GAIA N°15, LlSBOAlLISBON, DEZEMBRO/DECEMBER 1998, pp. 339-353 (ISSN: 0871-5424) THERANGOSPODUS: TRACKWAY EVIDENCE FOR THE WIDESPREAD DISTRIBUTION OF A LATE JURASSIC THEROPOD WITH WELL-PADDED FEET Martin G. LOCKLEY Geology Department, Campus Box 172, University of Colorado at Denver. P.O. Box 173364, DENVER, COLORADO 80217-3364. USA E-mail:
[email protected] Christian A. MEYER Universitat Basel, Geologisch-palaontologisches Institut. Bernoullistrasse, 32, BASEL, CH-4056. SWITZERLAND E-mail:
[email protected] Joaquin J. MORATALLA Unidad de Paleontologia, Departamento Biologia, Universidad Aut6noma de Madrid. CANTOBLANCO 28049, MADRID. SPAIN ABSTRACT: Assemblages of distinctive, medium·sized theropod tracks indicative of animals with well-padded feet are known from large Upper Jurassic samples of well-preserved mate rial from North America and Asia. These tracks, herein named Therangospodus pandemi cus, always reveal a lack of distinct, separate digital pads, regardless of whether they are preserved as casts or molds. We interpret this as consistent ichnological evidence of a fleshy fool. Similar tracks from the ?Upper Jurassic-?Lower Cretaceous of Spain, first na med Therangospodus onca/ensis, are also formally described. Although the fleshy nature of the foot of this trackmaker, makes it's individual digits appear comparable with those of ornithopod trackmakers, the elongate track and asymmetric postero-medial indentation and narrow trackway, indicate that it is probably of theropod an affinity. Tracks of this type provide an instructive lesson for ichnologists and paleontologist in general because they reveal that tracks are a record of flesh on foot bones, and need not necessarily be an accura te reflection of the morphology of foot skeletons.