HORTAX Cultivated Plant Taxonomy
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BIOL/APBI 324 – Introduction to Seed Plant Taxonomy
This syllabus is a general representation of the course as previously offered and is subject to change. BIOL/APBI 324 – Introduction to Seed Plant Taxonomy General Course Syllabus (as of September 2019) About the Course: Course Description: An introduction to seed plant taxonomy emphasizing descriptive morphology and identification. Each student will be required to submit a plant collection. Correct understanding of the actual relationships between plants, rather than superficial resemblance, is the basis of comparative biology required to analyze the diversity of plant forms. The species diversity of plants is considerable, and this course will pay particular attention to very diverse and important groups such as the grasses and the orchids, focusing on reasons for their evolutionary success. Identification skills will be inculcated by the lectures working in tandem with the laboratory sessions. This course aims to give students a good working knowledge of plant taxonomy as a preparation for work in any biological discipline. Course Format: Lecture and Laboratory Credits: 3 Pre-requisites: BIOL 121 Course Learning Objectives: By the end of this course, students will be able to: • Achieve a good working knowledge of concepts, principles, and recent discoveries in plant taxonomy. • Gain an overview of seed plant diversity, including the most species-rich plant families and so be able to place any botanical information in the overall context of plant diversity. • Learn ways in which their knowledge can be applied to ecology and evolutionary biology. • Gain an appreciation of how current research in the field is being done by reading recent research papers. Textbooks and Additional Resources: Laboratory fee: $25; please bring to first lab. -
Taxonomic Review of the Genus Rosa
REVIEW ARTICLE Taxonomic Review of the Genus Rosa Nikola TOMLJENOVIĆ 1 ( ) Ivan PEJIĆ 2 Summary Species of the genus Rosa have always been known for their beauty, healing properties and nutritional value. Since only a small number of properties had been studied, attempts to classify and systematize roses until the 16th century did not give any results. Botanists of the 17th and 18th century paved the way for natural classifi cations. At the beginning of the 19th century, de Candolle and Lindley considered a larger number of morphological characters. Since the number of described species became larger, division into sections and subsections was introduced in the genus Rosa. Small diff erences between species and the number of transitional forms lead to taxonomic confusion and created many diff erent classifi cations. Th is problem was not solved in the 20th century either. In addition to the absence of clear diff erences between species, the complexity of the genus is infl uenced by extensive hybridization and incomplete sorting by origin, as well as polyploidy. Diff erent analytical methods used along with traditional, morphological methods help us clarify the phylogenetic relations within the genus and give a clearer picture of the botanical classifi cation of the genus Rosa. Molecular markers are used the most, especially AFLPs and SSRs. Nevertheless, phylogenetic relationships within the genus Rosa have not been fully clarifi ed. Th e diversity of the genus Rosa has not been specifi cally analyzed in Croatia until now. Key words Rosa sp., taxonomy, molecular markers, classifi cation, phylogeny 1 Agricultural School Zagreb, Gjure Prejca 2, 10040 Zagreb, Croatia e-mail: [email protected] 2 University of Zagreb, Faculty of Agriculture, Department of Plant Breeding, Genetics and Biometrics, Svetošimunska cesta 25, 10000 Zagreb, Croatia Received: November , . -
Principles of Plant Taxonomy, V.*
THE OHIO JOURNAL OF SCIENCE VOL. XXVIII MARCH, 1928 No. 2 PRINCIPLES OF PLANT TAXONOMY, V.* JOHN H. SCHAFFNER, Ohio State University. After studying the taxonomy of plants for twenty-five years the very remarkable fact became evident that there is no general correspondence of the taxonomic system with the environment, but as the great paleontologist, Williams, said in 1895: "environmental conditions are but the medium through which organic evolution has been determinately ploughing its way." Of course, the very fact that there is a system of phylogenetic relationships of classes, orders, families, and genera and that these commonly have no general correspondence to environment shows that, in classifying the plant material, we must discard all notions of teleological, utilitarian, and selective factors as causative agents of evolution. The general progressive movement has been carried on along quite definite lines. The broader and more fundamental changes appeared first and are practically constant, and on top of these, .potentialities or properties of smaller and smaller value have been introduced, until at the end new factors of little general importance alone are evolved. These small potentialities are commonly much less stable than the more fundamental ones and thus great variability in subordinate characters is often present in the highest groups. We must then think of the highest groups as being full of hereditary potentialities while the lower groups have comparatively few. As stated above, there is a profound non-correspondence of the .taxonomic system and the various orthogenetic series with the environment. The system of plants, from the taxonomic point of view, is non-utilitarian. -
