ave. Elev. 246 ft. WTU-34238 45° 41.2' N, 122° 28.1' W; T2N R3E S5 Naturalized, edge of riparian zone, full sun; associated with Fraxinus, Quercus, Rhamnus. White flowers fading yellow; vine climbing 6 meters into trees. Phenology: Flowers. P. F. Zika 16192 7 Jun 2001 WTU-358220 Caprifoliaceae Caprifoliaceae Lonicera conjugialis Kellogg Lonicera utahensis S. Watson U.S.A., OREGON, JOSEPHINE COUNTY: U.S.A., IDAHO, BONNEVILLE COUNTY: Oregon Caves Nataional Park and vicinity, Sand Ridge, Lake Caribou Range, Anderson Gulch, along road to Caribou City (ghost Mountain Trail. town), 15 airline miles south of Palisades Dam. 42.086445°, -123.368135°; WGS 84, uncertainty: 800 m., Source: Elev. 6800 ft. Digital Map, Georef'd by Joseph Stormer T3S R44E S27; WGS 84, uncertainty: 10000 m., Source: Phenology: Fruits. GeoLocate, Georef'd by WTU Staff One shrub in lodgepole pine forest. 3 to 4 feet tall; fruit salmon pink. William H. Baker 163 30 Jun 1949 Phenology: Fruits. Origin: Native. with George C. Ruhle Noel H. Holmgren 5463 17 Jul 1971 WTU-298314 with Vernon Marttala WTU-276380 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., IDAHO COUNTY: Lonicera sp. Palouse Country and about Lake Coeur d' Alene. -116.747304°, 0°; uncertainty: 24000 m. U.S.A., OREGON, JACKSON COUNTY: Valley. Phenology: Fruits. Origin: Native. Huckleberry Mountain, Rogue River National Forest. Elev. 4500 ft. E. A. Shumway s.n. Jun 1892 Alpine meadows. Phenology: Fruits. WTU-188496 J. William Thompson 13014 1 Jul 1936 WTU-2892JWT Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON COUNTY: Lonicera periclymenum L. Nason Creek. Elev. 2150 ft. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Phenology: Fertile. Origin: Native. Route 109, near Condra Road and Copalis River, Copalis Beach. Elev. 16 ft. J. H. Sandberg 610 30 Jul 1893 47° 7.1' N, 124° 10.2' W with J. B. Leiberg Adventive in thickets with Rubus armeniacus, Salix hookeriana, Pinus contorra. White flowers fade dull yellow, tube always purple as WTU-116036 are buds, fruit dark red; twining vine, no fused leaves. Phenology: Flowers. Caprifoliaceae P. F. Zika 17694 27 Aug 2002 Lonicera conjugialis Kellogg WTU-354174 U.S.A., OREGON COUNTY: Castle Rock Gardens, Goverment Camp, Crater Lake National Park. Caprifoliaceae Elev. 6500 ft. Rocky hillside. Phenology: Flowers & Fruits. Lonicera involucrata (Richardson) Banks ex Spreng. W. G. Sclheim 701 31 Jul 1933 U.S.A., MONTANA, DEER LODGE COUNTY: Anaconda, Pintlar Range, east slope of Mount Tiny. WTU-47592 Elev. 8500 ft. Still in the krummholz of alpine fir, near watercourse. Low bushes. Phenology: Fruits. Origin: Native. Caprifoliaceae Klaus H. Lackschewitz 5310 21 Jul 1974 Lonicera periclymenum L. WTU-266453 U.S.A., WASHINGTON, PACIFIC COUNTY: South side of Route 4, circa 0.3 road miles east of junction with Route 401, Naselle River basin, Naselle. Caprifoliaceae Elev. 32 ft. 46° 22.6' N, 123° 47.7' W; T10N R9W S4 Lonicera conjugialis Kellogg Adventive in thickets, full sun, by beaver wetland. White flowers fade purplish, fruit bright red juicy berry; climbing vine, common escape U.S.A., OREGON, KLAMATH COUNTY: from cultivation in area. Phenology: Flowers. On west side of Crater Lake. Elev. 5000 ft. P. F. Zika 19218 1 Oct 2003 42.941677°, -122.144249°; WGS 84, uncertainty: 5000 m., with Cathy Maxwell Source: Digital Map, Georef'd by Joseph Stormer Lower alpine meadows. Phenology: Fruits. WTU-356444 J. William Thompson 12265 23 Jul 1935 WTU-33115 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera periclymenum L. U.S.A., IDAHO, IDAHO COUNTY: U.S.A., WASHINGTON, KING COUNTY: Clearwater River. North half of Mercer Slough, Bellevue. 46.0225°, -115.89°; WGS 84, uncertainty: 10000 m., Source: Elev. 16 ft. GeoLocate, Georef'd by WTU Staff T24N R5E S5 E 1/2; NAD 27, uncertainty: 805 m., Source: Arborvitae-hemlock zone. Phenology: Flowers. Origin: Native. TRS2LL, Georef'd by Ben Legler Local adventive, damp ground thickets. Twining, red fruit. R. F. Daubenmire 37193 7 May 1937 Phenology: Flowers. WTU-271539 P. Zika 14546 6 Oct 1999 with F. Weinmann Caprifoliaceae WTU-342080 Lonicera conjugialis Kellogg Caprifoliaceae U.S.A., OREGON, KLAMATH COUNTY: On west side of Crater Lake. Lonicera involucrata (Richardson) Banks ex Spreng. Elev. 5000 ft. 42.941677°, -122.144249°; WGS 84, uncertainty: 5000 m., U.S.A., WASHINGTON, YAKIMA COUNTY: Source: Digital Map, Georef'd by Joseph Stormer Tieton Ranger Station, forks of Bethel Ridge Road. Lower alpine meadows. Phenology: Flowers & Fruits. Elev. 2400 ft. 46.75833°, -120.97528°; WGS 84, uncertainty: 10000 m., Source: J. William Thompson 12265 23 Jul 1935 GeoLocate, Georef'd by WTU Staff Along creeks; deep sandy soil; roadside; associated plants: willow. WTU-4099JWT Fairly common. Phenology: Fruits. Origin: Native. E. H. Reid 70 21 Jun 1936 Caprifoliaceae WTU-322723 Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, OKANOGAN COUNTY: Caprifoliaceae Valley of Myers Creek, Chesaw. Elev. 3000-4500 ft. Lonicera periclymenum L. 48.946236°, -119.049966°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., OREGON, LANE COUNTY: Damp mossy woods. Phenology: Flowers & Fruits. Origin: Native. In the Coast Range along State Highway 36, just east of milepost 32, 1.7 miles east of Blachly. Harold St. John 5227 4 Jul 1921 Elev. 747 ft. with W. D. Courtney, Charles S. Parker T16S R7W S11; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler WTU-112822 Woody vine climbing in a thicket of brambles and willows with Rubus discolor, Rubus laciniatus, Salix, Phalrais. Flowers fragrant, varying from pinkish to yellow to white. Phenology: Flowers. Caprifoliaceae Richard R. Halse 5223 9 Jun 1997 Lonicera conjugialis Kellogg WTU-337485 U.S.A., OREGON, JOSEPHINE COUNTY: Cliffs above Bolan Lake, Siskiyou Mountains, near the California border. Caprifoliaceae Elev. 6000 ft. Crevices of rocks, cliffs. Phenology: Flowers & Fruits. Lonicera involucrata (Richardson) Banks ex Spreng. C. L. Hitchcock 5252 7 Jul 1939 U.S.A., IDAHO, VALLEY COUNTY: with J. S. Martin Big Creek Summit, 17 miles east of Cascade. 44.624284°, -115.80231°; WGS 84, uncertainty: 1000 m., Source: WTU-53555 GeoLocate, Georef'd by WTU Staff Rocky hillside, open woods. Phenology: Flowers & Fruits. Origin: Native. Caprifoliaceae William H. Baker 11791 12 Jul 1954 Lonicera involucrata (Richardson) Banks ex Spreng. WTU-297787 U.S.A., MONTANA, DEER LODGE COUNTY: Anaconda; Pintlar Range; east slope above Hicks Lake. Elev. 8850 ft. Between big boulders. Flowering in unbranched shoots, less than one foot high. Phenology: Flowers & Fruits. Origin: Native. Klaus H. Lackschewitz 6325 14 Aug 1975 WTU-266474 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, VALLEY COUNTY: U.S.A., IDAHO, ELMORE COUNTY: Along Mormon Creek at the Junction of the south fork of the Salmon Big Trinity Lake, Trinity Mountains. River, 10 miles south of Warm Lake. 43.62417°, -115.42722°; WGS 84, uncertainty: 10000 m., Source: 44.508514°, -115.666071°; WGS 84, uncertainty: 1000 m., GeoLocate, Georef'd by WTU Staff Source: GeoLocate, Georef'd by WTU Staff Edge of meadow. Phenology: Flowers & Fruits. Origin: Native. Wet seepage slope, shady woods. Phenology: Flowers. Origin: Native. William H. Baker 11198 27 Aug 1953 William H. Baker 11811 13 Jul 1954 WTU-297783 WTU-297786 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera periclymenum L. U.S.A., WASHINGTON, KING COUNTY: North Seattle. CANADA, BRITISH COLUMBIA: 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: Queen Charlotte Islands; 2.5 miles south of Jungle Beach between GeoLocate, Georef'd by WTU Staff Tlell and Skidegate, Graham Island; an escape at Port Clements, In a damp thicket near road. Phenology: Flowers & Fruits. Origin: Jumdis Creek and Queen Charlotte City. Native. Climbing over walls and roof of abandoned homestead. Phenology: Flowers. W. J. Eyerdam s.n. Jun 1933 J. A. Calder 36678 8 Aug 1964 WTU-292973 with R. L. Taylor WTU-241041 Caprifoliaceae Lonicera periclymenum L. Caprifoliaceae U.S.A., OREGON COUNTY: Lonicera utahensis S. Watson Florence. Thicket. Well established. Phenology: Flowers. U.S.A., IDAHO, LATAH COUNTY: Thayuna Hills. Morton E. Peck 13538 1 Aug 1924 Forest Openings Probable Climax: Thuja Plicata. Phenology: Fruits. Origin: Native. WTU-19319JWT R. F. Daubenmire 37515 10 Jul 1937 Caprifoliaceae WTU-271538 Lonicera utahensis S. Watson Caprifoliaceae U.S.A., IDAHO, IDAHO COUNTY: At headwaters of Red River, Red River Hot Springs. Lonicera involucrata (Richardson) Banks ex Spreng. 45.809081°, -115.473482°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., IDAHO, ADAMS COUNTY: Creek bottom. Phenology: Fruits. Origin: Native. 6 miles north of Cuprum. T21N R3W; WGS 84, uncertainty: 6000 m., Source: GeoLocate, William H. Baker 12411 11 Aug 1954 Georef'd by Joseph Stormer Moist ground about spring. Phenology: Flowers & Fruits. Origin: WTU-297991 Native. William H. Baker 11328 3 Sep 1953 Caprifoliaceae WTU-297785 Lonicera utahensis S. Watson U.S.A., IDAHO, SHOSHONE COUNTY: Caprifoliaceae West of Orphan Peak near Buzzard's Roost Junction. In climax forest dominated by Abies Lasiocarpa, Tsuga mertensiana, Lonicera involucrata (Richardson) Banks ex Spreng. and Xerophyllum. Phenology: Vegetative. Origin: Native. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: R. F. Daubenmire 46327 17 Aug 1946 Near Hoquiam. 46.98111°, -123.88806°; WGS 84, uncertainty: 5000 m., Source: WTU-271505 GeoLocate, Georef'd by WTU Staff Marshy ground. Phenology: Flowers. Origin: Native. Sister Mary Milburge 1377 15 May 1937 WTU-10917JWT Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., WASHINGTON, KING COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Seward Park, Seattle. Elk Mountain. 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: 45.91278°, -115.06111°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Moist bank. Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. J. William Thompson 5971 11 Apr 1931 J. H. Christ 51-602 15 Jul 1951 WTU-19375JWT WTU-294433

Caprifoliaceae Caprifoliaceae Lonicera bracteosa Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, KING COUNTY: Renton. Seattle. 47.48306°, -122.21583°; WGS 84, uncertainty: 10000 m., Source: 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., GeoLocate, Georef'd by WTU Staff Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers. Origin: Native. Phenology: Flowers & Fruits. Origin: Native. W. V. Lovitt s.n. 16 Apr 1910 E. S. Meany s.n. 1898 WTU-16474 WTU-37172

Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., MONTANA, FLATHEAD COUNTY: U.S.A., WASHINGTON, KING COUNTY: East bank of South fork of the Kootenai River, 0.5 miles south of the Seattle. Highway #2. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., At edge of woods. Plant 6 decimeters tall. Phenology: Vegetative. Source: Digital Map, Georef'd by Joseph Stormer Phenology: Flowers & Fruits. Origin: Native. C. L. Hitchcock 24028 11 Jul 1965 E. S. Meany s.n. 1898 WTU-224163 WTU-37176 Caprifoliaceae Caprifoliaceae Lonicera pileata Oliv. Lonicera sp. U.S.A., WASHINGTON, CLALLAM COUNTY: East end of Lake Crescent, Vicinity Storm King Ranger Station, U.S.A., WASHINGTON, KING COUNTY: Olympic Peninsula. Alki Point, Seattle. Elev. 640 ft. 47.57639°, -122.40861°; WGS 84, uncertainty: 10000 m., Source: T30N R9W S28 SE 1/2; NAD 27, uncertainty: 805 m., Source: GeoLocate, Georef'd by WTU Staff TRS2LL, Georef'd by Ben Legler Phenology: Flowers & Fruits. Scattered in clearings and woods, associated with Tsuga heterophylla, Abies grandis, Alnus rubra, Acer macrophyllum, Rubus Emma A. Shumway s.n. 17 May 1892 ursinus. Purple berries, escaped from cultivation, deer-browsed shrubs, bird-sown, 2 meters tall. Phenology: Flowers. WTU-110069 P. F. Zika 13408 22 Nov 1999 Caprifoliaceae WTU-342458 Lonicera utahensis S. Watson Caprifoliaceae U.S.A., IDAHO, SHOSHONE COUNTY: Sunset Peak. Lonicera americana 47.56333°, -115.83361°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., WASHINGTON, KING COUNTY: Phenology: Flowers. Origin: Native. Seattle. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., J. H. Christ 51-365 8 Jul 1951 Source: Digital Map, Georef'd by Joseph Stormer Woods. S - 297. Phenology: Flowers. WTU-296289 Geo. B. Rigg s.n. 26 May 1908 WTU-20316 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, SNOHOMISH COUNTY: Seattle. Everett. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., 47.97917°, -122.20083°; WGS 84, uncertainty: 10000 m., Source: Source: Digital Map, Georef'd by Joseph Stormer GeoLocate, Georef'd by WTU Staff Phenology: Flowers & Fruits. Origin: Native. [No collection date given on label]. Phenology: Flowers. Origin: Native. Emma A. Shumway 1535 May 1892 W. Bailey s.n. WTU-11930 WTU-16485 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., WASHINGTON, KING COUNTY: Seattle. U.S.A., IDAHO, BONNER COUNTY: 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Luby Bay, Priest Lake. Source: Digital Map, Georef'd by Joseph Stormer T60N R5W S24; WGS 84, uncertainty: 10000 m., Source: Phenology: Fruits. Origin: Native. GeoLocate, Georef'd by WTU Staff Woods. Phenology: Flowers. Origin: Native. Chas. V. Piper 120 20 May 1888 William H. Baker 13611 28 Jul 1955 WTU-16480 WTU-296117 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, SHOSHONE COUNTY: Summit of Glidder Pass, Idaho State Boundary, 5 miles east of U.S.A., IDAHO, VALLEY COUNTY: Burke. 3 miles north of Upper Payette Lake, Squaw Meadows. 47.52023°, -115.712001°; WGS 84, uncertainty: 500 m., Source: 45.154553°, -116.02556°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Thickets, in Spruce-Fir woods. Phenology: Flowers. Origin: Native. Thickets. Phenology: Flowers. Origin: Native. William H. Baker 13527 18 Jul 1955 W. H. Baker 10331 9 Jul 1953 WTU-296053 WTU-296043 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., WASHINGTON, KING COUNTY: Seattle. U.S.A., IDAHO, ADAMS COUNTY: 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Above the north side of Payette Lake. Source: Digital Map, Georef'd by Joseph Stormer On slope; growing in very brushy undergrowth in Pine forest. Phenology: Flowers & Fruits. Origin: Native. Phenology: Flowers. Origin: Native. C. V. Piper 120 20 May 1888 Sidney C. Beinke 482 29 May 1938 WTU-16479 WTU-194874

