Diptera: Limoniidae)

Gonomyia (Gonomyia) Iucidula de Meijere and allies (Diptera: Limoniidae)

Jaroslav Stary

Stary, J. 2010: Gonomyia (Gonomyia) Iucidula de Meijere and allies (Diptera: Limoniidae). — Entomol. Fennica 21: 232—242. Gonomyia (Gonomyia) lucidula de Meijere, 1920 and G. (G.) abscondita Lackschewitz, 1935 are redescribed, and the lectotypes ofboth species are designated. Desciptions ofanother two species are provided, viz. G. (G.) securiformis sp. n. (Bulgaria, Czech Republic, Italy, Switzerland) and G. (G.) undiformis sp. n. (Slovakia). Male terminalia are illustrated and distributions are outlined for all the species.

J. Stary, Department ofZoology and Laboratory ofOrnithology, Faculty ofSci— ence 0ftlze Palacky University, tr. Svobody 26, CZ—77146 Olomouc, Czech Re— public; E—mail.' stary@prfnw. up0l. cz Received 10 February 2010, accepted 19 April 2010

1. Introduction regarded as a somewhat doubtful taxon, although mentioned in the literature from time to time, es- Gonomyia (Gonomyia) lucidula de Meijere, pecially, recently, by British authors (e.g. Stubbs 1920 was recorded from many European coun- 2001, Boardman 2007). tries (Oosterbroek 2009). Lackschewitz (1935) As early as in the 1970’s, I had examined the described G. (G.) abscondita, with emphasis on a types ofboth G. (G.) Iucidula and G. (G.) abscon— close relationship to G. (G.) Iucidula, but without dita and found details in the structure ofthe male specially comparing their male terminalia. A dis- terminalia that distinguished the two as valid spe- tinct pattern on the pleuron was mentioned by cies. The pigmentation of the parameres proved him as the only character distinguishing G. (G.) to be one of several species-specific differences, abscondita from G. (G.) Iucidula. Yet the respec- perhaps the most conspicuous, but by far not the tive descriptions (de Meijere 1920: 83, Figs 82a— most significant taxonomically. Having exam- b; Lackschewitz 1935: 13—14, Figs 7a—e) and the ined additional material, however, I discovered a key by Lackschewitz (1940: 5 8) to the European few variant specimens, which destroyed the un- Gonomyia Meigen, 1818 suggested at least one perturbed taxonomic situation, as it had appeared genital difference between the two species, initially. As a result, an almost finished manu- namely that the parameres are both darkly pig- script was postponed for further study, and it, in mented in G. (G.) Iucidula, whereas, in G. (G.) fact, remained somewhat forgotten. abscondita, the right, shorter paramere is pale. The variants in my collection gradually in- The first time since its description, G. (G.) creased in number, and it eventually turned out abscondita was reported by Stary & Rozkosny that these represent two additional species closely (1970) from Slovakia, but its validity was related to G. (G.) Iucidula. Thus, in the present doubted because of possible synonymy with G. paper, I redescribe G. (G.) Iucidula and G. (G.) (G.) Iucidula. G. (G.) abscondita has since been abscondita and designate the lectotypes of both ENTOMOL. FENNICA Vol. 21 ° Gonomyia (Gonomyia) lucidula and allies 233 species. I describe another two species, G. (G.) Gonomyia microdentata Bangerter 1947: 376 securiformis sp. n. and G. (G.) undz’formis sp. n. (description), Figs 19A—B (male terminalia). All the species are extremely similar in general appearance and variable in the pleural patterning, Redescription. Male. Small species within so external characters are only briefly mentioned