Annotated Checklist of Vascular Flora, Arches National Park

Home , Achnatherum aridum, Achnatherum coronatum, Ageratina herbacea, Agoseris, Aliciella leptomeria, Allenrolfea

National Park Service U.S. Department of the Interior

Natural Resource Program Center

Annotated Checklist of Vascular Flora Arches National Park

Natural Resource Technical Report NPS/NCPN/NRTR—2009/220 ON THE COVER Double Arch, Arches National Park, Utah. Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Arches National Park

Natural Resource Technical Report NPS/NCPN/NRTR—2009/220

Authors Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741

Sarah Topp Northern Colorado Plateau Network National Park Service P.O. Box 848 Moab, UT 84532

Mary Moran Southeast Utah Group National Park Service P.O. Box 907 Moab, UT 84532

Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network National Park Service P.O. Box 848 Moab, UT 84532

June 2009

U.S. Department of the Interior National Park Service Natural Resource Program Center Fort Collins, Colorado The National Park Service, Natural Resource Program Center publishes a range of reports that address natural resource topics of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, conservation and environmental constituencies, and the public.

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All manuscripts in the series receive the appropriate level of peer review to ensure that the information is scientiﬁ cally credible, technically accurate, appropriately written for the intended audience, and designed and published in a professional manner. This report received informal peer review by subject-matter experts who were not directly involved in the collection, analy- sis, or reporting of the data.

Views, statements, ﬁ ndings, conclusions, recommendations, and data in this report are those of the author(s) and do not necessarily reﬂ ect views and policies of the National Park Service, U.S. Department of the Interior. Mention of trade names or commercial products does not constitute endorsement or recommendation for use by the National Park Service.

This report is available from the Northern Colorado Plateau Network website, http://www. nature.nps.gov/im/units/NCPN, as well as at the Natural Resource Publications Management web site, http://www.nature.nps.gov/publications/NRPM.

Please cite this publication as:

Fertig, W., S. Topp, and M. Moran. 2009. Annotated checklist of vascular ﬂ ora: Arches Nation- al Park. Natural Resource Technical Report NPS/NCPN/NRTR—2009/220. National Park Service, Fort Collins, Colorado.

NPS 100065, June 2009 ii Annotated Checklist of Vascular Flora: Arches National Park Contents

Figures ...... v Tables ...... v Abstract ...... vii Acknowledgements ...... ix

1 Introduction ...... 1 1.1 Background and justiﬁ cation ...... 1 1.2 Objectives ...... 1 1.3 Study area ...... 1 1.3.1 History and setting ...... 1 1.3.2 Geology ...... 3 1.3.3 Vegetation ...... 3 1.3.4 Previous botanical studies ...... 4

2 Methods ...... 7

3 Results ...... 9 3.1 Summary of the ﬂ ora of Arches National Park ...... 9 3.2 Life form and biogeographic diversity ...... 10 3.3 Plants of special concern ...... 10 3.4 Non-native and noxious weeds ...... 10 4 Discussion ...... 15

5 Literature Cited ...... 17

Appendices ...... 21 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes...... 23 Appendix B. Potential (unconfi rmed) taxa of Arches National Park...... 75 Appendix C. Rejected (falsely reported or questionable) taxa of Arches National Park...... 85 Appendix D. Confi rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes...... 91

Contents iii

Figures

Figure 1.3.1. Location of Arches National Park, Grand County, Utah...... 2 Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Arches National Park, 1940– 2006...... 6

Tables

Table 1.3.4. Chronology of plant-collecting efforts in Arches National Park...... 5 Table 3.1. Statistical summary of the ﬂ ora of Arches National Park...... 9 Table 3.3. Plant species of conservation concern from Arches National Park...... 11 Table 3.4. Noxious weed species of Arches National Park...... 13

Contents v

Abstract

In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Arches National Park. This entailed a thorough review of existing literature, re-examination of specimens from the Arches National Park herbarium and other state and national museum collections, and additional fi eld work to corroborate unverifi ed species reports or locate new species. Based on this study, 522 vascular plant taxa have been confi rmed or reliably reported as occuring in the park (485 are known from validated herbarium specimens and 37 from unvouchered literature reports). Another 50 species formerly considered part of the park fl ora have now been shown to be falsely reported. The current fl ora of Arches National Park represents 14.5% of the known fl ora of Utah, and 30.8% of the fl ora of the Colorado Plateau ecoregion within Utah. At least 30 of these species are considered species of concern by the state natural heritage program. Introduced plants make up 14.4% of the park fl ora (75 species), of which six species are considered noxious weeds by the State of Utah. Further surveys in Arches National Park are likely to fi nd additional new species (146 are known from similar habitats outside the park boundary), especially among late fl owering, weedy, or wetland taxa.

