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Nakedstem Sunray (Enceliopsis Nudicaulis [A. Gray] A. Nelson)
NAKEDSTEM SUNRAY Enceliopsis nudicaulis (A. Gray) A. Nelson Asteraceae – Aster family Corey L. Gucker and Nancy L. Shaw | 2020 ORGANIZATION NOMENCLATURE Nakedstem sunray (Enceliopsis nudicaulis) (A. Names, subtaxa, chromosome number(s), hybridization. Gray) A. Nelson belongs to the Ecliptinae subtribe and Heliantheae tribe within the Asteraceae family. Nomenclature follows Welsh et al. (2016). NRCS Plant Code. ENNU (USDA NRCS 2019). Range, habitat, plant associations, elevation, soils. Synonyms. Encelia nudicaulis A. Gray Common Names. Nakedstem sunray, Ash Life form, morphology, distinguishing characteristics, reproduction. Meadows sunray, nakedstem, naked-stemmed daisy, sunray (USFWS 1983; Curtis 2006; Welsh et al. 2016; USDA NRCS 2019). Growth rate, successional status, disturbance ecology, importance to Subtaxa. Some systematists (Cronquist 1972; animals/people. Welsh et al. 2016) recognize nakedstem sunray varieties: corrugata and bairdii, although others do not support varietal distinctions (Sanders and Current or potential uses in restoration. Clark 1987; Curtis 2006). Chromosome Number. Reported chromosome numbers include: 2n = 32, 34, and 36; but most Seed sourcing, wildland seed collection, seed cleaning, storage, consistently reported are 2n = 34 and 36 (Reveal testing and marketing standards. and Styer 1974; Hickman 1993; Curtis 2006; Welsh et al. 2016). Recommendations/guidelines for producing seed. Hybridization. The possibility of hybrids within the Enceliopsis genus have been suggested based on DNA evidence. Sequencing of two nuclear and two chloroplast regions suggests that a single Recommendations/guidelines for producing planting stock. plant collected from co-occurences of nakedstem sunray and Panamint daisy (E. covillei) in California may represent a hybrid or backcrossed individual (Fehlberg and Ranker 2007). Hybrids of nakedstem Recommendations/guidelines, wildland restoration successes/ failures. -
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report
Annotated Checklist of the Vascular Plant Flora of Grand Canyon-Parashant National Monument Phase II Report By Dr. Terri Hildebrand Southern Utah University, Cedar City, UT and Dr. Walter Fertig Moenave Botanical Consulting, Kanab, UT Colorado Plateau Cooperative Ecosystems Studies Unit Agreement # H1200-09-0005 1 May 2012 Prepared for Grand Canyon-Parashant National Monument Southern Utah University National Park Service Mojave Network TABLE OF CONTENTS Page # Introduction . 4 Study Area . 6 History and Setting . 6 Geology and Associated Ecoregions . 6 Soils and Climate . 7 Vegetation . 10 Previous Botanical Studies . 11 Methods . 17 Results . 21 Discussion . 28 Conclusions . 32 Acknowledgments . 33 Literature Cited . 34 Figures Figure 1. Location of Grand Canyon-Parashant National Monument in northern Arizona . 5 Figure 2. Ecoregions and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 8 Figure 3. Soil types and 2010-2011 collection sites in Grand Canyon-Parashant National Monument in northern Arizona . 9 Figure 4. Increase in the number of plant taxa confirmed as present in Grand Canyon- Parashant National Monument by decade, 1900-2011 . 13 Figure 5. Southern Utah University students enrolled in the 2010 Plant Anatomy and Diversity course that collected during the 30 August 2010 experiential learning event . 18 Figure 6. 2010-2011 collection sites and transportation routes in Grand Canyon-Parashant National Monument in northern Arizona . 22 2 TABLE OF CONTENTS Page # Tables Table 1. Chronology of plant-collecting efforts at Grand Canyon-Parashant National Monument . 14 Table 2. Data fields in the annotated checklist of the flora of Grand Canyon-Parashant National Monument (Appendices A, B, C, and D) . -
Bibliography
