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(966t)toz-t8I :(Z) 0t ersfnN t82 NuytsiaVol.10, No.2 (1995) related to certain bipinnateleaved speciesof seclior,Botrycephalqe (Benth.) Taub. (seeTindale & Roux 1969 & 1974, Andersonet al. 197| & 1984, Anderson1978, Vassal 1972,Chappill & Maslin 1995,Brain 1987,and Brain & Maslin in prep.).

While phyltode nerve number and inflorescenceform are useful charactersfor recognizing ,,uninerves-Racemosae"species these are not completelyreliable charactersfor ascribingtaxa to this (artificial) group. For example, A. difformis R. Baker has 2 longitudinal nerveson each face of its "Acaciq phyllodes yet its inflorescence and carpological characterssuggest its inclusion in the microbotrya Group"; similarly, A. binen'ata DC. and A. wardellii Tind have Prominently 2- or 3-nervedphyllodes but are clearly relatedto A. bancrofrii (l-nerved phyllodes)which is placed in the "Acacia microbotrya Group" (seenotes below underA. brumalis sp.nov). There are also a number oftaxa, especiallyftom southwestwestern Austxalia,with 2 nervesalong the (often thickened)upper margin of the phyllodes, thus making a total of 5 nervesper phyllode (2 along the upper margins, I on each face and I along the lower margin); thesetaxa uncommonly have racemoseinflotescences (e.g. A. scleroclada sp. nov. below). Similarly there are some caseswhere species with simple "Acaciq inflorescencesare closely related to racemosetaxa, e g. A synchronicia Maslin in the victoriae cfor]p,, (Maslin 1992). The length of the raceme varies considerablybut within section Phyllodineae thereseems to be a significanttaxonomic discontinuity between those taxa having very reducedracemes (the axesgenerally not aboveabout I mm long and supportingI or 2 heads)and those having more elongatedracemes with more numerousheads. A recent study by Brain and Maslin (in prep.) using serologicaldata suggeststhat specieswith very reducedracemes are most closely related io non-ru""-or" speciesand that there is little or no relationshipbetween this combined group and the ,,Uninerves-Racemosae",These relationshipsbased on biochemical data wele an unexpected result ofthe Brain and Maslin study,however, ifthey arecoroborated by other evidencethen a major rearrangementof the classificationof sectionPhyllodineae will be needed.

"uninerves-Racemosae"; This paperis not a revision ofthe the intentionhere is simply to describe "Flora new speciesand lectotypify namesahead of their inclusion in the forthcoming of Australia' volumes.

Methods

The taxa included in this paper are ananged alphabeticallyby speciesname. unless oth€rwise statedall plant measurementsare taken from dried herbadummaterial. Botanical Distdcts referred to under the Distribution of eachnew speciesare defined by Beard ( 1980). speciesdisfiibutions ale also expressedby 1: 250 000 map number (seeMaslin and Pedley 1982 for maps of the Ausfalian Acdcia flo'a published in this sameformat).

My approachto both typification and the application of rank are discussedelsewhere (Maslin & Cowan 1994a and Cowan & Maslin 1995 respectively) --1

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2.5 km NW of Carnamahon Midlands Road to Three Springs,B.R. Maslin 6583 (MEL, PERTH); Mouroubra Station, 68.5 km S of PaynesFind, B.R. Maslin 6666 (PERTH); East Yuna Reserve, A28415,29231, c. 70 km NE of Ceraldton, B.G. Muir 158(2.7)(PERTH); 5 km E of Kanoun Hill, 2 P. Roberts223 (PERTH); Cowcowing, F. Stoward 370 (PERTH).

Distribution, Southw€st Westem Australia in the Irwin and Avon Botanical Districts with one collection from the extremenorthwest ofthe CoolgardieBotanical Distlict (l:250,000 mapsG50-13, H50-1,2,6,7,10,11). Occurs from near Yuna (which is c. 35 km eastof Northampton) southeastto a Cowcowing (c. 20 km north-northeastof Wyalkatchem), also near Galena (which is c. 35 km p northwest of Yuna) and Kanoun Hill (c. 75 km northwestof Cowcowing). a I Hqbitqt, Grows in flat, low-lying areason red-brown sand or loam in woodland or shrubland. a c I FLoweringand period^s.Flowering has been recordedfrom August to December,however, fruiting l, the main flush occurs in September.Pods with matureseeds have been collectedfiom November to U January,

Conservations/atus. Widespread,not known to be threatened.

Etymology. From the Creek, anthos -f1ower,and chairo - rejorce.Alluding to the bright goldenheads clusteredin sholt terminal racemes.The nameis also a very appropriatecommemoration of the birth of my daughter, Kimberly Sarah, in March 1991, around the time that the original draft of rhis description was prepared,

"Acacia Affinities. Acacia anthochaera belongs to the prainii Group" (see discussion under A. dorsenna sp. nov. below) and until now was regardedas a long phyllode yariant of A, hemiteles Benth. While thesetwo specieshave very similar inflorescenceand carpological featuresthey are readily distinguished by their growth habit, bark characteristicsand phyllode proportions. The principal characterswhich set A. hemiteles apart fiom the new speciesare the following: shrub 0.5-2(3) m tall; bark smooth, sometimesgrey and fissuredat the extremebase of the relatively thin stems;phyllodes narrowly elliptic to a narrowlyoblong or linear,4-10 cm long,4-9(11) mm wide, length to width ratio 5-15, coriaceousand somewhatrigid, straight to recurved, rarely shallowly incurved, mucro commonly coarsely pungent. The dist butions of the two speciesoverlap in the region betweenabout Beacon (c. 55 km northeastof Koorda) and Morawa, and the morphology of a few herbariumspecimens suggest that hybridizationmay occur(e.g. c. l5 km NW of Kalannietowards Dalwallinu, B.R. Maslin 3977, PERTH; 13 km E of Coorow towards Latham, B.R. Masltn 4128 & 4128A.PERTH).

Lrke A. pruinii Maidet, A. anthochaeragrows to a beautiful,shapely tree with a densecanopy and massesof iiagrant, bright golden heads. Occasionally the two are sympatric, for example, on Mouroubra Station, c. 70 km south of Paynes Find; B.R. Maslin 6666 (A. anthochaera) and B.R. Maslin 6667 (A. prainii). At this locality the general facies of the two specieswas similar, however,A. anthochaerq occwred as a tree 7 m tall (which seemsto representthe maximum height attainedby the spccies)whereas,4. 2ralnii reachedonly 4 rn;'A. prainii was further distinguishedby its shorter (to c. 5 cm long), pungentphyllodes.