Contents vii

Acknowledgements

We would like to thank the following individuals for their assistance with this project: Dr. Angela Evenden, botanist and former leader of the Northern Colorado Plateau Network (NCPN) program in Moab, Utah, was an active proponent of this project and helped secure funding for ﬁ eld work and herbarium studies in 2006. Vicki Webster, curator of museum collections for the National Park Service Southeast Utah Group (SEUG), provided access to all mounted and unmounted herbarium specimens from Arches National Park, as well as work space within the museum vaults. Charlie Schelz, biologist with SEUG, provided research permits for work in 2005–2006, and shared his unmounted plant collections. Libby Nance, former database specialist with the NCPN, assisted with development and quality control of the park’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories on the park. Drs. Stanley Welsh and Duane Atwood provided access to herbarium collections at Brigham Young University. Dr. Michael Piep provided access to collections at the Utah State University (Intermountain) herbarium. Dr. Ron Hartman and Ernie Nelson provided access and specimen loans from the Rocky Mountain Herbarium at the University of Wyoming. Thanks to the technical reviewers (Dr. Angela Evenden, Dr. Dusty Perkins, and Libby Nance) for their comments and suggestions. To any others who we may have inadvertently forgotten, please accept our apologies (and thanks).

Research was conducted under permit # ARCH-2006-SCI-0007 for study number ARCH-00033. Funding was provided by the National Park Service through the University of Wyoming.

Contents ix

(Evenden et al. 2002). This eff ort has entailed re- 1 Introduction viewing existing species checklists, corroborating the identifi cation of specimens in park museum In 2006, the National Park Service (NPS) estab- and herbarium collections, locating additional lished a cooperative agreement with the Univer- specimen records from other museums, mining sity of Wyoming, through the Rocky Mountains relevant literature and datasets, and conducting Cooperative Ecosystem Studies Unit, to produce fi eld surveys to locate new taxa or confi rm un- a series of technical reports summarizing the vas- substantiated reports.1 All of these data have been cular plant fl ora of each of the 16 national park synthesized in NPSpecies, the park service’s stan- units of the Northern Colorado Plateau Network dardized database for documenting park status (NCPN). These reports are based on existing in- (present, probably present, unconfi rmed, falsely formation in the park service’s NPSpecies data- reported), abundance, residency, nativity, and base system and augmented by further data min- rarity. With the production of a technical series of ing, reviews of state and regional herbaria, and reports on the fl ora of each park, the NPSpecies new fi eld surveys to resolve the status of many re- data will now be available to a broader audience. ported and unconfi rmed species. The objective of these reports is to summarize the current state of knowledge of the fl ora of each park for the bene- 1.2 Objectives fi t of NPS biologists and interpretive staff , outside The objectives of this project were to: researchers, and the general public. The following report addresses the vascular fl ora of Arches Na- 1. Document the current status (present, tional Park, in southeastern Utah. Other NCPN reported, potential, or falsely reported) of all park units are addressed in separate reports. vascular plant species that have been cited as occurring in Arches National Park, based on a review of herbarium collections and past 1.1 Background and justifi cation literature. One of the primary missions of the NPS is “to conserve unimpaired the natural and cultural 2. Develop an annotated checklist of the fl ora resources and values of the national park sys- of Arches National Park, with supplemental tem for the enjoyment of this and future genera- information on taxonomic synonyms, dis- tions” (NPS 2000). Due to their high degree of tribution, within-park abundance, fl owering protection and distribution across a wide variety period, growth form, nativity, habitat, and of habitat types, the nation’s national park units sources for each record. make a signifi cant contribution to the protection of biological diversity (Noss and Cooperrider 1.3 Study area 1994). Unfortunately, the full contribution of the National Park System is not known, as many park 1.3.1 History and setting units have traditionally lacked comprehensive The slickrock country north of Moab, Utah (Fig- biological inventories or monitoring programs to ure 1.3.1) contains the world’s largest concentra- identify the composition and condition of their tion of natural stone arches, as well as numerous biota. Knowing the status of park fl ora and fauna rock spires, balanced rocks, windows, and other is critical for eff ective park management and for geologic features carved by the forces of ero- systematically identifying and fi lling gaps in the sion (Chronic 1988). In the late 1920s, local civic nation’s protected area network (Margules and leaders promoted the area for recognition as a Pressey 2000). national park to encourage tourism to what was then one of the most remote areas of the United To rectify this shortcoming, Congress passed the States. In April 1929, President Herbert Hoover National Parks Omnibus Management Act in invoked the Antiquities Act to establish Arches 1998, to provide guidance and funding for an in- National Monument. Over the ensuing decades, ventory and monitoring program across the entire the monument’s boundary was expanded on sev- National Park System. Since 2000, the NCPN In- eral occasions (most recently growing to 30,979 ventory and Monitoring Program has been devel- hectares, with the addition of the Lost Spring area oping and verifying species lists of vertebrates and in 1999) and, in 1971, Congress passed legislation vascular plants for each park unit in the network