Gonomyia. Wing length 4.3—5 .4 mm. Prescutum in the descriptions, and emphasis is laid on the brown, pleuron yellow with orange tinge, without structure of the male terminalia (see Figs 1—12). any distinct brown pattern, restrictedly patterned Females are so far indistinguishable and only with sulphur yellow. Central spot on pleuron, of- listed occasionally with syntopic and synchro- ten distinct in both G. (G.) abscondita and G. (G.) nous males in the Material examined sections. G. securz'formis sp. n., sometimes only indicated as (G.) securlformis sp. n. is confined to high alti- obscure yellow stain. In general resembling small tudes, and G. (G.) undz’formis sp. n. seems to oc- specimens of G. (G.) recta Tonnoir, 1920. cur on limestone substrates. Although ecological Male terminalia (Figs 1a—c). Outer gonosty- affinities of G. (G.) lucidula and G. (G.) abscon— lus pale, flattened laterally, comparatively short, dita were characterized as somewhat different only slightly exceeding inner gonostylus, gradu- (Boardman 2007), the two species are largely ally tapered to subacute tip. Inner gonostylus of syntopic based on the material available to me. It complex structure, in general divided into two is likely that additional species may belong to this parts: inner (medial) part pale, not especially complex. broad, with short beak-like projection tipped with The following acronyms ofmuseums and col- a few long setae; outer (lateral) part represented lections are used in the text: BMNH — The Natu- by irregularly bilobed blade, slightly darkened at ral History Museum [formerly British Museum tips ofboth lobes and tilted over inner part of in- (Natural History)], London, England, UK; JSO — ner gonostylus. Sharp dark tooth at outer base of

Collection ofJ . Stary, Olomouc, Czech Republic; inner gonostylus, provided with basal seta. Both MMBC — Moravské zemské muzeum, Brno, parameres darkly pigmented distally and pointed Czech Republic; NHMW — Naturhistorisches at tips. Aedeagus flattened laterally in distal part, Museum, Vienna, Austria; SDEI — Senckenberg forming rounded apical portion separated from Deutsches Entomologisches Institut, Munche- rest of aedeagus by deep subapical excision ven- berg, Germany; SMOC — Slezské zemské mu- trally; obtuse low projection proximal to zeum, Opava, Czech Republic; ZMAN — Zoolo- subapical excision (less developed than in G. (G.) gisch Museum, Universiteit van Amsterdam, abscondita). Distal part of aedeagal duct only Amsterdam, The Netherlands; ZMHB — Museum slightly arched, running along dorsal margin of fiir Naturkunde der Humboldt-Universitat, Ber- aedeagus and ending at posterodorsal edge ofapi- lin, Germany. cal portion. Female. Resembles male in general appearance, not reliably distinguished from females of 2. Descriptions other species. Type material examined. The species was de- 2.1. Gonomyia (Gonomyia) lucidula scribed from an unspecified number ofspecimens de Meijere, 1920 (Figs la—c) from Valkenburg (L.), Juni; Bunde, Juni; Oldenzaal, Juli; Nieuwersluis, Juni.” (de Meij ere Gonomyia lucidula de Meijere 1920: 83 (descrip- 1920: 83). I have examined four male syntypes tion), P1. 10, Figs 82a—b (male terminalia). from ZMAN, ofwhich only one, from the locality Valkenburg, here designated as lectotype, be- Gonomyia (Gonomyia) lucidula: Savchenko longs to G. (G.) lucidula. The other three speci- 1982: 290 (redescription) [not Figs 139/5, 140/5, mens (paralectotypes oflucidula), one from each copied from Edwards 1938]; Savchenko et al. remaining locality, proved to be G. (G.) abscon— 1992: 303 (Palaearctic catalogue); Stubbs 2001: 5 dita and are listed in the Other material examined (key), Fig. (male terminalia); Oosterbroek 2009 under the latter species. The lectotype was listed (electronic World catalogue). by de Jong (2000: 122). This, however, is not a