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: U.S.A., MONTANA, RAVALLI COUNTY: Near Aloha. 4 to 5 miles along Watchtower Creek Trail from its junction with road 47.201944°, -124.1525°; NAD 27, uncertainty: 3000 m., Source: #473 about 20 miles west of Highway 93, 65 miles south of Georeferenced, Georef'd by WTU Staff Missoula, Bitterroot Mountains, Bitterroot River drainage. Along roadside. Phenology: Flowers & Fruits. Origin: Native. Phenology: Flowers. Origin: Native. Harold E. Helmrich 135 9 Jul 1933 Patrick Elvander 697 12 Jun 1976 WTU-306007 WTU-267761 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, LATAH COUNTY: U.S.A., IDAHO, LATAH COUNTY: East of Troy. East Moscow Mountain. 46.73694°, -116.76861°; WGS 84, uncertainty: 10000 m., Source: 46.80472°, -116.83889°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff GeoLocate, Georef'd by WTU Staff Opening in pinus contorta forest probable climax; Abies Grandis. Woods and thickets. Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. William H. Baker 5925 22 May 1949 R. F. Daubenmire 3853 6 May 1938 WTU-302099 WTU-271513 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, LATAH COUNTY: U.S.A., OREGON COUNTY: Moscow Mountain, ca. [circa] five miles northeast of Moscow. Eagle Cap Wilderness, Wallowa Whitman National Forest; adjacent 46.7325°, -116.99917°; WGS 84, uncertainty: 10000 m., Source: to Echo Lake. GeoLocate, Georef'd by WTU Staff Elev. 7400 ft. Woods. Phenology: Flowers. Origin: Native. On rocky north slope. Associated with Abies lasiocarpa, Phyllodoce empetriformis. Uncommon. Phenology: Flowers. Origin: Native. William H. Baker 8955 29 May 1952 Brenda Bafus 327 21 Jul 1979 WTU-302098 WTU-278653 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, KOOTENAI COUNTY: One mile northeast of Chilco. U.S.A., IDAHO, IDAHO COUNTY: 47.86389°, -116.74556°; WGS 84, uncertainty: 10000 m., Source: Summit of Pilot Knob. GeoLocate, Georef'd by WTU Staff Elev. 7100 ft. Forming thickets, on rocky hillside. Phenology: Flowers. Origin: T30N R6E S28; WGS 84, uncertainty: 10000 m., Source: Native. GeoLocate, Georef'd by WTU Staff Among rocks. Shrub 1- 2 feet high. Phenology: Flowers. Origin: William H. Baker 7361 2 May 1951 Native. WTU-302097 Winnifred Newell Elwood 38 13 Jul 1947 WTU-121225 Caprifoliaceae Lonicera utahensis S. Watson Caprifoliaceae U.S.A., IDAHO, VALLEY COUNTY: Lonicera utahensis S. Watson At summit of divide between Warm Lake and Cascade. 44.51639°, -116.04083°; WGS 84, uncertainty: 10000 m., Source: U.S.A., OREGON, WALLOWA COUNTY: GeoLocate, Georef'd by WTU Staff South Fork of the Lostine River, 18 miles southeast of Lostine. Near Stream. Phenology: Flowers & Fruits. Origin: Native. 45.227376°, -117.364494°; WGS 84, uncertainty: 18000 m., Source: Digital Map, Georef'd by Joseph Stormer C. L. Hitchcock 14015 26 Jun 1946 Shady woods. Phenology: Flowers. Origin: Native. with C. V. Muhlick William H. Baker 11482 4 Jun 1954 WTU-105103 WTU-196230 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, LATAH COUNTY: Bald Mountain. U.S.A., IDAHO, LATAH COUNTY: T43N R2W; WGS 84, uncertainty: 10000 m., Source: GeoLocate, [No locality given on label] Georef'd by WTU Staff 46.811276°, -116.691521°; uncertainty: 43000 m. In cutover forest associated with Larix, Pseudotsuga, and Pinus on In woods. Phenology: Fruits. Origin: Native. the northern slope. Phenology: Flowers. Origin: Native. C. V. Piper s.n. 4 Jul 1893 E. L. Richards 137 26 May 1956 WTU-16789 WTU-296247 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, IDAHO COUNTY: U.S.A., WASHINGTON, YAKIMA COUNTY: Below Heaves Gate, Seven Devils Mountains. (YIN) Along north Satus Road at 12 miles southeast Toppenish. 45.30917°, -116.55389°; WGS 84, uncertainty: 10000 m., Source: Elev. 705 ft. GeoLocate, Georef'd by WTU Staff 46.254514°, -120.129435°; WGS 84, uncertainty: 500 m., Source: Steep rocky banks. Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff Yard; upper sonoran house yard. Phenology: Flowers. Origin: William H. Baker 12086 21 Jul 1954 Native. WTU-298008 E. Hunn 728 9 Aug 1977 WTU-271080 Caprifoliaceae Caprifoliaceae Lonicera utahensis Wats. f. ebractulata (Rydb.) St. J. Lonicera utahensis S. Watson U.S.A., IDAHO, SHOSHONE COUNTY: Little Baldy Lookout. U.S.A., IDAHO COUNTY: T50N R5E S8 SE 1/4; WGS 84, uncertainty: 10000 m., Source: Near south shore of Priest Lake. GeoLocate, Georef'd by WTU Staff Elev. 3000 ft. Phenology: Flowers. In rather deep shade of lodgepole forest. Phenology: Flowers. Origin: Native. Jean G. Witt s.n. 26 Jun 1946 C. L. Hitchcock 2913 14 May 1936 WTU-119183 WTU-6213JWT Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, VALLEY COUNTY: Warm Lake Summit. U.S.A., WASHINGTON, GARFIELD COUNTY: Elev. 7286 ft. Umatilla National Forest, summit of Diamond Peak. 44.64639°, -115.66889°; WGS 84, uncertainty: 10000 m., Source: Elev. 6376 ft. GeoLocate, Georef'd by WTU Staff 46° 5' N, 117° 35' W; T7N R42E S3; WGS 84, uncertainty: 275 Woods. Phenology: Flowers. Origin: Native. m., Source: GeoLocate, Georef'd by Richard Olmstead Summit with open rocky meadow dominated by Artemisia tridentata; William H. Baker 11750 10 Jul 1954 krummholz Abies lasiocarpa also present; west of summit, small patch of woods - mix of Abies lasiocarpa, Pseudotsuga menziesii, WTU-297989 Picea engelmannii, Pinus contorta; eastern edge of wooded area. Paired light yellow flowers. Phenology: Flowers. Origin: Native. Caprifoliaceae Tracy Fuentes 38 28 Jun 1997 Lonicera utahensis Wats. f. ebractulata (Rydb.) St. J. WTU-339018 U.S.A., MONTANA, SWEET GRASS COUNTY: At Four Mile Ranger Station, Boulder Canyon. Caprifoliaceae 45.341419°, -110.233338°; WGS 84, uncertainty: 1000 m., Source: Digital Map, Georef'd by Joseph Stormer, Invalid Lonicera utahensis S. Watson coordinates (located outside county bounds). In grass pasture. [Collector number Suffix written in pencil over U.S.A., IDAHO, LATAH COUNTY: typed suffix. Difficult to is either e or l]. Phenology: Flowers & Fruits. 5 miles south of Deary. 46.727057°, -116.555°; WGS 84, uncertainty: 500 m., Source: C. L. Hitchcock 16335E 11 Jul 1947 GeoLocate, Georef'd by WTU Staff Shrub 1 inch tall. Phenology: Flowers. Origin: Native. WTU-115540 Marion Ownbey 2017 14 Apr 1940 with Ruth P. Ownbey Caprifoliaceae WTU-73618 Lonicera utahensis S. Watson U.S.A., IDAHO, IDAHO COUNTY: Caprifoliaceae Headwaters of Salamander Creek, 3 miles northwest of Salmon Mountain, Bitterroot National Forest. Lonicera utahensis S. Watson T27N R13E S; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., WASHINGTON COUNTY: Talus slope. Phenology: Fruits. Origin: Native. 14 miles north of Metaline Falls. Elev. 2600 ft. William H. Baker 12675 17 Aug 1954 Open woods near highway. Phenology: Flowers. Origin: Native. WTU-297992 C. Leo Hitchcock 2929 15 May 1936 WTU-6215JWT Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., MONTANA, BEAVERHEAD COUNTY: U.S.A., WASHINGTON, GARFIELD COUNTY: Big Hole River Canyon, [circa] 15 miles west of Wisdom on North rim of Tucannon River canyon, 5 miles south of Clearwater Gibbonsville Pass Road. Ranger Station, Blue Mountains. Wet meadow. Phenology: Flowers. Origin: Native. Elev. 4500 ft. 46.13994°, -117.52036°; WGS 84, uncertainty: 1000 m., Source: C. L. Hitchcock 9193 22 Jun 1944 GeoLocate, Georef'd by WTU Staff with C. V. Muhlick Small shrub, edge of woods. Phenology: Flowers. Origin: Native. WTU-92850 L. Constance 1752 13 Jun 1936 with H. F. Clements Caprifoliaceae WTU-5733JWT Lonicera utahensis S. Watson Caprifoliaceae U.S.A., WASHINGTON, WHATCOM COUNTY: Mount Baker National Forest; Church Mountain. Lonicera utahensis S. Watson 48.92694°, -121.88083°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff U.S.A., WASHINGTON, OKANOGAN COUNTY: On cliff near timberline. Phenology: Flowers. Origin: Native. Bonaparte Lake. Elev. 3700 ft. W. C. Muenscher 9844 9 Jun 1939 48.79333°, -119.06°; WGS 84, uncertainty: 1600 m., Source: GeoLocate, Georef'd by WTU Staff WTU-56048 Evergreen woods. Phenology: Flowers. Origin: Native. Harold St. John 5286 8 Jul 1921 Caprifoliaceae with W. D. Courtney, Charles S. Parker Lonicera utahensis S. Watson WTU-88660 CANADA, BRITISH COLUMBIA: Sunup Cabin, southwest of Mount Baldy, northwest of Ross Spur. Caprifoliaceae Elev. 4200-4500 ft. Open coniferous forest, decomposed granite soil; nearly level. Lonicera utahensis S. Watson Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, OKANOGAN COUNTY: George G. Hedgcock s.n. 14 Jun 1939 At Salmon Meadows, above Conconully. 48.65556°, -119.83806°; WGS 84, uncertainty: 5000 m., Source: WTU-102267 GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. Caprifoliaceae Chas. B. Fiker 1274 18 Jun 1933 Lonicera utahensis S. Watson WTU-19357JWT U.S.A., MONTANA COUNTY: About 14 miles north of Metaline Falls. Caprifoliaceae Elev. 2600 ft. Open woods near highway. Phenology: Flowers. Origin: Native. Lonicera utahensis S. Watson C. Leo Hitchcock 2929 15 May 1936 U.S.A., IDAHO, VALLEY COUNTY: Upper Payette Lake. WTU-37714 Elev. 5500 ft. 44.91167°, -116.125°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Caprifoliaceae On the bank just above high water mark. Lodgepole pine forest. Coarse sandy-gravelly soil with thin humus layer. With Vaccinium Lonicera utahensis S. Watson globulare, V. scoparium. Phenology: Fruits. Origin: Native. U.S.A., OREGON, UMATILLA COUNTY: C. Davidson 1614 24 Jun 1972 Umatilla National Forest, at Morning Creek Sno Park. Elev. 4851 ft. WTU-259900 45° 46' N, 118° 4' W Associated genera: Ribes, Pinus, Carex, Sambucus, Polemonium. Flowers white shrub 1 meter tall. Phenology: Flowers. Origin: Native. Caprifoliaceae Richard R. Halse 6196 18 Jun 2002 Lonicera utahensis S. Watson WTU-356283 U.S.A., OREGON, WALLOWA COUNTY: Wallowa Mountains. Bowman Trail, about 15 miles south of Lostine. Elev. 5500 ft. With Spiraea, Calochortus and Pachistima in moist shady coniferous woods. Shrub 6 decimeters tall; berries red. Phenology: Fruits. Origin: Native. Georgia Mason 8982 6 Aug 1971 WTU-257860 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO COUNTY: U.S.A., IDAHO, VALLEY COUNTY: Spirit Lake. Above head of Payette Lake. Woods. Phenology: Flowers. Origin: Native. 44.91167°, -116.125°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff Sister Mary Milburge s.n. Apr 1931 Rocky slopes. Phenology: Flowers. Origin: Native. WTU-30762 C. L. Hitchcock 8548 30 May 1944 with C. V. Muhlick Caprifoliaceae WTU-93740 Lonicera utahensis S. Watson Caprifoliaceae U.S.A., IDAHO, BONNER COUNTY: Hagen Lake Bog, trail to Bismark Mountain, west of Kalispel Creek, Lonicera utahensis S. Watson Priest Lake. 48.59528°, -116.99611°; WGS 84, uncertainty: 10000 m., Source: U.S.A., MONTANA, FERGUS COUNTY: GeoLocate, Georef'd by WTU Staff Half Moon Canyon, 3.5 miles from mouth, Big Snowy Mountains. Woods. Phenology: Flowers & Fruits. Origin: Native. 46.800083°, -109.311897°; WGS 84, uncertainty: 3500 m., Source: Digital Map, Georef'd by Joseph Stormer William H. Baker 16603 5 Jun 1964 Leaves glabrous. Phenology: Flowers. Origin: Native. WTU-261815 C. L. Hitchcock 12003 5 Jul 1945 with C. V. Muhlick Caprifoliaceae WTU-99714 Lonicera utahensis S. Watson Caprifoliaceae U.S.A., OREGON, WALLOWA COUNTY: Wallowa Mountains, about 1-1.5 miles up the Bowman Trail south of Lonicera utahensis S. Watson Lostine. Elev. 5600 ft. U.S.A., IDAHO, IDAHO COUNTY: With Vaccinium Ceanothus and Spiraea in moist shady coniferous West of Lolo Pass. woods. Shrub about 1 meter tall. Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. Georgia Mason 8791 28 Jun 1971 Fred A. Barkley 2412 27 May 1938 WTU-258177 WTU-51558

Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, LATAH COUNTY: U.S.A., IDAHO, KOOTENAI COUNTY: [No locality given on label] Coeur d'Alene. 46.811276°, -116.691521°; uncertainty: 43000 m. 47.67778°, -116.77944°; WGS 84, uncertainty: 10000 m., Source: Phenology: Fruits. Origin: Native. GeoLocate, Georef'd by WTU Staff Phenology: Flowers. Origin: Native. C. V. Piper 1719 21 Jun 1894 Unknown s.n. Apr 1912 WTU-37165 WTU-16769 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., MONTANA COUNTY: Hyalite Creek. U.S.A., IDAHO, LATAH COUNTY: Habitat damp wood. Phenology: Fruits. Origin: Native. [No locality given on label] T40N R5W S14; NAD 27, uncertainty: 805 m., Source: TRS2LL, D. B. Swingle s.n. 24 Aug 1933 Georef'd by Ben Legler Moist burned-over ground. Abies-Thuja-Pseudotsuga forest. WTU-37098 Canadian Life Zone. Phenology: Fruits. Origin: Native. Louis A. Dillion 663 22 Jun 1936 with Louis A. Dillon WTU-52023 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO COUNTY: U.S.A., MONTANA, LAKE COUNTY: Hope Idaho. High above and several miles southeast of McDonald Lake; Mission Copses. [4214 written in pen in upper right corner] Herbarium, Range. University of Washington Presented by C. E. Burglehaus. On cliffs. Phenology: Flowers. Origin: Native. Phenology: Flowers. Origin: Native. C. L. Hitchcock s.n. 12 Jul 1948 N.A.S. s.n. May 1893 WTU-125448 WTU-16771 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson CANADA, BRITISH COLUMBIA: U.S.A., IDAHO, NEZ PERCE COUNTY: Between Cascade and Rossland. [No locality given on label] Elev. 4000 ft. Copses. [4214 written in pen in upper right corner]. Phenology: 49.040529°, -118.0093°; WGS 84, uncertainty: 14442 m., Source: Flowers. Origin: Native. Digital Map, Georef'd by Joseph Stormer Crest of high mountain. Legit: J. W. Thompson. Phenology: Fruits. N.A.S. 221 May 1892 Origin: Native. WTU-16770 J. W. Thompson 14412 18 Jul 1939 WTU-57169 Caprifoliaceae Lonicera utahensis S. Watson Caprifoliaceae U.S.A., IDAHO COUNTY: Lonicera utahensis S. Watson Near south shore of Priest Lake. Elev. 3000 ft. U.S.A., MONTANA, GALLATIN COUNTY: In rather deep shade of lodge-pole forest. Phenology: Flowers. Spring Hill Canyon. Origin: Native. Elev. 5300 ft. Moist open woods. [Collection year difficult to read]. Phenology: C. L. Hitchcock 2913 14 May 1936 Fruits. Origin: Native. WTU-38096 D. B. Swingle s.n. 16 Jul 1934 WTU-43903 Caprifoliaceae Lonicera utahensis S. Watson Caprifoliaceae U.S.A., MONTANA, PARK COUNTY: Lonicera sp. At the falls of the Clarks Fork of the Yellowstone River, about 4 miles east of Cooke City. U.S.A., IDAHO, VALLEY COUNTY: Glaciated granite knobs between the dam and the old sawmill. Deep canyon on Gold Fork Lookout, Payette National Forest, Phenology: Fruits. Origin: Native. Sawtooth Mountains. Elev. 7000 ft. Jean G. Witt 1357 8 Aug 1948 44.67194°, -115.83667°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-129062 Phenology: Fruits. J. William Thompson 13906 19 Jul 1937 Caprifoliaceae WTU-38709 Lonicera utahensis S. Watson U.S.A., MONTANA, MISSOULA COUNTY: Caprifoliaceae 1/2 mile east of Upper Holland Lake, Flathead Range, Flathead National Forest. Lonicera utahensis S. Watson Phenology: Flowers. Origin: Native. CANADA, BRITISH COLUMBIA: C. L. Hitchcock 18464 15 Jul 1948 Cheakamus River Trail, Garibaldi Park. Elev. 2500 ft. WTU-125239 Garibaldi Park. Chekamus River Trail. Herb. M.B.G. 903,327 (Wyoming) Same as 4185 [Written above collection label, in pencil:Garman 118] [Written to left of collection label in pencil: 9] [Written to left of collection label in pencil: 271]. Phenology: Fruits. Origin: Native. F. Comte 5514 11 Aug 1967 WTU-318963 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, IDAHO COUNTY: U.S.A., OREGON, UNION COUNTY: Fog Mountain Saddle, above Selway River Falls 5800-6200 feet or 200 yards south of E. E. [East Eagle] Creek Falls. more. Elev. 4600 ft. Elev. 5800-6200 ft. T5S R44E S35; NAD 27, uncertainty: 805 m., Source: TRS2LL, 46.051227°, -115.220657°; WGS 84, uncertainty: 10000 m., Georef'd by Ben Legler Source: GeoLocate, Georef'd by WTU Staff Good #. [as printed on label] south, semi open slope. Wet, good Phenology: Flowers. Origin: Native. depth soil, with Vaccinium, Acer, Pyrola, Abies grandis, Sorbus, Anemone. Distributed by Oregon State University Herbarium. Brian O. Mulligan s.n. 18 Jun 1980 Phenology: Flowers. Origin: Native. WTU-281662 S. C. Head 871 15 Jun 1957 WTU-224159 Caprifoliaceae Lonicera utahensis S. Watson Caprifoliaceae U.S.A., MONTANA COUNTY: Lonicera utahensis S. Watson Teepee Ridge, 2 miles south of Wagon Mountain Road; 25 miles southwest of Missoula, Montana. U.S.A., MONTANA, FLATHEAD COUNTY: Elev. 5200 ft. Along South Fork Flathead River, 1/4 mile west of Hungry Horse. Open forest slope with mixed conifers. Scare. Phenology: Fruits. In deep woods. Shrub about 2 feet tall. Phenology: Fruits. Origin: Origin: Native. Native. Peter F. Stickney 680 1 Jul 1962 C. L. Hitchcock 24500 4 Jul 1966 WTU-237517 WTU-232202

Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, IDAHO COUNTY: U.S.A., MONTANA, MISSOULA COUNTY: Fifteen miles south of Grangeville. At west end of Upper Holland Lake, Swan Range, about 80 miles 45.76583°, -115.48889°; WGS 84, uncertainty: 10000 m., Source: northeast of Missoula. GeoLocate, Georef'd by WTU Staff Elev. 7000 ft. Clearings in fir forest. Phenology: Flowers. Origin: Native. T20N R15W S29; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler C. L. Hitchcock 23508 25 May 1964 In low ground at west. Shrub 1-2 meters tall. Flowers ochroleucous or nearly white. Phenology: Flowers. Origin: Native. WTU-218095 Arthur Cronquist 7909 9 Jul 1955 Caprifoliaceae WTU-162517 Lonicera utahensis S. Watson Caprifoliaceae U.S.A., MONTANA, MEAGHER COUNTY: Harley Park, circa 6 miles west of Neihart, Little Belt Mountains. Lonicera utahensis S. Watson 46.927562°, -110.847355°; uncertainty: 3200 m. Plants circa 1 foot tall; leaves pilose beneath and ciliate. Phenology: U.S.A., IDAHO, CLEARWATER COUNTY: Flowers. Origin: Native. Several miles west of Elk River. T40N R2E; WGS 84, uncertainty: 10000 m., Source: GeoLocate, C. L. Hitchcock 12305 13 Jul 1945 Georef'd by WTU Staff with C. V. Muhlick In open second-growth coniferous woods. Phenology: Vegetative. Origin: Native. WTU-99715 Arthur Cronquist 5890 23 Jun 1949 Caprifoliaceae WTU-169579 Lonicera utahensis S. Watson Caprifoliaceae U.S.A., MONTANA, SWEET GRASS COUNTY: By spring Half Moon Camp, Big Timber Creek, Crazy Mountains. Lonicera utahensis S. Watson In deep shade. Berries deep red, leaves pilose beneath as well as ciliate. Phenology: Fruits. Origin: Native. U.S.A., OREGON, WALLOWA COUNTY: At Horseshoe Lake, Lake Basin, Wallowa Mountains. C. L. Hitchcock 13199 4 Aug 1945 Phenology: Flowers. Origin: Native. with C. V. Muhlick A. R. Kruckeberg 2404 21 Jul 1950 WTU-100961 WTU-181174 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis Wats. f. ebractulata (Rydb.) St. J. U.S.A., MONTANA, PARK COUNTY: CANADA, BRITISH COLUMBIA: Vicinity Cooke Guard Station, about 2 miles east of Cooke City. One mile southwest of South Slocan; Nelson-Trail Highway. Elev. 8000 ft. Occasional at edge of mixed woods. Shrub to 1 1/2 feet; fruit red. 45.025336°, -109.877994°; WGS 84, uncertainty: 2000 m., Phenology: Fruits. Source: Digital Map, Georef'd by Joseph Stormer Rocky knobs. Phenology: Flowers. Origin: Native. J. A. Calder 9440 20 Jun 1953 with D. B. O. Savile Jean G. Witt 1670 26 Jun 1949 WTU-187030 WTU-178813 Caprifoliaceae Caprifoliaceae Lonicera utahensis Wats. f. ebractulata (Rydb.) St. J. Lonicera utahensis S. Watson U.S.A., IDAHO, IDAHO COUNTY: U.S.A., MONTANA, GRANITE COUNTY: Along French Creek, 2 miles south of Salmon River, 20 miles east of Skalkaho Pass. Riggins. Elev. 7200 ft. T24N R3E; WGS 84, uncertainty: 1000 m., Source: GeoLocate, 46.244419°, -113.771014°; WGS 84, uncertainty: 988 m., Source: Georef'd by WTU Staff Digital Map, Georef'd by Joseph Stormer Among granite boulders. Shrubs 1-2 meters high. Flowers fragrant, Phenology: Flowers. Origin: Native. whitish, weakly tinted with yellowish green and dull lavender. Phenology: Flowers. C. L. Hitchcock 14471 7 Jul 1946 with C. V. Muhlick Arthur Cronquist 6179 21 Apr 1950 WTU-104234 WTU-144272

Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera utahensis S. Watson U.S.A., IDAHO, FREMONT COUNTY: U.S.A., IDAHO, IDAHO COUNTY: Found a few feet from Howard Springs, one mile east of Highway Road to Red River Hot Springs, 7 miles southeast of Elk City. 20, T28N R9E S8; WGS 84, uncertainty: 1000 m., Source: Elev. 6900 ft. GeoLocate, Georef'd by WTU Staff 44° 40' N, 111° 17' 30" W; T16N R44E S32C Along creek, in shade. Shrub to 1.5 meters. Phenology: Flowers. Growing in shaded area on the bank of a mountain spring. Shrub Origin: Native. approximately 4 feet tall. Phenology: Flowers. Origin: Native. Quentin Jones 23 14 Jun 1950 Heather Haile 10 14 Jun 1993 WTU-142813 WTU-330714 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, NEZ PERCE COUNTY: U.S.A., IDAHO, IDAHO COUNTY: [No locality given on label] At headwaters of Red River, Red River Hot Springs. Copses. Phenology: Flowers. Origin: Native. 45.809081°, -115.473482°; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff E. A. Shumway s.n. Jun 1892 Creek bottom. Phenology: Fruits. Origin: Native. WTU-188497 William H. Baker 12398 11 Aug 1954 WTU-298009 Caprifoliaceae Lonicera ciliata (Pursh) Poir. Caprifoliaceae U.S.A., MINNESOTA, CARLTON COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. [No locality given on label] Woods. Presented by E. A. Shumway. Phenology: Flowers. U.S.A., OREGON, WALLOWA COUNTY: 5 miles southeast of Lostine along the South Fork of Lostine River. unknown s.n. Jun 1891 Streamside. Phenology: Flowers & Fruits. Origin: Native. WTU-188499 William H. Baker 11505 4 Jun 1954 WTU-298007 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. CANADA, BRITISH COLUMBIA: U.S.A., IDAHO, LATAH COUNTY: Mountains between Cascade and Rossland. Circa 5 miles east of Moscow. Moist woods. Phenology: Fruits. Origin: Native. 46.732451°, -116.893562°; WGS 84, uncertainty: 500 m., Source: GeoLocate, Georef'd by WTU Staff J. William Thompson 14426 19 Jul 1939 Moist thickets. Phenology: Flowers. Origin: Native. WTU-57148 William H. Baker 6628 20 May 1950 WTU-302100 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., OREGON, DESCHUTES COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Just east of Little Cultus Lake. Elev. 4800 ft. U.S.A., IDAHO, BEAR LAKE COUNTY: Small meadow. Corolla yellow. Phenology: Flowers & Fruits. Origin: Along Saint Charles Creek, 3.5 miles west of Saint Charles. Native. T15S R43E S20; WGS 84, uncertainty: 1000 m., Source: GeoLocate, Georef'd by WTU Staff C. L. Hitchcock 4922 25 Jun 1939 Thickets; douglas fir zone. Phenology: Fruits. Origin: Native. with J. S. Martin William H. Baker 9430 22 Jul 1952 WTU-54297 WTU-302095 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., OREGON, CLATSOP COUNTY: Near Skipanon. U.S.A., MONTANA, MISSOULA COUNTY: Phenology: Flowers & Fruits. Origin: Native. Rattlesnake Valley 6 miles north of Missoula. Rich loam soil, north slopes. Phenology: Flowers. Origin: Native. G. E. Morrill 26 15 Jun 1933 Phil Smith 253 4 May 1938 WTU-27796 WTU-51488 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis Wats. f. ebractulata (Rydb.) St. J. U.S.A., IDAHO, LATAH COUNTY: Along the Troy Road 4 miles east of Moscow. U.S.A., IDAHO, VALLEY COUNTY: 46.732469°, -116.914684°; WGS 84, uncertainty: 500 m., Source: Payette lake. GeoLocate, Georef'd by WTU Staff 44.91167°, -116.125°; WGS 84, uncertainty: 10000 m., Source: Thickets. Phenology: Flowers. Origin: Native. GeoLocate, Georef'd by WTU Staff Mounted duplicate from the Herbarium of the University of California. William H. Baker 5868 17 May 1949 The Ira W. Clokey Herbarium. Phenology: Flowers. WTU-302096 R. J. Davis 763 12 May 1939 WTU-98727 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., OREGON COUNTY: Lonicera utahensis S. Watson Crater Lake National Park, near Government Camp. Phenology: Flowers. Origin: Native. U.S.A., IDAHO, NEZ PERCE COUNTY: Craig Mounains, near Lake Waha. A. A. Heller 13475 24 Jul 1920 46.21333°, -116.85111°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff WTU-88464 Woods. Phenology: Flowers. Origin: Native. J. H. Sandberg 221 21 May 1992 Caprifoliaceae with D. T. Macdougal, A. A. Heller Lonicera involucrata (Richardson) Banks ex Spreng. WTU-243784 U.S.A., OREGON COUNTY: Crater Lake National Park; southwest corner of the Park in the Canyon of Redblanket Creek. Phenology: Flowers & Fruits. Origin: Native. William H. Baker 6437 28 Aug 1949 WTU-131409 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, IDAHO COUNTY: U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Above Canteen Meadows, just below Fenn Mountains, Crags Near Aloha. Mountains, Middle fork Selway river ranger district. Nez Perce 47.201944°, -124.1525°; NAD 27, uncertainty: 3000 m., Source: National Forest. Georeferenced, Georef'd by WTU Staff Elev. 7000 ft. Along roadside. Phenology: Flowers & Fruits. Origin: Native. 46.082289°, -115.425324°; WGS 84, uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff Harold E. Helmrich 135 9 Jul 1933 On steep subalpine granitic slopes. Phenology: Flowers. Origin: Native. WTU-306007 A. R. Kruckeberg 4121 8 Jul 1957 Caprifoliaceae WTU-191995 Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., IDAHO, IDAHO COUNTY: Opposite Kamiah. Lonicera utahensis S. Watson 46.22722°, -116.02833°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff, Invalid coordinates (located U.S.A., WASHINGTON, KING COUNTY: outside county bounds). Near west shore of Lake Washington, Washington Park Arboretum. Open woods. Arid Transition Life Zone. Climbing on shrubbery and Elev. 49 ft. trees. Topotype. Phenology: Flowers. Origin: Native. 47° 38' N, 122° 18' W; T25N R4E S21 Rubus spectabilis, Scripus microcarpous, Ranunculus reptans. Fred G. Meyer 926 6 Jun 1937 (equals collector number 19883). Phenology: Flowers. Origin: Native. WTU-17056JWT Peter F. Zika 19417 23 Apr 2004 Caprifoliaceae WTU-358840 Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, CHELAN COUNTY: Lonicera utahensis S. Watson Lake Chelan National Recreation Area. North end of Coon Lake along trail. 1 kilometer northeast of High Bridge in Stehekin River U.S.A., IDAHO, LATAH COUNTY: Valley. Mineral Mountain rest area on U.S. 95 at milepost 371. Elev. 2180 ft. 47.06694°, -116.88194°; WGS 84, uncertainty: 10000 m., Source: 48° 23' 27.4" N, 120° 50' 3.3" W; UTM Zone 10, 660330E, GeoLocate, Georef'd by WTU Staff 5362022N; WGS 84, uncertainty: 9 m., Source: GPS Woody area. This shrub is very abundant in this woody area. Mixed conifer forest with Pseudotsuga, ,Tsuga, Abies, Pinus and Phenology: Flowers & Fruits. Origin: Native. Thuja. Understory of Vaccinium, Smilacina, Disporum and Clintonia. Corolla bright orange. Scattered clambering vines. Phenology: Jenesta Nettles 2 28 Apr 2003 Flowers. Origin: Native. WTU-362821 Barbara Smith 06-24 29 Jun 2006 WTU-366854 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata CANADA, BRITISH COLUMBIA: Maple Ridge Trunk Road. U.S.A., WASHINGTON, KING COUNTY: Roadside meadow. ACC# v177504; students collection for Botany Alki Point, Seattle. 311. Phenology: Flowers & Fruits. 47.57639°, -122.40861°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, Georef'd by WTU Staff J. P. Luitjens 690169 22 Apr 1976 Phenology: Flowers. Origin: Native. with students Emma A. Shumway s.n. 17 May 1892 WTU-362820 WTU-110069 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera involucrata (Richardson) Banks ex Spreng. var. U.S.A., WASHINGTON, KING COUNTY: involucrata West Seattle. 47.60639°, -122.33083°; WGS 84, uncertainty: 10000 m., Source: U.S.A., WASHINGTON, KING COUNTY: GeoLocate, Georef'd by WTU Staff Seattle. Phenology: Fruits. Origin: Native. 47.613617°, -122.310098°; WGS 84, uncertainty: 15750 m., Source: Digital Map, Georef'd by Joseph Stormer W. V. Lovitt s.n. 19 Nov 1909 Woods. Phenology: Flowers. Origin: Native. WTU-16611 Geo. B. Rigg s.n. 26 May 1908 WTU-20316 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. var. Lonicera utahensis S. Watson involucrata U.S.A., IDAHO, SHOSHONE COUNTY: U.S.A., WASHINGTON, KING COUNTY: Clearwater National Forest. Hoodoo Pass, South Trailhead parking Renton. lot. 47.48306°, -122.21583°; WGS 84, uncertainty: 10000 m., Source: Elev. 5993 ft. GeoLocate, Georef'd by WTU Staff 46.97483°, -115.02681°; WGS 84, uncertainty: 500 m., Source: Phenology: Flowers. Origin: Native. Nat. Geog. TOPO!, Georef'd by John Bassett Rocky meadow surrounded by forest of Tsuga, Abies and Picea. W. V. Lovitt s.n. 16 Apr 1910 Artemisia species and Penstemon dominant. Flowers creamy white. Phenology: Flowers. Origin: Native. WTU-16474 Mary Johanson 05-02 8 Jul 2005 with Robert Goff, Doug Williams, Joe Johanson, Jessie Johanson, Caprifoliaceae Nomi Odano, Julia Odano, Ben Legler, David Giblin Lonicera involucrata (Richardson) Banks ex Spreng. var. WTU-369328 involucrata U.S.A., WASHINGTON, CHELAN COUNTY: Caprifoliaceae North Cascades National Park. North shore of Dagger Lake, circa 3 air miles southeast of Rainy Pass. Lonicera involucrata (Richardson) Banks ex Spreng. var. Elev. 5508 ft. involucrata 48° 28.129' N, 120° 39.34' W; UTM Zone 10, 673280.642E, 5371063.857N; WGS 84, uncertainty: 4 m., Source: GPS, UTM U.S.A., MONTANA, MINERAL COUNTY: calculated from Lat-Lon. Clearwater National Forest. Heart Lake Trail. 3/8 mile below lake. Wet subalpine meadows, mostly flat with Eriophorum, Carex Elev. 5710 ft. species, Viola palustris, scattered Abies lasiocarpa, Picea 46.95318°, -114.96813°; WGS 84, uncertainty: 1000 m., Source: engelmannii. Fruits deep maroon-black; shrubs to 2 meters tall. Nat. Geog. TOPO!, Georef'd by John Bassett Phenology: Fruits. Origin: Native. Wet area along stream; meadow. 10 degree west facing slope. With Alnus, Rudbeckia occidentallis, Picea engelmannii, Senecio Nathan Walker 06-19 28 Jul 2006 triangularis. Flowers yellow. Phenology: Flowers. Origin: Native. WTU-366895 Don Knoke 838 8 Jul 2005 WTU-365748 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, CHELAN COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Lake Chelan National Recreation Area; Northeast side of Coon Lake where creek enters lake. One kilometer northeast of High Bridge in U.S.A., IDAHO, CLEARWATER COUNTY: Stehekin River Valley. Clearwater National Forest. Hidden Creek Camp. Elev. 2170 ft. Elev. 3410 ft. 48° 23' 26.8" N, 120° 49' 57.5" W; UTM Zone 10, 660447E, 46.83177°, -115.17851°; WGS 84, uncertainty: 500 m., Source: 5362004N; WGS 84, uncertainty: 8 m., Source: GPS Nat. Geog. TOPO!, Georef'd by John Bassett Moist deciduous forest and brushy thickets, with Acer macrophyllum, Level damp soil; few rocks. With Thuja plicata, Symphoricarpos alba, Alnus sp., Cornus sericea, Rubus parviflorus, Viola sp. Berries dark Trillium ovatum, Acer glabrum. Black fruit. Shrubs 2 meters tall. red-black; bracts red; occasional shrubs to 3 meters tall. Phenology: Uncommon. Phenology: Fruits. Origin: Native. Fruits. Origin: Native. Don Knoke 885 9 Jul 2005 Barbara Smith 06-29 29 Jun 2006 WTU-374192 WTU-366873 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., IDAHO, CLEARWATER COUNTY: U.S.A., IDAHO, CLEARWATER COUNTY: Clearwater National Forest. Hidden Creek Camp. Clearwater National Forest. Black Canyon on Clearwater River. Elev. 3410 ft. National Forest Road 250. Roadside stream to river. 0.35 miles 46.83177°, -115.17851°; WGS 84, uncertainty: 500 m., Source: south of Pete Ott Creek. Nat. Geog. TOPO!, Georef'd by John Bassett 46.74512°, -115.24091°; WGS 84, uncertainty: 500 m., Source: Level damp soil; few rocks. With Thuja plicata, Symphoricarpos alba, Nat. Geog. TOPO!, Georef'd by John Bassett Trillium ovatum, Acer glabrum. Shrubs 1 meter tall; berries bright Wet roadside ditch with Marchantia, bog orchids and mosses. Low red. Phenology: Fruits. Origin: Native. shrubs and dense herbaceous cover mostly on uphill side of road. Shrubs collected on river side of road. Dark purple/black fruit flanked Don Knoke 899 9 Jul 2005 by maroon bracts. Phenology: Fruits. Origin: Native. WTU-365824 Maria Gerace 05-34 9 Jul 2005 with Robert Goff, Doug Williams, Dale Blum WTU-369293 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Ripple Island. San Juan Islands. Skull Island. 48° 39.415' N, 123° 7.896' W Elev. 40 ft. Oblong (north-south) island with extensive shrub cover and a stand 48° 38.343' N, 122° 59.168' W; WGS 84, Source: GPS of mostly dead Populus tremuloides. Large shell and gravel beach A 40 foot high island with many Juniperus trees, abundant Camassia on west side, with broad shelving bedrock bench draped with leichtlinii and Plectritis congesta. Numerous bedrock outcrops with seaweed. A few scattered plants. Phenology: Flowers. Origin: ledges. Occasional trailing over shrubs. Phenology: Flowers. Origin: Native. Native. Rod Gilbert 06-84 27 Apr 2006 Pam Camp 06-05 30 May 2006 WTU-385660 WTU-388936 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., IDAHO, CLEARWATER COUNTY: Clearwater National Forest. 0.2 miles northeast of Mush Saddle. U.S.A., WASHINGTON, SAN JUAN COUNTY: Forest Service Road 711. San Juan Islands. Skull Island. Elev. 5287 ft. Elev. 40 ft. 46.75606°, -115.3646°; WGS 84, uncertainty: 500 m., Source: 48° 38.343' N, 122° 59.168' W; WGS 84, Source: GPS Nat. Geog. TOPO!, Georef'd by John Bassett A 40 foot high island with many Juniperus trees, abundant Camassia Southeast facing rocky roadcut. Open forest and brush. Common. leichtlinii and Plectritis congesta. Numerous bedrock outcrops with Bush about 1 meter tall. Phenology: Fruits. Origin: Native. ledges. Two plants on island. Phenology: Flowers. Origin: Native. Wendy McClure 05-40 10 Jul 2005 Pam Camp 06-45 30 May 2006 with Suzanne Bagshaw, Ben Legler, Don Knoke, Jessie Johanson, Joe Johanson, Mary Johanson WTU-388889 WTU-370609 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Oak Island. U.S.A., WASHINGTON, SAN JUAN COUNTY: Elev. 10 ft. San Juan Islands. Flattop Island. Weed Point on northeast end of 48° 36.463' N, 122° 57.283' W; WGS 84, Source: GPS Flattop Island; northern part of San Juan Archipelago. Very small, 0.19 acre (measured) island with Holodiscus/Rosa Elev. 0-10 ft. shrubs, Sanicula understory; one goose nest; otter evident 48° 38.871' N, 123° 4.707' W; WGS 84, uncertainty: 2 m. abundantly; 3 Pseudotsuga menziesii, 2 Juniperus scopulorum, and Rocky open point, treeless, with thin soil, with Rosa nutkana, 1 Quercus garryana. A few vines trailing across shrubs. Phenology: Grindelia, Lathyrus japonicus, grasses. Corolla rich orange; a Flowers. Origin: Native. occasional vine among shrub thickets. Phenology: Flowers. Origin: Native. Peter Dunwiddie 06-174 30 May 2006 Carrie Marschner 06-33 31 May 2006 WTU-389247 WTU-388682 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Fawn Island. U.S.A., WASHINGTON, SAN JUAN COUNTY: Elev. 50 ft. San Juan Islands. Flattop Island. Weed Point on northeast end of 48° 36.853' N, 123° 0.421' W; Source: GPS Flattop Island; northern part of San Juan Archipelago. Three acre island is developed with 3 houses, trails, and Elev. 0-10 ft. considerable disturbance of understory; forest (open) of 48° 38.871' N, 123° 4.707' W; WGS 84, uncertainty: 2 m. Pseudotsuga menziesii, Arbutus, Quercus, and Juniperus. A few Rocky open point, treeless, with thin soil, with Rosa nutkana, shrubs, low and trailing, on bedrock outcrop. Phenology: Flowers. Grindelia, Lathyrus japonicus, grasses. Flowers pink-purple; vine Origin: Native. clambering over rock and low shrubs. Phenology: Flowers. Origin: Native. Peter Dunwiddie 06-230 31 May 2006 Carrie Marschner 06-71 31 May 2006 WTU-389805 WTU-388713 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ×bella Zabel U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, OKANOGAN COUNTY: San Juan Islands. McConnell Island off southwest side of Orcas E end of Conners Lake, Sinlahekin Wildlife Refuge. Island; north end of island near small wetland. Elev. 1508 ft. Elev. 5 ft. 48° 45' N, 119° 39.5' W 48° 35.902' N, 123° 1.444' W; WGS 84, uncertainty: 5 m., Source: Lakeshore, dike. Two meter tall shrubs, fruit bright red, juicy, GPS occasional adventive, with Sonchus arvensis, Cirsium arvense. Small rocky bald by wetland; mostly grassy with scattered Phenology: Fruits. Pseudotsuga, Juniperus, and Quercus. Corolla bright orange; uncommon, climbing up trees. Phenology: Flowers. Origin: Native. Peter F. Zika 22416 25 Sep 2005 Victoria Wyllie-Echeverria 06-20 30 May 2006 WTU-364244 WTU-389942 Caprifoliaceae Caprifoliaceae Lonicera ×bella Zabel Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, SKAGIT COUNTY: Under powerlines just north of Summer Lake U.S.A., WASHINGTON, SAN JUAN COUNTY: along Lake Cavanaugh Road; 3 miles NE of San Juan Islands. McConnell Island. Lake McMurray. Elev. 0-25 ft. Elev. 560 ft. 48° 35.747' N, 123° 1.469' W 48° 20' 7" N, 122° 9' 58" W; NAD 27 Forest edge/rocky outcropping habitat along west and south sides of Brushy cleared area under powerlines, with island. Common and abundant along south and west-facing rocky small perennial stream dominated by Phalaris outcroppings. Phenology: Fruits. Origin: Native. arundinacea. Corolla strongly bilabiate, deep pink; single shrub ca. 2 meters tall near stream. Pam Camp 06-173 8 Sep 2006 Flowers collected 4/30/2005; fruit collected 7/4/2005 from the same shrub. Reference WTU-392654 specimen for illustration in Revision of Flora of the Pacific Northwest WTU Herbarium, 2017. Phenology: Flowers & Fruits. Origin: Introduced. Caprifoliaceae Ben Legler 2605 30 Apr 2005 Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray WTU-367410, WTU-V-034836 U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Sentinel Island. Elev. 134 ft. Caprifoliaceae 48° 38.381' N, 123° 9.049' W; WGS 84, Source: Digital map Frequent at forest edge, trailing and climbing. Phenology: Fruits. Lonicera involucrata (Richardson) Banks ex Spreng. var. Origin: Native. involucrata Phil Green 06-72 7 Sep 2006 U.S.A., WASHINGTON, KING COUNTY: Along Forest service Road 55 0.4 road miles SE of where it WTU-395008 intersects Interstate 90 at Exit 42. Elev. 1480 ft. 47° 24' 35" N, 121° 35' 9" W; WGS 84, uncertainty: 50 m., Caprifoliaceae Source: Digital map Mixed conifer forest with open brushy area around perennial stream, Lonicera ciliosa (Pursh) Poir. ex DC. with Lonicera involucrata, Cornus sericea. Shrubs to 3 meters tall forming thickets by creek; stems light brown. Phenology: Buds. U.S.A., WASHINGTON, SKAGIT COUNTY: Origin: Native. Padilla Bay National Estuarine Research Reserve. Saddlebag Island. Ben Legler 3151 10 Feb 2006 Elev. 17 ft. 48° 32.117' N, 122° 33.333' W; WGS 84, Source: GPS Herbarium: WTU High island with rocky shoreline cliffs; Douglas fir/Madrone forests with Snowberry, Mahonia; forest and understory. Occasional trailing over shrubs. Phenology: Flowers. Origin: Native. Caprifoliaceae Ben Legler 3357 23 May 2006 Lonicera ciliosa (Pursh) Poir. ex DC. WTU-371862 U.S.A., WASHINGTON, SNOHOMISH COUNTY: Along Highway 9 1/4 mile northwest of where highway crosses Harvey Creek, 1.7 miles northwest of Arlington. Elev. 130 ft. 48° 13' 16" N, 122° 8' 23" W; WGS 84, Source: Digital map Roadside backed by Pseudotsuga forest with brushy understory. Vine climbing over other shrubs; bark tan-orange. Phenology: Buds. Origin: Native. Ben Legler 3202 4 Mar 2006 Herbarium: WTU Caprifoliaceae Caprifoliaceae Lonicera tatarica L. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, YAKIMA COUNTY: U.S.A., WASHINGTON, KITTITAS COUNTY: Yakima. Between arboretum and west bank of Blewett Pass. Yakima River, north of W. Birchfield Road. Elev. 4217 ft. Elev. 984 ft. 47° 20.914' N, 120° 40.971' W 46° 35.3' N, 120° 28' W On crest. Dry rocky soil. Slope gentle to 30 degree. Open forest, Partially shaded riparian thickets with Morus, pine/fir. Associated plants; Pinus contorta, Pachystima myrsinites, Populus, Toxicodendron, Crataegus douglasii. Arctostaphylos nevadensis, Spiraea betulifolia. Orange. Uncommon. Adventive shrubs, flowers pink - fading pink; Open area among trees. Phenology: Flowers. Origin: Native. glabrous, bracts << ovary. Reference specimen for illustration in Revision of Flora of the Pacific Robert Goff 06-53 8 Jul 2006 Northwest, WTU Herbarium, 2016. Phenology: Flowers. Origin: Introduced. WTU-370955 Peter F. Zika 22727 Caprifoliaceae 12 May 2006 Lonicera involucrata (Richardson) Banks ex Spreng. var. WTU-365040, WTU-V-031917 involucrata U.S.A., WASHINGTON, KITTITAS COUNTY: Wenatchee National Forest. Vicinity of Swauk Creek Camground. Caprifoliaceae Elev. 3257 ft. 47° 19.708' N, 120° 39.196' W Lonicera tatarica L. In fruit; shrub to 2 meters; fruit black with red bracts. Phenology: Fruits. Origin: Native. U.S.A., WASHINGTON, YAKIMA COUNTY: On east side of Yakima River, north of Route 24, near Yakima. Sheila Olmstead 2006-04 8 Jul 2006 Elev. 1000 ft. with Richard Olmstead, Richard Robohm 46° 35.6' N, 120° 24.4' W Partial shade of Acer saccharinum, Populus balsamifera; riparian WTU-370981 zone. Adventive shrubs 1.5 meters tall, flowers pink and fading, to pink, wholly glabrous, spread from cultivation to shoreline. Phenology: Flowers & Fruits. Origin: Introduced. Caprifoliaceae Peter F. Zika 22723B 12 May 2006 Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata WTU-365039 U.S.A., WASHINGTON, KITTITAS COUNTY: Esmeralda Basin. Caprifoliaceae Elev. 4571 ft. 47° 26.475' N, 120° 56.387' W; WGS 84 Lonicera ×bella Zabel Wet meadow. 10 degree south facing slope. Saturated black soil. Associated; Pinus monticola, Rhododendron albiflorum, Lonicera U.S.A., WASHINGTON, SPOKANE COUNTY: involucrata, Alnus sinuata. Edge of meadow in moist soil. Spokane; Finch Arboretum near Woodland Boulevard and Garden Phenology: Flowers & Fruits. Origin: Native. Springs Creek. Elev. 1984 ft. Barbara Williams 06-21 9 Jul 2006 47° 38.6' N, 117° 27.7' W with David Giblin Woods near irrigated lawns. Adventive shrubs 1.5 meters tall; flowers pale pink; spread from cultivation, now a rare escape to WTU-371883 woods near irrigated lawns. Phenology: Flowers. Origin: Introduced. Peter F. Zika 22735 15 May 2006 Caprifoliaceae WTU-365122 Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, CHELAN COUNTY: Caprifoliaceae Wenatchee National Forest. 5 miles from U.S. Highway 97 on Forest Road 7200 about 2 miles south of Camas Meadows. (Camas Land). Lonicera ×bella Zabel Elev. 3319 ft. 47° 26.987' N, 120° 35.272' W U.S.A., WASHINGTON, YAKIMA COUNTY: Open Pine/Fir woodland. Orange flowers; vine. Phenology: Flowers. On east side of Yakima River, north of Route 24, near Yakima. Origin: Native. Elev. 1000 ft. 46° 35.6' N, 120° 24.4' W Doug Williams 2006-28 9 Jul 2006 Edge of clearing, near shoreline, partial shade of Populus with Robert Goff, Dick Olmstead, Sheila Olmstead, Maria Gerace, balsamifera. Adventive shrubs 0.7-1.5 meters tall, flowers white and Cindy Spurgeon, Barb Smith fading, to yellow. Phenology: Flowers & Fruits. Origin: Introduced. WTU-372371 Peter F. Zika 22725A 12 May 2006 WTU-365434 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., WASHINGTON, SKAGIT COUNTY: Coon Island. Padilla Bay National Estuarine Research Reserve. Dot Island. 48° 35.725' N, 123° 1.115' W Elev. 69 ft. Common amongst many of the grasslands on the island. Phenology: 48° 32.0412' N, 122° 33.2112' W; Source: Digital map Fruits. Origin: Native. Sloping island with high end at north end. Rich slopes of Festuca rubra, Plectritis, Cerastium, Camassia, and Zygadenus. Juniperus Megan A. Jensen 530 20 Sep 2007 are common, very shaped and stunted by salt spray, little deer with Peter Dunwiddie, David Giblin, Ted Thomas, Phil Green browse. No geese nesting; otter use on southwest side. Occasional; climbing over Juniperus scopulorum. Flowers orange/yellow. WTU-375478 Phenology: Flowers. Origin: Native. Ben Legler 3780 1 May 2007 Caprifoliaceae WTU-377226 Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, SAN JUAN COUNTY: Caprifoliaceae Reef Island. 48° 36.179' N, 123° 1.021' W Lonicera involucrata (Richardson) Banks ex Spreng. var. Large island dominated by Pseudotsuga menziesii; open understory involucrata with Elymus and Holodiscus. Occasional in Pseudotsuga menziesii understory. Phenology: Fruits. Origin: Native. U.S.A., WASHINGTON, YAKIMA COUNTY: Wenatchee National Forest; Forest Service Road 1502. Pond on Peter Dunwiddie 07-96 19 Sep 2007 edge of road adjacent to dry coniferous forest. with Phil Green, Megan Jensen, David Giblin, Ted Thomas Elev. 5488 ft. 46.74905° N, 121.12782° W; WGS 84, Source: GPS WTU-375776 Small pond adjacent to Pinus contorta and Abies lasiocarpa forest. Carex species dominant around pond. On forest edge; occasional. Phenology: Flowers. Origin: Native. Caprifoliaceae Megan A. Jensen 319 6 Jul 2007 Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray with David Giblin, Don Knoke U.S.A., WASHINGTON, SAN JUAN COUNTY: WTU-369824 Reef Island. 48° 36.179' N, 123° 1.021' W Large island dominated by Pseudotsuga menziesii; open understory Caprifoliaceae with Elymus and Holodiscus. Common in open grasslands around island. Phenology: Fruits. Origin: Native. Lonicera ciliosa (Pursh) Poir. ex DC. Peter Dunwiddie 07-159 19 Sep 2007 U.S.A., WASHINGTON, SKAGIT COUNTY: with Phil Green, Megan Jensen, David Giblin, Ted Thomas Padilla Bay National Estuarine Research Reserve. Dot Island. 48° 32.0412' N, 122° 33.2112' W; Source: Digital map WTU-376206 Sloping island with high end at north end. Rich slopes of Festuca rubra, Plectritis, Cerastium, Camassia, and Zigadenus. Juniperus are common, very shaped and stunted by saltspray, little deer Caprifoliaceae browse. No geese nesting. Otter use on southwest side. A couple of plants in Symphoricarpos on east side. Phenology: Flowers. Origin: Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Native. U.S.A., WASHINGTON, SAN JUAN COUNTY: Barbara Williams 07-04 7 Jun 2007 San Juan Islands National Wildlife Refuge and Wilderness Area. Turn Island. Large island 0.25 miles off the east shore of San Juan WTU-378494 Island. 48° 31.97' N, 122° 58.22' W; NAD 83, Source: Digital map Caprifoliaceae Large island 1/4 mile off the east shore of San Juan Island. Common around entire perimeter, mostly in open grasslands. Phenology: Lonicera periclymenum L. Fruits. Origin: Native. U.S.A., WASHINGTON, KING COUNTY: David Giblin 1258 20 Sep 2007 Mercer Island, Lake Washington. Near Island Crest Park and Island with Peter Dunwiddie, Megan Jensen, Phil Green, Ted Thomas Crest Way. Elev. 328 ft. WTU-375649 47° 33.1' N, 122° 13.3' W Partly shaded thickets, with Pseudotsuga, Acer macrophyllum, Caprifoliaceae Rubus armeniacus, Corylus cornuta. Adventive vine climbing 8 meter up Arbutus, spread from cultivation in area. Phenology: Lonicera ciliosa (Pursh) Poir. ex DC. Flowers. Origin: Introduced. U.S.A., WASHINGTON, SAN JUAN COUNTY: Peter F. Zika 22820 25 Jun 2006 San Juan Islands National Wildlife Refuge and Wilderness Area. Turn Island. Large island 0.25 miles off the east shore of San Juan WTU-366186 Island. 48° 31.97' N, 122° 58.22' W; NAD 83, Source: Digital map Occasional and widely scattered. Phenology: Fruits. Origin: Native. David Giblin 1241 20 Sep 2007 with Phil Green, Megan Jensen, Peter Dunwiddie, Ted Thomas WTU-375709 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera japonica Thunb. U.S.A., WASHINGTON, KITTITAS COUNTY: U.S.A., WASHINGTON, ASOTIN COUNTY: Summit of Lion Rock on west side of Table Mountain. South bank of reservoir on Snake River (Lower Granite Lake), 3 air Elev. 6377 ft. kilometers northeast of Silcott. 47° 15.138' N, 120° 35.138' W; WGS 84 Elev. 934 ft. Dry subalpine rocky slopes, basalt rock with Abies lasiocarpa, Pinus 46° 25.4' N, 117° 9.8' W; T11N R45E S22 contorta, Lupinus sp., Microseris nutans, Erigeron linearis. Corolla With Verbascum thapsus, Elaeagnus angustifolia. White flowers yellow; single shrub 1 meter tall on rock talus. Phenology: Flowers. fade yellow, common vine spread from original introduction and now Origin: Native. covering parts of pasture and fences. (=coll. #16661). Phenology: Flowers. Origin: Introduced. Don Knoke 1097 7 Jul 2006 with Ben Legler, Richard Robohm, Barbara Smith, Joe Arnett, Peter F. Zika 22528 7 Oct 2005 Suzanne Bagshaw WTU-368322 WTU-377899 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SAN JUAN COUNTY: COUNTY: North Finger Island. Lake Armstrong Road north of Arlington. Elev. 0-65 ft. T32N R5E S35; NAD 27, uncertainty: 805 m., 48° 45' 24.18" N, 122° 53' 27.64" W; WGS 84 Source: TRS2LL, Georef'd by Ben Legler Primarily Pseudotsuga/Arbutus forest with exposed areas along the Edge of mixed conifer woods. Woody vine perimeter, especially at east end of island; sandstone substrate. climbing over other shrubs. Flowers orange. Phenology: Flowers & Fruits. Origin: Native. [Located in teaching collection]. Phenology: Flowers. Origin: Native. David Giblin 1361 27 Sep 2007 M. Arnot 461 WTU-381032 8 May 1992 Caprifoliaceae WTU-323833, WTU-V-005734 Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: North Finger Island. Lonicera involucrata (Richardson) Banks ex Spreng. Elev. 0-65 ft. 48° 45' 24.18" N, 122° 53' 27.64" W; WGS 84 U.S.A., WASHINGTON, SNOHOMISH Primarily Pseudotsuga/Arbutus forest with exposed areas along the COUNTY: perimeter, especially at east end of island; sandstone substrate. Near Marysville. Common throughout island in forest understory; berries 48.05194°, -122.17583°; WGS 84, reddish-orange. Phenology: Fruits. Origin: Native. uncertainty: 5000 m., Source: GeoLocate, Georef'd by WTU Staff David Giblin 1360 27 Sep 2007 Upper edge of tide flats. With stunted Picea sitchensis and Rosa nutkana amid open areas WTU-381523 of Aster sp., Juncus and grasses. Area bordered to the north with Acer macrophyllum and Alnus rubra. In isolated clumps. Scrawly shrub 1.5 Caprifoliaceae meters tall. [Located in teaching collection]. Phenology: Flowers & Fruits. Origin: Native. Lonicera maackii (Rupr.) Herder M. Arnot 196 U.S.A., WASHINGTON, KING COUNTY: North end of Washington Park Arboretum, Seattle. 3 Jun 1990 Elev. 32 ft. 47° 38.5' N, 122° 17.7' W WTU-317064, WTU-V-005767 Near Duck Bay, partial shade, with Acer macrophyllum, Acer circinatum, Rubus armeniacus, Anus rubra. Pith brown, hollow between nodes, slightly immature fruit subglobose to depressed Caprifoliaceae globose, shiny, orange-red, 5.5-6.3 x 5.8-6.8 millimeters wide; bird-sown adventive (not cultivated in vicinity). Phenology: Fruits. Lonicera ciliosa (Pursh) Poir. ex DC. Origin: Introduced. U.S.A., WASHINGTON, ASOTIN COUNTY: P. F. Zika 23551 8 Oct 2007 Fields Spring State Park. Adjacent to dirt road that runs to southwest with A. L. Jacobson corner of park. T7N R45E S21; WGS 84, uncertainty: 10000 m., Source: WTU-367828 GeoLocate, Georef'd by WTU Staff Shady Pinus ponderosa forest with Amelanchier alnifolia, Symphoricarpos oreophilus, and Rosa in understory. Scattered; flowers orange. Phenology: Flowers. Origin: Native. Jeff Walker 200 26 Jun 2004 WTU-370042 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray var. Lonicera ciliosa (Pursh) Poir. ex DC. hispidula U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S.A., OREGON, LANE COUNTY: Cliff Island. Eugene; The Nature Conservancy Willow Creek Preserve, south Elev. 22 ft. side of W 18th Avenue. Near East Fork Willow Creek. 48° 35.391' N, 123° 0.826' W Elev. 400 ft. Island is forested heavily with second growth Pseudotsuga with 44° 2.2' N, 123° 9.9' W; T18S R4W S4 extensive Gaultheria shallon understory. Heavily deer grazed. Many Partly shaded damp thickets, shrub and tree-invaded wet prairie species relatively uncommon around periphery on cliffs. remnant, with Franxinus latifolia, Crataegus suksdorfii, Crataegus Occasionally widespread in forest. Phenology: Flowers. Origin: monogyna, Pyrus communis, Amelanchier alnifolia. Flowers purple; Native. glabrous, glaucous, upper leaves fused; vine. Phenology: Buds. Origin: Native. David Giblin 1613 29 May 2008 Peter F. Zika 19575 14 May 2004 WTU-396155 WTU-369130 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: Lonicera morrowii A. Gray San Juan Islands. Little McConnell Island; small island (1.3 acres) connected to McConnell Island by beach at low tide. U.S.A., WASHINGTON, OKANOGAN COUNTY: Elev. 0-25 ft. West side of Sinlahekin Creek, Sinlahekin Wildlife Area; circa 1.5 air 48° 35.927' N, 123° 1.547' W miles north of headquarters. Open grassland with scattered Juniperus, madrone. One flowering Elev. 1476 ft. shrub/vine on north side of island. Phenology: Flowers. Origin: 48° 46' N, 119° 39' W; T39N R25E S4 Native. Thickets on bank, with Apocynum, Crataegus douglasii, Cornus sericea, Symphoricarpos albus, Rosa woodsii. Adventive shrubs Carrie Marschner CM 08-30 29 May 2008 1.5-2 meters tall, fruit bright red, naturalized and common. Phenology: Fruits. Origin: Introduced. WTU-394119 Peter F. Zika 22155 21 Jul 2005 Caprifoliaceae WTU-369735 Lonicera myrtillus Hook. f. & Thomson CANADA, BRITISH COLUMBIA: Caprifoliaceae Asian Garden, University of British Columbia Botanical Garden, Vancouver. Lonicera ciliosa (Pursh) Poir. ex DC. Mounded shrub circa 1 meter tall with stiff, wiry stems. Flowers pale cream. Garden #24945-508-86. Phenology: Flowers. Origin: U.S.A., WASHINGTON, SAN JUAN COUNTY: Introduced. Cultivated. Ewing Island. Long, narrow island off NE end of Sucia Island. Steep sandstone ridges run length of island. Gerald B. Straley 8994 15 May 1996 48.76455° N, 122.88093° W; WGS 84, Source: Digital map Mostly forested with PSME, ARME, and JUMA. Grassy balds along WTU-371287 south shore and at east end of island. Frequent clambering across shrubs. Widespread. Phenology: Flowers. Origin: Native. Caprifoliaceae Janneke Hille-Ris Lambers JHRL 08-42 28 May 2008 Lonicera involucrata (Richardson) Banks ex Spreng. WTU-395995 CANADA, BRITISH COLUMBIA: Waglisla, Campbell Island, by health centre at roadside. Caprifoliaceae 52° 10' N, 128° 8' W Phenology: Flowers. Origin: Native. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Brian D. Compton 99 2 Aug 1987 U.S.A., WASHINGTON, SAN JUAN COUNTY: Ewing Island. Long, narrow island off NE end of Sucia Island. Steep WTU-371315 sandstone ridges run length of island. 48.76455° N, 122.88093° W; WGS 84, Source: Digital map Mostly forested with PSME, ARME, and JUMA. Grassy balds along Caprifoliaceae south shore and at east end of island. Occasional clambering vine, widespread. Phenology: Flowers. Origin: Native. Lonicera involucrata (Richardson) Banks ex Spreng. Janneke Hille-Ris Lambers JHRL 08-67 28 May 2008 U.S.A., OREGON, CURRY COUNTY: Circa 1.5 miles south of Floras Lake. WTU-396141 Elev. 150 ft. T31S R15W S29; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler On moist sandy margins of cultivated cranberry fields, full sun, with Holcus lanatus, Equisetum arvense. Flowers yellow, rare. Phenology: Flowers. Origin: Native. P. F. Zika 14310 8 Sep 1999 WTU-372437 Caprifoliaceae Caprifoliaceae Lonicera japonica Thunb. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, KING COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Near shore of Lake Washington, Clarke Beach Park, southeast end San Juan Islands National Wildlife Refuge and Wilderness Area. of Mercer Island. East Sucia #7. Elev. 16 ft. Elev. 50 ft. T24N R5E S30; NAD 27, uncertainty: 805 m., Source: TRS2LL, 48.76398° N, 122.88579° W; Source: GPS Georef'd by Ben Legler The largest island in the chain. Forested with Arbutus, Juniperus, Twining vines spreading from plants in cultivated area, ca. 40 plants Pseudotsuga, with shrub understory. Height of 50 feet or so. A few seen; with Rubrus armeiacus, Hedera helix, Populus balsamifera. plants trailing over shrubs. Phenology: Fruits. Origin: Native. Fragrant cream-yellow flowers, shiny black fruit 5-6 millimeters long. Phenology: Flowers & Fruits. Origin: Introduced. Peter Dunwiddie 08-168 26 Sep 2008 Peter F. Zika 15723 12 Dec 2000 WTU-393600 with A. L. Jacobson WTU-373513 Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Caprifoliaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray San Juan Islands National Wildlife Refuge and Wilderness Area. East Sucia #7. U.S.A., WASHINGTON, SAN JUAN COUNTY: Elev. 50 ft. San Juan Islands National Wildlife Refuge and Wilderness Area. Cliff 48.76398° N, 122.88579° W; Source: GPS Island. The largest island in the chain. Forested with Arbutus, Juniperus, Elev. 22 ft. Pseudotsuga, with shrub understory. Height of 50 feet or so. A 48° 35.391' N, 123° 0.826' W couple of plants trailing over shrubs. Phenology: Fruits. Origin: Island is forested heavily with second growth Pseudotsuga with Native. extensive Gaultheria shallon understory. Heavily deer grazed. Many species relatively uncommon around periphery on cliffs. Occasional Peter Dunwiddie 08-183 26 Sep 2008 and widely sscattered. Phenology: Fruits. Origin: Native. WTU-393572 Phil Green 08-87 24 Sep 2008 Caprifoliaceae WTU-393672 Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands National Wildlife Refuge and Wilderness Area. Lonicera ciliosa (Pursh) Poir. ex DC. East Sucia #4. Elev. 40 ft. U.S.A., WASHINGTON, SAN JUAN COUNTY: 48.76435° N, 122.88718° W; Source: GPS San Juan Islands National Wildlife Refuge and Wilderness Area. 40 feet high sandstone rock with mostly overhanging cliffs around all Willow Island. A large, 100' high rocky island 1/3 mile west of Blakely sides. Heavily vegetated across top, with diverse meadows, thickets, Island. and some Pseudotsuga and Juniperus. 0.21 acres. One plant trailing Elev. 100 ft. over Rosa nutkana. Phenology: Fruits. Origin: Native. 48.5402°, -122.82242°; WGS 84, Source: Georeferenced, Georef'd by WTU Staff Peter Dunwiddie 08-193 26 Sep 2008 Partially treed with Pseudotsuga and Arbutus. Open understory dominated by Elymus glaucus and Symphoricarpos albus. East side WTU-393685 has extensive cliffs and ledges and is particularly rich with ferns. Considerable deer browse. Rock appears to be serpentine. Climbing over holodiscus and Pseudotsuga. Phenology: Fruits. Origin: Native. Caprifoliaceae Steve Hahn 08-90 25 Sep 2008 Lonicera ×bella Zabel WTU-392979 U.S.A., WASHINGTON, DOUGLAS COUNTY: Columbia River. 47.71576°, -119.626904°; WGS 84, uncertainty: 65000 m., Caprifoliaceae Source: Digital Map, Georef'd by Joseph Stormer Slope 0%. Phenology: Flowers. Origin: Introduced. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Kathryn A. Beck 200508 19 May 2005 U.S.A., WASHINGTON, SAN JUAN COUNTY: U.S BLM land. Victim Island. WTU-374947 Elev. 50 ft. 48° 36.831' N, 122° 58.472' W An open, forested, 3 acre island with Arbutus, Quercus, Juniperus, and Pseudotsuga; Holodiscus dominant shrub understory; deer evident. Occasional, climbing over shrubs. Phenology: Fruits. Origin: Native. Peter Dunwiddie 08-155 24 Sep 2008 WTU-393779 Caprifoliaceae Caprifoliaceae Lonicera tatarica L. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, DOUGLAS COUNTY: U.S.A., WASHINGTON, SAN JUAN COUNTY: Columbia River. San Juan Islands. West Matia #1. Large island T30N R24E; uncertainty: 7200 m., Georef'd by Joseph Stormer, at north side of harbor, west end of Matia Island. Invalid coordinates (located outside county bounds). 48.74946° N, 122.84378° W; WGS 84, Plant's growing near river's edge along abandoned, partially Source: Digital map submerged highway. Associated with Amelanchier alnifolia, Rosa Cliffy sandstone island rising steeply out of cove multiflora, Hypericum perforatum, Dactylis glomerata, Asclepius at west end of Matia Island. Approximately 50 speciosa, Purshia tridentata, Rubus leucodermis. Phenology: Fruits. feet high, with Pseudotsuga menziesii and Origin: Introduced. Arbutus menziesii forest across top. Common in understory across island. Phenology: Fruits. Kathryn A. Beck 200512 28 Jun 2005 Origin: Native. WTU-374952 Brianne Cohen 09-93 24 Sep 2009 Caprifoliaceae WTU-403733, WTU-V-012806 Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Little Sucia Island, off the southwest corner of Caprifoliaceae Sucia Island. Elev. 80 ft. Lonicera ciliosa (Pursh) Poir. ex DC. 48° 45.283' N, 122° 55.198' W Includes forested uplands (circa 80 feet high) with shrub understory, U.S.A., WASHINGTON, SAN JUAN COUNTY: steep sandstone cliffs, eroding dirt bluffs. Linked by a low, San Juan Islands. West Matia #1. Large island shrub-covered (Rosa nutkana) shell-midden berm. Psuedotsuga at north side of harbor, west end of Matia Island. menziesii and arbutus forest with dense shrub understory. Isthmus 48.74946° N, 122.84378° W; WGS 84, connects two higher upland areas. Trailing vine on very dry Source: Digital map south-facing dirt bank. Phenology: Flowers. Origin: Native. Cliffy sandstone island rising steeply out of cove at west end of Matia Island. Approximately 50 David Giblin 2866 28 May 2009 feet high, with Pseudotsuga menziesii and Arbutus menziesii forest across top. Occasional WTU-396745 on north side, understory. Phenology: Fruits. Origin: Native. Caprifoliaceae Brianne Cohen 09-102 Lonicera ciliosa (Pursh) Poir. ex DC. 24 Sep 2009 U.S.A., WASHINGTON, SAN JUAN COUNTY: WTU-403618, WTU-V-012868 San Juan Islands. Little Sucia Island, off the southwest corner of Sucia Island. Elev. 80 ft. 48° 45.283' N, 122° 55.198' W Caprifoliaceae Includes forested uplands (circa 80 feet high) with shrub understory, steep sandstone cliffs, eroding dirt bluffs. Linked by a low, Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray shrub-covered (Rosa nutkana) shell-midden berm. Psuedotsuga menziesii and arbutus forest with dense shrub understory. Isthmus U.S.A., WASHINGTON, SAN JUAN COUNTY: connects two higher upland areas. Trailing vine with bright orange San Juan Islands. Ripple Island. flowers. Frequent and widespread. Phenology: Flowers. Origin: 48° 39.415' N, 123° 7.896' W Native. Oblong (north-south) island with extensive shrub cover and a stand of mostly dead Populus David Giblin 2870 28 May 2009 tremuloides. Large shell and gravel beach on west side, with broad shelving bedrock bench WTU-396733 draped with seaweed. Several plants, local, on north end of island. Phenology: Fruits. Origin: Native. Amy Dearborn 09-03 23 Sep 2009 WTU-403686, WTU-V-012099 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata U.S.A., WASHINGTON, SAN JUAN COUNTY: San Juan Islands. Matia Island. Forested wetland on southeast side U.S.A., MONTANA, GRANITE COUNTY: of island. Butte Creek, two miles up creek from Rock Creek. Trail ends at Elev. 32 ft. 46.51519 N, 113.73225 W. 48.7466° N, 122.8438° W; WGS 84, uncertainty: 100 m., Source: Elev. 4453-4842 ft. Digital map 46.51036° N, 113.73225° W Shrub to 2 meters on edge of wetland. Scattered. Phenology: Fruits. Uneven area between creek and talus slopes. Soil moist, rocky. Origin: Native. Associated species: Picea engelmannii, Aster radulinus, Philadelphus lewisii, Symphoricarpos albus, Physocarpus. Black Phil Green 09-94 24 Sep 2009 fruit; common. Phenology: Fruits. Origin: Native. WTU-398275 Hannah Marx 08-6 12 Jul 2008 WTU-395890 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae CANADA, BRITISH COLUMBIA: Lonicera involucrata (Richardson) Banks ex Spreng. var. Meares Island: Opsitsaht. involucrata 49° 11' N, 125° 50' W Coniferous oldgrowth forest west of village site. Growing in full sun U.S.A., MONTANA, GRANITE COUNTY: at forest margin/edge of rocky shoreline. Phenology: Flowers. Origin: Lolo National Forest. Rock Creek Canyon. Turnout for Microburst Native. interpretive sign. 46° 26' 39.03" N, 113° 45' 41.42" W; WGS 84, uncertainty: 10 m., A. R. McCutcheon C-M-2 22 Jun 1994 Source: Digital map Mixture of mesic and dry meadows along Rock Creek, bordered by WTU-375931 south-facing, dry rocky slope. Locally common in mesic area of meadow near river. Phenology: Fruits. Origin: Native. Caprifoliaceae Hanh-La Phan 08-36 12 Jul 2008 Lonicera japonica Thunb. WTU-392970 U.S.A., WASHINGTON, ASOTIN COUNTY: Chief Timothy habitat management area near shore of Snake River, Caprifoliaceae about 11 air kilometers wet of Clarkston, near Silcott Road off Route 12. Lonicera involucrata (Richardson) Banks ex Spreng. Elev. 656 ft. 46° 25' N, 117° 11.2' W U.S.A., WASHINGTON, KITTITAS COUNTY: Weed. Phenology: Flowers. Origin: Introduced. Along the west fork of Caseknife Creek about 1 mile from Taneum Creek. Richard Old s.n. 5 Oct 2009 Elev. 3599 ft. 47° 5.322' N, 120° 58.76' W; T18N R16E S4; WGS 84 WTU-377268 Dry, gravelly soil; 20° southeast-facing slope. Other plants: Pseudotsuga menziesii, Spiraea betulifolia, Calamagrostis rubescens. Immature fruit; scattered. Phenology: Fruits. Origin: Caprifoliaceae Native. Lonicera involucrata (Richardson) Banks ex Spreng. Don Knoke 1863 23 Jun 2009 U.S.A., OREGON, HARNEY COUNTY: WTU-381085 Steens Mountain. Fish Lake Campgrounds. Area above slope, southeast side of campground. Caprifoliaceae Elev. 7390 ft. 42° 44.307' N, 119° 38.638' W; WGS 84, uncertainty: 500 m., Lonicera ciliosa (Pursh) Poir. ex DC. Source: Digital map, Georef'd by John Bassett Area above slope; Southeast side of C.G. Gentle slope; dry soil; U.S.A., WASHINGTON, WHATCOM COUNTY: large rock at rim covered w/ Atremisia tridentata. Other plants: Ross Lake National Recreation Area. Random walk from Lightning grasses, Antennaria microphylla. Black fruit; scattered; shrub. Creek Campground to Cat Island along east shore of lake. Phenology: Fruits. Origin: Native. Elev. 1646 ft. 48.88402° N, 121.03805° W; WGS 84, uncertainty: 10 m., Don Knoke 1437 21 Jul 2007 Source: GPS Mixture of seasonally inundated lake bottom, and rocky balds. Pinus WTU-392613 contorta. Locally common on west-facing rocky outcropping. Phenology: Flowers. Origin: Native. Becky Peace 10-4 13 May 2010 WTU-381920 Caprifoliaceae Caprifoliaceae Lonicera ×bella Zabel Lonicera chamissoi Bunge ex P. Kir. U.S.A., WASHINGTON, KING COUNTY: RUSSIAN FEDERATION, SAKHALIN REGION: North end of Lake Park Drive South. Mt. Baker, East/central Sahkalin; Longri River basin. Seattle. 54° 1.297' N, 142° 57.009' E Elev. 22 ft. Creek bed and draw running east to west; dense herbaceous T24N R4E S10; NAD 27, uncertainty: 805 m., understory with scattered Betula, Larix, Sorbus and Pinus. In Source: TRS2LL, Georef'd by Ben Legler meadow next to creek bed; plants with loose, bushy habit, growing Slightly overgrown thickets, partial shade. Pink with Pinus and Sorbus; 2-3 feet tall. Phenology: Fruits. Origin: flowers, cultivated shrubs 2-2.5 meters tall. Native. Reference specimen for illustration in Revision of Flora of the Pacific Northwest, WTU Kari Stiles 2001-311 16 Aug 2001 Herbarium, 2016. Phenology: Flowers. Origin: Native. Cultivated. WTU-390666 P. F. Zika 15953 Caprifoliaceae with A. L. Jacobson Lonicera sachalinensis (Fr. Schmidt) Nakai 20 Apr 2001 RUSSIAN FEDERATION, SAKHALIN REGION: WTU-376090, WTU-V-031918 Southeast Sakhalin Island; Cape Menaputsy. 46° 23.177' N, 143° 36.511' E Edge of Betula/Picea forest and steep slope covered with herbs and forbs. In fruit. Phenology: Fruits. Origin: Native. Caprifoliaceae Kari Stiles 2001-348 21 Aug 2001 Lonicera chamissoi Bunge ex P. Kir. WTU-391797 RUSSIAN FEDERATION, SAKHALIN REGION: Molchiga Bay. 54° 16.803' N, 142° 26.343' E Caprifoliaceae Pinus, Larix forest with low herb/forb understory. Phenology: Fruits. Origin: Native. Lonicera sachalinensis (Fr. Schmidt) Nakai Kari Stiles 2001-82 6 Aug 2001 RUSSIAN FEDERATION, SAKHALIN REGION: Southern part of Sakhalin Island. Environs of Yuzhno-Sakhalinsk. WTU-391109 Approximately 6 kilometers east of town on road past Pionerskaya and reservoir. Rogatka River valley. Elev. 1630 ft. Caprifoliaceae 46° 59.049' N, 142° 49.614' E Betula and Fraxinus forest. Sasa dominant in understory. Occasional Lonicera chamissoi Bunge ex P. Kir. Pinus pumila in understory. In fruit (green.) Growing as shrub next to foot path. Phenology: Fruits. Origin: Native. RUSSIAN FEDERATION, SAKHALIN REGION: Tum River mouth, Nadezhda Bay; northwest Sakhalin Island. Suzanne Joneson 2798 22 Jul 2001 54° 12.135' N, 142° 24.383' E Two drainages south of campsite (1 kilometer south?), sloping WTU-391059 meadows running up from beach to Pinus/Larix-covered bluff; herbs/forbs with Sorbus, Salix and junipers, soil sandy and muddy. In meadow at edge of Larix forest; with Sorbus, Salix. Phenology: Caprifoliaceae Fruits. Origin: Native. Lonicera chamissoi Bunge ex P. Kir. Kari Stiles 2001-125 9 Aug 2001 RUSSIAN FEDERATION, SAKHALIN REGION: WTU-391913 Southern part of Sakhalin Island. Sea of Okhotsk side. Environs of Starodubskoye Village and mouth of Nayba River. Elev. 39 ft. Caprifoliaceae 47° 25.826' N, 142° 46.294' E River delta area. Flat area with ferns and grasses; Hemerocallis and Lonicera chamissoi Bunge ex P. Kir. Cirsium dominant. Main vegetation over 1 meter tall. With red berries. Common in large area of burned trees and low-lying herbs. RUSSIAN FEDERATION, SAKHALIN REGION: Phenology: Fruits. Origin: Native. Northwest Sakhalin Island; near Moskal'vo Village; environs of Bolshoye Lake. Suzanne Joneson 2888 23 Jul 2001 Elev. 60 ft. 53° 37.698' N, 142° 29.079' E WTU-391026 Dunes and sandy meadows; plants have large, moundy habit. Incredible specimen; only one plant, huge fruits; growing in dunes alongside (in shadow of) 6 foot high Sorbus. Phenology: Fruits. Origin: Native. Kari Stiles 2001-222 13 Aug 2001 WTU-391725 Caprifoliaceae Caprifoliaceae Lonicera sachalinensis (Fr. Schmidt) Nakai Lonicera involucrata (Richardson) Banks ex Spreng. RUSSIAN FEDERATION, SAKHALIN REGION: U.S.A., OREGON, GRANT COUNTY: Southern part of Sakhalin Island. Approximately 25 kilometers west North Fork Malheur Campground. of Yuzhno-Sakhalinsk; north side of Tiobut River. Approximately 407 Elev. 4785 ft. kilometers east of junction with Lyntoga River. 44.20833° N, 118.38258° W Elev. 656 ft. Pine, larch, Douglas fir forest and wet meadows 46° 58.639' N, 142° 14.876' E along river. In fruit (black) and with red bracts; Hillside of dominantly Sasa, with occasional patches of Populus and streamside. Phenology: Fruits. Origin: Native. Betula. In fruit. Growing as short shrub in understory of Populus, and above Sasa. Phenology: Fruits. Origin: Native. Sheila Olmstead 2010-3 Suzanne Joneson 2923 26 Jul 2001 18 Jul 2010 WTU-391090 WTU-401450, WTU-V-8351

