PERSOONIA
Published by the Rijksherbarium, Leiden
Volume Part 245-265 10, 2, pp. (1979)
Type studies on entolomatoidspecies in the
Velenovský Herbarium—I.
in the and Species described genera Nolanea, Leptonia Telamonia.
M.E. Noordeloos
Rijksherbarium, Leiden
(With 29 Text-Figures)
All types of entolomatoid fungi available in the Velenovský Herbarium at Prague
have been studied. In this first report the types of 30 species described by Velenovský
in Nolanea, Leptonia and Telamonia (one species) are described. Species accepted
been transferred which resulted in 18 here have, if necessary, to Entoloma, new
combinations and 4 new names. One described in Nolanea had to be transferred to
Pluteus.
In the course ofhis long life JosefVelenovsky (1858-1949)described a great number of new taxa
in in various groups of fungi. Kotlaba (1977) in his revision of the polypores Velenovsky's
life work. herbarium gives a thorough survey of Velenovsky's and
Untillrecently only a minor part ofVelenovsky's new taxa in the Agaricales have been revised:
Harmaja(1970,1974)published on those in Clitocybe and Omphalina, Hornicek (1957) on those in Tricholoma in and Svrcek (1966) on those various small genera.
The work deals with in the and and present Velenovsky's species genera Nolanea Leptonia with one entolomatoid species erroneously described in Telamonia. A revision of his entolomatoid
in species described the genera Entoloma, Eccilia, Claudopus, Clitocybe and Arenicola is in preparation. All entolomatoid agarics are considered by the present author to belong to one single genus, viz. Entoloma (Fr.) Kumm. emend. Donk (syn. Rhodophyllus Quel.).
As to the Velenovsky's library was restricted works ofFries, Saccardo and Ricken, onecannot be surprised that many species described as new by Velenovsky appeared to be identical with species previously described in literature unknown to him. This was explicitely shown in
Kotlaba's revision of Velenovsky's polypores mentionedabove. In the entolomatoidfungi this phenomenon is not so manifestbecause this group of fungi is rather neglected in Europe, except in France, thanks to the excellend work of Romagnesi and Kiihner. Moreover Velenovsky's collecting sites are rather favourablefor species ofEntoloma. In some cases however itappeared
be for to necessary to create new names Velenovsky's taxa, as many combinations in Entoloma were preoccupied.
Velenovsky's types are depositedin the Herbarium ofthe National Museum, Prague, (PRM)
Instituteof Charles as exsiccata, and in the Botanical the University, Prague, (PRC) in liquid.
The latter collection contains most species described in Ceske Houby (1921).
245 246 P E R s o o N i A -Vol. 10, Part 2, 1979
In the presentpaper the infragenerictaxonomy of Romagnesi(1974,1978) is followed ingreat
lines. However, as 1 use the generic name Entoloma instead of Rhodophyllus some new com-
binations are necessary:
Entoloma subgenus Alboleptonia (Largent & Benedict) Noordeloos, comb. & stat. nov
(Basionym: Alboleptonia Largent & Benedict in Mycologia 62: 439. 1970.)
Entoloma sect. Papillati (Romagn.) Noordeloos, comb. nov. (Basionym: Rhodophyllus Quel,
sect. Papillati Romagn. in Bull. Soc. linn. Lyon 43: 330. 1974.)
Entoloma sect. Endochromonema (Largent & Thiers) Noordeloos, comb. nov. (Basionym:
Nolanea (Fr.) Kumm. sect. EndochromonemaLargent & Thiers in Northwest Sci. 46: 35. 1972.)
Velenovsky's species accepted here will be treated more extensively in another publication
(Noordeloos, 1980) and future papers on Entoloma by the present author.
METHODS AND PRESENTATION
and drawn in The microscopical structures of the exsiccata were observed, measured am-
moniac Red solution after in NH OH. The liquid preserved 1% Congo remoistening 10% 4
collections were observed in their own liquid, and usually also after staining with Congo Red.
Drawings were made with the aid of a camera lucida.
The magnifications of the figures are: spores, x 1000; all other microscopical details, x 670.
The following abbreviations are used:
Q.—Length-width ratio, usually given as follows: Q = 1.2 1.3-1.4, which means 'Q between 1.2
1.4 the and with an average of 1.3'. The size of spores relates to the largest length and width,
excluding the apiculus.
L-D. —Length minus width, usually given as follows: L-D = 1-2-3 pm, which means 'length
1 minus width between and 3 pm with an average of 2 /mi.'
ACKNOWLEDGEMENTS
Sincere thanks are due to Dr. Z. Urban, headof the Mycological Departmentof the Charles
University, Praha, for providing working facilities, and to Dr. J. Klan at the same institutefor his
assistance and Dr. Z. Dr. M. Svrcek the generous hospitality. Pouzar and at Botanical
ofthe National for Department Museum, Praha are greatly acknowledged fortheir help and the
loan of specimens. I am very grateful to Dr. C. Bas for critically reading the manuscript ofthis
paper.
Species described in Nolanea
alba. — Nolanea alba Velen., Ceske Houby: 629. 1921. — Neotype ( design. mihi): J. Vele-
novsky, july 1940, Mnichovice, collis Rlechac (PRM 154381). —Fig. 10. NOORDELOOS: Velenovsky'S Entoloma types-I 247
There is no original type materialpreserved in PRM or PRC. In PRM, however, there are under this name two later collections, gathered and named by J. Velenovsky, viz.:
PRM 154381 Mnichovice, collis 1940. This is the collection cited , Rlechac, July obviously by
Velenovsky in 1947 (: 78).
PRM 154481, Mnichovice, Vsësimy, in piceto vetere, 16 Sept. 1940. This collection contains three dark of pigmented specimens a species which is different from PRM 154381.
