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A CATALOG OF THE COLEÓPTERA OF AMERICA NORTH OF MEXICO

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./S^ UNITED STATES AGRICULTURE PREPARED BY (fyj) DEPARTMENT OF HANDBOOK AGRICULTURAL AGRICULTURE NUMBER 529-105 RESEARCH SERVICE FAMILIES OF COLEóPTERA IN AMERICA NORTH OF MEXICO

Fascicle ' Family Year issued Fascicle ' Family Year issued Fascicle ' Family Year issued

1 Cupedidae 1979 45 Chelonariidae 98 Endomychidae __ 2 Micromalthidae 46 Callirhipidae 100 Lathridiidae 3 Carabidae 47 Heteroceridae 1978 102 Biphyllidae 4 Rhysodidae 48 Limnichidae 103 Byturidae 5 Amphizoidae 49 Dryopidae 104 Mycetophagidae 6 Haliplidae 50 Elmidae 105 Ciidae 1982 8 Noteridae 51 Buprestidae 107 Prostomidae 9 Dytiscidae 52 Cebrionidae 10 Gyrinidae 53 Elateridae 109 Colydiidae 13 Sphaeriidae 54 Throscidae 110 Mononunatidae 14 Hydroscaphidae 55 Cerophytidae 111 Cephaloidae 15 56 Perothopidae 112 Zopheridae 16 Hydrophilidae 57 Eucnemidae 115 Tenebrionidae _ _ 17 Georyssidae 58 —Telegeusidae 116 Alleculidae 18 Sphaeritidae 61 Phengodidae 117 -Lasriidae 20 Histeridae 62 Lampyridae 118 Salpingidae 21 Ptiliidae 63 Cantharidae 119 Mycteridae 22 Limulodidae 64 Lycidae 120 Pyrochroidae _— 23-__-Dasycendae 65 121 Othniidae 24 Micropeplidae 66 Nosodendridae 122 Inopeplidae 25 -Leptinidae 67 Dermestidae 123 Oedemeridae ___ 26 Leiodidae 69 Ptinidae 124 Melandryidae __ 27 Scydmaenidae 70 Anobiidae 125 Mordellidae 28 Silphidae 71 Bostrichidae 126 Rhipiphoridae __ 29 Scaphidiidae 72 Lyctidae 127_ _- Meloidae 30 Staphylinidae 74 Trogositidae 128 Anthicidae 31 Pselaphidae 76 129 Pedilidae 32 Lucanidae 78 Melyridae 130 Euglenidae 33 Passalidae -.-^ 79 131 Cerambycidae __ 34 Scarabaeidae 81 Sphindidae 132 Bruchidae 35 Eucinetidae 82 Nitidulidae .,_. 133 Chrysomelidae — 36 Helodidae 83 Rhizophagidae 134 Nemonychidae _- 37 86 Cucujidae 135 Anthribidae 38 Dascillidae 90 Cryptophagidae 138 Allocorynidae 39 Rhipiceridae 92 Languriidae 140 Brentidae —__> 40 Byrrhidae 93 Erotylidae 141 Platypodidae _— -.- 1979 41 Psephenidae 94 Phalacridae 142 Scolytidae > 42 Brachypsectridae 95 Cerylonidae 143 Curculionidae -_ 43 Artematopidae 96 Corylophidae 144 Stylopidae 44 97 Coccinellidae 145 Fossil Coleóptera

' Missing numbers are those assigned in the computer program to families not found in the United States and Canada.

For salo by the Superintendent of Documents, U.S. Government Printing Office, Washington, D.C. 20402 /A CATALOG OF THE COLEÓPTERA OF >A,MERICA NORTH OF MEXICO FAMILY: CIIDAE

BY J. F.ÍLAWRENCE DEPARTMENT OF ENTOMOLOGY, CSIRO CANBERRA, AUSTRALIA

UNITED STATES AGRICULTURE PREPARED BY DEPARTMENT OF HANDBOOK AGRICULTURAL V AGRICULTURE NUMBER 529-105 RESEARCH SERVICE

May 1982

FOREWORD

Many of are important pests of agricultural crops, stored food products, forests, wood products and structures, and fabrics. Many other species, in contrast, are beneficial in the biological sup- pression of pest and weeds, as well as in the decomposition of plant detritus, carcasses, and dung. Part of our national responsibility to American agriculture is to provide correct identification of species of American beetles so that appropriate controls can be applied. Most information about animal species, whether agricultural, biological, or experimental, is filed under the species' scientific names. These names are therefore the keys to retrieval of such information. Because some species have been known by several names, a complete listing of these names for each species is neces- sary. For the user of scientific names, an up-to-date taxonomic catalog providing currently accepted names and pertinent bibliographic and distributional data is an indispensable tool. Although taxonomic literature is constantly changing to reflect current work, the traditional published taxonomic catalog remains static with updating left to the individual user until it is revised. Production of catalogs in the past has been laborious with long printing delays resulting in data that are obsolete before being published. However, the computer now provides the capability of storing, updating, and retrieving taxonomic data; rapid publica- tion through computer-driven typesetting machinery; and a greater degree of currentness and flexibility. All 124 fascicles in this catalog of the beetles of America north of Mexico are produced by an original group of computer programs, designed and written during a pilot project by personnel of the Systematic Entomology Laboratory, Agricultural Research Service, and the Communications and Data Services Divi- sion, Science and Education Management Staff. The published information is stored on computer tape, is updated periodically to reflect taxonomic progress in the family, and is available in a data base for computer searching.

T. B. Kinney, Jr. ^ Administrator Agricultural Research Service

PREFACE The Coleóptera, or beetles, are represented in the world by about 220,000 described species, of which about 24,000 occur in ithe United States and Canada. A comprehensive taxonomic catalog of beetles for this area has not been available except the series of world-based "Coleopterorum Catalogus" volumes (1909-present, Junk, Berlin). The Leng "Catalogue of the Coleóptera of America North of Mexico" (J. D. Sherman, Jr., Mt. Vemon, NY), which was published in 1920 with supplements to the end of 1947, is a checklist. However, it has served professional and amateur alike for nearly 60 years as the principal source of scientific names of beetles. Since 1947, many new taxa have been described and many changes in status and nomenclature have appeared in numerous scattered publications, but little effort has been made to summarize these changes. This catalog will supplant the Leng catalog and supply additional essential information. It is produced by an original suite of storage, retrieval, and printing programs written especially for automated taxonomic catalogs. The catalog for each family is published as a separate fascicle with its introductory text, bibliography, and index. Each family is numbered as listed, but the order of issuance of fascicles is not necessarily in numerical sequence. The publishing of separate fascicles makes data available shortly after they are assembled. Computer tapes for each fascicle are maintained for updating and necessary reprinting. The information on each family is the responsibility of the respective author or authors. The editors modify it only to correct obvious errors and to make it conform to the requirements of the computer programs. No original proposal for a new name, taxon, status, or classification is given, such data having been previously published, but new host and distributional data are often listed. The rules of "The International Code of Zoological Nomenclature" are followed. The geographic scope of this catalog includes the continental United States, Canada, Alaska, Greenland, and the associated continental islands. Names of taxa found only in other regions are excluded. If the range of a species extends outside these geographic limits, this fact is indicated. Inside the back cover is a map of the 12 faunal regions based on historical and faunal criteria to simplify distribution recordings. Two-letter Postal Service style abbreviations are used for States and Provinces, and faunal regions are indicated in each distribution record by a diagonal line between groups of abbreviations. It is not the purpose of this catalog to present a complete scheme of higher classification within the order. The familial makeup is somewhat intermediate between that of R. H. Arnett in "The Beetles of the United States" (1960-62, Catholic University Press, Washington, DC) and that of R. A. Crowson in "The Natural Classification of the Families of Coleóptera" (1967, Biddies Ltd., Guildford, England). Modifications of these two systems are largely those advocated by J. F. Lawrence based in part on suggestions by taxonomic specialists for certain families. Generic groups and higher categories within the family are arranged phylogenetically as indicated by the author of the particular fascicle, and species group names with their respective synonyms are arranged alphabetically. Names referable to incertae sedis and nomen dubium are listed separately at the end of the nearest applicable taxon with notations as to their status. Each available name is followed by its author, date proposed, and page number referring to the complete bibliographic citation containing the original description. Following each generic name are

VII the type-species and method of its designation, necessary explanatory notes, and pertinent references on immature stages, , redescription, ecology, and keys. After the specific name entry are the original (if different from the present placement), type-locality, geographical distribution by State, Province, and broad extralimital units, explanatory notes, pertinent references to immature stages, taxonomy, redescription, and ecology, depository of type-specimen and its sex, and hosts. In addition to the list under the map of faunal regions (inside back cover), the following abbrevia- tions are used in this catalog:

ABBREVIATIONS, GENERAL

Amer. Bor.—America Borealis Mus.—Museum Amer. Sept.—America Septentrionalis N. Amer.—North America Autom.—Automatic Orig. des.—Original designation C. Amer.—Central America Preocc.—Preoccupied Co.—County S. Amer.—South America Cosmop.—Cosmopolitan Sp.—Species Design.—Designated Subseq. monot.—Subsequent monotypy F.—Female Subsp.—Subspecies Holarc.—Holarctic Taut,.—Tautonymy Isl.—Island Univ.—University M.—Male USA—United States of America Mex.—Mexico Var.—Variety Monot.—Monotypy W. Ind.—West Indies

