Survey of the Herbarium Resources of Ohio1

SURVEY OF THE HERBARIUM RESOURCES OF OHIO1

ALLISON W. CUSICK, Division of Natural Areas and Preserves, Ohio Department of Natural Resources, Fountain Square, Columbus, OH 43224 J. A. SNIDER, Department of Biological Sciences, University of Cincinnati, Cincinnati, OH 45221

ABSTRACT. More than one million specimens of vascular and nonvascular plants are housed in Ohio herbaria. These collections are an invaluable scientific resource. Only 13 Ohio herbaria are listed in the Index Herbariorum. This survey identifies 37 Ohio herbaria and summarizes their holdings. Details given for these herbaria include: names and addresses of curators, hours of access, strengths of the collections, and research activities of the staff.

OHIO J. SCI. 84 (4): 175-188, 1984

INTRODUCTION Only 13 Ohio herbaria are listed in the Herbaria are invaluable scientific re- Index Herbariorum (Holmgren et al. 1981). sources for study of the more than one- Thirty-seven Ohio herbaria are represented quarter million plant species in the world. in our survey. Our listing also attempts In essence, herbaria are reference libraries. to provide the researcher with enough de- The specimens housed in these collections tails on the holdings of the herbarium so help answer questions on the disappearance that his or her time and energies may of species, changing distributions, and be used to the best advantage. It is hoped evolutionary patterns of divergence and that these descriptions will allow a student reticulation. Herbarium records lead re- of a particular area or taxonomic group to searchers down new pathways toward identify holdings which deserve critical development and utilization of food and examination. medicinal plants, economically signifi- The specific stimulus for the survey cant plant materials, and aesthetic horti- was the organization of a symposium on cultural innovations. Ohio's herbarium resources sponsored The herbaria of Ohio are numerous and by The Ohio Academy of Science. This contain a diversity of holdings. These symposium was held at The Ohio State collections constitute a significant sci- University in conjunction with the annual entific resource documenting the flora of meeting of the Academy, 24 April 1982. Ohio and the world and embracing all ele- Our survey at that time appeared in a ments of the plant kingdom. Collectively separate publication which was distri- they contain more than one million speci- buted to attendees at the symposium mens with a conservative value of over (Cusick and Snider 1982). The present $7,500,000. version is slightly revised from the pre- The present survey of these collections vious publication. stems from the absence of any single listing for all of them and from the need for METHODS AND MATERIALS communication among these institutions. In January and February 1982 the authors sent questionnaires to 52 Ohio institutions. The list of Revised and updated from a presentation at a institutions was assembled from four sources: symposium, "The Herbarium Resources of Ohio: Holmgren et al. (1981), Ohio Natural Heritage Present Status and Future Directions," organized for Report (1977), Peterjohn and Walker (1978), and the meetings of The Ohio Academy of Science, the personal knowledge of the authors. Positive Columbus, Ohio, 24 April 1982. Manuscript re- responses were received from 37 different herbaria. ceived 26 April 1984 and in revised form 6 August A summary of these replies constitutes the bulk of 1984 (#84-20). this paper.

175 176 A. W. CUSICK AND J. A. SNIDER Vol. 84

We are aware that this survey is not complete. A Strengths: Late 19th century collections few institutions with herbaria are not represented, by Edward Claypole from the Akron area; either becuase they failed to respond to the authors' questionnaires and follow-up efforts, or because the local flora. existence of the collection was not known to the NONVASCULAR HOLDINGS: None reported. authors. Large private collections also may not be ACCESS: By appointment. included Ar similar reasons. Some important col- Teaching and reference collection. lections of fossil plants probably have been over- looked since these may be housed in departments of ANTIOCH COLLEGE geology rather than botany. Department of Biology Thirty-one of the herbaria included in our survey Yellow Springs, OH 45387 are academic collections and six are housed at non- academic institutions. This proportion is representa- STAFF: Walter Tulecke, Professor of Biology tive of the disposition of herbaria nationally. Of the (513) 767-7331 approximately 230 public herbaria in the United VASCULAR HOLDINGS: States about 75% are academic collections (Beaman 1965). Beaman arbitrarily classified herbaria into Ohio—500 three broad groups: small, with fewer than 100,000 Strengths: Local flora; flora of Clifton specimens; medium, with between 100,000 and Gorge and Glen Helen, Greene County, 500,000 specimens; and large, with more than Ohio. 500,000 specimens. Using these criteria, Ohio has but a single large herbarium, The Ohio State NONVASCULAR HOLDINGS: None reported. University, one medium-sized herbarium, Miami ACCESS: By appointment. University, and 35 small collections. EQUIPMENT: Dissecting microscopes. RESULTS Teaching and reference collection. The listings which follow are arranged BALDWIN-WALLACE COLLEGE alphabetically by institution. The code Department of Biological Sciences is the accepted acronym from 7th ed. Berea, OH 44017 Index Herbariorum. Information given in STAFF: Donald S. Dean, Department any category is that supplied by the re- Chairman spondents. The headings largely are (216) 826-2262 self-explanatory. Absence of a heading in- VASCULAR HOLDINGS: dicates that no information on that subject Ohio —3,900 was provided. The heading "Equipment" North America—300 lists what the visitor may expect to find Total —4,200 available for his or her use. An affirmative Strengths: Flora of northern Ohio. response to "Mail Loans" means that the NONVASCULAR HOLDINGS: usual herbarium procedures for such loans Algae: None reported. are followed by that institution. At the end Fungi: Approximately 300 speci- of each listing is a sentence or two sum- mens of myxomycetes forming the foun- marizing the general use and activities of dation for E. L. Fullmer's study of the that herbarium and any special research slime molds of Ohio. Maintained in boxes. interests mentioned by the respondent. All Lichen: None reported. replies to the questionnaires have been Bryophytes: Approximately 130 edited for conciseness. specimens from Ohio; in packets. Un- UNIVERSITY OF AKRON known number of specimens from outside Department of Biology Ohio; mounted on cards. Akron, OH 44325 ACCESS: By appointment. STAFF: W. P. Stoutamire, Curator EXCHANGES: Yes. (216) 375-7162 Teaching and reference collection. VASCULAR HOLDINGS: BOWLING GREEN STATE Ohio —2,328 UNIVERSITY North America— 1,170 Edwin Lincoln Moseley Herbarium Eastern Hemisphere — 97 Department of Biological Sciences Total—3,595 Bowling Green, OH 43403 OhioJ. Sci. SURVEY OF OHIO HERBARIUM RESOURCES 177

