Transactions and Proceedings

of the Palaeontological Society of

New Series No. 88

Palaeontological Society of Japan December 20, 1972 ------·-·

Editor: Takashi HAMADA Associate Editor: Yasuhide IWASAKI

Officers for 1971 - 1972

President : Tokio SHIKAMA Councillors(* Executives): Kiyoshi ASAND*, Kiyotaka CHINZEI*, Takashi HAMADA*, Tetsuro HANAI*, Kotara HAT AI, ltaru HAY AMI, Koichiro lCHIKA WA, Taro KANAYA, Kametoshi KANMERA, Tamio KOTAKA, Tatsuro MATSUMOTO*, Hiroshi OZAKI*, Tokio SHIKAMA*, Fuyuji TAKA!*, Yokichi TAKAYANAGI Secretaries: Wataru HASHIMOTO, Saburo KANNO Executive Committee General Affairs: Tetsuro HANAI, Naoaki AOKI Membership : Kiyotaka CHINZEI, Toshio KOIKE Finance: Fuyuji T AKAI, Hisayoshi lGO Planning: Hiroshi OZAKI, Kazuo AsAMA Publications Transactions: Takashi HAMADA, Yasuhide IWASAKI Special Papers: Tatsuro MATSUMOTO, Tomowo OZAWA '' Fossils" : Kiyoshi AsANO, Toshiaki TAKAYAMA

All communications relating to this journal should be addressed to the PALAEONTOLOGICAL SOCIETY OF JAPAN c/o Business Center for Academic Societies, Japan Yayoi 2-4-16, Bunkyo-ku, 113, Japan. Sole agent: University of Tokyo Press, Hongo, Tokyo Trans. Proc. Palaecint. Soc. Japan, N. S., No. 88, pp. 447-457, pl. 54, December 20, 1972

607. UPPER FROM THE TANOURA DISTRICT, PREFECTURE, JAPAN*

T AKESHI ISHIBASHI

Department of Geology, Faculty of Science, Kyushu University

Jm;fs:~llliinit!!JrRO)J:ll!I.:::.~*~~.:Em = lm**~HI\. mim!i!Wll::~:tn-t7.> .=:.t~l±liilll! :t,;.tCJ'!liili"'JI.t IJ imllit...t-:. \!J:.lEID~#it-tt..~ifi*· ;;t-l,~W nt. ~:Pii stm~~ll'.t.. t-:., ? ~~1i 1 f!H1¥rf<1l't'oo .:.tt,~O)iji.lEmri'l'ft~'t'li.Oil:, 8*0).:::.:\ilf..t 1HJJII.> "Cffl'.!i' ~lim t 1J, i!ti.lEfflitJ#JtlfO)lVf~i;:l;tiJtli:"t' <1; .o. .:.tt. 'f "t':i':l::::. =tHHt:ril: t c-? ~~~~ttn:~n.: Mt. -c~.o,t~n:. .:. .:. "t'iii.lEmft1i~:. J: .o IF.fttl::::. ··:w-cfi=f~ t:.. llliitl/1 O)J~t'lKiiHil-~7:--'l::i~-t.ORfi'mM:~mmit...t=, :Pt~ *:1

upper members. The lower member Introduction and aclmowledgments yields fossils, which were The cephalopods described in this ar· collected from the following localities ticle were collected from the Upper denoted by numbers** : Triassic Tanoura (TAMURA et a!., 1958) · Loc. 85 Buchites kumamotoensis n. sp. and Takagochi (ORITA, 1962) Formations Loc. 43 Phormedites sp. distributing in the Tanoura district, Loc. 46 Rhacophyllites sp. . TAMURA (1959) Loc. 25 Buchites kumamotoensis n. sp. and ORITA (1962) studied the stratigra· Arcestes sp. cf. A. (Proarcestes) phy and molluscan fossils of the Tano­ ausseeanus (HAUER) ura Formation. Almost all the speci­ Ussuritidae or Discophyllitidae mens used in this study were collected gen. et sp. indet. by them, except for a few which were Loc. 44 Holconautilus sp. collected by myself. ORITA (1962) listed several ammonoids The Takagochi Formation is about which were preliminarily examined by 400 m in thickness (ORITA, op. cit.), and MATSUMOTO. Based on the bivalves and yields molluscan fossils through out the ammonoids, the Tanoura Formation was formation. Ammonoid was found in its referred to as the Carnian and the Ta· middle part (Joe. 52 and 55), and is re­ kagochi Formation as the Carnian to presented by only a single species, Pia­ Norian in geologic age (TAMURA 1959; cites sp. aff. P. oxyphyllus (MOJSISOVICS). 0RITA. 1962). This ammonite is associated with Pleu­ According to ORITA (op. cit.), the Ta­ romysidia kanmerai TAMURA, Paleoneil~ noura Formation is about 250m in thick· iwaiensis lCHIKA wA, Halobia sp ., etc .. ness and is divided into the lower and Pleuromysidia kanmerai TAMURA and * Received May 25, 1972; read Jan. 23, •• The numbers of locality correspond with 1972 .at . those of 0RITI. (1962). 447 448 Takeski ISHIBASHI

0 3KM

Text-fig. 1. Location of samples collected.

Halobia (?) sp. occur in the lower part of Takagochi Formation is suggestive (loc. 54), whereas 1Honotis (Entomonotis) of the Norian, conservatively the Carno­ sp. cf. M. (E.) typica (KIPARISOVA) is Norian age. known from the upper part (toe. 56). The cephalopod fauna of the present On the basis of the molluscan fossils study generally resembles that of Sara­ ORITA (op. cit.) concluded that the upper gai Group of Northeast Japan which in­ part was early Norian, and the middle cludes species of Placites, Rhacophyllites and lower parts were Carnian in age. and Arcestes (SHIMIZU and MABUCHI, NAKAZAWA (1964b) also considered that 1932; NAKAZAWA, 1964a). ISHIBASHI the middle horizon (locs. 52, 55) of Taka­ (1970) reported numerous Carnian am­ gochi Formation was referable to the monites from the Okinawa-jima, but Late Carnian. little genera are common between the I investigated the cephalopod fossils Tanoura district and the Okinawa-jima, collected from the Tanoura and Taka­ except for Arcestes. Two species of gochi Formations, and identified one Buchites and Phormedites to be described nautiloid species and five ammonoid spe­ in this article are known from the Ja· cies, of Which one is new. On the basis panese Islands for the first time. of this ammonoid faunule the lower I wish to express my sincere thanks member of the Tanoura Formation is to Professor Tatsuro MATSUMOTO of correlated with the L~te Carnian. Ac­ Kyushu University for his arrangements cordingly the upper member probably of the material. criticisms and reading extends to the Early Norian. Placites of the manuscript, and to Professor sp. aff. oxyphyllus from the middle part Ryuzo TORIY AMA of the same university 607. Triassic cephalopods from Tanoum 449

"'for kindly reading preliminary draft and Buchites kwrlamotoensis ISHIBASHI, -encouragement. Thanks are also due to sp. nov . . Professor Minoru TAMURA of Kumamo· ·to University for the offer of his am- Pl. 54, Figs'. 1-9, 10? and 11?

monoid collections. I am obliged to Miss 1962. Helictites? sp., ORITA, Sci. Rept., Fac. :Seiko 1-IAYAKAWA for typewriting the Sci., Kyushu Univ., Vol. 6, No. 1, p. 4, .manuscript. listed . Repository.-All specimens illustrated ··in this article are deposited· in the De­ Material.-The holotype (GK. F 458) _partment of Geology, Kyushu University and some of paratypes (GK. F 459-462), ·with the registered number using a collected by Dr. TAMURA from the Umi­ :symbol of GK. F. noura area (loc. 25), Kumamoto · Prefec· ture. Two others (GK. F 374, 375) listed already as Helictites ? sp. collected by Systematic paleontology Mr. ORITA and the rest (GK. F 463-466) by myself. Class Cephalopoda CUVIER, 1797 Description.-Shell considerably invo­ :Subciass AGASSIZ, 1847 lute and compressed, increasing gra­ dually its height. Whorl embracing Order Ceratitida HYATT, 1884 about a half or one-third of the inner whorl; umbilicus narrow and moderately Superfamily Clydonitaceae deep; ribs strong, rounded, flexiradiate MOJSISOV!CS, 1879 on the flank and projected on the ven· Family Buchitidae HYATT, 1900 trolateral part, with irregularly inter· calated secondaries, without crossing Genus Buchites MOJSISOVICS, 1893 over the venter ; venter round, smooth where ribs gradually disappear ; suture Type-species.-Ceratites (Buchites) al· ceratitic. with serrated lobes. cdrovandii MOJSISOVICS, 1893. Remarks.-The present species is some· Remarks.-The genus Buchites was what similar to Helictites beneckei re· ·originally established by MOJSrsovrcs ported by MOJSISOvrcs (1893) from the •(1893) as one of the subgenera of Cera· Alps in lateral view, but the ribs of the tites. The species belonging to Buchites latter cross straight over venter . . are known only from the Carnian to The present new species rather resem· Norian of the Tethys region, such as bles Buchites aldrovandii {MOJSISOV!CS) ·the Alps, Greece, Sicily, Himalayas and from the Alps but the whorl of the ·Timor. Buchites has flexuous ribs which latter is more evolute than that of the :are projected at the ventrolateral shoul· former. -der and gradually disappear on the The other known species from the ·rounded venter but occasionally cross Alps and Himalayas are characterized by ·over it in the last whorl of a mature the appearance of the flexuous short ·specimen. Whorls are moderately in­ lateral ribs at the ventrolateral area. B. volute, increasing its height at a mo­ kumamotoensis has bifurcating ribs some ··derate rate. Ribs are distinct and in· of ~hich diverge from the main ribs at ·.tercalated with the short ones. or near the umbilical shoulder, and its suture-line has the serrated accessary 450 '71alcesh'i [SffiBASfll lobes. The suture:iine of B. kumamo­ CKAS (1928) from Timor. On the ·other toensis is rather· similar to B. modestus hand, the Norian one, Ph. juvavicus was~ (BUCH) .from the Alps, but the latter is described . by MOJSisovrcs (1893) from, evidently different in that the ribs sli· the Alps and by PAKUCKAS (1928) from .ghtly cross over the venter and has a Timor. Ph. fascia/us was reported bY' ·broad U:rhbilicus. B. emersoni and B. cf. MOJSISOVICS (1893), DIENER (1906) ana hilaris described by DIENER (1906) have PAKUCKAS (1928) from the Alps, Hima-­ evolute whorls and primitive sutures. layas and Timor, respectively. Buchites heteroplichus reported by Based on the original descriptions of' GEMMELLARO (1904) is very similar to these species, species of Phormedites are" the present new species with respects divided into the following two groups:. to only the whoi! volution. but is clif· 1. The group of Ph. schopeni-Ph. ferent from the latter in the characteris­ juvavicus tics of ribs, whorl size, venter, etc. It · Shell is consideral::lly involute with. is resonable to establish a new species fine, prorsiradiate ribs. The ribs cross .. in the genus Buchites for the present over the venter. The venter is round .. form. The type-species, Ph. juvavicus, is about. Occurrence.-Locs. 25 and 85. With Ar­ 28 mm in diameter. cestes sp. cf. A. (Proarcestes) ausseeanus 2. The group of Ph. pygmaeus-Ph. (HAUER) and Halobia ? sp. (Loc. 25). ani:! fasciatus Pleuromysidia kanmerai TAMURA (Loc. The shell is evolute with fine, prorsi-­ 85). radiate ribs. The ribs are bundled. The' venter is roundt:d with a faint. groove .. Genus Phormedites MOJSISOVICS, 1893 Ph. fasciatus is about 32 mm in diameter.. The characteristics of this group are Type-species.-Ceratites (Phormedites) very similar to those of the genus Da­ juvavicus MOJSISovrcs, 1893 Phinites as pointed out by MOJSISOVICS. (1893). Remarks.-MOJSrsovrcs (1893) establi· shed the subgenus Phormedites in the Phormedites sp. indet. genus Ceratites of the family Ceratitidae. ZITTEL (1900). GEMMELLARO (1904) and Pl. 54, Figs. 13 and 14 DIENER (1906) also treated it as a sub· 1962. Phormedites (?) sp .. OR ITA, Sci. Rept ., . genus of Ceratites. DIENER (1923) ranked Fac. Sci., Kyushu Univ., Vol. 6, No. 1, it as a genus of the same family. but p. 4, listed. SPATH (1951) included the genus Phorme­ dites in the family Buchitidae. SPATH's Material.-A part of outer whorl of' classification seems to be reasonable at rubber cast which was made from an present. The species referable to this external mould (GK. F 377a), and a frag­ genus, including the type-species desig­ mental part of ventral part (GK. F 377b). nated by DIENER (19.15). have been These specimens were collected by Mr. known from the Carnian and Norian ORITA. ages. The Carnian species. Ph, pyg­ Descriptive remarlzs.-The shell is maeus and Ph. schopeni were described considerably involute and compressed. by GEMMELLARO (1904) from Sicily, and Judging from outer whorl. it is at least Ph. aff. pygmaeus was reported by PAKU· about 60 mm in diameter, though the 607. Triassic cephalopods [rom Tanotwa 4,51

,umbilical part is l.ost.. The venter is Type-species.-Arcestes bramantei ·narrowly arc,!}ed. The l~teral surface MOJSISOVICS, 1869 ·of sh6ll .has, numerous, g~ntly fiexira­ ·diate· ri~~',which: are· ~t'Qji~te.d· sharply Arcestes (Proarcestes). sp. cf. A. (P.) for~war\f.r at. the~ ventrolateral shoulder. ausseeanus (HAUER) The rib.$. are: sl.ender, keep, almost the ·same ~i'dth fro'm:'·the umbilital shoulder Pl. 54, Fig. 15, Text-figs. 2 and 3 ·to the. ve.qtral· part, and· cross over the Compars- venter. ·The i~tercostal space is v.ery 1847. Ammonites ausseeanus HAUER, Hai· narrow·near: u.mbilical shoulder. These dinge1·s Nat. Abhandl., I, p. 268. pl. 8, · ch~ra.cters ·s\ligest that the present spe· figs. 6-8. · cim.ens .beJon'g- to the genus Plzormedites. 1875. llrcestes (Proarcestes) ausseeanus HA· 1\;Tiq~·g: the sp.ecies of Phormedites, UER: MoJSISOVICS, ]ahrb. Geol. Rei· -only two, Ph. pygmaeus and Ph. schopeni, chsanst., Wien, No. 6, Heft. 1, pl. 51, have been known from the Carnian of figs. 1 and 4, pl. 53, figs. 28 and 31. :Sicily. Others hav.e been reported. from 1907. Arcestes (P.) ausseeanus HAUER: FRE· ·the Norian of the Alps, Himalayas and CH, N. ]ahrb. f. Min. Festbd., p. 20. pl. ·Timor. As mentioned above, all these 4. fig. 5. ·species are smaller than the species 1914. Arcestes (P.) bicarinatus-ausseeanus HAUER: WELTER, Paliiont. von Timor. from Tanoura, being one·third or a half I, p. 180. in diameter. There are many differences 1927. Arcestes (P.) ausseeanus HAUER: AR­ in respect to ribs, volution, size and THABER, ]ahrb. Mijnw. Nederl. Ind .. venter between the species from the Vol. 55. No. 2, p. 51, pi 5, figs. 1-6. Alps, Sicily and Japan. The specimens at hand are so poor Material.-One external mould (GK. F ·that the present species has been left 467) and its plaster cast (GK. F 468) are unnamed until better preserved material examined here. The external mould :becomes available. was collected by Dr. M. TAMURA of Occurrence.-Loc. 43. Shale bed of . upper Lower Member of Tanoura For· Description.-Shell, involute. globose, mation. With Halobia sp., Halobia mo· about 30 mm in diameter ; whorl. broad, .lukkana WANNER, and Pleuromya fors­ 24 mm in width deeply embracing and .bergi (BOHM). indented by the inner volutions ; helmet· shape in section : surface with three Superfamily Arcestaceae oblique, periodic constrictions cross over the ventral part, no striae and ribs; MOJS!SOV!CS, 1875 venter narrow. its shoulder abruptly Family Arcestidae MoJsrsovrcs, rounded; suture, ammonitic, deeply di· gitate, having numerous lobes and sad­ 1875 dles, with accessary lobes. Genus Arcestes SUESS, 1865 Remarfls.-Arcestes (Proarcestes) aus· seeanus has been known from the Car· 1ype-species.-Ammonites galeiformis nian of the Northern Alps, Timor. HAUER, 1850 Greece, Himalayas. etc. This species was originally described by HAUER Subgenus Proarcestes MOJS!SOVICS, 1893 (1847). MOJSISOV!CS (1873) included this 452 Takeshi ISHIBASHI

species into the group of A. (P.) bicari· natus (MUNSTER), and RENZ (1910) and WELTER (1914) reported ausseeanus as a variety of A. (P.) bicm·inatus. A. (P.) ausseeanus has three constrictions on the lateral part. but the specimens des­ cribed by MOJS!SOVICS (1875, pl. 51. figs. 4a-b) have evidently four constrictions. ARTHABER (1927) described A. (P.) aus· seeanus from Timor. One of the Timor specimens (pl. 5, fig. 5) is immature and has two constrictions, but the other mature ones have four constrictions. A. (P.) hanieli, which was originally des­ cribed by WELTER (1914) from Timor, was also reported by ARTHABER (1927) from the same region. A. (P.) hanieli Text-fig. 3. Suture-line preserved at­ described by ARTHABER closely resem· the external mould of Arcestes (Proarce .. bles A. (P.) ausseeanus, though the !at· stes) cf. au sse anus (HAUER). x 5. ter has a higher whorl than the former. SMITH (1927) described several new Arcestes (A.) aff. oligosarcus MOJSISOVICS .. species of Proarcestes from the Carnian SIIIMIZU and MABUCHI (1932) (Saragail of California. of which A. (P.) pacificus Group, Kitakami) is similar to the present specimen in Arcestes sp. NAKAZAWA (1964) (Saragai~ whorl section, but the former has a Group, Kitakami) smaller shell and a simpler, flared suture· Arcestes (Stenarcestes) sp. NAKAZAWA (1959)· (Jito Formation, Nariwa, ) line in which the accessary lobes. are not Arcestes (Proarcestes) aff. hanieli WELTER~ developed. SHIMIZU (1931) (Proarcestes Bed, !CHI~ The following species of arcestids have KAWA (1951)) been known from the Japanese Islands: Arcestes (P.) aff. bicarinatus (MUNsTER) SHIMIZU (1931) (Kochigatani Group, Shi-­ _...- -.__ / ' koku) / I The first two species came from the, I Norian strata, whereas the last thre~ I from the Carnian. The present species. \ is easily distinguished from these spe­ \ I ... 't"',....' cies by whorl section, suture-line ancl __ other characteristics. \ ', To sum up the present specimen is. \ best comparable to and may be iden­ \ ,_ tified with A. (P.) ausseeanus with res­ ' pects to whorl section, ornamentation,. 0 10 ----- mm suture-line and other characteristics. 'Text-fig. 2. Whorl section of Arcestes Occurrence.-Loc. 25. Shale bed of (Proarcestes) cf. ausseeanus (HAUER). upper Lower Member of Tanoura Forma- 607. Triassic cephalopods from Tanoura 453