MUSEUMS IN THE CONTINENTAL UNITED STATES AND CANADA ^

AMNH—American Museum of Natural History, FSCA—Florida State Collection, Gainesville New York HAHC—H. & A. Howden Collection, Ottawa, ANSP—Academy of Natural Sciences, Phila- Canada delphia, PA ICCM—Carnegie Museum, Pittsburgh, PA BYUC—Brigham Young University, Provo, UT INHÍ5—Illinois Natural History Survey, Urbana CASC—Cahfornia Academy of Sciences, San JGEC—J. G. Edwards Collection, San Jose, CA Francisco KMFC—K. M. Fender Collection, McMinnville, CISC—University of California, Berkeley OR CNCI—Canadian National Collections, Ottawa KSUC—Kansas State University, Manhattan CUIC—Cornell University, Ithaca, NY LACM—Los Angeles County Museum, CA CWOB—C. W. O'Brien Collection, Tallahassee, LSUC—Louisiana State University, Baton Rouge FL MCZC—Museum of Comparative Zoology, Har- DHKC—D. H. Kistner Collection, Chico State vard University, Cambridge, MA College, CA MSUC—Michigan State University, East Lansing ELSC—E. L. Sleeper Collection, Long Beach, NCSM—North Carolina State University, Raleigh CA NYSM—New York State Museum, Albany FMNH—^Field Museum of Natural History, Chi- OSEC—Oklahoma State University, Stillwater cago, IL OSUC—Ohio State University, Columbus OSUO—Oregon State University, Corvallis

^ Abbreviations for U.S. and Canadian museums abridged from Arnett, R. H., Jr., and Samuelson, G. A., 1969, "Di- rectory of Coleóptera Collections of North America (Canada Through Panama)," Cushing-Malloy, Ann Arbor, MI, 123 pp. vni PMNH—Peabody Museum, Yale University, New SMSH—Stovall Collection, University of Okla- Haven, CT homa, Norman PSUC—Pennsylvania State Museum, University TAMU—Texas A. & M. University, College Park Station PURC—Purdue University, West Lafayette, IN UCDC—University of California, Davis RUIC—Rutgers University, New Brunswick, NJ UMMZ—University of Michigan, Ann Arbor SEMC—Snow Museum, University of Kansas, UMRM—University of Missouri, Columbia Lawrence USNM—^U.S. National Museum of Natural His- SJSC—San Jose State College, CA tory, Washington, DC SLWC—S. L. Wood Collection, Provo, UT WSUC—Washington State University, Pullman

MUSEUMS IN FOREIGN COUNTRIES

BMNH—British Museum (Natural History), NHRS—Naturhistoriske Riksmuseet, Stockholm NMPC—Narodni Museum, Prague, Czechoslo- London BPBM—Bernice P. Bishop Museum, Honolulu vakia GUHC—Glasgow University, Hunterian College, SCUT—Spinola College, University of Turin, Scotland Italy HMOX—Hope Museum, Oxford, England SMTD—Staatliches Museum für Tierkunde, HNHM—Hungarian Natural History Museum, Dresden, East Germany UNAM—Universidad Nacional Autónoma, Mexi- Budapest IPZE—Institut Pflanzenschutzforschung Zweig- co City stelle, Eberswalde, East Germany UZMC University Zoological Museum, Copen- IRSB—Institut Royal Sciences Belgique, Brus- hagen, Denmark sels UZMH—University Zoological Museum, Hel- MFNB—Museum für Naturkunde (Humboldt), sinki, Finland Berlin ZMAS—Zoological Museum, Academy of Sci- MGFT—Museum G. Frey, Tutzing, Munich, ences, Leningrad West Germany ZMPA—Zoological Museum, PoUsh Academy of MHNL—Museum d'Histoire Naturelle, Lyon, Sciences, Warsaw France ZMUL—Zoological Museum, University of Lund, MNHP—Museum National d'Histoire Naturelle, Sweden Paris ZMUM—Zoological Museum, University of MNSL—Museum of Natural Sciences, Leipzig, Moscow East Germany ZSBS—Zoologische Sammlung Bayerischen MZBS—Museum Zoología, Barcelona, Spain Staates, Munich, West Germany ACKNOWLEDGMENTS We are indebted to many individuals who contributed to the planning and development of this ca- talog. We are especially grateful to the following specialists who helped to make it as complete and accurate as possible: Richard H. Foote, Systematic Entomology Laboratory (SEL.), Agricultural Research Service (ARS), for his suggestions, guidance, and encouragement; C. W. Sabrosky, SEI,, for valuable counsel on nomenclatural problems; J. P. I.awrence, Division of Entomology, Com- monwealth Scientific and Industrial Research Organization, Canberra, Australia, for his recommenda- tions on higher categories; and more than 50 coleopterists in Canada, the United States, and Mexico for voluntarily contributing information about their specialty groups. We thank the following members of the Communications and Data Services Division, SEMS: San- dra Strauss and Marianne Kingston for designing and writing the computer programs, and Margaret Seldin for developing the editing system.

J. M. Kingsolver, editor in chief Systematic Entomology Laboratory, Agricultural Research Service Washington, D.C.

Editorial Board

J. M. Campbell, Biosystematics Research Institute, Agriculture Canada, Ottawa, ON T. L. Erwin, Department of Entomology, Smithsonian Institution, Washington, DC H. F. Howden, Department of Biology, Carleton university, Ottawa, ON P. J. Spangler, Department of Entomology, Smithsonian Institution, Washington, DC T. J. Spilman, Systematic Entomology Laboratory, ARS, Washington, DC R. E. White, Systematic Entomology Laboratory, ARS, Washington, DC

