Russian Entomol. J. 15(4): 409–417 © RUSSIAN ENTOMOLOGICAL JOURNAL, 2006

To the knowledge of Chrysomelidae (Coleoptera) described by V.Motschulsky

Ê ïîçíàíèþ æóêîâ ëèñòîåäîâ (Coleoptera: Chrysomelidae), îïèñàííûõ Â.Ìî÷óëüñêèì

L.N. Medvedev Ë.Í. Ìåäâåäåâ

Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospect 33, Moscow 119071, Russia. Институт проблем экологии и эволюции РАН, Ленинский проспект 33, Москва 117071, Россия. KEY WORDS: Chrysomelidae, types, collection of V. Motschulsky, new synonyms, new combinations, new names, status ressurect. КЛЮЧЕВЫЕ СЛОВА: Chrysomelidae, типы, коллекция В. Мочульского, новые синонимы, новые комбинации, новые замещающие названия, восстановленный статус.

ABSTRACT. 41 poorly known type species of leaf chulsky, 1860), comb. n. = C. paviei Lefevre, 1890, syn. described by V. Motschulsky were studied. n.; Pseudoides coeruleipennis (Motschulsky, 1860), Transferred to other : Orsodacne indica Mots- comb. n. = P. flavicollis Jacoby , 1903, syn. n.; Atra- chulsky, 1866 to Hyphaenia Baly, 1865; Melixanthus chya bimaculata Hornstedt, 1788 = A. maculicollis flaveolus (Motschulsky, 1866), M. acutungulus (Mots- (Motschulsky, 1858), comb. et syn. n.; Chaetocnema chulsky, 1866), M. nigrolimbatus (Motschulsky, 1866), (Tlanoma) gracilis Motschulsky, 1859 = Ch. indica M. suturalis(Motschulsky, 1866) — to Coenobius Suf- Weise, 1916, syn. n.; Chaetocnema (Tlanoma) puncti- frian, 1857; Scelodonta aenea Motschulsky, 1866 — collis Motschulsky, 1859 = Ch. discreta Baly, 1876, to Lefevre, 1884; rugipennis syn. n.; suturellus (Motschulsky, 1866) = Motschulsky, 1860 — to Laporte, 1833; L. rangoonensis Jacoby, 1892, syn. n.; Aphthona al- Trichochrysea rufula Motschulsky, 1866, T. fuscula bescens (Motschulsky, 1866), comb. n. = A. opaca Motschulsky, 1866 — to Aoria Baly, 1860; Monolepta Allard, 1889, syn. n.; Nisotra gemella Erichson, 1834 = maculicollis Motschulsky, 1858 — to Atrachya De- N. javana (Motschulsky, 1866), syn. n.; Dactylispa jean, 1837; ; Atrachya nigrocincta Motschulsky, 1858, longicornis (Motschulsky, 1861) = D. severini Gestro, A. basalis Motschulsky, 1858 — to Monolepta Erich- 1897, syn. n.; Dactylispa nigromaculata (Motschul- son, 1843; Longitarsus albescens Motschulsky, 1866 — sky, 1861) = D. xanthospila Gestro, 1890, syn. n.; to Aphthona Chevrolat, 1837; Luperus coeruleipennis Dactylispa nigripennis (Motschulsky, 1861) = D. xan- Motschulsky, 1860 — to Pseudoides Jacoby, 1892, thospila Gestro, 1890, syn. n.; Gonophora zinzibaris Anisodera nigricauda Motschulsky, 1863 — to Gono- (Motschulsky, 1861) = G. akalankita Maulik, 1919, phora Baly, 1858; Tricliona oblonga (Motschulsky, syn. n.; Gonophora nigricauda (Motschulsky, 1863) 1866) removed back to Lefevre, 1885. New = G. taprobane Gestro, 1902, syn. n. Dactylispa fili- synonymy are established: Coenobius flaveolus cornis (Motschulsky, 1861), stat. rest. is a good spe- (Motschulsky, 1866), comb. n. = C. acutangulus (Mots- cies. Renamed because of homonymy: Monolepta ni- chulsky, 1866), comb. et syn. n. = C. nigrolimbatus grocincta Jacoby, 1900 to Monolepta jacobyana (Motschulsky, 1866), comb. et syn. n. = C. suturalis Medvedev, 2006, nom. n.; Monolepta basalis Harold, (Motschulsky, 1866), comb. et syn. n.; Nodina subdi- 1880 to Monolepta haroldiana Medvedev, 2006, nom. latata Motschulsky, 1858 = N. rotundata Motschul- n. Redescription of 1 genus and 16 species are given. sky, 1858, syn. n.; Basilepta viridipenne (Motschul- For 55 species with clear taxonomical status lectotypes sky, 1860) = B. frontalis (Baly 1867), syn. n.; Basilepta and paralectotypes are designated. sculpturata (Motschulsky, 1860) = B. bhamoense (Ja- coby, 1892), syn. n.; Pagria restituens Walker, 1859 = РЕЗЮМЕ. Изучен 41 тип малоизвестных видов P. aenea (Motschulsky, 1866), comb. et syn. n. = P. жуков листоедов, описанных В. Мочульским. Пере- costatipennis Jacoby, 1887, syn. n.; Scelodonta dillw- несены в другой род: Orsodacne indica Motschulsky, yni Stephens, 1831 = S. strigicollis (Motschulsky, 1866 в Hyphaenia Baly 1865; Melixanthus flaveolus 1866), syn. n.; Aoria nigripes Baly, 1860 = A. rufula (Motschulsky, 1866), M. acutungulus (Motschulsky, (Motschulsky, 1866), comb. et syn. n. = A. fuscula 1866), M. nigrolimbatus (Motschulsky, 1866), M. (Motschulsky, 1866), comb. et syn. n.; lat- suturalis(Motschulsky, 1866) — в Coenobius Suffrian eralis (Motschulsky, 1866), comb. n. = C. variabilis 1857; Scelodonta aenea Motschulsky, 1866 — в Pagria Baly, 1874, syn. n.; Colaspoides rugipennis (Mots- Lefevre, 1884; Colasposoma rugipennis Motschulsky, 410 L.N. Medvedev

