The Green Machine Understanding the Seed Plants (Gymnosperms) 22
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Chemical Element Concentrations of Cycad Leaves: Do We Know Enough?
horticulturae Review Chemical Element Concentrations of Cycad Leaves: Do We Know Enough? Benjamin E. Deloso 1 , Murukesan V. Krishnapillai 2 , Ulysses F. Ferreras 3, Anders J. Lindström 4, Michael Calonje 5 and Thomas E. Marler 6,* 1 College of Natural and Applied Sciences, University of Guam, Mangilao, GU 96923, USA; [email protected] 2 Cooperative Research and Extension, Yap Campus, College of Micronesia-FSM, Colonia, Yap 96943, Micronesia; [email protected] 3 Philippine Native Plants Conservation Society Inc., Ninoy Aquino Parks and Wildlife Center, Quezon City 1101, Philippines; [email protected] 4 Plant Collections Department, Nong Nooch Tropical Botanical Garden, 34/1 Sukhumvit Highway, Najomtien, Sattahip, Chonburi 20250, Thailand; [email protected] 5 Montgomery Botanical Center, 11901 Old Cutler Road, Coral Gables, FL 33156, USA; [email protected] 6 Western Pacific Tropical Research Center, University of Guam, Mangilao, GU 96923, USA * Correspondence: [email protected] Received: 13 October 2020; Accepted: 16 November 2020; Published: 19 November 2020 Abstract: The literature containing which chemical elements are found in cycad leaves was reviewed to determine the range in values of concentrations reported for essential and beneficial elements. We found 46 of the 358 described cycad species had at least one element reported to date. The only genus that was missing from the data was Microcycas. Many of the species reports contained concentrations of one to several macronutrients and no other elements. The cycad leaves contained greater nitrogen and phosphorus concentrations than the reported means for plants throughout the world. Magnesium was identified as the macronutrient that has been least studied. -
Plant Evolution an Introduction to the History of Life
Plant Evolution An Introduction to the History of Life KARL J. NIKLAS The University of Chicago Press Chicago and London CONTENTS Preface vii Introduction 1 1 Origins and Early Events 29 2 The Invasion of Land and Air 93 3 Population Genetics, Adaptation, and Evolution 153 4 Development and Evolution 217 5 Speciation and Microevolution 271 6 Macroevolution 325 7 The Evolution of Multicellularity 377 8 Biophysics and Evolution 431 9 Ecology and Evolution 483 Glossary 537 Index 547 v Introduction The unpredictable and the predetermined unfold together to make everything the way it is. It’s how nature creates itself, on every scale, the snowflake and the snowstorm. — TOM STOPPARD, Arcadia, Act 1, Scene 4 (1993) Much has been written about evolution from the perspective of the history and biology of animals, but significantly less has been writ- ten about the evolutionary biology of plants. Zoocentricism in the biological literature is understandable to some extent because we are after all animals and not plants and because our self- interest is not entirely egotistical, since no biologist can deny the fact that animals have played significant and important roles as the actors on the stage of evolution come and go. The nearly romantic fascination with di- nosaurs and what caused their extinction is understandable, even though we should be equally fascinated with the monarchs of the Carboniferous, the tree lycopods and calamites, and with what caused their extinction (fig. 0.1). Yet, it must be understood that plants are as fascinating as animals, and that they are just as important to the study of biology in general and to understanding evolutionary theory in particular. -
Number of Living Species in Australia and the World
Numbers of Living Species in Australia and the World 2nd edition Arthur D. Chapman Australian Biodiversity Information Services australia’s nature Toowoomba, Australia there is more still to be discovered… Report for the Australian Biological Resources Study Canberra, Australia September 2009 CONTENTS Foreword 1 Insecta (insects) 23 Plants 43 Viruses 59 Arachnida Magnoliophyta (flowering plants) 43 Protoctista (mainly Introduction 2 (spiders, scorpions, etc) 26 Gymnosperms (Coniferophyta, Protozoa—others included Executive Summary 6 Pycnogonida (sea spiders) 28 Cycadophyta, Gnetophyta under fungi, algae, Myriapoda and Ginkgophyta) 45 Chromista, etc) 60 Detailed discussion by Group 12 (millipedes, centipedes) 29 Ferns and Allies 46 Chordates 