In phyllodes and sometimesin habit A. anthochaerq may resembleA. aestiy.tlisE. Pritz. which has light-golden, appressed-puberulousraceme axes, united sepalsand has seedswith a long, red- brown (brittle) funicle and a thick aril. 'rupJSoJnald 'epl^\ '3uo1ruut 'cgd111e eql SurrepJoq,{urqs-qns sarupeuos lnq llnp unu g-7 g-tr o13uo1qo 'pulpnl€uo1 'snorqr13'el€lnclleJ spaas qsDlo?lqo1 u,ro.rqqsqdrnd {Jep lou lqSrcJls ssel Jo elou q 'sno€c"leqc ,{lruJrJo1 snoecer:oc ,{1urqt 'snolnpued 'SuuJncco sarxlleuos suollculsuocdeep luopuEJ ',tlel"Jeporu ^\el e qlr,r speesuee,rleq pelcrJlsuos(aoleq I lu"I.r€A Jepunolou ees ,{1a:cr),(pq311s 'spees ot pa8pa-tq8r€rs,{llunsn eql ueo,,'r1aqpalctrlsuoc ,(lelEJepou seulnetuos lo ,{llq8lIs'speesJe^o I sPod eltq,tro1 ,rto1le,( epd srrrq pepunor ,41snor,rqoo1 ,{pq8r1s'epr,|^u]lx 9-g'3uo1 urc A1 ol'lEeuII li '^Jreq-pesseJdd€ ,{yalureporuol snoJq?I8'8uoyurur g 1-g 1 sptad ui\\oJg8ur,4rp eqnl'snolnleqnd 'l 'ql8uel 'EIIoroc 'snoledasorue8rtrlo, uepyo8elud uego seqol slr 9/I , JoJpepl^Ip ro q'fiuayy1g-711 p 'snoJJur-E sJrd4olJ speaq papucdxaunur sJJ,4\oUueJ^\leq alqBAJesqo puu umolq sno.rqr13 u 'elElorJqurJ-elrq,r 'ept,n 'ctldllle esr,rJeqlo ro -,tolle,{ eled turu g 0-t 0 ^lesJa^suerlol JelncJlcssol p ro eJou e?ururul:rueurl srAEIc'3uo1tutu 1 ,'elelled-qns ol elElnqtl?dssapapDtg pere,\\ou-St-rl '(petnlrlsuoceJ) 'ueplo8 deep o1 uep1o8-prrutq8uq slseqlw tt ruvlp txul g-g'pto8uolqo o1 re1nqol8 'crJIIoJd '(peln]tlsuoceJ) puc At^ogsspDaH sexe eurecsruo s[ runluaunput txl?lpulx I-9 0'(fup) 'uurp ujru g-1g '3uoy rtrlu (t)S-€(Z) sapunpad ur sno:qu1asetulletuos 'srleq allq,t fJe^lls 4 lrnr; SI io /'\olle,{ eled pesserddzgo runluaunPureql 'esPqlu eluetfiJqo salL'DawaJDJ:spEeq (/l)ZI-t(Z-I) 'snut,rlnd l,' qll.a Suol tutu (0t)SZ-9(0) .saurarrll snul^lnd;o pue letsrp1t ,{lpuorscrco a,roqe 'lueurruoJd ruru (0g)0g-Z pelEnlrslsoura^\ol eql'€,(leJuJ'Z Jo I tou spuolD pel{uu,,!\,{lesJe^suEr '3uol 'luept^e 'ultl?tu ,(l uu 6-1 snur,rlnd :eJncsqo,{Je se^reu leJelel lnq ,\\otmu se^JeuIuuISJsru 'J 'snolql?13'Pe^JncoJ reddn sprvrrrol,{11q3rls Jo l€Jluaclueuruord JeqlpJqupltu :snocne13otr uee:31q8tl 'SulpuecsE 'snoeccuoc ul ro pa^Jncur ,t1,no1pqs,fte,1 ro 1q3rc4s:snolnpuad Je^eu ol lualud ,{Dsoru ' '8uol qr pue uuu JeqluJLz-g ottvr qlpr.r ol qfuel'epl^\ ulur oz-t(g) (uc tl-9(99)'xedB elturcun -qns Jo elEurcunoql l3 pa^\olrgu ,(11unpur3pq,rrreruos Jo,{lldnJqB JeqluJ'esEqle pe,,\\oJJEu'J?euII ,{lpeorq sauqeuos 'crldrlle-Suolqo^l,to.uuu Jo ct1dt11eXlmo:ruu o1 cpdrlle 'e1uloecuu1qo,{1,,rorreu t'; ol elBloecuBlqo'ozrs pus eduqs uI elqptJE^sapo {qd snocnpcosalrdtjs esoulnJd,{lelEJapou JE ,{leJEJ'peJ Jo u,roJq-,\\olle,{'snoJqulS 'peqqu ,{leuIJ'selllueJlxe lg pelSu€lnq elanl slalqcuotg ueetB ',{er8'peJnssrtr -ezuoJq(Ireq ^ eu a) setuqeuosJo per ^leutl (s{unJl uIEtuJo es"q lB lseel ls) Jo qlootus 'peqcuurq '1p1 )rDB' JrlnqlpunJurselurleuos ,(1uedoleq,teulos ro ,{qsnq ut €-z saall Jo sqntqs sl 'uuuruqe'l 'JznD ql t(8t81) EgZ :Z ssle.Id Id C C I ulussrehl C J .'qtue11uou o &ldool Vf

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(s6 98r t | ,{!rBllo)srW D,r,"J y urlsar{ U g NuytsiaVol. 10, No.2 (1995) black; pleurogramcontinuous or with a narrow opening at hilar end; arcole2.5-3 mm long, c. 1 mm ph wide1.funicle filiform, yellow when fresh but &ying light brown to light reddish brown, sometimes (r with a fold near attachmentto pod then encircling the seedin a single fold and terminatingin a thick, Pa dull yellow-brown (dry) aril. en A. Distribution. SouthwestWestern Australia, principally in the Avon Botanical District but extending to peripheries of the adjoining Roe and Darling and kwin Botanical Districts (1:250,000 maps H50-1,2,6,10,11,14,15,16and 150-4). Most commonin theregion from RegansFord (on theMoore gl River north of Perth) and Wubin (c. 20 km north of Dalwallinu) south to near Hyden, but also A. extending north to near Morawa and near Northampton. pe tlr

Flowering and fruiting periods. Flowers from late May to September.Pods with matureseeds have been collected from late November to earlv January. di la ConservationJlatr.J. Not consideredrare or endangered. pl Etymology. The epithet brumaLis(Latin for wintry) refels to the predominantly winter flowering phenology of the species. te