1860 — в Colaspoides Laporte, 1833; Trichochrysea Introduction rufula Motschulsky, 1866, T. fuscula Motschulsky, 1866 — в Aoria Baly, 1860; Monolepta maculicollis Motschulsky’s collection, deposited in the Zoolog- Motschulsky, 1858 — в Atrachya Dejean, 1837; Atrachya ical Museum of the Moscow State University, includes nigrocincta Motschulsky, 1858, A. basalis a lot of unclear Chrysomelid species, which were never Motschulsky, 1858 — в Monolepta Erichson, 1843; throughoutly revised, except one Ogloblin’s publica- Longitarsus albescens Motschulsky, 1866 — в tion [1930] concerning rather large amount of Alticinae. Aphthona Chevrolat, 1837; Luperus coeruleipennis In the publication proposed I have studied 41 type Motschulsky, 1860 — в Pseudoides Jacoby, 1892, species from this collection, partly with redescriptions, Anisodera nigricauda Motschulsky, 1863 — в partly with synonymization and taxonomical notes and Gonophora Baly, 1858; Tricliona oblonga (Motschul- also designated lectotypes and paralectotypes for spe- sky, 1866) перенесена обратно в Basilepta Lefevre, cies with clear taxonomical status. 1885. Установлены новые синонимы: Coenobius Bad safety of Motschulsky collection not always flaveolus (Motschulsky, 1866), comb. n. = C. acut- allows to determine a sex of some specimens, therefore angulus (Motschulsky, 1866), comb. et syn. n. = C. nigro- it is specified not for all types. limbatus (Motschulsky, 1866), comb. et syn. n. = C.suturalis (Motschulsky, 1866), comb. et syn. n.; Nodina subdilatata Motschulsky, 1858 = N. rotundata Taxonomical part Motschulsky, 1858, syn. n.; Basilepta viridipenne (Motschulsky, 1860) = B. frontalis (Baly 1867), syn. n.; Coenobius flaveolus (Motschulsky, 1866), comb. n. Basilepta sculpturata (Motschulsky, 1860) = B.bhamo- = Coenobius acutangulus (Motschulsky, 1866), comb. et syn. n. ense (Jacoby, 1892), syn. n.; Pagria restituens Walker, = Coenobius nigrolimbatus (Motschulsky, 1866), comb. et syn. n. 1859 = P. aenea (Motschulsky, 1866), comb. et syn. n. = = Coenobius suturalis (Motschulsky, 1866), comb. et syn. n. P. costatipennis Jacoby, 1887, syn. n.; Scelodonta = Coenobius basalis (Motschulsky, 1866), comb. n. TYPE MATERIAL. Lectotype of Monachus flaveolus Mots- dillwyni Stephens, 1831 = S. strigicollis (Motschulsky, chulsky, 1866 (designated here), “Ceylon”. 3 paralectotypes 1866), syn. n.; Aoria nigripes Baly, 1860 = A. rufula with same label as lectotype; (Motschulsky, 1866), comb. et syn. n. = A. fuscula Lectotype of Monachus acutangulus Motschulsky, 1866 (Motschulsky, 1866), comb. et syn. n.; Cleoporus (designated here), “Ceylon”; Lectotype of Monachus nigrolimbatus Motschulsky, 1866 lateralis (Motschulsky, 1866), comb. n. = C. variabilis (designated here), “Ceylon”. 2 paralectotypes with same label Baly, 1874, syn. n.; Colaspoides rugipennis as lectotype; (Motschulsky, 1860), comb. n. = C. paviei Lefevre, 1890, Lectotype of Monachus suturalis Motschulsky, 1866 (designat- syn. n.; Pseudoides coeruleipennis (Motschulsky, 1860), ed here), “Ceylon”. 4 paralectotypes with same label as lectotype. comb. n. = P. flavicollis Jacoby , 1903, syn. n.; Atrachya NOTES. We have studied 4 Motschulsky’s species placed in the genus Melixanthus [Clavareau, 1913] and found that bimaculata Hornstedt, 1788 = A. maculicollis they are identical morphologically and belong to the genus (Motschulsky, 1858), comb. et syn. n.; Chaetocnema Coenobius Suffrian 1857. Monachus basalis Motschulsky, (Tlanoma) gracilis Motschulsky, 1859 = Ch. indica 1866 was already synonymized with M. flaveolus as colour Weise, 1916, syn. n.; Chaetocnema (Tlanoma) variation [Clavareau, 1913]. We have not seen the type of puncticollis Motschulsky, 1859 = Ch. discreta Baly, M.basalis, but it seems to be correct. A short description of 1876, syn. n.; Longitarsus suturellus (Motschulsky, the species is given below. 1866) = L. rangoonensis Jacoby, 1892, syn. n.; Aphthona REDESCRIPTION. Colour very variable. Body entirely albescens (Motschulsky, 1866), comb. n. = A. opaca fulvous (typical C. flaveolus), or elytra darkened along basal, Allard, 1889, syn. n.; Nisotra gemella Erichson, 1834 = lateral and sutural margin (C. suturalis), and might be rather N. javana (Motschulsky, 1866), syn. n.; Dactylispa broad and almost reaching apex (C. nigrolimbatus) or elytra longicornis (Motschulsky, 1861) = D. severini Gestro, dark brown with apex often pale (C. acutangulus); underside sometimes dark brown, legs always fulvous. 1897, syn. n.; Dactylispa nigromaculata (Motschulsky, Body short ovate. Head distinctly punctuate, eyes large, 1861) = D. xanthospila Gestro, 1890, syn. n.; Dactylispa contiguous along all their length in both sexes. Antennae nigripennis (Motschulsky, 1861) = D. xanthospila Gestro, short, segments 3–6 small and subequal, next segments clear- 1890, syn.n.; Gonophora zinzibaris (Motschulsky, ly widened. Prothorax short, strongly narrowed anteriorly, 1861) = G. akalankita Maulik, 1919, syn.n.; Gonophora with side margins almost straight, basal margin feebly arcu- nigricauda (Motschulsky, 1863) = G. taprobane Gestro, ate, with basal lobe very feeble, broadly rounded. Surface 1902, syn. n. Dactylispa filicornis (Motschulsky, 1861), with distinct collar on anterior margin, strongly punctate, stat. rest. является самостоятельным видом. Пере- without any depressions, basal row of punctures not very именованы в связи с омонимией: Monolepta regular in middle. Scutellum lanceolate, impunctate, twice as nigrocincta Jacoby, 1900 в Monolepta jacobyana long as wide. Elytra regularly punctate striate throughout, interspaces feebly convex and with a row of very fine punc- Medvedev, 2006, nom. n.; Monolepta basalis Harold, tures. Prosternum more than twice as wide as long, with a 1880 в Monolepta haroldiana Medvedev, 2006, nom. n. few strong punctures. Length 1.8–2 mm. Сделаны переописания 1 рода и 16 видов. Для 55 DIAGNOSIS. The species is near C. basalis Jacoby, видов с ясным таксономическим статусом обозна- 1908 from South India and C. seminigris Jacoby, 1908 from чены лектотипы и паралектотипы. Tenasserim, but these both have prothorax impunctate, with distinct impressions. To the knowledge of Chrysomelidae (Coleoptera) described by V.Motschulsky 411

4

3 1 2 6 5

11

7 12 8 9 10

Figs 1–12. Details of Chrysomelidae spp. from collection of V. Motschulsky: 1–10 — aedeagus; 11 — anterior margin of clypeus; 12 — mandible; 1, 11–12 — Nodina pusilla; 2 — N. laevicollis; 3 — N. subdilatata; 4–5 — Bathseba ferruginea; 6 — Pseudoides coeruleipennis; 7–8 — Monolepta nigrocincta; 9–10 — M. basalis; 1–3, 4, 6, 9 — dorsal view; 7 — ventral view; 5, 8, 10 — lateral view. Ðèñ. 1–12. Äåòàëè ñòðîåíèÿ Chrysomelidae spp. èç êîëëåêöèè Â. Ìî÷óëüñêîãî: 1–7 — ýäåàãóñ; 8 — ïåðåäíèé êðàé íàëè÷íèêà; 9 — ìàíäèáóëà; 1, 11-12 — Nodina pusilla; 2 — N. laevicollis; 3 — N. subdilatata; 4–5 — Bathseba ferruginea; 6 — Pseudoides coeruleipennis; 7–8 — Monolepta nigrocincta; 9–10 — M. basalis; 1–3, 4, 6, 9 — ñâåðõó; 7 — ñíèçó; 5, 8, 10 — ñáîêó.