Multiflora Rose, Rosa Multiflora Thunb. Rosaceae
REGULATORY HORTICULTURE [Vol. 9, No.1-2] Weed Circular No. 6 Pennsylvania Department of Agriculture April & October 1983 Bureau of Plant Industry Multiflora Rose, Rosa multiflora Thunb. Rosaceae. Robert J. Hill I. Nomenclature: A) Rosa multiflora Thunb. (Fig. 1); B) Multiflora rose; C) Synonyms: Rosa Dawsoniana Hort., R. polyantha Sieb. & Zucc., R. polyanthos Roessia., R. thyrsiflora Leroy, R. intermedia, Carr., and R. Wichurae Kock. Fig. 1. Multiflora rose. A) berrylike hips, B)leaf, note pectinate stipules (arrow), C) stem (cane). II. History: The genus Rosa is a large group of plants comprised of about 150 species, of which one-third are indigenous to America. Gray's Manual of Botany (Fernald 1970) lists 24 species (13 native; 11 introduced, 10 of these fully naturalized) for our range. Gleason and Cronquist (l968) cite 19 species (10 introductions). The disagreement in the potential number of species encountered in Pennsylvania arises from the confused taxonomy of a highly variable and freely crossing group. In fact, there are probably 20,000 cultivars of Rosa known. Bailey (1963) succinctly states the problem: "In no other genus, perhaps, are the opinions of botanists so much at variance in regard to the number of species." The use of roses by mankind has a long history. The Romans acquired a love for roses from the Persians. After the fall of Rome, roses were transported by the Benedictine monks across the Alps, and by the 700's AD garden roses were growing in southern France. The preservation and expansion of these garden varieties were continued by monasteries and convents from whence they spread to castle gardens and gradually to more humble, secular abodes. -
Principles of Plant Taxonomy Bot
PRINCIPLES OF PLANT TAXONOMY BOT 222 Dr. M. Ajmal Ali, PhD 1 What is Taxonomy / Systematics ? Animal group No. of species Amphibians 6,199 Birds 9,956 Fish 30,000 Mammals 5,416 Tundra Reptiles 8,240 Subtotal 59,811 Grassland Forest Insects 950,000 Molluscs 81,000 Q: Why we keep the stuffs of our home Crustaceans 40,000 at the fixed place or arrange into some Corals 2,175 kinds of system? Desert Others 130,200 Rain forest Total 1,203,375 • Every Human being is a Taxonomist Plants No. of species Mosses 15,000 Ferns and allies 13,025 Gymnosperms 980 Dicotyledons 199,350 Monocotyledons 59,300 Green Algae 3,715 Red Algae 5,956 Lichens 10,000 Mushrooms 16,000 Brown Algae 2,849 Subtotal 28,849 Total 1,589,361 • We have millions of different kind of plants, animals and microorganism. We need to scientifically identify, name and classify all the living organism. • Taxonomy / Systematics is the branch of science deals with classification of organism. 2 • Q. What is Plant Taxonomy / Plant systematics We study plants because: Plants convert Carbon dioxide gas into Every things we eat comes Plants produce oxygen. We breathe sugars through the process of directly or indirectly from oxygen. We cannot live without photosynthesis. plants. oxygen. Many chemicals produced by the Study of plants science helps to Study of plants science helps plants used as learn more about the natural Plants provide fibres for paper or fabric. to conserve endangered medicine. world plants. We have millions of different kind of plants, animals and microorganism. -
Cupressaceae Et Taxodiaceae
AVERTISSEMENT Ce document est le fruit d'un long travail approuvé par le jury de soutenance et mis à disposition de l'ensemble de la communauté universitaire élargie. Il est soumis à la propriété intellectuelle de l'auteur. Ceci implique une obligation de citation et de référencement lors de l’utilisation de ce document. D'autre part, toute contrefaçon, plagiat, reproduction illicite encourt une poursuite pénale. Contact : [email protected] LIENS Code de la Propriété Intellectuelle. articles L 122. 4 Code de la Propriété Intellectuelle. articles L 335.2- L 335.10 http://www.cfcopies.com/V2/leg/leg_droi.php http://www.culture.gouv.fr/culture/infos-pratiques/droits/protection.htm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
Some Rose Species