Afrtnities. Acacia brumalis is a member of a large Australia-wide group of specieswithin section " Phyllodineae which is informally referredto hereas t e Acacia microbotrla Croup". The following v charactersare found in speciesof the Group but not all characte$are present in all species.Phyllodes c( l-nerved per face (occasionally 2-nerved, e,g. A. difformis), glabrous; inflorescencesnormally al racemose;heads globular, sometimesshortly oblongoid; bracteolespeltate or sub-peltate,commonly S{ golden-fimbriolate; pods normally firmly chartaceousto thinly coriaceous;seeds normally black; pleurogram commonly continuous;funicle filiform, normally partially or completely €ncircling the seed in a single or double fold, usually dark red-brown to light brown, occasionally pale yellow (e.g.A. jennerae,A. steedmaniiand A. vaLidinenia). A provisionallist of speciesrefered to the Group includes the following; A. aestivaLisE. Pritz. (W.A.), A. alcockii Maslin & Whibley (S.A.), A. ambLyophyllaF. Muell. (W.A., perhaps better treated as an infraspecific taxon under A. microbotrya), A. amoenaH.L.Wendl. (N.S.W.),A. angastaMaiden & Blakely (Qld), A. araneosa Whibley (S.A.), A. anceps DC, (W.A., S.A.), A. attenuata Maiden & Blakely (Qld), A. ban\oftii ( Maiden (Qld), A. brumalisMaslin (W.A.), A. caLamifuliaSweet ex Lindley (S.A., Vic., N.S.W.), A. chqmaeleonMaslin (W.A., see below), A. chalkeri Maiden (N.S.W.), A. chryselLaMaiden & L Blakely (W.A.), A. confluens Maiden & Blakely (S.A.), A. uetacea Maslin & Whibley (S.A.), ! A. deuteroneuraPedley(Qld), A. diffornis R. Baker (N.S.W.,Yic.), A. falcataWilld. (Qld, N.S.W.), A. flocktoniae Maiden (N.S.W.), A. /orsytlrli Maiden & Blakely N.S.W.), ,4. glllll Maiden & Blakely (S.A.),A. gLadiiformisCunn. ex Benth.(N.S.W.), A. x grayanaJ.H. Willis (S.A., Vic., a hybrid involvingA. brachybotryaand A. cqlamifoliq),A. harvsli Benth.(W.A.), A. jenneraeMaiden (W.A., S.A., N.T., N.S.W.), A. kydrensis Tind. (N.S.W.), A. leichhardtii Benth. (Qld, precise affinities unknown but the funicle encircling the seedssuggests inclusion in the "A. microbotrya Group"), A. leiophyllaBenth.(S.A.),A. leptopetqlaBenth. (W.4.), A. mabellaeMaiden (N.S.W.),A. meisneri Lehm. ex Meissn. (W.A.), A. merrickiae Ma;Lden& Blakely (W.A.), A. microbotrya Benth. (W.A.), A. nematophyllaF.Muell.ex Benth.(S.A.), A. notabilisF.Muell. (S.A.,N.S.W., Vtc.), A. quomensis J. Black (S.A.), A. retinodes Schldl. (S.A., Vic.), A. rlvalls J. Black (S.A., also recordedfor N.S.W. but possibly not native in this State),A. rubida Cunn. (Qld, N.S.W., Vrc.), A. steedmaniiMaiden & Blakely (W.A.), A. subulataBonpl. (N.S.W.), A. vq.lid.inerviaMaidelt'& Blakely (W.A., N.T., S.A.) andA. wattsiana F. Muell. ex Benth. (5.4.). Acacia bineryata DC. (N.S.W., Qld) and A. wardellii Tind- (Qld) appearto be closelyrelated to certainmembers ofthis Group (e.g.A. bancroflii) on account of their seedshaving encircling funicles, however,they arereadily distinguishedby their 2-3-nerved '(HJUAd '.tr'N OI I "I[I/.I'>D 'fer'rq8rg I L8 {a4los y y'3u1I1,119op uq g ureqrrop leerg uo 3edelllu gg I :(HJuad) l0I 'uolduequoN 'C) ,s1141 uu,(a 3 g Jo r\\ uq gZ , :(HIUAd ZL6l ulFDn y.g ue8uol6 ol peol eql uo EruqtrdJoSrul8 r:(HJUAd'lN'.Ight')'SNVJ) /l0Z ut\soltJy.g,luf,pooJ spr€,rol3urll"urooD 'a^reseu ,a ,CXO ,.IAn ,){ ,D Jo .,!\S uq 8 8 :(HJdAd) yy79 ntldog q g s{rod uo3€rc :0[ 'htt 'd 'CXO 'CINn-I.)'htg) l6Z puE(,4N ZttTpuoltwue T,r€^ru ue^\S:(HJUAd.SNVJ) Z60Z .A SulwunC V'proisuuSeg spre,nolrequrnSol lJo I[ [uDJZ 9Z] se|ru g.9I :(HIUA d) VVgl SulwtunJ 'xnop€J ,.IAht .) ,SNVJ) .H.S spJe^\ol urre^\o( Jo N lrul 9 t] elnu I :(HJUgd g1g undv J 'rueqclDllu,{lA .pautwora Jo N [uDl Z e] selru Z :VI.jVdJSnV NdAJSAA suawlJadspapaps

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'I {uclle.srwrDrry'urlsel I dg 188 NuytsiaVol.10, No.2 (1995) B,I

Varietion. Specimensfrom nearNorthampton have pods which are a little more consffictedbetween 1 the seedsthan normal but otherwisethey seemtypical of this variant.

Variant 2 fili (rn Bcrt dark red throughoutor only on upper branches.Branchlets sometimespruinose. Phyllodes int narrowly oblanceolate,5-10 cm long, 6-20 mm wide, lengthto width ratio 5-15, staight or sometimes pu very shallowly recurved, glaucous to subglaucous. Gland wually 2-10 mm above pulvinus, gl: occasionallyto 20 mm or at distal end of pulvinus. Heads globular or oblongoid, 6-8 mm diam. at ve anthesis,(26)30-45-fl owered. Le ad ru Distribution. Most common fiom Cowcowing (c. 20 km north-northeastof Wyalkatchem)southeast to near Meffedin, but also occurring further southeastto the Hyden area. 2a Habitat. Loam, sand or clay, sometimeswith laterite in low woodland or open shrubland.

Selectedspecimens examined. WESTERN AUSTRALIA: I 1.6 km E of Merredin on Great Eastern M Highway to Kalgoorlie, N. HallHT6/41(PERTH); 8.5 miles [13.6 km] E ofHyden on the Lake Varley ,.ll road, B.R. Maslin 535 (AD, PERTH); 4.5 miles [c. 7 km] W of Rabbit Proof Fence No. 1 on the Narembeento SouthernCross Road, B. R. Maslin 573 (PER'IH, NSW, NT,); 10 km N of Wyalkatchem towards Koorda, B.R. Maslin 4103 (CANB, K, MEL, PERTH); Cowcowing, E. Wittwer I21l (PERTH). \l P Affinities. This variant resemblesVariant 2 of A. chamaeLeon(see below for discussion). It may also P rcsembleA. jennerae which is distinguishedby its broader pods (6-8 mm wide), glabrous to sub- glabrous racemeaxes and peduncles,frequently obviously acuminatephyllodes and smaller seeds,

II Variant 3 r( b Phyllodes broadly linear to narrowly oblanceolate,8-13 cm long, 4-8 mm wide, length to width b incurved,green ratio 12-27,usually shallowly and uniformly to a subglaucous.Gland l0-3O(50) mm lt pulvinus. globular, golden,5-6 above Heads deep mm diam. at anthesis,18-30-flowered. Pods and n JeedJnot seen. tr t Distribution. Occurs mainly in the Wubin-Pithara-Kalanniearea (near Dalwallinu), but also north c to near Morawa. I e Habitat. Usually in loam, sometimessand or clay in shrubland. t ( Selectedspecimens examined. WESTERN AUSTR{IA: alongrabbit-proof fence, 15 miles [24 km] t NNW of Kalannie, T.E.H. ApLin 543 (MO, NSW, PERTH); 6 km N of Wubin towards Latham, l B.R. Maslin 4131 (CANB, K, MEL, PERTH): 10.5 km SW of Pithara on the road to Miling, B.R.Mqslin 6128 (PERTH);Gutha, KW. McLean,5 April 1934(PERTH 00861693); 35 km E of Perenjori, A.A. MitcheLl 1532 (PERTH); Buntine R€serve, 10.5 km N of Wubin, B.G. Muir 188 I (PERTH). I

'lhrs , Affinitie& variant is closely related to A. chrysella, especially the form that occurs between Wyalkatchem and Holt Rock (c. 60 km southeastof Hyden), which is distinguishedby its shorter, narrower phyllodes (5-7 cm long and 1.5-3 mm wide). -l