Genus Nodina Motschulsky, 1858 middle, than widened again to apex, with long and acute Motschulsky described 4 species, including the type of apical process (Fig. 1). Length — 1.9 mm. genus, N. pusilla. We give below additional data for each of Female: Anterior margin of clypeus feebly concave, these species. They all are metallic bronze with fulvous legs and very alike, differing mostly with sexual characters and mandibles without basal lobe, elytra with very feeble and short ridge behind humerus. inhabit in Burma. Length — 2.1 mm. Nodina pusilla Motschulsky, 1858 DIAGNOSIS. N. rufipes Jacoby, 1908 seems to be near Figs 1, 11–12 to this species. TYPE MATERIAL. Lectotype of Nodina pusilla Motschul- Nodina laevicollis Motschulsky, 1858 sky, 1858 (designated here), (#), “Ind. or.”. Paralectotype, ($) Fig. 2 with same label as lectotype. TYPE MATERIAL. Lectotype of Nodina laevicollis Mots- REDESCRIPTION. Male: Antennae fulvous with slightly chulsky, 1858 (designated here), (#), “Ind. or.”. darkened apical segments, labrum fulvous, mandibles fulvous REDESCRIPTION. Antennae and labrum fulvous, mandi- with black base. Anterior margin of clypeus biemarginate, bles piceous. Anterior margin of clypeus subtriangularly emar- with tooth on each side (Fig. 11). Mandibles large, flat, with ginated, mandibles simple. Elytra without lateral ridge, punc- strongly elevated basal lobe (Fig. 12). Elytra without lateral tured rows obliterated on sides and behind middle. Aedeagus ridges, rows of punctures very feeble laterally and in apical gradually narrowed apically, with extreme apex obtuse (Fig. 2). part, but not obliterated. Aedeagus slightly narrowed before Length — 2.2 mm. 412 L.N. Medvedev