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Winterswijk, 19.vi.1921, 2 6; Ommen, 11.vi. based on overall shape of the aedeagus, despite 1921, 1 6; Lochem, vii.1916, 1 6 (all J.C.H. de the fact that both parameres were depicted as Meijere leg.) (ZMAN). Slovakia: Sagan, darkly pigmented; the aedeagal complex was de- 16.vi.1978, 1 6; Belianske Tatry Mts, Tatranska scribed correctly in the text as having the shorter Kotlina, Bela valley (750 m a.s.l.), 27.vii. 1974, 1 paramere pale (Edwards 1938: 1 1 1). The illustra- 6, 26.vi.1975, 1 6, 19.vi.1977, 1 6, 21.vi.1977, tions under G. (G.) lucidula by Coe (1950, Fig. 4 a, 22.vi. 1977, 1 a; Pol’ana Mts, Oiemy Potok 22b) and Podenas et al. (2006, Figs 14.12—14.13) (700 m a.s.l.), 15.vi.2007, 1 6 (at light); Ocova are just copies of the drawings by Edwards nr. Zvolen, Hucava valley (500 m a.s.l.), (1938). Savchenko distinguished G. (G.) lucidula 22.vi.2004, 1 6; Hostovice, Udava shores, and G. (G.) abscondita, copying, however, Ed- 16.vi.1986, 3 6; Bukovské vrchy [hills], Rusky wards’s (1938) figures and those of Stary & Potok, valley, 11.vii.1993, 1 6; Bukovské vrchy Rozkosny (1970), respectively (Savchenko [hills], Zboj, Zbojsky brook, 8.vii.1993, 2 6, 1982, Figs 139/5, 140/5 and 139/6, 140/6), thus, 10.vii.1993, 1 6 [the latter three localities listed in fact, presenting in both cases, what I think is G. by Stary 1995] (all J. Stary leg.) (JSO). (G.) abscondita (see above and Discussion of Discussion. Externally, G. (G.) lucidula can abscondita). Yet his records from Ukraine proba- be distinguished from the other related species bly are correct for both species. Stubbs (2001: 5) (except for G. (G.) undiformis sp. n.) by a yellow illustrated some distinguishing characters be- pleuron without any distinct brown pattern. This tween G. (G.) lucidula and G. (G.) abscondita. character, however, is variable to a certain degree Based on the figures by Bangerter (1947, Figs and cannot serve as a reliable criterion. The struc- 19A—B), G. (G.) microdentata Bangerter, 1947 is ture of the male terminalia is essential for identi- here confirmed, in accordance with Savchenko et fying all the species treated here. The following al. (1992), as a junior synonym of G. (G). luci— combination of characters is species-specific for dula. G. (G.) lucidula (Figs 1a—c): outer gonostylus Distribution. Gonomyia (Gonomyia) lucidula short, only slightly exceeding inner gonostylus, is reported as widely distributed in Europe, rang- gradually tapered to subacute tip; inner gonosty- ing to Transcaucasia (Oosterbroek 2009). Based lus with distinct tooth at outer base; both on the material examined here, it is known from parameres darkly pigmented distally and pointed Bulgaria, Czech Republic, Germany, Nether- at tips; aedeagus with apical portion generally lands, and Slovakia. Original figures confirm its rounded and aedeagal duct ending at its postero- occurrence in Switzerland (Bangerter 1947) and dorsal edge. It should be emphasized that the Great Britain (Stubbs 2001). Records from aedeagus alone clearly distinguishes the species Ukraine (Savchenko 1982) likewise are probably treated here. On the other hand, it appears that the correct (see Discussion of lucidula). Other re- pigmentation of the shorter paramere is less con- cords (Savchenko et al. 1992, Oosterbroek 2009) stant and less stable than that ofthe longer one. In need confirmation. the material of G. (G.) lucidula preserved in ethanol, for example, the shorter paramere is rather pale. 2.2. Gonomyia (Gonomyia) abscondita The original figures of G. (G.) lucidula by de Lackschewitz, 1935 (Figs 2a—c) Meijere (1920, P1. 