1Taxa refer to all named species, subspecies, and varieties.

Chapter 1: Introduction 1 Arches National Park Utah

To 70

i e 191 ado R v r lor Co 128

Visitor Center

To Moab

Established April 12, 1929 Legend 1,219–1,585 m elevation (4,000–5,200 ft elevation) Intermittent streams Park boundary (NPS) 30,966 ha (76,519 ac) Perennial streams U.S. Bureau of Land Management Rivers Private 0123 Kilometers Paved roads State Unpaved roads 0123 Other state or local ownership Miles Unimproved/4WD roads The Nature Conservancy

MAP CREATED BY ANETH WIGHT, NCPN (DERIVED FROM EVENDEN ET AL. 2002)

Figure 1.3.1. Location of Arches National Park, Grand County, Utah.

2 Annotated Checklist of Vascular Flora: Arches National Park converting it to national park status (Evenden et plastic. Ultimately, the upwelling dome of the Par- al. 2002). adox layers breached the surface in the Salt Valley and Klondike Hills area (exposing gypsum de- In addition to its namesake 200 stone arches, posits), as overburden sediments were removed Arches National Park (NP) contains signifi cant by erosion and faulting. Gradually, cracks in the archaeological and cultural sites, wildlife habitat, faulted Entrada sandstone blocks became wider and representative examples of native Colorado through erosion, forming parallel rock fi ns. Some Plateau vegetation. The park is characterized by of these fi ns, in turn, were sculpted by wind- hot, dry summers and cool winters, with aver- borne grit into balanced rocks, or hollowed out to age annual precipitation of 202 mm (8 in). Peak form windows or stone arches (Doelling 2003).2 precipitation occurs in March and August. Mean The greatest concentration of arches is located annual temperature is 56° F, but can range from in the Windows and Devils Garden areas of the –16° F in winter to 112° F in mid-summer (Even- park. Most of the south end of the park consists den et al. 2002). of outcrops of Navajo Sandstone, with the Wing- ate exposed on the cliff s above the Colorado Riv- 1.3.2 Geology er. Extensive areas of the west and north ends of the park are covered by Quaternary wind-blown The geology of Arches NP consists of hundreds sand. Mancos Shale and Dakota Sandstone are of feet of late Paleozoic to Cretaceous-age sedi- exposed mostly in the Salt Wash–Cache Valley mentary layers deposited over a thick bed of salt Wash area (Doelling 2003). and gypsum of the Paradox Formation. These salt deposits originated during the Pennsylvanian Pe- riod (about 300 million years ago) through repeat- 1.3.3 Vegetation ed cycles of evaporation of shallow sea water in a Harrison and others (1964) recognized fi ve main landlocked coastal bay (Chronic 1988; Doelling vegetation types within Arches NP; additional data 2003). Over time, the salt beds were buried by san- on the vegetation of the park is available in Allan dy limestones of the late Pennsylvanian Honaker (1977) and the NCPN (in ed.). Rocky slopes of Trail Formation, sandstone rubble of the Permian washes and rims of sandstone ridges are typically Cutler Formation, fl oodplain and delta sandstones occupied by pygmy conifer woodlands of Utah and shales of the Triassic Moenkopi and Chinle juniper (Juniperus osteosperma) and two-needle formations, and sand-dune and stream deposits pinyon (Pinus edulis). Commonly associated spe- of the Jurassic Wingate, Kayenta, and Navajo for- cies include singleleaf ash (Fraxinus anomala), mations. In the mid-Jurassic, the Arches area was Gambel oak (Quercus gambelii), shinnery oak (Q. inundated by a series of sandstone deposits, in- welshii), and Utah serviceberry (Amelanchier uta- cluding the Dewey Bridge Member of the Carmel hensis). Juniper-pinyon forests may also occur in (sometimes included in the Entrada), the main valley bottoms, but usually are restricted to sites body of the Entrada, and the Moab member of with shallow, rocky soils. the Curtis Formation (Doelling 2003). During the late Jurassic, the area was covered by fl oodplain Upland valleys and sandy plains support exten- deposits of the Morrison Formation, followed by stive stands of blackbrush (Coleogyne ramosissi- Cretaceous beach (Dakota Sandstone) and deep ma), green ephedra (Ephedra viridis), and Indian marine shales (Mancos Formation). ricegrass (Stipa hymenoides), especially where soils are relatively shallow to bedrock. Areas with The signature arches and other unusual rock fea- deeper sandy soils may be locally dominated by tures of Arches NP occur primarily within the sparse grasslands or sand sagebrush (Artemisia Jurassic-age Entrada Sandstone and the Dewey fi lifolia). Basin big sagebrush communities are Bridge and Moab members of the Carmel and prevalent mostly on rocky toeslopes or on alluvial Curtis formations. These massive sandstones be- terraces in valley bottoms on moderately deep came cracked and fi ssured during the uplift of soils near the water table. Salt desert scrub, domi- the Salt Wash anticline along an old fault system. nated by shadscale (Atriplex confertifolia), Fre- The anticline formed from pressure exerted by mont’s mahonia (Mahonia fremontii), prince’s- the deep Paradox salt deposits, which, under the plume (Stanleya pinnata), and galleta (Hilaria weight of their overlying sediments, became more jamesii) occurs on outcrops of the Mancos Shale,

