DOCSLIB.ORG

Skamienia³oœci Antiarcha (Vertebrata, Placodermi) W Dewonie Gór Œwiêtokrzyskich

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Skamienia³oœci Antiarcha (Vertebrata, Placodermi) W Dewonie Gór Œwiêtokrzyskich Przegl¹d Geologiczny, vol. 54, nr 7, 2006 Skamienia³oœci Antiarcha (Vertebrata, Placodermi) w dewonie Gór Œwiêtokrzyskich Piotr Szrek* Fossils of Antiarcha (Vertebrata, Placodermi) in the Devonian of the Holy Cross Mountains (central Poland). Prz. Geol., 54: 610–614. Summary. Thepaper indicates a similarity between remains of various Palaeozoic fishes thus hindering identifi- cation of fossil of Antiarcha — rare placoderms in the Holy Cross Mountains. A comparison of the vertebrate fossil assemblage from Lode Famennian delta deposits in Latvia with that from the Lower Devonian “Placoderm Sand- stone” of the Pod³azie hill in the Holy Cross Mountains (Poland) shows their similarity. It indicates a fluvial (?deltaic) origin of the “Placoderm Sandstone” from Pod³azie. Additionally, fragmentary antiarch plates are revealed from the Frasnian of the Wietrznia Quarry in Kielce (Holy Cross Mountains). Key words: Placodermi, Antiarcha, “Placoderm Sandstone”, Devonian, Holy Cross Mountains Celem niniejszego artyku³u jest przedstawienie aktual- Dolnodewoñskie skamienia³oœci Antiarcha nego stanu badañ nad rybami pancernymi, szczególnie z rzêdu Antiarcha, z dewonu Gór Œwiêtokrzyskich. Poruszo- Stwierdzona obecnoœæ przedstawicieli antiarchów w ne s¹ g³ównie zagadnienia zwi¹zane z rozpoznawaniem „piaskowcach plakodermowych” (Szrek, 2003a) zgodna skamienia³oœci tej grupy i wynikaj¹ce z tego problemy. Ma jest z genez¹ tych ska³. Œrodowisko ich powstawania to du¿e znaczenie, gdy¿ publikacje na temat tego rzêdu z zosta³o okreœlone przez Tarnowsk¹ (1976) jako zmienne: Gór Œwiêtokrzyskich, s¹ bardzo nieliczne, a stwierdzenie od strefy przybrze¿nej p³ytkiego morza do œrodowiska pla- w osadzie skamienia³oœci tej grupy jest bardzo pomocne ¿owego. Ponadto zaznacza siê tu ewidentnie wp³yw œrodo- przy konstruowaniu wniosków facjalnych. Obecnoœæ ska- wiska rzecznego, zw³aszcza w przypadku œrodkowego mienia³oœci antiarchów pozwala bowiem na okreœlenie kompleksu piaskowcowego (formacja piaskowców z Porê- pewnych w³aœciwoœci, jakimi ska³a w trakcie powstawania by; vide Tarnowska 1976), gdzie „piaskowce plakodermo- siê charakteryzowa³a, m.in. zawartoœci substancji orga- we” (sensu Szrek 2003a) wystêpuj¹ najczêœciej. Brak nicznej, która stanowi³a pokarm antiarchów (Denison, jakiejkolwiek fauny bezkrêgowców i charakterystyczny 1978; Long, 1996; Stensiö, 1931, 1948). sk³ad taksonomiczny fauny krêgowców z „piaskowców W dewonie Gór Œwiêtokrzyskich dotychczas stwier- plakodermowych” najlepiej wskazuje na wp³yw œrodowi- dzono cztery rzêdy plakodermów: Arthrodira (artrodiry), ska rzecznego. Bardzo dobrym odniesieniem mo¿e byæ tu Antiarcha (antiarchy), Ptyctodontida (ptyktodonty) i Phyl- porównanie stanowiska na górze Pod³azie ko³o Daleszyc lolepida (filolepidy). Trudnoœci w rozró¿nieniu skamie- (ryc. 1A) (stanowisko to jest, z punktu widzenia stanu nia³oœci poszczególnych grup miêdzy sob¹ mo¿na zachowania i sk³adu taksonomicznego fauny krêgowców, napotkaæ jedynie w przypadku artrodirów i antiarchów. reprezentatywne dla ca³ego dewonu dolnego regionu kie- Ostateczne potwierdzenie obecnoœci antiarchów w leckiego Gór Œwiêtokrzyskich, w miejscach wystêpowania utworach dewoñskich Gór Œwiêtokrzyskich da³y dopiero „piaskowców plakodermowych” sensu Szrek 2003a) ze wyniki badañ ostatnich lat (Szrek, 2003a, b, 2004). Do tej stanowiskiem w Lode we wschodniej £otwie (ryc. 1B), pory z Gór Œwiêtokrzyskich zosta³y opisane trzy okazy gdzie ods³oniête s¹, w du¿ym kamienio³omie, fameñskie szcz¹tków antiarchów w dewonie dolnym (Szrek, 2003a) i osady proksymalnej czêœci delty, zawieraj¹ce bardzo jeden (prawie kompletny) okaz