to the alba According protologueNolanea is a pale colouredspecies common in early summer in I deciduous forests. Considering this select collection PRM 154381 as a neotype for Nolanea alba Velen. The neotype collection contains one specimen in a relatively bad state, partly attacked by a mould, showing the following microscopical characters:
x Spores (9.0—)9.6—10.8(—11.3) 6.8-7.9 pm, Q= 1.2-1.35-1.4,L-D =2.4-3.5(-4.0) pm, 5- 6-
scattered angled in side-view. Basidia about 30-36 x 9-10 4-spored. Cheilocystidia (?; parts of gill-edge destroyed.) 40-55 x 7-9 / Clamp-connections numerous in hymenium, rare in trama. Nolaneaalba is close to N. mammosa Fr. sensu P. D. Orton (1960: 329) but differs in the lack of Moser and pigment. Entoloma cuneatum (Bres.) differsin the more pronouncedlypigmentedcap lack of Entoloma album Hesler is different in the cheilocystidia. quite many characters and belongs to subgenus Alboleptonia. Entoloma cinerascens Hesler is more robust and more typically a member of subgenus Entoloma. to Nolaneaalba seems represent a distinct species, which still has to be transferredto Entoloma. E. however is I Entoloma The name album already occupied. Therefore propose the new name Ceske nivescens Noordeloos, nom. nov. (Basionym Nolaneaalba Velen., Houby: 629. 1921; non Entoloma album Hesler, 1967.) — autumnalis. Nolanea autumnalis Velen. in Mykologia 6: 28. 1929.— No type material is left at PRM, nor at PRC. betulina. — Nolanea betulina Velen., Novitates mycologicae novissimae: 79. 1947. — Ho- lotypus: J. Velenovsky, 20 July 1943, Mnichovice (PRM 154455) — Fig. 17. The type consists of one specimen, partly damagedby a mould, with subglobose to broadly thin-walled 7.5-8.6 x 5.0-6.8 ellipsoidal, spores, pm. doubt this the Fr. The Without any specimen represents a species belongingto genus Pluteus following transfer is made: Pluteus betulinus (Velen.) Noordeloos, comb. nov. (Basionym: Nolanea betulina Velen., Novitates mycologicae novissimae: 79. 1947.) Ceske — crassipes. — Nolanea crassipes Velen, Houby: 627. 1921. Holotypus: O. Zverina, aprilii 1920, Hvezda, Pragae. (PRC; bottle 123a). — Fig. 20. 248 PE RSO ONI A-Vol. 10, Part 2, 1979 consists of well about 30 conico-convex. The type one preserved specimen. Cap mm broad, Lamellaefairly crowded, ventricose. Stipe about 70 x 5-6 (at apex), x 12 mm (at base). Spores = L D =2-3.2 in side- 10.3-12.4 x(7.5-)8.3-10.2 /mi, Q (1.2-) 1.2-1.25-1.35, /mi, 5-6-angled view. Basidia (30-)35-47 x 9-10.5 pm, Q= 3.6-4.4, 4-spored. Cystidia none. Hymenophoral colour- trama regular, composed of long, inflated elements, 230-435 x 12-21 /mi, thin-walled, 4— less. Pileipellis a thin, poorly differentiatedcutis of 10(—12) /ml wide, cylindrical hyphae. Pileitramaregular; hyphae composedofcylindrical to inflated, about 300-520x 20-32 /mi cells. Pigmentation inconspicious, probably diffusely intracellular; also some hyphal walls pale yellow; no encrusting pigments seen. Stipitepellis a differentiated cutis with loosely arranged, 4-10 wide with cylindrical, /mi hairs rounded, sometimes subcapitate tip. Clamp-connections present in hymenium. in Nolaneacrassipes is very close to Entoloma cuneatum (Bres.) Moser, which differs a typical yellow papilla on the pileus and a more or less glabrous stipe. Nolanea holoconiota Largent & Thiers differs in having cheilocystidia. As the name Entoloma crassipes already exists, the following new name has to be introduced: Entoloma lanuginosipes Noordeloos, nom. nov. Ceske (Basionym: Nolanea crassipes Velen., Houby: 627, 1921; non Entoloma crassipes Petch, 1924.) — depressa. — Nolanea depressa Velen., Novitates mycologicae: 148. 1939. Holotypus: J. Velenovsky, Aug. 1939, in formationeCalamagrostis Epigeios in colle silv. prope Miresovice (PRM 154453) — Fig. 3. with the The type consists of one partly damaged specimen, following microscopical charac- ters: Spores 9—10.7(—11.3) x 6.8-7.9 pm, Q= 1.15—1.3—1.45, L-D =2.4-3.5 /mi, rather pro- nouncedly 4-5-angled in side-view. Basidia 27-39x9-12 pm, 4-spored. Pileipellis a cutis of in narrow, slightly encrusted hyphae. Vascular hyphae present. Clamp-connections present hymenium. The convex-umbilicatepileus, the colouration, the cottony base of the stipe and, microscopi- of well with cally, the size and the shape the spores and the pigmentation agree perfectly Entoloma minutum (P. Karst.) Noordeloos, comb. nov. var. polymorphum (Romagn.) Noor- deloos. comb. nov. (Basionyms; Nolaneaminuta P. Karst, Hattsv. 1: 281,1879,and Rhodophyllus 19: 7. minutus (P. Karst.) Romagn. var. Polymorphus Romagn. in Rev. Mycol. 1954.) fracta. — Nolaneafracta Velen., Novitates mycologicae: 146. 1939. — Holotypus: J. Velen- ovsky, July 1936, Mnichovice, Struharov. (PRM 154483)—Fig. 1. The type consists of 6 well-preserved specimens with the following microscopical characters: L-D= in Spores 6.8-7.9 x(4.5-)5.7-6.7 pm, Q= 1.1—1.2—1.4, 1.2-1.8-2.4 /mi, (4-) 5-angled side-view. Basidia 25-32 x 7.6-10.2 pm, slenderly clavate, 4-spored. Cystidia none. Hymeno- phoral trama regular, cells cylindrical or inflated, up to 270 pm long and 10-24pm wide, hyaline, thin-walled, colourless. Pileipellis a cutis with transitions to a trichodermium, composed of inflated cylindrical to slightly hyphae, 7—17(—22) pm wide, with at centre of pileus tufts of clavate, terminal cells, 32-60 x 8-22 pm, with brown-encrusted walls and brown-olivaceous intracellular pigment granules. Pileitramaregular with cylindrical to slightly inflated brown- encrusted hyphae. Clamp-connections absent in all tissues studied. NOORDELOOS: Velenovsky's Entoloma types-I 249 — — 3. Nolanea — Fig. 1. Nolanea fracta, spores. Fig. 2. Nolanea tristis, spores. Fig. depressa, spores. Fig. — 4. Nolanea — 5. Nolanea 6. Nolanea 7. robiniae, spores. Fig. pusilla, spores. Fig. zonata, spores. — Fig. basidia. — 8. Nolanea — 9. Nolanea Leptonia omphaliaeformis spores and Fig. inodora, spores. Fig. and basidia. — 10. Nolanea and from globispora, spores Fig. alba, spores cheilocystidia. (All figs. type.) 250 PERSOON I A-Vol. 10, Part 2, 1979 Entoloma Nolaneafracta comes very close to fernandae (Romagn.) Noordeloos, comb. nov. (Basionym: Rhodophyllus fernandae Romagn. in Rev. Mycol. 