2 By contrast, stone bridges are formed by the erosion of moving water in a stream or wash.

Chapter 1: Introduction 3 Paradox, and Morrison formations (Harrison et The fi rst botanist to conduct extensive fi eld- al. 1964). work in Arches NM was Bertrand F. Harrison, of Brigham Young University (BYU). From 1941 Riparian vegetation occurs along the banks of pe- to 1952, Harrison made several collecting trips to rennial streams and the Colorado River, and may the monument and added at least 96 new species consist of Fremont cottonwood (Populus fremon- to the local fl ora (Table 1.3.4, Figure 1.3.4). Har- tii), yellow willow (Salix eriocephala var. watsonii), rison returned to Arches NM in 1963, with BYU coyote willow (S. exigua), Goodding’s black wil- colleagues Stan Welsh, Glen Moore, and others, low (S. gooddingii), cattails (Typha domingensis to conduct a systematic inventory of the fl ora. The and T. latifolia), sedges, rushes, and fi ve-stamen BYU team added 236 new species to the monu- tamarisk (Tamarix chinensis). A small, desert- ment checklist, more than tripling the size of the saltgrass (Distichlis spicata) wet meadow occurs documented fl ora (Harrison et al. 1964). in the vicinity of Wolfe Ranch on salt-encrusted soils (Harrison et al. 1964). Hanging gardens oc- Over the next three decades, collecting eff orts cur infrequently in shady alcoves associated with within the newly renamed Arches National Park wet seeps or springs, and are locally dominated were more sporadic. John Allan, a graduate stu- by Eastwood’s monkeyfl ower (Mimulus eastwoo- dent from BYU, added seven new species to the diae), cave primrose (Primula specuicola), alcove fl ora in 1972–1973, while conducting a study on columbine (Aquilegia micrantha), western posion the composition and distribution of the park’s ivy (Toxicodendron rydbergii), southern maiden- major vegetation types (Allan 1977). In the late hair fern (Adiantum capillus-veneris), and giant 1980s, Jayne Belnap and colleagues established helleborine orchid (Epipactis gigantea) (Harrison permanent vegetation transects within the park et al. 1964; Springer et al. 2006). and added nearly 40 new taxa to the fl ora (Table 1.3.4). Ken Heil and associates from San Juan 1.3.4 Previous botanical studies College conducted a rare plant and fl oristic survey of Colorado Plateau national parks from The earliest known botanical studies in the Arch- 1989 to 1991, and added 19 new species to the es NP area were conducted by pioneer western park checklist. Fowler and others (2007) added taxonomist Alice Eastwood, who collected the four additional species reports during a survey of type specimens of Wetherill’s cryptanth (Cryp- hanging-garden fl ora along the Colorado River. tantha wetherillii), slender cryptanth (C. tenuis), and creeping rush-pea (Caesalpinia repens) from Since 1997, the fl ora of Arches NP has increased the Courthouse Wash area, north of Moab, in by 79 taxa through the eff orts of Southeast Utah 1892. Given the remoteness of the future Arches Group (SEUG) biology staff and projects spon- NP site and lack of access roads, it is probable sored by the NCPN (Table 1.3.4, Figure 1.3.4). that Eastwood’s collections were made outside NPS biologists Charlie Schelz and Mary Moran the modern park boundary (Table 1.3.4). Per Axel have added 24 new species in the past decade Rydberg and Albert O. Garrett collected the types (Schelz et al. 2006). Dewey and Andersen (2005) of Eastwood’s amsonia (Amsonia eastwoodiana, reported four new noxious and invasive weed now considered a synonym of A. tomentosa var. species from their inventory of the park. Re- stenophylla) and Garrett’s saltbush (Atriplex becca Harms collected two new riparian species garrettii) from the vicinity of Moab in 1911, but while conducting a survey of seeps and springs again were probably beyond today’s park borders (Springer et al. 2006). Sarah Topp and Lisa Hahn (Welsh 1982). The earliest confi rmed collection added 32 new species through their eff orts with from Arches National Monument (NM) was in the NCPN vegetation mapping project (NCPN in 1940, when plant chemist O.A. Beath and Leslie ed.). Lastly, Walter Fertig added 17 new species in Goodding documented at least one milkvetch the course of revising the Arches NP checklist in specimen (crescent milkvetch, Astragalus am- 2006 (Topp 2007). phioxys var. amphioxys) as part of their research on selenium-bearing plants (Table 1.3.4).