w dewonie górnym (Szrek, dobrze zachowan¹, bogat¹ i zró¿nicowan¹ faunê krêgow- 2004). Powodem takiego ubóstwa s¹ trudnoœci w identyfi- ców. Brak zaanga¿owania tektonicznego i dobry stopieñ kacji niekompletnych skamienia³oœci przedstawicieli tej ods³oniêcia, jak te¿ trwaj¹ce od kilkudziesiêciu lat inten- grupy, co przyczynia³o siê w przesz³oœci do powstania nie- sywne badania fauny sprawi³y, ¿e stanowisko mo¿na uznaæ zgodnoœci w oznaczeniach. Informacje o znalezieniu za wzorcowe i do niego odnosiæ takie miejsca jak ods³oniê- szcz¹tków antiarchów podawali: Gürich (1896), Czarnocki cie na górze Pod³azie. Stwierdziæ mo¿na podobieñstwo (1919, 1936), Gorizdro-Kulczycka (1934, 1949) i Kul- sk³adu fauny krêgowców w Lode, w którym dominuj¹ czycki (1957), jednak zdaniem Ivanova i Gintera (1997) kolejno: ostrakodermy (g³ównie psammosteidy — ryc. szcz¹tki opisane przez powy¿szych autorów nale¿¹ w rze- 3A), ryby kostnoszkieletowe (Sarcopterygii — miê- czywistoœci do Brachythoraci. Autor sk³ania siê ku œniop³etwe) oraz pancerne, a wœród nich antiarchy (ryc. 2, pogl¹dowi, i¿ te i póŸniej znalezione szcz¹tki reprezentuj¹ 3C; Lukševiès, 1992, 2001), do sk³adu fauny w „piaskow- jednak przedstawicieli rzêdu Antiarcha, co zosta³o stwier- cach plakodermowych”. Oczywiœcie taksonomicznie, na dzone na podstawie analizy morfologii i ornamentacji poziomie gatunków, oba stanowiska nie maj¹ ze sob¹ nic powierzchni tarcz. wspólnego, co jest wynikiem oddalenia stratygraficznego osadów z obu ods³oniêæ. Mamy natomiast do czynienia z podobieñstwem proporcji iloœciowej poszczególnych grup krêgowców i/lub form morfologicznych. W piaskowcach Pod³azia stwierdzono mianowicie obecnoœæ zró¿nicowa- *Wydzia³ Geologii, Uniwersytet Warszawski, ul. ¯wirki i nego taksonomicznie zespo³u ryb miêœniop³etwych (Sar- Wigury 93, 02-089 Warszawa; [email protected] copterygii: Porolepis sp. Holoptychius cf. nobilissimus – 610 Przegl¹d Geologiczny, vol. 54, nr 7, 2006 na podstawie Kulczyckiego (1960), jak równie¿ plakoder- Jedyn¹ cech¹, która decydowa³a o oznaczeniu danego mów, wœród których wiêcej stwierdza siê szcz¹tków okazu jako przedstawiciela rzêdu Antiarcha by³ u Güricha antiarchów (Szrek, 2003a, b); bogat¹ faunê psammoste- (1896), Gorizdro-Kulczyckiej (1934, 1949) i Kulczyckie- idów opracowa³ Tarlo (1962, 1964, 1965). Podobne aso- go (1957), charakterystyczny wygl¹d ornamentacji cjacje zosta³y stwierdzone tak¿e na platformie powierzchni tarcz. Istotnie, urzeŸbienie mo¿e byæ zasad- wschodnioeuropejskiej w innych stanowiskach g³ównego nicz¹ cech¹ diagnostyczn¹ w przypadku, gdy nie ma mo¿li- pola dewonu (Moloshnikov, 2004; Lukševiès, 1992, 1999, woœci badania histologicznego, a w „piaskowcach 2001). plakodermowych” mamy do czynienia praktycznie jedynie Elementem w³aœciwym dla „piaskowców plakodermo- z negatywami zewnêtrznej (ryc. 4–6) lub wewnêtrznej wych” z Pod³azia, nieobecnym w Lode, jest czêste wystê- strony p³yty, b¹dŸ obu, w przypadku pustki po tarczy. powanie kolców p³etwowych przedstawicieli gromady Kawa³ki koœci zachowane s¹ w postaci szcz¹tkowej i sta- fa³dop³etwych (Acanthodii), ujmowanych pod nazw¹ nowi¹ rzadkoœæ (Tarnowska, 1981; Szrek, 2003a). Machaeracanthus polonicus Gürich. Ich rola w tym eko- Odciski tarcz ró¿nych krêgowców w „piaskowcach systemie, jak te¿ opisanie tych niezilustrowanych, a zró¿- plakodermowych” ju¿ na pierwszy rzut oka wykazuj¹ ró¿- nicowanych morfologicznie szcz¹tków, stan¹ siê jednak nice taksonomiczne (porównaj ryc. 4–6). Wynikaj¹ one z przedmiotem osobnego opracowania paleontologicznego. rodzaju i budowy guzków pokrywaj¹cych ich powierzch- Rodzaj œrodowiska silnie determinowa³ obecnoœæ lub niê. Ornamentacja powierzchni pancerza heterostraków absencjê przedstawicieli rzêdu Antiarcha (Stensiö, 1931, (psammosteidów), licznie wystêpuj¹cych w „piaskowcach 1948; Denison, 1978; Long, 