Caprifoliaceae Caprifoliaceae Lonicera caerulea L. Lonicera involucrata (Richardson) Banks ex Spreng. RUSSIAN FEDERATION, SAKHALIN REGION: Northwest part of Sakhalin Island. East of Cape Mani. U.S.A., OREGON, GRANT COUNTY: Elev. 91 ft. Malheur National Forest. Big Creek Campground. Off Road 815 on 54° 16.899' N, 142° 26.912' E Forest Service Road 16. Beach bluff, with patches of windblown Larix and meadows. Small Elev. 5150 ft. shrub. 2 fruits present. Not common in area. Phenology: Vegetative. 44.18672° N, 118.61591° W Origin: Native. Ponderosa pine/lodgepole pine woodland, open campsites, dry soil; also wetter, shady area by creek. Medium-sized shrub to 2-3 meters. Suzanne Joneson 3129 6 Aug 2001 Rare in moist riparian bank. Corollas gone, bracts red, fruits black. Phenology: Fruits. Origin: Native. WTU-391568 Ian Shelley 10-23 17 Jul 2010 Caprifoliaceae WTU-388553 Lonicera chamissoi Bunge ex P. Kir. Caprifoliaceae RUSSIAN FEDERATION, SAKHALIN REGION: Northern tip of Sakhalin Island. From unnamed river north of Muni Lonicera involucrata (Richardson) Banks ex Spreng. var. River, to Cape Bakianiy. involucrata Elev. 0 ft. 54° 23.624' N, 142° 39.734' E U.S.A., OREGON, GRANT COUNTY: Meadows and cliffs along beach. In fruit (yellow.) Small shrub. Malheur National Forest. Mount Baldy Research Natural Area, on Growing in small sloughing meadow along beach. Phenology: Fruits. south side of Pine Creek adjacent to the RNA parcel, just east of Origin: Native. Malheur National Forest Road 1430 after creek crossing. Elev. 6340 ft. Suzanne Joneson 3136 7 Aug 2001 44.331771° N, 118.80673° W; WGS 84 Mostly closed canopy conifer forest surrounding large fen complex WTU-391571 on hillside. Open, wet shelves with only herbaceous cover frequent. Small stream adjacent complex drains into creek below. Associates: Picea engelmannii, Abies lasiocarpa, Abies grandis/concolor Caprifoliaceae dominant. Shrubs to about 1 meter, growing at margins on stream among woody debris. Flowers yellow-green. Phenology: Flowers. Lonicera caerulea L. Origin: Native. RUSSIAN FEDERATION, SAKHALIN REGION: C. E. Hinchliff 1043 18 Jul 2010 Tum River mouth, Nadezhda Bay; northwest Sakhalin Island. 54° 12.135' N, 142° 24.383' E WTU-388826 Two drainages south of campsite (1 kilometer south?), sloping meadows running up from beach to Pinus/Larix-covered bluff; herbs/forbs with Sorbus, Salix and junipers, soil sandy and muddy. Growing in shade of Betula, Salix at edge of Larix forest. Phenology: Fruits. Origin: Native. Kari Stiles 2001-122 9 Aug 2001 WTU-390961 Caprifoliaceae Caprifoliaceae Lonicera japonica Thunb. Lonicera japonica Thunb. U.S.A., WASHINGTON, ASOTIN COUNTY: U.S.A., OREGON, LANE COUNTY: Chief Timothy Wildlife Management Area, West end of Skinner Butte summit, north of Legacy Oaks, Eugene. United States Bureau of Reclamation. Elev. 672 ft. Elev. 731 ft. 44° 3.5' N, 123° 5.7' W 46.42314° N, 117.16623° W; WGS 84, At forest edge, with Oemleria, Prunus avium, Hedera. White flowers uncertainty: 10 m., Source: GPS fade yellow, presumably naturalized. Phenology: Flowers. Origin: Degraded shoreline area on south bank of Introduced. Snake River with small pond area. Massive patch near roadside and sprawling; flowers P. F. Zika 25599 13 Jul 2011 white and yellow. Reference specimen for with E. R. Alverson, B. Newhouse illustration in Revision of Flora of the Pacific Northwest WTU Herbarium, 2016. Phenology: WTU-387456 Flowers. Origin: Introduced. David Giblin 3537 Caprifoliaceae 9 Jun 2010 Lonicera maackii (Rupr.) Herder WTU-397254, WTU-V-030496 U.S.A., OREGON, MULTNOMAH COUNTY: Portland Women's Forum State Park at Chanticleer Point; 1.5 air miles east of Corbett. Forested area northwest of parking lot. Caprifoliaceae Elev. 840 ft. UTM Zone 10, 557681E, 5042590N; WGS 84 Lonicera involucrata (Richardson) Banks ex Spreng. var. Shrubby understory of moist deciduous forest, northerly aspect. With involucrata Acer macrophyllum, Alnus rubra, Symphoricarpos albus, Hedera helix, Urtica dioica. Fruits red; naturalized, numerous plants U.S.A., WASHINGTON, STEVENS COUNTY: spreading in understory. Phenology: Fruits. Origin: Introduced. Western Colville National Forest and vicinity: Little Round Top Mountain: Middle Fork Mill Creek, circa 9 road miles northeast of R.E. Brainerd 1588 14 Aug 2009 Colville on Aladdin Route. Elev. 2300-3300 ft. WTU-388195 T36N R40E S15 NW; T36N R40E S16 NE; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Cedar at creekside; mixed Douglas fir forest, and aspen groves on Caprifoliaceae lower slopes; Douglas fir, ponderosa pine and larch with pinegrass and ninebark on upper slopes; rock outcrops of shale/slate. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, SKAGIT COUNTY: Jean Wood 872 5 Jun 1995 Padilla Bay National Estuarine Research Reserve. Hat Island. Southeast and southern end of island. WTU-386264 48° 31.326' N, 122° 32.728' W; WGS 84 Douglas fir forested high island with steep grassy balds (Elymus glaucus, Festuca rubra, Koeleria macrantha) on south side, with Caprifoliaceae mostly native bunch grasses and abundant Zigadenus venenosus. Common in forest understory. Phenology: Fruits. Origin: Native. Lonicera involucrata (Richardson) Banks ex Spreng. David Giblin 3207 17 Sep 2009 U.S.A., WASHINGTON, COLUMBIA COUNTY: [No locality given on label] WTU-388941 T7N R39E S3 NE of NE; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Open ridge, bordered by ponderosa pine, Douglas and grand fir, Caprifoliaceae slight slope, west facing. Phenology: Flowers. Origin: Native. Lonicera maackii (Rupr.) Herder Susan Whittlesey s.n. 18 Jun 1980 U.S.A., OREGON, MULTNOMAH COUNTY: WTU-387315 Sauvie Island, Oak Island Road at the end of the pavement. Elev. 10 ft. UTM Zone 10, 513623E, 5060904N; WGS 84 Caprifoliaceae Shrubby roadside. With Fraxinus latifolia, Crataegus monogyna, Rosa multiflora, Rubus armeniacus. Shrub 2.5 meters tall; fruits red. Lonicera involucrata (Richardson) Banks ex Spreng. var. Phenology: Fruits. Origin: Introduced. involucrata R.E. Brainerd 1639 15 Oct 2009 U.S.A., WASHINGTON, YAKIMA COUNTY: with N. Otting Okanogan-Wenatchee National Forest. Below and to northeast of Little Bald Mountain on west side of Forest Road 1600. WTU-388422 46.91046° N, 121.15371° W; WGS 84, uncertainty: 100 m., Source: GPS North-south running drainage with basalt talus basin; Pseudotsuga/Abies/Xanthocyparis forest and open seasonally wet meadow with southeastern exposure. Uncommon along edge of basalt talus; corollas yellow. Phenology: Flowers. Origin: Native. David Giblin 3942 14 Jul 2011 with Alex D. Harkness, Don Knoke WTU-388502 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, THURSTON COUNTY: U.S.A., WASHINGTON, SKAGIT COUNTY: Maytown Road, approximately 2 miles east of Interstate-5. Vendovi Island. Near cove on north side of Elev. 226 ft. island. In general vicinity of house. 46° 53.929' N, 122° 55.471' W Elev. 16-65 ft. Orange flowers; vine. Phenology: Flowers. Origin: Native. 48° 36' 46.44" N, 122° 36' 50.4" W; WGS 84, uncertainty: 50 m., Source: Digital map Richard G. Olmstead 2006-2 19 May 2006 Open, mowed area around house and with Sheila Olmstead, Valerie Soza, Yuan Yaowu outbuildings. Abundant trailing vine, widespread in forests and shrub thickets. Phenology: WTU-388519 Flowers. Origin: Native. Peter Dunwiddie 2011-47 Caprifoliaceae 1 Jun 2011 Lonicera involucrata (Richardson) Banks ex Spreng. WTU-404325, WTU-V-012390 U.S.A., WASHINGTON, THURSTON COUNTY: Junction of Maytown Road with Tilley Road Southwest. Elev. 186 ft. 46° 53' 59" N, 122° 54' 31" W Caprifoliaceae Roadside near wet area. Yellow flowered shrub to 3 meters. Phenology: Flowers. Origin: Native. Lonicera ciliosa (Pursh) Poir. ex DC. Richard G. Olmstead 2006-11 19 May 2006 U.S.A., WASHINGTON, WHATCOM COUNTY: with Sheila Olmstead, Valerie Soza, Yuan Yaowu Above Clayton Bay, circa 9 miles south of Bellingham. In thin soil over steep sandstone bank. Shrub circa 2 meters tall, WTU-388511 flowers orange. Phenology: Flowers. Origin: Native. Carl A. Linde Jr. s.n. 7 May 1957 Caprifoliaceae WTU-390390 Lonicera utahensis S. Watson U.S.A., IDAHO, ADAMS COUNTY: Caprifoliaceae Payette National Forest: along Forest Service Road 149 at head of Beaver Creek, circa 9 air Lonicera ciliosa (Pursh) Poir. ex DC. miles northeast of Council. Elev. 5346 ft. U.S.A., WASHINGTON, THURSTON COUNTY: 44.855916° N, 116.314045° W; WGS 84, Glacial Heritage Preserve. uncertainty: 15 m., Source: GPS Elev. 37 m. Dry slope with small basalt pinnacles; soil of 46.86484° N, 123.03631° W; UTM Zone 10N, 497234E, fine-textured gravelly basalt; with Artemisia 5190145N; NAD 83 tridentata, Allium, Mimulus. Corolla pale yellow, Woodland. Quercus garryana, Symphoricarpos albus, Cardamine easily detached; shrubs. Phenology: Flowers. nuttallii, Claytonia sibirica, Galium triflorum. Orange flowers, vine, Origin: Native. scattered. Phenology: Flowers. Origin: Native. Barbara Smith 2011-54 Pat Montague 98 27 May 2008 with Dan Montague 26 Jun 2011 WTU-391287 WTU-406305, WTU-V-17018 Caprifoliaceae Caprifoliaceae Lonicera periclymenum L. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, GRAYS HARBOR COUNTY: Chehalis River, south bank. U.S.A., WASHINGTON, KING COUNTY: Elev. 4 m. Carkeek Park, Ballard, Seattle. 46.94756° N, 123.75713° W Elev. 200 ft. Saltwater marsh dominated by Carex lyngbyei and Deschampsia 47.709893°, -122.37026°; WGS 84, uncertainty: 1000 m., Source: cespitosa; adjacent railroad grade. Shrub to 3 meters tall; flowers Digital Map, Georef'd by Ibro Polovina pink and white, fading to orange. Phenology: Flowers. Origin: In marsh. Woody shrub with hairy leaves; two large bracts Introduced. surrounding yellow petals. Collected for Biology 317 (Plant Identification and Classification). Phenology: Flowers. Origin: Native. David Giblin 3144 1 Sep 2009 June Landenburger 1209 22 Apr 2012 WTU-393005 WTU-390039 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata U.S.A., WASHINGTON, CHELAN COUNTY: Okanogan-Wenatchee National Forest. About 1 U.S.A., WASHINGTON, PEND OREILLE kilometer below Mad River Trailhead junction on COUNTY: Forest Service road. Colville National Forest. Along Round Top Elev. 3800 ft. Mountain Trail #512 (Shedroof Divide trail). 47.82581° N, 120.59873° W; WGS 84, Elev. 1830 m. uncertainty: 100 m., Source: Digital map 48.8061° N, 117.1298° W; WGS 84, Roadside cut along Forest Service road; uncertainty: 1000 m., Source: Digital map, southwestern aspect; scattered rocky seeps. Georef'd by Maria Yousoufian Occasional; flowers orange; growing on forest Mixed coniferous open forest along trail. Some edge. Phenology: Flowers. Origin: Native. rocky outcrops and seeps. Phenology: Flowers. Origin: Native. David Giblin 2971 with Ben Legler Susan Kephart 752 23 Jun 2009 21 Jul 2012 WTU-399104, WTU-V-005489 WTU-407865, WTU-V-021662

Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, PEND OREILLE U.S.A., WASHINGTON, PEND OREILLE COUNTY: COUNTY: Colville National Forest. Along Round Top Colville National Forest. Road to Salmo Mountain Trail #512 (Shedroof Divide trail). Mountain. Approximately 1.5 kilometers below Elev. 1830 m. summit. 48.8061° N, 117.1298° W; WGS 84, Elev. 1900 m. uncertainty: 1000 m., Source: Digital map, 48.9633° N, 117.105° W; WGS 84, Georef'd by Maria Yousoufian uncertainty: 100 m., Source: Digital map, Mixed coniferous open forest along trail. Some Georef'd by Maria Yousoufian rocky outcrops and seeps. Phenology: Fruits. South-facing slope bisected by road; small Origin: Native. seasonal stream along edge of road; slope with seasonally moist open areas. Occasional on Susan Kephart 783 edge of meadow; corollas yellow. Phenology: Flowers. Origin: Native. 21 Jul 2012 Amy Cabral 12-37 WTU-407870, WTU-V-021655 21 Jul 2012 Caprifoliaceae WTU-406770, WTU-V-17819 Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, PEND OREILLE COUNTY: Lonicera involucrata (Richardson) Banks ex Spreng. Colville National Forest. Sullivan Creek Road, 10.2 miles from junction with Sullivan Lake U.S.A., WASHINGTON, PEND OREILLE Road. COUNTY: Elev. 887 m. Colville National Forest. Along old road bed 48.88888° N, 117.09759° W converted to trail between trailhead and start of Flowers yellow. Phenology: Flowers. Origin: trail to Hall Mountain at end of Forest Service Native. Road 500 off Sullivan Creek Road. Elev. 1670 m. Kathleen Whitson 2012-37 48.80119° N, 117.231037° W; WGS 84, uncertainty: 18 m., Source: GPS, Georef'd by 20 Jul 2012 Maria Yousoufian Moist mossy soil of road bank and bed under WTU-408307, WTU-V-021513 Alnus sinuata and conifer forest. Flowers yellow; shrubs. Phenology: Flowers. Origin: Native. Don Knoke 2311 21 Jul 2012 WTU-406843, WTU-V-019565 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, PEND OREILLE U.S.A., WASHINGTON, CHELAN COUNTY: COUNTY: Camas meadows. Colville National Forest. Bear Pasture trailhead Elev. 2000 ft. and along road near trailhead. 47° 28.774' N, 120° 37.172' W; T23N R18E sec. 20 Elev. 5620 ft. Roadside near Camas meadow. Douglas fir, Ponderosa pine forest 48.91724° N, 117.13914° W with Acer grandidentatum in the understory. Phenology: Flowers. Flowers white. Phenology: Flowers. Origin: Origin: Native. Native. P. Camp 3300 13 Jun 2013 Doug Williams 2012-1 WTU-394249 22 Jul 2012 WTU-407568, WTU-V-020624 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Caprifoliaceae U.S.A., WASHINGTON, PEND OREILLE Lonicera utahensis S. Watson COUNTY: Colville National Forest. Slate Creek; on Forest U.S.A., WASHINGTON, STEVENS COUNTY: Road 3155 at junction with 100. In streambanks By Red Marble Quarry. and roadsides. Elev. 4500 ft. Elev. 2836 ft. T31N R38E S25; NAD 27, uncertainty: 805 m., Source: TRS2LL, 48.94306° N, 117.28383° W Georef'd by Ben Legler Gravel road, moist streambank, mixed Douglas fir and larch community. Serviceberry and ninebark in the coniferous forest. In fruit; fruits black. understory. Phenology: Flowers. Origin: Native. Phenology: Fruits. Origin: Native. P. Camp 349 27 May 1980 Drew Parker 18 WTU-394212 20 Jul 2012 WTU-408312, WTU-V-021520 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, STEVENS COUNTY: Lonicera utahensis S. Watson By Red Marble Mine. T31N R38E S24; NAD 27, uncertainty: 805 m., Source: TRS2LL, U.S.A., WASHINGTON, PEND OREILLE Georef'd by Ben Legler COUNTY: Douglas fir and larch habitat. Snowberry and pinegrass in the Colville National Forest. Slate Creek; on Forest understory. Some cedar in the vicinity. Phenology: Flowers. Origin: Road 3155 at junction with 100. In streambanks Native. and roadsides. Elev. 2836 ft. P. Camp 445 24 Jun 1980 48.94306° N, 117.28383° W Gravel road, moist streambank, mixed WTU-394234 coniferous forest. In fruit; fruits red. Phenology: Fruits. Origin: Native. Caprifoliaceae Drew Parker 24 Lonicera involucrata (Richardson) Banks ex Spreng. 20 Jul 2012 U.S.A., WASHINGTON, CHELAN COUNTY: WTU-408882, WTU-V-022204 Camas meadows. Elev. 2810 ft. 47° 28.236' N, 120° 35.145' W; T23N R18E sec. 21 Caprifoliaceae Forb meadow rimmed by snowberry, serviceberry, some aspen, Spiraea and Douglas hawthorn. Douglas fir, grand fir forest Lonicera utahensis S. Watson surrounding the area. Common. Phenology: Fruits. Origin: Native. U.S.A., WASHINGTON, PEND OREILLE P. Camp 3299 13 Jun 2013 COUNTY: Colville National Forest. Round Top Mountain. WTU-394243 Top of mountain on Shedroof Divide Trail #512. Elev. 1975 m. 48° 48.618' N, 117° 7.79' W Open grassy meadow with Arenaria on lower slope approach and diversity of species below. Occasional Abies even at top. Phenology: Flowers. Origin: Native. Ruth Pelz 12-5 21 Jul 2012 WTU-408924, WTU-V-022243 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, COWLITZ COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Kress Lake, just east of Interstate-90. South of Kapowsin. Elev. 10 m. Elev. 652 ft. 46.0462° N, 122.8531° W; WGS 84, 46° 1.518' N, 122° 21.961' W; Source: uncertainty: 20 m., Source: GPS, Georef'd by Specimen Label, Invalid coordinates (located Maria Yousoufian outside county bounds). Apparent man-made lake approximately 5 Roadside, forested wetland ditch. Oemleria hectares in size. Aquatic plant diversity low with cerasiformis, Rubus sp., Dicentra formosa, Potamogeton amplifolius, Myriophyllum Maianthemum dilatatum, Galium sp., Rubus spicatum, and Elodea nuttallii most common. spectabilis, Claytonia sibirica, Vaccinium Locally common in upland adjacent to lake; parvifolium. Flowers cream yellow, 6 foot shrub. berries red-orange. Phenology: Fruits. Origin: Phenology: Flowers. Origin: Native. Native. Etsuko Reistroffer 17 David Giblin 5115 with Cathy Maxwell, Peter Zika 25 Apr 2014 28 Aug 2013 WTU-399898, WTU-V-006188 WTU-409048, WTU-V-023046 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata U.S.A., WASHINGTON, THURSTON COUNTY: Millersylvania State Park. U.S.A., MONTANA, GRANITE COUNTY: Elev. 188 ft. Beaverhead-Deerlodge National Forest. Coles 46° 54.656' N, 122° 55.15' W; Source: Gulch, road 5156. Specimen Label Elev. 6100 ft. Along a trail in wetland. Spiraea douglasii, 46.4078° N, 113.55693° W; WGS 84, Rubus spectabilis, Maianthemum dilatatum, uncertainty: 500 m., Source: GPS Lysichiton americanus, Physocarpus capitatus, Wet meadow with 3 sedges, scattered Picea, Hypochaeris radicata, Prunella vulgaris var. Lonicera, and Geum, transitioning to dry forest vulgaris. Cream yellow corolla with soft hair. with Lupinus, Vaccinium scoparium, Carex, Phenology: Flowers. Origin: Native. small scattered Populus tremuloides, Arctostaphylos uva-ursi. Scattered low-growing Etsuko Reistroffer 11 clumps in wet meadow. Phenology: Flowers. Origin: Native. 21 Apr 2014 Ryan Boyle 7 WTU-399903, WTU-V-006216 12 Jul 2008 WTU-403633, WTU-V-012853 Caprifoliaceae Lonicera maackii (Rupr.) Herder Caprifoliaceae U.S.A., WASHINGTON, KING COUNTY: Lonicera maackii (Rupr.) Herder Washington Park Arboretum, Seattle. Elev. 15 m. U.S.A., WASHINGTON, CLARK COUNTY: 47.637° N, 122.298° W; Source: Specimen Salmon Creek Trail, approximately 250 meters Label east of the east end of Salmon Creek Park. Very close to base of large Acer macrophyllum. 45.70814° N, 122.66145° W; WGS 84, White flowers, adventive shrub 3 meters tall. uncertainty: 50 m., Source: Digital map Reference specimen for illustration in Revision Phenology: Fruits. Origin: Introduced. of "Flora of the Pacific Northwest." WTU Herbarium, 2017. Phenology: Flowers. Origin: Michael Monfort s.n. Introduced. Nov 2014 Peter F. Zika 26518 WTU-418227, WTU-V-039575 1 May 2014 WTU-400222, WTU-V-006463 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray var. Lonicera caerulea L. hispidula U.S.A., MONTANA, BEAVERHEAD COUNTY: U.S.A., OREGON, CLACKAMAS COUNTY: Beaverhead-Deerlodge National Forest. Madrone Wall Park, Damascus. Mussigbrod Lake. Trail along east side of lake at 45.389764° N, 122.456849° W; Source: north end of lake. Specimen Label Elev. 1997 m. Opening in forest near cliff. Phenology: 45.80231° N, 113.62111° W; WGS 84, Vegetative. Origin: Native. uncertainty: 100 m., Source: GPS West-facing rocky slope in broken canopy of J. Lesh s.n. once-burned coniferous forest and adjacent willow swamp. Uncommon in willow meadow; 7 Aug 2014 shrub to 1 meter tall. Phenology: Fruits. Origin: Native. WTU-400723, WTU-V-007231 Rene Dufour 14-23 13 Jul 2014 Caprifoliaceae WTU-414414, WTU-V-030052 Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, KLICKITAT COUNTY: Junction Nester Creek and Nester Peak Roads. Caprifoliaceae 45.790207°, -121.536625°; WGS 84, uncertainty: 5000 m., Source: Digital Map, Lonicera involucrata (Richardson) Banks ex Spreng. Georef'd by Dan Post Shady roadside with grass, forbs, Douglas-fir U.S.A., MONTANA, BEAVERHEAD COUNTY: forest. Phenology: Flowers. Origin: Native. Beaverhead-Deerlodge National Forest. Circa 2 miles south of Steel Creek Campground on David Gross 996 Forest Road 2420 near South Fork Steel Creek. Elev. 2133 m. 21 May 2014 45.57967° N, 113.34314° W; Source: GPS Phenology: Flowers. Origin: Native. WTU-400767, WTU-V-007284 Jared Grummer 2014-14 12 Jul 2014 Caprifoliaceae WTU-411775, WTU-V-026312 Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, KITTITAS COUNTY: Along Jeep road off Forest Road 1115 0.1 miles Caprifoliaceae west of United States Highway 97. Elev. 750 m. Lonicera ciliosa (Pursh) Poir. ex DC. 47.24805° N, 120.69647° W; T20N R17E S3; WGS 84 U.S.A., WASHINGTON, MASON COUNTY: Fairly moist, loamy soil; 10° east-facing slope. Olympic National Forest. Cushman Lake Road. Other plants: Abies grandis, Berberis repens, Elev. 241 m. Calamagrostis rubescens. Flowers orange; 47.49646° N, 123.30341° W; WGS 84, common. Phenology: Flowers. Origin: Native. uncertainty: 200 m., Source: GPS Disturbed south-facing slope with loose, gravelly Don Knoke 2443 soil. Hypericum perforatum and Hypochaeris radicata common. Locally common vine along 4 Jul 2013 road; corollas orange. Phenology: Flowers. Origin: Native. WTU-409474, WTU-V-023486 David Giblin 5538 26 May 2015 WTU-417787, WTU-V-038981 Caprifoliaceae Caprifoliaceae Lonicera tatarica L. Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata U.S.A., WISCONSIN, WAUSHARA COUNTY: Pine Lake, Hancock. U.S.A., WASHINGTON, SNOHOMISH 44.13008°, -89.509878°; WGS 84, COUNTY: uncertainty: 715 m., Source: GeoLocate, Happy Valley Tree Farm. Georef'd by Maria Yousoufian 48.187751°, -122.262495°; WGS 84, Margin of wetlands. Sandy uplands. [Attached uncertainty: 264 m., Source: Digital Map, packet labeled 7-9-2005.] Phenology: Fruits. Georef'd by Maria Yousoufian Origin: Introduced. Phenology: Flowers. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 4 Jul 2001 20 May 1988 WTU-402698, WTU-V-010587 WTU-402692, WTU-V-010593

