Comments on the GSSP for the Basal Emsian Stage Boundary: the Need for Its Redefinition
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A New Species of Tiaracrinus from the Latest Emsian of Morocco and Its Phylogeny
A new species of Tiaracrinus from the latest Emsian of Morocco and its phylogeny CHRISTIAN KLUG, KENNETH DE BAETS, CAROLE JUNE NAGLIK, and JOHNNY WATERS Klug, C., De Baets, K., Naglik, C.J., and Waters, J. 2014. A new species of Tiaracrinus from the latest Emsian of Morocco and its phylogeny. Acta Palaeontologica Polonica 59 (1): 135–145. We describe a new species of the unusual crinoid Tiaracrinus, T. jeanlemenni sp. nov. from the latest Emsian of the fa- mous mudmound locality Hamar Laghdad, Morocco. It differs from the previously known species in the higher number of ribs and the vaulted rib-fields, which is corroborated by the comparison of simple quantitative characters and ratios as well as by the results of a cluster analysis and a Principal Component Analysis. Based on the new material and the published specimens, we discuss the phylogeny of the genus and suggest that T. oehlerti and T. moravicus represent the ancestral forms of this small clade. Key words: Crinoidea, mudmounds, phylogeny, morphometry, symmetry, Devonian, Morocco. Christian Klug [[email protected]] and Carole J. Naglik [[email protected]], Paläontologisches Institut und Museum, Universität Zürich, Karl Schmid-Strasse 4, CH-8006 Zürich, Switzerland; Kenneth De Baets [[email protected]], GeoZentrum Nordbayern, Fachgruppe PaläoUmwelt, Universität Erlan- gen, Loewenichstr. 28, D-91054 Erlangen, Germany; Johnny Waters [[email protected]], Department of Geology, Appalachian State University, ASU Box 32067, Boone, NC 28608-2067, USA. Received 30 November 2011, accepted 4 June 2012, available online 6 June 2012. Copyright © 2014 C. Klug et al. This is an open-access article distributed under the terms of the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. -
Lower Devonian (Emsian) Rugose Corals from the Cantabrian Mountains, Northern Spain
Lower Devonian (Emsian) rugose corals from the Cantabrian Mountains, northern Spain STEFAN SCHRÖDER and FRANCISCO SOTO Schröder, S. and Soto, F. 2003. Lower Devonian (Emsian) rugose corals from the Cantabrian Mountains, northern Spain. Acta Palaeontologica Polonica 48 (4): 547–558. Two species of Lower Devonian rugose corals are described from the Cantabrian Mountains of Spain: Cantabriastraea cantabrica gen. et sp. nov. and Tabulophyllum bonarense sp. nov. The development of root− or buffer−like attachment structures (“rhizoid processes”) in T. bonarense indicates adaption to the soft substrate and supports corallite stabilisa− tion. Because of their strongly everted calices and a corresponding arrangement of trabeculae, the colonies of the genus Cantabriastraea are assigned tentatively to the Paradisphyllinae, constituting the first record of the subfamily in Western Europe. Some specimens give information on colony−formation of this taxon, which is induced by strong lateral budding (nonparricidal increase) of a remarkable large and long−lasting protocorallite. Key words: Rugosa, Cantabriastraea, Tabulophyllum, Devonian, Emsian, Cantabrian Mountains, ecology. Stefan Schröder [[email protected]], Geologisches Institut, Zülpicher Strasse 49A, D−50674 Köln, Germany; Francisco Soto [[email protected]],Departamento de