1996; Lukševiès, 1992, 1999, plakodermowych”, zaznacza siê w najbardziej zewnêtrznej 2001). Mo¿na wiêc zamiennie: okreœlaæ rodzaj œrodowiska warstwie tarcz i stanowi³y j¹ guzki zbudowane z dentyny posi³kuj¹c siê faun¹ antiarchów i typowaæ mo¿liwoœæ ich (ryc. 3A). Guzki na powierzchni ich tarcz s¹ gêsto roz- obecnoœci w ska³ach okreœlonej genezy, zale¿nie od tego, mieszczone i powszechne jest stykanie siê s¹siaduj¹cych który z tych dwóch czynników jest widoczny lub bardziej ze sob¹ guzków. W przypadku najpospolitszego i reprezen- wyrazisty. tatywnego dla grupy rodzaju Psammosteus guzki s¹ tak bardzo blisko siebie, ¿e ich zarys jest wielok¹tny (najczê- œciej szeœciok¹tny; ryc. 3A, 4) i uk³adaj¹ siê one B ESTONIA w ci¹gi, których przebieg jest niekiedy determi- ROSJA nowany zarysem tarczy (porównaj Obruchev, A Ba³tyk Baltic Sea 1964; tabl. VI, fig. 2, 3, 6). Wy¿ej opisana orna- mentacja wyraŸnie ró¿ni siê od wystêpuj¹cej u Lode Górno Ryga RUSSIA Opató antiarchów. w Riga Wiêksz¹ trudnoœæ sprawia odró¿nienie od Brzechów Góra Pod³azie Œwinia Góra siebie odcisków koœci pancerza plakodermów i Kranów Suków ryb koœcistych (miêœniop³etwe — Sarcoptery- Daleszyce LITWA gii) ze wzglêdu na ich du¿e podobieñstwo. 05km S³opiec LITHUANIA Piñczów ROSJA Dotyczy to, spoœród plakodermów, przede RUSSIA BIA£ORUŒ wszystkim artrodirów, gdy¿ ich koœci skórne, POLSKA 0 100km BELARUS POLAND podobnie jak niektóre tarcze pokrywowe u miê- œniop³etwych (ryc. 6), s¹ pokryte nieregularnie Ryc. 1. Lokalizacja stanowiska na górze Pod³azie ko³o Daleszyc (A) i stanowi- rozmieszczonymi guzkami (ryc. 3B, 5). U ska w Lode we wschodniej £otwie (B) antiarchów uk³ad guzków jest bardziej Fig. 1. Location of the outcrop on the Pod³azie Hill near Daleszyce, central Poland (A) and the quarry in Lode in the eastern Latvia (B) uporz¹dkowany. Ponadto guzki u antiarchów s¹ wyraŸnie mniejsze ni¿ u artrodirów i miê- œniop³etwych i ich œrednica u podstawy rzadko AMD PMD nie przekracza 1 mm (porównaj ryc. 3C, 5, 6). O ile, w przypadku analizy budowy koœci, rozró¿- ADL MxL nienie przedstawiciela Arthrodira od Sarcop- terygii nie sprawia trudnoœci, z
Load more

Recommended publications
  • 'Placoderm' (Arthrodira)
    Jobbins et al. Swiss J Palaeontol (2021) 140:2 https://doi.org/10.1186/s13358-020-00212-w Swiss Journal of Palaeontology RESEARCH ARTICLE Open Access A large Middle Devonian eubrachythoracid ‘placoderm’ (Arthrodira) jaw from northern Gondwana Melina Jobbins1* , Martin Rücklin2, Thodoris Argyriou3 and Christian Klug1 Abstract For the understanding of the evolution of jawed vertebrates and jaws and teeth, ‘placoderms’ are crucial as they exhibit an impressive morphological disparity associated with the early stages of this process. The Devonian of Morocco is famous for its rich occurrences of arthrodire ‘placoderms’. While Late Devonian strata are rich in arthrodire remains, they are less common in older strata. Here, we describe a large tooth-bearing jaw element of Leptodontich- thys ziregensis gen. et sp. nov., an eubrachythoracid arthrodire from the Middle Devonian of Morocco. This species is based on a large posterior superognathal with a strong dentition. The jawbone displays features considered syna- pomorphies of Late Devonian eubrachythoracid arthrodires, with one posterior and one lateral row of conical teeth oriented postero-lingually. μCT-images reveal internal structures including pulp cavities and dentinous tissues. The posterior orientation of the teeth and the traces of a putative occlusal contact on the lingual side of the bone imply that these teeth were hardly used for feeding. Similar to Compagopiscis and Plourdosteus, functional teeth were pos- sibly present during an earlier developmental stage and have been worn entirely. The morphological features of the jaw element suggest a close relationship with plourdosteids. Its size implies that the animal was rather large. Keywords: Arthrodira, Dentition, Food web, Givetian, Maïder basin, Palaeoecology Introduction important to reconstruct character evolution in early ‘Placoderms’ are considered as a paraphyletic grade vertebrates.