ASH ME A DOWS REFE R ENCES by Kathleen Capels and Terence Yorks, High Level Research http://www.hlresearch.org/ in cooperation with Frank J. Smith, Western Ecological Services http://ecocorridors.com/western-ecological.php Note: all hyperlinks were checked for validity in late July 2009. Some may require entering the desired genera or local site name into a search space on the linked page to generate appropriate information. Abrams, Leroy, and Roxana Stinchfield Ferris. 1960. Bignoniaceae to Compositae: Begonias to Sunflowers. Vol. 4 of Illustrated Flora of the Pacific States: Washington, Oregon, and California. Stanford, CA: Stanford University Press. [Enceliopsis] Baldwin, Bruce G., Steve Boyd, Barbara J. Ertter, Robert W. Patterson, Thomas J. Rosatti, and Dieter H. Wilken, eds. 2002. The Jepson Desert Manual: Vascular Plants of Southeasten California. Berkeley and Los Angeles: University of California Press. [Centaurium, Ivesia, Nitrophila] Barneby, R[upert] C. 1970 [published 1971]. A new Astragalus (Fabaceae) from Nevada. Madroño 20(8):395–398. [Astragalus] Beacham, Walter, Frank V. Castronova, and Suzanne Sessine. 2000. Dicots. Vol. 4 of Beacham’s Guide to the Endangered Species of North America. Detroit: Gale Group. [Nitrophila] Beatley, Janice C. 1969. Vascular Plants of the Nevada Test Site, Nellis Air Force Range, and Ash Meadows (Northern Mojave and Southern Great Basin Deserts, South-Central Nevada). Los Angeles: University of California Laboratory of Nuclear Medicine and Radiation Biology. [Astragalus, Centaurium, Cordylanthus, Enceliopsis, Grindelia, Ivesia, Mentzelia, Nitrophila, Spiranthes] ———. 1970. Additions to Vascular Plants of the Nevada Test Site, Nellis Air Force Range, and Ash Meadows. Los Angeles: University of California Laboratory of Nuclear Medicine and Radiation Biology. -
Clark County Multiple Species Habitat Conservation Plan Amendment Covered Species Analysis Report
Clark County Multiple Species Habitat Conservation Plan Amendment Covered Species Analysis Report Prepared For: Clark County Department of Air Quality Desert Conservation Program 4701 West Russell Blvd., Suite 200 Las Vegas, NV 89118 WRA Contact: Ken Sanchez (415) 578-3184 [email protected] Patricia Valcarcel (415) 524-7542 [email protected] Date: June 11, 2018 WRA Project: 26346 2169-G East Francisco Blvd., San Rafael, CA 94702 (415) 454-8868 tel [email protected] www.wra-ca.com Draft Covered Species Analysis Report – June 2018 This page intentionally left blank. Draft Covered Species Analysis Report – June 2018 TABLE OF CONTENTS 1.0 INTRODUCTION.................................................................................................................. 1 2.0 SPECIES REVISION PROCESS ......................................................................................... 2 2.1 Species Considered for Coverage ............................................................................ 2 2.2 Criteria for Covered Species ..................................................................................... 3 3.0 ANALYSIS ........................................................................................................................... 4 3.1 Species Range ......................................................................................................... 4 3.2 Species Status.......................................................................................................... 4 3.3 Impacts from Covered Activities .............................................................................. -
Annotated Checklist of Vascular Flora, Arches National Park
National Park Service U.S. Department of the Interior Natural Resource Program Center Annotated Checklist of Vascular Flora Arches National Park Natural Resource Technical Report NPS/NCPN/NRTR—2009/220 ON THE COVER Double Arch, Arches National Park, Utah. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Arches National Park Natural Resource Technical Report NPS/NCPN/NRTR—2009/220 Authors Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741 Sarah Topp Northern Colorado Plateau Network National Park Service P.O. Box 848 Moab, UT 84532 Mary Moran Southeast Utah Group National Park Service P.O. Box 907 Moab, UT 84532 Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network National Park Service P.O. Box 848 Moab, UT 84532 June 2009 U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The National Park Service, Natural Resource Program Center publishes a range of reports that address natural resource topics of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, con- servation and environmental constituencies, and the public. The Natural Resource Technical Report Series is used to disseminate results of scientifi c stud- ies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service mission. The series provides contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. All manuscripts in the series receive the appropriate level of peer review to ensure that the in- formation is scientifi cally credible, technically accurate, appropriately written for the intended audience, and designed and published in a professional manner. -