CONTENTS

^ ., , Page Family Ciidae I Subfamily Sphindociinae j Subfamily Ciinae j Tribe 2 Tribe Orophiini j 2 Bibliography J3 Index 2 y Family CIIDAE (Cisidae, Cissidae, Cioidae) By J. F. Lawrence The Ciidae is a relatively small family with about 40 genera and 550 described species oc- curring on all the major continents and most oceanic islands. The North American fauna con- sists of 85 species in 13 genera. The classification and generic arrangement used here are from Lawrence (1971, 1974b). According to Crowson (1960, 1966, 1967) and Lawrence (1971, 1974b), the ciids are primitive members of the Heteromera, with affinities to the Tetratomidae, Myce- tophagidae, Prostomidae, and Pterogeniidae, but the relationships among these heteromeran families are still not clear. Although the two major subfamilies, Sphindociinae and Ciinae, appear to be well founded, the tribal classification within the latter group needs further study, and new genera remain to be described, especially from the Neotropical and Oriental regions. Much more work remains at the species level; this author has seen over 400 undescribed forms in collections. The North American fauna has been revised (Lawrence, 1971), but a few more species remain to be described from the Southeast. Although numerous larvae have been col- lected, few have been described to date. Work is continuing by this author on the phylogeny of the Ciidae and related families and on the biosystematics of various ciid genera. Ciids are usually collected in the fruiting bodies of higher Basidiomycetes, especially the (s. lat.), but some may be collected under bark or beaten from rotten branches or vines. Host preferences in the group have been discussed by Lawrence (1973). This manuscript was received November 1975. Subfamily SPHINDOCIINAE Genus SPHINDOCIS Fall Sphindocis Fall, 1917: 170. Type-species: Sphindocis denticollis Fall (monot.). TAXONOMY: Lawrence, 1974b: 1. denticollis Fall, 1917: 171. CA: Alameda Co.; CA. TYPE DEPOSITORY: MCZC. IMMATURE STAGES: Lawrence, 1974b: 2-7. TAXONOMY: Lawrence, 1971: 426, table 1 and 1974b: 1-14. REDESCRIPTION: Lawrence, 1974b. ECOLOGY: Lawrence, 1974a: 24 and 1974b: 1-2. HOST: Antrodia sepium, Tyromyces cinerascens. Subfamily CIINAE Tribe CIINI Genus Latreille Cis Latreille, 1796: 50. Type-species: Dermestes boleti Scopoli (subseq. monot.: Latreille, 1802: 205). Eridaulus Thomson, 1863: 191. Type-species: Anobium nitidum Fabricius (design, by Lawrence, 1971: 437). Macrocis Reitter, 1878: 34. Type-species: Macrocis taurus Reitter (design, by Lawrence, 1971: 437). Xestocis Casey, 1898: 85. Type-species: Xestocis levettei Casey (design, by Lawrence, 1965: 282). TAXONOMY: Lawrence, 1965 and 1971. REDESCRIPTION: Crowson, 1938: pi. 8, fig. 11 (metendosternite). KEYS: Lawrence, 1971. acritus Lawrence, 1971: 443, figs. 52, 63, 73, 80. AZ: Cochise Co., Rustler Park, 8 mi W. Portal; CA/ AZ NM. TYPE DEPOSITORY: CASC. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Hirschioporus abietinus. ¿ COLEÓPTERA CATALOG americanus Mannerheim, 1852: 360. AK: Sitka; AK/ BC WA OR/ AB MB SD/ WI ON PQ/ NB NS/ CA NV/ UT CO/ NY PA/ ME NH VT MA/ TN NC. TYPE DEPOSITORY: UZMH. insolens Casey, 1898: 86 (Xestocis). PA. TYPE DEPOSITORY: USNM. frosti Dury, 1917: 9. ME: Orono. TYPE DEPOSITORY: CMNH (lost). serricollis Dury, 1917: 9. OR: Linn Co. TYPE DEPOSITORY: CMNH. minor Hatch, 1962: 232 (Xestocis). OR: Wheatland. TYPE DEPOSITORY: USNM. oweni Hatch, 1962: 232 (Xestocis). WA: Seattle. TYPE DEPOSITORY: USNM. strigulosus Hatch, 1962: 231 (Xestocis), WA: Seattle. TYPE DEPOSITORY: USNM. nitidus Hatch, 1962: 232 fZes^ocisXpreoccupied Fabricius, 1792). OR: Wheatland. TYPE DEPOSITORY: USNM. hatchi Lawrence, 1971: 445 (replacement name for nitidus Hatch not Fabricius). OR: Wheatland. forsten Abdullah, 1973: 214 (unneeded replacement name for nitidus Hatch not Fabricius). OR: Wheatland. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Bjerkandera adusta, Piptoporus betulinus, Polyporus squamosus, Fomitopsis pinícola, Ganoderma spp., various other Basidiomycetes. angustus Hatch, 1962: 230. BC: Stanley; BC WA/ CA. TYPE DEPOSITORY: CASC. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Fomitopsis pinicola, Laricifomes officinalis, Heterobasidion annosum. biarmatus Mannerheim, 1852: 360. AK: Sitka; AR/ BC WA OR/ CA. TYPE DEPOSITORY: UZMH. bicarinatus (error) LeConte, 1867: 58. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Fomitopsis pinicola. castlei (Dury), 1917: 17 (Xestocis). OH: Cincinnati; ON/ lA IL OH KY/ PA MD DC/ LA MS TN SC NC FL/ Mex., C. Amer. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973. HOST: Bjerkandera adusta, Polyporus mutabilis, Nigroporus vinosus, Trametes scabrosa, various other Basidiomycetes. cayensis Lawrence, 1971: 448. FL: Monroe Co., Big Pine Key; FL/ W. Ind. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Pogonomyces hydnoides, Inonotus porrectus, I. corrosus, Phellinus robiniae, Lopharia papyrina. congestus Casey, 1898: 82. LA; IL/ MD VA/ TX/ LA MS AL TN GA SC NC FL. TYPE DEPOSITORY: USNM. lodingi Dury, 1917: 6. AL: Mobile. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973. HOST: Coriolus hirsutus, C. pinisitus. cornelli Lawrence, 1971: 450, figs. 42, 50, 67, 76, 79. NC: Carteret Co., Atlantic Beach; SC NC FL. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Hirschioporus sector. CIIDAE à cornutus Blatchley, 1910: 898, fig. 353. IN: Posey Co., Grand Chain; IL IN OH/ NY PA NJ MD DC VA/ AR MS AL NC FL. TYPE DEPOSITORY: PURC. ECOLOGY: Lawrence, 1973. HOST: Coriolus versicolor, C. subectypus. creberrimus Mellie, 1848: 357, pi. 12, fig. 5. LA: New Orleans; MN WI MI ON/ CA/ NE KS lA MO IL IN OH KY/ NY PA NJ MD DC WV VA/ VT MA/ AZ NM TX/ AR LA MS AL TN GA SC NC FL/ Mex., C. Amer., S. Amer., W. Ind. TYPE DEPOSITORY: MNHP. puberulus Mellie, 1848: 358. W. Ind.: Virgin Isis., St. Thomas. TYPE DEPOSITORY: MNHP. nubillus Gorham, 1898: 331. W. Ind.: St. Vincent, south end. TYPE DEPOSITORY: BMNH. nubilus (error) Dalla Torre, 1911: 15. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Bjerkandera adusta, Laricifornes officinalis, Ganoderma spp., Fomes sclerodermeus, Pogonomyces hydnoides, Trametes scabrosa, various other Basidiomycetes. crinitus Lawrence, 1971: 453, fig. 29. FL: Monroe Co., Lignum Vitae Key; LA AL GA NC FL/ W. Ind. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Lopharia papyrina, Coriolopsis fulvocinerea, Gloeophyllum striatum, Pogonomyces hydnoides, Ganoderma zonatum. discolor Lawrence, 1971: 455, figs. 44, 56, 62, 68, 81. AZ: Cochise Co., Rustler Park, 8 mi W. Portal; AZ. TYPE DEPOSITORY: CASC. ECOLOGY: Lawrence, 1973. HOST: Fomitopsis cajanderi, Antrodia sepium. dunedinensis Leng, 1918: 207 (replacement name for pusillus Dury). FL: Dunedin; FL/ W. Ind. pusillus Dury, 1917: 10 (preoccupied Gorham, 1898). FL: Dunedin. TYPE DEPOSITORY: CMNH. duplex Casey, 1898: 82. CA; CA/ AZ NM/ Mex. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Dichomitus squalens, Fomitopsis pinicola, Laricifomes officinalis, Ganoderma oregonense. ephippiatus Mannerheim, 1853: 234. AK: Sitka; AK/ BC WA OR/ PQ/ CA NV/ WY CO/ NH VT. TYPE DEPOSITORY: UZMH?. ephippium (error) Dalla Torre, 1911: 10. moznettei Dury, 1917: 16 (Xestocis). OR: Corvallis. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Fomitopsis pinicola, Ganoderma applanatum, G. brownii, Heterobasidion annosum, Piptoporus betulinus, Ischnoderma resinosum, Laetiporus sulphureus, Poria uersipora. festivulus Lawrence, 1971: 459, figs. 47, 65, 72, 84. DC; MD DC VA. TYPE DEPOSITORY: USNM. floridae Dury, 1917: 9. FL: Key West; GA FL/ W. Ind. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973. HOST: Phellinus gilvus, Phellinus sp., Lopharia papyrina. fuscipes Mellie, 1848: 271, pi. 10, fig. 23. MA: Boston; NT/ BC WA OR ID/ AB SK MB MT ND/ MN MI ON PQ/ NS/ CA/ UT/ NE KS lA MO IL IN OH KY/ NY PA NJ MD DC WV VA/ ME NH VT MA RI CT/ TX/ AR LA MS AL TN GA SC NC FL/ W. Ind., Madeira. C. fuscipes is also known from Maui, Hawaii and from New Zealand. TYPE DEPOSITORY: MNHP. atripennis Mellie, 1848: 258, pi. 10, fig. 15. MA: Boston. TYPE DEPOSITORY: MNHP. 4 COLEÓPTERA CATALOG

