The Non-Isocrinid Crinoids of the Michelotti Collection
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Bollettino della Società Paleontologica Italiana, 44 (3), 2005, 211-218. Modena, 30 novembre 2005211 The non-isocrinid crinoids of the Michelotti Collection Riccardo MANNI R. Manni, Dipartimento di Scienze della Terra, Università degli Studi di Roma “La Sapienza”, P.le A. Moro 5, I-00185 Roma, Italy; [email protected] KEY WORDS - Crinoids, Michelotti, Eocene, Miocene, North Italy. ABSTRACT - For the first time Tertiary non-isocrinid crinoids of the Michelotti Collection are described. In particular, Conocrinus pyriformis, C. thorenti, C. seguenzai, Holopus spileccense, and Cyathidium gastaldii are recognized. The material comes from Eocene and Miocene sediments of Piedmont and Veneto outcrops (North Italy). RIASSUNTO - [I crinoidi non isocrinidi della Collezione Michelotti] - Sono descritti i crinoidi non isocrinidi della Collezione Michelotti. Tale collezione fu donata dallo stesso Michelotti al Museo di Paleontologia dell’Università “La Sapienza” di Roma nel 1879. Comprendeva oltre 20.000 esemplari che erano stati rinvenuti in sedimenti terziari (Eocene e Miocene) del nord Italia (soprattutto Piemonte e Veneto). Purtroppo, durante la II guerra mondiale (19 luglio 1943), una bomba colpì in pieno un salone del Museo arrecando notevoli danni sia alle strutture che ai fossili. Numerosi furono i reperti di Michelotti che andarono perduti e solo ora si sta provvedendo al riordino del materiale superstite. I crinoidi rappresentano solo una minima parte di questa collezione. In particolare i crinoidi non isocrinidi sono in pratica inediti; tra essi sono state riconosciute le seguenti specie: Conocrinus pyriformis, C. thorenti e C. seguenzai, Holopus spileccense e Cyathidium gastaldii. INTRODUCTION (S. Vito, 9), C. zignoi (S. Vito, 5), Bourgueticrinus thorenti (S. Vito, 42); from Middle Miocene: In 1879 Giovanni Michelotti gave to Rome Micropocrinus gastaldii (Colli di Torino, 1), Conocrinus University his palaeontological collection. At that time seguenzai (Gavi, 30), Bourgueticrinus italicus this collection was constituted by over 20,000 fossil (Serravalle, 20); and from Pliocene: Bourgueticrinus specimens that were collected by Michelotti from italicus (Pienza, 6). These localities are all near Turin several Tertiary outcrops of North Italy. (Piedmont), except Bolca, near Verona (Veneto). This collection is important for Tertiary corals, but Besides, among the specimens of this collection here also for molluscs. Besides, from the Michelotti published, there is a cup of Holopus without original Collection register, it is evident that Michelotti collected label and not cited in the original Michelotti register. and classified also about 200 crinoid specimens. Michelotti recorded all the columnals as Michelotti described or cited (1847, 1861) only some Bourgueticrinus. Rasmussen (1978) underlines that in crinoid specimens (e.g. Isocrinus gastaldii (Michelotti, the crinoid literature “indeterminable columnals of 1847)). Tertiary Bathycrinidae have been recorded under Unfortunately, during the second World War (1943, different generic name (Rhizocrinus, Conocrinus and 19th July) a bomb hit a room of the Museum of Bourgueticrinus)”. For that reason and because it is Paleontology of “La Sapienza” University of Rome very difficult to assign a columnal to a cup, considering (MPURLS). The damages were enormous and also the that bathycrinids have different types of columnals in Michelotti Collection was damaged: several specimens a same stem, only cups are considered in the present were destroyed and many labels were displaced or paper. burned. In the frame of rearrangement of the historical collections of this Museum, the crinoid specimens of the Michelotti Collection were re-examined. The Subclass ARTICULATA Zittel, 1879 specimens of this collection are easily recognizable Order BOURGUETICRINIDA Sieverts-Doreck, 1953 because most of the specimens, as well as the original Family BATHYCRINIDAE Bather, 1899 labels, are still ticked on wooden base (Pl. 1, figs. 14- 17). Small crinoids characterized by five high basals In this paper only the cups of non-isocrinids species superposed by shorter radials. These plates can be will be described. completely or partially fused. Sometimes the radials can number four, five, six or seven. The columnals (Pl. 1, figs. 11-12, 15-16), with SYSTEMATICS different morphology in the same stem, are characterized by wide axial canals. In proximal Michelotti recognized the following non isocrinid columnals the articulation is synostosial; in others it is crinoid species from Eocene sediments: Conocrinus synarthrial, with variable orientation of the fulcral ridge. thorenti (locality: Bolca; 15 specimens), C. pyriformis The stem was attached by cirri. ISSN 0375-7633 N4 211 09/01/06 16 00 212 Bollettino della Società Paleontologica Italiana, 44 (3), 2005 Bathycrinids are known from Maastrichtian to Material - Michelotti collected nine cups. At Recent. There are six genera: Bathycrinus Thomson, MPURLS are stored only three specimens labelled as 1872, Conocrinus d’Orbigny, 1850, Democrinus Perrier, MPURLS 2536. 