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Endodontidae, Euconulidae, Zonitidae, Eta Vitrinidae Marras

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Lurralde 13 1990 p: 173-192 ISSN 0211-5891 IBERIAR PENINTSULAKO MALAKOFAUNARI BURUZKO GOGORAMEN BIOGEOGRAFIKOAK: ENDODONTIDAE, EUCONULIDAE, ZONITIDAE ETA VITRINIDAE MARRASKILO-FAMILIEN KASUA EUSKAL HERRIAN ETA ALDAMENETAKO LURRALDETAN (*) Recibido: 1989-11-14 K.ALTONAGA, A.I.PUENTE & C.E.PRIETO Zoologia Laborategia Animali Biologia eta Genetika Departamentua Zientzi Fakultatea Universidad del Pais Vasco/Euskal Herriko Unibertsitatea P.K.: 644. 48080-BILBAO. Espainia. LABURPENA: lberiar Penintsulako malakofaunari buruzko gogoramen biogeografikoak: En­ dodontidae, Euconulidae, Zonitidae eta Vitrlnidae marraskilo-familien kasua Euskal Herrian eta aldamenetako lurraldetan. ' ldazlan honetan, eszenario penintsularra itxuratzen duten ela, beraz, bertako fauna malakolo­ giko konplexuaren agerpenean bitartekatu duten faktoreak aztertu dira, horrez gainera, fauna horren egungo ezagumenduaren egoera azalduz. Textu-inguru horretan, aipaturiko familiek erakutsi duten banaketa-mota adierazi da, zeina kantauritartasun-gradiente batckin erlaziona bait daiteke. Gako hltzak: biogeografia, kantauritartasuna, malakofauna, Zonitidae, Vitrinidae, Endodontidae, Euconulidae, lberiar Penintsula. * J;usko Jaurlaritzako Hezkuntza, Unibertsitate eta lkerketa Sailaren X-86.044 Projektuak finantza­ tu du ikerlan honen parte bat. Lurralde, 13: 1990 173 ------------------------------------------------------------------------------ K. AlTONAGA- A. 1. PU ENTE - C. E. PRIETO RESUMEN: Conslderaclones biogeogrâficas sobre la malacofauna de la Penin sula lbérica: el caso de las familias Endodontidae, Euconulidae, Zonitidae y Vitrinidae en el Pais Vasco y zonas adyacentes. En el presente trabajo se da cuenta de los distintos factores que configuran el escenario peninsular y que han mediado en la aparicién de una fau na malacolégica compleja, exponiéndose, ademâs, en que estado se halla actualmente el estudio de esta ultima. En ese marco, se explicita el tipo de distribucién presentado por las familias mencionadas, el cual se puede relacionar con un gradiente de cantabricidad. Palabras clave: biogeografîa, cantabricidad, malacofauna, Zonitidae, Vitrinidae, Endodontidae, Euconulidae, Penlnsula lbérica. SUMMARY: Biogeographie considerations on the malacofauna of the lberian Peninsula: the case of the families Endodontidae, Euconulidae, Zonitidae and Vitrinidae in the Basque Country and adjacent zones. ln the present work it is given account of the different factors that shape the penin sul ar scenario and have mediated in the appearance of a complexe malacological fau na; it is expound, furthermore, which is the actual state of this fauna's study. ln this context, it is explicited the distribution type shown by the above mentioned familias, which can be related with a •cantabricity" gradient. Key words: biogeography, cantabricity, malacofauna, Zonitidae, Vitrinidae, Endodontidae, Euconu­ lidae, lberian Peninsula. IBERIAR PENINTSULA MALAKO GEOGRAFIAREN ALDETIK Europako estatu gehienetan oso aurreratuta dago lur-moluskuen ikasketa, espezie­ zentsua eta espezie bakoitzaren banaketa geografikoa zehaztasun