; . . . ·.; '· . . ;., . . \ O______IOmm

Text-fig. 4. Suture-line of Placites aff. oxyphyllus (MOJSISOVICS). tion, where Buchites kumamotoensis Isru­ rather involute and smooth on· the sur­ BASI·II and Halobia ? sp. are associated. face. The essential part of the suture is well preserved on the present spe­ cimen, showing the typical pattern of Superfamily Pinacocerataceae Placites, which consists of numerous, MOJSISOV!CS, 1879 deeply incised elements (Text-fig. 4). The present species closely resembles Family Pinacoceratidae PL. oxyphyllus (MOJSISOV!CS, 1873), from MOJS!SOV!CS, 1879 the Norian of the Alps and Sicily, in Genus Placites MOJSISOVICS, 1896 that its external saddle has shallow lo­ bules. Placites perauctus (MOJSISOVICS, Placites sp. aff. P. oxyphyllus 1873), from the Norian of the Alps, has (MOJSISOV!CS) deep lobules on the external saddle but differs from Pl. oxyphyllus by the deep Pl. 54, Figs. 12a-c and Text-fig. 4 lobules in the lateral saddles. Two ammonoids, Placites aff. oxyphllus 1962. Placites? sp.; ORITA, Sci. Rept., Fac. Sci., Kyushu Univ., Vol. 6, No. 1, p. 9, and Arcestes aff. oligosarcus have been listed. listed by SHIMIZU and MABUCHI (1932) from the Norian Saragai Group, Kita· Material.-One individual formed of kami. but their specimens have been lost three parts (GK. F 379) is here examined. (NAKAZAWA, 1964a p. 25). So it is im­ They are two external moulds and an possible to compare the present speci­ internal mould collected by Mr. ORITA. men with them. The Takagochi Forma­ Descriptive remarks.-The shell is com· tion, in which the present specimen J pressed with a narrowly arched venter, occurs. has been considered by ORIT A 454 Takeshi ISHIBASHI

(1962) as the Carno-Norian in age. There is a possibility that the bed containing Placites aff. oxyphyllus belongs to the Norian age, though the present speci· men is not so well preserved that the specific name has not been determined with certainty. Occurrence.-Loc. 52. Shale in the middle part of Takagochi Formation, Kumamoto Prefecture. It occurs with Pleuromysidia kanmerai TAMURA, Paleo­ neilo iwaiensis ICHIKAwA, Halobia spp., Camptonectes sp., Oxytoma sp., etc. ac­ cording to ORITA (1962). o______;_:IO m m

Text-fig. 5. Suture-line of Rhacophyllites sp. Order Phyltoceratida ARKELL, 1950

Superfamily Phyttocerataceae to Discophyllites, in the whorl shape and ZITTEL, 1884 ornamentation. The difference of the two genera is mainly in the pattern of Family Discophyllitidae SPATH, 1927 the sutures. Namely Rhacophyllites is Genus Rhacophyllites ZITTEL, 1884 diphyllitic or triphyllitic, whereas Dis· cophyllites is monophytlitic. The species Rhacophyllites sp. indet. of Rh. are known from the Upper Car­ nian to Norian in the various part of Pl. 54, Fig. 16 and Text-fig. 5 the world. When NAKAZAWA (1964a) described 1962. Rhacophyllites? sp.; ORITA, Sci. Rept., Rhacophyllites sp. from the Saragai Fac. Sci., Kyusltu Univ., Vol. 6, No. 1, p. 4, listed. Group, the present specimen was exa­ 1964. ·Rhacophyllites sp.; NAKAZAWA, Mem. mined by him. He said that the chara­ Col!. Sci., Univ. , .Vol. 30, No. 4, cters of Rh. sp. of the Saragai Group p: 29, 4th line from the top. resembled especially those of the Car· nian Rhacophyllites sp. of the Tanoura JV!aterial.-One fragmental specimen Formation rather than those of Norian (GK. F 371) is examined here, which was species such as Rh. debilis and Rh. neo· collected by Mr. ORIT A. jurensis and those of Carnian species Descriptive remarks.-The shell is in· reported by TRECHMANN {1917) as Dis­ volute with a rounded venter. The urn· cophyllites cf. ebne1·i. bilical part is almost lost. The suture As mentioned above, the species of is exposed on the ventrolateral part of Rhacophyllites from the Tanoura Forma­ the shell, in which the first lateral sad· tion and the Saragai Group certainly dle shows a diphyllitic type (Text-fig. 5). resemble each other in the character of The phylloid leaf is slender for the suture. The sutural patterns of these genus, probably because of the immature two specimens are also quite similar to state of the shell. Rhacophyllites is close that of Rh. ·· neojurensis" illustrated by 607. Triassic cephalopods from Tanoura 455

WELTER (1914, pl. 30, fig. 6*, text-fig. 72) develop on the flank. The venter is from the Norian of Timor which was round and smooth. The suture-line runs ·excluded by WIEDMANN (1970) from the almost parallel to ribs and shows one ·synonym list of the true neojurensis of lateral lobe. ~The siphonal position is WELTER. I would support the opinion not observabie in the present specimen. ·of WIEDMANN. Unfortunately the spe· On the grounds of the observed cha· ·cific name was not given by WIEDMANN racters the present specimen is referable for the Timor specimens at that time. to the genus Holconautilus. The species On the basis of suture-line, the present of Holconautilus are known from the ·specimen may be suggestive of the Early Anisian to Carnian of the Alps, Ger· Norian in age. many, the Meditteranean region, Timor. Occurrence.-Loc. 46. Shale of the and Japan. The present specimen is ,upper Lower Member of Tanoura For· closely similar to H. semicostatus (BEY· mation. With Pleuromysidia kanmerai RICH) in volution. ribs and suture. The "TAMURA. other species of Holconautilus are dif· ferent from the present species in the characters of whorl. The whorls in the .Subclass Nautiloidea AGASSIZ, 1847 former species increase height quicker Order Nautilida AGASSIZ, 1847 than H. semicostatus and the present species, and sometimes have a ventro· Superfamily Tainoceratacea lateral tubercle on the rib. HYATT, 1883 BANDO (1964) described Pleuronautilus (Holconautilus) yabei BANDO from the Family Tainoceratidae HYATT, 1883 Ladinian Rifu Formation of Northeast Japan. The present specimen is evi· cGenus Holconautilus MOJSISOVICS, 1902 dently distinguished from it in that it Type-species.-Nautilus semicostatus has a quadrate whorl section, with a flattened venter, two spiral rows of 8EYRICH, 1867 nodes on the venter and less prominent Holconantilus sp. i ndet. ribs. Pleuronautilus(?) sp. reported by YABE Pl. 54, Fig. 17 and SHIMIZU (1927) without text-figure is ornamented with distant umbilical Material.-One specimen (GK. F 378) ·collected by Mr. ORITA, in 1962. is exa· tubercles. Occurrence.-Loc. 44. Shale bed of the mined here. Lower Member of Tanoura Formation Descriptive remarlzs.-Only outer whorl with Freguelliella (l(umatrigo11ia) tanou­ {ca. 60 mm in diameter) is preserved. rensis TAMURA, Balcevelliid sp., Costa­ The shell is very evolute.and presumably toria (?) sp. widely umbilicate. The whorl is nearly .as high as broad, increasing its height gradually. The rectiradiate, ·distant ribs References cited

* WIEDMANN did not exclude the figs. 5 ARTHABER, G. von (1927) : Ammonoidea ;and 7 in the same plate but the suture of Leiostraca aus der oberen Trias von .fig. 6 seemed to be drawn from the specimen Timor. 2 Nederl. Timor Expeditie 19~6 •Of figs. 5 and 7. onder Ieiding van Dr. H. G. ]ONK~R :· ~ . . :. ,.. . '. \ ;~ . :·. .~... ~

456 Talceshi ISHIBASHI

Utigegeven soor Dr. H. A. BROUWER, 4, Triassic stratigraphy of Japan. GeoL jaarb. Mijnw. Nederl. Ind., Vol. 55, Pt. Surv. japan, Rep Spec. No., pp. 108-110. 2, pp. 1-174, pis. 1-20 (1926) (in Japanese) BANDO, Y. (1964): On some Middle Trias­ ISHIBASHI, T. (1970) : Upper Triassic Am­ sic fossil cephalopods from Japan, with monites from Okinawa-jima, Part 1. Mem. a note on the Middle Triassic formations Fac. Sci., Kyushu Univ., Ser. D, Geo/.,. in Japan. japan. jour. Geol. Geogr .. Vol. Vol. 22, No. 2, pp. 195-223, pis. 26-29. 35, Nos. 2-4, pp. 122-137, pl. 5. MoJSISOV.ICS, E. von (1869) : Beitrlige zur DIENER, C. (1906) : Fauna of the Tropites· Kenntniss der Cephalopoden-fauna desc Limestone of Byans. Palaeont Indica, alpinen Muschelkalks (Zone des Arceste.s: Ser. 15, Vol. 5, No. 1, pp. 1-201, pis. 1-17. studeri). jahrb. Geol. Reichsanst., Wien,. -- (1915) : Cephalopoda triadica. Fossilium Vol. 19, pp. 567-594, pis. 15-19. Catalogus, Altimalia I, Pars 8, pp. 1-369, -- (1873) : Die l\4ollusken-Faunen der ZJa. Junk, Berlin. mbach· und Hallstatter-Schichten. Ab­ -- (1920) : Die Ceratitoidea der karnisch­ handl. Geol. Reichsanst., Wien, Vol. 6, norischen Mischfauna des Feuerkogels No. 1, pp. 1-82, pis. 1-32. bei Aussee. Sitzungsber. Akad. Wiss. -- (1893) : Die Cephalopoden der Halls­ Wien, Vol. 129, pp. 589-618, pis. 1-3. tatter Kalke. Abhand/. Geol. Reichsanst., -- (1923) : Ammonoidea Trachyostraca Wien, Vol. 6, Pt. 2, pp. 1-835, pis. 71- aus der mittleren und oberen Trias von 200. Timor. jaarb. Mijnw. Nederl. Ind., Vol. NAKAZAWA, K. (1959): Two cephalopod 49, pp. 73-276, pis. 1-23. species from the Norian Nariwa GrouP' FRECH, F. (1907): Die Hallstlitter Kalke in , West Japan. bei Epidauros (Argolis) und ihre cephalo· japan. jour. Geol. Geogr .. Vol. 30, pp. poden. Neues ]ahrb. Min. Geol. Paliiont. 127-133, pl. 11. (Fest Bd.), pp. 1-32, pis. 1-6. -- (1964a) : On· the Monotis typica Zone in GEMMELLARO, G. G. (1904) : I cefalopodi del Japan. Mem. Col/. Sci., Univ. Kyoto, Ser. Trias superiore della regione occidentale B, Vol. 30, No. 4, pp. 21-39, pis. 3-5. della Sicilia. Giornale Sci. Nat. Econ., -- (1964b) : On the Upper Triassic Monotis Palermo, Vol. 24, pp. 1-319, pis. 1-30. Beds, especially, on the Monotis typica I-lAUER, F. von (1847): Neue Cephalopoden Zone. jour. Geol. Soc. japan, Vol. 70, aus dem rothen Marmor von Aussee. No. 829, pp. 523-535. (in Japanese with /laidinger's Naturwiss. Abhandl., I, pp. English abstract) 257-277, pis. 7-9. ORITA, Y. (1962): Upper Triassic Series in IcHIKAWA, K. (1951): Triassic formation at the Tanoura district, Kumamoto Prefec· Taho district, . in the ture. Sci. Rep. Fac. Sci., Kyushu Univ.,

Explanation of Plate 54

Figs. 1-11. B11chites kumamotoensis IsHIBASHI, n. sp. Fig. 1 GK. F 463: Fig. 2 GK. F 375 ventral part x2: Fig. 3 GK. F 459 x2: Fig. -i GK..F 465: Fig. 5 GK. F 464: Fig. 6 GK. F 462: Fig. 7 GK. F 460: Fig. 8 GK. F 458, Holotype: Fig. 9 GK. F 461 : Fig. 10 GK. F 374: Fig. 11 GK. F 466 x 2. Figs. 12 a-c. Plactes sp. aff. P. oxyphyllus (MOJSISOVICS) GK. F 379 X 2. Figs. 13 and 14. Phormedites sp. Fig. 13 GK.F 377b ventral part: Fig. 14 GK.F 377a. Fig. 15. .4rcestes sp. cf. A. (Proarcestes) ausseeanus (HAUER) GK. F 468. Fig. 16. Rhacophyllites sp. GK. F 371 ventral part. Fig. 17. llolconautiltts sp. GK. F 378. (Figures in natural size except as indicated) ISHIBASHI : Upper Triassic cephalopods Plate 54 607. T?·iassic cepha~opods jrom Ta•noura 457

Ser. Geol., Vol. 6, No. 1, pp. 1-13, 2 maps, Tosapecten from the Carnic Tanoura for­ (in Japanese with English abstract) mation in Kyushu, Japan. Mem. Fac. PAKUCKAS, C. (1928) : Nachtrag zur mittel· Educ., Kumamoto Univ, Vol. 7, pp. 212- und obertriadischen Fauna der Ammonea 224, pl. 2 .. : trachyostraca, C. DIENER's aus Timor, -- (1960) : Stratigraphical study on the­ mit Einleitung und stratigraphischer Zu· Sakamoto Group in Kyushu. jour. Geol. sammenfassung von G. von ARTHABER. Soc. japan, Vol. 66, No. 777, pp. 371-383- jaarb. Mijnw. Nederl. Ind., Vol. 56. pp. (in Japanese with English abstract). 143-218, pls. 1-2. --, KANMERA, K., AKAZU, K. and YAMA·­ RENZ, C. (1910) : Die Mesozoischen Faunen SHITA, A. (1958): Notes on some new Griechenlands. Theil 1. Die Triadischen finds on the Mesozoic strata in the Kuma. Faunen der Argolis. Palaeontographica, and Districts. jour. Geol Soc. Vol. 58, pp. 1-104, pls. 1-7. japan, Vol. 64, p. 370 (in Japanese). SHIMIZU, S. (1931): Notes on two Carnic TRECHMANN, C. T. (1917): The Trias of· species of Proarcestes from . New Zealand. Quart. jour. Geol. Soc. japan. jour. Geol. Geogr., Vol. 8, No. 3, London, Vol. 73, pp. 165-245, pis. 17-25. pp. 117-121, pis. 12-13. WELTER, 0. A. (1914): Die obertriadischen. -- and MAsucHI, S. (1932): Upper Trias­ Ammoniten und Nautiliden von Timor. sic in the Kitakami Massif. jour. Geol. Paliiontologie von Timor, Vol. 1, pp. 1-· Soc. japan, Vol. 39, No. 465, pp. 313-317 258, pls. 1-36. (in Japanese). WIEDMANN, J. (1970) : Uber dem Ursprung SMITH, ]. P. (1927) : Upper Triassic marine der Neoammonoideen-Das Problem einer· invertebrate faunas of North America. Typogenes. Eclogae Geologicae Helvetiae,. U.S. Geol. Surv., Prof. Pap. 141, pp. 1- Vol. 63, No. 3, pp. 923-1020, pis. 1-10. 261, pis. 1-71. YABE, H. and SHIMIZU, S. (1927) : The­ SPATH, L. F. (1951): The Ammonoidea of the Triassic Fauna of Rifu near . Trias (II). Cat. Fossil Cephalopoda Brit. Tohoku Imp. Univ., Sci. Rep., 2nd, :Ser .. Mus. (Nat. Hist.), Pt. 5, 209 p. Geol., Vol. 11, No. 2, pp. 101-136, pls. TAMURA, M. (1959): On Kumatrigonia, a 10-14. new subgenus of Frenguelliella, and a