CODE: BGSU Lichens: Approximately 7,000 STAFF: N. William Easterly, Curator and specimens. Southeastern and southwestern Professor of Botany Ohio are especially well represented; also (419) 372-2434 Kentucky, Michigan. Other U.S. holdings VASCULAR HOLDINGS: are represented by several hundred speci- Ohio—19,000 mens. Important collection of the genus North America—1,000 Cladonia studied by Alexander Evans. Eastern Hemisphere—500 Maintained in packets. Total —20,500 Bryophytes: Approximately Strengths: Flora of the Oak Openings 22,000 specimens. Unknown number region of northwest Ohio; collections of yet to be accessioned. Worldwide repre- N. William Easterly and E. L. Moseley. sentation, strong in U.S. and Canada NONVASCULAR HOLDINGS: None reported. collections. Particularly strong in south- ACCESS: 8 AM to 5 PM, year-round, or by eastern and southwestern Ohio collections, appointment. Michigan and Kentucky. Especially strong EQUIPMENT: Dissecting microscope, in North American Sphagnum collections. manuals. Numerous Exsiccati represented. The per- MAIL LOANS: Yes. sonal herbarium of Dr. Margaret Fulford EXCHANGES: Yes, with other Ohio her- (ca. 8,000 specimens) is housed and main- baria. tained separately by Dr. Fulford (address RESEARCH INTERESTS: Flora of the Oak as above). This collection very strong in Openings of northwest Ohio. Carribbean and South American hepatics. Many type specimens. Specimens maintained in packets. UNIVERSITY OF CINCINNATI INCIDENTAL HOLDINGS: Assorted seed and Department of Biological Sciences wood collections. Cincinnati, OH 45221 ACCESS: Monday-Friday, 8 AM to 5 PM, CODE: CINC September—June, or by appointment. STAFF: Jerry A. Snider, Curator EQUIPMENT: Dissecting microscope, (513)475-3741 library. Victor Soukup, Assistant Curator MAIL LOANS: Yes. (513)475-3741 EXCHANGES: By arrangement. VASCULAR HOLDINGS: RESEARCH ACTIVITIES: Flora of southwest Ohio—5,000 Ohio and adjacent Kentucky; systematics North America—25,000 of Orchidaceae and Trillium; bryophyte Western Hemisphere— 1,800 studies; lichen studies. Eastern Hemisphere — 20,000 NOTE: The herbarium of the Lloyd Library Total —51,800 of Cincinnati, about 25,000 sheets, Strengths: Flora of southwest Ohio and is on permanent loan to this her- adjacent Kentucky; Kentucky collection of barium and is housed therein. The E. Lucy Braun; herbarium of Lloyd Library international moss exchange for of Cincinnati. members of the American Bryo- NONVASCULAR HOLDINGS: logical and Lichenological Society, Algae: Approximately 1,000 speci- Inc., is currently located here. mens. Mostly miscellaneous North Ameri- can collections. Maintained on sheets, CLEVELAND MUSEUM OF packets, slides. NATURAL HISTORY Fungi: Approximately 5,000 Wade Oval, University Circle specimens, about 4,300 of these from Cleveland, OH 44106 Ohio. Holdings include approximately CODE: CLM 2,000 pre-1937 Ohio collections of Wil- STAFF: James K. Bissell, Curator of Botany liam Bridge Cooke. (216) 231-4600 178 A. W. CUSICK AND J. A. SNIDER Vol. 84

