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Palaeontological Society of Japan ISSN 0031 -0204 Transactions and Proceedings of the Palaeontological Society of Japan New Series No. 147 Palaeontological Society of Japan October 30.1987 Co-Editors Hisayoshi IGO and Tsunemasa SAITO Officers for 1987-1988 President: Tatsuaki KIMURA Honorary President: Teiichi KOBA YASH I Councillors: Kiyotaka CHINZEI, Takashi HAMADA, Yoshikazu HASEGAWA, Itam HAYAMI, Hisayoshi lGO, Junji lTOIGAWA, Tadao KAMEl, Tatsuaki KIMURA, Tamio KOTAKA, Kei MORI, Ikuwo OBATA, Tsunemasa SAITO, Yokichi TAKAYANAGI, Kazushige TANABE, Toshimasa TANAI Members of Standing Committee: Kiyotaka CHINZEI (Membership), Takashi HAMADA (For­ eign Affairs), ltaru HAYAMI (Finance), Hisayoshi lGO (Co-Editor of Transactions), lkuwo OBATA (General Affairs), Sumio SAKAGAMI (Co-Editor of Special Paper), Kazushige TANABE (Planning), Yokichi TAKAYANAGI (Editor of "Fossils"), Juichi YANAGIDA (Co-Editor of Special Paper) Secretaries: Katsumi ABE (Planning), Takeshi ISHIBASHI (Special Paper), Kunihiro ISHIZAKI ("Fossils"), Tomoki KASE (General Affairs and Membership), Hiroshi NODA (Editor of Transactions), Terufumi OHNO (Membership), Tatsuo OJ! (Planning), Kazuhiko UEMURA (General Affairs), Toshiyuki YAMAGUCHI (Finance) Auditor: Noriyuki lKEYA The fossil on the cover is Vicarya yokoyamai Takeyama, an Early Middle Miocene gastropod from the Kurosedani Formation at Kakehata, Yatsuo-cho, Nei-gun, Toyama Prefecture, central Japan (Collected by T. Kotaka and K. Ogasawara, lGPS No. 99075, photo by S. Ohtomo and Y. Kikuchi, xO.9). All communication relating to this journal should be addressed to the PALAEONTOLOGICAL SOCIETY OF JAPAN % Business Center for Academic Societies, Yayoi 2-4-16, Bunkyo-ku, Tokyo 113, Japan Trans. Proc. Palaeont. Soc. Japan, N. S., No. 147, pp. 91-108,7 Figs., October 30,1987 832. CRETACEOUS EOMIODON AND COSTOCYRENA (BIVALVIA) FROM SOUTHWEST JAPAN MASA YUKI TASHIRO Department of Geology, Faculty of Science, Kochi University, Kochi 780 Abstract. In this paper 4 species of Eomiodon and 8 species and 2 subspecies of Costo­ cyrena (including Costa cyrena oh n ish ii, sp. nov., Costocyrena hojiensis, sp. nov. and Costo­ cyrena otsukai obsoleta, subsp. nov.) belonging to the Family Neomiodontidae are described from the Cretaceous at various localities in southwest Japan. Geographic distribution of Eomiodon and Costocyrena species seems to change through the Cretaceous time in southwest Japan. In the Lower Cretaceous, Eomiodon species restricted­ ly occur from the southern side of Kurosegawa Tectonic Belt, while all Costocyrella species are known from the northern side of that belt, Northern Chichibu Belt. In the Upper Cretaceous, the geographic distribution of the two genera becomes overlap, and they occur together even at the same localities especially in central Kyushu. This may not mean habitat change of these bivalves, but seems to support the hypothesis of the Early Cretaceous movement of a large lateral fault in the Chichibu Belt. Introduction da of Kochi Pref. for their supply of material. The studied specimens (KSG) are kept in the Many species of Eomiodon and Costocyrena Faculty of Science, Kochi University. belonging to the Family Neomiodontidae widely thrived during the Cretaceous times in Japan. Systematic descritpion They occur abundantly from the various strati­ graphic horizones characterized by the