In Vivo Bone Strain and Finiteelement Modeling of the Craniofacial Haft In
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JVP 26(3) September 2006—ABSTRACTS
Neoceti Symposium, Saturday 8:45 acid-prepared osteolepiforms Medoevia and Gogonasus has offered strong support for BODY SIZE AND CRYPTIC TROPHIC SEPARATION OF GENERALIZED Jarvik’s interpretation, but Eusthenopteron itself has not been reexamined in detail. PIERCE-FEEDING CETACEANS: THE ROLE OF FEEDING DIVERSITY DUR- Uncertainty has persisted about the relationship between the large endoskeletal “fenestra ING THE RISE OF THE NEOCETI endochoanalis” and the apparently much smaller choana, and about the occlusion of upper ADAM, Peter, Univ. of California, Los Angeles, Los Angeles, CA; JETT, Kristin, Univ. of and lower jaw fangs relative to the choana. California, Davis, Davis, CA; OLSON, Joshua, Univ. of California, Los Angeles, Los A CT scan investigation of a large skull of Eusthenopteron, carried out in collaboration Angeles, CA with University of Texas and Parc de Miguasha, offers an opportunity to image and digital- Marine mammals with homodont dentition and relatively little specialization of the feeding ly “dissect” a complete three-dimensional snout region. We find that a choana is indeed apparatus are often categorized as generalist eaters of squid and fish. However, analyses of present, somewhat narrower but otherwise similar to that described by Jarvik. It does not many modern ecosystems reveal the importance of body size in determining trophic parti- receive the anterior coronoid fang, which bites mesial to the edge of the dermopalatine and tioning and diversity among predators. We established relationships between body sizes of is received by a pit in that bone. The fenestra endochoanalis is partly floored by the vomer extant cetaceans and their prey in order to infer prey size and potential trophic separation of and the dermopalatine, restricting the choana to the lateral part of the fenestra. -
Proceedings of the United States National Museum
LIS T OF ILL US TRA TIONS. IX Pa«e. Apices of ovipositors : o, Polysphincta texaiui Cresson ; &, Eymenoepi- meois iciltii (Cresson); Mandible; c, H. tmltii (Cresson) 389 Apex of ovipositor of Theronia fulvesceiis Cresson 389 Apices of ovipositors: a, Itoplectis canqMisitor Say; b, Apechthis pictir cornis (Cresson) 389 Apex of abdomen of female of Coleocentrus occidentalis 390 Areolet of Lahena grallatoi- (Say) 390 Areolets: a, Tromatohia rufovariata (Cresson) ; b, Itoplectis conquisitor (Say) ; c, Epiurus alboricta (Cresson) 390 Sessile first tergite of Perithous pleuralis (Cresson) 390 Petioatel first tergite of Xorides yukonensis (Rohwer) 390 Mandible of Poemenva americana (Cresson) 391 Female of Labena confusa Rohwer 412 Female of Rhyssella nitida (Cresson) 424 Propodeum of Zonde* picea*u« (Rohwer) 434 Front view of head of Xoi'ides albopictus (Cresson) 437 Propodeum and basal abdominal segments of Xorides albopictus (Cresson) 437 Apical abdomen segments of Xorides albopictus (Cresson) 437 Female of Xorides rileyi (Ashmead) 440 Front view of head of Deuteroxorides caryae (Harrington) 446 Propodeum and basal abdominal segments of Deuteroxorides caryae (Harrington) 446 Apical abdominal segments of Z)c«feroa:ori<Ze« caryae (Harrington) 446 Female of Deuteroxorides caryae (Harrington) 447 Female of Odontomerus canadensis Provancher 459 Female of Phytodietus annulatus (Provancher) 467 Femaie of Phydotietus annulatus (Provancher) 467 Diagram showing relation of dike to vein, Hecla Mine. A, Dike; B, Crushed material with quartz and ore; C, Galena; D, Quartzite wall rock ; width of section 15 feet 481 Showing relation to dikes (A) to vein (B) on No. 1 level of Marsh Mine_ 484 Showing relation of dike (A) to vein (B) on 2,000-foot level of Standard- Mammoth Mine 492 Showing relation of dike (A) to vein (B) on 1,200-foot level of Standard- Mammoth Mine 492 Coleocentrus occidentalis. -
Fossil Primates