VASCULAR HOLDINGS: Reference and research collection of Ameri- Ohio —22,000 can Ivy Society. North America—5,000 Western Hemisphere—500 CUYAHOGA COMMUNITY Eastern Hemisphere— 1,500 COLLEGE Total —29,000 Eastern Campus Strengths: Flora of northeast Ohio; 4250 Richmond Rd. collections of Leslie D. Stair and James K. Warrensville Twp., OH 44122 Bissell. STAFF: Barbara K. Andreas, Professor of NONVASCULAR HOLDINGS: Biology Algae: None reported. (216) 292-2389 Fungi: 250 specimens. Local col- VASCULAR HOLDINGS: lections and European specimens included. Ohio—1,000 Maintained on sheets, packets and boxes. Strengths: Spring flora of northeast Lichens: 200 specimens. Mostly Ohio. local collections, some European speci- NONVASCULAR HOLDINGS: None reported. mens included. Maintained in boxes. EQUIPMENT: Dissecting microscopes, Bryophytes: None reported. manuals. INCIDENTAL HOLDINGS: Worldwide col- Teaching and reference collection. lection of 2,000 wood samples. ACCESS: Tuesday-Friday, 10 AM to 5 PM, THE DAWES ABORETUM year—round. 7770 Jacksontown Rd. SE EQUIPMENT: Binocular microscopes. Newark, OH 43055 MAIL LOANS: Yes. STAFF: Charles T. Greaves, Botanist EXCHANGES: Yes. (614) 323-2355; (614) 323-2990 RESEARCH INTERESTS: Vascular flora of VASCULAR HOLDINGS: Total estimated northeast Ohio. holdings —700. NOTE: Most nonvascular holdings in stor- Strengths: Cultivated woody ornamen- age and difficult of access. tals, ferns and prairie species, all from the Arboretum's living collection. COX ARBORETUM NONVASCULAR HOLDINGS: None reported. 6733 Springboro Pike ACCESS: Monday-Friday, 7:30 AM to Dayton, OH 45449 4:30 PM, year—round, or by appointment. STAFF: Marvin Olinsky, Director EQUIPMENT: Dissecting microscope. (513)434-9005 MAIL LOANS: Yes. Amy J. Munich, Education EXCHANGES: Yes, if related to Arboretum Supervisor living collection. Sabina Sulgrove, Director of Reference collection of species growing American Ivy Society and on grounds of Arboretum. Research Center Pat Trapp, Herbarium Specialist UNIVERSITY OF DAYTON VASCULAR HOLDINGS: No number Department of Biology provided. Dayton, OH 45469 Strengths: The genus Hedera; Ameri- STAFF: Joseph D. Laufersweiler, Associate can Ivy Society Collection; vouchers of Professor of Biology new cultivars. (513) 229-2521 NONVASCULAR HOLDINGS: None reported. VASCULAR HOLDINGS: ACCESS: By appointment. Ohio —600 EQUIPMENT: Dissecting microscope. North America— 100 MAIL LOANS: Yes. Western Hemisphere—50 EXCHANGES: Limited. Total —750 Ohio J. Sci. SURVEY OF OHIO HERBARIUM RESOURCES 179

Strengths: Flora of the Dayton area; mostly from southwestern and south- Swiss alpine flora. central Ohio. Maintained in packets. NONVASCULAR HOLDINGS: INCIDENTAL HOLDINGS: Miscellaneous Algae: Approximately 100 speci- collections of plant galls, seeds, and winter mens. Coastal Florida collection con- twigs. taining primarily red and brown algae. ACCESS: Monday-Friday, 9 AM to 6 PM, Mounted on sheets. year—round, except holidays. Fungi: None reported. EQUIPMENT: Binocular microscope; li- Lichens: None reported. brary of over 10,000 volumes. Bryophytes: None reported. MAIL LOANS: Yes. ACCESS: By appointment. EXCHANGES: Yes; Ohio material for that EQUIPMENT: Microscopes. from eastern North America. MAIL LOANS: Yes. RESEARCH INTERESTS: Flora of Mont- EXCHANGES: No. gomery County, Ohio. Teaching and reference collection. DAYTON MUSEUM OF NATURAL DENISON UNIVERSITY HISTORY Department of Biology 2629 Ridge Ave. Granville, OH 43023 Dayton, OH 45414 CODE: DEN CODE: DMNH STAFF: Juliana C. Mulroy, Assistant Pro- STAFF: Gary A. Coovert, Curator of Biology fessor of Biology (513) 275-7432 (614) 587-6327; VASCULAR HOLDINGS: (614) 587-0810, ext. 261 Ohio —4,116 VASCULAR HOLDINGS: North America — 2,449 Ohio —5,000 Western Hemisphere — 6,565 North America — 2,500 Eastern Hemisphere— 105 Western Hemisphere—500 Total—13,235 Eastern Hemisphere— 100 Strengths: Flora of Montgomery Total —8,100 County, Ohio and adjacent areas; col- Strengths: Worldwide fern collec- lections of John W. VanCleve and William tion of Wallace Cathcart; flora of Licking B. Werthner. County, Ohio. NONVASCULAR HOLDINGS: NONVASCULAR HOLDINGS: Algae: 35 specimens. Mostly algae Algae: Approximately 200 speci- from Florida. Maintained in jars. mens. Poorly documented, used primarily Fungi: 736 specimens. Collection as a teaching collection. Mounted on sheets. consists primarily of fleshy fungi collected Fungi: Approximately 50 speci- during the past 10—15 years in south- mens. Local collections representing stu- western and southcentral Ohio. Complete dent projects. Maintained in boxes. collection data, many specimens accom- Lichens: 92 specimens. All Ohio panied by spore prints. Maintained in collections. Maintained in packets. boxes, jars. Bryophytes: 43 specimens. All Lichens: 69 specimens. Mostly Ohio collections. Maintained in packets. unidentified collections with complete ACCESS: By appointment. collection data primarily representing EQUIPMENT: Laboratory space, micro- southwestern and southcentral Ohio. scopes. Maintained in packets. MAIL LOANS: Yes. Bryophytes: 176 specimens. Col- EXCHANGES: Yes. lections largely unidentified but with Teaching and reference collection; research complete collection data. Specimens on the Licking County flora. 180 A. W. CUSICK AND J. A. SNIDER Vol. 84