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tI ,{trellatsrl trrrty'uIIsnI U'g NuytsiaVol. 10, No. 2 (1995)

usually sparseand confined to exfemity of rachides,base ebracteate;peduncles 4-6 (rarely 9) mm AT long, solitary or occasionallytwinned, sparselyto denselyt appressedpuberulous with pale yellow A. to white hairs which aresometimes confined to the upperhalfofthe peduncle;bas aL peduncular bracts dir 2, tdangular, c. 0.5 mn long, brown, glabrousor ciliolate; heads globular,4-5 mm diam. just prior to anthesis,7-8 mm diam. at anthesis(fresh),5.5-7 mm diam. (dry), light to medium golden, Vc 23-37-floweredbut mostly 25-35, the flowers not very densely 4rranged;bracteoles spathulate to sub- VA peltate, l-l.5 mm long; claws linear but dilated at apex, glabrous to glabrescent;laminae broadly sh ovate,0.4-0.6 mm diam., minutely fimbriolate (hairsgolden or white) otherwiseglabrous, light brown when dry. F/o"€/"f, 5-merous;caLyx l/2-213 length of corolla, gamosepalous,divided for I/6_l/4 its length into broadly triangular,not or very slightly thickened,sparsely hairy, slightly inflexed lobes; Vr calyx tube glabrous and not obviously neryed:petals 1.5-2 mn long, joine

Distribution. southwest western Australia, confined to the southwestpart of the Roe Botanical District and the southernextremity of the adjacentAvon BotanicalDistrict (l:250,000 maps di 150-7,8,11).Restdcted to a small area boundedby Broomehill,Nyabing 1c. 60 km northeastof dt Broomehill)and Jerramungup. pl ol Hqbitqt. Grey-brown, (gravelly) clay or loam over clay in eucalypt shrubland. c(

Biology. A last-growing pioneer specieswhich regenerateswell from seedfollowing fire and also along disturbed road verges. It has a lifb-span of l0-15 years(K. Newbey, pers.comm.).

Flowering and fruiting periods. Flowering specimens coll€cted throughout the year with a predominancein May and December. Pods with matureseeds collected in December.

Conservqtionstatrs. Not consideredrare or endangered. J u Etymology. The epithetis derived fiom the Greekword - chamaileon(alizard that is changeablewith 0 respectto its colours) and alludes to the phyllodes which are very variable in shapeand size. A, Typirtcaion. The type collection of A. stowardii at BM consistsof both flowering and fruiting d specimens. Although theseundoubtedly represent the sametaxon it seemsprobable that they were b gatheredfrom different plants. The lectotypeis one of the two flowering specimens. 0 p rl '(tsn8ny-,(py{ e1e1)porred 3uuds7:e1ur,reqt ur srseqlu€r{Jeor luql speeqpro8uolqo:o .le1nqo13pue selcunpedslr uo s:ruqmo11e,{ eled ro ueplo8 ,slelqcuulqesournJd 'sepoll,iqd .V sellllteuos snocnelSqnso1 snocnelS seq sJlawntq Jo 71uurru6 (,{renrqag-JeqlueJec) JelrJtunsur srseqtuerlcueJ qcrq,u speeq J€lnqolS puu selcunpedpeJruq-elrq,r ,{1pnsn ,slelqcuurq '(sueurceds esoulnrd ,{rp urory po8pnl Jnoloc) sepollfqd uaer8 ,{yqerunserds1t ,(q paqstn8uqsrp sI lnq (e^oq" ees) stlownJq y Jo Z luEr.rDApelnqrlsrp ,{y]aqpoueroru eql Eullqurasau salJrul.fy

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Funlle3srtllrrrDry'uqspl\l g tl d G6 Nuyrsiavol.10, No.2 (1995)

-lhe Note. type of A. leiophylla var. microcephala Meissnerseems retbrable to this entity. br' ler Variant 3 ml alt sli Phyllodes filiform, l-2 mm wide, drying sub-tereteto a quadrangular. na fil Dist bution. Found mostly around Nyabing. Ot Selectedspecimens examined. WESTERN AUSTRALIA: 3.3 miles [5.3 km] N of Nyabing Hotel towards Kukerin, R.J. Cumming 2697 (CANB, K, PERTH); 11 miles [17.6 km] SE of Nyabing, 2( K.R. Newbey 744 and 744D (both PERTH); 12 miles [19.2 km] E of Ongerup, K.R. Newbey 3003 (PERTH); 24 miles kml NE of Ongerup, K Newbey 3663 (CANB, K, PERTH); 5 km S of [38.5 D 21858 (AD, BM, C, MEL, NY, PERTH). Broomehill afong main road, A. Sffid (1

Note. This variant includes the type of A. stowardii S.Moorc. F, 4. Acacia dorsenna Maslin, sp. nov. pl tc Frnrex densustholifbrmis glaber usquead circa 1.5 m altum. PhyLlodiainaequilateraliter elliptica ad obovatos,margine supernovalde convexo,rotundato et mucronulatocum mucroneplus minusve laterali,10- l5 mm longaet 5-9 mm lata,subcarnosa la€via subglauca demum hebeti-viridia, costa et D nervislateralibus nullis vel obscuris,nervis marginalibus brunneolis. Glans parva,supra pulvinam 0.5-2 mm. RacemiI-2.5 cm longi,capitulis 7-l0juventuteabracteolis imbricatis parvis inclusi sed saepeultra crescentibu cum pedunculisultimis a phyllodiissecondariis subtentis; peduncuLi 4-16 mm TT fongi graciles; capitula globllart^ vivide medio-aurea, 15-21-floribus, bracteolis nullis. Flores 5-meri; sepalis discretis. Legumine (immatura) angusteoblonga, suprasemina secus medio-lineam conspicue rotundata alternatim in uterque lato, usque 6 cm longa et I I mm lata, firme chartacea A brrnnea. Semina (immatura) transversaliaoblongo-elliptica, usque ad 4 mm longa, vix arillata. h A c ?yprs: north of Norseman [precise locality withheld tbr conservationreasons], 3[ August 1986, B.R.MasLin 6112 (holo: PERTH 00769096;lso: CANB, K, MEL, NY). tr A Dense, domed, glabrous sftrabs to c. [.5 m tall and to 3 m diam. Bar& smooth, light grey. Branchlets terete, slightly angled at extremities,finely ribbed, ribs not resinous,light brownish to yellowish greenat extremities,cuticle often grey-white. Strplles early caducous,linear, to 2 mm long, t' scarious. Phyllodes inaequilaterallyelliptic to oboyate,abaxial margin more or less straight, upper f margin convex and abruptly roundedat apiceswith a a lateral,minute (0.3-0.4 rnm long), innocuous, r mm long, 5-9 mm wide, lengthto width ratio 1.5-2,sub-fleshy, smooth brown mucro, 10-15 but I sometimesvery finely wrinkled when dry, somewhatcrowded, subglaucousand slightly pruinose at I ends of branchlets, ageing dull green; midrib very obscure and situated near abaxial margin, sometimesa less prominent secondnerve diverging fiom near the pulvinus, lateral nervesabsent or yellow-brown pulvinus few and obscure,marginal nerveslight brown to and t shiny; 0.5 mm long, ( finely wrinkled when dty. Gland notprominent,situated on uppermargin of phyllode0.5-2 mrn above ( pulvinus, c. 0.5 mm long. Recemesconfined to upper axils, l-2.5 cm long with 7- 10 heads,enclosed when very young by small, scarious,commonly caducous,brown bracts,often growing out at anthesis I with secondaryphyllodes subtendingthe peduncles;rqceme axes slender and sl'la;tlht,peduncles I 4-16 mm long, slender; basal peduncular brqcts absent(normally replacedby minute resin hairs); lzeadsglobular, 9 mm diam.at anthesis(fresh), 5-6 mmdiam. (dry), bright mid-golden,15-21-flowered; -l