63, Figs 82a—b) clearly show a distinct tooth at the outer base of the inner gono- Gonomyia abscondita Lackschewitz 1935: 13 stylus, although, in the Dutch material, including (description), Pl. 3, Figs 7a—d (male terminalia), the type series of G. (G.) lucidula, both G. (G.) 7e (wing). lucidula and G. (G.) abscondita are represented (see Material examined ofboth species). As to the Gonomyia (Gonomyia) abscondiia: Stary & drawings by Edwards (1938, Fig. 20b) assigned Rozkosny 1970: 114 (faun. records, note), Figs to G. (G.) lucidula, l have evaluated them and de- 63, 66, 69 (male terminalia); Savchenko 1982: termined that they belong to G. (G.) abscondita 291 (redescription), Figs 139/6, 140/6 (male ter- (see references below under G. (G.) abscondita) minalia) [copied from Stary & Rozkosny 1970];

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Female. Resembles male in general appear- Czech Republic: Bohemia: Doksy env., Obora, ance, not reliably distinguished from females of wetlands, 12.vii.2007, 1 8; gumava Mts, Borova other species. Lada, 8.vii.1992, 1 8 (all J. Stary leg.) (JSO). Type material examined. The species was de- Moravia: Hruby Jesenik Mts (= Jeseniky Mts), scribed from three specimens that were listed by Branna, Klepacsky brook (700 m a.s.l.), Lackschewitz (1935: 14) as follows: “Waliser 18.vii.2006, 1 8 (at light), “Dembauda” (900 m Alpen, Macugnaga, 6.VHI.00. 3 8 S2 (Typus!) a.s.l.), 8.viii.1972, 1 8, 18.vii.1973, 1 8, 14.vii. leg. Oldenberg”. This phrase, however, with the 1999, 2 8 (at light), 21.vii.1999, 2 8, 1 S? (at “Typus” (singular), does not make it possible to light), 9.vii.2002, 2 8 (at light); Hruby Jesenik distinguish whether one, and which one, or all Mts, Kouty nad Desnou, Divoka Desna valley, specimens were considered by Lackschewitz to “zamcisko” (970 m a.s.l.), 26.vi.2003, 2 8 (at be the “types”. In 1973, I examined one male light), 8.vii.2004, 2 8 (at light), 14.iX.2004, 1 8 from ZMHB with corresponding collecting data, (at light); Hruby Jesenik Mts, Vidly, “Skalni labelled by Lackschewitz as “Type”, now con- potok” (700 m a.s.l.), 20.vi.2000, 2 8 (at light), 3 firmed as being the single specimen of the type 8, 28.vi.2000, 1 8 (at light), 21.vi.2001, 4 8 (at series of G. (G.) abscondita in ZMHB (J. Ziegler, light), 2.vii.2002, 3 8 (at light); Hruby Jesenik pers. comm.). Recently, three additional, badly Mts, Praded (900—1,050 m a.s.l.), 26.vii.1994, 2 damaged specimens were traced in SDEI, from 8; Hruby Jesenik Mts, Filipovice (750 m a.s.l.), the same locality but with different dates of col- 17.viii.1982, 1 8; Velké Losiny, 13.vii.1977, 1 lection (”20.7.00” and “5.8.00”), all labelled by 8, 10.viii.1977, 1 8; Mala Moravka, Moravice Lackschewitz as “Type” (F. Menzel and J. shores, 28.viii.1997, 1 8; Moravicany, Ziegler, pers. comm.). Despite various discrepan- 1.vii.1972, 1 8; Jivova nr. Olomouc, 14.vi.1978, cies it is now clear that Lackschewitz had consid- 1 8; Smilov nr. Olomouc, 18.vi.1972, 1 8; Hruba ered equally several specimens (some possibly in Voda