FRANZ THEODORE Western Hemisphere— 10 STONE LABORATORY Total—1610 The Ohio State University Strengths: Flora of northeast Ohio; Put-in-Bay, OH 43456 spring flora. CODE: FTSL NONVASCULAR HOLDINGS: None reported. STAFF: Ronald L. Stuckey, Associate Direc- INCIDENTAL HOLDINGS: About 100 fos- tor for Instruction sil specimens, mostly from Ohio and Franz Theodore Stone Laboratory Pennsylvania. Professor of Botany, ACCESS: Monday-Friday, 9 AM to 5 PM, The Ohio State University, September—May; summer hours by Department of Botany appointment. 1735 Neil Ave. EQUIPMENT: Microscopes. Columbus, OH 43210 MAIL LOANS: Yes. (614) 422-6095 EXCHANGES: No. VASCULAR HOLDINGS: Teaching and reference collection. Ohio —2,500 to 3,000 THE HOLDEN ARBORETUM Strengths: Local flora of the Lake Erie 9500 Sperry Road Islands and adjacent Erie and Ottawa Mentor, OH 44060 Counties, Ohio. NONVASCULAR HOLDINGS: None reported. The former herbarium of Case Western ACCESS: 20 June to 30 August. Reserve University of Cleveland, Ohio, MAIL LOANS: No formal procedure. was donated to this institution. The col- EXCHANGES: No. lection is uncurated and at present is inac- Teaching collection; research on flora of the cessible to researchers. Lake Erie Islands. For information contact: Peter W. Bristol, Horticulturist (216) 946-4400. HEIDELBERG COLLEGE KENT STATE UNIVERSITY Biology Department Department of Biological Sciences Tiffin, OH 44883 Kent, OH 44242 STAFF: Percy L. Lilly, Professor of Biology CODE: KE (419) 448-2693 STAFF: Tom S. Cooperrider, Curator of VASCULAR HOLDINGS: Vascular Plants Ohio—1,000 (216) 672-3613 North America — 200 Samuel J. Mazzer, Curator of Non- Total—1,200 vascular Plants Strengths: Flora of Crawford and Seneca (216) 672-3613 Counties, Ohio; collections of Ronald L. Miwako K. Cooperrider, Herbar- Stuckey. ium Assistant NONVASCULAR HOLDINGS: None reported. (216) 672-3613 INCIDENTAL HOLDINGS: Special collection VASCULAR HOLDINGS: of 125 representative woody species by Ohio —43,000 R. E. Eshmeyer. North America — 6,000 ACCESS: By appointment. Rest of world— 1,000 Teaching and reference collection. Total —50,000 Strengths: Flora of Ohio; collections HIRAM COLLEGE of Barbara Andreas, Tom S. Cooperrider, Biology Department Allison W. Cusick, Almon N. Rood, and Hiram, OH 44234 Gene M. Silberhorn; flora of Cuyahoga VASCULAR HOLDINGS: Valley National Recreation Area; flora of Ohio—1,200 the following Ohio counties — Ashland, North America — 400 Geauga, Holmes, Jefferson, Knox, Me- OhioJ. Sci. SURVEY OF OHIO HERBARIUM RESOURCES 181 dina, Perry, Portage, Stark, Summit, STAFF: Stephen Diakoff, Curator Trumbull and Wayne. (216) 489-0800, ext. 499 NONVASCULAR HOLDINGS: VASCULAR HOLDINGS: Total estimated Algae: None reported. holdings—300 to 400. Fungi: Approximately 6,000 speci- Strengths: Unknown. mens. Collections consist primarily of NONVASCULAR HOLDINGS: None reported. fleshy fungi representing Great Lakes INCIDENTAL HOLDINGS: Miscellaneous region and eastern U.S. Especially good specimens of fossil plants. representation of fleshy fungi of Arcto— ACCESS: During school hours. Tertiary geoflora. Primary interest is in EXCHANGES: No. ectomycorrhizal taxa. Maintained in boxes. Teaching and reference collection. Lichens: Approximately 2,000 MARIETTA COLLEGE specimens. Collections mostly from North America with a small number Biology Department from Mexico, Central America, Japan and Marietta, OH 45750 Europe. Maintained in packets. STAFF: David Young, Professor of Biology Bryophytes: Approximately 2,000 (614) 373-4643 specimens. Collections mostly representa- Marilyn Ortt, Volunteer tive of Great Lakes Forest region and east- 701 Colegate, Marietta, OH 45750 ern U.S. Some western U.S. collections (614) 373-3372 also present. Maintained in packets. VASCULAR HOLDINGS: ACCESS: Monday-Thursday, 8:30 AM Ohio —3,000 to 11:30 AM, September—May, or by North America—200 appointment. Total—3,200 EQUIPMENT: Standard. Strengths: Local herbarium by John L. MAIL LOANS: Yes. Riddell (1830s); fern collection of George EXCHANGES: To a limited extent, chiefly Proctor; flora of southeast Ohio and adja- of Ohio specimens; also interested in cent West Virginia. exchanges for Central American and East NONVASCULAR HOLDINGS: None reported. Asian fungi. ACCESS: By appointment. RESEARCH INTERESTS: Flora of Ohio; my- EQUIPMENT: Worktable, microscopes. cological studies. MAIL LOANS: Yes. EXCHANGES: No. LAKELAND Teaching and reference collection; research COMMUNITY COLLEGE on the flora of southeast Ohio. Department of Biological Sciences MIAMI UNIVERSITY Rt. 306 and 1-90 Willard Sherman Turrell Herbarium Mentor, OH 44060 Department of Botany STAFF: Edward J. P. Hauser, Department Oxford, OH 45056 Chairperson CODE: MU (216) 953-7151 STAFF: W. Hardy Eshbaugh, Curator and VASCULAR HOLDINGS: Professor of Botany Ohio —500 Strengths: Flora of northeast Ohio. (513) 529-4934 NONVASCULAR HOLDINGS: None reported. Gene P. Williamson, Technician ACCESS: By appointment. (513) 529-2755 EXCHANGES: No. VASCULAR HOLDINGS: Total estimated Teaching and reference collection. holdings—131,000 Strengths: Non—Ohio material formerly MALONE COLLEGE in herbarium of Oberlin College; collec- Department of Science and Mathem? ICS tion of western U.S. plants of Percy Train; Canton, OH 44709 flora of Butler County, Ohio; New Zealand 182 A. W. CUSICK AND J. A. SNIDER Vol. 84