1883, Dunnicrinus Moore, 1967, Monachocrinus Clark, 1917, and Rhizocrinus Sars, 1868. Description - The cups are stout and pear-shaped. The basals are high and stout, the radials low and stout. The external surface is smooth. The sutures rarely are Genus Conocrinus d’Orbigny, 1850 evident. The interradial projections, low, are developed overall inside the central cavity. The radial facets are Type species - Conocrinus thorenti (d’Archiac, wide but moderately narrow. On each facet a fulcral 1846). ridge separates the aboral surface from the adoral one. The adoral surface is wider than aboral one. The aboral Diagnosis - Small cups with five stout basal plates surface, a little outwardly sloping, is characterized by and five stout radial plates. The basals are very high a narrow half moon-shaped ligament fossa; the ligament while the radials are low. Sutures between the plates pit is not too evident. The adoral surface, inwardly may be indistinct. The basals can be fused and can sloping, is characterized by two wide fossae. The axial cover partially the radials. Radial facets are relatively canal is placed, centrally, above the fulcral ridge. The small; they are placed generally between low interradial underside of the cup, very small and circular, is entirely projections. The central cavity is shallow and occupied by a shallow cavity. In this cavity is the moderately large. The underside of the cups is entirely articular surface for the stem. occupied by the articular surface for the stem. The columnals have large opening of axial canal. Remarks - The characters of this species are very clear only in two specimens, because one specimen is Remarks - The main characters of this genus are: deformed: probably it is not a C. pyriformis, because it the sutures among the plates little evident, the very high seems too oblong. Besides, due to poor preservation, basals, the radial facets sunken between the interradial its radial facets are not evident. The specimen figured projections, and the shallow ventral cavity. Some by d’Archiac as Bourgueticrinus thorenti in pl. 9, fig. Conocrinus can resemble in general external morphology 27 is here considered a C. pyriformis for its some Democrinus; but Conocrinus differs in the stout morphological characters. Besides C. handiaensis Roux, basals and stout radial plates. Numerous fossil and living 1978 (Lutetian of Biarritz, France), is considered species of Conocrinus are known. synonym of C. pyriformis due to the evident resemblance. Ecology - Living species of Conocrinus have a wide depth range. In the Gascogne Gulf, specimens of Stratigraphic distribution - The material of Conocrinus cherbonnieri are found at a minimum depth Michelotti comes from the Eocene sediments of San of -330 m (Roux & Plaziat, 1978) and specimens of Vito (Turin). C. cabiochi are found between -1975 m and -2070 m Cups of C. pyriformis are widespread in the Tertiary (Roux & Montenat, 1977); in Indonesia specimens of sediments of Italy: Meneghini (1976) cited specimens C. globularis are found at depth of -345 m (Roux & from Mossano (Berici Mts.); Rovasenda (1892) cited Montenat, 1977). specimens from the Bartonian sediments of Gassino Stratigraphic distribution - Fossil specimens of Conocrinus are known from Eocene and Miocene sediments of France, Hungary, Italy, Romania, Spain, Switzerland, Croatia (Istria) (Biese & Sieverts-Doreck, 1939). Conocrinus pyriformis (Münster, 1826-33) (Fig. 1; Pl. 1, figs. 1-3, 17) 1826-33 Eugeniacrinites pyriformis MÜNSTER, p.165, pl. 50, fig. 6. 1850 Bourgueticrinus thorenti d’Archiac - D’ARCHIAC, p. 418, pl. 9, fig. 27. 1876 Conocrinus pyriformis (Münster, 1826-33) - MENEGHINI, p. 46-50. 1876-80 Conocrinus (Rhizocrinus) pyriformis (Münster, 1826- 33) - ZITTEL, p. 392, fig. 281. 1924 Rhizocrinus pyriformis (Münster, 1826-33) - ZITTEL, p. 198, fig. 320. 1929 Conocrinus pyriformis (Münster, 1826-33) - PASOTTI, p. 72-75, pl. 1, figs. 1-2, 5, 7-8, 17, 20. Fig. 1 - Conocrinus pyriformis (Münster, 1826-33). a) lateral 1978 Conocrinus handiaensis ROUX, p. 265, fig. 1F. view; b) ventral view; c) dorsal view (all x 5). N4 212 09/01/06 16 00 R. Manni - Crinoids of the Michelotti Collection 213 (Turin); Zittel (1876-80) from the Eocene of Verona; The Jaekel specimen figured by Rasmussen (1978) Pasotti (1929) from the Eocene of Gassino and from as “old specimen” is here considered to pertain to C. the Palaeogene of Verona. At last, Albus (1930) cited veronensis (Jaekel,