© Zoological Institute, St.Petersburg, 2003

Type specimens of and described by F.A. Kolenati ()

I.M. Kerzhner

Kerzhner, I.M. 2003. Type specimens of Coreoidea and Pentatomoidea described by F.A. Kolenati (Heteroptera). Zoosystematica Rossica, 12(1): 93-98.

Lectotypes are designated for 23 Kolenati’s and varieties in the Museum of Natu- ral History, Vienna. The synonymies of Rhabdocoris arcuata Kolenati, 1845 with Lepto- corisa acuta (Thunberg, 1783) and of armeniacum Kolenati, 1846 with E. fieberi Fieber, 1837 are restored. The following new synonymies are established: Liorhyssus hyalinus (Fabricius, 1794) = Corizus rubescens Kolenati, 1845; Sehirus luctuosus Mulsant & Rey, 1866, nomen protectum = Cydnus affinis var. ibericus Kolenati, 1846, nomen oblitum; Geotomus elongatus (Herrich-Schaeffer, 1840) = Cydnus caucasicus Kolenati, 1846 (G. ciliatitylus Signoret, 1881 is a good species, not synonymous with C. caucasicus); Eurygaster integriceps Puton, 1881, nomen protectum = E. hottentotta var. obliqua Kolenati, 1846, nomen oblitum.

I.M. Kerzhner, Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, St.Petersburg 199034, Russia. E-mail: [email protected]

Introduction Kolenati published results of study of Cauca- sian , together with some other entomo- F.A. Kolenati (1812-1864), outstanding Czech logical studies, in the series “Meletemata Ento- zoologist, made a trip to Transcaucasia (1842- mologica”. The first five parts of this series were 1844), where he collected an extensive material published separately in St.Petersburg by the Rus- of (about 400 specimens of vertebrates, sian Academy of Sciences in 1845-1846; the next some molluscs and about 32 000 specimens of three parts (the last of them subdivided into 5 insects), many plants (about 12 000 specimens) sections and not finished) appeared as papers in and minerals (Flasar, 1965). The trip was finan- the “Bulletin de la Société impériale des natural- cially supported by the Russian Academy of Sci- istes de Moscou” (1857-1859). Heteroptera were ences. According to a report by Brandt (1845), considered in 3 parts of this series (1845, 1846a, Kolenati donated many of the collected animals 1857). It has long been known that Kolenati de- (47 specimens of mammals, 69 of birds, 48 of scribed or recorded from the Caucasus, along with amphibians, 3 of crustaceans, 12 600 of insects, true Caucasian species of Heteroptera, several 48 of molluscs), including all the unique speci- dozens of species not occurring there. They origi- mens, to the Zoological Museum of the Russian nated from Brazil (collected by Langsdorff), Si- Academy of Sciences, St.Petersburg. He worked beria (Gebler and others), Estonia (Obert), etc., in this museum as assistant during about two or were collected by Kolenati or his friends in years. In St.Petersburg, he made acquaintance environs of St.Petersburg. In some cases, Kolenati with local amateur entomologists and exchanged cited for such species both the true data on the specimens with them (Simashko, 1861). When locality and collector and false data on its col- returning to Prague, Kolenati apparently took lecting in the Caucasus, but in many cases he in- almost all his Caucasian collection with him, at dicated only a false Caucasian record, sometimes least only very few of his Heteroptera (some of with citation of “exact” Caucasian locality