13 Acknowledgements 63 Crustacea (crabs, lobsters, etc) 31 Bryophyta Mammalia (mammals) 13 Onychophora (velvet worms) 32 (mosses, liverworts, hornworts) 47 References 66 Aves (birds) 14 Hexapoda (proturans, springtails) 33 Plant Algae (including green Reptilia (reptiles) 15 Mollusca (molluscs, shellfish) 34 algae, red algae, glaucophytes) 49 Amphibia (frogs, etc) 16 Annelida (segmented worms) 35 Fungi 51 Pisces (fishes including Nematoda Fungi (excluding taxa Chondrichthyes and (nematodes, roundworms) 36 treated under Chromista Osteichthyes) 17 and Protoctista) 51 Acanthocephala Agnatha (hagfish, (thorny-headed worms) 37 Lichen-forming fungi 53 lampreys, slime eels) 18 Platyhelminthes (flat worms) 38 Others 54 Cephalochordata (lancelets) 19 Cnidaria (jellyfish, Prokaryota (Bacteria Tunicata or Urochordata sea anenomes, corals) 39 [Monera] of previous report) 54 (sea squirts, doliolids, salps) 20 Porifera (sponges) 40 Cyanophyta (Cyanobacteria) 55 Invertebrates 21 Other Invertebrates 41 Chromista (including some Hemichordata (hemichordates) 21 species previously included Echinodermata (starfish, under either algae or fungi) 56 sea cucumbers, etc) 22 FOREWORD In Australia and around the world, biodiversity is under huge Harnessing core science and knowledge bases, like and growing pressure. -
Identification of Conifer Trees in Iowa This Publication Is Designed to Help Identify the Most Common Trees Found in Iowa
Identification of Conifer Trees in Iowa This publication is designed to help identify the most common trees found in Iowa. It is based on vegetative characteristics including leaves, fruit, and bark. It is neither complete nor without possible oversights. Separate species are grouped by similar characteristics, mainly based on type and arrangement of leaves. These groups are; awl- or scale- like needles; single needles, flattened with rounded tips; single needles, square in cross section, with pointed tips; and needles in bundles or fasticles of two or more. Remember, vegetative character- istics are quite variable; use more than one specimen for comparison. Awl- or scale-like needles Juniperus Virginiana Eastern Red Cedar Leaves are dark green; leaves are both awl- and scale-like; cone is dark blue and berry-like. Thuja occidentalis Northern White Cedar Leaves are flattened and only of the scale type; cones have 4-6 scales; foliage is light green. Juniperus communis Common Juniper Leaves are awl shaped; cone is dark blue and berry-like. Pm-1383 | May 1996 Single needles, flattened with rounded tips Pseudotsuga menziesii Douglas Fir Needles occur on raised pegs; 3/4-11/4 inches in length; cones have 3-pointed bracts between the cone scales. Abies balsamea Abies concolor Balsam Fir White (Concolor) Fir Needles are blunt and notched at Needles are somewhat pointed, the tip; 3/4-11/2 inches in length. curved towards the branch top and 11/2-3 inches in length; silver green in color. Single needles, Picea abies Norway Spruce square in cross Needles are 1/2-1 inch long; section, with needles are dark green; foliage appears to droop or weep; cone pointed tips is 4-7 inches long. -
Chlorophyta Is a Division of Green Algae, Informally Called
Chlorophyta is a division of green algae, informally waters of the Sargasso Sea. Many brown algae, such as called chlorophytes. The name is used in two very members of the order Fucales, commonly grow along different senses so that care is needed to determine the rocky seashores. Some members of the class are used as use by a particular author. In older classification food for humans. systems, it refers to a highly paraphyletic group of all Worldwide there are about 1500–2000 species of brown the green algae within the green plants (Viridiplantae), algae.[4] Some species are of sufficient commercial and thus includes about 7,000 species [4] [5] of mostly importance, such as Ascophyllum nodosum , that they aquatic photosynthetic eukaryotic organisms. Like the have become subjects of extensive research in their own land plants (bryophytes and tracheophytes), green algae right.[5] [4] contain chlorophylls a and b, and store food as starch Brown algae belong to a very large group, the in their plastids. Heterokontophyta, a eukaryotic group of organisms In newer classifications, it refers to one of the two distinguished most prominently by having chloroplasts clades making up the Viridiplantae, which are the surrounded by four membranes, suggesting an origin chlorophytes and the streptophytes or charophytes.[6][7] from a symbiotic relationship between a basal In this sense it includes only about 4,300 species.[3] eukaryote and another eukaryotic organism. Most brown algae contain the pigment fucoxanthin, which is responsible for the distinctive greenish-brown color that The red algae, or Rhodophyta ( / r o ʊ ˈ d ɒ f ɨ t ə / or / gives them their name. -