Nodina subdilatata Motschulsky, 1858 Basilepta triangularis (Motschulsky, 1866) Fig. 3 TYPE MATERIAL. Lectotype of Nodostoma triangularis = Nodina rotundata Motschulsky, 1858, syn. n. Motschulsky, 1866 (designated here), “Ceylon”. Paralectotype, TYPE MATERIAL. Lectotype of Nodina subdilatata Mots- with same label as lectotype. chulsky, 1858 (designated here), (#), “Ind. or.”. 7 paralecto- REDESCRIPTION. Fulvous, 2 spots on prothorax, a types with same label as lectotype. spot on basal convexity of elytron and another in postbasal Lectotype of Nodina rotundata Motschulsky, 1858 (desig- groove, suture narrowly and lateral margin in anterior two nated here), (#), “Ind. or.”. 2 paralectotypes with same label thirds black. as lectotype In paralectotype lateral margin of elytra entirely fulvous, NOTES. Nodina rotundata Motschulsky, 1858 is a syn- suture indistinctly black. onym of N. subdilatata Motschulsky, 1858, being fully iden- Body elongate, almost parallel-sided. Head finely punc- tical, including structure of aedeagus. tuate, clypeus not separated from frons, delimited on sides REDESCRIPTION. Male: Antennae and labrum fulvous, with ridges, feebly concave on anterior margin. Frons with- mandibles fulvous with black base. Anterior margin of clypeus out impressions, with short ocular grooves. Antennae thin triangularly emarginated. Mandibles simple, not flattened, except two basal segments, proportions of segments are as without basal lobe. Prothorax strongly punctuate through- 11–9–11–19–19–17–15–13–16–18–19. Prothorax about 1.8 out. Elytra without lateral ridge, punctured rows obliterated times as wide as long, obtusely angulate in basal quarter, on sides behind middle. Aedeagus short, slightly narrowed lateral margin before angulation slightly arcuate, anterior and before middle, with rounded apex, having very small protu- posterior angles distinct, produced; collar sharp, not inter- berance (Fig. 3). rupted in middle. Surface finely and sparsely punctuate, Length — 1.8–2.0 mm. interspaces microsculptured. Elytra 1.4 times as long as Female: Practically identical with male, but elytra with wide, humeral tubercle high, basal convexity and postbasal very feeble and short lateral ridge. impression well developed, rows of punctures well devel- Length — 2.3 mm. oped and distinct to apex, interspaces flat or slightly convex, Basilepta viridipenne (Motschulsky, 1860) impunctate, with fine microsculpture. Propleurae punctuate. Length 3.2–3.4 mm. = Basilepta frontalis (Baly, 1867), syn. n. TYPE MATERIAL. Lectotype of Nodostoma viridipenne Basilepta oblonga (Motschulsky, 1866) Motschulsky, 1860 (designated here), “Ind. or.”. = Tricliona oblonga Weise, 1914 NOTES. I compared types of both species and found TYPE MATERIAL. Lectotype of Nodostoma oblonga Mots- that they are identical. This species is very common and chulsky, 1866 (designated here), “Ceylon”. Paralectotype (de- widely distributed in southern Asia. stroyed, without head, prothorax and elytra) with same label as lectotype. Basilepta sculpturata (Motschulsky, 1860) NOTES. Nodostoma oblonga Motschulsky, 1866 was = Basilepta bhamoense (Jacoby, 1892), syn. n. erroneously removed in the genus Tricliona Lefevre, 1885. TYPE MATERIAL. Lectotype of Nodostoma sculpturata REDESCRIPTION. Red fulvous, legs more pale, anten- Motschulsky, 1860 (designated here), “Ind. or.”. nae pale, fulvous with darkened apical segments. NOTES. I compared types of both species and found Head with distinct but very sparse punctures, clypeus that they are identical. Specimens from Himalaya (Sikkim) not divided from frons, delimited on sides with ridges, feebly determined by S. Kimoto as Basilepta viridipenne also belong concave on anterior margin. Frons with small longitudinal to this species. groove in middle and short ocular grooves. Antennae thin Basilepta suturalis (Motschulsky, 1866) except two basal segments, proportions of segments are as 11–9–11–13–15–15–15 (next segments absent). Prothorax TYPE MATERIAL. Lectotype of Nodostoma suturalis Mots- chulsky, 1866 (designated here), “Ceylon”. 2 paralectotypes 1.4 times as wide as long, broadest in basal quarter forming (partly destroyed) with same label as lectotype. here rounded angle, strongly narrowed anteriorly, anterior REDESCRIPTION. Fulvous, elytral suture very nar- and posterior parts of lateral margin straight, collar devel- rowly piceous; in one paralectotype suture narrowly black, oped on sides, absent in middle. Surface with strong and in another paralectotype suture and lateral margin in anterior moderately dense punctures, interspaces flat, shining, most- two thirds black. ly a little larger than diameter of punctures. Elytra 1.2 times Body elongate, almost parallel–sided. Head finely or as long as wide, basal convexity well developed, fine elytral moderately strong punctuate, clypeus not separated from rows more or less distinct in anterior part and along suture, frons, delimited on sides with ridges, feebly concave on absent behind middle. Propleurae punctate. anterior margin. Frons without impressions, with short Length — 2.6 mm. ocular grooves. Antennae thin except two basal segments, Basilepta flavescens (Motschulsky, 1866) proportions of segments are as 11–9–11–19–19–17–15– 13–16–18 (apical segment absent). Prothorax 1.6 times as TYPE MATERIAL. Lectotype of Nodostoma flavescens wide as long, obtusely angulate in basal quarter, lateral Motschulsky, 1866 (designated here), “Java”. REDESCRIPTION. Entirely fulvous. Head impunctate, margin before angulation practically straight, anterior and clypeus poorly delimited from frons, triangularly emarginat- posterior angles distinct; collar in middle feeble or inter- ed, on anterior margin, frons with small groove in middle and rupted. Surface with moderately strong but not densely sharp ocular grooves. Antennae absent. Prothorax 1.65 times punctuate, interspaces smooth and shining. Elytra 1.3 times as wide as long, broadest in basal two fifth, forming here as long as wide, humeral tubercle high, basal convexity and rounded angle, anterior half of lateral margin almost straight, postbasal impression well developed, rows of punctures collar on anterior margin well developed, sharp; surface shin- distinct anteriorly, very feeble on indistinct behind middle. ing, impunctate. Elytra 1.15 times as long as wide, with basal Propleurae punctuate. convexity delimited exteriorly and especially behind with Length 2.6–3.1 mm. impression, surface shining impunctate except a few punc- To the knowledge of Chrysomelidae (Coleoptera) described by V.Motschulsky 413 tures around basal convexity. Propleurae and metasternum impunctate. Length — 4.3 mm. Bathseba Motschulsky, 1866 A taxonomical position of this genus was unclear and it placed traditionally in “incertae sedis” [Clavareau, 1914; Seeno & Wilcox, 1982]. Because of this 1 give a redescription of the genus and the species. REDESCRIPTION. Body elongate ovate, not pubescent 13 14 15 above. Head not excavated above eyes, with narrow furrow along inner margin of eye. Clypeus divided from frons. An- tennae filiform, apical segments practically not thickened. Prothorax transverse, about twice as broad as long. Elytra regularly punctuate. Pygidium without longitudinal groove. Anterior margin of proepisterna feebly convex. Prosternum elongate, slightly narrowed between coxae and widened pos- teriorly. Anterior femora with strong acute tooth (Fig. 14). Middle and hind tibiae emarginated before apex. Claws sim- 16 17 18 ple, but with very broad base (Fig. 14). DIAGNOSIS. Near Cleorina Lefevre, 1885 and Mouho- Figs 13–18. Details of Chrysomelidae spp. from collection tina Lefevre, 1885, but with elongate prosternum, strongly of V. Motschulsky: 13–14 — Bathseba ferruginea; 15 — transverse prothorax and simple claws. Colaspoides rugipennis; 16 — Dactylispa filicornis; 17 — D. pallidipennis; 18 — D. nigripennis; 13 — anterior femur, 14 — Bathseba ferruginea Motschulsky, 1866 claw; 15 — spermatheca; 16–18 — lateral spines of prothorax. Figs 4–5, 10–11 Ðèñ. 13–18. Äåòàëè ñòðîåíèÿ Chrysomelidae spp. èç êîëëåêöèè Â. Ìî÷óëüñêîãî: 13–14 — Bathseba ferruginea; 15 — TYPE MATERIAL. Lectotype of Bathseba ferruginea Mots- Colaspoides rugipennis; 16 — Dactylispa filicornis; 17 — D. # chulsky, 1866 (designated here), ( ), “Ceylon”. Paralectotype, pallidipennis; 18 — D. nigripennis; 13 — ïåðåäíåå áåäðî, 14 — $ ( ), with same label as lectotype. êîãîòîê; 15 — ñïåðìàòåêà; 16–18 — áîêîâûå øèïû íà REDESCRIPTION. Red or reddish fulvous with pale ïåðåäíåãðóäè. flavous antennae. Head microsculptured, clypeus distinctly punctuate, frons finely punctuate, with longitudinal impres- Stephens, 1831. S. indica Duvivier, 1891 was correctly syn- sion. Antennae reach middle of elytra, segments 2–4 sub- onymized [Clavareau 1914] with S. strigicollis (Motschul- equal, next segments a little longer. Prothorax with maximal sky, 1866) I have compared the types of both species. width before base, narrowed anteriorly, side margins round- ed, surface densely microsculptured and finely punctuate. Aoria nigripes Baly, 1860 Elytra with fine rows of punctures and broad, flat and mi- = Aoria rufula (Motschulsky, 1866), comb. et syn. n. crosculptured interspaces. Middle and hind femora with small = Aoria fuscula (Motschulsky, 1866), comb. et syn. n. obtuse tooth. Aedeagus — Figs 4–5. TYPE MATERIAL. Lectotype of Eumolpus rufulus Mots- Length of male 3.5 mm, of female — 4.1 mm. chulsky, 1866 (designated here), “Ind. or.”; Lectotype of Eumolpus fusculus Motschulsky, 1866 (des- Pagria restituens Walker, 1859 ignated here), “Ind. or.”. = Pagria aenea (Motschulsky, 1866), comb. et syn. n. NOTES. Both species were described as Eumolpus sensu = Pagria costatipennis Jacoby, 1887, syn. n. Redtenbacher, 1858 (a synonym of Adoxus Kirby, 1837) and TYPE MATERIAL. Lectotype of Odontionopa aenea Mots- later placed under question in Trichochrysea Baly, 1860. chulsky, 1866 (designated here), “Ceylon”. They are however entirely identical with Aoria nigripes NOTES. Pagria aenea was described as Odontionopa Baly, 1860, the mostly abundant species in Oriental region. [Motschulsky, 1866] and placed in the genus Scelodonta Motschulsky’s types were compared with specimen of A. Westwood, 1837 according Lefevre’s opinion, but the type nigripes compared with Baly’s type. was never investigated. The species is however a typical Pagria Lefevre, 1884 because of appendiculate claws and is a Cleoporus lateralis (Motschulsky, 1866), comb. n. synonym of Pagria restituens Walker, 1859. I have com- = Cleoporus variabilis Baly, 1874, syn. n. pared Motschulsky’s type with a series of Pagria costati- TYPE MATERIAL: Lectotype of Nodostoma laterale Mots- pennis Jacoby, 1887, including type. The latter species has chulsky, 1866 (designated here), “Ind. or.”. Paralectotype with legs usually dark with apices of tibiae and tarsi fulvous, but a same label as lectotype. few specimens have middle and hind legs entirely fulvous. NOTES. Nodostoma laterale Motschulsky, 1866 has con- P.aenea has all legs entirely fulvous. So, P. costatipennis is vex anterior margin of propleurae, broad excavation above eye also a synonym of P. restituens. and toothed claws — typical characters of the genus Cle- oporus Lefevre, 1884. Species in question is identical with Scelodonta dillwyni Stephens, 1831 C.variabilis Baly, 1874 and represent aberration with dark = Scelodonta indica Duvivier, 1891 outer part of elytra. Types of both species were studied. = Scelodonta strigicollis (Motschulsky, 1866), syn. n. TYPE MATERIAL. Lectotype of Odontionopa strigicollis Colaspoides rugipennis (Motschulsky, 1860), comb. n. Motschulsky, 1866 (designated here), “Ind. or.”. 2 paralecto- Fig. 15 types, with same label as lectotype. = Colaspoides paviei Lefevre, 1890, syn. n. NOTES. Odontionopa strigicollis Motschulsky, 1866 is TYPE MATERIAL. Lectotype of Colasposoma rugipennis entirely identical in all characters with Scelodonta dillwyni Motschulsky, 1860 (designated here), $, “Ind. or.”. 414 L.N. Medvedev