ferns; collections from Brazil, South Africa, Total —23,500 and Tierra del Fuego. Strengths: Ferns and fern allies col- NONVASCULAR HOLDINGS: lected by Mr. and Mrs. William Adams, Algae: Approximately 2,000 speci- Jr.; flora of Muskingum County, Ohio. mens. U.S. mostly represented, some inter- NONVASCULAR HOLDINGS: None reported. national collections also present. Several ACCESS: Monday-Friday, 8 AM to 5 PM, exsiccati specimens represented. September—May, and by appointment. Fungi: 34,500 specimens, exclud- EQUIPMENT: Stereomicroscopes. ing exsiccati. Additional collections yet to MAIL LOANS: Yes. be accessioned. Worldwide representation, EXCHANGES: Yes; one—for—one basis. including approximately 1,000 specimens Teaching and reference collection. from Ohio collected by Bruce Fink and 6,000+ specimens from the collection of OBERLIN COLLEGE William Bridge Cooke. Maintained in Department of Biology packets, boxes. Kettering Hall of Science Lichens: 8,416 specimens. Addi- Oberlin, OH 44074 tional collections yet to be accessioned. CODE: OC Worldwide representation. A portion of STAFF: George T. Jones, Curator and Pro- Bruce Fink's lichen collection is housed fessor Emeritus here. Maintained in packets, boxes. (216) 775-8316 Bryophytes: 25,654 specimens. VASCULAR HOLDINGS: Additional collections yet to be acces- Ohio —26,066 sioned. Worldwide representation. Good North America—37 representation of specimens from Japan, Total —26,103 Micronesia. Harvey A. Miller collection Strengths: Flora of Lorain County, (1956-1967) deposited here. Numerous Ohio and adjacent areas; collections exsiccati specimens also present. Main- of F. O. Grover, George T. Jones and tained in packets. E. Wilkinson. INCIDENTAL HOLDINGS: Reference slide NONVASCULAR HOLDINGS: collection of anatomical materials; ca. Algae: 172 specimens, primarily 2,500 specimens of plant fossils. from Ohio. Maintained in packets, on ACCESS: Monday-Friday, 8 AM to 4:30 sheets. PM, year—round, or by appointment. Fungi: 5,347 specimens, 4,500 EQUIPMENT: Laboratory space, micro- from Ohio; 2,228 specimens of the family scopes, camera attachments, library. Polyporaceae. 300 specimens of myxo- MAIL LOANS: Yes. mycetes. Maintained in packets, on sheets. EXCHANGES: Yes; specimens also pur- Lichens: 346 specimens. Collec- chased. tions from Lorain, Erie and Huron Count- RESEARCH INTERESTS: Butler County, Ohio ies. In packets. flora; plant anatomy; systematics of Cap- Bryophytes: 1,047 specimens. sicum, Empetraceae and Myristicaceae. Mostly representative of Lorain, Erie and Huron Counties. In packets. MUSKINGUM COLLEGE INCIDENTAL HOLDINGS: Special ecological Biology Department collection of dominants and subdominants New Concord, OH 43762 of western U.S., about 600 in number; STAFF: Clement E. Dasch, Professor of miscellaneous seed and wood collections of Biology E. Wilkinson. (614) 826-8224 ACCESS: By appointment. VASCULAR HOLDINGS: MAIL LOANS: Yes. Ohio —21,000 Teaching and reference collection; research North America — 2,500 on flora of northern Ohio. OhioJ. Sci. SURVEY OF OHIO HERBARIUM RESOURCES 183

OHIO HISTORICAL SOCIETY CODE: OS Natural History Department STAFF: Tod F. Stuessy, Director 1982 Velma Ave. (614) 422-5200 Columbus, OH 43211 JohnJ. Furlow, Supervisor STAFF: David L. Dyer, Collections Manager (614) 422-3296 (614) 466-1500, ext. 299 Daniel J. Crawford, Curator VASCULAR HOLDINGS: (614) 422-2725 Ohio—1,000 Emanuel D. Rudolph, Curator North America—200 (614) 422-8952 Total—1,200 Ronald L. Stuckey, Curator Strengths: Flora of Leo Petroglyph, (614) 422-6095 Jackson County, and Fort Hill, Highland VASCULAR HOLDINGS: County, Ohio. Ohio—150,000 NONVASCULAR HOLDINGS: None reported. North America (excluding Ohio INCIDENTAL HOLDINGS: Wood collec- and Mexico)—145,000 tion, 300 specimens; seed collection, Mexico and Central 700 specimens. America— 10,000 ACCESS: Monday-Friday, 9 AM to 6 PM, South America—3,500 year—round. Eastern Hemisphere — 2,500 Total —311,000 EQUIPMENT: Laboratory space, microscopes. MAIL LOANS: No. Strengths: The OS Type Collection EXCHANGES: No. presently contains over 200 sheets of vas- Reference collection used for exhibits and cular plants. Some particularly important educational programs. collections preserved here include many holotypes and isotypes of Thomas H. OHIO NORTHERN UNIVERSITY Kearney from the years between 1890 and Department of Biology 1901, holotypes and isotypes of William Ada, OH 45810 A. Kellerman's Guatemalan collections STAFF: Charles C. Laing, Curator of 1905-1908, and isotypes and paratypes (419) 634-9921, ext. 2331 of Underwood and Selby for taxa collected VASCULAR HOLDINGS: Total estimated in Colorado in 1901 and published by holdings —250. P. A. Rydberg. Also included are William Strengths: Unknown. A. Kellerman specimens from Guatemala NONVASCULAR HOLDINGS: (1905-1908); the original specimens of Algae: None reported. Thomas H. Kearney of 1890-1901; Asa Fungi: None reported. Gray's Exsiccatae of North American Lichens: None reported. Gramineae and Cyperaceae (1834-1835); Bryophytes: Approximately 50 John H. Schaffner collection of Equisetum specimens from Hardin County, Ohio and (1900-1935); the pteridophyte herbarium vicinity. of Lewis S. Hopkins; the herbarium of INCIDENTAL HOLDINGS: Oliver Perry the Ohio Agricultural Experiment Station Medsger cone collection. at Wooster, Ohio; the North American ACCESS: By appointment. Appalachian Experimental Watershed EQUIPMENT: Dissecting microscopes. Herbarium prepared by Harold N. Mold- MAIL LOANS: No. enke in the 1940s; the James A. Duke Teaching and reference collection. et al. collection of Panamanian vascular THE OHIO STATE UNIVERSITY plants made during feasibility studies for Department of Botany an Atlantic-Pacific Interoceanic Sea-level 1735 Neil Ave. Canal in Central America (1966-1968); and Columbus, OH 43210 the Tod F. Stuessy et al. collection of Latin 184 A. W. CUSICK AND J. A. SNIDER Vol. 84