]ito j1J!. iiJi Rifu li'U lf.f Kitakami :lt J:: Saragai 1D1 ~ Kochigatani i"II*W~ Takagochi Al iiif r"l Nariwa $. ~~ Tanoura IE im Okinawa-jima i'P*'l.l% Uminoura iii iili Ota ~ [lJ "Trans. Proc. Palaeont. Soc. Jagan,. N. S.., No .. 88 .. pp. 458-461, pl. 55 .. December 20, 1972

-608 .. A FOSSIL WOEM TRAIL FROM THE UPPER NAKAMINATO FORMATION, 1BARAKJ". PREFECTURE*

KOTORA HAT AI and HIROSHI NODA

Institute of Geology and Paleontology, Faculty of Science, Tohoku University, Japan

$C!JI.i.)~U.:lltll3.illi*n11Jii.l~mffii ;:A,ii:-ff.JI::.:O~C."?v''L: ~!J£1'iHJFIJii.IIJfm 1HI!tiiUI;i!l: 5Hrd-~ .1·.:111!3!!~!*JK>Jii.l~m!fltlil<7)lf~il!T$/11i'€> ilE!±ll...t.: liv'i!JFft~lifil~:iXQ)R:~*"l 29 em. m 1.2-2.0 em Q)flji!llllilfi~ 1..-bb !..-, f~ 3 em Q)ll!ISJ'I;:r.J 8 JlJQ)!J~J:f*ti'(;;?: Ji!.€>:11.. Jf?-~(~l:::tn{J.:.l -n, Nentes Q){ljlfh,(J).t? tJ:t\':IA:$1~fffi(J)tl/i:iX~Wl~JJ!i~1:ffl:l/:-ff.t.!tt?:h.tJ:It'. -:J:t:. U'l~J:f* [!li): ~n!J,;j" '.) Arenicola, Balanoglossus, Chaetopterus c <7) lt~~iit M'TfJ:-:> t,:;?:, .I!'~ P1Jffll~ffi{Q)t11,.,~{1:t.il1 Hitachia nakaminatoensis fJ:'.}~~. f/i~"eif.,'.);: c ~~~t.:. ~ ; 1:::. *ll:.:r:ii!E!fll;: J:: ~ lf-~tl!IRhf:*ilmJtffl~Q)ffii~rf.HJJilJIQ)lGi¥ifii~Q)JftfJ'H~l? ~ '.>;: c '.i::~~l..-t.:. WJ:J:t-1J'11f. g!fiiJfc!iffi

comprises chiefly dark gray sandstone Introduction and acknowledgments with interbedded graded sandstone lay­ Exposed on t.he surface of a weather· ers." The Hiraiso-machi Member yielded -ed sandstone at the sea-side of Hiraiso. to SAITO (1961, p. 112), Didymoceras awa­ Nakaminato City, lbaraki Prefecture, jiense (Y ABE), D. nakaminatoense (SAITO), was found a trail of a worm, probably Nipponaster nakaminatoensis SAITO, be­ ·of marine origin. This trail, impressed sides several dther 'fossil molluscs. And ·on the sandstone of the Hiraiso-machi the discovery of the fossil worm trail Member of the Nakaminato Formation verifies that the Hiraiso-machi Member is the first discovery of a trail fossil was deposited in shflllOw water. from the member and is thus considered The worm trail uppon which the pre· important for interpretation of the con· sent article is based was collected by ·ditions under which that part of the the junior writer during the writer's formation was deposited. The Hiraiso­ field trip to the Nakaminato area with machi Member occupies the middle part the students majoring in geology of the ·of the Nakaminato Formation of Upper Faculty of Education, lbaraki University, ·Cretaceous age. guided by Associate Professor Toshio According to SAITO (1961) the type SAITO and Research Associate Haruyu­ locality of the Hiraiso·machi Member of ki TAKAHASIII of the same University. the Nakaminato Formation is "at the At this place the writers express their sea coast of Hiraiso-machi where it at· deep appreciation to Associate Professor tains about 510 meters in thickness and Toshio SAITO and Research Associate Haruyuki TAKAHASHI for their kind * Received June 3, 1972; read June 3, 1972, guidance to the Cretaceous deposits dis· .at . tributed along the coast of Hiraiso, Na· 458 608. Upper ·cretaceous worm trail

Tertiary System

...... unconformity ...... ~ lsoai Member, 730 m thick. Graded sandstone and siltstone. Inoceramus cf. shikotanensis, Nipponaster nakaminatoensis, Baculites inornatus, etc. Hiraiso-machi Member, 510 m. thick. Siltstone with inter· Nakaminato Formation I ca:tated sandstone. Didymoceras awajiense, Pravitoceras sp., etc. Chikko Member, 35m thick. Granule ·sandstone in lower and \ .coarse to fine grained sandstone in upper ...... conformity 1030 m thicl<. Conglomerate intercalated with thin sandstone Oarai Formation layers, coal seams, and plant fossils. · Base not exposed ·······--··-········--·························· kaminato City, and also for their infor­ posterior extremities, and thus its ori­ mation on the geology of the area. ginal length remains unknown. But, as. Thanks are due to Mr. Kimiji KUMAGAI shown in the annexed figure, features. for photographic work and to Mrs. Ki­ suggesting the original segm·ents of the miko SHIBUYA for typing the manus­ body, and the approximate thickness of cript. the rounded .body are still preserved_ Since the sandstone ·layer in Which the. Stratigraphic position of the worm trail is .preserved .is not over­ turned, the exposed q10rphological fea­ worm trail tures represent the original .ventral -sur· The stratigraphic position of the face of the ·worm. Hiraiso-machi Member in the geological The worm trail measures. about 29 em column of the Cretaceous deposits dis­ in length measured along its longitu· tributed along the sea coast of Nai

460 Kotora HATAI and Hiroshi NODA

-the mentioned genera, the present fossil taxonomic classification be emphasized. -worm trail shows resemblance with the Recent species Arenico/a crista/a STIM· Genus Hitachia HATAI & NODA, n. gen. · PSON, now living along the coast of .Japan. That species, according to OKU· Type species :-Hitachia nakaminato­ DA and I!ZUKA (1949) attains about 60- ensis HATAI and NODA, n. gen. et n. sp. "90 mm in length and about 18 mm in Type locality, formation, geological age, width. Crista/a lives in sandy mud or and depository:-Upper Part of the Hi­ muddy sand in shallow depths along the raiso-machi Member of the Nakaminato "coast of Japa_n. Arenicola claparedii Formation exposed along the sea coast LEVINSON (OKUDA and IIZUKA, 1949), at Hiraiso, Nakaminato City, lbaraki which attains about 100-200mm in length Prefecture, Cretaceous. Preserved in the :also lives in sandy mud or muddy sand collection of the Institute of Geology and in shallow water, and is distributed from Paleontology, Faculty of Science, Toho· ·Central Japan to the Kurile Islands. Of ku University (Reg. No. 91765). ·these two Recent species, the present Diagnosis :-Body rounded· elongate. ·fossil worm impression seems to be measuring .12-20 mm in minimum and more similar to A. cristata than to A. maximum widths, and 29 em in length, -claparedii, although differing from both. segments well developed. strong, num­ Among the fossils reported by HO· bering 8 in a distance of 30 mm. With­ ·WELL (1962), none expect possibly the out longitudinal mesial depression or ·_genus Tosalorbis proposed by KATTO podia-like structures. "(1962) based upon specimens from the Eocene Muroto Formation in Kochi Pre­ Hitachia nakaminatoensis HA TAl ·fecture, Shikoku, Japan, seem to be com· ·parable with the present fossil. How­ and NODA, n. sp. . ever, Tosalorbis differs from the pre· Pl. 55 ·sent specimen in having closer set seg­ . ments and much shorter body length. Description:-Worm trail impression :Specimens comparable with the present elongate, winding, 29 em in length; body . one were not noticed in the recently rounded, thicker at middle part than at published work "Trace Fossils" edited extremities, measuring 12 to 20 mm in :by CRIMES and HARPER (1970). minimum and maximum widths; seg· Under the circumstances mentioned ments well developed, strong, number •above it seems that a new name should 8 within a distance of 30 mm, broader _given to the present interesting worm than interspaces, indistinct at or near -trail, the first of its kind from the Cre- extremities; external morphology areni· -taceous of Japan, in order that its usage coloid. in biostratigraphy, paleoecology and Remarks :-Comparison and affinity of

Explanation of Plate 55

~Hitachia nakaminatoensis HAT AI and NODA, natural size. Locality: Sea coast near Hiraiso, Nakaminato City, , Upper Cretaceous Hiraiso-machi Member of the Na­ kaminato Formation, IGPS coli. cat. no. 91765. HAT A! and NODA: Upper Ctetaceous worm trail "Plate 55

K. KuMAG A I photo. ! ~.

608. Upper C1·etaceous worm. trail 461 the present worm trail with other spe­ tise on Invertebrate Palaeontology, Part -cies was given above and here it is to W, Worms, p. 144-177, figs. 85-108. be said that its similarity with the spe­ KATTO, J., 1960. Some Problematica from -cies of the genus Arenicola suggests the so-called unknown Mesozoic strata of that it may have led a similar life. It the southe~n part of Shikoku, Japan. Sci. Rep. Tohoku Univ., 2nd Ser., Spec. Vol., is judged that the fine-grained some­ (HAI'ZAWA Memorial Volume), no. 4, p. what muddy sandstone of the upper 323-334, 2 pis. part of the Hiraiso-machi Member in MACSOTAY, 0., 1967. Huellas problematicas which the present worm trail was found, y suvalor paleoecologico en Venezuela. and where some ammonite and pelecypod Geos. Escuela de Geologica, Minas y Me­ fossils have been found, was deposited talurgiu, Univ. Vent. Venezuela, no. 16, p. under a shallow water, open sea en· 7-38, 18 pis. vironment, and further it is inferred OKADA, S., and liZUKA, K., 1949. Polychaeta that the water temperature was mo­ in lllustrated Encyclopedia of the Fauna

Chikko Muroto ~ p Hirai so Nakaminato »11 Ji!I ~ Isoai Oarai * i')l; Trans. Proc. Palaeont, Soc. Japan, N. S .. No. 88, pp. 462-471, pl. 56 December 20, 1972

609. AMMONITES FROM THE MESO,ZOIC Y AMABU FORMATION, KYUSHU

MASAYUKlNODA

Wasada Junior High School, Oita

.7L.fflllHlllliM •€> Hii:U L.t.: 7 :/ ~ 7-1 1- 1::-::> "''"C : JL~1j0) ~5C~fl;:lfli Jtp':JI.::J'll:f1-n, S!lli ~iifl't~O)-·-::>-e ~ Q llJffil/lf';l:.: tt. ~ -c: ~~ifU/ltO) ~~{:ti*lt e: lj. t.t: ~ tt. -c ~ t:vi, Ul>t: rJ.7cO) ~Ul;~J: IJ.::f!lii7J7:/~'}-1'J-il:/*~~ttf.:. --::>f';l: 3-P ou'ti!i~!tl!~O)-'!J7'/7:/::a: ij'j7f-i" Q Berriasella "('. ftl!.i';l:J{ 7 :/-¥ :=. 7 :/J:.ffil ::a:t.\'~-?tt Q Pset~doosterella l:tt·~ f... ~QtO)~~~ • .:~~J:?-c~ma*~•~t~J:.ili!tl!O)•~· w•~·~•mw!tl!O)~ tLJIIliii::~Ht ~,? ~ .-.; ... 7;Y:I.::tt-~-::>?~t•-c:pp!t~ t... .f:'O)f!Jl.Wi :5.~¥Hit17J~l/Iic L.. '"C ~~II IJ 1.:: -:r Q c c t lc.. :Jl!:aij. fil!ttll.llt.t: I? rJ.I.::.'I'llJill/!~ t ~U>'"C. ;:O)f-:rJ.17J~.lM~ .-.;.,. 7*S!li!~17J.t;!fi.I:OO-tQ IE~I¥Jr,mml::~tt..-c6atJg -~1.::-::>~t•'t'-'f g~~~A-tQ, •mm~

and assigned the sequence to the Lower Introductoin Cretaceous Ryoseki Series. Further· The Mesozoic Yamabu formation is more, he pointed out that the forma· exposed in a narrow area of eastern tions described above were all of bra· Kyushu, which belongs to the Chichibu ckish water or non-marine environment. belt of the Outer Zone of Southwest Since then, MATSUMOTO (1954, 1967) Japan. It lies unconformably on the regarded this formation as the Ryoseki Paleozoic strata in the western part of Group, and he and TERAOKA (1970) as· the southern margin and is demarcated signed it to the Kochian Series. by faults in the other sides, occupying Anyhow, the Ryoseki Group thus un· the area of about 6 km from east to derstood was assigned to the lowest part west and about 1.2 km from north to of the Cretaceous deposits of Japan, and south. Its sequence is about 450 m in approximately correlated to the Wealden thickness. of Europ3. The stratigraphy and the structure of On December 29th, 1970. the author this formation were previously reported unexpectedly collected several speci­ by FUJII (1954) and TERAOKA (1970). mens of ammonites from the black shale FUJII considered that the basal to middle intercalated in the basal conglomerate, member of the Yamabu formation might of which the one is referable to a spe­ be correlated to the Ryoseki formation cies closely allied to Berriasella patula of south Shikoku, and also to the Kawa· from the various localities of France. guchi formation of the Yatsushiro area, There is furthermore another specie3. western Kyushu, for the reasons of the This paper deals with the paleontolo· resemblance in fossils and lithofacies, gical descriptions of the two species of

462 609. Ammonites front Yamabu .formation 463

EXPLANAliON shalt _./ laull sandston• fossil locality () co119lomoroto () tos

2km

Text.fig. 1. Geological map of the Yamabu area, showing lithostratigraphic division and distribution (adopted from TERAOKA, 1970).

ammonites from the Yamabu formation with a remark on the geological age and Stratigraphy of the Yamabu formation also some discussions on the problem of -Cretaceous boundary in ] a pan. The Yamabu formation structurally Before going further, I express my shows the syncline with axial trend from sincere thanks to Professor Tatsuro east to west. In its sequence four minor MATSUMOTO of Kyushu University for sedimentary cycles are distinguished. his kind guidance and supervision of (1) Basal member. About 20 m-90 m this study and also to Dr. Tadashi SA TO in thickness. Mainly exposed on the of the University of Tokyo for his southern synclinal wing and partly in useful suggestion about the identifica­ the northern area, generally thickened tion of the species. Acknowledgements westwards and northwards. Its lower are also due to Dr. Yoji TERAOKA of the part consists of ill-sorted conglomerate Geological Survey of Japan for his kind­ with intercalation of black shale about ness of permission to adopt his geolo­ 20 m in thickness near Shinkai, Honjo gical map in this paper. Village, Minamiamabe County, Oita Pre- 464 Masayulci NODA fecture, and generally becomingirich in (3) Middle member. Ab.out 200m in sandstone upwards. The upper part thickness. Its lower part consists of consists of thin bedded and moderately conglomerate about 10-15 m in thickness. thickened alternation of sandstone and The main,·middle part is fine to medium­ shale. Th'e··conglomerate is made up of grained sandstone and shale; the upper well-rounded boulders, cobbles and peb­ part. sandy shale. The features of this bles of granite, granite-porphyry, aplite, conglomerate is similar to those of the diorite, quartz-porphyry, porphyrite, basal one in grading, and kinds of sandstone, slate, chert and green-schist, gravel but distinguishable in its more commonly with sandy matrix and partly muddy matrix. Occasionally '·it is so with muddy one. Sandstone is well­ rich in matrix that it can be called the sorted and medium-grained. Thin bed­ pebbly mudstone. In the main part, the ded shale inserted in sandstone bears following brackish water mollusks of carbonaceous or poorly preserved plant the Ryoseki type are contained in the fossils. ' black. shale inserted in feldspathic­ (2) Lower member. About 80 m" in quartz-sandstone (FUJII, 1954; TERAOKA, thickness. Its lower part consists of 1970). white or grayish white feldspathic-quar­ tz-sandstone, well-bedded about 1 m in Brotiopsis kobayashii SUZUKI thickness. The main part consists of Polymesoda (Isodomella) otsukai (Y ABE hard, fine to medium-grained sandstone, and NAGAO) occasionally inserted with thin layers of Protocyprina cf. naumanni (NEUMA YR) shale, and gradually becoming more Eomiodon cf. sakawanus (KOBAYASHI fine-grained upwards. and SUZUKI)

