Perennial Centaurea for the Garden
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The Vascular Plants of Massachusetts
The Vascular Plants of Massachusetts: The Vascular Plants of Massachusetts: A County Checklist • First Revision Melissa Dow Cullina, Bryan Connolly, Bruce Sorrie and Paul Somers Somers Bruce Sorrie and Paul Connolly, Bryan Cullina, Melissa Dow Revision • First A County Checklist Plants of Massachusetts: Vascular The A County Checklist First Revision Melissa Dow Cullina, Bryan Connolly, Bruce Sorrie and Paul Somers Massachusetts Natural Heritage & Endangered Species Program Massachusetts Division of Fisheries and Wildlife Natural Heritage & Endangered Species Program The Natural Heritage & Endangered Species Program (NHESP), part of the Massachusetts Division of Fisheries and Wildlife, is one of the programs forming the Natural Heritage network. NHESP is responsible for the conservation and protection of hundreds of species that are not hunted, fished, trapped, or commercially harvested in the state. The Program's highest priority is protecting the 176 species of vertebrate and invertebrate animals and 259 species of native plants that are officially listed as Endangered, Threatened or of Special Concern in Massachusetts. Endangered species conservation in Massachusetts depends on you! A major source of funding for the protection of rare and endangered species comes from voluntary donations on state income tax forms. Contributions go to the Natural Heritage & Endangered Species Fund, which provides a portion of the operating budget for the Natural Heritage & Endangered Species Program. NHESP protects rare species through biological inventory, -
02. CENTAUREA.Indd
ActaCentaurea Botanica sect. MalacitanaCyanus en la 35. Península 23-55 Ibérica Málaga, 201023 REVISIÓN TAXONÓMICA DEL COMPLEJO DE CENTAUREA CYANUS L. (CENTAUREA SECT. CYANUS, ASTERACEAE) EN LA PENÍNSULA IBÉRICA Adolfo Francisco MUÑOZ1 y Juan Antonio DEVESA2* Dirección de los autores.1Departamento de Biología Ambiental y Salud Pública, Facultad de Ciencias Experimentales, Universidad de Huelva, Avenida de la Fuerzas Armadas s/n, 21071-Badajoz, España 2Departamento de Botánica, Ecología y Fisiología Vegetal, Facultad de Ciencias, Universidad de Córdoba, Campus de Rabanales, Edificio José Celestino Mutis, ctra. de Madrid km. 396, 14071-Córdoba, y Jardín Botánico de Córdoba, Avda. de Linneo s/n, 14004-Córdoba, España. *Autor para correspondencia: [email protected] Recibido el 1 de septiembre de 2010, aceptado para su publicación el 10 de octubre de 2010 RESUMEN. Revisión taxonómica del complejo de Centaurea cyanus L. (Centaurea sect. Cyanus, Asteraceae) en la Península Ibérica. Se ha efectuado la revisión taxonómica de Centaurea sect. Cyanus en la Península Ibérica. Se reconocen seis táxones y se hacen dos nuevas combinaciones: C. triumfettii var. semidecurrens (Jord.) Muñoz Rodr. & Devesa y C. graminifolia (Lam.) Muñoz Rodr. & Devesa. Se aporta una clave para la identificación de los táxones, así como su descripción morfológica e información sobre la distribución geográfica, apetencias ecológicas, fenología y número cromosómico. Palabras clave. Asteraceae, Centaurea, corología, taxonomía, Península Ibérica. ABSTRACT. Taxonomic revision of the Centaurea cyanus L. complex (Centaurea sect. Cyanus, Asteraceae) in the Iberian Peninsula. A taxonomic review of Centaurea sect. Cyanus in the Iberian Peninsula was done. Six taxa are recognized and two new combinations are proposed: C. triumfetti var. -
Squarrose Knapweed EXOTIC Centaurea Virgata Ssp