Geologia,Universidad de Oviedo,c/ Arias de Velasco s/n, E−33005 Oviedo, Spain. Introduction Sedimentological investigations by Truyols et al. (1984), Keller (1988) and Vera de la Puente (1989) suggested that During the last 150 years a wide range of Palaeozoic fossils the lower part of the La Vid Group (Felmin Formation) cor− has been described from the Cantabrian Mountains and with responds to a supra−tidal environment, while the upper part regard to Devonian faunas the so−called “Sabero−fossils” are (La Pedrosa and Coladilla Formations) was deposited in a especially famous. -
Evaluating the Frasnian-Famennian Mass Extinction: Comparing Brachiopod Faunas
Evaluating the Frasnian-Famennian mass extinction: Comparing brachiopod faunas PAUL COPPER Copper, P. 1998. Evaluating the Frasnian-Famennian mass extinction: Comparing bra- chiopod faunas.- Acta Palaeontologica Polonica 43,2,137-154. The Frasnian-Famennian (F-F) mass extinctions saw the global loss of all genera belonging to the tropically confined order Atrypida (and Pentamerida): though Famen- nian forms have been reported in the literafure, none can be confirmed. Losses were more severe during the Givetian (including the extinction of the suborder Davidsoniidina, and the reduction of the suborder Lissatrypidina to a single genus),but ońgination rates in the remaining suborder surviving into the Frasnian kept the group alive, though much reduced in biodiversity from the late Earb and Middle Devonian. In the terminal phases of the late Palmatolepis rhenana and P linguifurmis zones at the end of the Frasnian, during which the last few Atrypidae dechned, no new genera originated, and thus the Atrypida were extĘated. There is no evidence for an abrupt termination of all lineages at the F-F boundary, nor that the Atrypida were abundant at this time, since all groups were in decline and impoverished. Atypida were well established in dysaerobic, muddy substrate, reef lagoonal and off-reef deeper water settings in the late Givetian and Frasnian, alongside a range of brachiopod orders which sailed through the F-F boundary: tropical shelf anoxia or hypońa seems implausible as a cause for aĘpid extinction. Glacial-interglacial climate cycles recorded in South Ameńca for the Late Devonian, and their synchronous global cooling effect in low latitudes, as well as loss of the reef habitat and shelf area reduction, remain as the most likely combined scenarios for the mass extinction events. -
EDITORIAL NOTE Collection of Paleontology Papers in Honor of The
Anais da Academia Brasileira de Ciências (2019) 91(Suppl. 2): e20191434 (Annals of the Brazilian Academy of Sciences) Printed version ISSN 0001-3765 / Online version ISSN 1678-2690 http://dx.doi.org/10.1590/0001-3765201920191434 www.scielo.br/aabc | www.fb.com/aabcjournal EDITORIAL NOTE Collection of Paleontology Papers in honor of the Centenary of the Brazilian Academy of Sciences ALEXANDER W.A. KELLNER* and MARINA B. SOARES Laboratório de Sistemática e Tafonomia de Vertebrados Fósseis, Departamento de Geologia e Paleontologia do Museu Nacional/UFRJ, Quinta da Boa Vista, s/n, São Cristóvão, 20940-040 Rio de Janeiro, RJ, Brazil How to cite: KELLNER AWA AND SOARES MB. 2019. Collection of Paleontology Papers in honor of the Centenary of the Brazilian Academy of Sciences. An Acad Bras Cienc 91: e20191434. DOI 10.1590/0001-3765201920191434. The Brazilian Academy of Sciences