...r .0 > EXPLANATION z -·r------~ '18 301 ~ 0 •nal• i= E < M 0 ~ 0 a: .0 N lillillilll > ~ sandston• :::> _.. CD ~ < conglomfl'Qto ::L -l-E ~ 0.. p --+------0 VB 102 plant fossil bod E .0 0 > ___... L______it '18 101 x animal fossil bod Paleozoic

Text-fig. 2. Schematic columnar section of the Yamabu formation. 609. Ammonites from Yamabu formation 465

()strea ryosekiensis KOBAYASHI and Material.-One specimen from toe. SUZUKI YB101. the basal member of the Yamabu Aloides sp. formation, Oita Prefecture, Kyushu, col­ lected by M.: NODA (1970) is concerned (4) Upper member. More than 100m with the description. 'in thickness, mainly occupying the axial Descriptive remarks.-The specimen is :area of· the syncline. Its lower part so crushed secondarily that the original -consists of conglomerate. The main proportion of breadth and height is un­ part is dark gray coarse-grained sand­ certain. The umbilical wall is probably stone and thin bedded alternation of steep, forming a rounded shoulder with sandstone and shale. Upper limit is the flank. The whorl is fairly evolute, uncertain becaus.e of the synclinal struc­ probably less than one fourth of inner ture. Sandstone is commonly gray­ whorl being embraced by the outer one, wacke with hard matrix of aphanitic increasing comparatively slowly in silica. height. The original proportion of the The geological map and stratigraphic umbilicus and the diameter is not pre­ --column of this formation are shown in cisely known owing to the effect of ·Text-figs. 1 and 2, deformation, but it is probably larger than the measured value. The flanks are rather flat. on which, especially on Paleontological descriptions the outer two whorls, the ornamentation is fairly well-preserved. On inner Family Berriasellidae SPATH, 1922 whorls the ornamentation is unfortuna­ ·Subfamily Berriasellinae SPATH, 1922 tely indiscernible because of the poor preservation. In later stages more ~han Genus Berriasella UHLIG, 1905 33 mm in diameter, the whorls are orna· mented with sharp, distinct and some· Berriasella sp. aff. B. patula SCHNEID what sigmoidal ribs. They begin at the PL 56, Fig. la, b. umbilical margin with conspicuous ele­ vation and bifurcate at the middle or ·Compare- at the point a little below the middle .1915. Berriasella patula ScHNEID, Ceo/. Pal­ of the flank, curving considerably for· aeont. Abh. 13, (5), p. 66, pl. 7, fig. 5 ward on the ventrolateral part. There 1939. Berriasella patula, MAZENOT, Mem. are 41 ribs in the last whorl which is Soc. Geol. France, N. S. N•. 41, pp. 60, 61, pl. 4, fig. 5a, 5b. more than 116 mm in diameter. On the ..1962. Berriasella sp. aff. patula, CoL.LIGNON, last whorl. the ribs are as wide as in­ Atlas des fossils Caracteristiques de terspaces and on the inner half of the Madagascar, p. 6, pl. 117, fig. 772 flank, the interspaces are fairly wide

Measurements.- Diameter Umbilicus U!D (1) 105.1 mm 41.4mm 0.40 (2) 85.4mm 37.1mm 0.42 (1) At the end of visible whorl. .(2) About 65' back from (1). 466 Masayuki NODA

Table 1.

species material diameter umbilicusj u;o - B. priversensis MAZE NOT pl. 2, fig. 3 58mm 2lmm 0.36 B. priversensis MAZE NOT pl. 2, fig. 4 56mm 23mm 0.40 B. priversensis MAZE NOT pl. 2, ftg. 6 42mm 16mm 0.38 B. sp. aff. B. priversensis MAZE NOT pl. 2, fig. 5 38mm 12mm 0.32 B. collistoides MAZE NOT pl. 7, fig. 1 76mm 22mm 0.29 B. callistoides MAZE NOT pl. 7. fig. 2 81 mm 24mm 0.30 B. patula MAZE NOT pl. 5, fig. 5 73mm 33mm 0.45 B. patula ScHNElD pl. 7, fig. 5 74mm 36mm 0.48 present specimen 93. 5mm 38.8mm 0.42

and fiat or broadly concave. The suture B. morti MAZENOT, 1939 from the· lines are indiscernible. Upper Tithonian of Vogue, also resem­ These characteristics are comparable bles this specimen but differs from the with those of Berriasella patula SCHNEID, latter in more crowded ribs which bi­ 1915 from the Upper Tithonian of Neu­ furcate at somewhat higher point than burg and the Berriasian of Saint-Julien­ the middle of the flank, and absence of en Bochaine, Berrias and Bournet, the conspicuous elevation at the starting· France. But there are some differences point of the ribs. between the specimens concerned with Occurrence.-Locality YB101, at Ya­ the SCHNEID's original description (1915) mabu, Honjo Village, Minamiamabe and the photograph of the same species County, Oita Prefecture. Location: by MAZENOT (1939). That is, the MA­ Long. 131 "38'33"8E, Lat. 32.55'5"7N .. ZENOT's specimen has more rapidly en­ The specimen occurred in the black larging whorls and a less wide umbilicus shale inserted in the basal conglomerate. than the SCHNEID's. The Yamabu spe­ of the Yamabu formation. cimen is rather closer to the MAZENOT's in this respect, if the effect of deforma­ tion is taken into consideration. The present specimen also resembles B. priversensis (PICTET, 1867) and B. callistoides (8EHRENDSEN, 1891), but dif­ fers from them in the following chara­ cters. (1) The umbilicus of the present specimen as shown in Table 1, is fairly wide in proportion to diameter. (2) The ribs are flexuous in the present species but nearly straight in B. priversensis. (3) The ribs bifurcate near the middle of the flank in the present species but at much higher point up about a fourth of the flank in B. priversensis. Text-fig. 3. Locality map. 609. Ammonites from Yamabu formation 467

~Family Oosterellidae BREISTROFFER, 1940 is associated with Berriasella aff. patula.

Genus Pseudoosterella SPATH, 1924 On the geological age of the Pseudoosterella sp. Yam~bu formation

PI. 56, Figs. 2-4 The Yamabu formation has been con­ sidered as the non-marine deposits. be­ Material.-Three specimens from toe. cause of the occurrence of Protocyprina YB101. the basal member of the Yamabu cf. naumanni in its middle member. It has formation, Oita Prefecture, Kyushu, been set to the lowest part of the Lower ·(coli. M. NODA) are concerned with~ the Cretaceous in Japan, probable correlative ·description. of the Wealden of England (see remarks Descriptive remarks.-Although the in the introduction). The discovery of ·specimens are all· very incomplete frag­ ammonites from the basal member. ments. the original. characteristics are however, indicates that the early stage preserved in the convexity of whorl and of sedimentation of the Yamabu form­ -ornamentation. The specimen of NODA ation is probably of marine origin and collection JG. I-11003 is an adult indivi­ the occurrence of Berriase/la probably ·dual about one sixth of whorl with 19.3 indicates Berriasian age. According mm in height and other two specimens to MAZENOT (1939), Berriasella patula ·(JG. I-11004 and JG. I-11005) are both im~ occurs not only in the Lower Tithonian mature, of which the one, JG. I-11004, is of Nefiburg (SCHNEID regarded the same about one fourth of whorl and the other locality as the Upper Tithonian) but also is about one sixth . of whorl.. from the Berriasian of Saint-Julien-en· The umbilical wall is fairly steep, Bohain, Berrias and Bournet of Sou­ forming a round shoulder with the flank. theast France. Although there are some The whorl is compressed probably in~ questions about the range of that species ·Creasing slowly in height. The flank is described in MAZENOT, the occurrence 'broadly convex and forms a subangular of· Berriasella aff. patula from the Ya­ shoulder at the ventrolateral part. There mabu formation. at least, indicates the. is a finely serrate keel at the mid-venter. geological age corresponding to the The simple ribs bigin at the umbilical Berriasian. margin, running rectiradially on the SATO (1958, 1961) previously described flank. and sharply projected at the ven­ some Berriasian and Valanginian am­ trolateral shoulder. They are as wide as monites from the Lower Cretaceous de­ the concave interspaces, and regular in posits which comformably overlie the ~strength and distance. The rib density Jurassic strata in the Kitakami and the ranges from 8 to 9 per one sixth of Abukuma massifs. They are Berriasella whorl and 10 per one fourth of whorl. akiyamae SATO, Thurmanniceras isoku­ In these features, this species is re­ sensis KOBAYASHI and FUKADA from the ferable to Pseudoosterella. The specific Isokusa formation, Berriasella sp. ex gr. identification remains uncertain because B. berthei TOUCAS, Spiticeras (Spiticeras) -of the incomplete materials. cf. binodige1· UHLIG. Olcostephanus sp. Occurrence.-Loca!ity, YB10l, at Ya· and Kilianella juv. sp. from the Naga· mabu, Honjo Village, Minamiamabe saki formation and Parakilianella uma­ ·County, Oita Prefecture. The specimen zawensis SATO, Thurmanniceras sp. and 468 Masayuki NODA Berriasella sp. from the Koyamada for­ ,,__.,. BERIIAS mation. Based on the assemblage of ...... L-o< Up;tK """" - AHCllOCERATI DAE species he considered that there are - -- -,-- some marine deposits in Japan which Vb'go.toaplncllna• lo.. Dctsofi'Qnilina• ~ are contemporaneous with or somewhat Vigcllll"'l• ~ A•ph:toc•rotll'lato :l! ~ I z older than the non-marine Ryoseki Series SiMOCII't'GtlM• 'n I '::! and that they probably correspond to CICIIII•d•l•nae Gorni~trturallnae '~ the Berriasian in the international scale, follllna. i ------r-- ~------~ regardless the occurrence of Olcoste­ Splllootollnat I~ ." Olc.otl~tphclnlna~e I Polyptrc:hllina• f !il phanus sp., the genus from the Upper 1-, Valanginian of Europe, South Africa, &.rrloteiiJMe i= I Madagascar, Pakistan. Mexico and Peru, HlmQICififlncJit '~ NltOcmlllfi.Git ir being later than Berriasella in these _T_ f---- 1'1 I OOSlfAEUIDAE areas. r- ·t Hoploc:tralldcle The basal member of the Yamabu :: ?:;:.: '"""""""' Clpp•IIICia. ·m formation, therefore, may be correlated I DESWOC£JL\TAC£AE I to those formations of Northeast Japan. Text-fig. 4. Range chart of ammonites at But there is a question that Pseudoo­ Jr/Cr boundary (after WIEDMANN, 1967). sterella sp. which should normally chara­ cterize the Upper Valanginian occurs further problem. from the same bed with B. aft. patula. This is similar to the case of the Naga­ saki formation. The international problem of the At present, it is uncertain whether or Jr/Cr b9undary not the association of the Berriasian and the Valanginian species is the excep­ The geological age of the Yamabu•. tional case found only in Japan. The formation has some relations to the in­ solution of this problem is important ternational problem of the Jr/Cr boun­ for more precise world-wide correlation. dary, because there is no agreed opinion,. The upper part of the middle member at present, for the border-line by means­ of the Yamabu formation is characteri­ of paleontological evidence and sedi­ zed by brackish water mollusks listed mentary structure. The geological divi­ in p. 464, common in various localities sions of the Upper Jurassic and the Lower of the Ryoseki Series in Japan. The Cretaceous are interpreted differently by similar examples in the Lower Creta­ various researchers as shown in Table 2 .. cous, i. e. the marine deposits under the The PERGAMENT's division is based. non-marine ones are also found in the Um­ on paleontological evidence, notwith-· inoura formation, Kumamoto Prefecture standing it differs from that of HAUG, (TAMURA, 1960b; SATO, 1961 and MATSU­ TOUCAS and WIEDMANN. He has re­ MOTO. 1962) and the Kawaguchi forma­ garded the zones of Subthurmannia bois­ tion, Kumamoto Pref. (MATSUMOTO, per­ sieri and Riasanites rjasanensis as the· sonal information, 1971). However, due lowest limit of the Cretaceous. under the· to the insufficient data, more precise name of Valanginian and he excluded correlation of the Yamabu formation the name of Berriasian in his division. and those of Kumamoto Pref. is left In other words. the zones characterized. unsolved. I must investigate it as a by BerriastJlla spp. are included into the: 609. Ammonites from Yamabu formation 469

Table 2. Geological division of the Upper Jurassic and the Lower Cretaceous .(thickness not to scale).

COOUAHD 1871 ., toUAH TOUC4S YABE PICTET 1910 CA.SE't BREIST~OFF. 4Ll£N PERGJ\MENT HAJJ() 1890 1917 SATO 1887 MAZENOT 1953 1964 1955 1965 1898 HIS& Hl39 WIEDM.AHN MATSUMOtO BOSNARDO 196~ 19U HI 55

ValanQinlcn Valc.nglnla.n Vutonginlan Volanginl.ul) Valanginian Valanglnlo.n $.,:Sit, 1-:slt. Wealdtn ~ S..Sfr, W•1ld•n Ryosei:l tl (CO ...!.AIIE a. (URltl'fl ~LLJi"SllU u Valonglnla.n c Kochic.n Yalo.n.glnian i Ryczanlan B.rrlaslon Upo"P- .. (WAfSUICI.Oit4ll Bll'rriaskln IIOGC!l.OIISIUI J..s.lr. Purb.cldan Duris ton• "' 111S) (lfOIOIUtf Ceooulllelf'/11 un1 (ODOl 115-~· Bds hriaslan Infra UpJM-r Yatanglnian Tithonian c ~ ll!,t,lll ttl ~I Tithonian Lowe-r Purbtdl: (Ard•sdan) Upp•r crouc.u t.lto..J Volglan (Porllandlari (Lulworth hds) :s.l. J,.J.Ir Tlthor.ion Tithonian Volglan (Volglan) (~III.IWI 'NlJ Portlandian Portla.ndkln r.tori~ jlr~'fSOI (tOICQtN"'-ItM. Tithonian (TLthontan) (TIIhonoan) Low., Totlnosu (!M'I'nTU! I Lowu Tlthonlan (twi&Oilltol Porlla.ndian Volglan (Donuboan) s. ~tr. s. sl.~ (IIOUJUISO.I

I Volgian. While, TOUCAS and others TSUMOTO (1943, 1954) proposed the Ko­ included the two zones in the Tithonian, chian ( =Kotian) for the lower series of hence the top of the Tithonian in their the tripartite Lower Cretaceous in Japan. definition corresponds to the border-line He attempted to define it by the avail­ of 'Berriasian and Valanginian of MA­ able marine fossils (trigonians), but our ZENOT. BUSNARDO, BREISTOROFFER and knowledge of the taxonomy and strati­ KILIAN. graphic .occurrence of the trigonians at WIEDMANN's division is ba&ed on a that time was not. sufficient. SATO's range chart of ammonites (Text-fig. 4). (1958, 1961) discovery of ammonites from For the Cretaceous of Japan the di­ the lsokusa and the formations vision proposed by YABE (1927) had been has made it possible to correlate the used. He intended to use the term Ryo­ Japanese scale with the international seki Series as a time-stratigraphic unit one. From the fact that the ammonites but gave a definition that it is a series, of Upper Valanginian aspect are asso­ mostly of fresh and brackish water ciated with Berriasian species, it is deposits, which is the product of a time better to ascribe these formations to the of marine regresssion and that it is ap­ Lower Cretaceous Berriasian as defined proximately correlated with the Weal­ by MAZENOT and BUSNARDO rather than den, the time of transition Jrom the to the Tithonian. The facts newly ob­ Jurassic Period to the Cretaceous in the served in the Yamabu formation also European standard. Actually, what had support this. been recognized as the Ryoseki in va­ The apparent intermingling of Valan­ rious areas of Japan was rather a facies· ginian aspect ammonites with the un­ stratigraphic unit and accordingly MA- doubtedly Berriassian ones at several 609. Ammonites from Yamabu formation 469

Table 2. Geological division of the Upper Jurassic and the Lower Cretaceous .(thickness not to scale).