NOTES. This species, erroneously placed in Colasposoma twice as long as 2, but a little shorter than 4, next segment Laporte, 1833 is a typical Colaspoides Laporte, 1833. Species subequal to 4. Prothorax 1.4 times as wide as long, with lateral belongs to group 5 [Medvedev, 2003], having entirely metallic margins almost straight, surface with feeble oblique impres- underside and legs and unarmed femora. A form of spermatheca sion on each side, densely punctuate. Elytra with dense, but (Fig. 15) shows its identity with C. paviei Lefevre, 1890. not strong punctures, less shining than prothorax, with traces of longitudinal impressions. Apex of pygidium rounded trun- Pseudoides coeruleipennis (Motschulsky, 1860), comb. n. cate in both sexes. Anterior coxal cavities closed. Aedeagus Fig. 6 (Figs 7–8) is very complicate. Middle lobe of last abdominal = Pseudoides flavicollis Jacoby, 1903, syn.n. segment elongate, slightly concave, with very feeble longitudi- TYPE MATERIAL. Lectotype of Luperus coeruleipennis nal ridge. Motschulsky, 1860 (designated here), (#), “Ceylon”. Paralec- Length of male 5.2 mm, of female 6.2 mm. totype with same label as lectotype. DIAGNOSIS. The species described as Luperodes (= Atra- NOTES. Pseudoides flavicollis Jacoby, 1903 described chya nigrocincta Motschulsky, 1858 is transferred to Mono- from South India (Nilgiri Hills) identical with this species lepta Erichson ,1843 because of closed coxal cavities. It is near including form of aedeagus and represents a new synonym. M. marginipennis Jacoby, 1892 and M. atrimarginata Kimo- REDESCRIPTION. Fulvous, elytra metallic blue, under- to, 1989. M. nigrocincta Jacoby, 1900 is renamed to M. jacob- side blackish blue. yana Medvedev, 2006, nom. n., because of homonymy. Body elongate ovate. Head impunctate, frontal tubercles transverse, delimited behind with transverse line, interanten- Monolepta basalis (Motschulsky, 1858), comb. n. nal ridge narrow, frons broad, 2.5 times as wide as transverse Figs 9–10 diameter of eye. Proportions of antennal segments are as 16– TYPE MATERIAL. Lectotype of Luperodes basalis Mots- 9–7–14–13–17–15–15 (next segments absent), preapical seg- chulsky, 1858 (designated here), (#), “Ind.or.”. Paralectotype, ments about 3 times as long as wide. Prothorax 1.5 times as with same label as lectotype. wide as long, broadest behind middle, anterior and posterior REDESCRIPTION. Fulvous with elytra a little more borders distinctly margined, lateral ones arcuate; surface with- pale, hind part of vertex, labrum, antennae except basal seg- out any impressions, shining, with very fine microsculpture. ment, basal margin of elytra, including humerus, anterior Elytra 1/6 times as long as wide, without postbasal impres- third of lateral margin and very often basal part of suture, sion, strongly and densely punctuate, interspaces mostly breast basal half of femora, apical half of tibiae and tarsi smaller than diameter of punctures, shining, with very fine black. microsculpture. Prosternum not visible between coxae, coxal Head smooth, shining, finely and sparsely punctuate, cavities very narrowly closed. Apical part of aedeagus — frontal tubercles triangular, delimited behind with deep trans- Fig. 6 (basal part was destroyed). verse impression; frons broad. Antennae thin, about ¾ of Length 3.4–3.7 mm. body length, segment 3 a little longer than 2, segment 4 as DIAGNOSIS. This species has closed coxal cavities, long as two preceding together, next segments subequal to 4. spurs on mid and hind tibiae, thin antennae with fifth to Prothorax 1.4 times as wide as long, with maximal width near eighth segments more then twice as long as wide. Because of anterior margin, side margins feebly rounded, surface without all these characters it might be included in the genus any impressions, shining, finely and more or less densely Pseudoides Jacoby, 1892. punctuate. Elytra shining, finely punctuate. Anterior coxal cavities closed. Aedeagus — Figs 9–10. Atrachya bimaculata Hornstedt, 1788 Length of body 3.2–3.5 mm. = Atrachya maculicollis (Motschulsky, 1858), comb. et syn. n. DIAGNOSIS. The species was described as Luperodes TYPE MATERIAL. Lectotype of Cnecodes maculicollis (= Atrachya) basalis Motschulsky, 1859, but must be trans- # Motschulsky, 1858 (designated here), ( ), “Ind. or.”. ferred to Monolepta, as having closed coxal cavities. For NOTES. This species placed till now in Monolepta Erich- M.basalis Harold, 1880 I propose a new name M. haroldi- son 1843, was described as having an oblong patch in the ana Medvedev, 2006, nom. n., because of homonymy. middle of prothorax. In the single type specimen the protho- rax is damaged in middle, and this part looks like elongate Hyphaenia indica (Motschulsky, 1866 ), comb. n. asymmetrical dark spot. In all other characters this species is TYPE MATERIAL. Lectotype of Orsodacna indica Mots- fully identical with unspotted form of Atrachya bimaculata chulsky, 1866 (designated here), female, “Ceylon”. Paralecto- Hornstedt, 1788. Male of this species has a narrow elongate type, female, with same label as lectotype. groove on elytra near suture. This character seems to be REDESCRIPTION. Fulvous, side margins of prothorax overlooked by previous authors [see: Maulik, 1936] and narrow darkened, elytra more pale, with suture very narrow- firstly mentioned by Kimoto [1989]. ly darkened, antennae black with segments 1 and 2 fulvous. Body narrow, elongate. Head impunctate, frons as wide as Monolepta nigrocincta (Motschulsky, 1858), comb. n. diameter of eye. Antennae as long as body, without erect hairs, Figs 7–8 proportions of segments are as 9–2–7–10–10–10–10–9–9–9– TYPE MATERIAL. Lectotype of Luperodes nigrocincta 10, preapical segments about 6 times as long as wide. Protho- Motschulsky, 1858 (designated here), (#), “Ind.or.”. Paralecto- rax 1.3 times as wide as long, feebly cordiform, broadest behind type, ($), with same label as lectotype. anterior angles and narrowed to base, surface with broad and REDESCRIPTION. Fulvous; labrum, all margins of elytra very shallow transverse impression, shining, impunctate, with very narrowly (including suture), inner margin of epipleurae, traces of microsculpture. Elytra parallel–sided, times as long as tibiae and tarsi black; antennae except two basal segments wide, without distinct basal convexity and postbasal impres- and breast infuscate. Elytra a little more pale than prothorax. sion, but with shallow impression inside of humeral tubercle, Head smooth, shining, finely and sparsely punctuate, with surface shining, strongly and densely punctate. distinct transverse impression behind triangular frontal tuber- Length 4.2–4.3 mm. cles, frons broad, wider than transverse diameter of eye. An- DIAGNOSIS. This is a single species from Ceylon and tennae thin, about 2/3 of body length, third segment almost South India with fulvous colour. To the knowledge of Chrysomelidae (Coleoptera) described by V.Motschulsky 415