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera utahensis S. Watson U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, OKANOGAN COUNTY: COUNTY: Tunk Valley. Happy Valley Tree Farm, Happy Valley. 48.561819°, -119.486442°; WGS 84, 48.187751°, -122.262495°; WGS 84, uncertainty: 300 m., Source: GeoLocate, uncertainty: 264 m., Source: Digital map, Georef'd by Crystal Shin Georef'd by Maria Yousoufian Phenology: Flowers. Origin: Native. Drainage ditch. Phenology: Vegetative. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. May 1990 Feb 2010 WTU-402697, WTU-V-010590 WTU-402693, WTU-V-010594

Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Happy Valley Tree Farm, Happy Valley. Happy Valley Tree Farm, Happy Valley. 48.187751°, -122.262495°; WGS 84, 48.187751°, -122.262495°; WGS 84, uncertainty: 264 m., Source: Digital map, uncertainty: 264 m., Source: Digital map, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian Along drainage ditch. Phenology: Vegetative. Phenology: Flowers. Origin: Native. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 18 Apr 2006 Jan 2009 WTU-402690, WTU-V-010595 WTU-402694, WTU-V-010591 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera involucrata (Richardson) Banks ex Spreng. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: Happy Valley Tree Farm. Happy Valley Tree Farm, Happy Valley. 48.187751°, -122.262495°; WGS 84, 48.187751°, -122.262495°; WGS 84, uncertainty: 264 m., Source: Digital Map, uncertainty: 264 m., Source: Digital map, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian [Attached packet labeled 6-11-89.] Phenology: Side drainage. Phenology: Flowers. Origin: Fruits. Origin: Native. Native. Clayton Haberman s.n. Clayton Haberman s.n. 10 Jun 1989 3 May 2009 WTU-402691, WTU-V-010596 WTU-402695, WTU-V-010592 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: East side Port Susan, north Tulare Beach. East shore Port Susan, north Tulare Beach. 48.105718°, -122.34509°; WGS 84, 48.105718°, -122.34509°; WGS 84, uncertainty: 367 m., Source: Digital map, uncertainty: 367 m., Source: Digital Map, Georef'd by Maria Yousoufian Georef'd by Maria Yousoufian Base of sandy-clay bank. Phenology: Fruits. Phenology: Flowers. Origin: Native. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 29 May 1987 22 Sep 2005 WTU-402684, WTU-V-010601 WTU-402688, WTU-V-010597 Caprifoliaceae Caprifoliaceae Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray Lonicera periclymenum L. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: East shore Port Susan. Happy Valley. 48.145716°, -122.338629°; WGS 84, Fence row. Phenology: Flowers. Origin: uncertainty: 10000 m., Source: Digital Map, Introduced. Georef'd by Ibro Polovina Bank. Phenology: Fruits. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 11 Jun 1989 2 Nov 1987 WTU-402689, WTU-V-010598 WTU-402685, WTU-V-010602

Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SNOHOMISH COUNTY: COUNTY: "Tulalip Pass" Marine Drive. East shore Port Susan, north Tulare Beach. 48.068429°, -122.29181°; WGS 84, 48.105718°, -122.34509°; WGS 84, uncertainty: 500 m., Source: GeoLocate, uncertainty: 367 m., Source: Digital Map, Georef'd by Ibro Polovina Georef'd by Maria Yousoufian Phenology: Flowers. Origin: Native. Shoreline. [Attached packet is labeled 10-9-95.] Phenology: Flowers. Origin: Native. Clayton Haberman s.n. Clayton Haberman s.n. 26 May 1988 4 May 2005 WTU-402501, WTU-V-010694 WTU-402686, WTU-V-010599 Caprifoliaceae Caprifoliaceae Lonicera dioica L. Lonicera hispidula (Lindl.) Douglas ex Torr. & A. Gray U.S.A., WISCONSIN, WAUSHARA COUNTY: North shoreline Pine Lake, Hancock. U.S.A., WASHINGTON, SNOHOMISH 44.132544°, -89.509792°; WGS 84, COUNTY: uncertainty: 667 m., Source: Digital Map, East shore Port Susan, north Tulare Beach. Georef'd by Maria Yousoufian 48.105718°, -122.34509°; WGS 84, Sandy-mud. Phenology: Flowers. Origin: Native. uncertainty: 367 m., Source: Digital Map, Georef'd by Maria Yousoufian Ellen Gennrich s.n. Base of sandy-clay bank. Phenology: Flowers. Origin: Native. 24 May 2009 Clayton Haberman s.n. WTU-402578, WTU-V-010858 6 Jun 2005 WTU-402687, WTU-V-010600 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera conjugialis Kellogg U.S.A., WASHINGTON, SNOHOMISH U.S.A., WASHINGTON, SKAMANIA COUNTY: COUNTY: Southwest side of Mount Adams. Stagman Ridge. Warm Beach Senior Community, Stanwood. Elev. 1394 m. 48.181878°, -122.35046°; WGS 84, 46° 8.977' N, 121° 35.21' W; Source: Field notes uncertainty: 360 m., Source: Digital Map, Sprawling shrub 1.5 meters tall. Found second shrub at 4600 feet Georef'd by Maria Yousoufian elevation. Phenology: Flowers and Fruits. Origin: Native. Roadside, mixed woods. Phenology: Fruits. Origin: Native. David Biek 05-79 25 Jun 2005 with Susan McDougall Clayton Haberman s.n. Herbarium: WTU 20 Sep 2008 WTU-402577, WTU-V-010859 Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Caprifoliaceae U.S.A., WASHINGTON, YAKIMA COUNTY: Lonicera ciliosa (Pursh) Poir. ex DC. Southeast side of Mount Adams. Forest Road 170. Elev. 1113 m. U.S.A., WASHINGTON, SNOHOMISH 46° 2.933' N, 121° 27.251' W; Source: Field notes COUNTY: Phenology: Flowers. Origin: Native. Tulare Beach. 48.101763°, -122.343202°; WGS 84, David Biek 05-113 17 Jul 2005 uncertainty: 650 m., Source: GeoLocate, with Susan McDougall Georef'd by Maria Yousoufian Base of clay bluff, "seep" area. Phenology: Herbarium: WTU Fruits. Origin: Native. Clayton Haberman s.n. Caprifoliaceae 22 Sep 2005 Lonicera involucrata (Richardson) Banks ex Spreng. WTU-402574, WTU-V-010862 U.S.A., WASHINGTON, OKANOGAN COUNTY: Mount Chopaka area. On the west side of Mount Chopaka. Caprifoliaceae T40N R24E S23; NAD 27, uncertainty: 805 m., Source: TRS2LL, Georef'd by Ben Legler Lonicera involucrata (Richardson) Banks ex Spreng. var. Moist seep. Growing with the roots in water, involucrata shady. Douglas fir and Engelmann spruce forest. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, SKAMANIA COUNTY: West side of Mount Adams. Forest Road 23, at the Pacific Crest P. Camp 1108 Trail crossing. Elev. 1173 m. 25 Jun 1982 46° 9.77' N, 121° 37.94' W; Source: Field notes Several shrubs, in a wet ditch below the road; shaded part of day by WTU-405113, WTU-V-013536 Picea engelmannii and Thuja plicata. Branch from shrub 2 meters tall. Phenology: Flowers. Origin: Native. David Biek 04-245 18 Jul 2004 Caprifoliaceae with Susan McDougall Lonicera utahensis S. Watson Herbarium: WTU U.S.A., IDAHO, VALLEY COUNTY: Payette National Forest. Profile Creek Road. Caprifoliaceae Approximately 4 kilometers north from junction with Stibnite Road. From GPS coordinate to 300 Lonicera conjugialis Kellogg meters south along road. Elev. 1796 m. U.S.A., WASHINGTON, YAKIMA COUNTY: 44.99526° N, 115.40882° W; WGS 84, Southeast side of Mount Adams. Logging road along northern slopes uncertainty: 10 m., Source: GPS of ridgetop dividing the Dairy Creek and Cougar Creek drainages. East-facing slope adjacent to road on edge of Elev. 1227 m. Picea engelmannii forest. Scattered at base of 46° 7.949' N, 121° 20.048' W; Source: Field notes embankment in dry soils. Phenology: Fruits. Phenology: Flowers. Origin: Native. Origin: Native. Paul Slichter 28 12 Jun 2005 Veanna Willard 2015-6 with Mike Carlson, John Bassett, David Giblin Herbarium: WTU 10 Jul 2015 WTU-415397, WTU-V-033981 Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata U.S.A., IDAHO, VALLEY COUNTY: Payette National Forest. End of old mining road U.S.A., WASHINGTON, SKAGIT COUNTY: leading to Crater Lake. Sullivan Slough by the round-about in La Elev. 2200 m. Conner. 45.04193° N, 115.42029° W; WGS 84, 48.39304° N, 122.48461° W; Source: GPS uncertainty: 100 m., Source: Digital map Wetland backwater of slough dominated by West-facing subalpine meadow bordered by Typha latifolia, Urtica dioica, and Solanum forested riparian zone and dry, rocky dulcamara. Lonicera is the dominant shrub. outcroppings. Scattered on slope in loose, open Abundance: Common. Growth Habit: Fruiting soil. Phenology: Fruits. Origin: Native. shrub-branch. Site ID: NWCA16-4855. Visit #2. Collection #Q5 (QA Voucher). Phenology: Mike Carlson 15-11 Fruits. Origin: Native. with Veanna Willard, John Bassett, David Giblin Patricia Johnson NWCA16-4855-V2-Q5 10 Jul 2015 20 Jul 2016 WTU-415330, WTU-V-032437 WTU-411144, WTU-V-025443