southwestlearning.org AME R ICAN SOUTHWEST SPECIES FACT SHEET Asteraceae (Sunflower family) Squarrose Knapweed EXOTIC Centaurea virgata ssp. squarrosa At a Glance • Perennial • Highly branched stems that grow one to three feet tall. SITY • Lower leaves are deeply dissected, but the upper leaves R are entire and linear. • Flowers are pink to pale-purple. • Flowerheads have four to eight florets and appear more slender than the flowerheads of other knapweeds. • The bracts underneath the flower have a central spine that curves backwards. • Fruit is a golden to dark-brown achene. UNIVE STATE UTAH / DEWEY STEVE The bracts underneath the flower of squarrose knapweed have Habitat and Ecology a central spine that curves backwards. Squarrose knapweed (Centaurea virgata ssp. squarrosa) is present in Utah, Nevada, California, Oregon, Wisconsin, spotted (Centaurea maculosa) or diffuse knapweed (Cen- and Michigan. How it was introduced to North America is taurea diffusa). Squarrose knapweed is not palatable to unknown. Squarrose knapweed often grows on degraded livestock and can form monocultures. Its taxonomic status rangeland soils and tolerates drought and cold better than is uncertain: it is also known as Centaurea squarrosa and Centaurea virgata. Description Squarrose knapweed is a long-lived perennial with highly branched stems that grow one to three feet tall. The stems grow above a woody crown and stout taproot. Under unfa- vorable conditions, the plants remain as basal rosettes be- fore developing flowering stems. Squarrose knapweed re- produces by seed. The seed head falls near the parent plant, but the backwards-curved spines under the seed head can cling to hair, wool, fur, and clothing, allowing the seeds to disperse over greater distances. -
Weed Risk Assessment: Centaurea Calcitrapa
Weed Risk Assessment: Centaurea calcitrapa 1. Plant Details Taxonomy: Centaurea calcitrapa L. Family Asteraceae. Common names: star thistle, purple star thistle, red star thistle. Origins: Native to Europe (Hungary, Switzerland, Czechoslovakia, Russian Federation, Ukraine, Albania, Greece, Italy, Romania, Yugoslavia, France, Portugal, Spain), Macaronesia (Canary Islands, Madeira Islands), temperate Asia (Cyprus, Lebanon, Syria, Turkey) and North Africa (Algeria, Egypt, Morocco, Tunisia) (GRIN database). Naturalised Distribution: Naturalised in New Zealand, South Africa, Central America, South America, the United States of America (eg. naturalised in 14 states, mostly in northwest including California, Idaho, Washington, Wyoming, New Mexico, Oregon, Arizona) (USDA plants database), and Australia (GRIN database). Description: C. calcitrapa is an erect, bushy and spiny biannual herb that is sometimes behaves as an annual or short-lived perennial. It grows to 1 m tall. Young stems and leaves have fine, cobweb-like hairs that fall off over time. Older stems are much-branched, straggly, woody, sparsely hairy, without wings or spines and whitish to pale green. Lower leaves are deeply divided while upper leaves are generally narrow and undivided. Rosette leaves are deeply divided and older rosettes have a circle of spines in the centre. This is the initial, infertile, flower head. Numerous flowers are produced on the true flowering stem and vary from lavender to a deep purple colour. Bracts end in a sharp, rigid white to yellow spines. Seed is straw coloured and blotched with dark brown spots. The pappus is reduced or absent. Bristles are absent. Seeds are 3-4mm long, smooth and ovoid. The root is a fleshy taproot (Parsons and Cuthbertson, 2001) (Moser, L. -
Agri-Environment Nectar Chemistry Suppresses Parasite Social Epidemiology in an 2 Important Pollinator
bioRxiv preprint doi: https://doi.org/10.1101/2021.01.30.428928; this version posted February 1, 2021. The copyright holder for this preprint (which was not certified by peer review) is the author/funder, who has granted bioRxiv a license to display the preprint in perpetuity. It is made available under aCC-BY-NC-ND 4.0 International license. 