Chaetocnema (Tlanoma) gracilis Motschulsky, 1859 NOTES. Many authors, following the original descrip- = Chaetocnema minuta Jacoby, 1896 tion of this genus accept anterior coxal cavities open and = Chaetocnema indica Weise, 1916, syn. n. prothorax without impressions [Maulik, 1926; Scherer, 1969]. TYPE MATERIAL. Lectotype of Tlanoma gracilis Mots- However Ogloblin [1930] correctly indicated that this genus chulsky, 1859 (designated here), “Ind. or.”. The length of type has closed coxal cavities and short longitudinal grooves on (not mentioned in the original description) is 1.5 mm. base of prothorax. I can confirm these characters after study- NOTES. Chaetocnema indica Weise, 1916 (new name for ing the type. Therefore this genus must be placed near Poda- Chaetocnema minuta Jacoby, 1896) is a new synonym of grica Chevrolat, 1837 and Kamala Maulik, 1926. It differs Chaetocnema gracilis Motschulsky, 1859. Both species are from the first with rounded and convex body form and very entirely identical morphologically, have densely microsculp- regular rows on elytra, from the latter in body much larger tured prothorax and smallest size among Oriental species and wings developed. (1.4–1.7 mm). Nisotra gemella Erichson, 1834 Chaetocnema (Tlanoma) puncticollis = Nisotra javana (Motschulsky, 1866), syn. n. Motschulsky, 1859 TYPE MATERIAL. Lectotype of Sphaeroderma javana = Chaetocnema discreta Baly, 1876, syn. n. Motschulsky, 1866 (designated here), (#), “Java”. 3 paralecto- TYPE MATERIAL. Lectotype of Tlanoma puncticollis Mots- types, ($$), with same label as lectotype. chulsky, 1859 (designated here), “Ind. or.”. 3 paralectotypes NOTES. Ogloblin [1930] investigated already this spe- with same label as lectotype. The length of type specimens is cies but had only 2 females and therefore can not make 1.9–2.2 mm. conclusion about its taxonomical status. Happily I found a NOTES. Chaetocnema discreta Baly, 1876, very usual in male specimen; a study of aedeagus showed that it is a new Oriental region, is entirely identical with Motschulsky’s synonym of Nisotra gemella Erichson, 1834. species in all characters. Psylliodes palleola Motschulsky, 1866 Longitarsus suturellinus Csiki, 1940 TYPE MATERIAL. Lectotype of Psylliodes palleola Mots- = Longitarsus suturellus (Motschulsky, 1866) [preoccupied chulsky, 1866 (designated here), “Ceylon”. by Longitarsus suturellus Duftschmid, 1825] REDESCRIPTION. Pale flavous, including lateral mar- = Longitarsus rangoonensis Jacoby, 1892, syn. n. gins of elytra (in the original description they are mentioned TYPE MATERIAL. Lectotype of Teinodactyla suturella as darkened). Seems to be immature specimen. Motschulsky, 1866 (designated here), “Ind. or.” (not Ceylon, as Body narrow, elongate. Frontal ridge developed, frontal was given in the original description). 3 paralectotypes with tubercles very inner margins of eyes, widened posteriorly. same label as lectotype. NOTES. Teinodactyla suturella Motschulsky, 1866 was Vertex densely microsculptured, without punctures. Protho- renamed for Longitarsus suturellinus Csiki, 1940 because rax transverse with sides almost straight and parallel, ob- homonymy with L. suturellus Duftschmid, 1825. It is identi- liquely truncate anteriorly, but without distinct angulation; cal with L. rangoonensis Jacoby, 1892 (same colour and surface densely punctuate with shagreened interspaces and structure, including punctured stripe on frons) which must distinct longitudinal groove on each side of basal margin. be accepted as a new synonym. Elytra parallel, moderately convex, with segment 1 about 2/5 of tibia length. Aphthona albescens (Motschulsky, 1866), comb. n. Length 3.7 mm. = Aphthona opaca Allard, 1889, syn. n. DIAGNOSIS. Morphologically identical with P. viri- TYPE MATERIAL: Lectotype of Teinodactyla albescens dana Motschulsky, 1866, differs only in coloration and small Motschulsky, 1866 (designated here), “Ind. or.”. 3 paralecto- size. It might be an immature specimen of the latter species. types with same label as lectotype. NOTES. Aphthona albescens (Motschulsky, 1866) de- Dactylispa filicornis (Motschulsky, 1861), stat. rest. scribed as Teinodactyla Chevrolat 1842 was placed in the Fig. 16 genus Longitarsus [Maulik, 1926], but is fully identical with TYPE MATERIAL. Lectotype of Hispa filicornis Motschul- Aphthona opaca Allard, 1889, which is therefore a new sky, 1861 (designated here), “Ind. or Nepal”. synonym of A. albescens. NOTES. Hispa filicornis Motschulsky, 1861 was synon- A. opaca is known from China and Indochina, but I have ymized under question with Dactylispa brevispinosa Chapuis, in my collection a few specimens from Burma. This species 1877, but is a good species near D. singularis Gestro, 1888. has comparatively long the first segment of hind tarsus and The type is strongly damaged: only two antennal segments are looks as transitional between Aphthona Chevrolat, 1842 and present, prothoracic spines mostly broken, elytra absent. Nev- Longitarsus Berthold, 1827. ertheless its systematic position is quite clear. REDESCRIPTION. Body entirely black, upperside of Hyphasis cyanipennis (Motschulsky, 1866) abdomen pitchy red. Antennae (two basal segments) with TYPE MATERIAL. Lectotype of Aphthona cyanipennis deep longitudinal striation. Head very uneven, with acute Motschulsky, 1866 (designated here), “Ind. or.”. 2 paralecto- process between antennae, deep round groove in middle of types with same label as lectotype. vertex and deep furrow along inner margin of eye. Prothorax NOTES. Aphthona cyanipennis Motschulsky, 1866 was has on each side of anterior margin one stout spine, having studied by Ogloblin [1930], who synonymized Hyphasis additional small spine near base (Fig. 16), side margin with 3 bevani Baly, 1878 with this species. But this publication spines arising independently, anterior two are rather short was missed by Heikertinger & Csiki [1940] and later by and stout; the third spine thin and at least twice as short as G.Scherer [1969], who used again a name H. bevani. the first. Surface with very large and deep round punctures (practically grooves) divided with very narrow interspaces; Thrylaea flavipennis (Motschulsky, 1866) middle part transversely elevated, not smooth, divided in TYPE MATERIAL. Lectotype of Hypnophila flavipennis middle with impressed line and delimited behind with deep Motschulsky, 1866 (designated here), “Ceylon”. transverse depression. 416 L.N. Medvedev