is a non-profit organization (ABC 2019) that has completed one century of existence in 2016. A series of special publications was organized by the Annals of the Brazilian Academy of Sciences (AABC) in celebration of this important date (e.g., Kellner 2017, Crespilho 2018, Cavaleiro 2018). Here we have the pleasure to introduce the final of these volumes gathering 20 original contributions in paleontology, the science dedicated to the study of all evidences of life that have been preserved in layers of deep time. The topics presented here vary from the description of new species and specimens of flying reptiles, dinosaurs, and crocodylomorphs to studies on biogeography, osteohistology, and specific contributions provided by microfossils. Over 70 authors from different countries were involved in this volume, showing the increasing international integration of Brazilian paleontologists. -
GEOLOGIC TIME SCALE V
GSA GEOLOGIC TIME SCALE v. 4.0 CENOZOIC MESOZOIC PALEOZOIC PRECAMBRIAN MAGNETIC MAGNETIC BDY. AGE POLARITY PICKS AGE POLARITY PICKS AGE PICKS AGE . N PERIOD EPOCH AGE PERIOD EPOCH AGE PERIOD EPOCH AGE EON ERA PERIOD AGES (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) HIST HIST. ANOM. (Ma) ANOM. CHRON. CHRO HOLOCENE 1 C1 QUATER- 0.01 30 C30 66.0 541 CALABRIAN NARY PLEISTOCENE* 1.8 31 C31 MAASTRICHTIAN 252 2 C2 GELASIAN 70 CHANGHSINGIAN EDIACARAN 2.6 Lopin- 254 32 C32 72.1 635 2A C2A PIACENZIAN WUCHIAPINGIAN PLIOCENE 3.6 gian 33 260 260 3 ZANCLEAN CAPITANIAN NEOPRO- 5 C3 CAMPANIAN Guada- 265 750 CRYOGENIAN 5.3 80 C33 WORDIAN TEROZOIC 3A MESSINIAN LATE lupian 269 C3A 83.6 ROADIAN 272 850 7.2 SANTONIAN 4 KUNGURIAN C4 86.3 279 TONIAN CONIACIAN 280 4A Cisura- C4A TORTONIAN 90 89.8 1000 1000 PERMIAN ARTINSKIAN 10 5 TURONIAN lian C5 93.9 290 SAKMARIAN STENIAN 11.6 CENOMANIAN 296 SERRAVALLIAN 34 C34 ASSELIAN 299 5A 100 100 300 GZHELIAN 1200 C5A 13.8 LATE 304 KASIMOVIAN 307 1250 MESOPRO- 15 LANGHIAN ECTASIAN 5B C5B ALBIAN MIDDLE MOSCOVIAN 16.0 TEROZOIC 5C C5C 110 VANIAN 315 PENNSYL- 1400 EARLY 5D C5D MIOCENE 113 320 BASHKIRIAN 323 5E C5E NEOGENE BURDIGALIAN SERPUKHOVIAN 1500 CALYMMIAN 6 C6 APTIAN LATE 20 120 331 6A C6A 20.4 EARLY 1600 M0r 126 6B C6B AQUITANIAN M1 340 MIDDLE VISEAN MISSIS- M3 BARREMIAN SIPPIAN STATHERIAN C6C 23.0 6C 130 M5 CRETACEOUS 131 347 1750 HAUTERIVIAN 7 C7 CARBONIFEROUS EARLY TOURNAISIAN 1800 M10 134 25 7A C7A 359 8 C8 CHATTIAN VALANGINIAN M12 360 140 M14 139 FAMENNIAN OROSIRIAN 9 C9 M16 28.1 M18 BERRIASIAN 2000 PROTEROZOIC 10 C10 LATE -
Tage。。,,。。。Ev。Nia。。Ystem by W
鹭。tage。。,,。。。ev。nia。。ystem by W . Ziegler and G .K lapper Th e S ubc om m is sio n on D ev on ian S tr at ig rap h y h a s f orm a ll y ag re ed up on th e n om en c la tu re f or sev en S tan d ard S t ag es o f th e D e von ian S ys tem . A ba sa l b oun d ar y str a to typ e is d e scr ib ed in th i s ar ti c le f or th e E if e lian S t ag e , c o in ci de n t wi th th e b as e of th e M idd l e D evon i an S er i es . C r it er ia f or r ec ogn i zin g a Givetian-Frasnian Stage boundary (=base of Upper M O D e von ian Se ri es ) an d a F ra sn i an -F am enn i an S ta ge bo un d ar y are al so o u tl in ed . F u tu re w ork wi ll c om - p le te th e d ef in it ion o f ba sa l b oun d ar y st ra to t yp es f o r e a c h S t a g e . In trod uc tio n A nnual m eetings of the Subcom m ission on D evonian Stra- tigraphy (SD S) have been held since its foundation in 1973, w ith im portant decisions sum m arized here on stratigraphic standards being m ade in 1979, 1981, 1982 and 1983; these have now been form ally ratified by lU G S. -