Inventory of Vascular Plants at Mojave National Preserve & Manzanar Historic Site
Inventory of Vascular Plants at Mojave National Preserve & Manzanar Historic Site For U.S. National Park Service Inventory and Monitoring Program Camissonia boothii ssp. boothii Prepared by James M. André University of California Riverside, Granite Mountains Desert Research Center November 15, 2006 Contract Number: P2128020178 TABLE OF CONTENTS 1.0 INTRODUCTION 1.1 National Inventory and Monitoring Program and Mojave Network 1 1.2 Program Overview 1 1.3 Regional Context and Significance 2 1.3.1 Mojave National Preserve 2 1.3.2 Manzanar Historic Site 4 1.4 Objectives 5 2.0 METHODS 2.1 Herbarium Surveys 7 2.1.1 Database and Bibliographic Query 7 2.2 Field Surveys 7 2.2.1 Mojave National Preserve 7 2.2.2 Manzanar Historic Site 8 2.3 Voucher Specimens 8 2.4 Survey Timing 9 2.5 Field Investigators 9 3.0 RESULTS 3.1 Level of Effort 10 3.2 Summary of Findings - Manzanar Historic Site 10 3.3 Summary of Findings - Mojave National Preserve 14 3.3.1 Field Surveys 14 3.3.2 New Vascular Plant Taxa for MOJA 14 3.3.3 Special-Status Plants 33 3.3.4 Non-Native Alien Plants 40 4.0 DISCUSSION 4.1 Summary 41 4.2 Recommendations 41 Acknowledgements 43 5.0 REFERENCES 44 i List of Maps, Tables and Appendices Map 1. Distribution of all Area Searches in Priority Locations, Targeted Surveys, and Opportunistic Surveys conducted in the Mojave National Preserve, 2002 – 2005. 15 Table 1. Checklist of vascular plant taxa known to occur in the Manzanar Historic Site. -
101 Appendix 3. Federally Listed Threatened and Endangered
101 Appendix 3. Federally listed Threatened and Endangered species in the family Asteraceae that occur in the continental United States.1 Regions Listing of Scientific Name Common Name Current Range Species tested Status2 Occur- rence3 SUBFAMILY CICHORIOIDEAE Tribe Cardueae Cirsium fontinale var. fountain thistle CA E 8 Cirsium fontinale var. fontinale fontinale Cirsium fontinale var. Chorro Creek bog thistle CA E 8 obispoense Cirsium hydrophilum var. Suisun thistle CA E 8 Cirsium hydrophilum var. hydrophilum vaseyi Cirsium loncholepis La Graciosa thistle CA E 8 Cirsium loncholepis Cirsium pitcheri Pitcher's thistle IL, IN, MI, WI, T 3 Cirsium brevistylum, Canada (Ont.) C. ciliolatum, C. cymosum, C. occidentale var. venustum Cirsium vinaceum Sacramento Mountains NM T 2 Cirsium vinaceum thistle Tribe Lactuceae Malacothrix indecora Santa Cruz Island CA E 8 Agoseris grandiflora malacothrix Malacothrix squalida island malacothrix CA E 8 Stephanomeria Malheur wire-lettuce OR E 1 Stephanomeria cichoriacea malheurensis Taraxacum californicum California taraxacum CA E 8 Lactuca sativa Tribe Vernonieae Vernonia proctorii (no common name) PR E 4 Stokesia laevis 102 Appendix 3 Scientific Name Common Name Current Range Status2 Region3 Species tested SUBFAMILY ASTEROIDEAE Tribe Anthemideae Artemisia campestris ssp. northern wormwood OR, WA C 1 Artemisia californica borealis var. wormskioldii Tribe Astereae Baccharis vanessae Encinitas baccharis CA T 8 Symphyotrichum (=Aster) chilense Boltonia decurrens decurrent false aster IL, MO T 3 Chrysopsis floridana -
Astragalus Phoenix Barneby