chevrolatii Mellie, 1848: 249. LA: New Orleans. TYPE DEPOSITORY: MNHP. dubius Mellie, 1848: 273. LA: New Orleans. TYPE DEPOSITORY: MNHP. carolinae Casey, 1898: 78. NC: Asheville. TYPE DEPOSITORY: USNM. impressus Casey, 1898: 79. BC: Vancouver Isl. TYPE DEPOSITORY: USNM. pallens Casey, 1898: 78. MT: Missoula. TYPE DEPOSITORY: USNM. IMMATURE STAGES: Boeving and Craighead, 1931: pi. 92. TAXONOMY: Lawrence, 1967a. ECOLOGY: Lawrence, 1967a, 1973 and 1974a. HOST: Coriolus versicolor, C. hirsutas, C. conchifer, C. pubescens, Lenzites betulina, various other Polyporaceae. hirsutus Casey, 1898: 83. FL: Lake Worth; FL/ W. Ind. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973. HOST: Ganoderma zonatum, Ganoderma sp., Pogonomyces hydnoides, Trámeles scabrosa, Fomes sclerodermeus, Hirschioporus sector, various other Basidiomycetes. horridulus Casey, 1898: 81. PA: Westmoreland Co.; NT/ BC/ MB/ MN WI MI ON/ UT CO/ NY PA/ ME NH VT MA/ AZ NM/ NC. TYPE DEPOSITORY: USNM. mormonicus Casey, 1898: 81. UT: southwestern. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973. HOST: Hirschioporus pargamenus, H. abietinus. huachucae Dury, 1917: 8. AZ: Huachuca Mts., Miller Caxiyon; AZ TX/ Mex. TYPE DEPOSITORY: CMNH. hystriculus Casey, 1898: 82. CA: Lake Tahoe; BC WA OR/ CA. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Hirschioporus abietinus, Tyromyces cinerascens, Poria versipora. krausi Dalla Torre, 1911: 13 (replacement name for bimaculatus Kraus not Germain, not Sharp). TX: Victoria; TX/ GA FL. bimaculatus Kraus, 1908: 76 (preoccupied Germain, 1855; Sharp, 1885). TX: Victoria. TYPE DEPOSITORY: USNM. bicolor Dury, 1917: 7 (preoccupied Sharp, 1879). GA: Tybee Isl. TYPE DEPOSITORY: CMNH. duryi Leng, 1918: 207 (replacement name for bicolor Dury not Sharp). laminatus Mellie, 1848: 318, pi. 11, fig. 16. France: Montrieux, near Toulon; NY/ Europe. TYPE DEPOSITORY: MNHP. ECOLOGY: Lawrence, 1973. HOST: Crytoporus volvatus. levettei (Casey), 1898: 85 (Xestocis). IN?; BC WA/ AB SK MB/ MN WI MI ON PQ/ NS NF/ CA/ CO/ KS lA MO IL IN KY OH/ NY PA NJ MD WV VA/ ME NH VT MA CT/ TX/ AL TN SC NC. TYPE DEPOSITORY: USNM. levellei (error) Dalla Torre, 1911: 20. IMMATURE STAGES: Peterson, 1957: C5, C54. ECOLOGY: Lawrence, 1973. HOST: Ganoderma applanatum, G. tsugae, G. lucidum, Fomes fo,nentarius, Fomitopsis pinícola, Oxyporus populinus, various other Basidiomycetes. maritimus (Hatch), 1962: 233 (Xestocis). OR: Ocean Park; BC WA OR/ MB/ CA. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Phaeolus schweinitzii, Phellinus pini. CIIDAE ö megastictus Lawrence, 1971: 465, figs. 39, 55, 64, 74, 78. CA: Plumas Co., Bucks Lake; CA. TYPE DEPOSITORY: CASC. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Heterobasidion annosum, Fomitopsis pinícola, Laetiporus sulphureus. miles (Casey), 1898: 85 (Xestocis). PA: Westmoreland Co., St. Vincent; MO IL IN OH/ NY PA MD DC VA/ AR LA MS AL GA SC NC FL. TYPE DEPOSITORY: USNM, ECOLOGY: Lawrence, 1973. HOST: Lenzites hetulina, Coriolus versicolor, C. subectypus, Fomitella supina, Stereum ostrea. niedhauki Lawrence, 1971: 467, fig. 5. FL: Monroe Co., Lignum Vitae Key; FL. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Phellinus robiniae. pistoria Casey, 1898: 79. RI: Boston Neck; NT/ AB SK MB/ MN WI ON PQ/ NY/ ME NH VT MA. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973. HOST: Coriolus versicolor, C. pubescens, C. hirsutus, Bjerkandera adusta. quadridentatus (Dury), 1917: 17 (Xestocis). MA: Framingham; lA IL/ NY DC VA/ VT MA/ TX/ MS NC FL. TYPE DEPOSITORY: CMNH. blatchleyi Dury, 1917: 7. FL: Dunedin. TYPE DEPOSITORY: PURC. robiniophilus Lawrence, 1971: 470, figs. 49, 61, 71, 83. OH: Hamilton Co., Preston; OH KY/ MD. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973. HOST: Trametes robiniophila. rotundulus Lawrence, 1971: 471, figs. 41, 51. SC: CoUeton Co., Walterboro; TX/ AR LA MS AL GA SC NC FL. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: , G. curtisii, Cyclomyces iodinus, Phellinus gilvus, Stereum ostrea. stereophilus Lawrence, 1971: 472, figs. 40, 53, 66, 77, 82. NC: Carteret Co., Atlantic Beach; PA NJ MD DC/ MA/ NC FL/ Mex. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Stereum ostrea, Stereum sp. striolatus Casey, 1898: 79. CO: Salida; NT/ BC OR/ MB ND SD/ MN MI ON PQ/ NS/ CA/ UT CO/ KS IL IN/ VA/ NH VT MA/ FL. TYPE DEPOSITORY: USNM. fraternus Casey, 1898: 80. UT: southwestern. TYPE DEPOSITORY: USNM. macilentus Casey, 1898: 80. CA: Lake Tahoe. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Hirschioporus abietinus, H. pargamenus, Cerrena unicolor, Coriolus hirsutus. subfuscus Gorham, 1886: 357. Mex.: Cordova, Veracruz; TX/ Mex. TYPE DEPOSITORY: BMNH. ECOLOGY: Lawrence, 1973. HOST: Coriolus hirsutus, Panellus stipticus, Lenzites elegans. subtilis Mellie, 1848: 353. N. Amer.; WI MI/ IL IN OH/ NY PA NJ MD DC VA/ NH MA CT/ TX/ AR LA MS AL GA SC NC FL. TYPE DEPOSITORY: MNHP. confusus Blatchley, 1910: 899. IN: Perry Co. TYPE DEPOSITORY: PURC. julichi Dury, 1917: 7. NY: New York City. TYPE DEPOSITORY: CMNH. o COLEÓPTERA CATAUXÎ wenzeli Dury, 1917: 8. PA: Delaware Co. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973. HOST: Hirschioporus pargamenus, H. sector, H. abietinus, Ganoderma applatum, Phellinus gilvus, Bjerkandera adusta, Stereum ostrea. tetracentrum Gorham, 1886: 357. Mex.: Northern Sonora; CA/ AZ/ Mex. TYPE DEPOSITORY: BMNH. arizonae Dury, 1917: 5. AZ: Santa Rita Mts., Madera Canyon. TYPE DEPOSITORY: OSUC?. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Coriolus versicolor. tridentatus Mannerheim, 1852: 360. AK: Sitka; AK/ BC WA OR/ CA. TYPE DEPOSITORY: UZMH. ednae Hatch, 1962: 232 (Xestocis). WA: Seattle. TYPE DEPOSITORY: USNM. reßexus Hatch, 1962: 232 (Xestocis). WA: Coupeville (Sunnyside). TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Fomitopsis pinícola, Ganoderma oregonense, Tyromyces cinerascens, Laetiporus sulphureus, various other Basidiomycetes. tristis Mellie, 1848: 343. LA: New Orleans; WI MI/ CO/ NE KS MO IL IN OH/ NY NJ DE MD WV VA/ MA/ TX OK/ LA MS AL GA SC NC FL/ Mex. TYPE DEPOSITORY: MNHP. setulosus Mellie, 1848: 257. N. Amer. TYPE DEPOSITORY: MUS. Nat. His., Geneva. falli Blatchley, 1910: 898. IN: Marion Co. TYPE DEPOSITORY: PURC. ECOLOGY: Lawrence, 1973. HOST: Coriolus versicolor, C. maximus, C. hirsutus, Lenzites betulina,L. elegans, Funalia hispida, various other Basidiomycetes. ursulinus Casey, 1898: 83. AL; IN/ AR LA MS AL SC NC FL. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973. HOST: Phellinus gilvus, Ganoderma tsugae, Hirschioporus versatilis, Bjerkandera adusta, Laetiporus sulphureus. versicolor Casey, 1898: 80. CA: Calaveras Co.; OR/ CA/ AZ NM TX/ Mex. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Coriolus versicolor, C hirsutus, Funalia hispida, Lenzites betulina, Pycnoporus cinnabarinus, Ganoderma brownii, Schizophyllum commune. vitulus Mannerheim, 1843: 299. CA; OR/ CA/ AZ. TYPE DEPOSITORY: UZMH?. illustris Casey, 1898: 81 (preoccupied Broun, 1880). CA: Humboldt Co. TYPE DEPOSITORY: USNM. caseyi Dalla Torre, 1911: 8 (replacement name for illustris Casey not Broun). ECOLOGY: Lawrence, 1973 and 1974a. HOST: Coriolus versicolor, Lenzites betulina, Bjerkandera adusta, Schizophyllum commune. Genus Mellie Ennearthron Mellie, 1847: 110. Type-species: Cis cornutus Gyllenhal (design, by Desmarest, 1860:261). Ennearthrum (error) Bach, 1852: 111. aurisquamosum Lawrence, 1971: 481, fig. 8. NC; OH KY/ NC. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Bjerkandera adusta. CIIDAE ' spenceri (Hatch), 1962: 233 (Plesiocis), BC: Vancouver; BC. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973. HOST: Cryptoporus volvatus. Genus Dury Dolichocis Dury, 1919: 158. Type-species: Dolichocis manitoba Dury (monot.). Ennearthron, of authors, in part, not Mellie. TAXONOMY: Lawrence, 1965 and 1971. KEYS: Lawrence, 1971. indistinctus Hatch, 1962: 234. BC: Stanley; BC/ PQ/ CA/ CO/ VT/ AZ. TYPE DEPOSITORY: CASC. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Fomitopsis pinícola, Antrodia serialis, Laricifomes officinalis. manitoba Dury, 1919: 158. MB: Aweme; NT/ BC OR ID/ AB MB SD/ WI ON PQ/ NB/ CA/ WY UT/ NY PA/ ME NH VT MA. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973 and 1974a. ^ , i . HOST- Fomitopsis pinícola, Píptoporus hetulínus, Heterobasidion annosum, Ganoderma applanatum, Laetiporus sulphureus, Laricifomes officinalis, various other Polyporaceae.