AccessScience from McGraw-Hill Education Page 1 of 16 www.accessscience.com Fossil primates Contributed by: Eric Delson Publication year: 2014 Extinct members of the order of mammals to which humans belong. All current classifications divide the living primates into two major groups (suborders): the Strepsirhini or “lower” primates (lemurs, lorises, and bushbabies) and the Haplorhini or “higher” primates [tarsiers and anthropoids (New and Old World monkeys, greater and lesser apes, and humans)]. Some fossil groups (omomyiforms and adapiforms) can be placed with or near these two extant groupings; however, there is contention whether the Plesiadapiformes represent the earliest relatives of primates and are best placed within the order (as here) or outside it. See also: FOSSIL; MAMMALIA; PHYLOGENY; PHYSICAL ANTHROPOLOGY; PRIMATES. Vast evidence suggests that the order Primates is a monophyletic group, that is, the primates have a common genetic origin. Although several peculiarities of the primate bauplan (body plan) appear to be inherited from an inferred common ancestor, it seems that the order as a whole is characterized by showing a variety of parallel adaptations in different groups to a predominantly arboreal lifestyle, including anatomical and behavioral complexes related to improved grasping and manipulative capacities, a variety of locomotor styles, and enlargement of the higher centers of the brain. Among the extant primates, the lower primates more closely resemble forms that evolved relatively early in the history of the order, whereas the higher primates represent a group that evolved more recently (Fig. 1). A classification of the primates, as accepted here, appears above. Early primates The earliest primates are placed in their own semiorder, Plesiadapiformes (as contrasted with the semiorder Euprimates for all living forms), because they have no direct evolutionary links with, and bear few adaptive resemblances to, any group of living primates. -
The Mechanical Function of the Postorbital Bar in Eulemur Fulvus Analyzed Using Finite Element Analysis
University at Albany, State University of New York Scholars Archive Anthropology Honors College 4-2010 The Mechanical Function of the Postorbital Bar in Eulemur fulvus Analyzed Using Finite Element Analysis Danielle Parisi University at Albany, State University of New York Follow this and additional works at: https://scholarsarchive.library.albany.edu/honorscollege_anthro Part of the Anthropology Commons Recommended Citation Parisi, Danielle, "The Mechanical Function of the Postorbital Bar in Eulemur fulvus Analyzed Using Finite Element Analysis" (2010). Anthropology. 23. https://scholarsarchive.library.albany.edu/honorscollege_anthro/23 This Honors Thesis is brought to you for free and open access by the Honors College at Scholars Archive. It has been accepted for inclusion in Anthropology by an authorized administrator of Scholars Archive. For more information, please contact [email protected]. The Mechanical Function of the Postorbital Bar in Eulemur fulvus Analyzed Using Finite Element Analysis Danielle Parisi Senior Thesis Dr. David Strait April 27, 2010 1 INTRODUCTION This study examines the functional anatomy of the primate postorbital bar. Postorbital bars are “bony arches that encompass the lateral aspect of the eye and form part of a circular orbit,” (Heesy 2005). They are formed by the dorsal part of the frontal bone and the ventral part of the zygomatic. Postorbital bars are not unique to primates. In fact, historically, they have evolved many times in many different mammalian species. However, they are found in all primates and, thus, are considered a defining characteristic of the family. Nevertheless, while their importance is undeniable, their reason for evolving and their functional role is highly debated among scholars. -
Paleontology Lab: Mammalian Diversity
Paleontology Lab: Mammalian diversity Objectives: • Be able to recognize a mammal. • Be able to describe mammalian traits and evolutionary adaptations. • Be able to identify the major orders of mammals. • Be able to identify the characteristics of primates. Materials needed for this lab : Preserved specimens and skeletons of various mammals including primates Human brain and sheep brain for comparison Class Mammalia: Mammals are warm-blooded vertebrates with a fully divided (four-chambered) heart that nourish their young with milk. Most mammals are furry, but some marine mammals have a thick layer of subcutaneous fat instead. Living mammals can be divided into a few egg- laying species (the Monotremes, including the platypus and echidna), the pouched mammals (Marsupials, including opossums, kangaroos, koalas, and others), and the placental mammals ( Eutheria ). If we include fossil lineages, we might more appropriately divide mammals into Protheria and Theria. Subclass Prototheria: These are mammals whose molar teeth have cusps in antero-posterior rows and whose petrosal bone is exposed within the eye socket. Most of these mammals are extinct. Extinct prototherians include the Triconodonta, Docodonta, and Multituberculata. Order Monotremata: Egg-laying mammals, including only the platypus and the echidna ("spiny anteater"), both now confined to Australia and New Guinea. Examine any monotreme specimens that may be available. Subclass Theria: Therian mammals, with molar teeth whose molar cusps were originally arranged in opposing triangles with shear surfaces between them, the so-called tribosphenic tooth pattern. The petrosal bone is never exposed inside the eye socket (orbit). Therian mammals are divided into infraclasses Pantotheria, Metatheria, and Theria. Infraclass Pantotheria includes the extinct orders Symmetrodonta and Eupantotheria. -