    [Show full text]
  • Contributions in BIOLOGY and GEOLOGY
    MILWAUKEE PUBLIC MUSEUM Contributions In BIOLOGY and GEOLOGY Number 51 November 29, 1982 A Compendium of Fossil Marine Families J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions in BIOLOGY and GEOLOGY Number 51 November 29, 1982 A COMPENDIUM OF FOSSIL MARINE FAMILIES J. JOHN SEPKOSKI, JR. Department of the Geophysical Sciences University of Chicago REVIEWERS FOR THIS PUBLICATION: Robert Gernant, University of Wisconsin-Milwaukee David M. Raup, Field Museum of Natural History Frederick R. Schram, San Diego Natural History Museum Peter M. Sheehan, Milwaukee Public Museum ISBN 0-893260-081-9 Milwaukee Public Museum Press Published by the Order of the Board of Trustees CONTENTS Abstract ---- ---------- -- - ----------------------- 2 Introduction -- --- -- ------ - - - ------- - ----------- - - - 2 Compendium ----------------------------- -- ------ 6 Protozoa ----- - ------- - - - -- -- - -------- - ------ - 6 Porifera------------- --- ---------------------- 9 Archaeocyatha -- - ------ - ------ - - -- ---------- - - - - 14 Coelenterata -- - -- --- -- - - -- - - - - -- - -- - -- - - -- -- - -- 17 Platyhelminthes - - -- - - - -- - - -- - -- - -- - -- -- --- - - - - - - 24 Rhynchocoela - ---- - - - - ---- --- ---- - - ----------- - 24 Priapulida ------ ---- - - - - -- - - -- - ------ - -- ------ 24 Nematoda - -- - --- --- -- - -- --- - -- --- ---- -- - - -- -- 24 Mollusca ------------- --- --------------- ------ 24 Sipunculida ---------- --- ------------ ---- -- --- - 46 Echiurida ------ - --- - - - - - --- --- - -- --- - -- - - ---
    [Show full text]
  • Redescription of Yinostius Major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the Interrelationships of Brachythoraci
    bs_bs_banner Zoological Journal of the Linnean Society, 2015. With 10 figures Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci YOU-AN ZHU1,2, MIN ZHU1* and JUN-QING WANG1 1Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China 2University of Chinese Academy of Sciences, Beijing 100049, China Received 29 December 2014; revised 21 August 2015; accepted for publication 23 August 2015 Yinosteus major is a heterostiid arthrodire (Placodermi) from the Lower Devonian Jiucheng Formation of Yunnan Province, south-western China. A detailed redescription of this taxon reveals the morphology of neurocranium and visceral side of skull roof. Yinosteus major shows typical heterostiid characters such as anterodorsally positioned small orbits and rod-like anterior lateral plates. Its neurocranium resembles those of advanced eubrachythoracids rather than basal brachythoracids, and provides new morphological aspects in heterostiids. Phylogenetic analysis based on parsimony was conducted using a revised and expanded data matrix. The analysis yields a novel sce- nario on the brachythoracid interrelationships, which assigns Heterostiidae (including Heterostius ingens and Yinosteus major) as the sister group of Dunkleosteus amblyodoratus. The resulting phylogenetic scenario suggests that eubrachythoracids underwent a rapid diversification during the Emsian, representing the placoderm response to the Devonian Nekton Revolution. The instability of the relationships between major eubrachythoracid clades might have a connection to their longer ghost lineages than previous scenarios have implied. © 2015 The Linnean Society of London, Zoological Journal of the Linnean Society, 2015 doi: 10.1111/zoj.12356 ADDITIONAL KEYWORDS: Brachythoraci – Heterostiidae – morphology – phylogeny – Placodermi.