Genus Casey Orthocis Casey, 1898: 84. Type-species: Orthocis aterrima Casey (design, by Lawrence, 1965: 288) punctatus (MeUie). Mellieicis Lohse, 1964: 122 (as subgenus). Type-species: Cis alni Gyllenhal (orig. des.). TAXONOMY: Lawrence, 1965 and 1971; Lohse, 1964. KEYS: Lawrence, 1971. huesanus Kraus, 1908: 77, pi. 3, fig. 3. FL: Key West; FL. TYPE DEPOSITORY: USNM. longulus Dury, 1917: 13. PA: Linglestown; KS/ PA/ TX/ LA MS AL FL/ Mex. TYPE DEPOSITORY: CMNH. pulcher Kraus, 1908: 78, pi. 3, fig. 4. FL: Key West; FL. TYPE DEPOSITORY: USNM. punctatus (Mellie), 1848: 337 (Cis). N. Amer.; AK NT/ BC WA OR/ AB MB SD/ WI m PQ/ NF/ CA/ KS IN OH/ NY PA NJ MD DC WV VA/ ME NH MA/ NM TX/ AR LA MS AL TN GA SC NC FL. TYPE DEPOSITORY: MNHP. aterrimus Casey, 1898: 84. CA: Alameda Co. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Auricularia auricula. transversatus (Kraus), 1908: 79, pi. 3, fig. 7 (Ennearthron). FL: Crescent City; VA/ TX/ LA AL GAFL. TYPE DEPOSITORY: USNM. pallidus Kraus, 1908: 79, pi. 3, fig. 6 (Ennearthron). FL: Haw Creek. TYPE DEPOSITORY: USNM (lost). Genus Dury Strigocis Dury, 1917: 18. Type-species: Strigocis opacicollis Dury (monot.). TAXONOMY: Lawrence, 1971: 488. KEYS: Lawrence, 1971: 489. bilimeki (Reitter), 1878a: 33 (Cis). Mex.; AZ/ Mex. Arizona record needs confirmation. TYPE DEPOSITORY: MNHP. ECOLOGY: Lawrence, 1973. HOST: Lenzites betulina, Coriolus versicolor. 8 COLEÓPTERA CATALOG

opacicoUis Dury, 1917: 20. OH: Cincinnati; MI ON/ KS MO IL IN OH KY/ NY PA NJ MD DC VA/ NH VT MA CT/ AR MS AL TN GA SC NC FL/ Mex. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973. HOST: Coriolus versicolor, C. tenais, C subectypus, C. pinisitus, C. maximus, C. hirsutas, Lenzites betalina, Pycnoporus sangaineas, various other Polyporaceae. opalesçens (Casey), 1898: 86 (Xestocis). PA: Westmoreland Co., St. Vincent; MI/ OH KY/ NY PA MD DC WV VA/ MS NC/ Mex. TYPE DEPOSITORY: USNM. davisi Dury, 1917: 16 (Xestocis). NY: Staten Island. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973. HOST: Coriolas versicolor.

Genus HADRAULE Thomson Hadraule Thomson, 1863: 182. Type-species: Cis elongatulus Gyllenhal (monot.). Maphoca Casey, 1900: 165. Type-species: Maphoca blaisdelli Casey (monot ) Maphaea (error) Dalla Torre, 1911: 21. Diphyllocis, of authors, in part, not Reitter. Pityocis Peyerimhoff, 1918: 141. Type-species: Pityocis coarctatus Peyerimhoff (monot.) elongatula (Gyllenhal). Knablia Roubal, 1936: 53. Type-species: Cis elongatulus Gyllenhal (monot.). TAXONOMY: Lawrence, 1965 and 1971. KEYS: Lawrence, 1971. blaisdelli (Casey), 1900: 165 (Maphoca), CA: Calaveras Co., Mokelumne Hill; BC OR/ MI/ CA/ UT/ lA OH/ MA CT/ AZ NM TX/ FL/ Mex. Records from lA, MA, and CT are based on herbarium infestations. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Gloeophyllam saepiariam, Daedaleopsis confragosa, Fanalia hispida, Coriolas biformis, Dichomitas sqaalens, Bjerkandera adasta, various other Polyporaceae. elongatula (Gyllenhal), 1827: 627 (Cis). Sweden: Sparrsaetra Westrogothiae; NB/ Old World. TYPE DEPOSITORY: Zool. Univ. Mus., Uppsala. striata Sahlberg, 1901: 11 (Ennearthron). Finland: southern. TYPE DEPOSITORY: UZMH?. coarctata Peyerimhoff, 1918: 142 (Pityocis). Algeria: Ain-Haouas, near Djelfa. TYPE DEPOSITORY: MNHP. ECOLOGY: Lawrence, 1973. HOST: Piptoporas betalinas. explanata Lawrence, 1971: 493, fig. 21. ME: Washington Co., E. Machias; ME. TYPE DEPOSITORY: CASC.

Genus PLESIOCIS Casey Plesiocis Casey, 1898: 87. Type-species: Plesiocis cribrum Casey (monot.). TAXONOMY: Lawrence, 1971. cribrum Casey, 1898: 87. CA: Calaveras Co., Mokelumne Hill; BC WA OR ID/ AB MB MT/ WI MI PQ/ CA NV/ WY UT CO/ NY PA VA/ ME MA/ AZ NM. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Cryptoporas volvatas, Dichomitas sqaalens, Hapalopilas albolateas. Genus Mellie Ceracis Mellie, 1848: 375 (as subgenus). Type-species: Ennearthron sallei Mellie (design bv Lawrence, 1967b: 95). * ^ TAXONOMY: Lawrence, 1967b. KEYS: Lawrence, 1967b and 1971. CIIDAE y californicus (Casey), 1884: 36 (Ennearthron). CA; WA OR/ CA NV/ UT CO/ NE/ DC/ CT/ AZ NM/ Mex. TYPE DEPOSITORY: USNM. convergens Casey, 1898: 89 (Ennearthron). CA: Los Angeles. TYPE DEPOSITORY: USNM. discolor Casey, 1898: 89 (Ennearthron), CA: Sonoma Co. TYPE DEPOSITORY: USNM. grossulus Casey, 1898: 89 (Ennearthron). CA: southern. TYPE DEPOSITORY: USNM. coloradensis Dury, 1917: 22 (Ennearthron). CO: Grand Lake, Middle Park. TYPE DEPOSITORY: CMNH. oregonus Dury, 1917: 22 (Ennearthron). OR: Corvallis. TYPE DEPOSITORY: CMNH. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b, 1973 and 1974a. HOST: Ganoderma browniU G. applanatum, G. lobatum, Ganoderma spp., Coriolus versicolor, Bjerkandera adusta, Funalia hispida, various other Basidiomycetes. curtus (Mellie), 1848: 367, pi. 12, fig. 15 (Ennearthron). Cuba: Havana; PA/ TX/ FL/ W. Ind. TYPE DEPOSITORY: MNHP. obesus Mellie, 1848: 335, pi. 11, fig. 13 (Cis). N. Amer. TYPE DEPOSITORY: MUS. Nat. His., Geneva. compactus Dury, 1917: 21 (Ennearthron). FL: Key West. TYPE DEPOSITORY: CMNH. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Pogonomyces hydnoides, Fomes sclerodermeus. dixiensis (Tanner), 1934: 47 (). UT: Zion N. P.; CA/ UT/ AZ NM TX/ Mex. TYPE DEPOSITORY: BYUC. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b, 1973 and 1974a. HOST: Funalia hispida, Ganoderma sp. magister Lawrence, 1971: 498. FL: Monroe Co., Lignum Vitae Key; FL. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Phellinus rohiniae. minutissimus (Mellie), 1848: 334, pi. 11, fig. 12 (Cis). MA: Boston; MI/ IN OH KY/ PA NJ/ NH MA/ AL. TYPE DEPOSITORY: MNHP. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. HOST: Cerrena unicolor, Coriolus versicolor, Bjerkandera adusta. minutus Dury, 1917: 25. FL: Palm Beach; TX/ FL/ W. Ind. TYPE DEPOSITORY: CMNH. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Coriolus pinisitus, C. maximus, Pogonomyces hydnoides, Coriolopsis occidentalis, Pycnoporus sanguineus, various other Basidiomycetes. monocerus Lawrence, 1967b: 115 (replacement name for unicorne. Casey not Gorham). FL; LA FL/ W. Ind. unicornis Casey, 1898: 90 (EnnearthronXpreoccupied Gorham, 1898). FL. TYPE DEPOSITORY: USNM. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Pycnoporus sanguineus. 10 COLEÓPTERA CATALOG