The Effects of Sulphur Dioxide on Selected Hepatics" (1978)
Eastern Illinois University The Keep Masters Theses Student Theses & Publications 1978 The ffecE ts of Sulphur Dioxide on Selected Hepatics Steven L. Gatchel Eastern Illinois University This research is a product of the graduate program in Botany at Eastern Illinois University. Find out more about the program. Recommended Citation Gatchel, Steven L., "The Effects of Sulphur Dioxide on Selected Hepatics" (1978). Masters Theses. 3192. https://thekeep.eiu.edu/theses/3192 This is brought to you for free and open access by the Student Theses & Publications at The Keep. It has been accepted for inclusion in Masters Theses by an authorized administrator of The Keep. For more information, please contact [email protected]. PAPF-:R CERTIFICATE #2 TO: Graduate Degree Candidates who have written formal theses. SUBJECT: Permission to reproduce theses. The University Library is receiving a ' number of requests from other institutions asking permission to reproduce dissertations for inclusion in their library holdings. Although no copyright laws are involved, we feel that professional courtesy demands that permission be obtained from the author before we allow theses to be copied. Please sign one of the following statements: Booth Library of Eastern Illinois University has my permission to lend my thesis to a reputable college or university for the purpose of copying it for inclusion in that institution's library or research holdings. Date Author I respectfully request Booth Library of .Eastern Illinois University not allow my thesis be reproduced because---------------- Date Author pdm THE EFFECTS OF SULPHUR DIOXIDE ON SELECTED HEPATICS (TITLE} BY Steven L. Gatchel THESIS SUBMITTED IN PARTIAL FULFILLMENT OF THE REQUIREMENTS FOR THE DEGREE OF Master of Science IN THE GRADUATE SCHOOL, EASTERN ILLINOIS UNIVERSITY CHARLESTON, ILLINOIS 1978 I HEREBY RECOMMEND THIS THESIS BE ACCEPTED AS FULFILLING THIS PART OF THE GRADUATE DEGREE CITED ABOVE ol}-~ d2/, 19"/f DATE ADVISER I/ 'Ouue~2\ 1\l~ DATE ' ~RTMENT WHEAD THE EFFECTS OF SULPHUR DIOXIDE ON SELECTED HEPATICS BY STEVEN L. -
Cycad Forensics: Tracing the Origin of Poached Cycads Using Stable Isotopes, Trace Element Concentrations and Radiocarbon Dating Techniques
Cycad forensics: Tracing the origin of poached cycads using stable isotopes, trace element concentrations and radiocarbon dating techniques by Kirsten Retief Supervisors: Dr Adam West (UCT) and Ms Michele Pfab (SANBI) Submitted in partial fulfillment of the requirements for the degree of Masters of Science in Conservation Biology 5 June 2013 Percy FitzPatrick Institution of African Ornithology, UniversityDepartment of Biologicalof Cape Sciences Town University of Cape Town, Rondebosch Cape Town South Africa 7701 i The copyright of this thesis vests in the author. No quotation from it or information derived from it is to be published without full acknowledgement of the source. The thesis is to be used for private study or non- commercial research purposes only. Published by the University of Cape Town (UCT) in terms of the non-exclusive license granted to UCT by the author. University of Cape Town Table of Contents Acknowledgements iii Plagiarism declaration iv Abstract v Chapter 1: Status of cycads and background to developing a forensic technique 1 1. Why are cycads threatened? 2 2. Importance of cycads 4 3. Current conservation strategies 5 4. Stable isotopes in forensic science 7 5. Trace element concentrations 15 6. Principles for using isotopes as a tracer 15 7. Radiocarbon dating 16 8. Cycad life history, anatomy and age of tissues 18 9. Recapitulation 22 Chapter 2: Applying stable isotope and radiocarbon dating techniques to cycads 23 1. Introduction 24 2. Methods 26 2.1 Sampling selection and sites 26 2.2 Sampling techniques 30 2.3 Processing samples 35 2.4 Cellulose extraction 37 2.5 Oxygen and sulphur stable isotopes 37 2.6 CarbonUniversity and nitrogen stable of isotopes Cape Town 38 2.7 Strontium, lead and elemental concentration analysis 39 2.8 Radiocarbon dating 41 2.9 Data analysis 42 3. -