Astragalus phoenix Barneby Status: ASH MEADOWS MILKVETCH Global/State ranks G2, S2 Fabaceae NNHP: At-risk list NNPS: Threatened State: Critically endangered BLM: Sensitive USFWS: Threatened Habitat: Seasonally moist to dry flats, knolls and washes with alkaline calcareous soils. 671 - 716 meters (2,201 - 2,349 feet) Counties: Nye Life Form: Dicot perennial herb Bloom period: February - May © James. M. Andre © James. M. Andre Nevada Natural Heritage Program Rare Plant Fact Sheet CalPhotos Database Results NNHP Taxon Report SEINet Taxon Information UNR Herbarium Records © Karen Tanner © Karen Tanner © Sarah Kulpa Last revised 08/21/2015 Astragalus phoenix Barneby An older plant with living tissue restricted to the periphery on one side Fresh and drying flowers © Karen Tanner Aseed pod after seed dispersal Ayoung plant © Karen Tanner © Karen Tanner Old herbivory - clipped inflorescence Fresh herbivory - tissue may retain stems are dry and gray-white in color. a greenish cast and will be pliable. Old tissues are stiff and prone to Leaves may also be clipped by snapping off when moved. herbivores. © Karen Tanner © Karen Tanner Adead individual © Karen Tanner © Karen Tanner © Karen Tanner © Karen Tanner Last revised 08/21/2015 Plant associates: Atriplex confertifolia, Distichlis spicata, Enceliopsis nudicaulis corrugata, Haplopappus acradenius, Mentzelia leucophylla Jepson eFlora Treatment (Astragalus genus only; A. phoenix not listed): Martin F. Wojciechowski & Richard Spellenberg 2012. Astragalus, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi-bin/get_IJM.pl?tid=54612, accessed on Jan 24 2014. Reprinted with permission from the Jepson Herbarium, UC Berkeley. Note that these treatments and keys describe California taxa and may not be a perfect match for Nevada material. -
Enceliopsis Nudicaulis Var. Corrugata (Ash Meadows Sunray)
Enceliopsis nudicaulis var. corrugata (Ash Meadows sunray) 5-Year Review: Summary and Evaluation Photo by Sarah Kulpa, U.S. Fish and Wildlife Service U.S. Fish and Wildlife Service Nevada Fish and Wildlife Office Reno, Nevada September 2011 5-YEAR REVIEW Enceliopsis nudicaulis var. corrugata (Ash Meadows sunray) I. GENERAL INFORMATION Purpose of 5-Year Reviews: The U.S. Fish and Wildlife Service (Service) is required by section 4(c)(2) of the Endangered Species Act (Act) to conduct a status review of each listed species at least once every 5 years. The purpose of a 5-year review is to evaluate whether or not the species’ status has changed since it was listed (or since the most recent 5-year review). Based on the 5-year review, we recommend whether the species should be removed from the list of endangered and threatened species, be changed in status from endangered to threatened, or be changed in status from threatened to endangered. Our original listing of a species as endangered or threatened is based on the existence of threats attributable to one or more of the five threat factors described in section 4(a)(1) of the Act, and we must consider these same five factors in any subsequent consideration of reclassification or delisting of a species. In the 5-year review, we consider the best available scientific and commercial data on the species, and focus on new information available since the species was listed or last reviewed. If we recommend a change in listing status based on the results of the 5-year review, we must propose to do so through a separate rule-making process defined in the Act that includes public review and comment. -
Ash Meadows National Wildlife Refuge Plants I Perhaps Owe Having Become a Painter to Flowers
U.S. Fish & Wildlife Service Ash Meadows National Wildlife Refuge Plants I perhaps owe having become a painter to flowers. -Claude Monet Introduction Plant Guide Key Welcome Ash Meadows National Wildlife Refuge Symbols Refuge location where plant can be Refuge, established in 1984, is the most commonly seen: largest oasis in the Mojave Desert, supporting an incredible diversity of Crystal Spring boardwalk plants and wildlife year-round. Over 24,000 acres of alkali seeps, springs Point of Rocks boardwalk and other unique habitats make Ash Meadows a biological wonder for Longstreet boardwalk everyone to enjoy and protect. Peterson Reservoir Enjoying the With such rare habitats, Ash Devils Hole Refuge's Plants Meadows has some of the most unique plant communities in the world. Unlike some desert areas where flowers bloom simultaneously Conservation * an Ash Meadows endemic plant in spectacular spring-time displays, Status Symbols (only grows in the Ash Meadows the blooms at Ash Meadows are much area) more