American Compositae (1969- )• Ohio plants of the Great Lakes region; flora of collections include those of William S. Ohio; Lichenological studies. Sullivant (1840), Floyd Bartley, Erwin THE OHIO STATE M. Herrick, Allison W. Cusick, Clara G. UNIVERSITY —LIMA CAMPUS Weishaupt and Ronald L. Stuckey. 4240 Campus Drive NONVASCULAR HOLDINGS: Lima, OH 45804 Algae: Approximately 16,500 specimens. Maintained on slides, some STAFF: Roy E. Snyder, Curator mounted specimens. Approximately (419) 228-2641, ext. 367 4,000 slides of fresh water algae of North VASCULAR HOLDINGS: America; approximately 12,000 slides of Ohio—1,000 international diatoms, primarily from North America—2,000 the diatom collection of Fletcher and Total —3,000 Edward Hodges. NONVASCULAR HOLDINGS: Fungi: Approximately 10,000 col- Algae: 150 specimens of brown, lections housed separately. Gilbert slime green and red algae, primarily from Ore- mold collection; W. G. Stover and W. A. gon, mounted on sheets and in liquid. Kellerman fungi collections, numbering Fungi: 175 specimens from Wash- about 5,000 Ohio specimens, of which ington, West Virginia and Canada, main- about 3,000 are plant pathogens; Keller- tained in packets. man collection of Guatemalan fungi; Con- Lichens: None reported. tact Roland Seymour, Aquatic Mycology, Bryophytes: 150 specimens. and C. Wayne Ellett, Parasitic Fungi. ACCESS: By appointment. Lichens: Approximately 14,300 MAIL LOANS: No. specimens. Collections primarily from EXCHANGES: No. Ohio; several from the polar regions. Col- Teaching and reference collection. lections forming part oi Lichens of Ohio by OHIO UNIVERSITY Fr. Conan Taylor housed here. Maintained Bartley Herbarium in packets. Department of Botany Bryophytes: Approximately 10,500 Athens, OH 45701 specimens. Approximately 75% is Ohio material, remainder from North America. CODE: BHO Several exsiccati represented. STAFF: Philip D. Cantino, Curator of Vas- INCIDENTAL COLLECTIONS: Miscellaneous cular Plants wood of Compositae of Latin America; (614) 594-6344 large, but unorganized, seed collection of Warren A. Wistendahl, Curator of North-American species. Nonvascular Plants ACCESS: Monday-Friday, 9 AM to 5 PM, (614) 594-5860 year—round, or by appointment. VASCULAR HOLDINGS: EQUIPMENT: Work tables, dissecting mi- Ohio —25,800 croscopes, annotation labels, library. North America—8,000 MAIL LOANS: Yes. Rest of world — 700 Total —34,500 EXCHANGES: Yes, especially for Latin Strengths: Flora of southeast Ohio; American Compositae; about 35 active collections of Floyd Bartley and Charles exchanges at present. R. Goslin. RESEARCH ACTIVITIES: Taxonomic and NONVASCULAR HOLDINGS: revisionary studies in the Compositae, Algae: None reported. Chenopodiaceae, and Betulaceae; chemo- Fungi: Acrasiomycetes of Ohio, systematics; numerical phenetics; cladis- collections and cultures. Contact James tics; the history of botany; aquatic vascular C. Cavender. OhioJ. Sci. SURVEY OF OHIO HERBARIUM RESOURCES 185