multipunctatus (Mellie), 1848: 368, pl. 12, fig. 16 (Ennearthron). Cuba; AL FL/ W. Ind. TYPE DEPOSITORY: ?. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Ganoderma zonatum, G. applanatum, G. lucidum, Fomes sclerodermeus, Pogonomyces hydnoides, Fomitella supina. nigropunctatus Lawrence, 1967b: 118, fig. 16. LA: Iberville Par., 4 mi S. Grosse Tete; TX/ LA/ Mex., C. Amer. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Pogonomyces hydnoides, Fomes sclerodermeus, Ganoderma sp., Coriolus hirsutus. obrieni Lawrence, 1967b: 119, fig. 25. AZ: Santa Cruz Co., 2 mi S. W. Patagonia; AZ/ Mex. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Phellinus gilvus. pecki Lawrence, 1971: 499. FL: Jackson Co., Florida Caverns State Park; IL/ MD VA/ FL. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Melanoporia nigra. powelli Lawrence, 1967b: 120, fig. 23. AZ: Cochise Co., Rustler Park, 8 mi W. Portal; AZ/ Mex. TYPE DEPOSITORY: MCZC. ECOLOGY: Lawrence, 1973. HOST: Hirschioporus abietinus, H. pargamenus. puUuIus (Casey), 1898: 90 (Ennearthron). FL; LA AL TN GA SC NC FL/ W. Ind. TYPE DEPOSITORY: USNM. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Phellinus gilvus, P. licnoides, Inonotus porrectus, Inonotus spp., Melanoporia nigra, Cyclomyces iodinus, Ganoderma zonatum, various other Polyporaceae. punctulatus punctulatus Casey, 1898: 90. FL; FL. TYPE DEPOSITORY: USNM. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Pogonomyces hydnoides, Phellinus licnoides, P. gilvus, Ganoderma zonatum, Ganoderma sp. punctulatus rubriculus Lawrence, 1967b: 124, fig 13. MO: Boone Co., 10 mi S. Columbia; MI/ KS lA MO IL IN OH KY/ NY PA NJ MD DC VA/ VT MA/ TX/ AR LA MS AL TN GA SC NC FL/ W. Ind. TYPE DEPOSITORY: MCZC. sallei, of authors, in part, not Mellie. ECOLOGY: Lawrence, 1973. HOST: Phellinus gilvus, Pogonomyces hydnoides, Fomes fomentarius, Inonotus radiatus, Ganoderma spp., various other Polyporaceae. quadricornis Gorham, 1886: 359. Mex.: Tuxtla; TX/ Mex., C. Amer. TYPE DEPOSITORY: BMNH. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Coriolus maximus, C. hirsutus, Coriolopsis occidentalis, C. crocata, Trametes scabrosa, T. cirrifer, Pogonomyces hydnoides. sallei (Mellie), 1848: 377, pl. 12, fig. 22 (Ennearthron), LA: New Orleans; WI MI ON PQ/ NE KS lA MO IL IN OH KY/ NY PA NJ MD WV VA/ VT MA CT/ TX OK/ AR LA AL TN GA NC FL. TYPE DEPOSITORY: MNHP. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Ganoderma applanatum, G. curtisii, G. lobatum, G. zonatum, Pogonomyces hydnoides, Fomes sclerodermeus, Oxyporus populinus, various other Polyporaceae. CIIDAE 11 schaefferi Dury, 1917: 25. TX: Brownsville; TX/ Mex. TYPE DEPOSITORY: CMNH. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Ganoderma sp. singularis (Dury), 1917: 14 (XestocisX OH: Cincinnati; MN WI MI ON/ NE lA IL IN OH KY/ NY PA MD DC VA/ MA/ TX OK/ AR LA MS AL TN SC NC FL/ C. Amer. TYPE DEPOSITORY: CMNH. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Phellinus gilvus, P. igniarius, P. robiniae, P. ribis, Phellinus spp., Melanoporia nigra, Ganoderma spp., Gloeophyllum saepiarium, various other Polyporaceae. thoracicornis (Ziegler), 1845: 270 (Cis). Carolina; MB/ MN WI MI ON PQ/ NE KS lA MO IL IN OH KY/ NY PA NJ MD DC VA/ ME NH VT MA RI CT/ TX/ AR LA MS AL TN GA SC NC FL. TYPE DEPOSITORY: MCZC. mellyi Mellie, 1848: 369 (Ennearthron). N. Amer. TYPE DEPOSITORY: MUS. Nat. His., Geneva. pumicatus Mellie, 1848: 333, pi. 11, fig. 11 (Cis). LA: New Orleans. TYPE DEPOSITORY: MNHP. unicolor Casey, 1884: 37 (Ennearthron). NY: Long Isl., Willet's Point. TYPE DEPOSITORY: USNM. laminifrons Casey, 1898: 89 (Ennearthron). LA: Morgan City. TYPE DEPOSITORY: USNM. piceus Casey, 1898: 88 (Ennearthron). TX: Columbus. TYPE DEPOSITORY: USNM. oblongus Blatchley, 1910: 900 (Ennearthron). IN: Marion Co. TYPE DEPOSITORY: PURC. thoracicus (error) Dalla Torre, 1911: 19. bifoveatus Dury, 1917: 26. OH: Cincinnati. TYPE DEPOSITORY: CMNH. thoracicinus (error) Sherborn, 1931: 6493. IMMATURE STAGES: Boeving and Craighead, 1931: pi. 92. TAXONOMY: Lawrence, 1967b. REDESCRIPTION: Lawrence, 1967b. ECOLOGY: Lawrence, 1967b and 1973. HOST: Hirschioporus pargamenus, H. abietinus, H. sector, Lenzites betulina, L. elegans, Coriolus versicolor, Fomitella supina, Bjerkandera adusta, various other Basidiomycetes. Genus Dury Sulcacis Dury, 1917: 20. Type-species: Sulcacis lengi Dury (design, by Lawrence, 1965: 278). Entypus Redtenbacher, 1847: 350 (preoccupied Dahlbom, 1843). Type-species: Cis affinis Gyllenhal (monot.). Type species was misidentified as Apate fronticornis Panzer. Entypocis Lohse, 1964: 121 (as subgenus). Type-species: Cis bidentulus Rosenhauer (orig. des.). TAXONOMY: Lawrence, 1965 and 1971. KEYS: Lawrence, 1971. curtulus (Casey), 1898: 83 (Cis). NY; BC WA OR ID/ AB SK MB MT ND/ WI MI PQ/ CA NV/ WY UT CO/ NE lA MO IL IN/ NY PA NJ MD/ NH MA CT/ AZ NM/ NC. TYPE DEPOSITORY: USNM. montana Casey, 1898: 82 (Cis). MT: Missoula. TYPE DEPOSITORY: USNM. sorror Casey, 1898: 83 (Cis). BC: Vancouver Isl., Victoria. TYPE DEPOSITORY: USNM. cylindricus Dury, 1917: 8 (Cis). OR: Umatilla Co. TYPE DEPOSITORY: CMNH. 12 COLEÓPTERA CATALOG