and them received with Kuschakewitsch’s collection) host plant. are now found in St.Petersburg. He took also Kolenati’s specimens in the Naturhostorisches some specimens of Heteroptera from the collec- Museum in Wien bear narrow labels with short tion of the St.Petersburg Museum, especially text (one word). They are easily recognizable. those from Brazil and Siberia. Soon after return During my work in this museum, I was able to to Prague, Kolenati has sold his Caucasian col- examine the collection and have found type speci- lection together with specimens taken from mens of most of the species and varieties of St.Petersburg to the Museum of Natural History Coreoidea and Pentatomoidea described by in Vienna (Naturhistorisches Museum in Wien). Kolenati. Their lectotypes are designated below. 94 I.M. Kerzhner: Kolenati’s Coreoidea and Pentatomoidea • ZOOSYST. ROSSICA Vol. 12 Recognition of Kolenati’s specimens in the lehmannii Kolenati, 1857: 496 collection of Zoological Institute, St.Petersburg (“in desertis Kirghisorum, praecipue deserto is more difficult. Specimens labelled “Caucasus” Kisil-Kum … (Lehmann!) (Coll. Acad. sc. and “Kolenati” are found for Jalla anthracina Petropol.)”). Type specimens are found neither (lectotype) and Carpocoris bilunulata (para- in NHMW nor in ZIN. Kolenati cited in syn- lectotype), the latter with a handwritten label. The onymy “Syromastes Corallinus. Mus. Berolin. same handwriting is revealed in labels of exem- Syromastes damietta. Mart.” If a specimen la- plars of Centrocoris variegata listed by Kiritshen- belled “Syromastes corallinus” is present in the ko (1916) as Kolenati’s specimens, the lectotype Museum of Natural History, Humboldt Univer- of Corizus caucasicus, and a specimen of Alydus sity, Berlin, it can be considered a syntype. Cur- tragacanthae. They have the identification label rent name: Cercinthus lehmannii (Kolenati, only, and their belonging to the Kolenati’s speci- 1857). mens is therefore more or less problematic. In Coreus (Palethrocoris) disciger Kolenati, addition, the same handwriting is found in a la- 1845: 49, pl. 7, fig. 4 (“in … Karabagh”). bel “Raphigaster Germarii Kolenati an virens Lectotype (present designation; NHMW): F, Klug” under a male of Brachynema germarii Karabach/ Kolenati/ 4 [figure number]/ Enoplops from Samarkand collected by Lehmann. The lat- disciger Kol. [handwritten by Fieber]. Current ter was possibly identified by Kolenati, who par- name: Enoplops disciger (Kolenati, 1845). ticipated in identification of Lehmann’s insects. Hebecerus persicus Kolenati, 1845: 66, pl. 8, In the paper, information on the type localities fig. 9 (“Karabagh, in confinio Persiae”). from the original descriptions is cited in brack- Lectotype (present designation; NHMW): M, ets. Labels are separated by oblique stroke; those Karabagh/ Kolenati/ Hebeceris Persicus Kole- not stated as handwritten are the printed ones. nati, patria false. Mel. [handwritten]. Current The following abbreviations are used: NHMW, name: Merocoris tristis Perty, 1833. – Note. This Naturhistorisches Museum, Wien; ZIN, Zoologi- is a Neotropical species. cal Institute, St.Petersburg. Coreus (Gonocerus) marginatus var. inermis Kolenati, 1845: 52. No Kolenati’s specimens are Family found. Current name: Coreus marginatus (Lin- naeus, 1758). Centrocoris westwoodii Kolenati, 1845: 46, pl. 7, fig. 1 (“in … provinciae … Karabagh et in Family confinio Persiae”). Lectotype (designated by Dolling, 1977; NHMW): F, Kolenati/ Karabach/ Alydus hirsutus Kolenati, 1845: 64, pl. 8, fig. Westwoodi det. Kolenati [handwritten]/ Lecto- 8 (“prope Elisabethopolin et in provincia Kara- type/ Lectotype Centrocoris westwoodi Kol. det. bagh, nec non in provinciis Caspicis”). Lecto- W.R. Dolling, 1972. Current name: Vilga west- type (present designation; NHMW): M, Cauc. occ. woodii (Kolenati, 1845). – Note. It is a Neotro- / Kolenati/ Alydus hirsutus Kol. [handwritten by pical species. F.X. Fieber made to this species a Fieber]. Current name: Alydus calcaratus (Lin- note in the copy of Kolenatis’s work from the naeus, 1758). library of the Vienna Museum: “Habitat in Alydus tragacanthae Kolenati, 1845: 65, pl. Brasilia, rapacit in Museo imp.” (Distributed in 8, fig. 7 (“in Astragalo Caucasico (Pall.) etiam Brazil, stolen from the Imperial Museum [in Transcaucasiae et in confinio Turciae, prope St.Petersburg]). Dolling (1977) incorrectly stated Alexandrapol”). Lectotype (present designation; that the LT is in ZIN; it is actually in NHMW. NHMW): M, Karabach / Kolenati/ 7 [figure num- Centrocoris variegata Kolenati, 1845: 46, pl. ber]/ Camptopus tragacanthae [handwritten by 7, fig. 2 (“in … provinciae Elisabethopoleos”). Fieber]. Paralectotype (NHMW): M, Kolenati/ Lectotype (present designation; NHMW): F, Karabach. Current name: Camptopus traga- Elisabethop/ Kolenati. Current name: C. va- canthae (Kolenati, 1845). – Note. In the ZIN riegatus Kolenati, 1845. – Note. The lectotype collection, there is a female labelled “Alydus Tra- is clearly the specimen figured. In ZIN, there are gacanthae Kolenati N 20 fig. 7”; the handwrit- 3 specimens (on one pin) labelled “Centrocoris ing is the same as in the lectotype of Corizus Klti variegata” [handwritten]; they are listed by caucasicus and specimens of Centrocoris va- Kiritshenko (1916) as Kolenati’s specimens, but riegatus ascribed to Kolenati by Kiritshenko I cannot confirm or refute this attribution. (1916). Centrocoris pallescens Kolenati, 1845: 47, pl. Rhabdocoris arcuata Kolenati, 1845: 68, pl. 7, fig. 3 (“in … provinciae Karabagh”). Lectotype 8, fig. 11 (“in … montis Ssalwat, Caucasi (present designation; NHMW): F, Karabach/ Lhesgici”). Lectotype (present designation; Kolenati/ 3 [figure number]. Current name: C. NHMW): M (genitalia examined), Ssalwat/ spiniger (Fabricius, 1781). Kolenati/ Rabdocoris [sic] Arcuata [handwrit- ZOOSYST. ROSSICA Vol. 12 • I.M. Kerzhner: Kolenati’s Coreoidea and Pentatomoidea 95 ten]. Current name: Leptocorisa acuta (Thunberg, naeus, 1761). In NHMW, there is a female of 1783) = Rhabdocoris arcuata Kolenati, 1845, Sehirus luctuosus Mulsant & Rey, 1866 labelled syn. rest. (not syn. of L. oratoria (Fabricius, “Karabach”. It fits well the original description 1794)). – Note. Kolenati (1846b: 170) placed of var. ibericus, except the differing locality in Rhabdocoris in synonymy with Leptocorisa the label. Because of this contradiction, I abstain Latreille, 1829, and the species was subsequently from designation of lectotype. However, the origi- synonymized with L. acuta (Thunberg, 1783). nal description of var. ibericus (large, strongly However, Ahmad (1965) placed Rh. arcuata in shining body, with white membrane) fits S. synonymy with the Oriental species L. oratoria