nr. Olomouc, 12.vii.1977, 1 8; Krcman nr. error) to be the types of G. (G.) abscondita. Olomouc, 30.vi.1977, 1 8, 2.viii.1978, 1 8; Hence, the specimens listed by him in his descrip- Slatinice, Kosir, 3.vii.1974, 1 8; Novy Dvfir nr. tion (Lackschewitz 1935: 14) are to be regarded Opava, 12.vi.1967, 1 8; Kletné nr. Suchdol nad as syntypes, and the one examined by me is desig- Odrou, 25.vii.1971, 1 8, 21.viii.1977, 1 8, nated here as lectotype. Lectotype 8 (by present 11.viii.1978, 1 8; Nové Oldrfivky, Odra valley, designation): Italy, Piemonte, Macugnaga, 22.vi.1993, 2 8; Libava env., Stara Voda, 6.viii. 1900 (L. Oldenberg leg.) (ZMHB), labelled 27.viii.1978, 1 8; Libava env., Slavkov, 17.vii. “Macugnaga / Oldenberg / 6.8.00”, “coll. Olden- 1997, 1 8; Nydek, Hluchova valley, 20.vi.1996, berg”, “G. abscondita / Type nov. sp. / det. 1 8; Ticha nr. Frenstat pod Radhostem, 30.vi. Lacksch.” (red label). Labelled as lectotype by 1994, 1 8; Moravskoslezské Beskydy Mts, the present author (”LECTOTYPE / Gonomyia Kosariska, 26.vi.1997, 1 8 (at light); Morav- (s. str.) / abscondita Lacksch. 8 / J. Stary 2009”, skoslezské Beskydy Mts, Tanecnice (900—1,000 red label). The specimen is pinned, in bad condi- m a.s.l.), 24.vii.1984, 1 8, 12.vii.1990, 1 8; tion, without legs and with right wing glued to the Moravskoslezské Beskydy Mts, Prostredni pin (state of 1973). Terminalia dissected by Becva (500—600 m a.s.l.), 30.vii.1992, 1 8, Lackschewitz and placed in Canada balsam be- 18.vi.1993, 1 8, 23.vi.1993, 2 8, 30.vi.1994, 1 tween celluloid slides, pinned with the specimen. 8; Moravskoslezské Beskydy Mts, Malinova The lectotype is designated here to stabilize the (700—800 m a.s.l.), 12.vii. 1990, 2 8, 24.vii.1990, concept of the name with respect to G. (G) 2 8, 13.viii.1990, 1 8; Hostynské vrchy [hills], lacidula and other closely related species. “Valaska” (500—600 m a.s.l.), 23.vii.1992, 1 8; Other material examined (141 8, 1 S2). Aus- Hrobice nr. Zlin, 14.vii.1982, 5 8 (at light); tria: Oberosterreich: Hammem nr. Freistadt, Lesna nr. Zlin, 19.vii.1977, 1 8 (all J. Stary leg.) 28.vi.1874, 1 8, 25.vii.1875, 2 8 [all listed as (JSO); Adamov [Adamstal], 3.vii.1914, 1 8; lacidula by Lackschewitz 1940 and Franz 1989] Brno, Ricka valley [Rzickatal], 25.vii., 1 8 (both (J. Mik leg.) (NHMW). Bulgaria: Lilianovo nr. K. Czizek leg.) [both listed as lacidala by Stary Sandanski, 15.vi.1990, 1 8 (J. Stary leg.) (JSO). 1971] (MMBC); Brno-Bystrc, “Kocici z1eb”, 238 Star)? ' ENTOMOL. FENNICA Vol. 21

4.ix.1981, 1 8; Brno, Bobrava valley, Discussion. Gonomyia (Gonomyia) abscon— 29.vi.1974, 2 8 (all J. Stary leg.) (JSO); Vranov dita is distinctive in having the pleuron patterned nad Dyji [Frain], 2.ix.1883, 1 8 (A. Handlirsch with brown, with mostly a distinct central spot on leg.) [listed as lucidula by Lackschewitz 1940 the upper part of the katepistemum. This charac- and Franz 1989] (NHMW); Rohatec nr. ter, not well pronounced in commonly occurring Hodonin, 8.vi.1976, 1 8, 4.viii.1976, 1 8; teneral specimens, moreover variable in extent of Rohatec nr. Hodonin, Panov, 8.vii.1977, 1 8; the patterning, is, however, also shared by G. (G) Radejov, Lucina, 8.ix.1978, 1 8; Bile Karpaty securiformis sp. n. The following combination of Mts, Javornik, “Machova”, 1.vii.1993, 1 8; Bile characters is decisive for G. (G) abscondita (Figs Karpaty Mts, Suchovské Mlyny, Jamny, 2a—c): outer gonostylus short, only slightly ex- 5 .vii. 1994, 1 8 (all J. Stary leg.) (JSO). Germany: ceeding inner gonostylus, gradually tapered to Baden-Wiirtemberg: Greffern, Rheinaue, subacute tip; inner gonostylus with only tiny 19.vi.1995, 1 8, 20.vi.1995, 1 8, 27.vi. 1995, 1 bump at outer base; both parameres pointed at 8, 7.vii.1995, 2 8 (J. Stary leg.) (JSO). Bayem: tips, left one longer, darkly pigmented distally, Kreuzthal nr. Kempten/Allgau, 17.