COQUA!IO 1111 --~ KILIAN TOUC45 YABE PfCTET tltO J CASEY 8R£1 ST AOFP'. AU£N PEAG.UCNT HAUG 1110 1U7 SATO 18117 MAZENOT 1963 1964 19115 1885 1188 1956 IUQ WIEDioiANN MA~SUI«lTO BUSIWIOO 1987, liU 1865

Vq1a.noinian R)'oukl (r..a£ 1tvt Volcnglnlon 1----+----1

(OU~ UHI

Tithonian Low., Purbtdl (ArdneiCW'I) Upp~r Vololon (for!~~ft lfOUC.AS ltM.I--i..-~---1 (l.utwo.rth Beds ) J,lfr nthonlon lllhonlon Yotgfan ,..... "".''' (Vo\gian) Porllcnd\Qn Potttan41Gn (O'o.o.rrli1GI r.torl­ l'tOICR'MUIIMI Tithonian 1---~ (TIIhonlonJ (TIIhonlon) Low« Torlnos-u (~11UI Lowor Tlth!M'an ti'WU.OI. '"'' Portlandian Yal;lon (Oorublun) .s.~lr, .l.Jir. IIIOLUU1tOII ' I I Volgian. While, TOUGAS and others TSUMOTO (1943, 1954) proposed the Ko· ~ included the two zones in the Tithonian, chian ( =Kotian) for the lower series of f· hence the top of the Tithonian in their the tripartite Lower Cretaceous in Japan. definition corresponds to the border-line He attempted to define it by the avail· of 'Berriasian and Valanginian of MA· able marine fossils (trigonians), but our ' ZENOT. BUSNARDO, BREISTOROFFER and knowledge of the taxonomy and strati· KILIAN. graphic .occurrence of the trigonians at WIEDMANN's division is ba~ed on a that time was not_ sufficient. SA TO's range chart of ammonites (Text-fig. 4). (1958, 1961) discovery of ammonites from For the Cretaceous of Japan the di­ the Isokusa and the Nagasaki formations vision proposed by Y ABE (1927) had been has made it possible to correlate the used. He intended to use the term Ryo­ Japanese scale with the international seki Series as a time-stratigraphic unit one. From the fact that the ammonites but gave a definition that it is a series, of Upper Valanginian aspect are asso­ mostly of fresh and brackish water ciated with Berriasian species, it is deposits, which is the product of a time better to ascribe these formations to the of marine regresssion and that it is ap· Lower Cretaceous Berriasian as defined proximately correlated with the Weal· by MAZENOT and BUSNARDO rather than den, the time of transition from the to the Tithonian. The facts newly ob­ Jurassic Period to the Cretaceous in the served in the Yamabu formation als() European standard. Actually, what had support this. been recognized as the Ryoseki in va­ The apparent intermingling of Valan­ rious areas of Japan was rather a facies­ ginian aspect ammonites with the un· stratigraphic unit and accordingly MA· doubtedly Berriassian ones -at several 470 Masayuki NODA

localities in Japan is unusual and left MATSU~tOTO, T. and others, 1967. Historical to be solved in the future. geology. Asakura, Japan. (in Japanese). MAZE NOT, G., 1939. Les Palaeohoplitidae To sum up, the discovery of new data Tithoniques et Berriasiens du Sud-Est in Japan as described in this paper, not de Ia Fr~nce. Mem. Soc. Geol. France, only enables us to attempt the world· N•. 41, pp. 1-303, pls. 1-40. wide correlation but also may give some MOORE, R. C. (ed.), 1957. Treatise on. in· contribution to the international problem vertebrate paleontology, 4. Geol. of the Jr/Cr boundary. Soc. America, Univ. Kansas Press. PERGAMENT, M.A., 1965. Zonal scale of the Cretaceous in the North-east· of Asia and its correlation with the American and Reference cited · European scales. Report to the II I sym­ CASEY, R., 1963. The dawn of the Creta· posium on the development of oil and gas ceous Period in Britain. SE Union Sci. resources in ECAFE countries, Tokyo. Soc. Bull. no. 117, pp. 1-15. 1965. Fuj(l, K., 1954. Stratigraphy and geological SATO, T., 1958. Presence du Berriasien dans structure of the Usuki area, Oita Pre­ Ia stratigraphie du plateau de Kitakami. fecture, Kyushu (1). j~ur. Geol. Soc. Boll. Soc. Geol. France, ser. 6, vol. 8, pp. japan, vol. 60, pp. 413-427. (in Japanese 585-599, pl. 28. with English abstract). --, 1961. La limit Jurassico-Cretacee dans KAMBE, N. and TERAOKA, Y., 1968. Geology Ia stratigraphie Japonaise. japan. jour. of the Usuki district. Quad. Ser. 1: 50000, Geol. Geogr., vol. 32, pp. 533-541, pl. 12. (14), no. 88, Geol. Surv. japan. --, 1961. Faune Berriasienne et Tithonique (in Japanese). Superieure Nouvellent Decouverte au MATSUMOTO, T., 1943. Fundamentals in the Japon. Ibid., pp. 543-551, pl. 13. Cretaceous stratigraphy of Japan, Parts ScHNEID, T., 1915. Die Ammoniten fauna II & III. Mem. Fac. Sci., Kyushu Univ., der obertithonischen Kalke von Neuburg Ser. D (Geol.), vol. 2, pp. 97-237. A. D. Geol. Palaeqnt. Abh., N. F. Bd. 13, -- (ed.), 1954. The Cretaceous System in pp. 305-413, pis. 17-29. the japanese Islands. Japan Sci. Promor­ TAMURA. M .. 1960a. A stratigraphic study tion Soc., Tokyo. (for 1953). of the Torinosu group and its relatives. -- and KANMERA, K., 1952. The lower Mem. Fac. Educ. Kumamoto Univ., vol. 8, valley of the Kuma. Guidebook of the Supplement, pp. 1-40. (in Japanese). geological excursions. Dep. Geol., Fac. --, 1960b. Stratigraphical study on the Sa­ Sci., Kyush1t Univ. (in Japanese). kamoto group in Kyushu. four. Geol. -- and --, 1964. Hinagu, Kagoshima-49. Soc. japan, vol. 66, pp. 371-383. (in Japa­ Explanatory text of the geological map of nese with English abstract). japan. 1: 50000, Geol. Surv. japan. (in TERAOI

Explanation of Plate 56 Fig. la, b. Berriasella sp. aff. B. patula ScHNEID, natural size. Loc. YBlOl, Shinkai, Honjo Village, Minamiamabe County, Oita Prefecture. (Coil. NooA, 1970) Figs. 2-4. Pseudoosterella sp. 2. natural size, 3, 4. x 1.5. Loc. YBlOl. (Coil. Noo·A, 1971) J<: NODA: A1nmonites j?·om Yamabtt fonnation Plate 56 609. Ammonites from Yamabu formation 471

'WIEDMANN, J., 1968. Das Problem strati- Y ABE, H., 1927. Cretaceous stratigraphy of graphischer Grenzziehung und die Jura/ the Japanese Islands Sci. Rep., Tohoku Kreide-Grenze. E'cologae Geologicae Hel­ Imp. Unilj., 2nd ser., vol. 11, pp. 27-100, vetiae, vol. 6l, pp. 321-386. pis. 3-9. '

Abu kuma ~iiJ ~AI! Nagasaki -Bi: Uf Honjo [!r. Ryoseki ~ :fi lsokusa *Til 1ii Shinkai tiT 1m Kawaguchi )II IJ Uminoura i1il I iili Kitakami ;f~ ..!=. Yamabu LlJ all Koyamada ,j, LlJ 1B Yatsushiro i\ tt Minamiamabe iliiliJllll Trans. Proc. Palaeont, Soc. Japan, N. S., No. 88, pp. 472-484, pl. 57, December 20, 1972

610. SOME FOSSIL PTEROPODA FROM MIYAZAKI AND OKINAWA PREFECTURES, SOUTHWEST JAPAN*

HIROSHI NODA

Institute of Geology and Paleontology, Faculty of Science, Tohoku University

i!Yi~IFJ *i

According to the authorities just men­ Introduction tioned the Pteropoda are cosmopolitan. in distribution. There are only few studies on the Fortunately the writer collected some­ fossil pteropods in and outside Japan, Miocene and Pliocene species from Mi­ probably because of their rare occur­ yazaki and Okinawa Prefectures. Sou­ rence due to thin and fragile shell. thwest Japan. These records are im­ Although the Pteropoda range from portant for regional correlation and· the Paleogene (HARRIS, 1894; CURRY, reconstruction of the paleo-environment 1965) to Recent, in Japan the oldest of the deposits. record is a Miocene species. Cavolina raritatis NOMURA and ZINBO, 1935, from the Yanagawa Shell Bed in and several Pleistocene spe­ Acknowledgments cies from the Semata Shell Bed by Y A­ MAKAWA and ISHIKAWA (1912a, b), No­ The writer wishes to express his deep· MURA and HATAI (1936) and KOBAYASHI gratitude to Professor Kotora HATAI of (1956), whereas there are no records the Institute of Geology and Paleon­ from the Pliocene in Japan. The syste­ tology, Faculty of Science. Tohoku Uni·· matic classification of the shell bearing versity, for his continuous encourage­ pteropods has been based mainly on the ment and supervision during the present soft parts (RANG, 1825 ; PELSENEER, study. Acknowledgments are due to· 1888b : TESCH, 1904; WENZ, 1960; TAYLOR Associate Professor Tamio KOTAKA,. and SOHL, 1962; SPOEL, 1967; HYMAN, Tohoku University; Associate Prof. To­ 1967; BOL TOBSKOY, 1971 and others). mohide NO!-IARA, Ryukyu University;: Drs. Taijiro KONISHI, Taisuke SUZUKI and • Received June 20, 1972; read June 3, Kazuo MORI, Geological Survey of Japan. 1972, at Utsunomiya. for their kind information and sugges-· 472 610. Fossil Pteropoda, Miyazaki and Okinawa 47~

tions on the stratigraphy of Okinawa-ji· been revised by KONISHI et al. (1970. rna. Thanks are also due to the members 1971) and FUKUDA et al. (1970). Accord· of the Research Group of Cenozoic Mol· ing to them, the stratigraphical sequence !uses (Profs. Koichiro MASUDA, Miyagi of the sou~h to central parts of Okina· University, Kotora HATAI, Tohoku Uni· wa-jima is: versity, Tokio SHIKAMA, Uni· versity, Shozo HAYASAKA, Kagoshima Ryukyu Group Naha Limestone University, Associate Professors Tamio KOTAKA, Tohoku University, Takehiko Chinen Sand \Kunigami Gravel lWAI, Hirosaki Unive·rsity, Taisuke TA· Shinzato Formation KAYASU, Akita: University, Saburo KAN· Shimajiri Group Upper NO, Tokyo University of Education, Ki· Yonabaru Formation Middle yotaka CHINZEI. Tokyo University, Hiro· {Lower yuki OTSUKA, Kagoshima University, Naha Formation Oroku Sand Junji ITOIGAWA, University, Sakae OIIARA, Chiba University, Drs. Base unexposed ----- Yasuhide IWASAKI, Tokyo University, Kazuo OKAMOTO, University, The lower part of the Yonabaru For· Messrs. Kimihiko OKI, Kagoshima Uni· mation (FUKUDA et al.. 1969, 1970) is versity, and Kenshiro OGASAHARA, To· nearly the same as the lower part of hoku University) for their kind informa· the Yonabaru Clay of MACNEIL (1960)­ tion on the biostratigraphy of the Miya· in the western part of island. The for· zaki Basin and discussions in the field. mation dips eastwards with low dips Thanks are expressed to Mr. Michio and is composed mainly of massive dark KATO, Tohoku University for picking gray to brownish gray clayey siltstone up the fossil pteropods under the mic· sometimes intercalated with fine-grained roscope and to Mr. Kimiji KUMAGAI for tuffaceous sandstone layers which yiel· the photographic work. ded Cavolinia telemus LINNAEUS, Cleo­ dora okinawana n. sp., and Creseis aci­ cula RANG. The first mentioned species. Stratigraphic occurrence of is rather common but the other two are fossil Pteropoda rather rare in occurrence. These ptero­ pods are sometimes associated with The fossil pteropods treated in this molluscs, foraminifers and fish otoliths. article were collected from the two iso· The Shinzato Formation, first pro­ lated areas (Miyazaki and Okinawa) as posed by MACNEIL (1960) for the white described below. acidic tuffs intercalated in the massive A. Okinawa District. clayey siltstone around Shinzato, Sashi· The fossil pteropods were collected ki·son in the eastern part of island. from some localities in the area of dis· under the name of the Shinzato Tuff tribution of the Shinzato and Yonabaru Member, also yielded fossil pteropods. formations (Text-fig. 2) of the Shimajiri The formation was revised by FUKUDA Group (HANZAWA, 1935). The classifica· et al. (1970), and stated to be composed tion of the Neogene stratigraphy of the mainly of massive dark gray to brow­ Okinawa-jima proposed by HANZAWA nish gray siltstone intercalated with (1935) and MACNEIL (1960). has recently very fine, layered acidic tuffs and partly- 474 Hiroshi NODA sandy siltstone. The formation is dis­ Yonabaru Formation (FUKUDA et al., tributed mainly on the eastern side of 1970). the island and in a small isolated area B. Miyazaki District.

As already stated by PELSENEER (1888b) the Pteropoda differ from the Cephalo­ poda, Tentaculites, Conularia and Hyoli­ thes though the external sculptures and morphology are somewhat similar. PEL­ SENEER {1888b), TESCH (1904), CHEN and B~ (1964), HYMAN (1967) and others have made a historical review of the Ptero­ poda and further reworks are not ne­ cessary. At this place, remarks are given on the first records from the Miyazaki and Okinawa Prefectures, sou­ thwest Japan because of their being first from the Japanese Pliocene and· the second from the Miocene. As known at Text-fig. 1. Map showing fossil localities. present the earliest record of the shell- 610. Fossil Pteropoda, Miyazaki and Okinawa 475 bearing Pteropoda is from the Eocene above are known to be distributed in of England (HARRIS, 1894; CURRY, 1965). the subtropical to temperate seas as The Koyu Formation in Miyazaki Pre­ stated by PELSENEER (1888a, b), TESCH fecture is Middle Miocene in age accord­ (1904), PILSBRY (1922) TOKIOKA (1955, ing to SHUTO (1961), who studied the 1960 in OKADA et al.), CHEN and BE marine molluscan fossils from the forma­ (1964) and others. From the occurrence tion. SHUTO (1961) subdivided the forma­ of the pteropod species the Shinzato tion into three members, among which Formation is inferred to have been de­ the lowest or the Kawabaru Member posited under an open sea condition but yielded Cleodora hataii, n. sp., a species whether of rather deep water analogous resembling Clio pulcherrina reported to the depth of pteropod ooze may be from the Cessole Formation (Langhian). questionable. The correlation of se­ Italy by ROBBA (1971). This Middle parated -geological formations by the Miocene record is the oldest in the pteropod fossils is at present difficult Japanese Neogene. owing to the few occurrences, but should The Shimajiri Group originally des­ their records be increased their value in cribed by HANZAWA (1935) was revised biostratigraphic correlation and paleo­ by MACNEIL in 1960. Recently, FUKUDA biogeographic analysis may become im­ et al. (1969, 70) published a geological map portant. of the Shimajiri Group distributed in the central southern part of the Okinawa­ ~ystematic description jima from the view point of the natural gas resources. The Yonabaru Forma­ Family Cavolinidae FISCHER, 1833 tion. the middle part of the group. ac­ cording to FUKUDA et al. (1969, 1970) Genus Cleodora PERON and underlies the Shinzato Formation with LESUEUR, 1810 conformity and yielded Amussiopecten praesignis, Hawaiarca uwaensis, Limop­ Cleodora hataii NonA. n. sp. sis tajimae, L. tokaiensis, Glycymeris Pl. 57, figs. 9-11 pilsbryi and many other molluscan spe­ cies besides Cavolinia telemus which was Type locality :-Road side cliff, east originally described from the Mediter­ of the tunnel northeast of Yamaji, Saito ranean Sea and also reported from the City, , Kawabaru Miocene Bowden and Pliocene Man­ Member of the Koyu Formation, Mio­ chioneal beds of Jamaica and Jacnel cene. IGPS coli. cat. no. 92579. Pliocene Formation in Tahiti. The geo­ Description .·-Shell very thin, rather logical age of the Yonabaru Formation large for the genus, triangular, some­ may be Upper Miocene from the macro­ what inequilateral in form. Ventral fossils cited above. The Shinzato For­ side convex with a single broad, slightly mation yielded seven pteropod species elevated median ridge with concentric among which three are new species, and equi-spaced growth bands. Dorsal side Diacrina bisulcata, Cavolinia telemus and slightly convex with medium rib near Creseis acicula have been recorded from apex, branching into three longitudinal the Middle Miocene Bowden of Jamaica, ribs ; middle rib narrower than the Santa Rosa. Vera Cruz, Mexico and others with concentric growth lines. Virginia. All of the species mentioned Both sides of radial ridge on both sides ~ m -.::1

~"'i 0 ~ ""· ~ 1::::1 ( 8) ( c ) :to buni ~ ~') km Upper } li?77:1;1 Naha Limestone 8... . Middle Yonabaru Key Bed Formation • Lower p: .::q Chinen Sand