luctuosus, and the following synonymy should (Fabricius, 1794). Ahmad (1965: 94) regarded as be established: Sehirus luctuosus Mulsant & Rey, holotype of Rh. arcuata a male from the ZIN 1866, nomen protectum = Cydnus affinis var. collection labelled “k. A. Kushakevicha” [in ibericus Kolenati, 1846, syn. n., nomen oblitum. Cyrillic characters; = collection of A. Kusha- – Note. Kolenati’s name has not been used as kevitsh] and “Rhabdocoris Klti armatus Klti Mel. valid after 1899. The name S. luctuosus was used II, p. 67” [handwritten]. This specimen originates in many dozens of publications in the last 50 not from Kolenati’s collection and is markedly years. A selected list of 25 works (for full refer- wider and darker than that figured by Kolenati; ences see Aukema & Rieger, 1995) is given be- certainly, it does not belong to the type series. low to fulfill the requirements of Art. 23.9 of the The specimen from NHMW fits well the pub- Code: Andersen & Gaun, 1974; Aukema, 1989; lished figure and is apparently the specimen from Cmoluchowa & Mielewczyk, 1990; Coulianos & which Kolenati’s description was made. L. acuta Ossiannilsson, 1976; Gidayatov, 1982; Günther is a species widely distributed in Oriental and & Schuster, 1990; Halászfy, 1959; Hoberlandt, Australian regions and the SE Palaearctic (China, 1956a, 1959a, 1977a; Huldén & Heikinheimo, Japan), but not occurring in the Caucasus. 1989; Josifov, 1981; Kerzhner & Jaczewski, 1964; Kiritshenko, 1964; Kis, 1984; Kloet & Family RHOPALIDAE Hincks, 1964; Lammes & Rinne, 1990; Lin- navuori, 1967; Lis, 1989a; Putshkov, 1961, 1965; Corizus caucasicus Kolenati, 1845: 59, pl. 7, Servadei, 1967; Southwood & Leston, 1959; fig. 5 (“in … Caucasi”). Lectotype (designated Wagner, 1961a, 1966a). by Putshkov & Kerzhner, 1983; ZIN): M, Cydnus caucasicus Kolenati, 1846a: 70, pl. caucasicus Kolenati N. 19. tab. 4. fig. 5 [hand- 15, fig. 34 (“montis Kasbek, … ad pagum written]. Paralectotype (NHMW): M, Cauc/ osseticum Görgeti”). Lectotype (present designa- Kolenati. Current name: Maccevethus errans tion; NHMW): F, Kasbek/ Kolenati/ caucasicus caucasicus (Kolenati, 1845). – Note. There is Kol. [handwritten]. Current name: Geotomus some doubt about belonging of the lectotype to elongatus (Herrich-Schaeffer, 1840) = Cydnus the type series. The handwriting of the label un- caucasicus Kolenati, 1846, syn. n. (not senior der the lectotype is identical to that of 3 speci- synonym of Geotomus ciliatitylus Signoret, mens of cited by Kiri- 1881). – Note. The lectotype fits well the illus- tshenko (1916) as Kolenati’s specimens. As fur- tration, except the setation of head (there are no ther evidence is needed to confirm or refute the setae or setiferous pores on clypeus) and sides of authenticity of the lectotype, I refrain from hemelytra (there are only 2 setiferous pores on changes. Anyhow, the lectotype and the paralecto- each hemelytron). Owing to the inexactly illus- type (which is obviously a Kolenati’s specimen) trated setation, Geotomus ciliatitylus Signoret, are conspecific. 