Aquatic and Wet Marchantiophyta, Order Metzgeriales: Aneuraceae
Glime, J. M. 2021. Aquatic and Wet Marchantiophyta, Order Metzgeriales: Aneuraceae. Chapt. 1-11. In: Glime, J. M. Bryophyte 1-11-1 Ecology. Volume 4. Habitat and Role. Ebook sponsored by Michigan Technological University and the International Association of Bryologists. Last updated 11 April 2021 and available at <http://digitalcommons.mtu.edu/bryophyte-ecology/>. CHAPTER 1-11: AQUATIC AND WET MARCHANTIOPHYTA, ORDER METZGERIALES: ANEURACEAE TABLE OF CONTENTS SUBCLASS METZGERIIDAE ........................................................................................................................................... 1-11-2 Order Metzgeriales............................................................................................................................................................... 1-11-2 Aneuraceae ................................................................................................................................................................... 1-11-2 Aneura .......................................................................................................................................................................... 1-11-2 Aneura maxima ............................................................................................................................................................ 1-11-2 Aneura mirabilis .......................................................................................................................................................... 1-11-7 Aneura pinguis .......................................................................................................................................................... -
Ecophysiology of Four Co-Occurring Lycophyte Species: an Investigation of Functional Convergence
Research Article Ecophysiology of four co-occurring lycophyte species: an investigation of functional convergence Jacqlynn Zier, Bryce Belanger, Genevieve Trahan and James E. Watkins* Department of Biology, Colgate University, Hamilton, NY 13346, USA Received: 22 June 2015; Accepted: 7 November 2015; Published: 24 November 2015 Associate Editor: Tim J. Brodribb Citation: Zier J, Belanger B, Trahan G, Watkins JE. 2015. Ecophysiology of four co-occurring lycophyte species: an investigation of functional convergence. AoB PLANTS 7: plv137; doi:10.1093/aobpla/plv137 Abstract. Lycophytes are the most early divergent extant lineage of vascular land plants. The group has a broad global distribution ranging from tundra to tropical forests and can make up an important component of temperate northeast US forests. We know very little about the in situ ecophysiology of this group and apparently no study has eval- uated if lycophytes conform to functional patterns expected by the leaf economics spectrum hypothesis. To determine factors influencing photosynthetic capacity (Amax), we analysed several physiological traits related to photosynthesis to include stomatal, nutrient, vascular traits, and patterns of biomass distribution in four coexisting temperate lycophyte species: Lycopodium clavatum, Spinulum annotinum, Diphasiastrum digitatum and Dendrolycopodium dendroi- deum. We found no difference in maximum photosynthetic rates across species, yet wide variation in other traits. We also found that Amax was not related to leaf nitrogen concentration and is more tied to stomatal conductance, suggestive of a fundamentally different sets of constraints on photosynthesis in these lycophyte taxa compared with ferns and seed plants. These findings complement the hydropassive model of stomatal control in lycophytes and may reflect canaliza- tion of function in this group. -
X. the Conifers and Ginkgo