1: 162. 1936.) The spores, pi- leipellis and pigmentation are similar. The colours, however, are much darker in N. fracta, especially the pileus which is 'atro-fusco, centro nigro', whereas E. fernandae has the pileus 'umbrino-griseo'. Also the shape ofthe pileus is different:N. fracta has a pileus which is 'acute- conico'; in E.fernandae the pileus is 'aplani, a peine mamelonne'. Entolomapsilopus Arnolds & Noordeloos (1979) has a glabrous, convex, only slightly umbonate moderately grey-brown, Therefore distinctly hygrophanous pileus. Nolaneafracta seems to represent a distinct species. the following transfer is made: Entolomafractum (Velen.) Noordeloos, comb. nov. (Basionym: Nolaneafracta Velen., Novitates mycologicae: 146. 1939.) galeraeformis. — Nolaneagaleraeformis Velen., Novitates mycologicae novissimae: 78. 1947. — No type collection is left at PRM or PRC. Ceske — globispora. — NolaneaglobisporaVelen., Houby: 628.1921. Lectotype ( design. mihi): J. Velenovsky, July 1919, Mnichovice (PRC bottle 48a).—Fig. 9. In Velenovsky's herbarium in PRC there are two collections that agree with the protologue: Mnichovice, July 1918 (bottle 400a) and July 1919 (bottle 48a). Bottle 48a contains two well preserved specimens and bottle 400a only a fragment of one specimen in a bad state, but of Nolanea conspecific with the first collection. Therefore I selected bottle 48a as the lectotype globispora. 0-1.2 Spores (8.5—)9.0—11.3 x 7.7—9.3(—10.3) pm, 0=1.0-1.1-1.2, L-D = pm, (4-)5-6- angled in side-view, isodiametrical- subglobularwith rounded angles. Basidia 32-44 x 8.2-13.5 = cells in pm, Q 3.0-3.8, 4-spored. Cystidia none. Hymenophoral trama regular with medios- thin tratum 200-350 x 11-35 /mi, colourless, inflated, hyaline and thin-walled.Pileipellis a and poorly differentiatedcutis of 3.5—7(—11) pm wide, cylindrical hyphae with brown, encrusted walls. Pileitramaregular with cylindrical to slightly inflatedhyphae with brown, encrusted walls. Clamp-connections frequent in hymenium, elsewhere rare or absent. The additionalcollections:bottle PRC 154377 all 400a at and no 154484,154380and atPRM, collected by J. Velenovsky in the vicinity of Mnichovice are all conspecific with the lectotype. Nolanea globispora is undoubtfully identical with Rhodophyllus juncinus Kiihn. & Romagn. discussion the of (See following description Nolanea zonata Velen., p. 255). hirta. — Nolanea hirta Velen. in Mykologia 6: 28. 1928. — Entoloma hirtum (Velen.) Noordeloos in Persoonia 10: 223 1979.— Holotype: J. Velenovsky, June 1926, Radotin (PRM 148449). The type consists of 5 well-preserved specimens with the following microscopical characters: x Spores (10.2—)10.7—13.0(—14.7) (6.3—)6.8—7.9(—9.0) pm, Q= 1.4-1.55-1.75, L-D =(3-)3.5- NOORDELOOS: Velenovsky's Entoloma types-I 251 4-5.7 brownish in 34-50 pm, irregularly nodulose-angular, slightly thick-walled, water. Basidia x 10.2-11.5 pm, 4-spored. Cheilocystidia 36-60x 16.5—24(—36) pm, slenderly to broadly cla- vate with brown-encrusted walls. Subhymeniumcellular, colourless. Hymenophoral trama with distinctly developed hymenopodium of2.5-7 pm wide, coarsely encrusted hyphae, and medios- tratum composed of cylindrical to slightly inflated cells, 70-220 x 8-16 /mi. Pileipellis a cutis with transitions to a trichodermiumwith repent or ascending long, multiseptate, attenuatehairs with heavily yellowish brown encrusted walls. Pigment encrusting all hyphaeof covering layers and trama. Clamp-connections absent. A redescription and discussion on the taxonomical position of E. hirtum is given in another paper (Noordeloos, I.e.). inodora. — Nolanea inodora Velen., Ceske Houby: 629. 1921. — Holotype: J. Velenovsky, Julio 1919, in collibus insolatis prope Myslin (PRC; bottle 120a).—Fig. 8. The consists of in bad the collectiondried and type two specimens a very state. Obviously out has been remoistened again, whichcaused considerable damage to the tissues. Spores 10.2-12.4 x 8.3-9.7 L-D= 1-2 cuboid. in such bad pm, Q= 1.1-1.15-1.25, /mi, irregularly Hymenium a state that no intact basidia could be found. Some clamp-connections were seen at the base of young basidiaand at several septa in the hymenophoraltrama. Pigment granular-intracellular in upper layers of pileus only. No incrustations seen. Nolanea inodora is rather well characterised by the irregular cuboid spores and the in- tracellular in pigments. The other species Europe with similar spores, viz. Entoloma rhombis- E. Horak differ in porum (Kiihn. & Bours.) Horak and prismatospermum (Romagn.) having encrusting pigments and considerably smaller spores. The extra-European Entoloma incertum Horak differs in (Romagn.) having a non-hygrophanous pileus. In my opinion Entoloma inodorum (Velen.) Noordeloos, comb. nov. (basionym: Nolanea inodora Velen., teské Houby: 629. 1921) is a distinct species which awaits rediscovery. majalis. — Nolanea majalis Velen., Novitates mycologicae: 147. 1939. — Holotype: J. PRM 11. Velenovsky, 12 may 1937, Mnichovice ( 154375).—Fig. The type consists of fragments oftwo specimens with the following microscopical characters: = L-D = 6- Spores (9—) 10.2—12.4 x (6.8-)7.4-7.9 /mi, Q (1.3-) 1.4-1.45-1.6, (2.4-)3-4.5 /mi, 35-46 9-1+.5 = angled in side-view. Basidia x pal, Q 3.7-4.2, 4-spored. Cystidia none. Hy- menophoral trama regular, composed of long, strongly inflated, thin-walled, colourless cells. Pileipellis a poorly differentiatedcutis of 4—10(—12) pm wide cylindrical hyphae, here and there with made a clavate terminalcell up to 18 pm wide. Pileitramaregular; hyphae up oflong, slightly In inflated cells up to 17.5 pm wide. Pigmentation indistinct: pale diffusely intracellular. pileipellis and upper layer of pileitrama sometimes also hyphal walls pale-coloured, but never encrusted. Clamp-connections frequent in hymenium, but extremely rare in other parts of the carpophore. Nolanea with Entoloma majalis agrees perfectly pallescens (P. Karst.) Noordeloos, comb. nov. (Basionym: Nolaneapallescens P. Karst. in Meddn Soc. Fauna Flora fenn. 16: 94. 