    [Show full text]
  • The Earliest Phyllolepid (Placodermi, Arthrodira) from the Late Lochkovian (Early Devonian) of Yunnan (South China)
    Geol. Mag. 145 (2), 2008, pp. 257–278. c 2007 Cambridge University Press 257 doi:10.1017/S0016756807004207 First published online 30 November 2007 Printed in the United Kingdom The earliest phyllolepid (Placodermi, Arthrodira) from the Late Lochkovian (Early Devonian) of Yunnan (South China) V. DUPRET∗ &M.ZHU Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, P.O. Box 643, Xizhimenwai Dajie 142, Beijing 100044, People’s Republic of China (Received 1 November 2006; accepted 26 June 2007) Abstract – Gavinaspis convergens, a new genus and species of the Phyllolepida (Placodermi: Arthrodira), is described on the basis of skull remains from the Late Lochkovian (Xitun Formation, Early Devonian) of Qujing (Yunnan, South China). This new form displays a mosaic of characters of basal actinolepidoid arthrodires and more derived phyllolepids. A new hypothesis is proposed concerning the origin of the unpaired centronuchal plate of the Phyllolepida by a fusion of the paired central plates into one single dermal element and the loss of the nuchal plate. A phylogenetic analysis suggests the position of Gavinaspis gen. nov. as the sister group of the Phyllolepididae, in a distinct new family (Gavinaspididae fam. nov.). This new form suggests a possible Chinese origin for the Phyllolepida or that the common ancestor to Phyllolepida lived in an area including both South China and Gondwana, and in any case corroborates the palaeogeographic proximity between Australia and South China during the Devonian Period. Keywords: Devonian, China, Placodermi, phyllolepids, biostratigraphy, palaeobiogeography. 1. Introduction 1934). Subsequently, they were considered as either sharing an immediate common ancestor with the The Phyllolepida are a peculiar group of the Arthrodira Arthrodira (Denison, 1978), belonging to the Actin- (Placodermi), widespread in the Givetian–Famennian olepidoidei (Long, 1984), or being of indetermined of Gondwana (Australia, Antarctica, Turkey, South position within the Arthrodira (Goujet & Young,1995).
    [Show full text]
  • Facies Differentiation and Late Devonian Placoderms Fossils in the Holy Cross Mountains
    Przegl¹d Geologiczny, vol. 54, nr 6, 2006 Zró¿nicowanie facjalne a skamienia³oœci póŸnodewoñskich plakodermów w Górach Œwiêtokrzyskich Piotr Szrek* Facies differentiation and Late Devonian placoderms fossils in the Holy Cross Mountains. Prz. Geol., 54: 521–524. S u m m a r y. Main Late Devonian placoderm taxa known from the Holy Cross Mountains are characterised. The distribution of the Late Devonian placoderm fossils is described. Variation in their occurrences depend on the sedimentation environment of the rocks which contain fossils of this group. Connections between the Holy Cross Mountains placoderms development and the synsedimentary tectonic processes active in this area during the Late Devonian is discussed, and the local faunas compared to classic assemblages of the same age from Latvia. Key words: Placodermi, synsedimentation tectonic, Late Devonian, facies, Holy Cross Mountains Placodermi (ryby pancerne) to gromada krêgowców Plakodermy œwiêtokrzyskie wodnych, która pojawi³a siê w sylurze, a wymar³a z koñcem dewonu. Ich maksymalny stopieñ zró¿nicowania Znalezione i opisane dotychczas póŸnodewoñskie pla- przypada na póŸny dewon — wtedy to plakodermy zdomi- kodermy z Gór Œwiêtokrzyskich mo¿na podzieliæ na dwie nowa³y niemal wszystkie morskie nisze ekologiczne, zasadnicze grupy. Kryterium zastosowanego podzia³u jest wytwarzaj¹c ró¿norodne typy specjalizacyjne, umo¿li- sposób zdobywania po¿ywienia oraz stopieñ przywi¹zania wiaj¹ce ¿ycie zarówno w otwartym basenie morskim, œro- do okreœlonych œrodowisk, a tak¿e stopieñ opancerzenia dowisku przyrafowym, jak te¿ w œrodowiskach cia³a. brakicznych. W Górach Œwiêtokrzyskich grupa ta jest Do pierwszej grupy mo¿na zaliczyæ wszystkie pla- bogato reprezentowana w materiale kopalnym i by³a oma- kodermy, bêd¹ce aktywnymi myœliwymi, u których pancerz wiana w licznych pracach od ponad stu lat (m.in.