: 'serceds ,neu sql ol peteleJ,{lasolc ,(lprcedse 1ou eru 1nqezts puu adeqsepog(r,ld ut ouuastop y elqueseJos|r uoq oantnfin! .y puu .qtueg xe uunC .V I .orcnur s slwlolu n) y'sarreds uprpllsnv uJols€eeql Jo srlroJ euos luo8und + u qlr,r sepollrqd 'seuel"J 'pecnpor pue p€pEaq-E-I ,{leureJlxe,slrry e8u"Jo.ro ,.,,\ollo,{snonctdsuoc ,spod palroc Jo T po^Jnc'lpuls slr ,{qpeqsrnAurlsrp sr qcrq,r 1nqsarceds,neu eqlgo s8u€] I"crqdur8oa8eql urqlr.,rlsrncco .V ,: qclq,t IIantrl g lIlD aw y Jo sruJoletuos selquoseJouuastop ez.rspuv edeqsepo11,(qd u1 J 'pod eql ur [eurpnlfuol 'epr,r -ocuo ,speoq ll speaspu€ rutu 9-t puu pelroc-ecr^\tJo spod g-Z qll^\ Suol ruur t-[> setueo?J 'epo1l,{qd ,(t-Z 'ept,,n ll aqtgo elppru eqt ?^oqr .roJ"eu pue13lue8und + orcnru ollel qlpr,r ot q13ua1 urur 't ,esocrure^ g-g'3uo1urur /I-L) Jelpurssepo11,{qd Jo prcsl^,{pq311s ue1;o slelqcuurq :sfu,r Bur,lnollo; .ri eql ut Duuayop y ruo4 peqsrnSurlsrp{1peer sr opo1:toldwocotcocy .ctqedu(s Je,\o eru o,r} aql roqleq,{\ur ou{ tou sr 111nq setceds ,neu oqt go uor8et eq1ur s,ror8 qc rllr.opolco\clwoc .y lcraua8 o go uro; apo11,{qde8rel u ro1 uelelsru oq ty?tw ttuua op .y ,{11ercr;redng e1u11r:u,{gue1c ere selarunreql tnq secuecsalouurJulrurrs e^uq ,,{elpur.loalnoozqns y puc .llanl/\tr.go11&qdoay y 'serlp slr pue pllrlA (ltrutg) sua|oatonso?rrry .el€llrl?xeselcrunJ puE (snoecvlJllqc ,{1pnsn) pernlxel ^lulql spod seloetcElq,serJ slEdes ,stol?lq ,3uno{ ,pesolJuo luesqu u^\oJqsnouDrs ,{q uaq,r eJDqctq,r ,9 seueJEJuorls ur seruecsaJouur:Sur,rolloJ oql epnlcul sarcodssno.rqElS esaql ^q pa.tuqsslelJereqc .d .V luulrodurr eJou eqJ ueprylNllultlrd y pup qluag sap waq V.s,reJpuv DpDIJoJdwD) '(e^oq"ees)urlsewotaoqJorltuD'Vep\curdnor9aq1.;osrequtot{.,,dnoJDlll./112"/dDlcD)V,,"gtsBe]rqU.J ol perels: ,{lpruJojut sl rlrrq,,$rxel pelcla.r,{1eso1c 1o dnor8 u o1s8uoleq nauasvop orlt2Jv .saltlulgv ua tu 'sepo11,{qd SA eql Jo urBlPru raddn pepuno: ,{llueuruord eql ol Surpnllp ,pelcaqdunq - snuuastoputlu.l eql uoJC .t\olow(jg UI p€ 'enssr tul srql.lopuJ aJS lsrJ c.roll {tllolrd puuJJtru parclJrc l3 s,tueura3uuu141puul puBuone^losuoJ .Iolueu DdeCeql uo uoxul ,{grrorr4y .snw\s uolj,^J?suoC 1 eA 'reque^oN-prur ec ur pelcellof, ueeq wq spaoseJnluu ,{11un:udqlr^,\ ueurceds r puu requalde5 e1e1o1 .seraods,reu lsnBny e1u1uror; palcelloa uaaqe^"q sueturJedsSuue^ oU,Je^e,roH srqllo ,{Bolouaqd .spouad eq1ssesseflelu.rnccu ot llncr rptr salrru suoqcelJocIo ,ftrcned eq1 3ut11w!puD Buuaaog

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I ,-

194 Nuytsiavol. 10,No. 2 (1995) B.R.

5, Acacia gelasina Maslin sp. nov. Dist toa Frutex glabet 1-2.5 m altus. Phyllodia oblanceolataad lineari-oblanceolataversus basem It is angustataobtusa mucronata,4-10 cm longa et (3)5-8(10) nm lata, coriaceasubrigida erectaglauca exte ad subglauca,costa et nervis marginalibus luteis, nervis lateralibus obscuris. Glans obsaura sed plerumque supra pulvinam l-3 mm et etiam mucronisad basemornata. Racemi (l)2-20 mmlongil. HaL pedunculis 1-2 cm longis; capitula globularia dense 35-50-floribus. Flores 5-meri; sepalis plus minusve discretis, aliquando 2 ve 3 connatis. Legumina anguste oblonga plana, supra semina FIo rotundataet inter seminaleviter constricta,usque l5 cm longa et 12-14 mm lata, firme chartaceaad coll tenuiter codacea,plerumque luteo-brunnea. Semina longrtu.dinalia circularia ad late elliptica vel late ovata,7-8 mm longa, aliquantumcompressa leviter lucida atro-brunneaad nigra sed in centro leviter Cor depressapallida, exarillata. De(