ANTH 42: Primates in Nature Rules to a Story •What? •When? •Where
Rules to a story! … but today, not in that order:! •!What?! When? Dating methods and deep time! •!When?! Where? How do fossils form - ANTH 42: Primates in Nature! taphonomy! •!Where?! What? The primate fossil record! Lecture 2:! •!Why?! Why? Evolution and its Primate evolution! mechanisms (including natural selection) [later lecture!]! http://weber.ucsd.edu/~jmoore/courses/anth42web/! WHEN: Dating Bishop James Ussher: 1650 WHEN: Dating methods! Uniformitarianism methods! calculated from Bible that & the earth! Creation occurred 4004 BC! Strata laid down in layers, oldest are deepest.! Mary Ann Mantell: 1820 discovered “Iguanodon” tooth (Sussex, UK)*! Younger or older?! Richard Owen: 1842 coined Sometimes, word “Dinosaur”! geological 1854: Dinner in the Iguanodon! processes can Was hard to reconcile 6,000 years with creatures SO muddle that up.! different and so clearly extinct.! Uniformitarianism violated.! * Other giant bones known, but this specimen first to kick off ‘modern’ interest! But with more data, can usually figure it out.! WHEN: Dating methods! WHEN: Dating methods! 1566! " 100 years trying to date fossils. Absolute. ! Recognized fossil ‘stages’ - rocks with Dendrochronology! no crinoids or coral overlaid by rocks (tree rings)! Growth depends on with crinoids and coral, but no rainfall; annual rings.! dinosaurs, so could talk about “age of Count inward from crinoids” and “age of dinosaurs”. outermost ring, get age of tree.! Knew the order taxa showed up Bristlecone pine 5,000 yrs! (relative dating), but argue about when in years (absolute dating).! Buried log! WHEN: Dating methods! 1566! WHEN: Dating methods! Absolute. Dendrochronology! Absolute. ! River oaks, S. Radiometric! Dendrochronology! Germany: > (tree rings)! 10,000 yrs! Isotopic decay - Growth depends on measurable rainfall; annual rings.! Bristlecone rates! Pines, SW Count inward from USA: 8,500! outermost ring, get age of tree.! Bristlecone pine 5,000 yrs! Buried log! Buried log! WHEN: Dating methods! • total amount of WHEN: Dating methods! Absolute. -
Exceptional Skull of Huayqueriana (Mammalia, Litopterna, Macraucheniidae) from the Late Miocene of Argentina: Anatomy, Systematics, and Paleobiological Implications
EXCEPTIONAL SKULL OF HUAYQUERIANA (M AMMALIA, LITOPTERNA, M ACRAUCHENIIDAE) FROM THE L ATE MIOCENE OF ARGENTINA: ANATOMY, SYSTEMATICS, AND PALEOBIOLOGICAL IMPLICATIONS ANALÍA M. FORASIEPI, ROSS D.E. MacPHEE, SANTIAGO HERNÁNDEZ DEL PINO, GABRIELA I. SCHMIDT, ELI AMSON, AND CAMILLE GROHÉ BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY EXCEPTIONAL SKULL OF HUAYQUERIANA (MAMMALIA, LITOPTERNA, MACRAUCHENIIDAE) FROM THE LATE MIOCENE OF ARGENTINA: ANATOMY, SYSTEMATICS, AND PALEOBIOLOGICAL IMPLICATIONS ANALÍA M. FORASIEPI IANIGLA, CCT- Mendoza, CONICET ROSS D. E. MacPHEE Department of Mammalogy, American Museum of Natural History SANTIAGO HERNÁNDEZ DEL PINO IANIGLA, CCT- Mendoza, CONICET GABRIELA I. SCHMIDT Laboratorio de Paleontología de Vertebrados (CICYTTP-CONICET) ELI AMSON Paläontologisches Institut und Museum, Universität Zürich CAMILLE GROHÉ Department of Vertebrate Paleontology, American Museum of Natural History BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Number 404, 76 pp., 30 figures, 5 tables Issued June 22, 2016 Copyright © American Museum of Natural History 2016 ISSN 0003-0090 CONTENTS Abstract.............................................................................. 3 Introduction.......................................................................... 3 Geographical and geological contexts................................................... 5 Material and methods ................................................................ 7 Abbreviations ...................................................................... -
Subfossil Lemurs of Madagascar