1890 (Symb. will Myc. fenn. 29).) This species be discussed in another paper (Noordeloos, 1980). 252 P E R so o NI A —Vol. 10, Part 2, 1979 nitens. — Nolanea nitens Velen., Ceske Houby: 627. 1921. — Rhodophyllus nitens (Velen.) Kiihn & Romagn., Fl. anal.: 190. 1953. — No type-collection is left at PRM or PRC. of his of Romagnesi (1974b: 372) gives an excellent description interpretation Velenovsky's I species, which probably agrees with the original concept. Therefore introduce the name Ceské Entoloma nitens (Velen.) Noordeloos, comb. nov. (Basionym: Nolanea nitens Velen., Houby: 627. 1921.). Ceske — paludosa. — Nolaneapaludosa Velen., Houby: 628. 1921. No type-collection is left at PRM or PRC. perpusilla. — Nolaneaperpusilla Velen. in Mykologia 8: 22. 1931. — No type-collection is left at PRM or PRC. — pusilla. — Nolaneapusilla Velen.,Ceske Houby: 626. 1921. Holotypus: J. Velenovsky. July 1919, Mnichovice (PRC; bottle 85). —Fig. 5. of with the The type-collection consists one, well-preserved specimen following microscopical characters: Spores (9.3—) 10—12.4x9.3—11.2 /jm, Q= 1.0-1.1, L-D =0-1.5 //m, cruciform- stellate. Basidia 25-41 x 8-12/rm, 4-spored. Pileipellis a simple cutis of 4-11 pm wide, cylindric- al hyphae with pale diffusely intracellularpigment, sometimesin addition minutely encrusted. Clamp-connections none. charac- Nolanea pusilla represents a slender variety of Entoloma staurosporum (Bres.) Hesler, terized by its consistently slender nature and its habitat onrotten wood. Rhodophyllus xylophilus J. Lange (non Entoloma xylophilum Speg.) is identical. The following transfer is to be made: Entoloma staurosporum (Bres.) Hesler var. pusillum (Velen.) Noordeloos, comb. & stat. nov. (Basionym: Nolanea pusilla Velen., Ceske Houby: 626. 1921.) — robiniae. Nolanea robiniae Velen., Novitates mycologicae novissimae: 78. 1947. — Ho- lotypus: J. Velenovsky, July 1940, Mnichovice, in Robinieto (PRM 154379).—Fig. 4. The type consists of the pileus of one specimen, partly covered with a mould, with the following microscopical characters: Spores 9.0-10.2(—11) x 7.4-7.9 /im, Q= 1.2—1.3—1.4, L-D = 1.8-3.4 pm, (5-) 6-angled in side-view. Basidia 25-36 x 10-12 /im, clavate, 4-spored. Hy- menophoral trama regular, composedof cylindrical to slightly inflated, thin-walled,colourless cells, 180-220x6-14 pm. Pileipellis (difficult to interprète because of presence of mould) cutis of probably a thin, 3.5-8 /rm wide, cylindrical hyphae. Hypoderm strongly developed, 60-80 about /rm thick, composed of long chains of slightly inflated cells, 40-62 x 10-24 /im. Pileitrama under hypoderm regular; hyphae up to 15 pm wide, cylindrical. Pigment difficult to locate; hyphal wall uniformely coloured, not encrusted. Clamp-connections present in hy- menium, rare elsewhere. 253 NOORDELOOS: Velenovsky's Entoloma types-1 11. Nolanea — 12. basidia and — Fig. majalis, spores. Fig. Leptoniabrunnea, spores, cheilocystidia. Fig. 13. Leptonia conica, spores. —Fig. 14. Nolanea setulosa, spores and stipe-hairs. — Fig. 15. Leptoniaaurea, basidia and — 16. conica, basidia and spores, cheilocystidia. Fig. Leptonia spores, cheilocystidia. (All figs. from type.) P Part 1979 254 E R so oN I A -Vol. 10, 2, En- On account of its spores and its pigmentation Nolanea robiniae belongs to section dochromonema of subgenus Nolanea. Entoloma pallescens differs in slightly larger and more of stand for distinct elongate spores and the lack a hypoderm. Velenovsky's name seems to a species: Entolomarobiniae(Velen.) Noordeloos, comb. nov. (Basionym: NolanearobiniaeVelen., Novitates mycologicae novissimae: 78. 1974.) — setulosa. — Nolaneasetulosa Velen., Novitatis mycologicae: 147. 1939. Pouzarellasetulosa (Velen.) Mazzer in Bibltca. mycol. 46: 108. 1976. — Holotypus: J. Velenovsky, June 1931, PRM 14. Jidâsky prope Mnichovice ( 154485).—Fig. The type consists of two well-preserved specimens with the following microscopical charac- L-D = 4-4.5 ters: Spores 11—12.5(—14.7) x 6.8-7.9 pm, Q= 1.3-1.55-1.6(-l.8), pm, irregularly Basidia angular-gibbose, pale brown in water. 35-48 x 10-16 pm, broadly clavate, 4-spored, Cheilocystidia (difficult to find; only a few measured) 30-57 x 14-20 pan, slightly brown- of encrusted. Hymenophoraltrama with distinctly developed hymenopodium narrow, cylindric- al, coarsely encrusted hyphae and with mediostratum of inflatedhyphae with cells 55-110 x 12- 22 pm, brown andminutely encrusted. Pileipellis a cutis with transitionsto a trichodermiumwith repent or slightly ascending, long, attenuate,multiseptate, brown-encrusted hairs with cells 12- 25 pm wide (at base) to 6-14 pm wide (at apex). Stipitepellis a simple cutis with scatteredclavate with hairs in upper part of stipe and 2-3-celled, not encrusted, 8-20 pm wide hairs in lower part of stipe. Clamp-connections absent. Velenovsky stated in his diagnosis ofNolaneasetulosa that the spores are 15-18pm long. This mislead Mazzer (I.e.) in interpreting itas a distinct species. The present author places N. setulosa in the synonymy of Entoloma hirtum (see Noordeloos, 1979: 223). tristis. — Nolanea tristis Velen., Ceske Houby: 630. 1921. — Holotypus: J. Velenovsky, May 1920, Krc, Bohemia (PRC; bottle 370a).—Fig. 2. consists of in The type one specimen a very bad state. The hyphal wallsare partly desintegrated and the tissues are therefore difficultto study. The microscopical characters are: Spores 9.3-11.3 in Basidia x (6.2-)6.8-7.9 pm, Q= 1.25-1.35-1.55,6-7-angled side-view. 4-spored. pileipellis a cutis of radially arranged, thin-walled,6-11 pm wide, cylindrical hyphae, sometimesintermixed with fusoid, up to 15-18 pm wide terminalcells. Pileitramaregular, with long, inflated, fusiform cells measuring 250-310x27 Pigment abundant, dark brown, granular-intracellular in pileipellis and in broad adjacent zone in pileitrama. No encrusting pigment observed. Clamp- connections seen in hymenium. Nolanea tristis has to be placed in section Endochromonema of subgenus Nolanea. It differs from the other species in this section by its rather dark colour. It has some resemblance to Agaricuspiceus Kalchbr., but the latter species has a non-striatestipe which is concolorous with the pileus, and has a strong, cucumber-like or fish-like smell. (It is not unlikely that Kalchbrenner's species is identical with Macrocystidia cucumis (Pers. ex Fr.) Heim.) The following new combinationhas to be introduced: Entoloma tristis (Velen.) Noordeloos, comb, nov. (Basionym: Nolanea tristis Velen., Ceske Houby: 630. 