handiz ezagutzen direlarik (KERNEY, CAMERON & JUNGBLUTH 1983). Ostera, lberiar Penintsulak ezeza­ gun samar irauten du, nahiz eta azken hamarkadetan bertako malakologiak hartu duen aupada nabarmena izan. Egoera hari era bitako faktorek sorterazia da, faktore geografi­ koak eta historikoak alegia. Faktore historikoei, zeintzuak Penintsulako tradizioak jasan izan duan jarraitasun-eza aipatuz laburbildu ahal bai! dira (MANGA 1983; ALTABA & TRAVESSET 1985), faktore geografiko eta biohistorikoak gehitu behar zaizkie, berauen ondorioz konplexutasun malakologiko handia sortu delarik. Horren guztiorren emaitza, azken batean, espezieen ezagumendu urri samarra da, bai alde taxonomikotik nola biog eografikoki. lberiar Penintsula, Kaukasoarekin bata ra, Europako arearik interesgarrienetarikoa da ikuspegi biogeografikotik (SACCHI 1964). Autore horren arauera, Penintsula zubitzat jo daiteke, hain zuzen mendebaldeko Europatik, Frantziatik, Afrikaraino luzaturiko zubitzat, zeina, gainera, aipatutako kontinente biotatik geografi akzidente garrantzitsu biz isolatuta bait daga, Pirinioez ipar aldean eta Mediterranioaz hego aldean. Hortaz, trantsizio-gune bat da. Kokapen hain apartari eta gorago inplizituki "geografiko" hitzaren barruan seinalatu diren berezitasun litologiko, orografiko eta klimatikoei gehitu egin behar zaie, bestalde, GERMAINek (1930) Europako mendebalderako kontsideratu zituen hiru taldekamendu malakofaunistikoek Penintsularen gainazalean zehar konfluditzen dutela: taldekamendu septentrionala, taldekamendu atlantikolusitaniarra eta taldekamendu mediterraniarra. Malakofauna septentrionala, zeina estepa errusiarretik lberiar Penintsularaino Euro­ pa asean zehar hedatzen bait da, toki basatsu, heze eta hotzetako espezieez osotua da. Lurralde, 13: 1990 174 K. AL TONAGA • A. 1. PUENTE • C. E. PRIETO SACCHik (1964) adierazi duenez, taldekamendu malakofaunistiko honen banaketaren hegoko limitea,orohar, Ebro haranean kokatu beharda, nahiz eta hain bat espeziek barrera hori gainditzea lortu duten. Beraz, eta Penintsulari dagokiola, multzo bi bereiz daitezke taldekamendu septentrionalean. Alde batetik,ba daude zenbait espezie, erregio pirinia­ rrean bizi izateaz gainera lberiar Sisteman aurkitzen direnak, mendi-zerra horren koloni­ zazioa Euskal Mendietan barrena gertatu bide zelarik. Bigarren multzoan Pirinio eta aldamenetan bizi diren bare-marraskiloak inkludi ditzakegu, bai na, Ebro harana zeharkatu ez dutelarik. GERMAIN (1930) eta CASTILLEJOren (1981) arauera, malakofauna atlantikolusita­ niarra Portugal osotik, Galizia, Zirrinda Kantauriar et a Frantziako mendebaldeko kostalde­ tik Bretainiaraino eta lrla Britaniarren SWraino zabaltzen da, eta CAZIOTen (1915) eritziz, agian Marokotik ere. Ez da ezagun fauna horren jatorria, eta Atlantida populatu zuten espezieak izan litezkeelako ideiaz espekulatu da inoiz (CAZIOT 1915). Ziur aski, oso aintzinekoa eta lurralde epel eta hezetakoa zen fau na baten aztarna izan daiteke, zeina, glaziazio kuaternarioen kausaz, lehen aipatutako erregioen kostaldetara murriztu bait zen, non ez bait zen tenperaturaren beherakada bortizki pairatu itsasoarekiko hurbiltasunaga­ tik. Malakofauna mediterraniarra da espezie-kopururik handiena besarkatzen duena, eragin mediterraniarreko lurraldeak estaltzen bait du