    [Show full text]
  • Société Géologique Nord
    Société Géologique Nord 3 1 JAN. ?00D ANNALES Tome 7(2'me série), Fascicule 3 parution 1999 IRIS - LILLIAD - Université Lille 1 SOCIÉTÉ GÉOLOGIQUE DU NORD Extraits des Statuts Article 2 - Cette Société a pour objet de concourir à l'avancement de la géologie en général, et particulièrement de la géologie de la région du Nord de la France. - La Société se réunit de droit une fois par mois, sauf pendant la période des vacances. Elle peut tenir des séances extraordinaires décidées par le Conseil d'Administration. - La Société publie des Annales et des Mémoires. Ces publications sont mises en vente selon un tarif établi par le Conseil. Les Sociétaires bénéficient d'un tarif préférentiel 0 ). Articles Le nombre des membres de la Société est illimité. Pour faire partie de la Société, il faut s'être fait présenter dans l'une des séances par deux membres de la Société qui auront signé la présentation, et avoir été proclamé membre au cours de la séance suivante. Extraits du Règlement Intérieur § 7. - Les Annales et leur supplément constituent le compte rendu des séances. § 13. - Seuls les membres ayant acquitté leurs cotisation et abonnement de l'année peuvent publier dans les Annales. L'ensemble des notes présentées au cours d'une même année, par un auteur, ne peut dépasser le total de 8 pages, 1 planché simili étant comptée pour 2 p. 1/2 de texte. Le Conseil peut par décision spéciale, autoriser la publication de notes plus tangues. § 17. - Les notes et mémoires originaux (texte et illustration) communiqués à la Société et destinés aux Annales doivent être remis au Secrétariat le jour même de leur présentation.
    [Show full text]
  • Vertebrata: Placodermi) from the Famennian of Belgium
    View metadata, citationGEOLOGICA and similar papers BELGICA at core.ac.uk (2005) 8/1-2: 51-67 brought to you by CORE provided by Open Marine Archive A NEW GROENLANDASPIDID ARTHRODIRE (VERTEBRATA: PLACODERMI) FROM THE FAMENNIAN OF BELGIUM Philippe JANVIER1, 2 and Gaël CLÉMENT1 1. UMR 5143 du CNRS, Département «Histoire de la Terre», Muséum National d’Histoire Naturelle, 8 rue Buff on, 75005 Paris, France. [email protected] 2. 7 e Natural History Museum, Cromwell Road, London SW7 5BD, United Kingdom (9 fi gures, 1 table and 2 plates) ABSTRACT. A new species of the arthrodire genus Groenlandaspis is described from the upper part of the Evieux Formation (Upper Famennian), based on several specimens collected from quarries at Modave and Villers-le-Temple, Liège Province, Belgium. It is the fi rst occurrence of this widespread genus in continental Europe. O is new species is characterized by an almost smooth dermal armour, except for some scattered tubercles on its skull roof, median dorsal and spinal plates. Its median dorsal plate is triangular in shape and almost perfectly equilateral in lateral aspect and bears large, spiniform denticles on its posterior edge. All these Groenlandaspis remains occur in micaceous, dolomitic claystones or siltstones probably deposited in a subtidal environment. Outcrops of the same area have yielded other vertebrate remains, such as the placoderms Phyllolepis and Bothriolepis, acanthodians, various piscine sarcopterygians (Holoptychius, dipnoans, a rhizodontid, Megalichthys, Eusthenodon and a large tristichopterid), and a tetrapod that is probably close to Ichthyostega. O e biogeographical history of the genus Groenlandaspis is briefl y outlined, and the late Frasnian-Famennian interchange of vertebrate taxa between Gondwana and Euramerica is discussed.
