Quick Guide to Tribes of Compositae in California

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QUICK GUIDE TO TRIBES OF COMPOSITAE IN CALIFORNIA Subfamily Asteroideae Heliantheae — Chaffy receptacle; pappus of anything except (gen) capillary bristles Subtribe Ambrosiinae — Wind-pollinated; anthers free; female fls w/o corolla; florets dull-colored; easy Madieae — Heterogeneous group not united by obvious morphological features; common in CA Subtribe Madiinae — Phyllaries each enclosing a ray achene; tack-shaped “tar” glands; receptacular bracts gen limited to a series external to disc florets; easy Coreopsideae — Similar to Heliantheae but achenes obcompressed Chaenacteae — Mostly Chaenactis in CA, with typically radiant heads (exc Dimeresia w/o radiant heads) Perityleae — Desert plants with radiate heads; disc flts 4-merous; easy Bahieae — Mostly (in CA) discoid heads (exc Bahia) w/ scales or awns for pappus (exc some Bahia); most that you’ll see are in the desert; doesn’t jump out with obvious morph characters Helenieae — Hetrogeneous group not united by obvious morphological features, exc gen epaleate; common in CA Senecioneae — Pappus of capillary bristles; phyllaries appearing uniseriate and side-by-side Astereae — Pappus of capillary bristles; phyllaries appearing multiseriate and overlapping Anthemideae – Papery-margined phyllaries; compound or lobed leaves; truncate style branches; pappus gen 0; pls often aromatic Eupatorieae — Long style branches; corollas never yellow; head discoid Calenduleae — Radiate heads; tailed anthers; pappus 0; fr arcuate and warty Gnaphalieae — Each entire phyllary papery (not green, not herbaceous); inconspicuous florets/heads; tailed anthers; easy to recognize, but often overlooked in the field Tageteae – Aromatic, obvious oil glands on phyllaries, leaves, other parts; mostly desert in CA Subfamily Cichorioideae Arctoteae — Radiate heads; swollen area below style branches; stigmatic on adaxial surface Cardueae — Thistle-like, often prickly; heads discoid; corolla lobes long; styles long but style branches short; tailed anthers; pappus of capillary bristles; easy to recognize Cichoreae — All florets strap-shaped (ligulate) with 5 teeth; milky juice; the easiest to recognize Subfamily Mutisioideae Mutisieae — Bilabiate florets exclusively; tailed anthers; deserts; easy to recognize if you’re expecting them Subfamily Hecastocleoideae Hecastocleae — .

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