niger Dury, 1917: 21. IL: southern. TYPE DEPOSITORY: CMNH. criddlei Dury, 1919: 158 (Cis). MB: Aweme. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Coriolus versicolor, C. hirsutas, Lenzites betulina, Funalia hispida, Bjerkandera adusta, Pycnoporus sanguineus, various other Basidiomycetes. lengi Dury, 1917: 21. VT: Bennington Co.; MI/ KS lA MO IL IN OH/ NY PA NJ MD DC VA/ ME NH VT MA/ TX/ AR MS SC NC. TYPE DEPOSITORY: CMNH. ECOLOGY: Lawrence, 1973. HOST: Corio/ws versicolor, C. pubescens, C. hirsutus, Lenzites betulina. Genus MALACOCIS Gorham Malacocis Gorham, 1886: 356. Type-species: Malacocis championi Gorham (monot.). Brachycis Casey, 1898: 86. Type-species: Brachycis brevicollis Casey (monot.). TAXONOMY: Lawrence, 1971. brevicollis (Casey), 1898: 87 (Brachycis). NY: Ithaca; MB/ MN WI ON PQ/ KS lA MO IL IN OH KY/ NY PA NJ MD DC WV/ ME NH VT MA/ TX OK/ LA MS AL TN GA SC NCFL. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973. HOST: Phellinus gilvus, P. igniarius, P. laevigatus, P. licnoides, P. pini, P. robiniae, P. robustus, Phellinus spp., Inonotus corrosus, various other Polyporaceae. Tribe OROPHIINI Genus RHOPALODONTUS Mellie Rhopalodontus Mellie, 1847: 109 (justified emendation). Type-species: Cis perforatus Gyllenhal (monot.). Ropalodontus Mellie, 1847: 109 (original spelling, emended). Type-species: Cis perforatus Gyllenhal (monot.). TAXONOMY: Lawrence, 1971. americanus Lawrence, 1971: 507, figs. 25, 31, 59. WL Oneida Co., Woodruff; WL TYPE DEPOSITORY: FMNH. ECOLOGY: Lawrence, 1973. HOST: Piptoporus betulinus, Hapalopilus fibrillosus, Russula sp. Genus OCTOTEMNUS Mellie Octotemnus Mellie, 1847: 110. Type-species: Cis glabriculus Gyllenhal (design, by Jacquelin du Val, 1861: 239). TAXONOMY: Lawrence, 1971. laevis Casey, 1898: 91. HI; AK/ BC WA OR/ AB SK MB/ MN WI MI ON PQ/ NS/ CA/ KS lA IL IN OH KY/ NY PA NJ MD DC WV VA/ ME NH VT MA RI CT/ AL TN NC. denudatus Casey, 1898: 91. CA. TYPE DEPOSITORY: USNM. ECOLOGY: Lawrence, 1973 and 1974a. HOST: Coriolus versicolor, C. pubescens, C. hirsutus, C. conchifer, Lenzites betulina, Ganoderma applanatum, Bjerkandera adusta, various other Basidiomycetes. 13 BIBLIOGRAPHY Abdullah, M. 1973 The systematic position of Cisidae (Heteromera) including a catalogue of the world and comments on central European families of (Coleóptera). Zoolo- gische Beitraege, n. ser., vol. 19, pp. 189-246. Bach, M. 1852 Kaeferfauna fuer Nord- und Mitteldeutschland mit besonder Ruecksicht auf die preussischen Rheinlande. Hoelscher, Coblenz, Band II (first pt.), 148 pp. Blatchley, W. S. 1910 An illustrated descriptive catalogue of the Coleóptera or beetles (exclusive of the Rhynchophora) known to occur in Indiana with bibliography and descriptions of new species. Bulletin no. 1 Indiana Department of Geology and Natural Re- sources. Indianapolis, Indiana, 1386 pp., illus. Reprinted by Nature Publishing Co. in 1910. Boeving, A. G. and F. C. Craighead 1931 An illustrated synopsis of the principal larval forms of the order Coleóptera. Ento- mológica Americana, n. ser., vol. 11, pp. 1-351, illus. Casey, T. L. 1884 Contributions to the descriptive and systematic coleopterology of North America. Part I. Philadelphia, 60 pp., illus. Casey, T. L. 1898 Studies in the Ptinidae, Cioidae, and Sphindidae of America. Journal of the New York Entomological Society, vol. 6, pp. 61-93. Casey, T. L. 1900 Review of the American Corylophidae, Cryptophagidae, Tritomidae, and Dermesti- dae, with other studies. Journal of the New York Entomological Society, vol. 8, pp. 51-172. Crowson, R. A. 1938 The metendosternite in Coleóptera: a comparative study. The Transactions of the Royal Entomological Society of London, vol. 87, pp. 397-415. Crowson, R. A. 1960 The phylogeny of Coleóptera. Annual Review of Entomology, vol. 5, pp. 111-134. Crowson, R. A. 1966 Observations on the constitution and subfamilies of the family Melandryidae. EOS Revista Española de Entomología, vol. 41, pp. 507-513. Crowson, R. A. 1967 The natural classification of the families of Coleóptera. Addenda and corrigenda. The Entomologist's Monthly Magazine, vol. 103, pp. 209-214. Dalla Torre, K. W. von 1911 Cioidae. In Coleopterorum catalogus auspiciis et auxilio W. Junk editus a S. Schen- kling. Junk, Berlin, pt. 30, (vol. 16), 32 pp. Desmarest, E. 1860 Tome 17. Coléoptères. Partie 3. In J. C. Chenu (ed.). Encyclopédie d'histoire natur- elle ou traite complet de cette science. Didot, Paris, 360 pp., illus. Dury, C. 1917 Synopsis of the coleopterous family Cisidae (Cioidae) of America north of Mexico. Journal of the Cincinnati Society of Natural History, vol. 22, no. 2, pp. 1-27. Dury, C. 1919 A new ciside genus with new species from Manitoba. The Canadian Entomologist, vol. 51, p. 158. 14 COLEÓPTERA CATALOG

Fall, H. C. 1917 New Coleóptera, VI and VIL The Canadian Entomologist, vol. 49, pp. 163-171, 385- 390. Gorham, H. S. 1888 Cleridae (part), Ptinidae, Bostrychidae, Cioidae. In Biología Centrali-America, In- secta. Coleóptera. F. Godman and O. Salvin, eds. R. H. Porter, London, vol. 3, pt. 2, pp. 169-224, illus. Gorham, H. S. 1886 Supplement to Malacodermata (part). In Biología Centrali-Americana, Insecta. Co- leóptera. F. Godman and O. Salvin, eds. R. H. Porter, London, vol. 3, pt. 2, pp. 313-360, illus. Gorham, H. S. 1898 On the serricorn Coleóptera of St. Vincent, Grenada, and the Grenadines (Malaco- dermata, Ptinidae, Bostrychidae), with descriptions of new species. Proceedings of the Zoological Society of London, pp. 315-333, illus. Gyllenhal, L. 1827 Insecta Svecica. Classis I. Coleóptera sive Eleuterata. Tomus I. Pars. IV. Fleischer, Leipzig, 761 pp. Hatch, M. H. 1961 The beetles of the Pacific Northwest. University of Washington Publications in Bi- ology, vol. 16, pt. 3, ix and 503 pp., illus. Horn, G. H. 1894 The Coleóptera of Baja California. Proceedings of the California Academy of Sci- ences, ser. 2, vol. 4, pp. 302-449 Jacquelin du Val, P. N. C. 1861 Manuel entomologique. Genera des Coléoptères d'Europe comprenant leur classifi- cation en familles naturelles, la description de tous les genres, des tableaux syn- optiques destines a faciliter l'étude, le catalogue de toutes les espèces, de nom- breux dessins au trait de caracteres. Tome 3 (part). DeyroUe, Paris, pp. 236-240, cat. p. 168. Kraus, E. J. 1908 New bicolored Cioidae. (Coleóptera). Proceedings of the Entomological Society of Washington, vol. 10, pp. 74-81, illus. Latreille, P. A. 1796 Precis des caracteres génériques des Insectes, disposes dans un ordre naturel. Bor- deaux, 201 pp. Latreille, P. A. 1802 Histoire naturelle, genérale, et particulière des Crustacés et des Insectes. Dufart, Paris, vol. 3, 467 pp. Lawrence, J. F. 1965 Comments on some recent changes in the classification of the Ciidae (Coleóptera). Bulletin of the Museum of Comparative Zoology, vol. 133, pp. 273-293. Lawrence, J. F. 1967a Biology of the parthenogenetic fungus Cis fuscipes Mellie (Coleóptera: Ciidae), Breviora, no. 258, 14 pp. Lawrence, J. F. 1967b Delimitation of the genus Ceracis (Coleóptera: Ciidae) with a revision of North American species. Bulletin of the Museum of Comparative Zoology, vol. 136, pp. 91-144. Lawrence, J. F. 1971 Revision of the North American Ciidae (Coleóptera). Bulletin of the Museum of Comparative Zoology, vol. 142, pp. 419-522. CIIDAE 15

Lawrence, J. F. 1973 Host preference in ciid beetles (Coleóptera: Ciidae) inhabiting the fruiting bodies of Basidiomycetes in North America. Bulletin of the Museum of Comparative Zoo- logy, vol. 145, no. 3, pp. 163-212. Lawrence, J. F. 1974a The ciid beetles of California (Coleóptera: Ciidae). Bulletin of California Survey, vol. 17, 41 pp. Lawrence, J. F. 1974b The of Sphindocis denticollis Fall and a new subfamily of Ciidae (Coleóptera: Heteromera). Breviora, vol. 424, 14 pp. Leach, W. E. 1819 In Samouelle, The entomologist's useful compendium; or an introduction to the knowledge of British , comprising the best means of obtaining and pre- serving them, and a description of the apparatus generally used; together with the genera of Linne, and the modern method of arranging the classes Crustacea, Myriapoda, spiders, mites and insects from their affinities and structure, accord- ing to the views of Dr. Leach. Also an explanation of the terms used in entomol- ogy; a calendar of the times of appearance and usual situations of near 3,000 species of British insects; with instructions for collecting and fitting up objects for the microscope. Thomas Boys, London, 496 pp. LeConte, J. L. 1866 List of the Coleóptera of North America. Smithsonian Miscellaneous Collections, vol. 6, no. 140, pp. 1-78. Leng, C. W. 1918 Notes on some changes in the list of Coleóptera. Journal of the New York Entomo- logical Society, vol. 26, pp. 201-211. Lohse, G. A. 1964 Die in Mitteleuropa vertretenen Gattungen der Cisidae. (1. Beitrag zur Kenntniss der mitteleuropaeischen Cisidae). Entomologische Blaetter fuer Biologie und Sys- tematik der Kaefer, vol. 60, pp. 174-181. Mannerheim, C. G. von 1843 Beitrag zur Kaefer-Fauna der aleutischen Inseln, der Insel Sitkha und Neu-Cali- forniens. Bulletin de la Société Imperiale des Naturalistes de Moscou, vol. 16, no. 2, pp. 175-314. Mannerheim, C. G. von 1852 Zweiter Nachtrag zur Kaefer-Fauna der nord-amerikanischen Laender des russis- chen Reiches. Bulletin de la Société Imperiale des Naturalistes de Moscou, vol. 25, no. 1, pp. 283-387. Mannerheim, C. G. von 1853 Dritter Nachtrag zur Kaefer-Fauna der nord-amerikanischen Laender des russis- chen Reiches. Bulletin de la Société Imperiale des Naturalistes de Moscou, vol. 26, no. 3, pp. 95-273. Mellie, J. 1847 Melanges et nouvelles. Revue Zoologique, par la Société Cuvierienne, vol. 10, pp. 108-110. Mellie, J. 1848 Monographie de l'ancien genre Cis des auteurs. Annales de la Société Entomologi- que de France, ser. 2, vol. 6, pp. 205-274, 313-396, illus. Peterson, A. 1951 Larvae of insects. An introduction to Nearctic species. Part II. Coleóptera, Díptera, Neuroptera, Siphonaptera, Mecoptera, Trichoptera. Peterson, Columbus, Ohio, v and 416 pp., illus. 16 Peyerimhoff, P. de 1918 Nouveaux Coléoptères du Nord-Africain. (Vingt-huitième note: Faune du Pin d'Alep). Bulletin de la Société Entomologique de France, 1918, pp. 141-143. Redtenbacher, L. 1847 Fauna Austríaca. Die Kaefer. First edition (Part 3). Carl Gerold, Wien, pp. 321-480, illus. Reitter, E. 1878 Neue Cioidae. Mittheilungen des muenchener entomologischen Vereins., vol. 2, pp. 32-37. Roubal, J. 1936 Beschreibung einer neuen Untergattung, einer Varietaet und einer Form der Co- leopteren. Festschrift zum 60. Geburstage von Professor Dr. Embrik Strand, Riga, vol. 1, pp. 53-58. Sahlberg, J. 1901 Coleóptera nova vel minus cognita faunae Fennicae. Acta Societatis pro Fauna et Flora Fennica, vol. 19, no. 3, pp. 1-23. Sherborn, C. 1931 Index animalium. Section 2, Part 26. British Museum, London, pp. 6363-6582. Tanner, V. M. 1934 Coleóptera of Zion National Park, No. 2. Annals of the Entomological Society of America, vol. 27, pp. 43-49. Thomson, C, G. 1863 Skandinaviens Coleóptera, synoptiskt bearbetade, Tom. 5. Lundberg, Lund, 340 pp. Ziegler, D. 1845 Descriptions of new North American Coleóptera. Proceedings of the Academy of Natural Sciences of Philadelphia, vol. 2, pp. 266-272. INDEX