Length (to the apex of abdomen) 5.1 mm. Gonophora nigricauda (Motschulsky, 1863) DIAGNOSIS. Near D. singularis from Burma, but latter = Gonophora taprobane Gestro, 1902, syn. n. species has the second rudimental spine on anterior margin of TYPE MATERIAL. Lectotype of Anisodera nigricauda prothorax distinctly divided from main spine, two anterior Motschulsky, 1863 (designated here), “Ceylon”. lateral spines have common base and an elevated transverse NOTES. Motschulsky indicated size of his species as 4 area is smooth and shining. mm, in reality it is 4.7 mm. Maulik [1919] correctly included it in the genus Gonophora Baly, 1858, but indicated that “it Dactylispa longicornis (Motschulsky, 1861) is unlikely that this may be a small specimen of Gestro’s = Dactylispa severini Gestro, 1897, syn. n. G.taprobane” (the latter is 4.5–5 mm). I compared types of TYPE MATERIAL. Lectotype of Hispa longicornis Mots- both species and found that they are identical. chulsky, 1861 (designated here), “Ind. or Birma”. NOTES. Hispa longicornis Motschulsky, 1861 was united LECTOTYPES (LT) AND PARALECTOTYPES (PLT) DESIGNATED under question with Dactylispa severini Gestro, 1897 [Mau- FOR SPECIES WITH CLEAR TAXONOMICAL POSITION. lik, 1919] but later was replaced to unclear species [Uhmann, 1958]. I can confirm a full identity of both species. Criocerinae Dactylispa nigromaculata (Motschulsky, 1861) Lilioceris subpolita (Motschulsky 1860) — LT (Japan) = Dactylispa xanthospila Gestro, 1890, syn. n. Cryptocephalinae TYPE MATERIAL: Lectotype of Hispa nigromaculata Mots- chulsky, 1861 (designated here), “Ind. or Birma”. Cryptocephalus carneobifasciatus Motschulsky, 1866 NOTES. Hispa nigromaculata Motschulsky, 1861 was (=C. suillus Suffrian, 1860) — LT (Ceylon) correctly included in the genus Dactylispa Weise, but placed Cryptocephalus obliquostriatus Motschulsky, 1866 among unclear species [Uhmann, 1958]. I found that it is (=C. parvulus Mueller, 1776) — LT + PLT (Japan) identical with D. xanthospila Gestro, 1890, which is there- Melixanthus luridus (Motschulsky, 1866 (=M. hians fore a synonym of Motschulsky’s species. Suffrian, 1860) — LT (Ceylon) Dactylispa nigripennis (Motschulsky, 1861) Fig. 18 Basilepta uniformis (Motschulsky, 1866) — LT + 3 PLT = Dactylispa soror Weise, 1897, syn. n. TYPE MATERIAL. Lectotype of Hispa nigripennis Mots- (Ceylon) chulsky, 1861 (designated here), “Ind. or.”. Paralectotype, with Colasposoma viridicoeruleum Motschulsky, 1860 — same label as lectotype. LT (China: Hongkong) NOTES. Hispa nigripennis Motschulsky, 1861 fully Colasposoma viridifasciatum Motschulsky, 1860 — LT identical with melanistic specimens of Dactylispa soror Weise, (Philippines) 1897, which therefore is a new synonym of Motschulsky’s Colasposoma cyaneum Motschulsky, 1860 (=C. dauri- species. cum Mannerheim, 1849) — LT (Dauria) Dactylispa pallidipennis (Motschulsky, 1861) Colasposoma mongolicum Motschulsky, 1860 (=C. dau- Fig. 17 ricum Mannerheim, 1849) — LT (Mongolia) TYPE MATERIAL. Lectotype of Hispa pallidipennis Mots- Colasposoma auripenne Motschulsky, 1860 (=C. viridi- chulsky, 1861 (designated here), “Ind. or.”. 2 paralectotypes, coeruleum Motschulsky, 1860) — LT +8 PLT (Ind. or.) with same label as lectotype. Colasposoma purpuratum Motschulsky, 1860 — LT + REDESCRIPTION. Body fulvous, including antennae, PLT (Java) all elytral spines and spots around their bases black. Two Trichochrysea japana (Motschulsky, 1857) — LT (Ja- paralectotypes have antennae strongly darkened and protho- rax with two small and not very distinct black spots. pan) Morphologically this species is fully identical with pre- Chrysomelinae ceding, except other arrangement of prothoracic lateral spines. In D. pallidipennis (Motschulsky, 1861) the third small Ambrostoma chinense Motschulsky, 1860 (=A.fortunei spine placed just at base of common stock (Fig. 17); in Baly, 1860) — LT (China) D.nigripennis a small spine is distinctly removed from this Ambrostoma nepalense Motschulsky, 1860 (=A.mahesa stock (Fig. 18). In any case D.pallidipennis is very near to Hope, 1831) — LT (Nepal) D.nigripennis, D.kamarupa Maulik, 1919 and D. discicollis Plagiodera rufolimbata (Motschulsky, 1860) Gestro, 1890. (=P.egregia Gerstaecker ,1855) — LT (Mozambique) Gonophora zinzibaris (Motschulsky, 1861) Chrysolina cribellata (Motschulsky, 1860) (=Ch.herbacea Duftschmidt, 1825) — LT (Dalmatia) = Gonophora akalankita Maulik, 1919, syn. n. TYPE MATERIAL. Lectotype of Anisodera zinzibaris Mots- Chrysolina difficilis (Motschulsky, 1860) (strongly dam- chulsky, 1861 (designated here), “Ceylon, Mt. N. (=mount aged, but with aedeagus) — LT (Barnaul) Nuwara Eliya)”. 2 paralectotypes with same label as lectotype. Chrysolina instructa (Motschulsky, 1860) NOTES. This is very distinct and quite satisfactory (=Ch.aeruginosa Faldermann, 1835) — LT (Dauria) described species (only the length of body is 4.5 mm, not 3.5 Chrysolina tarda (Motschulsky, 1860) as it was given in the description). Gonophora akalankita (=Ch.aeruginosa Faldermann, 1835) — LT Sib. or.) Maulik, 1919 is a synonym of this species, types of both Gonioctena salicis Motschulsky, 1860 (=G.affinis Gyllen- species were compared and they are identical, only Maulik’s type is a little larger. hal, 1808) — LT (Sib.: Armenia) + 6 PLT (Sib. or. bor.) Gonioctena sorbi Motschulsky, 1860 (=G.sibirica Weise, 1893) — LT + 7 PLT (Dauria: Mt. Hamar-Daban) To the knowledge of Chrysomelidae (Coleoptera) described by V.Motschulsky 417