1881, sp. dist. (type locality: Teheran) was placed Corizus rubescens Kolenati, 1845: 59, pl. 7, fig. in synonymy with G. caucasicus by Horváth 6 (“in … provinciae Karabagh”). Lectotype (present (1918: 323) and treated subsequently as its jun- designation; NHMW): M, Caucasus/ Kolenati/ 6 ior synonym or a subspecies. This synonymy is [figure number]. Current name: Liorhyssus hyalinus erroneous. The lectotype of Cydnus caucasicus (Fabricius, 1794) = Corizus rubescens Kolenati, is not conspecific with G. ciliatitylus, and G. 1845, syn. n. – Note. C. rubescens was hitherto ciliatitylus even does not occur in the area from placed in synonymy with Rhopalus parumpuntatus which C. caucasicus was described: confirmed (Schilling, 1829), Rh. rufus (Schilling, 1829), or records of G. ciliatitylus in Transcaucasia are only Rh. lividus Fieber, 1861. from Armenia and Azerbaijan. Cydnus caucasicus var. rufulus Kolenati, Family CYDNIDAE 1846a: 70 (apparently from the same locality as f. typica). No type material found either in Vi- Cydnus affinis var. ibericus Kolenati, 1846a: enna or in St.Petersburg. Probably, a teneral speci- 69 (“Iberiae”). Cydnus affinis Herrich-Schaeffer, men of the preceding species. – Note. Kerzhner 1835 is a junior synonym of Sehirus morio (Lin- (1972) speculated about a female of Byrsinus sp. 96 I.M. Kerzhner: Kolenati’s Coreoidea and Pentatomoidea • ZOOSYST. ROSSICA Vol. 12 in ZIN labelled “Caucasus” and “Caucasicus name to Leach, but no such a name is found in Kolenati” [handwritten] that it is possibly a publications by Leach, and all subsequent authors syntype of C. caucasicus var. rufulus. This sup- credited the name to Kolenati. The name was hith- position is not substantiated, as the second label erto considered to be a synonym of E. austriacus is handwritten by E. Ménétriés, not by Kolenati. (Schrank, 1776). There are 9 Kolenati’s speci- mens of Eurygaster in the NHMW collection, all Family SCUTELLERIDAE belong to E. integriceps. The specimen desig- nated as lectotype is markedly darker than other Odontoscelis fuliginosa var. iberica Kolenati, specimens and with well developed colour pat- 1846a: 7 (“in … Iberiae”). Lectotype (present tern, thus clearly fitting the original description. designation; NHMW): F, Kolenati. Current The name obliqua has not been used after 1899 name: Odontoscelis fuliginosa (Linnaeus, 1761). as valid name, even for a variety. The name E. – Note. There are four Kolenati’s specimens of integriceps refers to an important pest of cereals O. fuliginosa in the NHMW collection, one of and is used in many hundreds of books and pa- them (female) belongs to f. typica, and other pers. In accordance with Art. 23.9 of the Code, three, to varieties described by Kolenati (their precedence is given to the name E. integriceps identity is determined from original descriptions). Puton. The name “Iberia” was used in Latin for both current Georgia and current Iberian Peninsula. Family Kolenati definitely had in mind the first sense, but Kirkaldy (1909: 265) and Göllner-Scheiding Odontotarsus notoceras Kolenati, 1857: 497, (1986) erroneously accepted that “Iberia” meant pl. 3, fig. 4a, b, c (“in Tauria et Caucaso (Kole- Spain. Göllner-Scheiding (1986) did not found nati!) (Mus. Caes. Viennense et Mus. auctoris”). the type, designated a neotype from Portugal, and Lectotype (present designation; NHMW): M, used the name Odontoscelis iberica Kolenati for Kolenati/ Odontotarsus Kol. Notoceras Kolen. a species from the Iberian Peninsula. Later Cauc. [handwritten]/ virescens [handwritten] det. (Göllner-Scheiding, 1987), she recognized the Reuter/ virescens H.S. sec. Reut. [handwritten]. mistake and proposed the name