1921.) NOORDELOOS: Velenovsky's Entoloma types-1 255 undulata. — Nolaneaundulata Velen., Ceske Houby: 627. 1921. —No type-collection is left at PRM of PRC. — variegata. — Nolanea variegata Velen., Ceske Houby: 630. 1921. No type-collection is left at PRM or PRC. zonata. — Nolanea zonata Velen.,Novitatesmycologicae novissimae: 78.1947. — Holotypus: J. Velenovsky, July 1940, Mnichovice, in palude inter gramine (PRM 154454).—Fig. 6. The The type consists oftwo specimens ina rather bad state, attacked by mouldsand by mites. characters 7.9-9x7.8-8.4 L-D = microscopical are: Spores fim, Q= 1.0-1.05, 0-0,5 /mi, 5- angled in side-view, subglobose-isodiametrical. Basidia 4-spored, clamped. Because of the bad state of the material other microscopical characters could not been chequed. convinced that this Judging from the microscopical characters and the protologue I am collection somewhat aberrant form of the rather Entoloma represents a common juncinum (Kiihn. & Romagn.) Noordeloos, comb. nov. (Basionym: Rhodophyllus juncinus Kiihn. & Romagn. in Rev. Mycol. 19: 5. 1954.) Synonyms: Nolanea globispora Velen., Ceske Houby: 628. 1921; non Entoloma globisporum — Morg.-Jones, 1971. Nolanea zonata Velen., Novitates mycologicae novissimae: 78. 1947; non Entoloma zonatum Hesler, 1967. Species described in Leptonia aurea. — Leptonia aurea Velen., Ceske Houby: 618. 1921. — Holotypus: J. Velenovsky, July 1918, Jevany (PRC; bottle 146b).—Fig. 15. The of type-collection consists one, well-preserved specimen with thefollowing microscopical characters: Spores (10.3—) 10.8—11.3(—11.8)x 7.2-8.2 pm, Q= 1.25-1.4-1.65, L-D = 2-3.2-4.5 in = pm, 6-angled side-view. Basidia 37-44 x 12.3-14.5 pm, Q 2.5-3, clavate, 4-spored. Cheilo- cystidia 65-84 x 12-22 x 3-7pm, lageniform, thin-walled,scattered, often in clusters. Pileipellis a trichodermium of broad, subcylindrical to clavate cells, 30-100 x 16-30 pm. Pileitrama regular, composed of short, broad elements. Pigment yellow-brown, intracellular, abundant in pileipellis and adjacent pileitrama. Clamp-connections present in hymenium. Leptonia aurea belongs to subgenus Alboleptonia and is identical with Entoloma kervernii (Gill.) Noordeloos, comb. nov. (Basionym: Leptonia kervernii Gill., Hymen. Fr.: 413. 1876.) brunnea. — Leptonia brunnea Velen., Ceske Houby: 618. 1921. — Holotypus: J. Velenovsky, coll. date & local, not indicated (PRC; bottle 320).—Fig. 12. 256 P E R so o N I A —Vol. 10, Part 2, 1979 The type-collection contains five well-preserved specimens with the following microscopical characters: Spores 11.2-12.8x8.2-8.7 //m, Q= 1.45—1.5—1.6, L-D = 3.7-4.1-4.7 nm, 5-6- in 12-18 angled side-view. Basidia 23-31 x 10.3-12.3 //m, 4-spored. Cheilocystidia 25-35 x /an, scattered, broadly-rounded clavate, thin-walled, colourless. Hymenophoral tramaregular; cells 49-87(-110)x 16-21 /an, cylindrical to inflated, thin-walled, colourless. Pileipellis a cutis of 10 cylindrical, up to /an wide hyphae with transitionsto a trichdermium,especially at centre of pileus, with cylindrico-clavate cells up to 27 fim broad. Pileitrama regular; cells cylindrical, 65- 140 (-200) x 8-16 /an. Pigment brown, granular-intracellular, abundant in pileipellis and adjacent pileitrama. Clamp-connections absent. Leptoniabrunnea is a true Leptonia close to Rhodophyllus asprellus Fr. sensu J. Lange,Quel., R. in Kiihn. & Romagn. The group of species around asprellus is not yet disentangled Europe. of different of with brown There are a lot interpretations Leptonia species a squamulose pileus For the time do brunnea Velen. and a blue-tinged stipe. being I not give Leptonia a new name, awaiting a revision of the R. asprellus-complex. (Velenovsky's name cannot be transferred to Entoloma because of the name Entoloma brunneum Petch, 1924). cinerascens. — Leptonia cinerascens Velen., Ceske Houby: 623. 1921. — Holotypus: J. Velenovsky, Julio 1919, in colle sicco graminis inter Strancice et Bozkov.' (PRC; bottle 48b).— Fig. 24. The type-collection consists ofone well-preserved specimen with the following microscopical 9-11.3x7.2-8.3 = L-D=1.7-3.2 in characters: Spores pm, Q 1.2—1.3—1.5, pm, 5-6-angled 34 = side-view. Basidia 45 x 10.3-13.4 pm, Q 2.7-3.7, clavate, 4-spored. Cystidia none. cells 320-360 15-26 Hymenophoral trama regular; x /mi, cylindrical to inflated, thin-walled, 4-9 inner colourless. Pileipellis a thin cutis of pm wide, cylindrical hyphae. Pileitrama regular, with 10-19 hyphae slightly inflated cells 270-350 x pm. Pigment minutely encrusting hyphae of pileipellis and pileitrama. Stipitepellis a cutis, at apex of stipe with clusters of cylindrical, sometimes subcapitate hairs, 4—7(— 10) pm wide. Clamp-connections present in hymenium at base of basidia. Leptonia cinerascens is both macro- and microscopically completely identical with Entoloma tenellum (Favre) Noordeloos, comb. nov. (Basionym: Rhodophyllus tenellusFavre, Assoc. fong. hauts-marais: 212. 1948.) Transfer ofVelenovsky's epithet to Entoloma wouldresult in a later homonym of E. cinerascens Hesler, 1967. Ceske cinerea. — Leptonia cinerea Velen., Houby: 623. 1921. — Holotypus: J. Velenovsky, Aug. 1919, Mnichovice (PRC; bottle 400b).