—22.vii. 1972, right shorter and pale; aedeagus with apical por- 5 8, 22.—27.vii.1972, 4 8, 27.vii.—3.viii.1972, 1 tion generally rounded and aedeagal duct ending 8, 10.—17.viii.1972, 2 8, 17.—24.viii.1972, 2 8; in middle of its posterior margin. Eichholz nr. Kempten/Allgau, 22.vii.—27.viii. The original figures of G. (G) abscondita by 1974, 1 8; Obergunzburg nr. Kempten/ Allgau, Lackschewitz (1935, Figs 7a—d) depict the char- 6.vii.—16.viii.1974, 2 8 (all light trap) (all H. acteristic shape of the aedeagus, but the outer Mendl leg.; in ethanol) (JSO). Great Britain: gonostyli appear rather long in Figs 7b—c. The Scotland: Angus, R. lsla, Den of Airlie, 4.vii. dissected terminalia ofthe lectotype, mounted be- 1977, 1 8 (AM. Hutson leg.) (JSO). England: tween celluloid slides (see Type material exam- Oxford, Sfl<., 14.vii.1908, 1 8 (J.J.F.X. King ined), show, however, the outer gonostyli of the leg.); S. Devon, Dartmouth, 28.—31.v.1920, 1 8; correct shape and length. The figures by Stary & Westmorland, Staveley, vi.1929, 1 8 (both F.W. Rozkosny (1970, Figs 63, 69) under G. (G) Edwards leg.) (BMNH). Wales: Flintshire, abscondita belong to that species based on the Ffrith, 7.—9.vi.1919, 2 8 (F.W. Edwards leg.) aedeagus and parameres. The illustrations given (BMNH, JSO). Netherlands: Bunde, 16.vi. 1919, under G. (G) abscondita by Podenas et al. (2006, 1 8; Nieuwersluis, 8.vi.1919, 1 8; Oldenzaal, Figs 14.1—14.2) were copied from Stary & 28.vi.1917, 1 8 [paralectotypes of lucidula; see Rozkosny (1970), as were those in Savchenko de Meijere 1920]; Winterswijk, 17.vi.1921, 1 8; (1982, Figs 139/6, 140/6) (see Discussion of ‘s Hage, vii.1903, 1 8 (all J.C.H. de Meijere leg.) lucidula). The figure by Agrell (1945, Fig. 2), (ZMAN). Slovakia: Stankovany, brook, based mainly on the aedeagus, clearly indicates 9.viii.1989, 1 8; Chocske vrchy [hills], Kvaci- that G. (G) sulphurea Agrell, 1945 is identical anska dolina [valley], 24.vi.1998, 1 8; Pol’ana with G. (G) abscondita, and the former is here Mts, Kyslinky, Hucava valley (750 m a.s.l.), confirmed, in accordance with Savchenko et al. 18.vi.2003, 1 8 (at light); Pol’ana Mts, (1992), as a junior synonym of the latter. “Hroncecky grun” (800—900 m a.s.l.), 10.vi. Distribution. Based on the material examined 1999, 1 8, 12.vi.1999, 3 8; Pol’ana Mts, Hroncek here, G. (G) abscondita is known from Austria, valley, (700 m a.s.l.), 12.Vi.1999, 1 8; Pol’ana Bulgaria, Czech Republic, Germany, Great Brit- Mts, Giemy Potok (700 m a.s.l.), 3.vii.2000, 3 6 ain, Italy, Netherlands, Slovakia, and Sweden. (at light), 4.vii.2000, 2 8 (at light), 6.vii.2000, 1 The data from Bulgaria and the Netherlands may 8 (at light), 7.vii.2000, 2 8 (at light), 13.vi.2007, be considered new country records. The record 1 8 (at light); Bukovské vrchy [hills], Ruské, from Ukraine (Savchenko 1982) is probably cor- Cirocha shores, 7.vi.1985, 1 8; Bukovské vrchy rect. Other records (Savchenko et al. 1992, [hills], Rusky Potok, valley, 10.vi.1985, 1 8 [the Oosterbroek 2009) need confirmation. latter two localities listed under lucidula by Stary 1995] (all J. Stary leg.) (JSO). Sweden: Sk., Kullaberg, Djupadal, 1 8 (B. Tj eder leg.) (JSO).