Lichens: None reported. EQUIPMENT: Microscopes, photocopying. Bryophytes: Approximately 2,800 MAIL LOANS: No. specimens. Collections mostly of local and EXCHANGES: No. regional areas; other U.S. also represented. Teaching and reference collection; research Collection includes approximately 200 in cytotaxonomy. hepatics. Floyd Bartley collection is de- URBANA COLLEGE posited here and includes approximately Department of Biology 400 accessioned specimens and nine car- College Way tons yet to be sorted and counted. Urbana, OH 43078 ACCESS: By appointment. EQUIPMENT: Dissecting microscopes, STAFF: Clara May Frederick, Professor of library. Biology MAIL LOANS: Yes. (513) 652-1301 EXCHANGES: Not at present time. Carolyn Circle, Administrative RESEARCH INTERESTS: Systematics of Labi- Assistant atae; flora of southeast Ohio; systematics VASCULAR HOLDINGS: of Ilex; ecological reference collection. Ohio—5,700 NOTE: There is a separate Ohio University North America— 190 Palaeobotanical Herbarium; for Total —5,890 information contact Gar W. Roth- Strengths: Nineteenth century collec- well, address as above. tions of Milo G. Williams, John Samples, and C. W. Short; flora of Cedar Bog, Champaign County, Ohio. OHIO WESLEYAN UNIVERSITY NONVASCULAR HOLDINGS: Department of Botany and Bacteriology Algae: Approximately 300 speci- 40 S. Henry St. mens. Collections mostly represent Cham- Delaware, OH 43015 paign County, Ohio and the western end STAFF: JaneM. Decker, Professor of Botany of Lake Erie; a few specimens from the (614) 369-4431 Caribbean and the Virgin Islands. Pre- VASCULAR HOLDINGS: served in jars. Ohio —4,000 Fungi: Approximately 100 speci- North America — 7,000 mens, mostly from Champaign County, Total—11,000 Ohio. Virginia and West Virginia speci- Strengths: Flora of Delaware County, mens also. Maintained in packets, boxes, Ohio; collections of Rufus Crane and jars. Robert W. Long; North American Com- Lichens: Approximately 75 speci- positae. mens. Collections from Champaign and NONVASCULAR HOLDINGS: Hocking Counties, Ohio. Maintained in Algae: Approximately 25 speci- packets. mens. East coast of U.S. represented. Bryophytes: Approximately 75 Maintained on sheets. specimens. Champaign County, Ohio Fungi: Approximately 700 speci- represented. Maintained in packets. mens. Worldwide representation includ- ACCESS: Monday-Friday, 8 AM to ing international Streptomyces collection of 4:30 PM, September—May, and by 600 taxa. Maintained as cultures. appointment. Lichens: None reported. EQUIPMENT: Dissecting microscopes. Bryophytes: Approximately 300 MAIL LOANS: No. specimens. Miscellaneous U.S. representa- EXCHANGES: No. tion. Maintained on sheets, in boxes. Teaching and reference collections; re- ACCESS: Monday-Friday, 8 AM to 5 PM, search on the flora of the bogs of west- September—June, or by appointment. central Ohio. 186 A. W. CUSICK AND J. A. SNIDER Vol. 84

WALSH COLLEGE CODE: WSFA Department of Biology STAFF: Sture F. Anliot, Assistant Professor 2020 Eastern Ave. of Biology Canton, OH 44720 (513) 382-6661, ext. 237 STAFF: Conrad Gutermuth, Associate Pro- VASCULAR HOLDINGS: fessor of Biology and Department Ohio—1,500 Chairman North America— 100 (216) 499-7090, ext. 22 Western Hemisphere — 200 VASCULAR HOLDINGS: Total—1,800 Ohio —429 Strengths: Flora of southwestern Ohio North America—6 and Clifton Gorge, Greene County, Ohio; Eastern Hemisphere — 3 uncurated collection of Patagonian species. Total —438 NONVASCULAR HOLDINGS: Strengths: Aquatic species, woody spe- Algae: Limited collection of ma- cies, especially Acer and Quercus. rine algae from the North Carolina coast. NONVASCULAR HOLDINGS: None reported. Mounted on sheets. ACCESS: By appointment. Fungi: Limited collection of local EQUIPMENT: Dissecting scopes. higher fungi. Maintained in boxes. MAIL LOANS: No formal procedures. Lichens: Approximately 300 speci- EXCHANGES: No formal policy. mens. Collections mostly of a local nature; NOTE: Specimens mounted on sheets some southeastern Alaska specimens also covered with clear self-adhere present. An unknown number of un- plastic. mounted specimens from Patagonia are Teaching and reference collection. also available. Maintained in packets. Bryophytes: Approximately 1,200 THE WILDERNESS CENTER, INC. specimens. Largely representative of south- Box 202 western Ohio; some southeastern Alaska Wilmot, OH 44689 and eastern U.S. collections also main- STAFF: Robert W. Hawes, Naturalist tained. An unknown number of collections (216) 359-5235 from Patagonia are available. Maintained VASCULAR HOLDINGS: in packets. Ohio — 800 ACCESS: By appointment. Strengths: Flora of The Wilderness EQUIPMENT: Microscopes, manuals, and Center properties and the Stark-Case space. prairie, Holmes and Stark Counties, Ohio. MAIL LOANS: Yes. NONVASCULAR HOLDINGS: None reported. EXCHANGES: Yes, of bryophytes only. INCIDENTAL HOLDINGS: Small teach- Teaching and reference collection; research ing collections of seeds of prairie plants, on vascular and nonvascular Floras of leaves of woody species, and herbaceous southwest Ohio. inflorescences. WITTENBERG UNIVERSITY ACCESS: Tuesday-Saturday, 9 AM to 5 Jacobs Herbarium PM, Sunday, 1 to 5 PM, except holidays. Department of Biology EQUIPMENT: Dissecting scopes. Springfield, OH 45501 MAIL LOANS: No. CODE: JHWU EXCHANGES: No. STAFF: Ronald A. deLanglade, Professor of Teaching and reference collection. Biology WILMINGTON COLLEGE (513) 327-7719 Wilmington College Herbarium and VASCULAR HOLDINGS: Herbarium of Sture F. Anliot Ohio—3,000 Department of Biology North America— 1,000 Wilmington, OH 45177 Western Hemisphere — 8,700 OhioJ. Sci. SURVEY OF OHIO HERBARIUM RESOURCES 187