—Fig. 23. The type consists of one specimen in very bad state; the base of the stipe is missing. The following microscopical characters have been observed: Spores 12.4-15x7.2-8.7 pm, irre- gularly nodulose-angular. Basidia clavate, 4-spored. (Most basidiaare damaged and impossible to 6-17 measure.) Hymenophoral trama regular; mediostratum regular; hyphae inflated, pm wide, minutely encrusted. Pileipellis a cutis with transitions to a trichodermiumof minutely encrusted, attenuate, multiseptate hairs. Pileitrama regular; hyphae brown-walled and minutely NOORDELOOS: Velenovsky's Entoloma lypes-l 257 — 17. Nolanea 18. — Fig. betulina, spores. Fig. Leptonia citrina, spores. Fig. 19. Leptonia involuta, spores. — Fig. 20. Nolanea crassipes, spores and basidia. — Fig. 21. Leptoniadensifolia, spores. —Fig. 22. Leptonia — mammillata,spores. Fig. 23. Leptonia cinerea, hairs ofpileus and spores. — Fig. 24. Leptonia cinerascens, spores and hairs of stipe-surface. (All figs. from type.) 258 P E R so o N I A — Vol. 10, Part 2, 1979 of hairs. encrusted. Stipitepellis a cutis cylindrical hyphae, with scarce, clavate, not encrusted Clamp-connections absent. from cinerea hirtum Judging the microscopical data Leptonia is very close to Entoloma (Velen.) But and Noordeloos. the spores seemto be slightly more elongate Velenovsky describes the stipe of of as being glabrous. Recombination Velenovsky's epithet would result in a later homonym Entoloma cinereum Hesler (1967). Because of the bad state of the type-collection I do not want to give Leptonia cinerea Velen. a new name before it has been rediscovered. citrina. — Leptonia citrina Velen.,Novitatesmycologicaenovissimae: 79.1947. —Holotypus: J. Velenovsky (leg. L. Hostânova), 10 Aug. 1940, Mnichovice (,PRM 152894).—Fig. 18. The type consists of threebadly dried specimens with the following microscopical characters: L-D = Spores 8.2-11.3 x (6.7—)7—8.2 pm, Q= 1.1-1.3-1.5, 0.5-2.3-3.7 pm, rather variable in shape. Basidia 4-spored (I was unableto find intact basidia). Cystidia not found.Hymenophoral trama regular, consisting of cylindrical cells, (60-) 100-240 x 20-35 pm. Covering layers dam- aged. Pigment probably intracellular, no trace of any encrusting pigment found. Clamp- connections not seen with certainty. The bad state of the material does not permit a certain identification. Judging from the diagnosis it could be a lemon yellow Alboleptonia close to Entoloma sericellum (Bull, ex Fr.) Kumm. Ceske conica. — Leptonia conica Velen., Houby: 623. 1921. — Lectotype (design, mihi): J Velenovsky, July 1919, Mnichovice (PRC; bottle 85). —Fig. 13, 16. Another collection inPRC, bottle 193a, is also labelledLeptonia conica Velen. Thatcollection was made at Libochovicky, a site quite a long distance from the type-locality Bozkov, situated quite near Mnichovice. Therefore collection 85 is selected here a lectotype. Spores 10.3-12.5 x 7.7-8.3 pm, Q= 1.2-1.4-1.6, L-D =2.4-5.3 pm, mostly 6-angled in side- 38-44 view. Basidia x 11-14pm, Q = 3.1-3.7, clavate, 4-spored. Cheilocystidia scattered, 55-90 x x and 11-20 2-5 pm, lageniform with moderately to distinctly swollen basal part long slender 250 neck, thin-walled. Hymenophoraltrama regular with cells up to (and more) pm long and 11- 27 wide pm wide, thin-walled, colourless. Pileipellis a thin cutis ofcylindrical 4-8 pm hyphae, loosely arranged, gradually passing into pileitrama. Pileitrama regular; hyphae cylindrical or slightly inflated, made up of thin-walled cells, 220-270 x 12-23 pm. Pigment exclusively in- tracellular, abundant in pileipellis and adjacent pileitrama. Clamp-connections frequent in hymenium, rare elsewhere"^ Leptonia conica belongs to subgenus Nolanea but it takes a rather isolated position in having lageniform cheilocystidia and intracellular pigment. As the combination Entoloma conicum is preoccupied, I introducein honour of J. Velenovsky for it the new name: Entoloma velenovskyi NOORDELOOS: Velenovsky's Entoloma types-l 259 Noordeloos Ceske non nom. nov. (Basionym: Leptonia conica Velen., Houby: 623, 1921; Entoloma conicum (Peck) Hesler 1967.) Noordeloos Entoloma velenovskyi var. longicystidiatum Arnolds & (1979: 296) differs in the and and the and more fusiform distinctly larger more irregularly shaped spores longer cheilocys- tidia. Ceske decurrens. — Leptonia decurrens Velen., Houby: 624. 1921. — No type-collection is left at PRM or PRC. — densifolia. — Leptonia densifolia Velen., Ceske Houby: 621. 1921. Holotypus: J. Velen- ovsky, July 1919, Myslin (PRC; bottle 54b).—Fig. 21. The type-collection contains onespecimen ina very bad state. Apparently the bottledried out and has been refilled afterwards, which made the trama 'glassy', very brittle and impossible to study. Spores (8.2-)8.7-10.3(-11.3)x7.2-7.7(-8.2) /mi, Q= 1.2-1.3-1.4, L-D= 1.7-2.0-3.7 with intracellular pm. Clamp-connections not seen with certainty. Some hyphae pigment. close It is impossible to identify this collection. It is, however, certainly a true Leptonia, to Entoloma griseocyaneum (Fr.) Kumm. sensu Kiihn. & Romagn. — Ceske involuta. Leptonia involuta Velen., Houby: 621. 1921. — Holotype: J. Velenovsky, July 1919, Mnichovice (PRC; bottle 54a).—Fig. 19. The type-collection contains two specimens in a caused of very bad state, by the process drying-out and remoistening. The hymenium is impossible to study. Spores 10—10.7(— 11.3) x L-D 2-4 7.2-7.9(-8.2) pm, Q= 1.25-1.42-1.7, = pm. 5-6-angled in side-view. Pileipellis a in trichodermiumof clavate cells up to 15 pm wide. Vascular hyphae seen trama ofpileus and hymenophore. Clamp-connections not seen. Leptonia involuta apparently belongs to subgenus Leptonia and is close to or identical with Entoloma ardosiacum (Fr.) Hesler ( =Rhodophyllus mougeotii (Fr.) Quel.) mammillata. - Leptonia mammillata Velen., Ceske Houby: 622. 1921. — Holotype: J. Velen- ovsky, July 1919, Mnichovice (PRC; bottle 54c). —Fig. 22. The type consists ofone partly damaged specimen, with the same brittle-glassy consistency as the typeofLeptonia densifolia has. Spores 10.8-13.4 x 7.2-8.7/tm, Q =(1.2—) 1.25-1.5-1.7, L-D =(2-)2.6-4-5.3 /im, rather irregular and assymetrical in side-view. Basidia 4-spored. Cystidia none. Hyphal walls in pileipellis, pileitrama and in hymenophoral trama brown and finely to coarsely encrusted. Clamp-connections frequent in hymenium, rare elsewhere. 260 PE R so oN i A —Vol. 10, Part 2, 1979 Leptonia mammillata is a form of Entoloma papillatum (Bres.) Hesler sensu Romagn. with will another slightly larger and more irregularly shaped spores. It be discussed in paper (Noordeloos, 1980). — minima. — Leptonia minima Velen., Novitates mycologicae novissimae: 79. 