Caprifoliaceae Caprifoliaceae Lonicera utahensis S. Watson Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, JEFFERSON U.S.A., WASHINGTON, PEND OREILLE COUNTY: COUNTY: Along Upper Big Quilcene Trail #833.1 ca 1.5 air Pend Oreille River, 10 miles south of Ione km west of trailhead. bridge, east side. Elev. 891 m. 48.590399°, -117.341292°; WGS 84, 47.826986° N, 123.062014° W; WGS 84, uncertainty: 1500 m., Source: Digital Map, uncertainty: 27 m., Source: GPS, Georef'd by Georef'd by Sara Shiina Ben Legler Phenology: Flowers. Origin: Native. Moist, montane conifer forest near river, with Tsuga heterophylla, Abies amabilis, Rubus Sandie Durand s.n. pedatus, Tiarella trifoliata, Clintonia uniflora, Hylocomnium splendens. Berries bright red, in 21 May 2016 pairs; small shrubs to ca 1 meter tall, local and uncommon in moss over a large boulder in WTU-411738, WTU-V-026271 slight forest opening. Phenology: Fruits. Ben Legler 14204 25 Jul 2016 Caprifoliaceae WTU-418967, WTU-V-041872 Lonicera ×bella Zabel Caprifoliaceae U.S.A., WASHINGTON, CLALLAM COUNTY: Sequim. Lonicera involucrata (Richardson) Banks ex Spreng. 48.079536°, -123.101844°; WGS 84, uncertainty: 4697 m., Source: GeoLocate, Georef'd by Ben Legler U.S.A., WASHINGTON, KING COUNTY: Introduced. Phenology: Flowers. Origin: Introduced. North Bend near Gardiners Creek. 47.50768° N, 121.80239° W; Source: GPS J. M. Grant s.n. Apr 1915 Scrub-shrub wetland with mucky saturated soils. Salix scouleriana and Douglas' spiraea WTU-16478 common. Athyrium filix-femina understory with Lysichiton americanus. Abundance: Common. Growth Habit: Woody, fruiting shrub. Site ID: Caprifoliaceae NWCA16-4884. Visit #1. Collection #Q5 (QA Voucher). Phenology: Fruits. Origin: Native. Lonicera morrowii A. Gray Patricia Johnson NWCA16-4884-V1-Q5 U.S.A., WASHINGTON, OKANOGAN COUNTY: Near bank of Okanogan River, west of Woody 31 May 2016 Island, south of Ione Street, Okanogan. Elev. 245 m. WTU-411087, WTU-V-025527 48.3503° N, 119.5998° W; Source: Specimen Label Shaded thicket, riparian forest, with Rosa, Populus trichocarpa, Ulmus glabra, Crataegus douglasii, Acer negundo. Shrubs 1 meter tall, pith brown and hollow between nodes, white flowers fade to yellow. Phenology: Flowers. Origin: Introduced. Peter F. Zika 27723 3 May 2016 WTU-410467, WTU-V-024469 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., IDAHO, SHOSHONE COUNTY: U.S.A., IDAHO, SHOSHONE COUNTY: Coeur d'Alene National Forest: where Forest Coeur d'Alene National Forest. Coeur d'Alene Service Road 805 crosses Eagle Creek. River Road approximately 1 kilometer east of Elev. 917 m. Independence Creek. 47.685794° N, 115.875966° W; WGS 84, Elev. 900 m. uncertainty: 9 m., Source: GPS 47.88597° N, 116.16918° W; WGS 84, Moist, open conifer forest and brushy stream uncertainty: 100 m., Source: Digital map bottoms bisected by gravel road, with Abies South-facing slope dominated by Holodiscus grandis, Thuja plicata, Tsuga heterophylla. discolor and Hypericum perforatum; rocky faces Shrubs circa 2-3 meters tall in brushy areas by throughout. Uncommon on forest edge; vine to 1 stream. Phenology: Fruits. Origin: Native. meter tall. Phenology: Fruits. Origin: Native. Terri Knoke 2016-06 John Bassett 2016-5 9 Jul 2016 9 Jul 2016 WTU-420521, WTU-V-047677 WTU-419240, WTU-V-042039

Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera maackii (Rupr.) Herder U.S.A., WASHINGTON, STEVENS COUNTY: RUSSIA: C-4 Road. Primorskiy territory, suburb of Vladivostok, Forest and thickets. Phenology: Flowers. Origin: vicinity of Academic town, slope of Amur bay. Native. 43.111746°, 131.879751°; WGS 84, uncertainty: 10000 m., Source: GeoLocate, unknown s.n. Georef'd by Dan Post Common. Reference specimen for illustration in 9 Jun 2004 Revision of "Flora of the Pacific Northwest", WTU Herbarium, 2017. Phenology: Flowers. WTU-405499, WTU-V-016494 S. Kharkevich 643 with T. Buch 7 Jun 1991 Caprifoliaceae WTU-336420, WTU-V-032496 Lonicera conjugialis Kellogg U.S.A., IDAHO, SHOSHONE COUNTY: Caprifoliaceae Coeur d'Alene National Forest. Border of Idaho and Montana (Sanders County). Forest Road Lonicera interrupta Benth. 430, 0.4 miles north of Jordan Summit. Elev. 1552 m. U.S.A., CALIFORNIA, SISKIYOU COUNTY: 47.94377° N, 116.00731° W; Source: GPS Along East Callahan Road, vicinity Callahan. In fruit; fruits red. Phenology: Fruits. Origin: 41.309588°, -122.801418°; WGS 84, Native. uncertainty: 10000 m., Source: GeoLocate, Georef'd by Dan Post Richard G. Olmstead 2016-31 On rocky outcrop with shrubs and forbs. with Sheila Olmstead, Evan Kelly, Veanna Phenology: Flowers. Origin: Native. Willard, Mike Carlson, Wendy McClure, Regina Johnson, Jim Duemmel, Richard Robohm, Terri David Gross 914 Knoke 28 May 2013 8 Jul 2016 WTU-413898, WTU-V-029349 WTU-420765, WTU-V-048925 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, PEND OREILLE U.S.A., WASHINGTON, PEND OREILLE COUNTY: COUNTY: Mill Creek Drainage, Selkirk Mountains. 0.5 Mill Creek Drainage, Selkirk Mountains. 0.5 miles east of LeClerc Creek Highway on Mill miles east of LeClerc Creek Highway on Mill Creek Road. Creek road. T35N R44E S33 SW; WGS 84, uncertainty: T35N R44E S33 SW; WGS 84, uncertainty: 1500 m., Source: Digital Map, Georef'd by 969 m., Source: Digital Map, Georef'd by Crystal Shin Crystal Shin Open east aspect with snowberry, Ponderosa Open east aspect with snowberry, Ponderosa pine, ninebark. Trumpet honeysuckle. pine, ninebark. Trumpet honeysuckle. Phenology: Flowers. Origin: Native. Cindy Talbott 266 Cindy Talbott 266 6 Jun 1978 6 Jun 1978 WTU-413252, WTU-V-028444 WTU-413258, WTU-V-028434 Caprifoliaceae Caprifoliaceae Lonicera utahensis Wats. f. ebractulata (Rydb.) St. J. Lonicera utahensis S. Watson U.S.A., WASHINGTON, STEVENS COUNTY: U.S.A., WASHINGTON, FERRY COUNTY: Between Rocky Creek and Meadow Creek. Western Colville National Forest and Vicinity: T37N R41E; WGS 84, uncertainty: 2000 m., Silver Bell Mine road, circa 7 air miles northwest Source: Digital Map, Georef'd by Crystal Shin of Republic; roadside on Forest road 514 at Growing in Abies grandis-Pachistima myrsinites Silver Bell Mine turnoff. habitat type. Phenology: Flowers. Origin: Native. Elev. 5100 ft. T37N R32E S30 SW1/4; WGS 84, T. E. Burke 71-11 uncertainty: 600 m., Source: Digital Map, Georef'd by Joseph Stormer 14 May 1971 Both sides of road; Abies lasiocarpa. Phenology: Flowers. Origin: Native. WTU-413260, WTU-V-028437 Jean Wood 471 28 May 1995 Caprifoliaceae WTU-413262, WTU-V-028435 Lonicera utahensis S. Watson U.S.A., IDAHO, BOUNDARY COUNTY: Kaniksu [National Forest] administered by Caprifoliaceae Colville [National Forest]. Little Snowy Top. Elev. 6400-6800 ft. Lonicera involucrata (Richardson) Banks ex Spreng. var. T65N R5W [S]20 NW1/4 RM involucrata Associated plants: Alpine fir/Luzula hitchcockii. Phenology: Flowers. Origin: Native. U.S.A., WASHINGTON, FERRY COUNTY: Western Colville National Forest and Vicinity: RCP 604 Sanpoil River Drainage: shores of Long Lake, circa 10 air miles of south-southwest of 23 Aug 1974 Republic. Elev. 3500 ft. WTU-413261, WTU-V-028433 T35N R32E S28. Also S33; WGS 84, uncertainty: 1000 m., Source: Digital Map, Georef'd by Sara Shiina Marly creek on lakeshore. Phenology: Flowers. Caprifoliaceae Origin: Native. Lonicera periclymenum L. Jean Wood 755 U.S.A., WASHINGTON, THURSTON COUNTY: 3 Jun 1995 Tumwater, Pioneer St. SW between Tumwater Hill and Barnes Lake. WTU-413251, WTU-V-028442 Elev. 200 ft. 47.0098° N, 122.91782° W; Source: Specimen Label Silt loam. With Alnus rubra, Populus trichocarpa, Hedera helix, Fraxinus latifolia, Spiraea douglasii. Uncommon. Flowers pink and white, fade to yellow. Phenology: Flowers. Origin: Introduced. Regina Johnson 2016-016 21 May 2016 WTU-414384, WTU-V-030586 Caprifoliaceae Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., OREGON, MULTNOMAH COUNTY: U.S.A., WASHINGTON, PIERCE COUNTY: Forest Park BPA Road. Lower section adjacent JBLM, Artillery Impact Area, Mortar Point 3, next to Nisqually Lake. to Hwy 30. Elev. 84 m. 45.6105° N, 122.7976° W; Source: Specimen 47.03032° N, 122.62275° W; WGS 84, Source: GPS Label Occasional under Quercus garryana, clambering over Relatively closed canopy in remnant Quercus Symphoricarpos albus. Flowers orange. Flowers: Bright orange. garryana woodland. Quercus garryana, Origin: Native. Pseudotsuga, Elymus glaucus, Geranium robertianum, Bromus carinatus, Rubus ursinus, Peter W. Dunwiddie PWD 16-97 23 May 2016 Hedera spp. Flowers. Small roadside with Sarah Krock, Amber Martens population damaged by herbicides. Phenology: Flowers. Origin: Native. Herbarium: WTU Dominic Maze 268 Caprifoliaceae 15 Jun 2016 Lonicera involucrata (Richardson) Banks ex Spreng. WTU-417340, WTU-V-037166 U.S.A., WASHINGTON, PIERCE COUNTY: JBLM, TA-2, Hanlet Marsh. Elev. 66 m. Caprifoliaceae 47.10414° N, 122.61572° W; WGS 84, Source: GPS Several shrubs at edge of wetland, with Phalaris arundinacea. Lonicera morrowii A. Gray Origin: Native. U.S.A., NEW YORK, DUTCHESS COUNTY: Peter Dunwiddie PWD-16-428 27 Jul 2016 1 mile south of Wassaic. with Sarah Krock, Peter Zika, Amber Martens 41.785511°, -73.55°; WGS 84, uncertainty: 2639 m., Source: GeoLocate, Georef'd by Herbarium: WTU Dan Post Wooded roadside. Bush 1 meter tall. Origin: Introduced. Caprifoliaceae Gregory E. Hyde 80 Lonicera ciliosa (Pursh) Poir. ex DC. 13 Aug 1968 U.S.A., WASHINGTON, MASON COUNTY: Olympic National Forest. Unknown spur from FS WTU-408588, WTU-V-024328 Road 23, approximately 5km from that junction. Elev. 491 m. 47.39626° N, 123.34125° W; WGS 84, Caprifoliaceae uncertainty: 100 m., Source: GPS Southeast-facing rocky bald with Polystichum Lonicera sp. munitum and flowering Amelanchier alnifolia and Castilleja hispida; sterile Sedum oreganum U.S.A., WASHINGTON, KING COUNTY: common. Climbing up Pinus monticola; corollas Arboretum, University of Washington. orange; climbing vine to 3 meters. Phenology: 47.636669°, -122.29551°; WGS 84, Flowers. Origin: Native. uncertainty: 1250 m., Source: Digital Map, Georef'd by Ibro Polovina David Giblin 5872 Phenology: Flowers. Cultivated. 28 Apr 2016 Bengt Oxelman 2331 WTU-420929, WTU-V-049770 1998 WTU-410610, WTU-V-024631 Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. var. Caprifoliaceae involucrata Lonicera tatarica L. U.S.A., IDAHO, SHOSHONE COUNTY: Panhandle National Forest. St. Joe Ranger U.S.A., WASHINGTON, SPOKANE COUNTY: District. Spruce Tree Campground. Along paved Centennial Trail State Park, west Elev. 699 m. of Harvard Road. 47.03416° N, 115.34964° W; WGS 84, Elev. 2020 ft. Source: GPS 47.68196° N, 117.1121° W; Source: Meadow and understory in campground. Specimen Label Flowers yellow; small shrub. Phenology: Fruits red. [Original label has Lonicera tartarica Flowers. Origin: Native. L.] Phenology: Fruits. Origin: Introduced. R. G. Olmstead 2018-17 Jeff Walker 1072 27 Jun 2018 10 Jul 2018 WTU-420587, WTU-V-048214 WTU-418456, WTU-V-039173 Caprifoliaceae Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. var. Lonicera ciliosa (Pursh) Poir. ex DC. involucrata U.S.A., WASHINGTON, KITSAP COUNTY: U.S.A., OREGON, BAKER COUNTY: Keyport Saltwater Park. Dogfish Bay. At outlet of Van Patten Lake. Elev. 1 m. Elev. 2252 m. 47.70068° N, 122.62716° W; WGS 84, uncertainty: 50 m., 44.95664° N, 118.184235° W; WGS 84, Source: GPS uncertainty: 10 m., Source: GPS Saltwater shoreline dominated by Jaumea carnosa and Salicornia Old, small, earthen dam at lake nutlet; mesic perennis. Scattered in adjacent forest edge; vine; corollas orange. soil, with Salix, Lonicera involucrata, Juncus Phenology: Flowers. Origin: Native. balticus, Trifolium repen, among subalpine conifer forest. Corolla yellowish; shrubs 1-2 David Giblin 6668 9 May 2019 meters tall. Phenology: Flowers. Origin: Native. Herbarium: WTU Makaila Wood 2018-1 13 Jul 2018 Caprifoliaceae WTU-421973, WTU-V-055322 Lonicera periclymenum L. U.S.A., WASHINGTON, PACIFIC COUNTY: Cranberry Road near Birch Street, 100 m N of Caprifoliaceae Clam Lake. Elev. 6 m. Lonicera utahensis S. Watson 46.3961° N, 124.0401° W; Source: Specimen Label U.S.A., OREGON, UNION COUNTY: Sunny damp thickets, locally common, with West side of FS Road 43 near Anthony Butte. Picea sitchensis, Alnus rubra, Malus fusca, Salix Elev. 2037 m. hookeriana, Gaultheria shallon, Ilex aquifolium, 45.04581° N, 118.23094° W; WGS 84, Rubus ursinus. Twining naturalized vines, uncertainty: 100 m., Source: GPS flowers dull pinkish-red outside, yellow-white South-facing slope dominated by Artemisia within, anthers orange, stigma green, 4 petal tridentata with Pinus contorta and Pseudotsuga lobes erect, 1 reflexed, fruit bright red, globose, menziesii forming the adjacent forest. Scattered shiny, 6-7 mm diameter. along forest edge; corollas white. Phenology: Flowers. Origin: Native. Peter F. Zika 29842 22 Oct 2019 David Giblin 6204 WTU-422998, WTU-V-059586 15 Jul 2018 Caprifoliaceae WTU-423125, WTU-V-060712 Lonicera periclymenum L. U.S.A., OREGON, LINN COUNTY: Caprifoliaceae South Santiam Wetland Mitigation Bank. 0.85 km south of Lacomb Drive; 8.2 km WSW of Lonicera ciliosa (Pursh) Poir. ex DC. Lacomb. Elev. 111 m. U.S.A., WASHINGTON, YAKIMA COUNTY: 44.56924° N, 123.84218° W; Source: Highway 410 approximately 1 kilometer east of Specimen Label, Invalid coordinates (located Indian Flat parking area; both sides of road. outside county bounds). Elev. 817 m. Farm field hedgerow; with Rosa nutkana, 46.98452° N, 121.12172° W; WGS 84, Crataegus gaylussacia, Rubus ursinus, Holcus uncertainty: 50 m., Source: GPS lanatus. Upright, shrubby, woody vine; South-facing slope with seep areas and scrambling on other vegetation and the adjacent dry upland; Populus trichocarpa, Pinus fenceline; two meters tall; flowers deep pink-red, ponderosa, and Eriogonum compositum maturing pinkish-white to cream-yellow. common. Uncommon on rocky outcropping; corollas orange. Phenology: Flowers. Origin: R. E. Brainerd 2749 Native. 21 May 2018 David Giblin 4885 WTU-422199, WTU-V-057755 18 Jun 2013 WTU-425285, WTU-V-069234 Caprifoliaceae Caprifoliaceae Lonicera periclymenum L. Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata U.S.A., WASHINGTON, KING COUNTY: Swift Avenue South, west side of road, approximately 5 meters U.S.A., WASHINGTON, ISLAND COUNTY: south of the intersection with the I-5 exit ramp. Hobbit Trail, Camano Island shrub wetland (former estuarine 47.550877° N, 122.313645° W; WGS 84, uncertainty: 8 m., wetland). Source: Digital map, Georef'd by David Giblin 48.213213° N, 122.448391° W Disturbed, open, roadside slope dominated by Rubus bifrons to 3 Saturated - moist, part sun, scrub-shrub swamp - former estuary. meters tall that is just coming into bloom. Sprawling, vining shrub Common, shrub, rhizomes, erect; yellow flowers. Phenology: climbing over top of the Rubus bifrons plants; flowers pleasantly Flowers. Origin: Native. fragrant; this is the southernmost sections of a population that occurs intermittently for 1 kilometer south along the chain-link fence Patricia Johnson NWC21-WA-10017-Q22 26 May 2021 on the west side of Swift Avenue South and South Albro Street. Phenology: Flowers. Origin: Introduced. Herbarium: WTU David Giblin 7411 21 May 2020 WTU-WTU

Caprifoliaceae Lonicera pileata Oliv. U.S.A., WASHINGTON, THURSTON COUNTY: Woodard Bay Natural Area, Weyer Point. Elev. 10 ft. 47.13283° N, 122.84014° W; WGS 84, Source: Digital map Lowland coniferous forest on Puget Sound bluff. Sandy soil. Uncommon - single large specimen, rooting branches. With Pseudotsuga menziesii, Tsuga heterophylla, Acer macrophyllum, Gaultheria shallon, Arbutus menziesii. Phenology: Vegetative. Origin: Introduced. Regina Johnson 2020-099 14 Aug 2020 Herbarium: WTU

Caprifoliaceae Lonicera involucrata (Richardson) Banks ex Spreng. var. involucrata U.S.A., WASHINGTON, FERRY COUNTY: Colville National Forest. Bangs Mountain Road approximately 5 km south of Highway 20. Elev. 865 m. 48.56171° N, 118.25319° W; WGS 84, uncertainty: 20 m., Source: GPS Mixed conifer forest with Adenocaulon bicolor and Linnaea borealis in the understory. Uncommon in understory; shrub to 2 meters tall; corollas yellow. Phenology: Flowers. Origin: Native. David Giblin 6758 22 May 2019 Herbarium: WTU

Caprifoliaceae Lonicera ciliosa (Pursh) Poir. ex DC. U.S.A., WASHINGTON, YAKIMA COUNTY: Okanogan-Wenatchee National Forest. West end of Clear Lake, west side of bridge, north side of small stream. Elev. 921 m. 46.70711° N, 121.19588° W; WGS 84, uncertainty: 20 m., Source: GPS Riparian corridor with Alnus rubra and Erythranthe guttata common. Scandent vine on downed log; corollas orange; uncommon. Phenology: Flowers. Origin: Native. David Giblin 7648 15 Jul 2020 Herbarium: WTU