Names are indexed as follows: ephippiam 3 CAPITALS: All names for taxa above the generic Eriaaalas Thomson 1 level; explanata Lawrence, Hadraule 8 Boldface: Valid generic and subgeneric names; falli Blatchley, Cis 6 Roman: Valid specific and subspecific names; . festivulus Lawrence, Cis 3 Italic: All invalid names such as synonyms, nomina floridae Dury, Cis 3 nuda, and extra-limital taxa even though valid. forsteri Abdullah, Cis 2 fraternas Casey, Cis 5 Parentheses around an author's name indicate that frosti Dury, Cis 2 the specific name has been transferred from its original fuscipes Mellie, Cis 3 genus. The generic name following the author's name in- grossalas (Casey), Ceracis 9 dicates the present placement of the species. Synonyms Hadraule Thomson 8 of species-group names are listed with the original spell- hatchi Lawrence, Cis 2 ing. hirsutus Casey, Cis 4 acritus Lawrence, Cis 1 horridulus Casey, Cis 4 americanus Lawrence, Rhopalodontus 12 huachucae Dury, Cis 4 americanus Mannerheim, Cis 2 huesanus Kraus, Orthocis 7 angustus Hatch, Cis 2 hystriculus Casey, Cis 4 arizonae Dury, Cis 6 illastris Casey, Cis 6 aterrimus Casey, Orthocis 7 impressas Casey, Cis 4 atripennis Mellie, Cis 3 indistinctus Hatch, Dolichocis 7 aurisquamosum Lawrence, Ennearthron 6 insolens (Casey), Cis 2 biarmatus Mannerheim, Cis 2 jalichi Dury, Cis 5 bicarinatus 2 Knablia Roubal 8 bicolor Dury, Cis 4 krausi Dalla Torre, Cis 4 bifoveatus Dury, Ceracis 11 laevis Casey, Octotemnus 12 bilimeki (Reitter), Strigocis 7 laminatus Mellie, Cis 4 bimaculatus Kraus, Cis 4 laminifrons (Casey), Ceracis 11 blaisdelli (Casey), Hadraule 8 lengi Dury, Sulcacis 12 blatchleyi Dury, Cis 5 levellei 4 Brachycis Casey 12 levettei (Casey), Cis 4 brevicollis (Casey), Malacocis 12 lodingi Dury, Cis 2 californicus (Casey), Ceracis 9 longulus Dury, Orthocis 7 Carolinas Casey, Cis 4 macilentas Casey, Cis 5 caseyi Dalla Torre, Cis 6 Macrocis Reitter 1 castlei (Dury), Cis 2 magister Lawrence, Ceracis 9 cayensis Lawrence, Cis 2 Malacocis Gorham 12 Ceracis Mellie 8 manitoba Dury, Dolichocis 7 chevrolatii Mellie, Cis 4 Maphaea 8 CIINAE 1 Maphoca Casey 8 CIINI 1 maritimus (Hatch), Cis 4 Cis Latreille 1 megastictus Lawrence, Cis 5 coarctata (Peyerimhoff), Hadraule 8 Mellieicis Lohse 7 coloradensis (Dury), Ceracis 9 mellyi (Mellie), Ceracis 11 compactas (Dury), Ceracis 9 miles (Casey), Cis 5 confusas Blatchley, Cis 5 minor (Hatch), Cis 2 congestus Casey, Cis 2 minutissimus (Mellie), Ceracis 9 convergens (Casey), Ceracis 9 minutus Dury, Ceracis 9 cornelli Lawrence, Cis 2 monocerus Lawrence, Ceracis 9 cornutus Blatchley, Cis 3 montana (Casey), Sulcacis 11 creberrimus Mellie, Cis 3 mormonicas Casey, Cis 4 cribrum Casey, Plesiocis 8 moznettei (Dury), Cis 3 criddlei (Dury), Sulcacis 12 multipunctatus (Mellie), Ceracis 10 crinitus Lawrence, Cis 3 niedhauki Lawrence, Cis 5 curtulus (Casey), Sulcacis 11 niger Dury, Sulcacis 12 curtus (Mellie), Ceracis 9 nigropunctatus Lawrence, Ceracis 10 cylindricas (Dury), Sulcacis 11 nitidas (Hatch), Cis 2 dauisi (Dury), Strigocis 8 nabillas Gorham, Cis 3 denticoUis Fall, Sphindocis 1 nabilas 3 denadatas Casey, Octotemnus 12 obesas (Mellie), Ceracis 9 Diphyllocis 8 oblongas (Blatchley), Ceracis 11 discolor (Casey), Ceracis 9 obrieni Lawrence, Ceracis 10 discolor Lawrence, Cis 3 Octotemnus Mellie 12 dixiensis (Tanner), Ceracis 9 opacicollis Dury, Strigocis 8 Dolichocis Dury 7 opalescens (Casey), Strigocis 8 dabias Mellie, Cis 4 oregonas (Dury), Ceracis 9 dunedinensis Leng, Cis 3 OROPHIINI 12 duplex Casey, Cis 3 Orthocis Casey 7 daryi Leng, Cis 4 oweni (Hatch), Cis 2 ednae (Hatch), Cis 6 pallens Casey, Cis 4 elongatula (Gyllenhal), Hadraule 8 paludas (Kraus), Orthocis 7 Ennearthron 7 pecki Lawrence, Ceracis 10 Ennearthron Mellie 6 piceas (Casey), Ceracis 11 Ennearthram 6 pistoria Casey, Cis 5 Entypocis Lohse 11 Pityocis Peyerimhoff 8 Entypas Redtenbacher 11 Plesiocis Casey 8 ephippiatus Mannerheim, Cis 3 powelli Lawrence, Ceracis 10 17 18 puberulus Mellie, Cis 3 Sphindocis Fall 1 pulcher Kraus, Orthocis 7 stereophilus Lawrence, Cis 5 pullulus (Casey), Ceracis 10 striata (Sahlberg), Hadraule 8 pumicatus (Mellie), Ceracis 11 Strigocis Dury 7 punctatus (Mellie), Orthocis 7 strigulosus (Hatch), Cis 2 punctulatus Casey, Ceracis 10 striolatus Casey, Cis 5 pusillus Dury, Cis 3 subfuscus Gorham, Cis 5 quadricornis Gorham, Ceracis 10 subtilis Mellie, Cis 5 quadridentatus (Dury), Cis 5 Sulcacis Dury H reßexus (Hatch), Cis 6 tetracentrum Gorham, Cis 6 Rhopalodontus Mellie 12 thoracicinus H robiniophilus Lawrence, Cis 5 thoracicornis (Ziegler), Ceracis 11 Ropalodontus Mellie 12 thoracicus H rotundulus Lawrence, Cis 5 transversatus (Kraus), Orthocis 7 rubriculus Lawrence, Ceracis 10 tridentatus Mannerheim, Cis 6 sallei 10 tristis Mellie, Cis 6 sallei (Mellie), Ceracis 10 unicolor (Casey), Ceracis 11 schaefferi Dury, Ceracis 11 unicornis (Casey), Ceracis 9 serricollis Dury, Cis 2 ursulinus Casey, Cis 6 setulosus Mellie, Cis 6 versicolor Casey, Cis 6 singularis (Dury), Ceracis 11 vitulus Mannerheim, Cis 6 sorror (Casey), Sulcacis 11 wenzeli Dury, Cis 6 spenceri (Hatch), Ennearthron 7 Xestocis Casey 1 SPHINDOCIINAE 1

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