lberiar Penintsulako zabalera gehie­ na. Erregio hori oso konplexuada faunistikoki (SACCHI1964), zeren bertan elkarrekin bizi bait dira, batetik, Europatik nahiz ipar Afrikatik iritsiriko espezie mediterraniarrak, bestalde, elementu zirkunmediterraniarrak eta, azkenik, Penintsulako endemikoak ere. Espezie zirkunmediterraniarren artean, batez ere kostaldean bizi direnak gailendu dira; hala ere, sarri askotan et a duten jasankortasun ekologikoaren arauera Penintsulako barne-alderago aurki daitezke. Europatiko elementu mediterraniarrak, bestetik, alde kataluniarretik iritsi dira; malakofauna iparrafrikarra, konkretukiago mintzatuz iberoma­ ghrebiarra (SACCHI1964), Afrikako NWko lurraldeko elementuez osotua da, zeintzuek, edo Mediterranioa barne-itsasoa zeneko garaietan kolonizatu bait zuten hegoalde iberia­ rra, edo aintzineko mazizo betiko-riftarra populatzen zuten, be rau geroago, Mediterranio eta Atlantiko arteko komunikazioa ezartzean, banatu egin zelarik. Azkenik, Penintsulako konplexutasun geografiko handiak, Euro pa eta Afrikatiko espezieakjasotzeko posibilitatea eskeini duan habitat-ugaritasunaz gainera, barrera geografiko asko aurkeztu ditu espezia­ zioa erraztu delarik, eta, batez ere, elementu mediterraniarretatik abiatutako espeziazioa. Deskribatzen ari garen dinamika biogeografiko horretan, Ebro haranak azpimarratua izatea merezi du, oso munta handiko zeregina betetzen bait du: esan denez, ezaugarri erlatiboki europearrak dituen Katalunia eta propioki iberiarra den gainontzeko Penintsula elkarretik bereizten ditu, elementu europearren hegoranzko eta iberiarren iparreranzko pasea galerazten duelarik. Halaber, espezie mediterraniarren Penintsulako NWrako iraganbidea da, Ebro arroak bizi dituen mediterraniartasun-baldintzak dire la eta (eguzkiz­ tapen handia,prezipitazio urria, idortasuna ... ). lberiar Penintsulako textu-inguruan Euskal Herriak garrantzi berezia du, zeren, hedadura geografiko erlatiboki txiki batetan, Europako mendebalderako aipatu diren hiru taldekamendu malakofaunistikoetako ordezkariak aurkitzen bait dira, eta horrez gainera, Euskal Herria bara da fauna horien teflakamendu-gunea. Horrela, bertatik gertatu da Penintsula barneranzko elementu europearren sarrera, Pirinio eta Mendi Kantauriarretan zehar, hain zuzen. Lurralde, 13: 1990 175 K. AL TONAGA - A. 1. PUENTE • C. E. PRIETO ~1 1 Mapatlberiar Penlntsularen ezagumendu malakologlkoaren adibidea 1etik 4raino higromiido-espezie aipaturik bibliografian. • 1 to 4 hygromiid species ched in the bibliography. 5etik Sraino higromlido-espezie aipaturik bibllograflan. e 5 to 8 hygromiid species ched in the bibliography. 9tik 12raino higromiido-espezie aipaturik bibllografian. • 9 to 12 hygromiid specles ched ln the bibliography. 13 baino hlgromiido-espezie geihago aipaturik bbliograflan. • More than 13 hygromiid species cited in the bibliography. Map 1/ Exarrple of the malacological Knowledge of the lverian Peninsula Lurralde, 13: 1990 176 K. AL TONAGA ·A. 1. PUENTE • C. E. PRIETO IBERIAR PENINTSULAKO IKERKETA MALAKOLOGIKOEN EGOERA Hasiera batetan aipatu den egoera hurrengo lerroetan itxuratuko dugu. Horretarako, ikus bedi 1 mapa (PU ENTE 1985). Bertan, lberiar Penintsulako
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  • ZM82 275-280 Gittenbergert.Indd