so-called Family Neomiodontidae Casey, 1955 brackish water facies, ranging from Hauterivian Subfamily Eomiodontinae Hayami, 1965 to Cenomanian, at many localities in Southwest Japan. Genus Eomiodoll Cox, 1935 Eomiodon and Costocyrena species described Eomiodon sakawanus (Kobayashi et Suzuki) in this paper seem to be important for the corre­ Figures 1-7-12 lation of the Cretaceous strata in Japan. Chang­ ing of the geographic distribution of Eomiodol1 1939. Astarte sakawana Kobayashi et Suzuki, Japan. and Co:.,tocyrena species through the Cretaceous Jour. Ceol. Ceogr., vol. 13, nos. 3-4, p. 219, pI. times also involve interesting problems both on 13, fig. 11 (non pI. l3, figs. 12 and 13). 1973. Eomiodon sakawanus (Kobayashi et Suzuki); palaeoecology and tectonic developments of Ohta, Bull. Fukuoka Univ. Educ., vol. 22, no. 3, eastern Asia. p. 250, pI. 1, figs. 1-11. I wish to express my sincere thanks to Miss 1975. Eomiodon sakawanus (Kobayashi et Suzuki); Takako Ohnishi of Kagawa Pref., Mr. Hideyuki Hayami, Univ. Mus., Ulliv. Tokyo, Bull., 10, p. Yanagisawa of Fukui Pref. and Mr. Seiji Matsu- 138. 1976. Eomiodon sakawanus (Kobayashi et Suzuki); *Received September 16, 1986; revised manuscript Ohta, Atlas of Japanese fossils, vol. 44, Meso­ accepted July 6, 1987; read June 15, 1985 at Osaka zoic bivalves (3), figs. 7-9. City University. 91 92 Masayuki TASHIRO Materials:-KSG 3803 and KSG3804,external figs. 1-7. 1981. Eomiodon matsumotoi Ohta; Ohta, Bull. Fuku­ moulds of right valves, from Uchiyama of Aioi, oka Univ. Educ., vol. 31, no. 3, p.103, pI. 7, figs. Tokushima Prefecture. KSG 3805 and KSG 1-8. 3806, external moulds of right and left valves, from Yonenomoto of Ittku, Kochi City. Materials:-KSG 3807-KSG 3810, external Measurements:-(in mm.) moulds of left valves; KSG 3811, external mould Specimens Length Height Thickness of right valve; all from Ohta of Tanoura, Ashi­ KSG 3803, r. ext. mol. 13.0 11.0 2.5 kita-gun in Kumamoto Prefecture. KSG 3812 - KSG 3804, r. ext. mol. 10.0 9.0 1.5 KSG 3813, external moulds of left valves, from KSG 3805, r. ext. mol. 14.1 11.5 2.5 Uchiyama of Aioi, Naka-gun in Tokushima KSG 3806, I. ext. mol. 18.0 15.0 2.0 Prefecture. Remarks:-The holotype was occurred from Measurements:-(in mm.) the Kaisekiyama Formation of Kamo, Sakawa in Specimens Length Height Thickness Kochi Prefecture. Two species, treated in this KSG 3807,1. ext. mol. 29.5 23.0 4.0 study (Figures 5-1, II and 12), preserved in the KSG 3809, I. ext. mol. 10.5 88.0 2.0 Fossil Museum of Sakawa (not registered), were KSG 3811, r. ext. mol. 22.0 16.0 3.0 also collected from the type locality. Remarks: - This species is characterized by the This species is characterized by more broadly very fine concentric striae which are developed spaced concentric ribs on the disk and less in­ on the interspaces of the concentric ribs on the flated valve than other Japanese Eomiodon disk. Although this species resembles Eomiodon species. About 5 concentric ribs are countable sakawanus (Kobayashi et Suzuki) in its roundly on the adult stage. They are narrower than their subtrigonal outline of the valve and distinct con­ interspaces and roof-shaped on the top. centric ribs on the disk, it difers from E. sakawa­ Occurrence and geological age: - Fine-grained nus in its more inflated valve, numerous and sandstone of the Nakaizu Formation at Uchi­ more or less irregularly spaced concentric ribs, yama of