X. The Conifers and Ginkgo Now we turn our attention to the Coniferales, another great assemblage of seed plants. First let's compare the conifers with the cycads: Cycads Conifers few apical meristems per plant many apical meristems per plant leaves pinnately divided leaves undivided wood manoxylic wood pycnoxylic seeds borne on megaphylls seeds borne on stems We should also remember that these two groups have a lot in common. To begin with, they are both groups of woody seed plants. They are united by a small set of derived features: 1) the basic structure of the stele (a eustele or a sympodium, two words for the same thing) and no leaf gaps 2) the design of the apical meristem (many initials, subtended by a slowly dividing group of cells called the central mother zone) 3) the design of the tracheids (circular-bordered pits with a torus) We have three new seed plant orders to examine this week: A. Cordaitales This is yet another plant group from the coal forest. (Find it on the Peabody mural!) The best-known genus, Cordaites, is a tree with pycnoxylic wood bearing leaves up to about a foot and a half long and four inches wide. In addition, these trees bore sporangia (micro- and mega-) in strobili in the axils of these big leaves. The megasporangia were enclosed in ovules. Look at fossils of leaves and pollen-bearing shoots of Cordaites. The large, many-veined megaphylls are ancestral to modern pine needles; the shoots are ancestral to pollen-bearing strobili of modern conifers. 67 B. -
Laurentian-Acadian Alkaline Conifer-Hardwood Swamp
Laurentian-Acadian Alkaline Conifer-Hardwood Swamp Macrogroup: Northern Swamp yourStateNatural Heritage Ecologist for more information about this habitat. This is modeledmap a distributiononbased current and is data nota substitute for field inventory. based Contact © Elizabeth Thompson (Vermont Land Trust) Description: A forested swamp of alkaline wetlands associated with limestone or other calcareous substrate in the northern part of the glaciated northeast. Northern white cedar is often present and may dominate the canopy or be mixed with other conifers or with deciduous trees, most commonly red maple or black ash. Some examples can be almost entirely deciduous and dominated by black ash. Red-osier dogwood is a common shrub. The herb layer tends to be more diverse than in acidic swamps, due to higher pH and nutrient level. Small open fenny areas may occur within the wetland. The moss layer is often extensive and diverse. Seepage may influence parts of the wetland, but the hydrology is State Distribution: CT, MA, ME, NH, NY, VT dominated by the basin setting. Total Habitat Acreage: 921,478 Ecological Setting and Natural Processes: Percent Conserved: 19.5% These forested wetlands are uncommon in the glaciated State State GAP 1&2 GAP 3 Unsecured northeast except in areas with extensive limestone or similar State Habitat % Acreage (acres) (acres) (acres) substrate. The substrate is typically mineral soil, but there ME 56% 520,121 14,203 60,307 445,611 may be some peat, and there is often direct contact with NY 38% 345,750 49,536 44,764 251,450 alkaline groundwater. VT 5% 43,899 1,177 4,786 37,935 NH 1% 7,363 2,054 1,013 4,295 MA 0% 4,261 643 1,267 2,350 CT 0% 86 0 0 86 Similar Habitat Types: Similar to North-Central Interior and Appalachian Rich Swamp, but with a flora characteristic of a cooler climate. -
Elemental Profiles in Cycas Micronesica Stems
plants Article Elemental Profiles in Cycas micronesica Stems Thomas E. Marler College of Natural and Applied Sciences, University of Guam, UOG Station, Mangilao, Guam 96923, USA; [email protected]; Tel.: +1-671-735-2100 Received: 24 August 2018; Accepted: 30 October 2018; Published: 1 November 2018 Abstract: Essential nutrients and metals have been quantified in stems of many tree species to understand the role of stems as storage and source organs. Little is known about stored stem resources of cycad tree species. Cycas micronesica tissue was collected from apical and basal axial regions of stems; and pith, vascular, and cortex tissues were separated into three radial regions. Leaves were also sampled to provide a comparison to stems. Minerals and metals were quantified in all tissues. Minerals and metals varied greatly among the six stem sections. Phosphorus varied more among the three radial sections than the other macronutrients, and zinc and nickel varied more than the other micronutrients. Stem carbon was less than and stem calcium was greater than expected, based on what is currently known tree stem concentrations in the literature. Elemental concentrations were generally greater than those previously reported for coniferous gymnosperm trees. Moreover, the stem concentrations were high in relation to leaf concentrations, when compared to published angiosperm and conifer data. The results indicated that the addition of more cycad species to the literature will improve our understanding of gymnosperm versus angiosperm stem nutrient relations, and that the non-woody cycad stem contains copious essential plant nutrients that can be mobilized and deployed to sinks when needed.