Total— 12,700 Strengths: Unknown Strengths: Flora of Iceland. NONVASCULAR HOLDINGS: NONVASCULAR HOLDINGS: Approximately Algae: Approximately 100 speci- 500 sheets, chiefly European algae, in mens of living cultures (liquid medium Stuckenberg collection (see below). and agar slants) representing U.S., west- INCIDENTAL HOLDINGS: J. H. W. Stucken- ern Canada and Great Britain. Complete berg collection from the late 19th century; collection of known genera, species and this is a group of portfolios containing strains of toxic freshwater blue-green algae about 2,000 sheets of European species. (cyanobacteria). ACCESS: By appointment. Fungi: Approximately2,000speci- EQUIPMENT: Workspace and microscopes. mens of myxomycetes, including collec- MAIL LOANS: Yes. tion of Harold W. Keller. Adams, Greene EXCHANGES: May be interested. and Scioto Counties, Ohio are especially Teaching and reference collection. represented in the collections. Lichens: None reported. COLLEGE OF WOOSTER Bryophytes: None reported. Department of Biology ACCESS: By appointment. Wooster, OH 44691 EQUIPMENT: Dissecting microscopes. STAFF: Gordon G. Whitney, Assistant MAIL LOANS: Yes. Professor of Biology Teaching and reference collection; research (216) 264-1234, ext. 379 into ecology of local plant communities. VASCULAR HOLDINGS: Ohio —2,200 YOUNGSTOWN STATE North America—2,800 UNIVERSITY Total—5,000 Department of Biological Sciences Strengths: Personal collections from 410 Wick Ave. the late 19th century of Gertrude Holden, Youngstown, OH 44555 George D. Hulst, R. H. Ingraham (north- CODE: YUO east Ohio) and A. D. Selby (Franklin STAFF: Carl F. Chuey, Associate Professor County); flora of Wayne County, Ohio. of Biological Sciences NONVASCULAR HOLDINGS: None reported. (216) 742-3601 ACCESS: Monday-Friday, 10 AM to 5 PM, Nicholas Sturm, Assistant Pro- September—May. fessor of Biological Sciences MAIL LOANS: No. VASCULAR HOLDINGS: EXCHANGES: No. North America—20,300 + Teaching and reference collection. Western Hemisphere—226 Eastern Hemisphere— 1,221 WRIGHT STATE UNIVERSITY Total —21,747 + Department of Biological Sciences Strengths: Worldwide ferns and fern Dayton, OH 45435 allies with emphasis on North America STAFF: James Runkel, Curator of Vascular (one-fourth of total vascular holdings); Plants, Assistant Professor of Bio- flora of Ohio, Pennsylvania, flora of logical Sciences Mahoning County, Ohio; 19th century (513)873-3199 collection from Siam by G. Culbertson; Wayne C. Carmichael, Curator of collections of John Laitsch, Almon Rood Nonvascular Plants, Associate and E. W. Vickers. Professor of Biological Sciences NONVASCULAR HOLDINGS: (513)873-3173 Algae: 65 specimens. Collections VASCULAR HOLDINGS: mostly marine, from New England, Ore- Ohio—100 gon and Washington. Mounted on sheets. North America—300 Fungi: 758 specimens. 625 basid- Total — 400 iomycetes, 133 ascomycetes and others. A. W. CUSICK AND J. A. SNIDER Vol. 84

Approximately 500 specimens are from ACKNOWLEDGMENTS. The authors are indebted Ohio, balance from Pennsylvania and West to the respondents to our survey who committed time and effort to make this publication as complete Virginia. Maintained in zip-lock bags, jars. as possible. Special thanks go to Dr. W. Hardy Lichens: 19 specimens. An addi- Eshbaugh, Miami University, for facilitating publi- tional 3000+ specimens from herbarium cation. The authors thank the Ohio Department of N. Sturm are yet to be accessioned. of Natural Resources, Division of Natural Areas and Preserves, and the Department of Biological Mahoning and Trumbull Counties, Ohio, Sciences, University of Cincinnati, for their gener- are represented, as well as Pennsylvania ous support. Special thanks are due Dr. Tod F. and West Virginia. Maintained in packets. Stuessy for his encouragement and assistance. Bryophytes: 477 specimens. An additional 400 specimens are yet to be accessioned. Ohio, Pennsylvania and West Virginia are represented. LITERATURE CITED INCIDENTAL HOLDINGS: 116 plant fossils. Beaman, J. H. 1965 The present status and ACCESS: Monday-Friday, 8 AM to 3 PM, operational aspects of university herbaria. Taxon September—May, or by appointment. 14: 127-133- MAIL LOANS: Yes; loans of nonvascular Cusick, A. W. and J. A. Snider 1982 Survey of the herbarium resources of Ohio. Organization materials restricted to bryophytes. of Herbaria in Ohio, Columbus. 43 p. EQUIPMENT: Microscopes. Holmgren, P. K., W. Keuken and E. K. Schofield EXCHANGES: Yes; also interested in dona- 1981 Index herbariorum. Part I. The herbaria tions from individuals or institutions. of the world. Regnum Vegetabile 106: 1-452. RESEARCH INTERESTS: Distributional stud- Ohio Natural Heritage Program Technical Report 1977 Columbus, Ohio Dept. of Natural Re- ies of ferns and fern allies; flora of West sources and The Nature Conservancy. 195 p. Virginia. Peterjohn, G. W. and P. B. Walker 1978 Aca- NOTE: Collections of the Ohio Mushroom demic biologists of Ohio, 1978-1979- Ohio Society are maintained here. Biol. Surv. Inform. Circ. No. 9. 20 p.