subtle and span the entire year. † an endangered species Natural fluctuations in weather can ‡ a threatened species affect bloom times as well. In any ▲ a non-native or invasive species season, something is blooming for insects and birds to eat, and people to admire. Look carefully and tread lightly—you will be amazed by what you discover! Alkali mariposa lily. Cyndi Souza/USFWS Using this Plant This list includes over 40 of the Guide approximately 340 plant species at Ash Meadows. Each of the plants listed are used by culturally affiliated American Indians for both food and medicines. These tribes have revealed select information and caution visitors not to collect, eat or prepare plants, possibly harming an individual or the land. -
Arctomecon Merriamii Coville
Arctomecon merriamii Coville Status: WHITE BEAR POPPY Global/State ranks G3, S3 Papaveraceae NNHP: At-risk list NNPS: Watch list State: unlisted BLM: Sensitive USFWS: unlisted Habitat: Basic soils ranging from dry to moist; alkaline sand and clay, gypsum, outcrops of carbonate rock, calcareous alluvial gravels. 610 - 1,585 meters (2000 - 5,200 feet) © Amber Swanson Counties: California Native Plant Clark, Lincoln, Society Nye. Also in CA. Life Form: Dicot perennial herb Bloom period: April - May © James M. Andre Nevada Natural Heritage Program Rare Plant Fact Sheet Jepson eFlora Taxon page NNHP Taxon Report CalPhotos Database Results SEINet Taxon Information Consortium of California Herbaria Records UNR Herbarium Records © Sarah Kulpa © Duncan S. Bell Last revised 02/18/2014 Arctomecon merriamii Coville © Duncan S. Bell © Duncan S. Bell © Duncan S. Bell Jepson eFlora Treatment and Key: Gary L. Hannan & Curtis Clark 2012. Arctomecon, in Jepson Flora Project (eds.) Jepson eFlora, http://ucjeps.berkeley.edu/cgi- bin/get_IJM.pl?tid=13901, accessed on Jan 22 2014. Reprinted with permission from the Jepson Herbarium, UC Berkeley. Note that these treatments and keys describe California taxa and may not be a perfect match for Nevada material. Artomecon Perennial herb; hairs generally ± 5–15 mm; taproot stout; sap colorless or yellow. Leaf: generally basal, obovate or wedge- shaped, long white-hairy; teeth rounded. Inflorescence: terminal, 1-flowered. Flower: bud nodding; sepals 2–3, long-hairy; petals 4 or 6, free, obovate, white [yellow], generally persistent after flower; © U.S. Fish & Wildlife Service stamens many, free; placentas 3–6, style 1, stigma lobes 3–6. Fruit: ovate to oblong, dehiscent from tip. -
Annotated Checklist of Vascular Flora Dinosaur National Monument
National Park Service U.S. Department of the Interior Natural Resource Program Center Annotated Checklist of Vascular Flora Dinosaur National Monument Natural Resource Technical Report NPS/NCPN/NRTR—2009/225 ON THE COVER Paria evening-primrose (Oenothera caespitosa var. navajoensis). Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Dinosaur National Monument Natural Resource Technical Report NPS/NCPN/NRTR—2009/225 Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741 Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532 July 2009 U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The National Park Service, Natural Resource Program Center publishes a range of reports that address natural resource topics of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, con- servation and environmental constituencies, and the public. The Natural Resource Technical Report Series is used to disseminate results of scientifi c stud- ies in the physical, biological, and social sciences for both the advancement of science and the achievement of the National Park Service mission. The series provides contributors with a forum for displaying comprehensive data that are often deleted from journals because of page limitations. All manuscripts in the series receive the appropriate level of peer review to ensure that the in- formation is scientifi cally credible, technically accurate, appropriately written for the intended audience, and designed and published in a professional manner. This report received informal peer review by subject-matter experts who were not directly involved in the collection, analy- sis, or reporting of the data.