ARNOLD ARBORETUM HARVARD UNIVERSITY BULLETIN OF POPULAR INFORMATION SERIES 4. VOL. VIII JUNE 28, 1940 NUMBER 6 SOME ROSE SPECIES are too little used in our present day landscaping. There mayROSES be several reasons for this, but after a careful examination of the types available, one is not surprised to find that there are cer- tain species which deserve a place in almost every garden. Attention is here called to some of the more outstanding rose species for use in modern gardens. The exact number of rose species has been a much debated point for many years. In 1892 Gandoger recognized 4266 species, while a few years before Bentham and Hooker recognized only 30-a rather striking difference of opinion! Be this as it may, Professor Alfred Rehder suggests that there are probably between 150 and 200 species of roses in the temperate and subtropical regions of the northern hem- isphere. On the basis of Rehder’s estimate, it is of interest to note that over one half of these occur in the living collections at the Arnold Arboretum, and that one nursery (Bobbink and Atkins) has recently listed 93 species and botanical varieties as being grown by them and ready for sale in the form of two year old, field grown plants. With these figures as a background, let us examine some of the im- portant reasons why more of these rose species should be grown in the garden. In the first place, as these are the wild roses of the world, they are entitled to a place in any wild garden, in the woodlands or at the border of roadways and woods. -
(Sarracenia) Provide a 21St-Century Perspective on Infraspecific Ranks and Interspecific Hybrids: a Modest Proposal* for Appropriate Recognition and Usage
Systematic Botany (2014), 39(3) © Copyright 2014 by the American Society of Plant Taxonomists DOI 10.1600/036364414X681473 Date of publication 05/27/2014 Pitcher Plants (Sarracenia) Provide a 21st-Century Perspective on Infraspecific Ranks and Interspecific Hybrids: A Modest Proposal* for Appropriate Recognition and Usage Aaron M. Ellison,1,5 Charles C. Davis,2 Patrick J. Calie,3 and Robert F. C. Naczi4 1Harvard University, Harvard Forest, 324 North Main Street, Petersham, Massachusetts 01366, U. S. A. 2Harvard University Herbaria, Department of Organismic and Evolutionary Biology, 22 Divinity Avenue, Cambridge, Massachusetts 02138, U. S. A. 3Eastern Kentucky University, Department of Biological Sciences, 521 Lancaster Avenue, Richmond, Kentucky 40475, U. S. A. 4The New York Botanical Garden, 2900 Southern Boulevard, Bronx, New York 10458, U. S. A. 5Author for correspondence ([email protected]) Communicating Editor: Chuck Bell Abstract—The taxonomic use of infraspecific ranks (subspecies, variety, subvariety, form, and subform), and the formal recognition of interspecific hybrid taxa, is permitted by the International Code of Nomenclature for algae, fungi, and plants. However, considerable confusion regarding the biological and systematic merits is caused by current practice in the use of infraspecific ranks, which obscures the meaningful variability on which natural selection operates, and by the formal recognition of those interspecific hybrids that lack the potential for inter-lineage gene flow. These issues also may have pragmatic and legal consequences, especially regarding the legal delimitation and management of threatened and endangered species. A detailed comparison of three contemporary floras highlights the degree to which infraspecific and interspecific variation are treated inconsistently. -
Anatomy and Go Fish! Background
Anatomy and Go Fish! Background Introduction It is important to properly identify fi sh for many reasons: to follow the rules and regulations, for protection against sharp teeth or protruding spines, for the safety of the fi sh, and for consumption or eating purposes. When identifying fi sh, scientists and anglers use specifi c vocabulary to describe external or outside body parts. These body parts are common to most fi sh. The difference in the body parts is what helps distinguish one fi sh from another, while their similarities are used to classify them into groups. There are approximately 29,000 fi sh species in the world. In order to identify each type of fi sh, scientists have grouped them according to their outside body parts, specifi cally the number and location of fi ns, and body shape. Classifi cation Using a system of classifi cation, scientists arrange all organisms into groups based on their similarities. The fi rst system of classifi cation was proposed in 1753 by Carolus Linnaeus. Linnaeus believed that each organism should have a binomial name, genus and species, with species being the smallest organization unit of life. Using Linnaeus’ system as a guide, scientists created a hierarchical system known as taxonomic classifi cation, in which organisms are classifi ed into groups based on their similarities. This hierarchical system moves from largest and most general to smallest and most specifi c: kingdom, phylum, class, order, family, genus, and species. {See Figure 1. Taxonomic Classifi cation Pyramid}. For example, fi sh belong to the kingdom Animalia, the phylum Chordata, and from there are grouped more specifi cally into several classes, orders, families, and thousands of genus and species. -
An Application to Biology