1 Agri-environment nectar chemistry suppresses parasite social epidemiology in an 2 important pollinator (1,#) (2) (2) (2,3) (1) 3 Arran J. Folly* , Hauke Koch , Iain W. Farrell , Philip C. Stevenson , Mark J.F. Brown 4 (1) Centre for Ecology, Evolution and Behaviour, Department of Biological Sciences, School of Life Sciences and the Environment, Royal 5 Holloway University of London, Egham, UK (2) Royal Botanic Gardens, Kew, UK (3) Natural Resources Institute, University of Greenwich, 6 Kent, UK 7 8 *Corresponding author: Arran J. Folly: [email protected] 9 #Current address: Virology Department, Animal and Plant Health Agency, Surrey, UK 10 11 Emergent infectious diseases are a principal driver of biodiversity loss globally. The population 12 and range declines of a suite of North American bumblebees, a group of important pollinators, 13 have been linked to emergent infection with the microsporidian Nosema bombi. Previous work 14 has shown that phytochemicals in pollen and nectar can negatively impact parasites in individual 15 bumblebees, but how this relates to social epidemiology and by extension whether plants can be 16 effectively used as disease management strategies remains unexplored. Here we show that 17 caffeine, identified in the nectar of Sainfoin, a constituent of agri-environment schemes, 18 significantly reduced N. -
Flowering Phenology As a Functional Trait in a Tallgrass Prairie
Research Flowering phenology as a functional trait in a tallgrass prairie Joseph M. Craine1, Elizabeth M. Wolkovich2, E. Gene Towne1 and Steven W. Kembel3 1Division of Biology, Kansas State University, Manhattan, KS 66502, USA; 2Ecology, Behavior & Evolution Section, University of California, San Diego, 9500 Gilman Drive #0116, La Jolla, CA 92093,USA; 3Center for Ecology & Evolutionary Biology, University of Oregon, Eugene, OR 97403, USA Summary Author for correspondence: • The timing of flowering is a critical component of the ecology of plants and has the poten- Joseph M. Craine tial to structure plant communities. Yet, we know little about how the timing of flowering Tel: +1 785 532 3062 relates to other functional traits, species abundance, and average environmental conditions. Email: [email protected] • Here, we assessed first flowering dates (FFDs) in a North American tallgrass prairie (Konza Received: 18 August 2011 Prairie) for 431 herbaceous species and compared them with a series of other functional traits, Accepted: 29 September 2011 environmental metrics, and species abundance across ecological contrasts. • The pattern of FFDs among the species of the Konza grassland was shaped by local climate, New Phytologist (2011) can be linked to resource use by species, and patterns of species abundance across the land- doi: 10.1111/j.1469-8137.2011.03953.x scape. Peak FFD for the community occurred when soils were typically both warm and wet, while relatively few species began flowering when soils tended to be the driest. Compared with late-flowering species, species that flowered early had lower leaf tissue density and were Key words: climate, community assembly, drought, grass, Konza Prairie. -
Asteraceae), Turkey
Mellifera 2018. 18(2):15-25 MELLIFERA RESEARCH ARTICLE Pollen and Achene Morphology of Some Cyanus L. Taxa (Asteraceae), Turkey Halime ATAR1*. Barış BANİ1. Talip ÇETER1 1 Kastamonu University, Arts and Sciences Faculty, Department of Biology, Kastamonu, Turkey *Corresponding author e-mail: [email protected] Received: 05th December, 2018; accepted: 15th December, 2018; published: 28th December, 2018 A B S T R A C T Cyanus is a genus of family Asteraceae. 20 taxa belonging to Cyanus distributes in Turkey and 9 of them are endemics. In this study, pollen morphology and achene micro and macro morphology of 4 taxa belonging to Cyanus (C. depressus, C. triumfettii, C. pichleri subsp. pichleri, C. lanigerus) were investigated. The aim of the study was to determine taxonomic value of pollen and achene micro characteristic. Pollen grains of Cyanus taxa observed as isopolar, radially symmetric, with tricolporate aperture and subprolate pollen shape. Polar axis range between 36 -55.8 µm while equatorial axis between 27.8-47 µm. C. lanigerus pollen measured as biggest pollen wile C. depressus pollen is the smallest. Pollen surface ornamentation determined as scabrate. The achene shape was observed as ovoid and oblong in studied taxa. SEM results showed tha t the achene surface had a striate-psilate sculpture and was depressed on both lateral surfaces, without rare hairs. The Achene length (4-5mm), width (1.6-2.4 mm) and pappus length (1-6.4 mm) ranges differed significantly. 15 C. triumfettii achene determined as smallest (4.0 mm) in length with the smallest (1.1 mm) pappus while C. -