USGS Professional Paper 1662, Chapter 4
Studies by the U.S. Geological Survey in Alaska, 2000 U.S. Geological Survey Professional Paper 1662 Late Triassic (Norian) Mollusks From the Taylor Mountains Quadrangle, Southwestern Alaska By Christopher A. McRoberts1 and Robert B. Blodgett2 Abstract Such paleobiogeographic data as those presented herein are extremely useful in constraining the past geographic positions We describe a diverse molluscan fauna of silicified fossils of these mobile terranes over time, and so are of utmost utility from two localities in the Taylor Mountains D–3 quadrangle of in unraveling the tectonic history of this part of Alaska. southwestern Alaska. The molluscan fauna consists of at least 8 species of bivalves, including 1 new species, Cassianella cordillerana McRoberts n.sp., and at least 11 species of gas- Geologic Setting tropods, including 2 new species, Neritaria nuetzeli Blodgett n.sp. and Andangularia wilsoni Blodgett n.sp. Bivalve and gastropod affinities suggest an early Norian age, with taxo- The Farewell terrane of southwestern and west-central nomic similarities to several southern Alaskan tectonostrati- Alaska (fig. 1) was established by Decker and others (1994) graphic terranes (for example, Alexander and Chulitna), as as a tectonostratigraphic entity incorporating three previously well as to the South American Cordillera of Peru. The mol- named, genetically related terranes (Nixon Fork, Dillinger, lusks are associated with numerous brachiopods that also sup- and Mystic) that are relegated the status of subterranes of the port a Norian -
International Chronostratigraphic Chart
INTERNATIONAL CHRONOSTRATIGRAPHIC CHART www.stratigraphy.org International Commission on Stratigraphy v 2014/02 numerical numerical numerical Eonothem numerical Series / Epoch Stage / Age Series / Epoch Stage / Age Series / Epoch Stage / Age Erathem / Era System / Period GSSP GSSP age (Ma) GSSP GSSA EonothemErathem / Eon System / Era / Period EonothemErathem / Eon System/ Era / Period age (Ma) EonothemErathem / Eon System/ Era / Period age (Ma) / Eon GSSP age (Ma) present ~ 145.0 358.9 ± 0.4 ~ 541.0 ±1.0 Holocene Ediacaran 0.0117 Tithonian Upper 152.1 ±0.9 Famennian ~ 635 0.126 Upper Kimmeridgian Neo- Cryogenian Middle 157.3 ±1.0 Upper proterozoic Pleistocene 0.781 372.2 ±1.6 850 Calabrian Oxfordian Tonian 1.80 163.5 ±1.0 Frasnian 1000 Callovian 166.1 ±1.2 Quaternary Gelasian 2.58 382.7 ±1.6 Stenian Bathonian 168.3 ±1.3 Piacenzian Middle Bajocian Givetian 1200 Pliocene 3.600 170.3 ±1.4 Middle 387.7 ±0.8 Meso- Zanclean Aalenian proterozoic Ectasian 5.333 174.1 ±1.0 Eifelian 1400 Messinian Jurassic 393.3 ±1.2 7.246 Toarcian Calymmian Tortonian 182.7 ±0.7 Emsian 1600 11.62 Pliensbachian Statherian Lower 407.6 ±2.6 Serravallian 13.82 190.8 ±1.0 Lower 1800 Miocene Pragian 410.8 ±2.8 Langhian Sinemurian Proterozoic Neogene 15.97 Orosirian 199.3 ±0.3 Lochkovian Paleo- Hettangian 2050 Burdigalian 201.3 ±0.2 419.2 ±3.2 proterozoic 20.44 Mesozoic Rhaetian Pridoli Rhyacian Aquitanian 423.0 ±2.3 23.03 ~ 208.5 Ludfordian 2300 Cenozoic Chattian Ludlow 425.6 ±0.9 Siderian 28.1 Gorstian Oligocene Upper Norian 427.4 ±0.5 2500 Rupelian Wenlock Homerian -
Paleogeographic Maps Earth History