Phratora longula Motschulsky, 1860 (=Ph.vulga- Nisotra viridipennis (Motschulsky ,1866) — LT + PLT tissima Linné 1758) — LT + PLT (Kamchatka) (Ceylon) Phratora obtusicollis Motschulsky, 1860 — LT (Amur) Parlina trancisa Motschulsky, 1866 — LT (Ceylon) Psylliodes viridana Motschulsky, 1858 — LT (Ceylon) Sangariola punctatostriata (Motschulsky, 1860) — LT Exosoma flaviventre (Motschulsky, 1860) — LT (Japan) + PLT (Japan) Galeruca extensa (Motschulsky, 1861) — LT (Japan) Sphaerodermella rufopicta (Motschulsky, 1866) — LT Hyphaenia pilicornis (Motschulsky, 1858) — LT + 2 (Ceylon) PLT (Ind. or.) Medythia suturalis (Motschulsky, 1858) — LT +2 PLT Hispinae (Ind. or.) Asamangulia tuberculosa (Motschulsky, 1861) — LT + Strobiderus albescens (Motschulsky, 1866) — LT (Cey- PLT (Nepal) lon) Hispa ceylonica Motschulsky, 1861 (=Hispa ramosa Gyllenhal, 1817) — LT + 2 PLT (Ceylon) Alticinae Hispellinus australis (Motschulsky, 1861) — LT + PLT Aphthona atripes (Motschulsky, 1866) — LT (Ceylon) (Nov. Hollandia) Aphthona nigrita Motschulsky, 1866– LT (Ceylon) Argopus punctipennis (Motschulsky, 1866) — LT + PLT Cassidinae (Japan) Cassida rugosopunctata Motschulsky, 1866 Chaetocnema nigrica Motschulsky, 1858 — LT + 5 PLT (=C.rubiginosa Mueller, 1776) — LT (Japan) (Ind. or.) Chaetocnema splendens (Motschulsky, 1845) — LT + References PLT (Sib. or.) Chaetocnema tarda (Motschulsky, 1845) — LT (Cauca- Clavareau H. 1913. Coleopterorum Catalogus. Chrysomelidae. sus: Pjatigorsk) Pars53. P.1–278. Berlin: W.Junk. Dibolia metallica (Motschulsky, 1845) — LT (Cauca- Clavareau H. 1914. Coleopterorum Catalogus. Chrysomelidae: sus: Pjatigorsk) Eumolpinae. Pars59. P.1–215. Berlin: W.Junk. Elytropachys dimidiata (Motschulsky, 1858) — LT Heikertinder F. & Csiki E. 1940. Coleopterorum Catalogus. (Ceylon) Chrysomelidae: Alticinae. Pars166,169. P.1–635. Graven- hage, Uitgeverij: Dr.W.Junk Elytropachys dorsalis Motschulsky, 1866 — LT (Cey- Kimoto S. 1989. Chrysomelidae of Thailand, Cambodia, Laos lon) and Vietnam. IV. Galerucinae // Esakia. No.27. P.1–241. Elytropachys latissima (Motschulsky, 1858) — LT + Maulik S. 1919. The fauna of British India, including Ceylon and PLT (Ceylon) Burma. Chrysomelidae: Hispinae and Cassidinae. London: Taylor and Francis. 439 pp. Elytropachys obscurata Motschulsky, 1866 — LT +2 Maulik S. 1926. The fauna of British India, including Ceylon and PLT (Ceylon) Burma. Chrysomelidae: Chrysomelinae and Halticinae. Elytropachys suturanigra (Motschulsky ,1866) London: Taylor and Francis. 442 pp. [=E.tropica (Motschulsky, 1866)] — LT (Ceylon) Maulik S. 1936. The fauna of British India, including Ceylon and Elytropachys tropica (Motschulsky, 1866) — LT (Cey- Burma. Chrysomelidae: Galerucinae. London: Taylor and Francis. 648 pp. lon) Medvedev L.N. 2003. Revision of the genus Colaspoides Elytropachys viridescens Motschulsky, 1866 — LT + Laporte (Chrysomelidae, Eumolpinae) from continental PLT (Ceylon) Asia // Russ. Entomol. Journ. Vol.12. No.3. P.257–297. Elytropachys viridifusca (Motschulsky, 1858) — LT Ogloblin D.A. 1930. De quelques especes de Halticinae de la collection de V. Motschulsky // Eos. Vol.6. P.83–112. (Ceylon) Scherer G. 1969. Die Alticinae des indischen Subkontinentes. Ivalia ruficollis (Motschulsky, 1866) — LT Pacific Monograph. Vol.22. P.1–251. Liprus rufotestaceus (Motschulsky, 1866) Seeno T. & Wilcox J. 1982. Leaf genera // Entomogra- (=L.punctatostriatus Motschulsky, 1860) — LT (Ja- phy. Vol.1. P.1–221. pan) Uhmann E. 1958. Coleopterorum Catalogus. Supplementa. Chrysomelidae: Hispinae. Pars35. P.1–490. Gravenhage, Nisotra orbiculata (Motschulsky, 1866) (=N.gemella Uitgeverij: Dr.W.Junk. Erichson, 1834) — LT (Ind. or.)