Physical Science & Biophysics Journal ISSN: 2641-9165 Information as Order Hidden within Chance: An Application to Biology Strumia A* Review Article Istituto Nazionale di Alta Matematica "Francesco Severi", Italy Volume 3 Issue 3 *Corresponding author: Alberto Strumia, Istituto Nazionale di Alta Matematica Received Date: August 12, 2019 Published Date: August 27, 2019 "Francesco Severi", Italy, Email: [email protected] Abstract We show, by didactical examples, how algorithmic information (coded e.g., into a computer program) is required to build the structure of an organized system (either simple or complex). Ordered structures can be obtained as attractors both by some dynamics starting from sequential initial conditions (order from order) and by some dynamics starting from random initial conditions (order from chance) provided that a leading algorithmic information is assigned to govern the evolution of the generating process. In absence of information emergence of some ordered structure, like e.g., an organ of a living system is so highly improbable to be impossible in practice. We provide didactical examples of static models of a human heart, each generated starting either from ordered initial conditions, or from random sparse initial conditions, or more realistically by random cellular automata (so that any mother cell is allowed to generate a daughter cell only in a random contiguous location). Significantly, as it was pointed out by Gregory Chaitin, not all algorithmic information can be compressed into a string shorter than the sequence of its original individual code digits (incompressible information string). A question is still open about the DNA and, more generally, any biological information: is it to be considered as a compressible or an incompressible code string? In our example of anatomic human heart model we have treated the sequence of the co-ordinates of each sphere (roughly modeling a cell) as an uncompressed string, while a compressed program string seems to be able to provide only less realistic models. -
Life History Variation Between High and Low Elevation Subspecies of Horned Larks Eremophila Spp
J. Avian Biol. 41: 273Á281, 2010 doi: 10.1111/j.1600-048X.2009.04816.x # 2010 The Authors. J. Compilation # 2010 J. Avian Biol. Received 29 January 2009, accepted 10 August 2009 Life history variation between high and low elevation subspecies of horned larks Eremophila spp. Alaine F. Camfield, Scott F. Pearson and Kathy Martin A. F. Camfield ([email protected]) and K. Martin, Centr. for Appl. Conserv. Res., Fac. of Forestry, Univ. of British Columbia, 2424 Main Mall, Vancouver, B.C., Canada, V6T 1Z4. AFC and KM also at: Canadian Wildlife Service, Environment Canada, 351 St. Joseph Blvd., Gatineau, QC K1A 0H3. Á S. F. Pearson, Wildl. Sci. Div., Washington Dept. of Fish and Wildl., 1111 Washington St. SE, Olympia, WA, USA, 98501-1091. Environmental variation along elevational gradients can strongly influence life history strategies in vertebrates. We investigated variation in life history patterns between a horned lark subspecies nesting in high elevation alpine habitat Eremophila alpestris articola and a second subspecies in lower elevation grassland and sandy shoreline habitats E. a. strigata. Given the shorter breeding season and colder climate at the northern alpine site we expected E. a. articola to be larger, have lower fecundity and higher apparent survival than E. a. strigata. As predicted, E. a. articola was larger and the trend was toward higher apparent adult survival for E. a. articola than E. a. strigata (0.69 vs 0.51). Contrary to our predictions, however, there was a trend toward higher fecundity for E. a. articola (1.75 female fledglings/female/year vs 0.91). -
Terrapene Carolina (Linnaeus 1758) – Eastern Box Turtle, Common Box Turtle
Conservation Biology of Freshwater Turtles and Tortoises: A Compilation Project ofEmydidae the IUCN/SSC — TortoiseTerrapene and Freshwatercarolina Turtle Specialist Group 085.1 A.G.J. Rhodin, P.C.H. Pritchard, P.P. van Dijk, R.A. Saumure, K.A. Buhlmann, J.B. Iverson, and R.A. Mittermeier, Eds. Chelonian Research Monographs (ISSN 1088-7105) No. 5, doi:10.3854/crm.5.085.carolina.v1.2015 © 2015 by Chelonian Research Foundation • Published 26 January 2015 Terrapene carolina (Linnaeus 1758) – Eastern Box Turtle, Common Box Turtle A. ROSS KIESTER1 AND LISABETH L. WILLEY2 1Turtle Conservancy, 49 Bleecker St., Suite 601, New York, New York 10012 USA [[email protected]]; 2Department of Environmental Studies, Antioch University New England, 40 Avon St., Keene, New Hampshire 03431 USA [[email protected]] SUMMARY. – The Eastern Box Turtle, Terrapene carolina (Family Emydidae), as currently understood, contains six living subspecies of small turtles (carapace lengths to ca. 115–235 mm) able to close their hinged plastrons into a tightly closed box. Although the nominate subspecies is among the most widely distributed and well-known of the world’s turtles, the two Mexican subspecies are poorly known. This primarily terrestrial, though occasionally semi-terrestrial, species ranges throughout the eastern and southern United States and disjunctly in Mexico. It was generally recognized as common in the USA throughout the 20th century, but is now threatened by continuing habitat conversion, road mortality, and collection for the pet trade, and notable population declines have been documented throughout its range. In the United States, this turtle is a paradigm example of the conservation threats that beset and impact a historically common North American species.