    ZM82 275-280 Gittenbergert.Indd

    Two problematic, troglophilous gastropods from the Peloponnese, Greece (Gastropoda: Pulmonata: Zonitidae) E. Gitt enberger Gitt enberger, E. Two problematic, troglophilous gastropods from the Peloponnese, Greece (Gastropoda, Pulmonata, Zonitidae). Zool. Med. Leiden 82 (28), 20.vi.2008: 275-280, fi gs 1-4.— ISSN 0024-0672. E. Gitt enberger, National Museum of Natural History 'Naturalis', P.O. Box 9517, NL 2300 RA Leiden, The Netherlands (gitt [email protected]). Key words: Gastropoda; Pulmonata; Oxychilidae; Pristilomatidae; new species; troglodytes; Greece. Two species of pulmonate gastropods are described as new to science although their generic classifi ca- tion remains uncertain since only shells are available for study. The species might be troglodytes. Their shells indicate the presence of subterraneous material. Introduction In 1985 and following years, during several excursions in the eastern part of the Peloponnese, Greece, empty shells of two unknown pulmonate species were found. Their subgeneric or even generic classifi cation is doubtful because of the absence of anatomical data. Probably these species occur in a for humans nearly inaccessible habitat. The snails are most probably troglophilous, if not troglodytic. Their shells may serve as indicators of deposits of subterranean material, which is an argument in favour of naming these species despite the fact that their systematic position remains unclear. Waiting for more data might become a too long wait. Both species were also found in a deposit together with shells of Tsoukatosia liae Gitt enberger, 2000 (Clausili- idae, Serrulininae), a clausiliid species with a glassy transparent shell, also indicative of a subterranean habitat and hitherto known from its type locality only.
  • Appendix ES-11 Alien Snail Control in Nurseries

    Appendix ES-11 Alien Snail Control in Nurseries

    Appendix ES-11 Alien Snail Control in Nurseries Alien snail control in nurseries Report to the Oahu Army Natural Resources Program (OANRP) Prepared by Norine W. Yeung and Robert H. Cowie 30 November 2012 Pacific Biosciences Research Center University of Hawaii 3050 Maile Way, Gilmore 408 Honolulu, Hawaii 96822 Phone: (808) 956 4909/0956 E-mail: [email protected] [email protected] 1 Appendix ES-11 Alien Snail Control in Nurseries BACKGROUND AND PURPOSE From the basic perspective of habitat restoration, non-native species should not be introduced to areas where they are not present. Specifically, non-native snails and slugs (hereafter ‘snails’) may impact both native vegetation (Joe & Daehler 2008) and native snails (Meyer & Cowie 2010a, b, Sugiura et al. 2011) in areas to which they are introduced. Therefore, it is important that plants to be outplanted, as well as equipment and vehicles used to transport plants, are free of non-native snails. This requires appropriate management practices and control measures to be adopted in the nurseries. For some time the OANRP has been planning to outplant native Hawaiian plants on military land on Oahu as part of its ecological restoration efforts. A major problem associated with this effort is that the plants grown in the nurseries for outplanting can become infested with non- native snails (Cowie et al. 2008), some of which, especially as juveniles, are very small (on the order of 1-2 mm). Preliminary discussions and visits to the Schofield baseyard and the nursery at the Nike site took place prior to the implementation of this agreement and the start of the project.
  • Fauna of New Zealand Website Copy 2010, Fnz.Landcareresearch.Co.Nz