    [Show full text]
  • Comments on the Late Devonian Placoderms from the Holy Cross Mountains (Poland)
    Comments on the Late Devonian placoderms from the Holy Cross Mountains (Poland) ALEXANDER IVANOV andMICHAŁ GINTER Ivanov, A. & Ginter,M. 1997. Comments on the Late Devonian placoderms from the Holy Cross Mountains (Poland).- Acta Palaeontologica Polonica 4,3,4I34f6. Taxonomy of the Late Devonian placoderm remains from the Holy Cross Mountains, Poland, described by Gorizdro-Kulczycka (L934,1950) and Kulczycki (1956, 1957),is revised. Several recently found specimens are also mentioned. The old collections are composed of representatives of Ptyctodontidae, Holonematidae, Plourdosteidae, Pholi- dosteidae, Selenosteidae, Titanichthyidae and Dinichthyidae, the latter with an unde- scribed species of Eastmanosteus. Newly found specimens belong to Ptyctodontidae, Plourdosteidae and Dinichthyidae. The occurrence of the Antiarcha in the Late Devonian of the Holy Cross Mountains, suggestedby former authors, has not been confirmed. K e y w o rd s : Placodermi,Late Devonian, Holy Cross Mountains, Poland. Alexander Ivanov [[email protected]], Laboratory of Paleontology, Institute of the Earth Crust, Sankt-Petersburg University, 16 Liniya 29, 199178 St.Petersburg, Russia. Michał Ginter [email protected]], InsĘtut Geologii Podstawowej, Uniwersytet War szaw ski, ul. Zw irki i Wi gury 9 3, 02 -089 War szaw a, P oland. Introduction The main goals of this study are to revise placodermsfrom the Late Devonian of the Holy Cross Mountains collected by Jan Czarnocki, Julian Kulczycki and Zinuda Gorizdro-Kulczycka in the first half of the century (the material now housed in the Museum of the Polish Geological Instituteand in the Museum of the Earth in Warsaw), and to describe a fęw new specimenscollected recently by Jerzy Dzik and Grzegorz Racki.
    [Show full text]
  • A New Lower Devonian Arthrodire (Placodermi) from the NW Siberian Platform
    Estonian Journal of Earth Sciences, 2013, 62, 3, 131–138 doi: 10.3176/earth.2013.11 A new Lower Devonian arthrodire (Placodermi) from the NW Siberian Platform Elga Mark-Kurik Institute of Geology at Tallinn University of Technology, Ehitajate tee 5, 19086 Tallinn, Estonia; [email protected] Received 24 August 2012, accepted 5 November 2012 Abstract. A new genus and species of arthrodires, Eukaia elongata (Actinolepidoidei, Placodermi), is described from the Lower Devonian, ?Pragian of the Turukhansk region, NW Siberian Platform. A single specimen of the fish, a skull roof, comes probably from the lower part of the Razvedochnyj Formation. The occurrence of an actinolepidoid arthrodire in the Early Devonian of this area of Siberia is unexpected. Eukaia shows some distant relationship with the genus Actinolepis, but several features indicate similarity to representatives of other arthrodires. Key words: actinolepidoid arthrodire, placoderm, Lower Devonian, ?Pragian, NW Siberian Platform. INTRODUCTION (Fig. 1). The first two units are Lochkovian in age, the lower part of the Razvedochnyj Fm belongs to the Pragian The northwestern part of the Siberian Platform in the and the upper part of the Razvedochnyj Fm plus the lower Russian Arctic is well known for rich and amphiaspidid- part of the Mantura Fm to the Emsian (Matukhin 1995). dominated Early Devonian fish faunas. One of the The Zub Fm (up to 150 m thick) consists of carbonaceous- important areas where these faunas have been discovered argillaceous and sulphate rocks. Invertebrates and fossil is the near-Yenisej zone of the Tunguska syneclise fishes, e.g. a cyathaspidid Steinaspis, are comparatively (Krylova et al.
    [Show full text]
  • Devonian Fish Remains from the Munabia Sandstone, Carnarvon Basin, Western Australia
    ------ ---_ - _- Ree. West. /lust. Mus. 1991,15(3) 501515 Devonian fish remains from the Munabia Sandstone, Carnarvon Basin, Western Australia John A. Long· Abstract Fish fossils have been recovered from two horizons at the base ofthe Munabia Sandstone, near the type section of Williambury Station. The fauna contains the antiarch Bothriolepis sp" the arthrodire Holonema sp., and an indeterminate osteolepidid crossopterygian. The presence of Holonema (Eifelian-Frasnian) with Bothriolepis(Givetian-Famennian), as well as microfossil dates from the underlying conformable Gneudna Formation (Early Frasnian), suggests a Frasnian age for the base of the Munabia Sandstone. Conodonts suggest that the top of the Munabia Sandstone is of Lower Famennian age. Introduction The Munabia Sandstone outcrops over a distance ofalmost 90 km from Mt. Sandiman homestead in the south to just north of Williambury Station as part of a linear belt of Devonian rocks at the base of the Carnarvon Basin sedimentary succession (Figure 1). The type section occurs about 8 km southeast of Williambury Station where it conformably overlies the lower Frasnian carbonates of the Gneudna Formation and underlies the coarser conglomerates of the Willaradie Formation (Condon 1954, 1965, Hocking et al. 1987). Although Condon (1965) favoured a marine depositional environment for the Munabia Sandstone and Willaradie Formation, Moors (1981) reported that only the base of the Munabia Sandstone was marine, the majority of the sequence representing distal fan to braided stream deposits, with minor marine incursions. The overlying Willaradie Formation is part of this depositional event, representing proximal alluvial fan deposits. None of the previous field studies of the Munabia Sandstone had found any body fossils, although trace fossils are common in the lower (marine) horizons, and the age of the unit was based entirely on extrapolation from the underlying Gneudna Formation, itself well-dated from marine invertebrates and palynomorphs (Seddon 1969, Dring 1980, Playford and Dring 1981).