CHAPTER TWENTY-ONE Subfossil Lemurs of Madagascar LAURIE R. GODFREY, WILLIAM L. JUNGERS, AND DAVID A. BURNEY Madagascar’s living lemurs (order Primates) belong to a radia- on steady, gradual diversifi cation would suggest. We believe tion recently ravaged by extirpation and extinction. There that the latter scenario is more consistent with the fossil are three extinct and fi ve extant families (two with extinct record. members) of lemurs on an island of less than 600,000 km2. The question of how lemurs got to Madagascar is still far This level of familial diversity characterizes no other primate from resolved (Godinot, 2006; Masters et al., 2006; Stevens radiation. The remains of up to 17 species of recently extinct and Heesy, 2006; Tattersall, 2006a, 2006b). It is clear that (or subfossil lemurs) have been found alongside those of still Madagascar (with the Indian plate) separated from Africa extant lemurs at numerous Holocene and late Pleistocene long before primates evolved and that it arrived at its present sites in Madagascar (fi gure 21.1, table 21.1). The closest rela- position relative to Africa by 120–130 Ma (Krause et al., 1997; tives of the lemurs are the lorisiform primates of continental Roos et al., 2004; Masters et al., 2006; Rabinowitz and Woods, Africa and Asia; together with the lemurs, these comprise the 2006). Most scholars favor chance rafting of an ancestral suborder Strepsirrhini. lemur from continental Africa to Madagascar (Krause et al., Most researchers have defended an ancient Gondwanan 1997; Kappeler, 2000; Roos et al., 2004; Rabinowitz and (African or Indo-Madagascan) origin for lemurs. -
Primate Evolution: Evidence from the Fossil Record, Comparative Morphology, and Molecular Biology
YEARBOOK OF PHYSICAL ANTHROPOLOGY 2757-72 (1984) Primate Evolution: Evidence From the Fossil Record, Comparative Morphology, and Molecular Biology PHILIP D. GINGERICH Museum of Paleontology, The University of Michigan, Ann Arbor, Michigan 48109 KEY WORDS Primate evolution, Phylogeny, Stratophenetics, Cladistics, Molecular clocks ABSTRACT Our understanding of primate evolution is ultimately based on patterns of phyletic relationship and morphological change documented in the fossil record. Stratophenetic interpretation of living and fossil primates yields an objective alternative to the arbitrary scala naturae assumed implic- itly in traditional comparative biology. Fossils provide an outline of primate history constraining comparative analyses incorporating taxa and morpho- logical characteristics not represented in the fossil record. Extant taxa with- out known prehistoric relatives may be interpolated into this outline using deductive cladistic analysis of morphological characteristics and overall mo- lecular similarity. Cladistic analysis provides a method for evaluating the relative strength of stratophenetic links between taxa. The phyletic node connecting Anthropoidea-Adapoidea-Lemuroidea is analyzed here as an ex- ample: the link between Eocene Adapoidea and primitive Anthropoidea ap- pears stronger than that between Adapoidea and Lemuroidea because it is based on shared-derived rather than shared-primitive characteristics. Full integration of molecular results with morphological information requires a better understanding of rates of molecular change over geological time. Rates of molecular evolution can be studied using paleontologically documented divergence times for Prosimii-Anthropoidea (ca. 55 m.y.B.P.1, Platyrrhini- Catarrhini (ca. 40 m.y.B.P.1, and Hominoidea-Cercopithecoidea (ca. 25 m.y. B.P.). Immunological distances combined with these divergence times indi- cate that primate albumin, widely used as a molecular clock in primatology, has evolved nonlinearly over geological time. -
5. Meet the Living Primates
5. Meet the Living Primates Stephanie Etting, Ph.D., Sacramento City College Learning Objectives • Learn how primates are different from other mammals • Understand how studying non-human primates is important in anthropology • Identify different types of traits that we use to evaluate primate taxa • Describe the major primate taxa using their key characteristics • Understand your place in nature by learning your taxonomic classification One of the best parts of teaching anthropology for me is getting to spend time at zoos watching primates. What I also find interesting is watching people watch primates. I have very often heard a parent and child walk up to a chimpanzee enclosure and exclaim “Look at the monkeys!” The parent and child often don’t know that a chimpanzee is not a monkey, nor are they likely to know that