E:"SS'-';1 Naha Formation ~~~~ Shinzato E !'ormation '\!EJ : Fossil locality

Text-fig. 2. Fossil localities of three separated areas (A; Naha district, B; Gushikami district, C; Chinen Peninsula district). Geological map from FUKUDA et a/., 1970. 610. Fossil Pteropoda, Miyazaki and Okinawa 477 Dimension:- Length 19.0 mm Width 12.0mm Height 4.2 mm (92579) Length 20.6 mm Width 17.9mm Height -mm (92580) I of shell become very fiat. Apex very of the Shinzato Formation, south cliff small, pointed. Aperture wide, narrow, of Shinzato, Sashiki-son, Okinawa Prefec­ middle part rather wide, both ends of ture, Shinzato Formation, Pliocene. aperture narrowly fiat. IGPS colt. cat. no. 92581. Comparison and affinities:-The pre­ Description :-Shell very fragile. moder· sent species resembles Cleodora bow­ ate in size, and of lozenge-form. Dorsal denensis COLLINS originally described side of shell ·sculptured with medium from the Middle Miocene of Jamaica rather narrowly elevated, longitudinal (COLLINS, 1934) in having radial ridge rib which becomes divided into three on the shell surface and in shell form similar longitudinal ribs. Interstices but differs ~rom the latter by the small flat, wider than longitudinal ribs width for shell· length, distinct ·radial sculptured with very fine concentric ridges from near a'pex to aperture and growth striations. Aperture sharply rather distinct concentric growth lines triangular in form. Lateral sides of on the external surface. Cleodora balan­ shell . very. thin rather fiat. Lateral tium RANG illustrated by DALL (1878) corner also sharp. Apical end prominent unqer the name of Balantinum recurvum and spinose. CHILDREN from the Recent sea of Aus­ Dimension :-(Holotype) Height 12.8 tralia resembles the present species in mm, width 6.0 mm. the undulated ridged growth lines, and Comparison and affinities:-The species three longitudinal ridges but the former is characterized by its rather distinct differs from the latter in the U -shaped three longitudinal ridges on the aper­ shell except for the sharp apical spine. tural margin and its lozenge-form. The latter species has a V-shaped tri­ Cleodora bowdenensis COLLINS, 1934 gonal shell form. Clio pulcherrina originally described from the Middle (MAYER) described recently by ROBBA Miocene Lower Zone at Bowden in (1971) from the Langhian Cessole For­ Jamaica by COLLINS (1934), resembles the mation in Italy resembles the present present new species but the former new species in the shell form and un­ differs from the latter in having straight, dulated growth lines crossing the longi­ very prominent spinose apical end, and tudinal ridges bu~ the former differs three distinct radial ridges on the dorsal from the latter by having five longi­ side of the shell. The latter species is tudinal ridges. characterized by its wider apertural Remarks:-At present, the present length, rather slight lateral elevation of species is the earliest record of Ptero­ medial ridge and recurved apex. Cleodora poda in Japan. pyramidata LINNAEUS (fide COLLINS, 1934, p. 201) is another species allied with the present one but differs in having rather Cleodora okinawana NODA, n. sp. inflated shell and distinct median ridge. Pl. 57, figs. 6, 12, 13 Cleodora hataii, n. sp. from the Miocene Koyu Formation in Miyazaki Prefecture Type locality :-Near the type locality resembles the present new species but 478 Hiroshi NODA

differs in the large size, inflated shell, Col .. Vcl. 3, p. 171, fig. 642. elevated medial part with three distinct Hyalocylix striata (RANG) : CHEr>: and BE~ longitudinal ridges on the dorsal side 1964, Bull. Mar. Sci. Gulf; Carrib., vol. and distinct growth lines. Cleodora 14, no. 2, p. 193, fig. 3g. australis o'ORBIGNY illustrated by DALL Hyalocylis striata (RANG): KEEl\, 1971, p .. 805, fig. 2285. (1878) resembles the present species in the pyramidal shell form but the former Description :-Shell very thin, fragile,. differs from the latter in having no small, conical tube, nearly straight but interstitial folds between the central slightly curved at end of apex. Aper­ costae. , ture round in cross section. Apex Locality and formation :-Loc. no. 13, narrow and rather spinose. Shell surface prefectural road (Route 44) side cliff sculptured with equispaced distinct near Kamima, Haebaru-son, Okinawa annulations. Prefecture, Middle Part of Yonabaru Dimension :-Length 6.2mm and width Formation, Miocene, IGPS col!. cat. no. 2.5mm. 92582; Loc. no. 15, south of Shinzato, Comparison and affinities:-The pre­ Sashiki-son, , Shin­ sent species is characterized by its zato Formation, Pliocene. IGPS coil. slightly curved apex and distinct external cat. nos. 92581, 92582 ; Loc. no. 12-06, sea annulations. This species resembles side cliff, west of Tobaru, Yonagusuku­ Hyalocylix haitensis COLLINS, 1934 in son, Okinawa Prefecture, Shinzato having similar external annulations but Formation, Pliocene, IGPS coil. cat. no. the former has finer and denser crenu­ 92596. Jations on the external surface and the apical end is more spinose, compared Genus Hyalocylix H. FOLKSKAL, 1875 with the latter. Remarks :-The present species, known Hyalocylix striata (RANG, 1828) as a Recent form of wide distribution in the warm to subtropical sea realm, Pl. 57, figs. 7, 8 has no previous record as fossil. The Creseis striata RANG, 1828. Ann. Sci. Nat., present discovery extends the geological Tom. 13, p. 315, pl. 17, fig. 7 (fide LA­ range of this species back to the Pliocene. MARCK, 1836). Locality and formation :-Loc. no. 12- Cleodora striata RANG: LAMARCK, 1836, Hist. 06, sea side cliff, west of Tobaru, Nat. Animaux, Verteb .. Tom. 7, p. 433. Miyagusuku-shima, Yonagusuku-son, Creseis striata RANG: DALL, 1878, pl. 5, figs. Okinawa Prefecture, Shinzato Formation, 30a, b. (non vidi). Pliocene, IGPS coil. cat. no. 92583. Clio (Hyalocylix) striata (RANG) : PELSENEER, 1888a, Rept. Sci., Voy. H.M.S. ChalleHgel", vol. 23, p. 54, pl. 2, fig. 3. Clio (Hyalocy/ix) striata (RANG) :TESCH, 1904, Genus Diacria GRAY, 1847 Siboga Exped. Monogr., vol. 52, p. 27, 28, pl. 1, figs. 16-17. Diacria bisulcata GABB, 1873 Hyalocy/ix striata (RANG) : TOK!OKA, 1955, Pl. 57, fig. lB. Publ. Seto Mar. Bioi. Lab., vol. 5, no. 1, p. 65, pl. 9, fig. 21. Diac1·ia bisnlcata GABB, 1873, Trans. Amer. Hyalocylix striata (RANG) : TOK!OKA in OKA· Phi los. Soc., n.s., vol. 15, p. 200. (non DA et al .. 1960, Encyclop. Zoo/. lllust. vidi) 610. Fossil Pteropoda, Miyazaki and Okinawa 479

CavoliiiQ (Diacria) bisulcata (GABB) : DALL, by WOODRING (1928, pl. 1, figs. 16-17) 1892, Trans. Wagner lnsl. Philad .. vol. and COLLINS (1934, pl. 10, figs. 1-3) re· 3, pt. 2, p. 430. (non vidi). semble Diacrina trispinosa s. s. by pos· Diacria bisulcata GABB: PILSBRY, 1922, Proc. sessing five longitudinal' ridges on the Acad. Nat. Sci., Phila, vol. 73, p. 309, dorsal surface. For this reason the fig. 4. species group should be re-examined. Diacria bisulcata GABB: WooDRING, 1928, Carnegie Inst. Washington, no. 385, p. 116, Locality and formation :-Loc. no. 12- pl. 1, figs. 14-17. 06, sea side cliff, west of Tobaru, Diacrina bisulcata (GABB) : CoLLINS, 1934, Miyagusuku-shima, Yonagusuku-son. john Hopkins Univ. Geol., no. 11, p. Okinawa Prefecture, Shinzato Formation, 197-200, pl. 10, figs. 1-3, pl. 9, figs. 13- Pliocene, IGPS coil. cat. no. 92584. 20. Genus Cavolinia ABILDGAAD, 1791 Description :-Shell small. equilateral, somewhat lozenge in form. Aperture Cavolinia telemus LINNAEUS, 1758 smoothly rounded, both sides broadly angulated, its external ends rather sharp, Pl. 57, figs. 14-17, 19-20, 22-26, 29-31 not spinose, apical area narrow and long. Cavolinia telemus LINNAEUS, 1758, Syst. Nat. External sculpture with slightly elevated ed. 10, p. 365, (fide CoLLINS, 1934). longitudinal ridge (medial) and secondary Cavolina telemus (LINN~) : WooDRING, 1928, ridges, outer sides become fiat, without Carnegie /nst. Washington, no. 385, p. 10, striations beside growth lines. Ventral p. 113, pl. 1, figs. 6-7. side unexamined at present. Height Cavolina telemus (LINNE) : CoLLINS, 1934, 4.8 mm and width 3.8 mm. john Hopkins Univ., Stud. Geol., no. 11, Comparison and affinities :-The pre· p. 188-190, pl. 8, figs. 4-7. sent species, only one shell showing the Cavolina (Cavolina) telemus (LINNE) : WENZ, dorsal side, resembles Diacria mbaensis 1959, p. 52, fig. 174. Cavolinia telemus (LINNE) : KuRODA, HABE LADD originally described by LADD (1934) and OYAMA in Biol. Lab. Imp. Household, from the Miocene Suva Formation, Fiji p. 300, pl. 64, fig. 12. Island in the external shell form but the latter is characterized by five subequal Description :-Shell rather large, rounded longitudinal ribs on the dorsal fragile, inequivalve and equilateral. surface, a character different from the Ventral valve much swollen, particularly present species. Diacrina trispinosa on apertural margin. External surface (BLAINV!LLE) illustrated by KURODA, rather smooth, with very fine concentric HABE and OYAMA (1971) from Sagami striations conspicuous near aperture and Bay, (1971 in Bioi. slightly bent backward medially. Dorsal Lab. Imp. Household) resembles the pre· valve rather flat. external surface sculp­ sent species by having rounded aperture, tured with three distinct elevated long­ sharply spinose ends and longitudinal itudinal ribs; one of them distinct and ridges on the dorsal surface but the situated on middle part of shell, other present species differs from the former two slightly weaker. Apertural margin by its low apertural margin, narrow with elevated rim, overhanging ventral concaved sides and distinct longitudinal part of shell, rather smoothly rounded. ridges on the dorsal surface. Some Posterior horn spinose, lateral ends not specimens identified as Diacrina bisulcata spinose but shouldered. Best preserved 480 · Hiroshi NODA and largest specimen is 12.9 mm in length Gushikami-son, Okinawa . Prefecture, and 10.5 mm in width. Shinzato Formation, Pliocene, IGPS, coiL Comparison and affinities:-The pre· cat. no. 92588; Loc. no. 129, west side sent species resembles Cavolinia globulosa cliff of Route 46 (prefectural road), (GRAY) and Cavolinia uncinata (RANG) in north of Gushikami, Gushikami-son, having similar shell form and growth Okinawa Prefecture, Shinzato Fo-rmation, lines but the latter two differ from the Pliocene, IGPS, coli. cat. no. 92589; Loc. present species by their .short and no. 12-06. sea side cliff, we~t of Tobaru, slightly incurved posterior horn and Miyagusuku'shima, Yonagusuku-soh, wide apertural~side. Cavolinia tridentata_ Okinawa Prefecture, Shinzato Formation, which was described from the Semat"' Pliocene, IGPS, coli. cat. no. 92590. Pleistocene Formation by Y AMAKAWA and ISHIKAWA (1912b) may be identified with the present species in being charac· Cavolinia okinawana NODA, n. sp. terize~ by one medial longitudinal ridge Pl. 57_, fi_gs. 27-28 with two on the dorsal shell, the shell form and posterior and lateral ends. Type locality :-Loc. no. 12-06, sea si.de Remarks :-The present species is. very cliff, west of Tobaru, Miyagusuku·shima, common in tuffaceous medium to fine Yonagusuku-son, Okinawa Prefecture, grained sandstone and tuffaceous silt­ Shinzato Formation, Pliocene. stone of the Shinzato Formation. The Description :-Shell fragile, rather small species has been discussed in detail by in size, subrounded, _inequiv'alve, equi­ COLLINS .(1934) who listed its many syno· laleral in shell form. Ventral shell nyms. The generic names Cavolinia and inflated, external surface smooth, covered Cavolina were treated by PELSENEER with dorsal apertufe. Dorsal valve not (1888a) who used Cavolinia in sa·me sense so swollen, sculptured with faint, ele· as Cavolina. Many previous authorities vated radial fold on apertural margin used the generic name Cavolina but and indistinct on middle to posterior recently SPOEL (1967) and KURODA, HABE side., Both sides. narrowly depressed and OYAMA (1971) used Cavolinia for with small elevated ridge. Aperture Cavolina. The nomenclature of the narrow, quadrate, apertural plate obli· generic name should be subjected to quely positioned with smooth surface.. further consideration. Lateral sides quadrate but not spinose. Locality and jormati011 :--'-Loc. no. 42, Posterior end small and spinose. east of Nesabe, Tomigusuku-son, Dimension :-Height 5.9 mm and width Okinawa Prefecture, lower part of 5.9mm. Yonabaru Formation, Miocene, IGPS, Compa,·ison and affinities:-The pre· coil. cat. no. 92585 : Loc. no. 15, south sent species resembles Cavolinia triden­ of Shinzato, Sashiki-son, Okinawa Pre· tata (FORKSKAL) . which is characterized fecture, Shinzato Formation, Pliocene, by faint radial ribs on the dorsal surface IGPS, coli. cat. no. 92586 ; Loc. no. 123, but differs from the latter in having small road side cliff, north of Hanagu· narrow radial depressed area, and smal! suku, Gushikami-son, Okinawa Prefec· elevated ridge on the lateral side, and ture, Shinzato Formation, Pliocene, IGPS, faint radial fold at the apertural margin coli. cat. hot. 92587; Loc. no. 126, small on dorsal surface. Cavolinia telemus road side cliff, north of Hanagusuku, LINNAEUS is considered a synonym of 610. Fossil Pteropoda, Miyazaki and Okinawa 481

1Cavolinia tridentata by KURODA, HABE tuberous in form and circular in cross :and OYAMA (in Bioi. Lab. Imp. House· sectiol).. Aperturai part widest, posterior hold, 1971) but the present writer con· end spinose put not curved. Shell sur· ·siders that both species are valid be­ face smooth but with transverse stria· ·cause the characteristics of the dorsal tion. surface are different from each other Comparison and affinity:-The present .as mentioned by TOKIOKA (1960) and species resembles Creseis virgula RANG . YAMAKAWA and ISHIKAWA (1912b). Ca­ Both Creseis acicula and C. virgula are volinia telemus differs from the present subdivided into subspecies, the former ·species in having only one mid-radial into two and the latter into four sub· ridge on the apertural margin and nar· species according to the morphologicai ·row apertural shell band. features of the posterior end and trans· Locality and formation:-Loc. no. 12- verse striation (s). The present species ·06, sea side cliff, west of Tobaru, Miya­ differs from the Creseis virgula group _gusuku·shima, Yonagusuku-son, Okina­ in having straight. posterior end without wa Prefecture, Shinzato Formation, Plio­ expansion near the posterior e'nd like eene, IGPS coil. cat. no. 92591. a knob. A subspecies, Creseis acicula clava RANG, has a shorter tuberous shell but resembles the· species. Genus Creseis RANG, 1828 Locality and formation :-Loc. no. 12, west of Y agena Harbour, Y agena, Yo· Creseis acicula (RANG, 1828) nagusuku-son, Okinawa Prefecture, Shi· Pl. 57, figs. 1-5. nzato Formation, Pliocene,· IGPS,' coil. cat. no. 92592; Loc.' no. "15, south of ·Clio acicula RANG, 1828, Ann. Sci. Nat. Ser., Shinzato, Sashiki-son, Okinawa Pre· vol. 13, p. 318, pl. 17, fig. 6, (fide PEL· fecture, Shinzato Formation, Pliocene, SENEER, 1888a) IGPS, coil. cat. no. 92593. ·Clio acicula RANG : PELSENEER, 1888a, C/ral­ lenger Rep., vol. 23, p. 51-53. ··Clio (Creseis) acicula RANG: YAMAKAWA and IsHIKAWA, 1912a, jour. Geol. Soc. Family Atlantidae Tokyo, vol. 19, p. 2-3, pl. 1, figs. la-b. Genus Atlanta LESUEUR, 1817 -Creseis acicula (RANG) : CoLLINS, 1934, john Hopkins Univ. Geol.. no. 11, · p. 207-208, Atlanta okinawana NODA, n. sp. pl. 9, figs. 6-7, pl. 13, figs. 7-8. -Creseis acicu{a (RANG): ABBOTT, 1954, p. Pl. 57, fig. 21 294, fig. 64-n. ·Creseis acicu{a (RANG) : TOKIOI