History of the Earth Age AGE Eon Era Period Period Epoch Stage Paleogeographic Maps Earth History (Ma) Era (Ma) Holocene Neogene Quaternary* Pleistocene Calabrian/Gelasian Piacenzian 2.6 Cenozoic Pliocene Zanclean Paleogene Messinian 5.3 L Tortonian 100 Cretaceous Serravallian Miocene M Langhian E Burdigalian Jurassic Neogene Aquitanian 200 23 L Chattian Triassic Oligocene E Rupelian Permian 34 Early Neogene 300 L Priabonian Bartonian Carboniferous Cenozoic M Eocene Lutetian 400 Phanerozoic Devonian E Ypresian Silurian Paleogene L Thanetian 56 PaleozoicOrdovician Mesozoic Paleocene M Selandian 500 E Danian Cambrian 66 Maastrichtian Ediacaran 600 Campanian Late Santonian 700 Coniacian Turonian Cenomanian Late Cretaceous 100 800 Cryogenian Albian 900 Neoproterozoic Tonian Cretaceous Aptian Early 1000 Barremian Hauterivian Valanginian 1100 Stenian Berriasian 146 Tithonian Early Cretaceous 1200 Late Kimmeridgian Oxfordian 161 Callovian Mesozoic 1300 Ectasian Bathonian Middle Bajocian Aalenian 176 1400 Toarcian Jurassic Mesoproterozoic Early Pliensbachian 1500 Sinemurian Hettangian Calymmian 200 Rhaetian 1600 Proterozoic Norian Late 1700 Statherian Carnian 228 1800 Ladinian Late Triassic Triassic Middle Anisian 1900 245 Olenekian Orosirian Early Induan Changhsingian 251 2000 Lopingian Wuchiapingian 260 Capitanian Guadalupian Wordian/Roadian 2100 271 Kungurian Paleoproterozoic Rhyacian Artinskian 2200 Permian Cisuralian Sakmarian Middle Permian 2300 Asselian 299 Late Gzhelian Kasimovian 2400 Siderian Middle Moscovian Penn- sylvanian Early Bashkirian -
2009 Geologic Time Scale Cenozoic Mesozoic Paleozoic Precambrian Magnetic Magnetic Bdy
2009 GEOLOGIC TIME SCALE CENOZOIC MESOZOIC PALEOZOIC PRECAMBRIAN MAGNETIC MAGNETIC BDY. AGE POLARITY PICKS AGE POLARITY PICKS AGE PICKS AGE . N PERIOD EPOCH AGE PERIOD EPOCH AGE PERIOD EPOCH AGE EON ERA PERIOD AGES (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) (Ma) HIST. HIST. ANOM. ANOM. (Ma) CHRON. CHRO HOLOCENE 65.5 1 C1 QUATER- 0.01 30 C30 542 CALABRIAN MAASTRICHTIAN NARY PLEISTOCENE 1.8 31 C31 251 2 C2 GELASIAN 70 CHANGHSINGIAN EDIACARAN 2.6 70.6 254 2A PIACENZIAN 32 C32 L 630 C2A 3.6 WUCHIAPINGIAN PLIOCENE 260 260 3 ZANCLEAN 33 CAMPANIAN CAPITANIAN 5 C3 5.3 266 750 NEOPRO- CRYOGENIAN 80 C33 M WORDIAN MESSINIAN LATE 268 TEROZOIC 3A C3A 83.5 ROADIAN 7.2 SANTONIAN 271 85.8 KUNGURIAN 850 4 276 C4 CONIACIAN 280 4A 89.3 ARTINSKIAN TONIAN C4A L TORTONIAN 90 284 TURONIAN PERMIAN 10 5 93.5 E 1000 1000 C5 SAKMARIAN 11.6 CENOMANIAN 297 99.6 ASSELIAN STENIAN SERRAVALLIAN 34 C34 299.0 5A 100 300 GZELIAN C5A 13.8 M KASIMOVIAN 304 1200 PENNSYL- 306 1250 15 5B LANGHIAN ALBIAN MOSCOVIAN MESOPRO- C5B VANIAN 312 ECTASIAN 5C 16.0 110 BASHKIRIAN TEROZOIC C5C 112 5D C5D MIOCENE 320 318 1400 5E C5E NEOGENE BURDIGALIAN SERPUKHOVIAN 326 6 C6 APTIAN 20 120 1500 CALYMMIAN E 20.4 6A C6A EARLY MISSIS- M0r 125 VISEAN 1600 6B C6B AQUITANIAN M1 340 SIPPIAN M3 BARREMIAN C6C 23.0 345 6C CRETACEOUS 130 M5 130 STATHERIAN CARBONIFEROUS TOURNAISIAN 7 C7 HAUTERIVIAN 1750 25 7A M10 C7A 136 359 8 C8 L CHATTIAN M12 VALANGINIAN 360 L 1800 140 M14 140 9 C9 M16 FAMENNIAN BERRIASIAN M18 PROTEROZOIC OROSIRIAN 10 C10 28.4 145.5 M20 2000 30 11 C11 TITHONIAN 374 PALEOPRO- 150 M22 2050 12 E RUPELIAN -
Alphabetical List