chimpanzees share more than 98% of their DNA with us. What strikes me as significant is that, although most people do not know the difference between a monkey, an ape, and a lemur, they nonetheless recognize something in the animals as being similar to themselves. What people probably mean when they say “monkey” is actually “primate,” a term that refers to all organisms classified within the Order Primates and also the subject of this chapter. You may be wondering why a field dedicated to the study of humans would include the study of non- human animals.Because humans are primates, we share a wide range of behavioral and morphological traits with the other species who also fall into this group. In Chapter 2, you learned about the nature of Linnaean classification, the system we use for organizing life-forms. -
Laboratory 1 Worksheet: the Skull Objective: Learn About The
Skull_Skeleton_Lab3.doc 09/15/09 page 1 of 48 Laboratory 1 Worksheet: The Skull Objective: Learn about the mammalian skull, and be able to define and/or identify on a specimen all underlined terms. Assignment: Turn in two photos/drawings of a skull with bones and structures labeled. Use a skull of a coyote (Canis latrans) or red fox (Vulpes vulpes) and identify the following bones and other features. Canids have a fairly "primitive" skull large enough to identify different bones. For comparative purposes other skulls are shown in the figures to illustrate differences among groups or features missing from the canid skull. After identifying features on a canid skull, you should be able to find the same bones or features on skulls of other mammals. Main features of mammal skulls are the zygomatic arches (the bars on both sides of the skull) under which the jaw muscles reach from the lower jaw to the back of the head, a secondary palate that separates the mouth from the nasal passages, and a mandible (lower jaw) consisting of a single dentary bone on the left and right sides. Cranium The cranium is divided into two regions: braincase and rostrum. The braincase, more developed in mammals than in other vertebrates, contains the brain. The rostrum corresponds to the snout or muzzle. Dorsal aspect of the cranium.—Bones seen from a dorsal aspect on a canid skull are shown in Fig. 3. The nasal bones are paired bones forming a “roof” over the nasal passages. The paired premaxillary bones form the lower margin of the nasal openings (nares) and the anteriormost part of the bony palate at the anterior upper jaw. -
Frontal Fusion: Collapse of Another Anthropoid Synapomorphy
THE ANATOMICAL RECORD 291:308–317 (2008) Frontal Fusion: Collapse of Another Anthropoid Synapomorphy 1,2,3 3 ALFRED L. ROSENBERGER AND ANTHONY S. PAGANO * 1Department of Anthropology and Archaeology, Brooklyn College, CUNY, Brooklyn, New York 2American Museum of Natural History, New York, New York 3Department of Anthropology, City University of New York Graduate Center, New York Consortium in Evolutionary Primatology (NYCEP), New York, New York ABSTRACT We test the hypothesis that the fused interfrontal suture of anthro- poids is a uniquely distinguishing feature and a derived characteristic indicative of their monophyletic origin. Our survey of nonanthropoid primates and several archontan families indicates frontal fusion is wide- spread. It is most variable (fused, open or partially fused) inter- and intra-specifically among strepsirhines. The frontal bone is more commonly fused in living lemuroids and indrioids than among lorisoids. It appears to be fused regularly among Eocene adapids. Among nonanthropoid haplorhines, the interfrontal is fused in Tarsius, even in neonates and invariably in adults, probably also in all fossil tarsiiforms preserving the frontal bone, and in the late Eocene protoanthropoid Rooneyia. The plesiadapiform pattern remains uncertain, but fusion is ubiquitous among living tree shrews, colugos and bats. Distributional evidence implies that interfrontal fusion was present in the last common ancestor (LCA) of haplorhine primates and possibly in the LCA of euprimates as well. Anthropoids, therefore, cannot be defined cladistically by interfrontal fusion, not out of concern for homoplasy but because it is probably a primitive feature inherited from other taxa related to anthropoids. Fusion of the large anthropoid frontal bone, which was extended anteriorly to roof the orbits and expanded laterally in connection with a wide forebrain in the LCA of anthropoids and protoanthropoids, may have been preadap- tive to the evolution of the postorbital septum.