O. hispanica for Current name: Tarisa virescens (Herrich-Schaef- the species discussed. As the original type mate- fer, 1851). rial is rediscovered, the neotype is set aside Eurydema oleraceum var. angularis (Code, Art. 75.8). Kolenati, 1846a: 23 (“prope Alexandropolin … Odontoscelis fuliginosa var. pallasii Kolenati, Petropoli unicum etiam exemplar inveni”). Lecto- 1846a: 7 (“in … Caucasi Lhesgici et prope mare type (present designation; NHMW): F, Petropol/ Caspium”). Lectotype (present designation; oleracea var. angularis Kolen. Petersb. [hand- NHMW): M (genitalia examined), Cauc/ Kole- written]. Current name: Eurydema oleracea (Lin- nati. Current name: Odontoscelis fuliginosa (Lin- naeus, 1758). – Note. In the lectotype, the abdo- naeus, 1761). men ventrally is dark brown, not yellow with dark Odontoscelis fuliginosa var. caucasica Kole- spots as stated in the original description. nati, 1846a: 7 (“in Caucasi occidentalis”). Lecto- Eurydema gebleri Kolenati, 1846a: 23, pl. 15, type (present designation; NHMW): F, Cauca- fig. 30 (“in provinciis Caspicis Scheki, Schirwan sus/ Kolenati. Current name: Odontoscelis et Baku. Etiam a Do. Gebler e Dauria”). Lectotype fuliginosa (Linnaeus, 1761). (present designation; NHMW): M, Nertschinsk/ Odontoscelis dorsalis Kolenati, 1857: 498 (“in Stschukin. Paralectotype (NHMW): F, Barnaul. desertis Kirghisorum et in Turcomania. (Leh- Current name: Eurydema gebleri Kolenati, 1846. mann!) (Mus. Caes. Acad. Petropol.)”). No types – Note. This species is common in Siberia (from are found either in ZIN or NHMW. Current name: which obviously all the Kolenati’s material origi- Odontoscelis dorsalis (Fabricius, 1798). – Note. nated), but does not occur in the Caucasus. The Kolenati’s specific name is a junior homonym specimen from Barnaul was apparently collected and synonym of Fabricius’s name. Much prob- by Gebler. The lectotype is the specimen figured ably the author’s name “Kolenati” was placed by Kolenati. In the paralectotype, the hind black after the species name in error. spots on the pronotum are fused; it was described Eurygaster hottentotta var. obliqua Kolenati, by Kolenati as a variety. 1846a: 12 (“cum precedentis [i.e. in Trans- Eurydema ornatum var. ventralis Kolenati, caucasia]”). Lectotype (present designation; 1846a: 26 (“in Caucaso et Transcaucasia”). The NHMW): M (genitalia examined), Transcauc/ lectotype will be designated by V.V. Derzhansky. Kolenati. Current name: Eurygaster integriceps Current name: Eurydema ventralis Kolenati, 1846. Puton, 1881, nomen protectum = E. hottentotta Eurydema lhesgicum Kolenati, 1846a: 28, pl. var. obliqua Kolenati, 1846, syn. n., nomen 15, fig. 31 (“prope montem Ssalwat et urbem oblitum. – Note. Kolenati attributed the varietal Nucha”). Lectotype (present designation; ZOOSYST. ROSSICA Vol. 12 • I.M. Kerzhner: Kolenati’s Coreoidea and Pentatomoidea 97 NHMW): M, Ssalwat/ Kolenati. Paralectotypes designation; NHMW): M, Elisabethop/ Kolenati. (NHMW): M, Transcauc.; M, Gebler; F, Sibir./ Paralectotype (NHMW): M, Elisabeth/ Kolenati. Gebler; M, Petropol/ Kolenati. Current name: Current name: varia (Fabricius, 1787). Eurydema dominulus (Scopoli, 1763). – Note. – Note. The lectotype is the specimen figured by The lectotype is the specimen figured by Kolenati. Kolenati. In the paralectotype, antennae are in- This species does not occur in the Caucasus; ap- complete and antennal segment 1 is reddish. parently all Kolenati’s specimens originated from Carpocoris bilunulata var. nubica Kolenati, Siberia and environs of St.Petersburg. 