A Structural Growth Model of the Invasive Weed Species Yellow Starthistle, Centaurea Solstitialis L
A Structural Growth Model of the Invasive Weed Species Yellow Starthistle, Centaurea solstitialis L. 1Thornby, D., 2J. Garren, 2R. Carruthers, 2D. Spencer and 2J.P. Weber 1Queensland Department of Primary Industries and Fisheries 2USDA Agricultural Research Service; Email: [email protected] Keywords: Centaurea solstitialis, yellow starthistle, structural model, LStudio, L-systems. EXTENDED ABSTRACT Yellow starthistle (Centaurea solstitialis L.) is a summer-flowering annual plant of the family Asteraceae, and a significant weed of roadsides, pasture and grasslands in California. It is also distributed, though with less economic and ecological significance, throughout the US and in other temperate parts of the world. Yellow starthistle grows vigorously during the dormant period for most native species in its grasslands habitat, is able to access deep soil moisture reserves, and being particularly spiny, reduces land value for both animal fodder and human recreation. It is a prolific seed producer and seed spreads readily through contact with animals, people, and vehicles. As with many weeds, yellow starthistle's structure and growth patterns have been relatively little- Figure 1. Virtual yellow starthistle studied. It is hoped that studying the ontogenesis and morphogenesis of yellow starthistle will help Stochasticity was added to the model, so that it uncover information that can assist in improving reproduces a range of plants of different sizes and control strategies, as well as providing a morphologies, within the range of likely sizes visualisation tool for disseminating detailed observed in the data for each field site. knowledge about this plant's growth and development to stakeholders in a readily understood way. -
Nuclear and Plastid DNA Phylogeny of the Tribe Cardueae (Compositae
1 Nuclear and plastid DNA phylogeny of the tribe Cardueae 2 (Compositae) with Hyb-Seq data: A new subtribal classification and a 3 temporal framework for the origin of the tribe and the subtribes 4 5 Sonia Herrando-Morairaa,*, Juan Antonio Callejab, Mercè Galbany-Casalsb, Núria Garcia-Jacasa, Jian- 6 Quan Liuc, Javier López-Alvaradob, Jordi López-Pujola, Jennifer R. Mandeld, Noemí Montes-Morenoa, 7 Cristina Roquetb,e, Llorenç Sáezb, Alexander Sennikovf, Alfonso Susannaa, Roser Vilatersanaa 8 9 a Botanic Institute of Barcelona (IBB, CSIC-ICUB), Pg. del Migdia, s.n., 08038 Barcelona, Spain 10 b Systematics and Evolution of Vascular Plants (UAB) – Associated Unit to CSIC, Departament de 11 Biologia Animal, Biologia Vegetal i Ecologia, Facultat de Biociències, Universitat Autònoma de 12 Barcelona, ES-08193 Bellaterra, Spain 13 c Key Laboratory for Bio-Resources and Eco-Environment, College of Life Sciences, Sichuan University, 14 Chengdu, China 15 d Department of Biological Sciences, University of Memphis, Memphis, TN 38152, USA 16 e Univ. Grenoble Alpes, Univ. Savoie Mont Blanc, CNRS, LECA (Laboratoire d’Ecologie Alpine), FR- 17 38000 Grenoble, France 18 f Botanical Museum, Finnish Museum of Natural History, PO Box 7, FI-00014 University of Helsinki, 19 Finland; and Herbarium, Komarov Botanical Institute of Russian Academy of Sciences, Prof. Popov str. 20 2, 197376 St. Petersburg, Russia 21 22 *Corresponding author at: Botanic Institute of Barcelona (IBB, CSIC-ICUB), Pg. del Migdia, s. n., ES- 23 08038 Barcelona, Spain. E-mail address: [email protected] (S. Herrando-Moraira). 24 25 Abstract 26 Classification of the tribe Cardueae in natural subtribes has always been a challenge due to the lack of 27 support of some critical branches in previous phylogenies based on traditional Sanger markers. -
Weed: Yellow Starthistle (Centaurea Solstitialis L.)