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Krzeminski leg.) (JSO). Czech Republic: Bohe- 2.4. Gonomyia (Gonomyia) undiformis sp. 11. mia: Krkonose Mts, Bile Labe (1,250 m a.s.l.), (Figs 4a—c) 14.—21.vi.2007, 1 8 (Malaise trap) (M. Bartak & J. Vanek leg.) (JSO). Moravia: Hruby Jesenik Description. Male. Small species within Gono— Mts, Branna, “Dembauda” (900 m a.s.l.), 15.vii. myia. Wing length 5.2—6.1 mm. Prescutum 1972, 1 8; Hruby Jesenik Mts, Kouty nad Des- brown. Pleuron pale yellow, without any distinct nou, Divoka Desna valley, “zamcisko” (970 m brown pattern, but obscure yellow stains some- a.s.l.), 2.vi.2003, 7 8, 3 S2, 26.vi.2003, 3 8, times apparent. In general resembling G. (G.) 8.vii.2004, 1 8, 1 $2, 27.vii.2006, 8 8, 1 SB (all at lucidula. light); Hruby Jesenik Mts, Velka kotlina (over Male terminalia (Figs 4a—c). Outer gonosty- 1,200 ma.s.l.), 15.vi. 1988, 4 8, 1 S2, 2.vii.2002, 5 lus shorter than that in G. (G.) securiformis sp. n., 8, 10 S2; Hruby Jesenik Mts, Vidly, “Jeleni exceeding inner gonostylus by about one-third its bucina” (900 m a.s.l.), 20.vi.2000, 1 8, 21.vi. length, otherwise much as in the latter species, 2001, 1 8 (all J. Stary leg.) (JSO). Italy: Prali pale, slightly expanded distally in dorsal aspect, (TO), Ribba, Miande Bont du Col (1,750 m apex rounded or obliquely truncate, with short in- a.s.l.), 3.Vii.1998, 1 8 (GB. Delmastro leg.); ner point. Inner gonostylus with its inner (medial) Castelmagno (CN), V.ne Siboleti (1,770 m a.s.l.), part similar to others; outer (lateral) part strongly 8.vi.2001, 1 8 (G.F. & G.B. Delmastro leg.) pigmented, with longer lobe produced into sharp (JSO). Switzerland: Canton Valais: Aletschwald, curved black hook and shorter lobe very broad. Teiffenwald (1,800 m a.s.l.), 4.vii.1996, 1 8 (J. Outer base of inner gonostylus with tiny pale Stary leg.) (JSO). bumb with projecting seta, similar to G. (G.) ab— Etymology. The name of this new species, scondita. Left, longer paramere darkly pigmented securiformis (= axe-shaped), refers to the shape distally and pointed at tip; right, shorter paramere ofits aedeagus. An adjective in nominative singu- pale, very stout and obtuse at tip. Aedeagus dif- lar. ferent from those of other species, not expanded Discussion. Gonomyia (Gonomyia) securi— apically, nor excised ventrally, or very shallowly formis sp. n. most resembles G. (G.) abscondita so, rather parallel-sided and irregularly undulate, in general appearance in having the pleuron dis- with numerous microscopic setulae dorsally near tinctly patterned with brown. It may be distin- tip, discernible at higher magnifications. Distal guished from the latter, as well as from the other part of aedeagal duct not reaching posterior mar- species, by its larger size, but this character varies gin of aedeagus. considerably, and the sizes overlap to a certain Female. Resembles male in general appear- degree. The following combination of characters ance, not reliably distinguished from females of is species-specific for G. (G.) securiformis sp. n. other species. (Figs 3a—c): outer gonostylus long, exceeding in- Type material. Holotype 8: Slovakia, Cer- ner gonostylus by nearly half its length, slightly veny Klastor, Dunaj ec valley, 30.v.2000 (J. Stary expanded distally, rounded or obliquely truncate leg.) (SMOC). The specimen is glued onto a tri- at apex; inner gonostylus with distinct tooth at angular cardboard point, with only right mid and outer base; both parameres darkly pigmented dis- hind legs missing; apex of abdomen cut off. Ter- tally and pointed at tips; aedeagus with apical minalia dissected by the present author and portion axe-shaped and aedeagal duct ending at placed in a sealed plastic tube with glycerine, its posteroventral edge. pinned with the specimen. Paratypes (3 8, 1 S2): Distribution. Bulgaria, Czech Republic, Italy, Slovakia: Belianske Tatry Mts, Tristarska dolina Switzerland. In the Czech Republic, the species [valley], 26.vi.1975, 1 8; Cerveny Klastor, was found to be syntopic with both G. (G.) Dunajec valley, 30.v.2000, 2 8, 1 SB (all J. Stary lucidula and G. (G.) abscondita at the locality leg.) (JSO). Hruby Jesenik Mts, Branna, “Dembauda” (see Etymology. The name of this new species, Material examined). undiformis (= undulate), refers to the shape of its aedeagus. An adjective in nominative singular. Discussion. This new species most resembles

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