1846a: 51, pl. 16, fig. 37 (“A Do. Awerow e Nubia”). Eurydema armeniacum Kolenati, 1846a: 28, Lectotype (present designation; NHMW): M, Nubia. pl. 15, fig. 32 (“prope Gumri (Alexandropolin) Current name: Codophila varia (Fabricius, 1787). Armeniae montuosae, in confinio Turciae et in – Note. The lectotype is the specimen figured by districtu Pambakiano”). Lectotype (present des- Kolenati. There are further 3 specimens labelled ignation; NHMW): F, Elisabeth./ Kolenati/ “Nubia” in NHMW; they belong to Codophila Strachia dominula Harr. Fieberi Schl. armeniaca maculicollis (Dallas, 1851) and do not fit the origi- Kol. [handwritten by Fieber]. Paralectotype nal description of C. bilunulata var. nubica. (NHMW): F, Karabach/ Kolenati/ Armeniacum Raphigaster germarii Kolenati, 1846a: 56, pl. Kolen. [handwritten]. Current name: Eurydema 16, fig. 38 (“in confiniis Persiae”). Lectotype fieberi Fieber, 1837 = E. armeniacum Kolenati, (present designation; NHMW): M, Karabach/ 1846, syn. rest. – Note. The above synonymy Kolenati. Current name: Brachynema germarii was long and universally accepted. Wagner (1959, (Kolenati, 1846). 1961, 1962, 1968) without substantiation listed E. armeniacum as a species separate from E. Acknowledgements fieberi in faunal lists for Turkey, Iran and Mac- I am very thankful to Dr. H. Zettel (Naturhistorisches edonia, but examination of large series and the Museum in Wien) for the opportunity to work with types of E. armeniacum did not reveal any sup- Kolenati’s collection in Vienna and for sending me some port for Wagner’s opinion. Kolenati’s type specimens for further study. Eurydema pictum var. conjuncta Kolenati, 1846a: 30 (“Duo exemplaria accepi e districtu References Eriwan a consorti meo Do. Fricke”). Type speci- mens are not found. Current name: Eurydema Ahmad, I. 1965. The Leptocorisinae (Heteroptera: Aly- ornata (Linnaeus, 1758). didae) of the world. Bull. Brit. Mus. (natur. Hist.) (Entomol.), Suppl. 5: 1-156. Jalla herrichii Kolenati, 1846a: 35, pl. 15, fig. Aukema, B. & Rieger, Chr. 1995. Catalogue of the Hete- 33 (“in provincia Elisabethopol”). Lectotype (des- roptera of the Palaearctic Region, 1. Netherlands ignated by Gidayatov, 1975; ZIN): M, Cauc/ Entomol. Soc. xxvi + 222 p. Kolenati. Paralectotypes (NHMW): F, Kolenati/ Brandt, J.F. 1845. Sur la récolte zoologique de voyage Elisabethop.; F, Elisabethop./ Kolenati; F, Kole- de M. Kolenati. Bull. Cl. phys.-math. Acad. Imp. Sci. nati. All paralectotypes with additional label St.Pétersbourg, 4: 172-174. Dolling, W.R. 1977. A revision of the Neotropical “Herrichii [handwritten by Horváth] det. Hor- Vilga Stål (: Coreidae). Syst. Entomol., 2: váth”. Current name: Agatharchus herrichii (Ko- 27-44. lenati, 1846). Flasar, I. 1965. Prof. Dr. Friedrich Anton Kolenati. Práce Carpocoris pusio Kolenati, 1846a: 48, pl. 16, odboru přirodních vìd Vlastivědného ústavu v Olo- fig. 39 (“in … provinciae Karabagh”). Lectotype movci: 1-28. Gidayatov, D.A. 1975. The synonymy of some bugs (He- (present designation; NHMW): M, Karabach/ teroptera) from the Caucasus. Entomol. Obozr., 54(3): Kolenati/ Typus P. pusio Kolen. [handwritten by 589-590. (In Russian). Tamanini]/ Codophila (Antheminia) pusio Kol. Göllner-Scheiding, U. 1986. Revision der Gattung Odon- [handwritten by Tamanini] det. L. Tamanini 1960. toscelis Laporte de Castelnau, 1832 (Heteroptera, Current name: Antheminia pusio (Kolenati, 1846). Scutelleridae). Deut. entomol. Z. (N. F.), 33: 95-127. Göllner-Scheiding, U. 1987. Ergänzung und Korrektur Carpocoris bilunulata Kolenati, 1846a: 51, zu der Revision der Gattung Odontoscelis Laporte pl. 16, fig. 35 (“in … provinciae Elisabethopol”). de Castelnau, 1832 (Heteroptera, Scutelleridae). Deut. Lectotype (present designation; NHMW): M, entomol. Z. (N. F.), 34: 217-218. Elisabethop/ Kolenati. Paralectotype (ZIN): F, Horváth, G. 1918. Ad cognitionem faunae hemipterorum Kolenati/ Caucasus/ bilunulata Kolenati [hand- balcanicae. Ann. hist.-natur. Mus. nat. Hung., 16: 321-340. written; same handwriting as in the label of the Kerzhner, I.M. 1972. New and little-known Heteroptera lectotype of Corizus caucasicus]. Current name: from Mongolia and adjacent regions of the USSR, I. Codophila varia (Fabricius, 1787). – Note. The Nasekomye Mongolii, 1: 349-379. Leningrad: Nauka. lectotype is the specimen figured by Kolenati. (In Russian). Carpocoris bilunulata var. persica Kolenati, Kiritshenko, A.N. 1916. Coreidae: . Fauna Ros- 1846a: 51, pl. 16, fig. 36 (“in provincia Karabagh sii i sopredel’nykh stran, Nasekomye poluzhestko- krylye (Insecta, Hemiptera), 6(2). Petrograd. v + 395 et Persia septemtrionali”). Lectotype (present p. (In Russian with Latin diagnoses). 98 I.M. Kerzhner: Kolenati’s Coreoidea and Pentatomoidea • ZOOSYST. ROSSICA Vol. 12 Kirkaldy, G.W. 1909. Catalogue of the Hemiptera (Hete- Putshkov, V.G. & Kerzhner, I.M. 1983. On the nomen- roptera). 1, Cimicidae. Berlin: Dames. xl + 392 p. clature of bugs of the family Rhopalidae (Heteroptera) Kolenati, F.A. 1845. Meletemata Entomologica II. Hemi- of the USSR. Entomol. Obozr., 62(1): 80-82. (In ptera Caucasi Tessaratomidae monographice dispo- Russian). sitae. Petropolis: Acad. Sci. 132 p. Simashko, Yu. 1861. On foundation of the Russian En- Kolenati, F.A. 1846a. Meletemata Entomologica IV. tomological Society. Horae Soc. entomol. Ross., 1: Hemiptera Caucasi Pentatomidae monographice 45-53. (In Russian). dispositae. Petropolis: Acad. Sci. ii + 72 p. Wagner, E. 1959. Beitrag zur Heteropterenfauna Anatol- Kolenati, F.A. 1846b. Meletemata Entomologica V. In- iens. Z. angew. Entomol., 44(1): 102-113. secta Caucasi. Coleoptera, Dermaptera, Lepidopte- Wagner, E. 1961. Beitrag zur Heteropteren-Fauna von ra, Neuroptera, Mutillidae, Aphaniptera, Anoplura. Iran. Ergebnisse der Österreichischen Iran-Expedi- Petropolis: Acad. Sci. iii + 170 p. tionen 1949/50 und 1956. Anz. math.-naturwiss. Kl. Kolenati, F.A. 1857. Meletemata Entomologica [VI]. Österr. Akad. Wiss., 1961(10): 156-164. Hemipterorum Heteropterorum Caucasi Harpago- Wagner, E. 1962. 2. Beitrag zur Heteropteren-Fauna Ma- corisae monographice dispositae. Additamenta ad cedoniens (Hem. Het.). Fragm. Balcan., 4: 115-122. Meletematum Entomologicorum Fasciculum II-dum. Wagner, E. 1968. Contribution a la faune de l’Iran. 7. Additamenta ad Meletematum Entomologicorum Hémiptères Hétéroptères (pro parte). Ann. Soc. ento- Fasciculum IV-tum. Additamenta ad Meletematum mol. France (N. S.), 4: 437-453. Entomologicorum Fasciculum V-tum. Bull. Soc. imp. Natur. Moscou, 29(4), 1856: 419-502. Received 20 August 2003