Typus: 46.5 km north of Murchison River, North West CoastalHighway, WesternAustralia, 23 July Etr (iolo: 1972,A.M. Ashby 4638 PERTH 00992844; iso: CANB, PERTH 00167975). sha Dense, spreading,shrubs 7-2.5 m tall, much-branchedat ground level. Bark smooth or finely Atr) fissured,grey. BranchLetstercte, ^ngl€d at€xtremities,finely ribbed,glabrous, reddish or light brown, A.L yellow on new growth. Stlpalesinsignificant, <0.5 mmlong, early caducous.PhylLodes oblanceolate isa to linear-oblanceolate,narrowed at base,obtuse, mucronate, 4-10 cm long, (3)5-8(10)mm wide, inc = I:w (6)8-15(25),thickly coriaceous,sub-rigid, erect, straight, glabrous, glaucous to subglaucous, sitr midrib and marginal nerves yellow, lateral nerves not prominent, two nerves on upper margin an( coalescingjust abovethe gland; pulvinus 0.5- I mm long,dilated at base,smooth or with a few coarse rhn wrinkles. G/andsnot prominent,situated on upper marginof phyllodenormally l-3 mm above Ac' pufvinus,usually a secondgland at baseof mucro. RacemesI or 2 per axil, (l)2-20 mm long with rts (l)2-6 heads,rarely growing out; racemeaxes glabrous,lightly pruinose,base ebracteatebut rec is present,peduncLes l-2 cm long, rarely few occasionally a solitary supra-basalbract a twinned, DI' yorng, peduncular persistent gfabrous,lightly pruinosewhen basaL bract or deciduous,0.5-l mm pel long, brown, thickened at base,solitary, occasionallyon uppermostpeduncles replaced by a pair of stipules with prophyll arising between them, the prophyll presumably expands with age; heads 6. gfobufar,densely 35-50 tlowered;bracteoLes 0.5-l mm long, claws linearand ciliolate,abruptly expandedinto a dark brown, cornmonly inflexed, sparselyciliolate lamina which is thickened in its lower half. Flowers 5-merous;sepals usually variablewithin a single flower, free or sometimes2 or Le 3 united,spathulate to oblong, l/3- l/2 lengthof petnls,petals 2.5-3 mm long, fiee, glabrous,nerveless. t8 Pods narrowly oblong, flat but obviously roundedover seeds,convexities rugulose and not extending to the margins, shallowly constrictedbetween seeds, occasionally deeply constdcted,to l5 cm long, to l2- 14 mm wide, up to l0 seedsper pod, pendulous,firmly chartaceousto thinly coriaceous,straight 1n to slightly curved, sometirnessparingly and obscurelytransversely nerved, glabrous, yellow-brown, di: sometimesnid-brown, commonly lightly pruinose,marginal nerves c. I mm wide. Seed.rlongitudinal pa in the pod, circular to widely elliptic or widely ovate,7-8 mm long, 6-7 mm wide, somewhatflattened IO (c. 3 rnm thick), slightly shiny, dark brown to blackish exceptfor the shallow, circular depressionin centre of seedwhich is usually yellow or light brown; plearogram obscure,open at hilar end; areole widely to transverselyelliptic, l-1.5 mm wide;/unlcle filiform, c. 5 rnm long, exarillate. pr n( Other specimensexamined. WESTERN AUSTRALIA: type locality, A.M. Ashby 3874 (K, MEL, R PERTH) and 19 November 1972, s.n. (PERTH 00993298,CANB: fruiting collection fiom the type plant); 15 km N of Murchison River on North West CoastalHighway, H. Demarz 10490 (PERTH); 48.5 km N of Murchison River on North West CoastalHighway, B.R. MasLin2786 (NSW, PERTH); id 48 km S ofBillabong Roadhouse,B.R. Maslln 4330 (PERTH);45 km N ofMurchison River on North D West CoastalHighway, B.R. Maslin 3346(BRI, G, NY, PERTH). 'urtuoq Sur,requr 1r se[quesel r{Jrq,{\pue atfiq V qlrt cr:ledurfs seurDeruossr qcrq,$ sercods€ suqndnt y sr lt pw g qJaqlE 116 ssord lo ueurcedsu peuruexe e^€q I eJnsunse,tr eq ,{tquepl 'rol)uJeqtuegftauoq esoq,^\Jo It6ssar./Jyo ueurcedse o1pe.neleJ(ilr yrapun uorssncsrpslq uI

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iI {uclle.sq,,{,rrrJy'urrset\' U g 196 Nuytsiavol. 10,No. 2 (1995) linear, l-nerved phyllodes and racemose inflorescences. Acacia cupuLaris,however, is readily distinguishedtiom A. harveyi by rts glabrous(not appressedpuberulous) raceme axes, golden heads SPT and its crustaceous,t erectpodscontaining seeds with shortfunicles and scarletarils (podscoriaceous brr and pendulousin A. hameyi, the long funicle l/2-3/4 encirclesthe seedin a single fold and ends in rnC a cream-white arrl). Acacia cupularis is describedand illustrated in Chapman& Maslin (1992). str ca{ 7, Acacia l€ptopetalaBenth., Linnaea 26: 619 (1855) cor Ph c. LectoD)pe(here selected):south-western Australia, J. Drummond 52 (K, upper righthand specimen c, on herb. Hooker sheet:,isolecto: BM, K - other specimenson lectotypesheet, MEL, OXF, P, PERTH sul OO'163144and 01941O70-fragmentex MEL and 01941O89-fiagment ex K). mi mt Typification. There are fbur specimensmounted on the lectotypesheet ,/ A. leptopetalaat Kew , ^11 "52 sli representingthe species and apparently part of the same collection. The sheet is labelled Pn J. Drummond. S.W. Australia 1850" lthis datepresumably being when the specimenwas receivedat "Herbarium "Acacia 1-, Kewl, stamped Hookerianum 1867" and annotated leptopetalq", in pencil, by "Swan SO Bentham. Another sheetat K is labelled River (Suppl. to 5th coll.) Drummond", is stamped Pe "Herbarium Benthamianum1854" and supportsa single specimenofA. Ieptopetalawhich Bentham "cf. br has annotated LeptopetaLa".As the phyllode dimensionson this specimenslightly exceedthose br given in the protologue,I do not regard it as a type ofA. Leptopetala. ab sn The type ofA. Ieptopetalawas prcsumablycollected on Drummond's 1847 expeditionto Mounts cl Caroline and Stirling which are situatedabout 15 km south of the present-daytownship of th "Supp. Kellerberrin. The specimenscollected on this trip were sentto Hooker and subscribersas to ar 5th Coll." (seeErickson 1969). During the journey liom Toodyay(where Drummond lived) he would stl most probably havecrossed the quite limited geographicrange ofthe conspicuouslypruinose element lo of A. Ieptopet(rlawhich accordswell with the type (i.e. the Bungalla-Kel)erberrinarea). m c. "Murchison Nomenclature. The river, Oldfield' specimencited by Bentham (1864: 370) under tn A. IeptopetaLais A. mut.ryana. This error led Bentham( I 875: 472) and Mueller ( 1887)to misapply the name A. leptepetala to A. murrayona and Maiden (1916: 495) to regard the two names as synonymous. ol A 8. Acacia sclerocladaMaslin, sp. nov. 0 N Fmlex aliquantum effusus et diffusus 0.5-1 m altus, ramulis +/- rectis et ascendentibusstdato- H costatisglabris subglaucis,interdum grossespinosis. PhyLlodiadistantia anguste linearia ad lineari- 0 elliptica, ad basemet apicemangustata, pungentia, 2-7 cm longa et plerumque1-3 mm lata, subrigida E ascendentiaad erectarecta vel incurvataglabra subglauca5-nervata, costa paulo prominenti et plus 0 minusve excentrica, margine supero binervato et leviter incrassato. Racemi l-20(30) mm longi capitulis 1-4, praecocijuventute a bracteolisimbricatis inclusi sedaliquando ultra crescentibu,axibus L glabris;pedunculisT - 12 mm longisglabris, capitula globdaria vivide aurea25-41-floribusl bracteolis F nullis. Flores plerumque5-meri; sepalisplus minusvediscretis. Legumina supra seminarotundata ( et inter semina leviter ad profunde constricta,usque [ 1.5 cm longa et 4-6 mrn lata, coriaceaglabra brunnea. Seminalongitudinalia elliptica, 5-6 nrn longa,turgida lucida atro-brunneaarillo longitudine T l/2-2l3 seminum partes aequantia. q I ?ypns: 9.6 km E of Edahtowards Mount Magnet,Western Australia, 1 August 1974,B.R. Mctslin36lS (holo: PERTH 00168939:lsor CANB, G, K, MEL, NSW, NY). I

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tI {uEllarsrt{ ,?Jrry'u!sEt{ U g G66 NuytsiaVol. 10,No. 2 (1995)