LIST E - GEOLOGIC AGE (STRATIGRAPHIC) TERMS - ALPHABETICAL LIST Age Unit Broader Term Age Unit Broader Term Aalenian Middle Jurassic Brunhes Chron upper Quaternary Acadian Cambrian Bull Lake Glaciation upper Quaternary Acheulian Paleolithic Bunter Lower Triassic Adelaidean Proterozoic Burdigalian lower Miocene Aeronian Llandovery Calabrian lower Pleistocene Aftonian lower Pleistocene Callovian Middle Jurassic Akchagylian upper Pliocene Calymmian Mesoproterozoic Albian Lower Cretaceous Cambrian Paleozoic Aldanian Lower Cambrian Campanian Upper Cretaceous Alexandrian Lower Silurian Capitanian Guadalupian Algonkian Proterozoic Caradocian Upper Ordovician Allerod upper Weichselian Carboniferous Paleozoic Altonian lower Miocene Carixian Lower Jurassic Ancylus Lake lower Holocene Carnian Upper Triassic Anglian Quaternary Carpentarian Paleoproterozoic Anisian Middle Triassic Castlecliffian Pleistocene Aphebian Paleoproterozoic Cayugan Upper Silurian Aptian Lower Cretaceous Cenomanian Upper Cretaceous Aquitanian lower Miocene *Cenozoic Aragonian Miocene Central Polish Glaciation Pleistocene Archean Precambrian Chadronian upper Eocene Arenigian Lower Ordovician Chalcolithic Cenozoic Argovian Upper Jurassic Champlainian Middle Ordovician Arikareean Tertiary Changhsingian Lopingian Ariyalur Stage Upper Cretaceous Chattian upper Oligocene Artinskian Cisuralian Chazyan Middle Ordovician Asbian Lower Carboniferous Chesterian Upper Mississippian Ashgillian Upper Ordovician Cimmerian Pliocene Asselian Cisuralian Cincinnatian Upper Ordovician Astian upper -
The Lochkovian-Pragian Boundary in the Lower Devo~Ian of the Barrandian Area (Czechoslovakia)
©Geol. Bundesanstalt, Wien; download unter www.geologie.ac.at Jb. Geol. B.-A. ISSN 0016-7800 Band 128 Heft 1 S.9-41 Wien, Mai 1985 The Lochkovian-Pragian Boundary in the Lower Devo~ian of the Barrandian Area (Czechoslovakia) By Ivo CHLUpAC, PAVEL LUKES, FLORENTIN PARIS & HANS PETER SCHÖNLAUB*) With 17 figures, 1 table and 4 plates Tschechoslowakei Barrandium Karnische Alpen Devon Stratigraphische Korrelation Lochkov-Prag-Grenze Tentaculiten Conodonten Graptolithen Chitinozoa Trilobita Brachiopoda Contents Summary, Zusammenfassung . .. 9 1. Introduction..... .. 9 2. Description of sections 10 2.1. Cerna rokle near Kosoi' 10 2.2. Trebotov - Solopysky 13 2.3. Praha - Velka Chuchle (Pi'fdol f) 14 2.4. Cikanka quarry near Praha-Slivenec 17 2.5. Radolfn Valley - Hvizaalka quarry 19 2.6. Oujezdce quarry near Suchomasty 22 3. Stratigraphic significance of some fossil groups in the Lochkovian-Pragian boundary beds of the Barrandian 22 3.1. Dacryoconarid tentaculites (P. LUKES) 22 3.2. Conodonts (H. P. SCHÖNLAUB) 24 3.3. Chitinozoans (F. PARIS) 27 3.4. Graptolites 28 3.5. Trilobites 28 3.6. Brachiopods 29 3.6. Some other groups 29 4. Proposal for a conodont based Lochkovian-Pragian boundary 30 5. Conclusion 30 References 32 Zusammenfassung Summary Im Barrandium Böhmens wurde die Lochkov/Prag-Grenze Six selected sections of the Lochkovian-Pragian boundary des Unterdevons an 6 ausgewählten Profilen in Hinblick auf ih- beds in the Barrandian area of central Bohemia were subject- ren Makro- und Mikrofossilinhalt biostratigraphisch untersucht. ed to investigations of mega- and microfossils. Joint occur- Für Korrelationszwecke sind in erster Linie Dacryoconariden, rence of different stratigraphically important fossil groups, par- Conodonten, Chitinozoen, Trilobiten, Graptolithen und Bra- ticularly dacryoconarid tentaculites, conodonts, chitinozoans, chiopoden geeignet.