    [Show full text]
  • Sepkoski, J.J. 1992. Compendium of Fossil Marine Animal Families
    MILWAUKEE PUBLIC MUSEUM Contributions . In BIOLOGY and GEOLOGY Number 83 March 1,1992 A Compendium of Fossil Marine Animal Families 2nd edition J. John Sepkoski, Jr. MILWAUKEE PUBLIC MUSEUM Contributions . In BIOLOGY and GEOLOGY Number 83 March 1,1992 A Compendium of Fossil Marine Animal Families 2nd edition J. John Sepkoski, Jr. Department of the Geophysical Sciences University of Chicago Chicago, Illinois 60637 Milwaukee Public Museum Contributions in Biology and Geology Rodney Watkins, Editor (Reviewer for this paper was P.M. Sheehan) This publication is priced at $25.00 and may be obtained by writing to the Museum Gift Shop, Milwaukee Public Museum, 800 West Wells Street, Milwaukee, WI 53233. Orders must also include $3.00 for shipping and handling ($4.00 for foreign destinations) and must be accompanied by money order or check drawn on U.S. bank. Money orders or checks should be made payable to the Milwaukee Public Museum. Wisconsin residents please add 5% sales tax. In addition, a diskette in ASCII format (DOS) containing the data in this publication is priced at $25.00. Diskettes should be ordered from the Geology Section, Milwaukee Public Museum, 800 West Wells Street, Milwaukee, WI 53233. Specify 3Y. inch or 5Y. inch diskette size when ordering. Checks or money orders for diskettes should be made payable to "GeologySection, Milwaukee Public Museum," and fees for shipping and handling included as stated above. Profits support the research effort of the GeologySection. ISBN 0-89326-168-8 ©1992Milwaukee Public Museum Sponsored by Milwaukee County Contents Abstract ....... 1 Introduction.. ... 2 Stratigraphic codes. 8 The Compendium 14 Actinopoda.
    [Show full text]
  • Family-Group Names of Fossil Fishes
    European Journal of Taxonomy 466: 1–167 ISSN 2118-9773 https://doi.org/10.5852/ejt.2018.466 www.europeanjournaloftaxonomy.eu 2018 · Van der Laan R. This work is licensed under a Creative Commons Attribution 3.0 License. Monograph urn:lsid:zoobank.org:pub:1F74D019-D13C-426F-835A-24A9A1126C55 Family-group names of fossil fishes Richard VAN DER LAAN Grasmeent 80, 1357JJ Almere, The Netherlands. Email: [email protected] urn:lsid:zoobank.org:author:55EA63EE-63FD-49E6-A216-A6D2BEB91B82 Abstract. The family-group names of animals (superfamily, family, subfamily, supertribe, tribe and subtribe) are regulated by the International Code of Zoological Nomenclature. Particularly, the family names are very important, because they are among the most widely used of all technical animal names. A uniform name and spelling are essential for the location of information. To facilitate this, a list of family- group names for fossil fishes has been compiled. I use the concept ‘Fishes’ in the usual sense, i.e., starting with the Agnatha up to the †Osteolepidiformes. All the family-group names proposed for fossil fishes found to date are listed, together with their author(s) and year of publication. The main goal of the list is to contribute to the usage of the correct family-group names for fossil fishes with a uniform spelling and to list the author(s) and date of those names. No valid family-group name description could be located for the following family-group names currently in usage: †Brindabellaspidae, †Diabolepididae, †Dorsetichthyidae, †Erichalcidae, †Holodipteridae, †Kentuckiidae, †Lepidaspididae, †Loganelliidae and †Pituriaspididae. Keywords. Nomenclature, ICZN, Vertebrata, Agnatha, Gnathostomata.
    [Show full text]