Flowering ond fruiting periods. Flowers tiom mid-June to sarly September. The single iruiting SIn specimenwas collected in December. wrl atl lor Conservqtion,rt l/rs. Not known to be under threat. slil a1 - - Etymology. The nam€ is derived from the Greek skleros,totgh or hard, and kkLdos t:ranch. les (1) A|finities. Acacia scleroc lada is perhapsmost closely allied to A. wiseonqC. Gardneron accountof fai the fbllowing characterswhich areshared by the two species:foliage rathersparse; branchlets glabrous pe, and striate; phyllodes distant, 5-nerved with the upper two close together and forming a slightly anl thick€ned upper margin; racemesshort and sometines resemblingthe branchlets;peduncle bases (re ebracteate(at least at anthesis);heads globular; sepalst fiee; bracteolesabsent; pods long; seeds cla longitudinal. Acacia wiseane is more widely distributedthan the new species(see Maslin & Pedley att 1982)and is clearlydistinguished in the followingways: it is a larger,intricate shrub to 4 m tall and spi hasdivaricate, light greenbranchlets; phyllodes 5-20 mm long, 1-1 .5 mm wide and patentto deflexed; pods6-12 mm wide; seedsglobose,5-9 mm diam. fla co The generalfhcies ofA. scleroclqda often resemblesthat of the typical variant ofA. inaequiLobL gk "spinifex" W. Fitzg. which grows on sand in country betweenQueen Victoria Spring and Streich lol Mound (well to the southeastof the rangeof the new species,c. 200 km east-northeastof Kalgoorlie). sli This variant of A. inaequiloba is most readily distinguishedfiom the new speciesin the fbllowing of ways: branchletsless prominently ribbed and t pruinose;upper margin of the phyllodes2-nerved cIr\ below the gland and l-nerved above it (2-nervedthroughout in A. scLerocLada);inUorescences comprisingvery short,l$eaded, racemessubtended by conspicuous,more or lesspersistent bracts; Ot headsI2-21-flowered with caducousbracteoles that are conspicuousin the buds;calyx inegularly 8., dissected;pods undulate; seeds c. 3 mm long and mottledblack and brown. 50 (P 9. Acacia subrigida Maslin,sp. nov. Di Frutex DSq\ead 2.5 m altum, ramulis glabris. PhyLlodia saepesectione rhombica subsessilia mi mucroneindurato atro-brunneorecto et erecto,8- l5 om longaet l- 1.5mm lata, planasubrigida erecta arr plusminusve recta ad leviterincurvata glabra viridia ad subglauca4-nervata sed nervo adaxiali prope M glandem basalembilurcato, nervis aureis. 6lans suprapulvinam perdiminutam3-17 mm et etiam ad basem mucronis glans obscura. Racemi plerumqte 2-10 mm longi capitulis 2-4, axibus glabris; Hr pedunculi 1-2 cm longi glabres;capitula globularia aurea l5-30-floribus, Flores S-meti sepalis on discretiset lineari-spathulatis.Legumina anguste oblonga, supra semina elevata, 6-11 cm longaet 7-10 mm lata, chartaceaglabra diluto-brunnea. .9?nir?alongitudinalia usque ad longitudinaliter FI obliquaovata, aliquando oblonga vel elliptica,5.5-7 mm longa,in centrovadose depressa, paulo lucida co nigra exarillata. AJ. Typus: 20.5 km E of Mukinbudin towardsBullfinch, WestemAustalia, l3 October 1975,B. R. Maslin A. 3965 (holo: PERTH 00167444; iso: CANB, C, K, MEL, NSW, NY). ph ol Erect shrubs to 2.5 m tall, single-stemmedor with 2-4 main stemsfrom ground level. Bark grey, ph fissured on main stems, smooth on branches. Branchlets glabrous,not pruinose, terete but often an slightly angledat the usuallybrown to red-brownextremities, penultimate branchlets usually invested AI with a light grey cuticle which becomeslongitudinally fissured and flaking with age;ribs often yellow, pc fine, scarcelyevident on penultimatebranchlets. Stipzles caducous. PhylLodesrhombic in section, ad rarely flat, 8 15 cm long but interspersedwith a few shorterones (to c. 5 cm long), 1-1.5 mm wide, Al

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tI {urllecsrll ,rrrr y'ullsrtll { B 200 Nuytsiavol. 10,No. 2 (1995) rhombic in section. Although the phyllodesof A. pachyacra arenormally:t rhombicthey arenarrower Ti and lessrigid than thoseof A. subrigida. Also, upon drying the phyllodesofA. pachyacra arc slightly longitudinally wrinkled or grooved whereas in A. subrigida they are pedectly smooth. Acacia subrigidais more southerlydistributed than eitherA. murrayana or A. pachyacra exceptfor the Shark Bay variant refened to below which occurs in the samegeneral region as A. murrayana, Acacia gelasina Mashn (see above) andA. praelongata F. Muell. fiom Northern Tenitory are also closely Br "Acacia related to the above taxa and togetherthese five speciescomprise the informal murralana Group". This Group is related to the'A. victoriae Grotp" (see Maslin 1992) but most readily distinguishedby non-spinosestipules and peduncleswhich lack a supra-basalbract. Ph and Land Conserv(ltionstatus. A Priority 2 taxon on the Departmentof Conservation Management's -li Declared Rare and Priority Flora List. Seeend of this issue.

Etymology. The epithet is derived tiom the Latin sabrigidus andreters to the textureof the phyllodes -s which is one of the charactersdistinguishing the new speciesfrom its close relatives,A. pachyacra and A. murrqyqna.

Voriation. A variant fiom Shark Bay and known only from the thrce collectionscited below has not been included in the above description. Thesespecimens were in late flower and one (8.R. MasLin -s 3682) possessedold pods and seedscollected liom the ground under the plant. Better material and additionalfield studiesare required in orderto ascertainth€ statusofthis variant. It diflers liom typical A. subrigidain th€ following ways. PhyLLodesI l-21 cm long, 2 mm wide, t flat but the midrib prominently |aised (and sometimesrendering the phyllodestransversely rhombic in section), occasionallywith afbw, obscure,longitudinally trendinglateral nerves (observ e when dry). Racemes (4-7)9-30(35-40)rnm Iong with (2)3-6(9)pedunculate heads, sometimes the peduncle fails to develop Ba within axil of lowermost bract on racemeaxes; peduncLes sometimes subtended by a minute phyllode at anthesis;heads 24-39-flowered, l0 mm diam. at anthesis(reconstituted), claws of bracteoles ancJ ,repal,rsparsely puberulous. Pod.r c. l3mmwide. Seelswidely oblong-elliptic, 8 mm long,7mm wide; p Ieurogram almost continuous. Nt

Specimensexamined. WESTERN AUSTRALIA: c. 6 krn W of Overlander-Denhamroad towards Pr Tamala, Shark Bay area, B.R. MasLin 3682 (CANB, K, MEL, PERTH); c. 9 km fiom Denham- Overlanderroad towards Tamala, B.R. Maslin 3714 (BRI, CANB, K, MEL, NSW, PERTH);7 km W Se of Denham-Overlanderroad on road to Tamala Station, B.R. Maslin 6296 (PERTH). p0

Distribution and haritar. WesternAustralia in the northernextremity ofthe Irwin Botanical District (l;250,000 map G49-12). Known only fron ths Shafk Bay area(south of Carnarvon)f'rom the collectionscited above. Occurson red-brownsandy loam in tall shrubland. t) isr 10. Acacia wilcoxii Maslin,sp. nov.