Weed: Yellow starthistle (Centaurea solstitialis L.) Family: Asteraceae (Sunflower family) Images: Brief Plant Description: (Summarized from Healy, E. and J. DiTomaso, Yellow Starthistle Fact Sheet, http://wric.ucdavis.edu/yst/biology/yst_fact_sheet.html) The seed leaves (cotyledons) are oblong to spatulate, 6-9 mm long and 3-5 mm wide, base wedge- shaped, tip +/- squared and glabrous. First few rosette leaves typically oblanceolate. Subsequent rosette leaves oblanceolate, entire to pinnate-lobed. Terminal lobes largest. Later rosette leaves to 15 cm long and are typically deeply lobed +/- to midrib and appear ruffled. Surfaces +/- densely covered with fine cottony hairs. Lobes mostly acute, with toothed to wavy margins. Terminal lobes +/- triangular to lanceolate. Mature plants have stiff stems, openly branched from near or above the base or sometimes not branched in very small plants. Stem leaves alternate, mostly linear or +/- narrowly oblong to oblanceolate. Margins smooth, toothed, or wavy. Leaf bases extend down the stems (decurrent) and give stems a winged appearance. Rosette leaves typically withered by flowering time. Largest stem wings typically to ~ 3 mm wide. Lower stem leaves sometimes +/- deeply pinnate-lobed. Foliage grayish- to bluish-green, densely covered with fine white cottony hairs that +/- hide thick stiff hairs and glands. Flower heads ovoid, spiny, solitary on stem tips, consist of numerous yellow disk flowers. Phyllaries palmately spined, with one long central spine and 2 or more pairs of short lateral spines. Insect- pollinated. Flowers mid-summer to fall. Corollas mostly 13-20 mm long. Involucre (phyllaries as a unit) ~ 12-18 mm long. Phyllaries +/- dense to sparsely covered with cottony hairs or with patches at the spine bases. -
Seed Set in a Non-Native, Self-Compatible Thistle on Santa Cruz Island: Implications for the Invasion of an Island Ecosystem
SEED SET IN A NON-NATIVE, SELF-COMPATIBLE THISTLE ON SANTA CRUZ ISLAND: IMPLICATIONS FOR THE INVASION OF AN ISLAND ECOSYSTEM JOHN F. BARTHELL1, ROBBIN W. THORP2, ADRIAN M. WENNER3, JOHN M. RANDALL4 AND DEBORAH S. MITCHELL5 1Department of Biology & Joe C. Jackson College of Graduate Studies & Research, University of Central Oklahoma, Edmond, OK 73034; [email protected] 2Department of Entomology, University of California, Davis, CA 95616 3Department of Ecology, Evolution & Marine Biology, University of California, Santa Barbara, CA 93106 4The Nature Conservancy, Invasive Species Program Initiative & Department of Vegetable Crops, University of California, Davis, CA 95616 5Animal Medical Center, Edmond, OK 73034 Abstract—In two previous studies we demonstrated a positive association between honey bee visitation and seed set or seed head weight in the invasive yellow star-thistle, Centaurea solstitialis. However, as reported here, we were unable to find similar evidence for the congeneric tocalote, Centaurea melitensis. Both seed head weights and percent seed set levels obtained from two plots showed no significant differences among four treatments. Indeed, no significant differences occurred between controls (open) and bagged flower heads that excluded honey bees (but allowed native bee visitation). There were also no differences between controls and complete exclusion of bee-pollinators, confirming self-compatibility reported for this species elsewhere. Unlike C. solstitialis, C. melitensis attracts relatively low numbers of honey bees. In addition, C. melitensis is generally more widespread on the Channel Islands than C. solstitialis. We discuss these patterns with reference to the invasiveness of both species on Santa Cruz Island. Keywords: Apis mellifera, Centaurea solstitialis, honey bee, invasive species, pollination, yellow star- thistle INTRODUCTION environments is subject to controversy, increasingly so during the last decade. -
Wildflowers of Scotland
Seed Origin and Quality Wildflowers of Scotland At Scotia Seeds we use our years of experience to ensure that the wildflower seed We are the leading producers of wildflower we supply is of the highest quality possible and seeds in Scotland and are committed to can be traced back to original collections in the providing the range and quality of seeds you wild. require. We have a wide range of species that we can Seed Origin provide. As well as the ones here in the All of the wildflower seeds we produce can be catalogue; please contact us if you are looking traced back to the sites where the original wild for a species not in our catalogue. plants grow. From these sites we collect a small amount of seed which is then sown on page Contents our farm to give us crops from which we can harvest a larger amount of seed. Seed Quality and Origin 2 We are signatories to the Flora Locale Code of Practice for Collectors, Growers and Suppliers Seed Packets 3 of Native Flora that ensures responsible collection and sale of native British plants. Establishing a Wildflower Meadow 11 Quality Yellow Rattle 13 We test samples of all our seed crops for germination and purity, to ensure that they have reached our stringent standards. Sowing Rates 14 Our quality laboratory specialises in testing the seeds of wildflowers and trees. For most of the Meadow Mixtures 15 species we test we have developed our own procedures in a research programme funded by How to Order 26 a Scottish Executive SMART Award.