LHYMA:-1, L. H .. 1967, The invertebrates. vol. vol. 73, pt 2, p. 305-435, pis. 16-47, figs. 1. Mollusca I. MacGratu Hill Book Co., 1-48. N.Y. 792 pp .. 249 figs. RANG, S., 1825, Description de deux generes KEEN, A.M., 1971, Sea shells of tropical nouveaux ~ppartenant a Ia classe des pte­ west America. Stanford Univ.. Press .. ropodes. Ann. d. Sci. Nat., Tom. 13, Stanford. 1064 pp., 22 pis. (non vidi). KoBAYASHI, T .. 1956, A paleo-meteorological REEVE, L., 1878, Monograph of the genus interpretation to the occurrence of the Pteropoda. Conch. Cab., vol. 20. pis. 1-6. Argonautinae in province Kaga, central ROBBA, E., 1971, Associazioni a pteropodi Japan. japan. jour. Geol. Geogr., vol. della formazione di Cessole (Langhiano). 27, nos. 2-4, p. 93-104, pl. 6, figs. a-d. Riv. /tal. Paleont., vol. 77, no. 1, p. 19- KoNISHI, T., et a{., 1970, Reports on the 126, pls. 1-5, fig. l-5. development of underground water re· SPOEL, S. van der, 1.967, Eutheocosomata. sources. nos. 1-3, Geol. Surv. japan, 77 A group with remarkable development pp., 27 figs. stages. (Gastropoda-Pteropoda). Zoo/. --. et al., 1971, Ditto .. no. 4, Geol. Surv. Mus., Amsterdam, ]. Noorduijin en Zoo11, japan, 70 pp., 15, figs. 9 tabs. N. V., p. 1-376. (fide OvAMA, K., 1970, LAI)D, H. S., 1934, Geology of Viti levu, Fiji. Venus, vol. 29, no. 1, p. 12). Bernice P. Bishop Mus., Bull .. 119, p. 1- TAYLOR, D. and SoHL, N., 1962, An outline 263, pis. 1-44, figs. 1-11, tabs. 1-7. of gastropod classification. Malacologia, LAMARCK, Par ]. B. P. A., 1836, Historie vol. 1, no. 1, p. 7-32. Naturelle des Animaux sans Vertebres. TESCH, ]. J .• 1904, The Thecosomata and Tom. 7, p. 1-735. Gymnosomata of the Siboga Expedition. MAcNEIL, F. S., 1960, Tertiary and Quater­ Siboga Exped. Mo11ogr., vol. 52, p. 1-92, nary Gastropoda of Okinawa. U.S. Geol. pis. 1-6. Surv. Prof. Paper, 339, p. 1-148, pis. 1- Toi

Aoshima }If lh Oroku 1], -*-~ Chinen ~:u $ Saito i!\'i Gushikami A:t:~ Sashi!d It ~·j( Haebaru i~ Hi. DR Semata if![ 3l. Hanagusuku 1E .1]£ Shimajiri Jfh fJl. Katsuren JIJI .iili Shinzato ~ .li':. Kawabaru Jll t.m Tobaru tit }]( Koyu .'R. ~ Tomigusuku ft.i.l!£ Miyagusuku 'S .1]£ Urazoe rm i~ Miyazaki 1r Yamaji 111 m Naha m ·~1m Yonabaru ~· »~ !* Okinawa i'l' "' Yonagusuku ~»>IIAlt

Explanation of Plate 57

Figs. 1-5, Creseis acicula RAr.;G, x 20, Loc. no. 15, Pliocene Shinzato Formation, IGPS coiL cat. no. 92593. Figs. 6, 12, 13, Cleodora okinawana NOD A, n. sp., x 3, figs. 6, 12, Loc. 15, IGPS·, coli. cat:­ nos. 92582, and 92581, Fig. 13, Loc. 12-06, Pliocene Shinzato Formation, IGPS coli. cat. no. 92596. Figs. 7, 8, Hyalocylix striata (RANG), x 5, Loc. no. 12-06, Pliocene Shinzato Formation, IGPS coli. cat. no. 92583. Figs. 9-11, Cleodora hataii NooA, n. sp., x2, Miocene Kawabaru Member of the Koyu Forma­ tion, IGPS coli. cat. no. 92579. Figs. 14-17, 19-20, 22-26, 29-31, Cavolinia telemus LINNAEUS, Figs. 14-16, 19--20, Loc. 123, x 2,. IGPS coli. cat. no. 92587; fig. 17, Loc. 129, x3. IGPS coli. cat. no. 92589,; fig. 22, Loc. 123, x 2, IGPS coli. cat. no. 92587; fig. 24, Loc. 123, X 3, IGPS coli. cat. no. 92789; fig. 23, x2, Loc. ~5. IGPS coli. cat. no. 92586; figs. 25, 26, x3, fig.27, x5, fig.30-31, x3,allfronlc Loc. 12-06, IGPS colt. cat. no. 92590; fig. 29, Loc. no. 126, IGPS colt. cat. no. 92588. All,. Pliocene Shinzato Formation. Fig. 18, Diacrina bisulcata GABB, x 5, Loc. 1206, Pliocene Shinzato Formation, IGPS coli. cat.. no. 92584. Fig. 21, Atlanta okinawana NooA, n. sp., x 8, Loc. 12-06, Pliocene Shinzato Formation, IGPS: coil. cat. no. 92594. Fig. 28, Cavolinia okinawana NoDA, n. sp., x3, Loc. 12-06, Plioc~ne Shinzato Formation, IGPS coli. cat. no. 92591. NODA.: Fossil Ptero1;oda, Miyazaki and Okinawa Plate 57

3 5 7

8

13 10 11

14 JS 16 17 18

19 20 21 22 23

24 25 26 27

28 2\J 30 31 PROCEEDINGS OF THE PALAEONTOLOGICAL SOCIETY OF JAPAN

r-1*i!ig:_:f'/1J~~m no IID"J~r:t. 1972 ~ 10 FJ ...... -3KUl :llr- 28 13 (±) ~~:k~Illl~§l\l;:::t;lt•"(f}~fil!~t'l.t.:. '''Ui1iH.Jl!WJ-c ~*• ~ n t.: ~ #il ~q:s Q)1f1L !:R <~1J!J~ so ,g). flf!~ 29 a

(New Series No. 81-No. 88)

PALAEOZOOLOGY

Protozoa Page CHOI, Dong Ryong: Colania douvillei (OZAWA), a fusulinid Foraminifera, from the Nor- thern Kitakami Mountains, NE Japan...... 369 KuRIHARA, Kenji: Foraminifera from the. Hayama Group, Miura Peninsula ...... 131 MATSUMARU, Kuniterui!' 1 The genera Nephrolepidina and Eulepidina from New Zealand.. 179 OKIMURA, Yuji: Permo-Carboni·ferous endothyraceans from Japan, Part 1. Biseriam- minidae ...... 414 SADA, Kimiyoshi: Fusulinids of the Profusulinella zone of the Taishaku limestone. (Studies of the stratigraphy and microfossil faunas of the Carboniferous and Permian Taishaku Limestone in West Japan, No. 2) ...... 436 YAMAGIWA, Nobuo and SAKA, Yukiyasu: On the Lepidolina zone discovered from the Shima Peninsula, Southwest Japan...... 260

Coelenterata

KATO, Makoto: J. FLEMING's species of British Lower Carboniferous corals...... 1

Bryozoa HAY AMI, Tomoko: Some Neogene Cheilostomata (Bryozoa) from Okinawa-jima ...... 73 ----: On some Bryozoa from near Namioka-cho, Minami-Tsugaru-gun, Aomori Pre- fecture, Japan...... 196 SAKAGAMI, Sumio: The Triassic Bryozoa from Kusaka, Sakawa basin, Shikoku, Japan.. 275

Mollusca IsHIBASHI, Takeshi: Upper Triassic cephalopods from the Tanoura district, Kumamoto Prefecture, Japan ...... 447 MAsUDA, Koichiro: On some Pathtopecten from North America ...... 166 ---: Amussiopecten from North America and northern South America...... 205 ---: Swiftopecten of the Northern Pacific ...... 395 MATSUMOTO, Tatsuro, MURAMOTO, Tatsuo and !NOMA, Akitoshi: Two small desmoce­ ratid ammonites from (Studies of the Cretaceous ammonites from Hokkaido and Saghalien-XXIV) ...... 377 MuRATA, Masafumi: Fusulinid biostratigraphy and molluscan fauna from the uppermost part of the Sakamotozawa formation, and the pre-Kanokura unconformity in the sou- thern part of the Kitakami massif, North-east Japan ...... 93

486 Systematic Index 487

'NooA, Hiroshi: New anadarid and associated molluscan fauna from the Haneji Forma- tion, Okinawa-jima, Ryukyu Islands ...... 27 ---: Some fossil Pteropoda from Miyazaki and Okinawa Prefectures, Southwest Japan ...... 472 NooA, Masayuki: Ammonites from the Mesozoic Yamabu formation, Kyushu ...... 462 ·SATO, Tadashi: Some Bajocian ammonites from Kitakami, Northeast Japan ...... 280 TASHIRO, Masayuki: Upper Cretaceous glycymerids in Japan ...... 225 ---: On the surface ornamentation of the pennatae trigoniids, and on three new species of the trigoniids from the Himenoura Group, Kyushu, Japan ...... 325

Arthropoda

Hu, Chung-Hung: Ontogeny of three Cedaria zone trilobites from Upper , Mon- tana...... 245 --- & TAN, Li-Lin: Ontogenies of two Upper Cambrian trilobites from northern Black Hills, South Dakota ...... 61

Vertebrata

HAT AI, Kotora and NooA, Hiroshi: A Problematica from the Mizuho-To of Pre- fecture ...... 319

Miscellanea

HATAI, Kotora and KoTAI

PALAEO BOT ANY

-(Megafossils) KIMURA, Tatsuaki and SEKIDO, Shinji: The discovery of the cycad-like leaflets with toothed margin from the Lower Cretaceous Itoshiro Sub-group, the Tetori Group, Central Honshu, Japan...... 190 --- & : Ctenis species from the Itoshiro Subgroup (Lower Cretaceous), the Tetori Group, Central Honshu, Japan ...... 360

(Microfossils) BuRCKLE, Lloyd H.: Correlation of Late Cenozoic marine sections in Japan and the Equatorial Pacific ...... 117 FUJI, Norio: Fossil spores and pollen grains from the Neogene deposits in Noto Penin- sula, Central Japan-IV. A palynologiccal study of the Late Miocene Tsukada Mem- ber ...... 295 Ko!ZUMI, Itaru: Marine diatom flora of the Pliocene Tatsunokuchi Formation in Fuku- shima Prefecture ...... 340 NisHIDA, Shiro: Nannofossils from Japan IV. Calcareous nannoplankton fossils from the Tonohama Group, Shikoku, Southwest Japan ...... -... 143 488 Nos. 81,.._88 (1971, 1972)

TAKAHASHI, Kiyoshi: Microfossils from the Pleistocene sediments of the Ariake· Sea area, west Kyushu...... llt TAKAYAMA, Toshi

PSEUDOFOSSIL

HAT AI, Kotora and NooA, Hiroshi: Peculiar markings on a sandstone layer of the Ila- gino Formation, ...... 162: 489>

INDEX OF FAMILIES, GENERA AND SPECIES

Notes: Words listed are names of families, genera and species, which are either described or illustrated in the Transactions and Proceedings of the Palaeontological Society of Japan,. New Series, Nos. 81-88; words in heavy type are names of new genera and species.

A Braarudosphaera bigelowi ...... 155, 43(} Buchites...... 449 Abies sp. 316 --- kumamotoensis ...... 449· Actinocyclus ellipticus ...... 350 Bulimina striata...... 142. elongatus ...... 350 Adeonellopsis arculifera ...... 87 --- sttbleres ...... 87 c --- yarrae11sis ...... 87 Caberea hataii ...... 8(} Amussiopectelt...... 209 Carya sp...... 312 Amussiopecten antiguensis...... 211 Aff. Carya sp ...... :...... 314 --- churuguarensis...... 213 Caryophy/lea (sic) a/finis ...... 4 ---- harrisi ...... 213 --- (sic) duplicata ...... ~ --- preglyptus ...... 212 (sic) fasciculata ...... 4 --- sp ...... 215 Catapsydrax dissimilis ...... 14Z --- va11vlecki...... 210 Cavolinia okinawana ...... 480 --- wooclringi ...... 214 ---- telemus . .'...... 479· Anadara (Hataiaraca) kogachiensis . . . . 37 Cellaria punctata ...... 80· Antropora tinct a ...... 77 Ceratolithus cristus ...... 155 Aphelaspis walcotti...... 62 --- aff. tricorniculatus ...... 15& Apiotrigonia...... 331 Cigclisula occlusa...... 83. --- minor ...... 326 Cladog ramma californica ...... 350 --- obsoleta ...... 326, 331 Cleidochasma bassleri ...... _ 82 --- cf. obsoleta...... 326 --- biavicularia ...... 81 --- postonodosa...... 326 --- aff. fa/lax...... 82 --- undulosa ...... 326 --- granifera ...... 82 --- utoensis ...... 333 Clementia (Clementia) vatheleti ...... 42 Arcestes (Proarcestes) cf. ausseeanus . . 451 Cleodora ltataii ...... 475. Asteromphalus darwinii ...... 350 --- okinawana ...... 477 Astrononion ltayamaense ...... 140 Coccolithus pelagicus ...... 149- Atlanta rniyagusukuana ...... 481 Codakia (]agonia) okinawazimana ...... 41 Aviculopecten sp...... 110 Colania douvill ei ...... 371 ? sp...... 111 Conescharellina cf. breviconica ...... 89- concava ...... 8~ B Copidozoum planum ...... 78. Bacteriosira fragilis...... 350 Coscinodiscus lineatus ...... J.50. Baltag nostus beltensis...... 254 nodulifer ...... 350· Baltisphaeridium sp...... 20 --- pustulatus ...... 35(} ? sp...... 21 Costazia radiata ...... 89- Batillaria zo1talis...... 45 ___ rota ...... 201 Bellerophon (Bellerophon) kitakamiensis 105 Cranosina coronata ...... 78. Biedulphia aurita ...... 350 Creseis acicula ...... 481 490 Nos. 81,.._88 (1971, 1972)

Cricolithus sp ...... 149 --- sp. B...... 423 Ctenis ...... 360 Globoquadrina dehiscens ...... 142 --- kaneharai ...... 364 Glycymeris ...... : ...... 226 --- nipponica ...... 361 Glycymeris (Glycymeris) amaktisensis . . 226 ---sp...... 365 (Glycymerita) aff. apletos...... 239 Cyclammina orbiwlaris ...... 142 --- ( ) himenourensis . . . . 229 Cyclococcolithus leptoporus ...... 149 --- ( ) japonica ...... 228 Cymatiosphaera globulosa ...... 21 --- ( ) multicostata...... 232 --- reticulosa ...... 21 --- ( ) veatchii .. ·...... 239 Cysti sella ? americana ...... 201 Glycymeris (Hanaia) densilineata . . . 234 --- midwaym1ica ...... 200 --- (---) haipensis ...... 234 --- ( ) hokkaidoensis ...... 235 D --- ( ) katsurazawensis . . 235 Denticula kamtschatica 350 --- ( ) matsumotoi . . . 233 --- seminae ...... 350 --- ( ) salida ...... 233 Desmoceratinae ...... 378 --- ( ) sublaevis ...... 234 Diacria bisulcata ...... 478 (Pseudoveletuceta) mifunensis 236 Diatosula marionense ...... 200 Glycymerita ...... 228 Dis coaster brouweri ...... 154 --- brouweri rutellus ...... 154 H --- challengeri ...... 154 Hanaia ...... 233 --- dilatus ...... 154 Helicoponlosphaera kamptneri ...... 152 --- kugleri ...... 154 Hemicystodinium cf. zoharyi ...... 18 --- pentaradiatus ...... 154 Hemidiscus cuneiformis ...... 351 --- stellulus ...... 155 --- weissftogi...... 351 --- surwlus ...... 155 Heterotrigonia ...... 334 Discolithina sp. A ...... : ...... 149 --- diversicostata ...... 332 Ditrupa ...... 412 --- himenourensis ...... 334 ------,. miyazakiensis 412 --- Sltbovalis ...... 332 Hippoporella spinigera ...... 83 E Hitachia ...... 460 Encephalartites leipzigii ...... 192 --- nakaminatoensis...... 460 Eulepidina dilatata dilatata ...... 184 Hoeglundina elegans...... 1•12 Holconautilus sp...... 455 F Holoporella palmata ...... 88 Fagus sp ...... ''I'...... 312 --- (?) sp...... 89 Fragilariopsis cyclindrus .... :...... 350 Hyc1locylix striata ...... 478 Fulvia sp...... 41 Hyalodiscus obsoletus ...... 351 Hystrichokibotium sp...... 19 G Hystrichosphaeridium ct. ferox ...... 17 Glaphyraspis parva ...... 66 --- sp. a ...... 18 Globigerinatella insueta ...... 142 --- sp. b ...... 18 ·Globiualvulina...... 417 cf. tiara ...... I 7 --- cf. gracea ...... 421 --- granulosa compressa ...... 420 I --- kamensis ...... 418 Cf. llex sp...... 314 --- cf. kantharensis ...... 421 Inapertipollent ites sp...... 312 --- mosquensis...... •119 --- 1·egularis ...... 422 J --- sp. A ...... 423 fuglans sp ..... 312 Index of Families, Genera and Species 491