    Fauna of New Zealand Website Copy 2010, Fnz.Landcareresearch.Co.Nz

    aua o ew eaa Ko te Aiaga eeke o Aoeaoa IEEAE SYSEMAICS AISOY GOU EESEAIES O ACAE ESEAC ema acae eseac ico Agicuue & Sciece Cee P O o 9 ico ew eaa K Cosy a M-C aiièe acae eseac Mou Ae eseac Cee iae ag 917 Aucka ew eaa EESEAIE O UIESIIES M Emeso eame o Eomoogy & Aima Ecoogy PO o ico Uiesiy ew eaa EESEAIE O MUSEUMS M ama aua Eiome eame Museum o ew eaa e aa ogaewa O o 7 Weigo ew eaa EESEAIE O OESEAS ISIUIOS awece CSIO iisio o Eomoogy GO o 17 Caea Ciy AC 1 Ausaia SEIES EIO AUA O EW EAA M C ua (ecease ue 199 acae eseac Mou Ae eseac Cee iae ag 917 Aucka ew eaa Fauna of New Zealand Ko te Aitanga Pepeke o Aotearoa Number / Nama 38 Naturalised terrestrial Stylommatophora (Mousca Gasooa Gay M ake acae eseac iae ag 317 amio ew eaa 4 Maaaki Whenua Ρ Ε S S ico Caeuy ew eaa 1999 Coyig © acae eseac ew eaa 1999 o a o is wok coee y coyig may e eouce o coie i ay om o y ay meas (gaic eecoic o mecaica icuig oocoyig ecoig aig iomaio eiea sysems o oewise wiou e wie emissio o e uise Caaoguig i uicaio AKE G Μ (Gay Micae 195— auase eesia Syommaooa (Mousca Gasooa / G Μ ake — ico Caeuy Maaaki Weua ess 1999 (aua o ew eaa ISS 111-533 ; o 3 IS -7-93-5 I ie 11 Seies UC 593(931 eae o uIicaio y e seies eio (a comee y eo Cosy usig comue-ase e ocessig ayou scaig a iig a acae eseac M Ae eseac Cee iae ag 917 Aucka ew eaa Māoi summay e y aco uaau Cosuas Weigo uise y Maaaki Weua ess acae eseac O o ico Caeuy Wesie //wwwmwessco/ ie y G i Weigo o coe eoceas eicuaum (ue a eigo oaa (owe (IIusao G M ake oucio o e coou Iaes was ue y e ew eaIa oey oa ue oeies eseac
  • The Canadian Field-Naturalist

    The Canadian Field-Naturalist

    The Canadian Field-Naturalist Tall grass prairie ecosystem management—a gastropod perspective Annegret Nicolai1, 2, *, Robert G. Forsyth3, Melissa Grantham4, and Cary D. Hamel4 1Université Rennes, UMR CNRS 6553 EcoBio, Station Biologique Paimpont, Paimpont 35380 France 2Western University, Department of Biology, 1151 Richmond Street North, London, Ontario N6A 5B7 Canada 3New Brunswick Museum, 277 Douglas Avenue, Saint John, New Brunswick E2K 1E5 Canada 4Nature Conservancy of Canada, Manitoba Region, Suite 200 - 611 Corydon Avenue, Winnipeg, Manitoba R3L 0P3 Canada *Corresponding author: [email protected] Nicolai, A., R.G. Forsyth, M. Grantham, and C.D. Hamel. 2019. Tall grass prairie ecosystem management—a gastropod perspective. Canadian Field-Naturalist 133(4): 313–324. https://doi.org/10.22621/cfn.v133i4.2217 Abstract Less than 5% of the original tall grass prairie in North America remains. A portion of this remnant, composed of wetland, grassland and forest, is protected by the Nature Conservancy of Canada (NCC) in southern Manitoba. This heterogene- ous ecosystem has rich biodiversity; however, gastropods have not been surveyed in Canada’s tall grass prairie. We studied gastropods in Prairie, Wet Meadow, Forest, and Wet Forest habitats of the Manitoba Tall Grass Prairie Preserve that vary with respect to land management practices (prescribed burning, grazing by cattle). Gastropod community composition was unique in the Prairie where mounds of grass litter form permanently moist cavities harbouring aquatic species, while dry-habitat species colonized the upper parts of these mounds. Gastropod communities in Prairie habitats were negatively affected by grazing and burning that occurred in the five years prior to our survey.