Frutex glaberper Iamosus2-4 m altus,habito Templetoniueegerce simili, rarnulisascendentibus Bt ad erectosteretibus, obscurenervatis, juventute diluto-viridibus sed aetatecinereis. saepe spinosis. ul Phyllodia ramulos simulans teretia acuta grossepungentia, plerumque 5-14 cm longa et 1.5-2 mm ol dianetro, crassa erecta, juventute diluto viridibus sed aetate cinereis, per obscure 4-nervatis. Pedunculi 5.25 mm longi, plerumque geminati, alteruter in racemis vel phyllodiorum in axillis in c. surculis novisi capltula globularia24-27 floribus. Flores 5-mer1,calyce gamosepalo et plus minusve trwcato. Legumina linearia, usquead 10.5 cm longa et 5 run lata, tenuiter coriaceaatro-brunnea. Semtna (leviter immatura) longitudinalia late elliptica usque late ovata, circa 4 mm longa, arillo a diluto-luteo. qsltollef se^rau)pe,lreu-y ,{1a:ncsqo ,(le^ 'alp:orJetepsller sr qsqlq,nSururnllnq u,{oJq uor8el parde '(oJcntu eql tdetxe slelqcu€Jquo sEJnoloc ldecxe) snorqelS1q3rc4s ssel lo erotu ,treJa ,,{lp ueq,,[ '{clql 'pl8u 'qloous ' '8uol 'elqeu€^ 'as€q 'e]elel epluq ruerpruut Z-9 1 urc tI -9(g) q13ua[ lE pe,ror.reu i 's1e1qcue:q 'u^\oJq'snorJEcs 'qlSuel ,3uo1 .t eqt 3ur1qst"s"Jsaponkl d JreqtJo lsoru JoJpesn; ruu g.y I 'snocnppc,{llea '(elcrlnr Frat sa1nd1ry BuDIEUue+o'qsDtq,l.r € ol enp) qsr{ar8 Sure8e1nq 3uno,4 ueq,,r 'sno:qr13'pe^Jeu 'eteJel's}urod uaar8epd ,4leJncsqo,{ls^ tue8undflasJEoc ^ts^ ut Surteullurelue*o l 'pr8u 'esonxeg '1cera leq,$euos ,{1tq8r1s,(ra,r o1 lq8re4s o1 SurpuecsusplqJuDJg .seqcuu:qreddn uo qsru^\oJqpuu r1toolxsSururoceq 'saqcueJq urcu pue $lunJl uo peueq8no:,{pq8r1s pue f,efi t1,ng 'oua8a ,ltEt .peqcuEJq-r{cnt{ oruolaldwal Sulquesar llquq w V-Z squqs pepunor ssel Jo eJou

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esourrud rtI asournrd (i(lpe{rEur,{larEr) DUt ,tlpo{reur ua$O ,{JturuJseu4euros esournrdloN slalqcuBJfl {le 012 eue,(e.r.rnru'y e.nu,tqrud'y 'Y BplBlrqns {lB D!Ji .y ,(}uEuB^ otmtottnw V puc-ono{IlJDd ,iD! 'y .1 ,fug lJ€qS ar;1Surpnlcxe) oprBuqns SulprnEupsrpsernlee; I€crSoloqdrou pdrcuu4 eyqe; Je,{1

tI,{ueuecsq,r,rry'ullset{ U'g NuytsiaVol. 10, No. 2 (1995) B.F and not or barelyraised); pulvinus c.2mmlong, slightly dilated at base,usually orange-brown;dPices col on young phyllodes variably hooked and possessinga glabrousor sparselyhairy mucro c. 1 mm long (tht with a glandJike structureat its baseon the upper surface,mucro lost with age resulting in straight, tw( coarselypungent apices(the gland-like structurerarely obseryableon matue Phyllodes). Gland on spe upper surfaceofphyllode to 3 mm aboYepulvinus, circular to oblong,c. 0.5 mm long, a minute gland- A. like structure at base of mucro (see above). Peduncles5-25.mm long, commonly twinned, either lari 2-5 pairs in determinateracemes l-3 cm long or axillary and arising synchronouslywith young phyllodes on new shoots,glabrous, base ebracteate; rac eme a&s with 4-5 minute caducousbracts at its base, sometimes growing out at apex; heads globular, 24-27-flowered; receptqcle papillose; bracteoles widely spathulate,c. 1 mm long, concave,sub-sessiie. Flowers 5-merous,glabrous; calyx 2/5 length of corolla, gamosepalous,very shallowly divided (for c. 1/6 or lessits length),into broadly triangular, non-thickenedlobes); p€fa/s c. 2 mm long, nerveless.Pods linear, to 10.5 cm long, 5 mm wide, with up to 9 seedsper pod, roundedover seedsand slightly to moderatelyconstricted between them, thinly coriaceous,more or less straight,neneless, glabrous,dark brown, margins nanow and yellowish. Seedr (slightly immature) longitudinal with aril facing apex of pod, widely elliptic to widely ovate,c. 4 mm long and 3.5 mm wide, brown; pleurogr.tm open athilar endi areoLec, 0.7 mm Bei long and 0.5 mm wide;funicle straightto shallowly curved,pale-yellow, reflexed abovea somewhat Ber thickened aril of the samecolour. Ber Bft Other specimensexanired. WESTERN AUSTRALIA (preciselocalities withheld for conseryation Bn reasons):Erong SpringsStation, R.J. Cranfield 5347 (PERTH); TangadeeStation, B.R. MasLin5287 (CBC, BRI, K, PERTH) and B.R. Maslfu 5288 (CANB, G, MEL, NSW, PERTH), A A. Mitchell26S (PERTH); Dalgety Downs Station,A.A. MitchelL 1552 (CANB, PERTH).

Distribution. Western Australia in the Ashburton Botanical District (l:250,000 maps G50-4, 6). Known only fiom Tangadee,Landor, Erong Springs and Dalgety Downs Stations,c. 250 km north to northwest of Meekatharra. Eri M! Hqbitat. Grows along watercoursesand adjacentstony plains and low stony hills, in open or closed scrub.

Flowering and fruiting periods. The few specimensto hand have been collected August and September. Slightly immature pods have beencollected in early October.

Conseryqtionstatus. A Priority I taxon on the DepartmentofConservation and Land Management's Declared Rare and Priority Flora List. Seeend of this issue. M Etymology. Named in honour of David G. Wilcox, lbrmerly of the WesternAustralian Department M of Agdcultur€, who has collected extensively since 1952 in the pastoral shrubland zone of W.A. M betweenthe Ashburton River and the Nullarbor Plain. Pe Pe Affinities. Acacia wilcorii is\ery distinctiyeby its thick, terete,relatively long, erectphyllodes which closely resemblethe branchletsand is not likely to be confusedwith any otherAcacia within its known resembleboth A. erocarpoidexW. Fitzg. and A. microcaly:r geographic range. It does, however, Ti Maslin in habit and in having coarselypungent, ascending to erect branchletswhich are frequently investedwith a whitish flaking cuticle. Thesetwo relatedspecies occur southand west of A. wilcoxii (seeMaslin and Pedley 1982). Although A. exocerpoideshas Erete phyllodesthey are much shorter than those of A. wiLcoxii, i.e. 8-l8 mm long, and unlike the new speciesare frequently shed upon

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