1947. Ho- lotype: J. Velenovsky, 6 Sept. 1940, Mnichovice, collis Plechâc, in verrimenti (Melampyrum) 153890 under the nivea —Fig. 27. (PRM , manuscript-name Leptonia Velen.). The type consists of a fragment of the pileus ofone specimen with the following microscopical = L-D = characters: Spores 9.3-10.7x (6.1-)6.6-7.7 pm,Q 1.2-1.35-1.4, 2-3.5 /rm, 5-6-angled in side-view. Basidia 30-40x9.5-15 clavate, 4-spored. Cystidia none. Pileipellis a tri- chodermium ofclavate, 7-12 fitn wide cells. Pigment not seen. Clamp-connections numerous in hymenium and trama. Leptonia minima belongs to subgenus Alboleptonia. It seemsto be a distinct species, for which the following transfer is made: Entolomaminimum (Velen.) Noordeloos, comb. nov. (Basionym: Leptonia minima Velen., Novitates mycologicae novissimae: 79. 1947.) Rhodophyllus omphaliformis Romagn. and Entoloma cephalotrichum (P. D. Orton) Noor- , , x ... v deloos, comb. nov. (basionym: Leptonia cephalotricha P. D. Orton in Trans. Br. mycol. Soc. 43: differ in 291, 1960), having considerably narrower spores. Entoloma olorinum (Romagn. & Favre) Noordeloos, comb. nov. (basionym: Rhodophyllus olorinus Romagn. & Favre in Rev. 3: in its Mycol. 75, 1938), differs omphaloid stature and more isodiametrical spores. Entoloma subsericellum Murr., as redescribed by Largent & Benedict (1970: 441-442), is possibly the closest related species, but differs in the far more robust stature. mycenoides. — Leptonia mycenoides Velen., Novitates mycologicae: 141. 1939. — Holotypus: J. Velenovsky, Oct. 1939, Mnichovice (PRM 153891).—Fig. 25. of The type-collection consists three badly dried specimens, covered by a mould, showing the following microscopical characters: Spores 8— 10.7(—11.2) x 6.7-7.2(-7.7) pm, Q= 1.1— 1.3—1.5, = in side-view. Basidia 0.5-2-3 pm, 5-6-angled 30-35x 8-11 pm, clavate, 4-spored. Cystidia not found.Hymenophoral trama regular, composedof cells measuring 100-120x 7-10 pm, hyaline, colourless. Pileipellis difficult to remoisten, damagedby mould; some cylindrico- clavate, up to 17 pm wide cells observed. Pigment none. Clamp-connections numerous in all tissues studied. Leptonia mycenoides belongsto subgenus Alboleptonia. It seems to be identicalwith Entoloma minimum Noordeloos. (Velen.) However, the edges of the lamellaeare now entirely overgrown with moulds. It is therefore possible that originally cheilocystidia were present. In that case L. mycenoides would represent small specimens of E. sericellum (Bull, ex Fr.) Kumm. nigrella. — Leptonia nigrella Velen., Ceske Houby: 623. 1921. — Holotypus: O. Zverina, July 1919, locis graminis prope Babice (PRC; bottle 120b).—Fig. 26. NOORDELOOS: Velenovsky's Entoloma types-I 261 Fig. 25. 26. Leptonia nigrella — Fig. 27. Leptoniaminima, Leptonia mycenoides, spores. — Fig. , spores. — 28. — 29. Telamonia from spores. Fig. Leptoniapapillata, spores. Fig. brevipes, spores. (All figs. type.) The type-collection consists of one partly damaged specimen. The tissues are impossible to reconstruct. The hymenial elements are also difficult to interprete. Pileus about 15 mm broad, but dark brown, glabrous. Lamellaeabout 30, emarginate, brown tingedpink. Stipe watery grey 10 = only the upper mm left. Spores 10—12.3 x 6.7-8.3 pm, Q= 1.3-1.5-1.7, L-D 3.2-4.3 /im, rather strongly asymmetrical-angular in side-view. Pigment in pileal surface layer brown, minutely encrusting. Clamp-connections observed in subhymenium. The type of spores and the pigmentation place this species in section Papillati of subgenus Nolanea. The macroscopical characters of the type-collection are contrary to Velenovsky's which of It is that the diagnosis suggests a species Pouzaromyces. very unlikely type-material really represents the species described by Velenovsky. Some error seems to be committed. consider Velen. dubium. Therefore I Leptonia nigrella a nomen omphaliaeformis. — Leptonia omphaliaeformis Velen., Novitates mycelogicae novissimae: 79. 1947. — Holotype: J. Velenovsky, 28 Sept. 1940, Mnichovice, Mencice, in colle (,Sarothamnus) (PRM 153893).—Fig. 7. The type consists of three specimens in bad condition, attacked by moulds. The following microscopical characters were observed: Spores (10.2—) 10.5—11.8(—12.4) x 7.2-8.2(-8.7) pm, Q = 1.3—1.45—1.5, L-D= 3-3.5-4 pm, 5-angled in side-view. Basidia 32-40x 10.5-12 pm, Q = 2.7-3.3, rather broadly clavate or cylindrical, 4-spored. Cystidia not found. Hymenophoral trama regular,consisting of cylindrical, sometimesinflated, hyaline, thin-walledcells, 40-100(- —Vol. Part 1979 262 PERSOONI A 10, 2, 150) x 12-17 /im. Pileipellis difficult to interprete, probably a cutis, but also some inflated with inflated elements seen. Pileitramaregular cylindrical to cells. Pigment diffuse, intracellular, pale brown in pileipellis and adjacent pileitrama; without a trace ofencrusting pigment. Clamp- connections absent. Vascular hyphae numerous in pileitrama. L. in The stature, pigmentation, clampless hyphaeand simple spores place omphaliaeformis subgenus Eccilia (Fr.) emend. Romagn. 1978. Entoloma parkensis (Fr.) Noordeloos, comb. nov. (basionym: Agaricusparkensis Fr. inKongl. Vitensk. Akad.Fôrh. 18:45. 1851)and E. leptonipes (Kiihn. & Romagn.) Moser both differ macro- and microscopically. Therefore the following recombinationis made:Entoloma omphaliaeformis (Velen.) Noordeloos, comb. nov. (Basionym: Leptonia omphaliaeformis Velen., Novitatis mycologicae novissimae: 79. 1949.) papillata. — Leptoniapapillata Velen., Ceske Houby: 622. 1921.— Holotypus: J. Velenovsky, July 1919, Mnichovice (PRC; bottle 85a).