K Navicula jamalinensis ...... 351 Kitchinites (Neopuzosia) haboroensis . . 384 Neozamitcs ...... 191 Knightites (Retispira?) hanzawai ...... 106 --- elongata ...... 192 Kubikichtlrys ...... 322 Nephrolepidina hornibrooki ...... 186 howchini ...... 187 --- rm·is ...... 322 orakeiensis ...... 1.85 L Neopuzosia ...... 384 Laevicardium sp...... 41 Nitzschia cf. exline/a ...... 351 Leiosphaeridia glolmlifera ...... 22 --- fossilis...... 352 Leopecten ...... 170 ---- reinholdii ...... 351 --- bakeri ...... 171 Nixonel/a ...... 250 --- bakeri diazi ...... 172 --- montanensis ...... 251 marquerensis ...... 173 Nodosaria longiscata ...... 142 Lepitlolina kumaensis ...... 267 Normannites (Itinsaites) cf. itinsae . . . . 283 --- multiseptata gigantea 265 Nuphar (?) sp...... 314 multiseptata multiseptata ...... 266 Limopsis kogata ...... 237 0 Lithostrotion jloriforme ...... 3 Osmund a sp...... 314 marginatum ...... 3 Ostrea (Ostrea) denselamellosa...... 40 striatum ...... 2 Otoites sp...... 28~ Lunella coronatus granulatus ...... 44 Ovoidites cf. microligneolus ...... 2~ --- sp ...... 44 --- ellipsoideus...... 22

M p Macoma (Macoma) praetexla ...... 42 Pachyphloia sp. 272 Martinottiella communis ...... 142 Parasmittina aviculoumbonata ...... 84 Melonis pacificus ...... 142 --- peristoaviculata ...... 84 albicans ...... 351 --- spathulata ...... 84 Mesopuzosia cf. pacifica ...... 391 --- trispinosa ...... 85 Metadoliolina gravitesla ...... 270 Patinopecten ...... 169 Micrhystridium ar·iakense ...... 19 Patinopecten (Patinopecten) skonunensis 169 ---densum ...... 20 Pelekodites cf. pelekus ...... 282 Microdesmoceras ...... 378 (Spatulites) spatians ...... 281 --- tetragonum ...... 378 Phormedites ...... 450 Microporina articulata ...... 198 --- sp ...... 450 Microtrigonia ...... 335 Picea sp ...... 316 amanoi ...... 326 Placites ...... 45~ --- imutensis ...... 335 Planktoniella sol ...... 352 Millerella sp. B ...... 437 Pododesmus (Monia) noharai ...... 39 Mizuhopecten ...... 170 Polinices cumingianus madioenensis ... . 46 --- warreni ...... 170 Pontosphaera ~alboranensis ...... 152 Modiolus sp ...... 38 --- japonica ...... 152 Mucronella sp ...... 84 --- sp. ··························· 153 "Murchisonia" sp...... 107 --- cf. vadosa ...... 152 Myriozoum coarctatum ...... 202 Porosira glacial is ...... 352 --- sub gracile ...... 202 Profusulinella fusiformis ...... 439 toriyamai...... 43& N Pseudoactinodontophora ...... 111 Nankinella yokoyamai ...... 441 --- yabei ...... 112 Nassarius (Zeuxis) caelatus ...... 46 Pseudobatostomella kobayashii ...... 275 492 Nos. 81,-...88 (1971, 1972)

Pseudopodosira elegans ...... 352 Stephanoceras sp. 284 Pseudostaffella taishakuensis ...... 440 Stephanopyxis lwrridus ...... 348 Pseudoosterella sp...... 467 --- schenckii ...... 353 Pseudoueletrtceta ...... 236 Striarca interp/ic~ta ...... 36 Pteria cf. coturnix ...... 38 Strigoceras cf. languidum ...... 285 Pullenia bulloides ...... : ...... 142 Swiftopecten ...... 398 Puzosiinae ...... 383 --- swi/tii ...... 399 Pyxidicula weyprechtii ...... 352 --- swiftii parmeleei ...... 401 Syracosphaera pulchra ...... 153 Q Syspacheilus dunoirensis ...... 246 ·Quercus sp. 314 T R Tainoceras kitakamiense...... 108 Rhabdosphaera claviger ...... 157 Thalamoporella novae·hollandiae ...... 79 Rhacophyllites sp...... 454 Thalassiosira antiqua ...... 353 Rhaphoneis ischaboensis ...... 352 convex a ...... 353 margaritalimbata ...... 352 gravida...... 353 ---- tatsunokuchiensis ...... 349 hyalina...... 353 Rhizosolenia bergonii ...... 353 kryophila ...... 354 Rhynchozoon aff. grandice/la ...... 86 nidulus ...... , ...... 354 sp ...... 86 nordenskioldi ...... 354 (?) sp ...... 86 --- zabelinae ...... 3S4 s Tilia sp...... 312 .Salix sp...... 312 Towapteria nipponica ...... 109 Schisrnopora chrysalis ...... 202 Tricellaria sp...... 199 .Schizobrachiella subhexagona ...... 200 Tricolpropollenites sp...... 316 Schizomavella ovoidea ...... 81 --- sp ...... 81 u Schizoretepora tumescens ...... 201 Umbilicosphaera cricota ...... 153 Schyphosphaera apsteini ...... 157 Umbonium (Suchium) moniliferum Siphonodendro11 affine ...... 5 decoratum ...... '...... 43. Smiltina sp...... 85 .Smittoidea acaroensis...... 85 w .Sphaeroidinlllla seminulina ...... 142 Waagenoperna (Permoperna) hayamii . . 109· Spiniferites ramosus ...... 16 y Spirosigmoilinella r:ompressa ...... 142 Sta.ffella akagoensis ...... 443 Yabeina columbiana...... 268 Steganoporella magnilabris:...... 79 aff. globosa...... 269 Transactions and Proceedings

of the Palaeontological Society of Japan

New Series

No. 81 ...... No. 88

1971-1972

Palaeontological Society of Japan The heading in Japanese commemorates the handwriting of Prof. Matajiro· YOKOYAMA, father of Japanese palaeontology, who W~S a professor of stratigraphy and palaeontology at the Geological Institute, Imperial University of Tokyo.

Fossil on the cover is left lower M2 of Palaeoloxodon naumamti (MAK!Y AMA, 1924) from the uppermost part of the Tokyo formati'on (Upper Pleistocene) at Ike· bukuro, Tokyo. CQNTEN.TS . ; ..., 'J 1 I r. I 1: j • f)'.' '1., , J!f;or' l • t • .... ,-,! j,

; I

Number 81 (Pubiish~d· April20, 1971) ., ',/ Transactions Article Page Plate 574. KATO, Makoto: J. FLEMING's species of British Lower Carboni- ferous corals ...... 1-10 1 575. TAKAHASHI, Kiyoshi: Microfossils from the Pleistocene sedi- ments of the · area, west Kyushu ...... 11-26 2-5 576. NODA, Hiroshi: New anadarid and associated molluscan fauna from the Haneji Formation, Okinawa-jima, Ryukyu Islands . ... . 27-51 6-7 ' .. 577. HATA!, Kotara and KOT:"-KA, Tamio: Some coprolites from Wa- kayama Prefecture ...... · ...... 52-58 Proceedings 59-60

Number 82 (Published June 30, 1971).

Transactions 578. Hu, Chung-Hung and TAN, Li-Lin: Ontogenies of two Upper Cambrian trilobites from northern Black Hills, South Dakota . . 61-72 8-9 579. HAY AMI, Tomoko: Some Neogene Cheilostomata (Bryozoa) from Okinawa-jima ...... ·...... 73-92 10-12 580. MuRATA, Masafumi: Fusulinid biostratigraphy and molluscan faj.lna from the uppermost p~u·t of the Sakamotozawa formation, and the pre-Kanokura unconformity, in the so-uthern part of the Kitakami massif, Northeast japan ...... ' ...... 93-116 13-14 581. BURCKLE, Lloyd H. : Correlation of Late Cenozoic marine sections in Japan and the Equatorial Pacific ...... 117-128

Number 83 (Published September 20, 1971) Transactions 582. KURIHARA, Kenji : Foraminifera from the Hayama Group, Miura Peninsula ...... 131-142 15 ii

583. NISHIDA, Shiro: Nannofossils from Japan IV. Calcareous nan­ noplankton fossils from the Tonohama Group, Shikoku, Southwest Japan ...... _ ... , ... , ...... , ...... 143-161 16-lS. f" o j j; I '- t '! I • ~ 584. HATAI, Kotora and NODA, Hiroshi: Peculiar markin~s on a sand- stone layer of the Hagino Formation, Nagano Prefecture ...... 162-165 585. MASUDA, K6ichir6: On some Patinopecten .. from No~th America ...... · ..... ·: ...... 166-178 19-21

Number 84 (Published December 30, 1971) Transactions 586. MA TSUMARU, Kuniteru : The genera Nephrolepidina and EulePi· dina from New Zealand ...... 179-189 22-23 587. KIMURA, Tatsuaki and SEKIDO, Shinji : The discovery of the cycad· like leaflets with toothed margin from the Lower Cretaceous ltoshiro Sub-group, the Tetori Group, Central Honshu, Japan .. .' .190-195 · 24 588. HAY AMI, Tomoko: On some Bryozoa from near Namioka-ch'o, Minami·Tsugaru:gun, , Japan ...... 196-204 589. MASUDA. K6ichir6: Amussiopecten from North America and nor· them South America ...... 205-224 25-26 590. TASHIRO, Masayuki: Upper Cretaceous glycrmerids in Japan .. 225-242 27-28- Proceedings ...... 243-244

Numper 85 (Published April 20, 1972) Transactions . . . .

591. Hu, Chung· Hung: Ontogeny of three Cedaria zone trilobites from Upper Cambrian, ...... ·...... 245-259 29-30 592. YAMAG!WA~ Nobuo and SAKA, Yukiyasu: Ori the Lepidolina zohe

discovered from. the Shima Peninsula, Southwest Japan ... ·..... 260-274. 31-32 593. SAKAGAMI, Sumio: The Triassic Bryozoa from Kusaka, Sakawa basin, Shikoku, Japan ...... : ...... 275..:279 33 594. SATO, Tadashi: Some Bajocian ammonites from Kitakami Nor· theast Japan ...... : ...... 280-292 34 Proceedings ...... 293 iii

! • Number 86. (Pt.Jl>lished june 30,1972) Transactions . . . - . _. ·I : :595. FUJI, Norio: Fossil spores and· pollen grains from the Neogene deposits in Noto Peninsula, Central· Japan-IV. A palynological study of. the Late Miocene 1.'!!ukada Memper ...... , ... ._ ..... 295-318 35-38 ' . ' .. '596. HAT AI, Kotora and NODA, Hiroshi: A Problematica from the ... . . ~ . Mizuho·To of ...... : ... 319-324, 39 .'597. TASHIRO, Masayuki: On the surface ornamentation of the pen· natae trigoniids, and on three new species of the trigoniids from the Himenoura Group, Kyushu, Japa_n ... : ...... 325-339 40-41 :598. KOIZUMI, Itaru: Marine diatom flora of the Plioce~e Tatsuno· kuchi·Formation,in Fukushima Pr~fecture ...... 340-359 41-43 599. KIMURA, Tats~aki and SEKIDO, Shi~ji : Ctenis species from the ltosh\ro Sub-group (Lower Cretaceous), the Tetori Group, Central Honshu, japan ...... 360-368 44-45 ·600. CHOI, Dong Ryong: Colania douvillei (OZAWA), a fusulinid Fora· minifera, from the Northern Kitakami Mountains, NE japan .... 369-374 46

Number 87 (Published September 30, 1972) Transactions ·601. MATSUMOTO, Tatsuro, MURAMOTO, Tatsuo and lNOMA, Akitoshi : Two small desmoceratid ammonites from Hokkaido (Studies of the Cretaceous ammonites from Hokkaido and Saghalien-XXIV) .. 377-394 47 ·602. MASUDA, Koichiro: Swiftopecten of the Northern Pacific ...... 395-408 48-49 ·603. HAT AI, Kotora, NODA, Hiroshi and OGASAHARA, Kenshiro: Sea· phopoda·like fossils from the Udo formation (Miocene) of Miya- zaki Prefecture, Japan ...... 409-413 ·604. OKIMURA, Yuji: Permo-Carboniferous endothyraceans from Japan, Part 1. Biseriarnminidae ...... 414-428 50-51 ·605. TAKAYAMA, Toshiaki: A note on the distribution of Braaru- dosphaera bigelowi (GRAN & BRAARUD) DEFLANDRE in the bottom sediments of Sendai Bay, Japan ...... 429-435 -606. SADA, Kimiyoshi : Fusulinids of the Profusulinella zone of the Taishaku limestone. (Studies of the stratigraphy and microfossil faunas of the Carboniferous and Permian Taishaku Limestone in iv

West japan, No. 2) ...... 436-445 52-53

:-::1'•!··-..;.:•:!t.•·' Number 88 (Published December 20, 1972) · F:~:-tt•".::/! ,d r r , • ·: j ·; ,,, ... :: ,,fl ·~. :1': ., • f,:·

!1. ··.··t :•. , • ,. •._ . T.. ransactio.n__ ... s. _ , _ ..•· ... . •• r ••• • ,;~ ( 1r; ': _· , :6o{'· I§ms.\sHr, Takeshi: Upper Triassic cephalopods'from the·Tanouta·'· ·, .. :- :.. district, 'Kum3:il1b'to Prefecture; Japan··:.'.~· .. .'.: ·.. ·. ::.' .. ·..... ·, :·! ·:: :447'-457 ' ·54 '608. 1-IATAI.' Kotora and NODA, Hiroshi: A fossil ·worin trail ·from the · Upper Cretaceous Nakaminato formati.on, lbaraki Prefecture .... 458-461 ·55 609. · NODA, Masayuki' :· Ammonites from the Mesozoic .Yamabu forma- '· · tion, Kyushu . : .... : .. : ...... , ...... ·...... :461-471 56- 610. NODA, Hiro'shi': Some fossil Pteropoda from Miyazaki and Oki .. n~rwa Prefectures, Southwest Japan ...... 472-484 57 Proceedings . : ...... 485 Systematic Index ...... 486-488 Index of Families, Genera and Species .. 489~492

•'' ·; rlfJ Hi! ffil r.n !1li B ~~ ilit $ ~ P.6 ~ B 1973~ *1}~. ~~ Jl ::lt ~ 1973 ~ 1 ~ 16·17 B .. ~ * 1973 ~ 6 Fl 23·24 B A 111 @I {?lj = frr i.!:; * ~ (,:Yj£) 1973~ 4 Fl 20 s•

li!!l ~ ~ ~ 0 IUGS ..(]) 'c~~mlssii>ti' on' Stratigraphy "t'fi, 1975 ~~~~ 1 @;t;:.~¥t:Fl:l!!.~~ti=*lll~~~~ B * "t'ml~-t;;,.:::. c ~f*j£ L-t.:, .:::.tt.K.BI\i! VC. li!ll*!"t'l'i:tll.f!l~J.il ~~--:> < ;;, t\l{;:O;i1tllt'.> ~ti.'Lv';;,. i!!*U::f'171hill~N1:. C*-~Jiirli*-) t;; .l: O:iRilllPrta Cll!::lt*),

0 1973~7Fl1B-8Bir.. Oslo l::.:t;lt·'L NATO Advanced Study Institute 17)-l;tcl.-'L, rEvolution and Morphology of the Trilobita, Trilobitoidea and Merostomata.t (])a~ilit. 8t~ ~;?:f#)tJ·tL Q, iili*ll5l:f:l: Dr. D. L. BRUTON Paleontologisk Museum, Sars gate 1, Oslo 5, Norway.

1 9 7 2 ~ 12 Fl 15 El f.IJ lM ~ft:tl" 8:;$:-cJ~~~~ 1 9 7 2 ~ 12 Fl 20 El :.c * 1&: iJfF $. 2-4-16 s *$~~J:tn-t!:; ~- ~*~ (~ ~ 1:1 ~ * * 8 4 7 8 0 1t) EB ± Wi t.:l ~ 88 ~ EIJ lilU :1f :ij( ll5 ~~ m1&: 11 35: :fl:; 2 I 13 900p:j * ~tfi~tfEIJiillt*~~tt "4 EB jt; Transactions and Proceedings of the Palaeontological Society of Japan

New Series No. 88 December 20, 1972

CONTENTS

TRANSACTIONS 607. lSHIBASIU, Takeshi: Upper Triassic cephalopods from the Tanoura district, Kumamoto Prefecture, Japan...... 447 608. HATAI, Kotara and NODA, Hiroshi: A fossil worm trail from the Upper Cretaceous Nakaminato formation, Ibaraki Prefecture...... 458 609. NODA, Masayuki: Ammonites from the Mesozoic Yamabu formation, Kyu· shu ...... 462 610. NODA, Hiroshi : Some fossil Pteropoda from Miyazaki and Okinawa Pre· fectures, Southwest Japan ...... 472 PROCEEDINGS ...... 485 Systematic Index ...... 486 Index of Families, Genera and Species...... 489 Editor: Takashi HAMADA Associate Editor: Yasuhide IWASAKI

Officers for 1971 - 1972

President: Tokio SHIKAMA Councillors (* Executives) : Kiyoshi ASANO*, Kiyotal

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All communications relating to this journal should be addressed to the PALAEONTOLOGICAL SOCIETY OF JAPAN c/o Business Center for Academic Societies, Japan Yayoi 2-4-16, Bunkyo-ku, Tokyo 113, Japan. Sole agent: University of Tokyo Press, Hongo, Tokyo