Aioi, Naka-gun in Tokushima Prefec­ and fine concentric striae on their in terspaces. ture; Aptian. Very fine-grained sandstone of the Occurrence and geological age: -Gray siltstone Funadani Formation at Ittku of Kochi City, of the Kawaguchi Formation at Ohta of Tanoura, Kochi Prefecture; Aptian. Sandy mudstone of Ashikita-gun, Kumamoto Prefecture; Hauterivian. the Kaisekiyama Formation at Kaisekiyama of Fine-grained sandstone of the Shobu Formation Sakawa, Kochi Prefecture. This species is also at Uchiyama of Aioi, Naka-gun in Tokushima known from the upper part of the Yamabu Prefecture; Hauterivian. This species had been Formation at Yamabu, Minami-Amabe-gun in reported from the lower member of Doganaro Oita Prefecture; Aptian(?). Formation of the Shimanto Belt in Kochi Prefec­ ture (Katto and Tashiro, 1979) and from the Eomiodon matsumotoi Ohta lower part of Yamabu Formation of the Chichi­ Figures 1-1-6 bu Belt in Ohita Prefecture (Tashiro, 1985a), both in the Hauterivian in age. This is also known 1973. Eomiodon matsumotoi Ohta, Bull. Fukuoka from the Uminoura Formation in central Kyushu Univ. Educ., vol. 22, no. 3, p. 245, pI. 2, figs. l­ (Ohta and Monji, 1976). The geological age of B. 1975. Eomiodon matsumotoi Ohta; Hayami, Univ. the Uminoura Formation is probably the earliest Mus., Univ. Tokyo, Bull., 10, p. 139. Cretaceous. 1976. Eomiodon l1latsul1lotoi Ohta; Ohta, Atlas of Japanese fossils, vol. 44, Mesozoic bivalves (3), Eomiodon matsubasensis Tamura figs. 10-15. 1979. Eomiodon matsul1lotoi Ohta; Katto and Tashiro, Figures 1-13-25 Res. Rep., Kochi Univ., vol. 27, Nat. Sci., pI. 2, 832. Cretaceous Eomiodon and Costocyrena 93 1977. Eomiodon matsubasensis Tamura, Mem. Fac. gun in Kumamoto Prefecture. This is also known Educ. Kumamoto Univ., no. 26, Nat. Sci., p. 107, from the Fukigoshi Formation of the Sotoizumi pI. 6, figs. 9-19. Group at Nagase of Odochi, Monobe-mura of 1982. Eomiodon matsubasensis Tamura; Tashiro and Matsuda, Trans. Proc. Palaeont. Soc. Japan, N.S., Kami-gun in Kochi Prefecture (Lower Ceno­ no. 127, p. 410, pI. 65, figs. 14-15. manian) (Tashiro and Matsuda, 1982). 1985. Eomiodon matsubasensis Tamura; Matsuda, Fos­ sils, no. 39, pI. 2, figs. 9-10. Eomiodon? kumamotoensis Tamura Materials:-KSG 3814-KSG 3815, external 1959. Eomiodon kumamotoensis Tamura, Trans. Proc. moulds of left and right valves, from Higashi­ Palaeont. Soc. Japan, N.S., no. 33, p. 115, pI. 12, machi (Shishijima island) in Kagoshima Prefec­ figs. 17-18. ture. KSG 3816-KSG 3818, external moulds of 1960. Eomiodon kumamotoensis Tamura; Tamura, left and right valves, from Karakizaki (Goshono­ Mem. Fac. Educ. Kumamoto Univ., vol. 8, Nat. Sci., p. 240. ura island) in Kumamoto Prefecture. KSG 3819 1975. Astarte? kumamotoensis (Tamura); Hayami, - KSG 3810, external moulds of left and right Univ. Mus., Univ. Tokyo, Bull., 10, p. 128. valves, from Chikaraishi, Kami-mashiki-gun in 1986. Eomiodon kumamotoensis Tamura; Tashiro and Kur-.amoto Prefecture, KSG 3821-KSG 3830, Katto, Res. Rep., Kochi Univ., vol. 34, Nat. Sci., external moulds of left and right valves, from pI. I, figs. 15-17. Furuyashiki (Goshonoura island) in Kumamoto Material:-KSG 3931, conjoined valves, from Prefecture. Tainokawa of Obama, Susaki City in Kochi Measurements:- (in mm.) Prefecture. Specimens Length Height Thickness Remarks:-The typical specimen of this KSG 3816, r.
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