—Fig. 28. The type consists of one specimen in relatively good state with the following microscopy: in Spores 9.3—10.7(—11.3) x 7.2-8.3 pm, Q= 1.2-1.3-1.4, L-D= 1.7-3.2 pm, 5-6-angled side- view. Basidia 33-44x7.2-12.4 pm, Q =3-4.6(-5.7), 4-, rarely 2-spored. Cystidia none. Hy- menophoral trama regular; hyphae cylindrical to inflated, composed of 270-500 x 20-30 pm mixed with differentiated of cells, 4-7 pmwide, cylindrical hyphae. Pileipellis a poorly cutis 6-10 pm wide, cylindrical hyphae, gradually passing intopileitrama. Pileitrama regular,consisting of to cylindrical inflated up to 20 pm wide hyphae, intermixed with very narrow (1.5-4 pm wide) hyphae. Pigment membranal and encrusting in hyphae of pileipellis and pileitrama and in narrow hyphae of the hymenophoral trama. Clamp-connections abundant in hymenium. Leptonia papillata is identical with Entolomapapillatum (Bres. 1887) Hesler. (see also Noor- deloos, 1980). — Ceske xanthopa. Leptonia xanthopa Velen., Houby: 622. 1921. —No type-collection is left in PRM or PRC. Kiihn. & 1953: Romagnesi (in Romagn., 205) introduced Rhodophyllus poliopus as a new name for Leptonia xanthopa Velen. He called attention to the very unsuitable name given by ' ' Velenovsky: xanthopa means with 'yellow stipe'. In the diagnosis the stipe is described 'chalybaeo-coerulescenti vel plumbeo-griseo'. Later Romagnesi described R. poliopus as a new Kiihn. & 1954: As is left, consider Velen. species (in Romagn. 8). no type I Leptonia xanthopa a nomen dubium. As it will never be certain whether L. xanthopa is identical with R. poliopus it Romagn. or not, seems to be better to use the lattername for the typical Leptonia, excellently described Therefore the combinationis made: by Romagnesi. following new Entolomapoliopus (Romagn.) Noordeloos, comb. nov. (Basionym: Rhodophylluspoliopus Romagn. in Rev. Mycol. 19: 8. 1954.) NOORDELOOS: Velenovsky's Entoloma types-I 263 A species described in Telamonia brevipes. — Telamonia brevipes Velen., teske Houby: 458. 1921. — Holotype: J. Velenovsky, Aug. 1915, Mnichovice (PRC; bottle 183).-—Fig. 29. ' The label of the type-collection bears the following note: Leptonia brevipes Velen., Mni- 8. 458 chovice, 1915, p. sub Telamonia' The type-collection contains five well preserved specimens. Spores 8.2—9.7(—10.3) x (7.7-)8.2-8.7(-9.3) pm,Q = 1.0-1.05-1.1(-1.15), L-D =0- 0.5-1 /zm, isodiametrical-subglobose, 5(-6)-angled in side-view. Basidia 35-42 x 11-15.5/zm, Q = clavate 2.5-3.2, 4-spored. Cystidia none. Hymenophoral trama composed of cylindrical to elements, 160-310x8-24 /zm, thin-walled, colourless. Pileipellis a thin, hardly differentiated cutis of 2-6 fim wide, cylindrical hyphae, gradually passing into trama. Pileitrama regular, 20 with 2-5 composed of cylindrical to inflated, up to /zm wide hyphae intermixed /zm wide, narrow-cylindrical hyphae. Hyphal walls, brown in pileipellis and adjacent pileitrama, also encrusted in the narrow hyphae of pileipellis and pileitrama. Stipitepellis a simple cutis of parallel, 3.5-7 /zm wide, thin-walled, yellow-brown, not encrusted hyphae. Vascular hyphae numerous in pilei- and stipitetrama. Clamp-connections frequent in hymenium, rare in trama. Telamonia brevipes differs from all members of subgenus Nolanea section Papillati with isodiametrical spores in the dark pigmented, non-striatepileus and the habitat, viz. coniferous forest. Therefore introduce the new name Entoloma Noordeloos, 1 pseudotelamonia nom. nov. (Basionym: Telamoniabrevipes Velen.,Ceske Houby: 458,1921; non Entolomabrevipes Murrill, 1917.) Index Agaricus Entoloma (cont.) parkensis Fr. 262 juncinum (Kiihn. & Romagn.) Noordeloos 205 piceus Kalchbr. 254 kervernii (Gillet) Noordeloos 205 Entoloma lanuginosipesNoordeloos 248 album Hesler 247 leptonipes (Kiihn & Romagn.) Moser 262 ardosiacum (Fr.) Hesler 259 minimum (Velen.) Noordeloos 260 brevipes Murrill 263 minutum (P. Karst.) Noordeloos 248 Petch brunneum 256 var. polymorphum (Romagn.) Noordeloos248 cephalotrichum (P. D. Orton) Noordeloos 260 nitens (Velen.) Noordeloos 252 cinerascens Hesler 256 nivescens Noordeloos 247 conicum (Peck) Hesler 258 olorinum (Romagn. & Favre) Noordeloos 260 crassipes Petch 248 omphaliaeformis (Velen.) Noordeloos 262 248 cuneatum (Bres.) Moser 247, pallescens (P. Karst.) Noordeloos 251, 254 fernandae (Romagn.) Noordeloos 250 papillatum (Bres.) Hesler 260, 262 250 fractum (Velen.) Noordeloos parkensis (Fr.) Noordeloos 262 globisporum Morg.-Jones 205 poliopus (Romagn.) Noordeloos 262 griseocyaneum (Fr.) Kumm. 259 prismatospermum (Romagn.) Horak 251 hirtum (Velen.) Noordeloos 250, 254, 258 pseudotelamonia Noordeloos 263 incertum (Romagn.) Horak 251 psilopus Arnolds & Noordeloos 250 inodorum (Velen.) Noordeloos 251 pusillum, (var. of staurosporum) 252 264 PERSOONI A—Vol. 10, Part 2, 1979 Entoloma (cont.) Nolanea (cont.) rhombisporum (Kiihn. & Boursier) Horak 251 betulina Velen. 247 robiniae (Velen.) Noordeloos 254 crassipes Velen. 247 Kumm. Velen. 248 sericellum (Bull, ex Fr.) 258, 260 depressa fracta 248 staurosporum (Bres.) Hesler 252 Velen. Velen. 250 var. pusillum (Velen.) Noordeloos 252 galeraeformis subsericellum Murrill 260 globispora Velen. 250, 255 tenellum (Favre) Noordeloos 256 hirta Velen. 250 holoconiota tristis (Velen.) Noordeloos 254 Largent & Thiers 248 velenovskyi Noordeloos 259 inodora Velen. 251 majalis Velei 251 var. longicystidiatum nitens Velen. Arnolds & Noordeloos 259 252 P. xylophilum Speg. 252 pallescens Karst. 251 Velen. zonatum Hesler 255 paludosa 252 Leptonia perpusilla Velen. 252 Velen. aurea Velen. 255 pusilla 252 brevipes Velen. 263 robiniae Velen. 252 setulosa Velen. brunnea Velen. 255 254 cephalotricha P. D. Orton 260 tristis Velen. 254 cinerascens Velen. 256 undulata Velen. 255 Velen. 255 cinerea Velen. 256 variegata citrina Velen. 258 zonata Velen. 255 conica Velen. 258 Pluteus decurrens Velen. 259 bctulinus (Velen.) Noordeloos 247 densifolia Velen. 259 Rhodophyllus involuta Velen. 259 asprellus (Fr.) J. Lange 256 mammillata Velen. 259 fernandae Romagn. 250 minima Velen. 260 juncinus Kiihn. & Romagn. 250, 255 mycenoides Velen. 260 minutus (P. Karst.) Romagn. 248 nigrella Velen. 260 var. polymorphus Romagn. 248 omphaliaeformis Velen. 261 mougeotii (Fr.) Quel. 259 papillata Velen. 262 olorinus Romagn. & Favre 260 xanthopa Velen. 262 omphaliformis Romagn. 260 Nolanea poliopus Romagn. 262 alba Velen. 246 tenellus J. 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