4 Annotated Checklist of Vascular Flora: Arches National Park Table 1.3.4. Chronology of plant-collecting efforts in Arches National Park. # of new Year Collector(s) Comments taxa Alice Eastwood collected the types of Wetherill’s cryptanth (Cryptantha wetherillii), slender cryptanth (C. tenuis), and creeping rush-pea (Caesalpinia 1892 A. Eastwood 0 repens) from “Courthouse Wash, Grand County”, probably outside the future boundaries of Arches National Park. Per Axel Rydberg and Albert Garrett collected the types of Eastwood’s amsonia P. A. Rydberg and 1911 0 (Amsonia eastwoodiana) and Garrett’s saltbush (Atriplex garrettii) from the A. O. Garrett vicinity of Moab, probably outside the park. Beath and Goodding collected crescent milkvetch (Astragalus amphioxys var. O. A. Beath and L. 1940 1 amphioxys) in Arches National Monument as part of their work on seleniferous N. Goodding plants of western North America (specimen at NY). Brigham Young University (BYU) botanist Bertrand F. Harrison collected Mormon 1941 B. F. Harrison 1 needlegrass (Stipa arida) from the monument (specimen at BRY). B. F. Harrison and colleagues from BYU collected 80 new species for Arches 1947– National Monument over three successive years. C. V. Morton and P. A. Munz B. F. Harrison 80 1949 also collected specimens in 1949 for the New York Botanical Garden, but apparently did not add any new species to the monument fl ora. B. F. Harrison added 15 new species, including the paratype of Arnow’s ricegrass 1952 B. F. Harrison 15 (Stipa arnowiae) (originally determined as Stipa hymenoides, but later described as a new taxon by Welsh in Welsh et al. 2003). A. H. Holmgren Arthur Holmgren and Loran Anderson discovered the fi rst population of white- 1955 1 and L.C. Anderson stem blazingstar (Mentzelia albicaulis) for the monument. Curtis Hanson confi rmed the presence of Atriplex garrettii var. garrettii in Arches 1961 C. Hanson 1 National Monument (the type is from the Moab area). S. L. Welsh, G. Stan Welsh, B. F. Harrison, and colleagues from BYU conducted a thorough Moore, G. L. fl oristic survey of Arches National Monument, more than tripling the number of 1963 Pyrah, L. M. Pitts, 236 species known for the area. Harrison and others (1964) lists 320 species for the B. F. Harrison, and monument. G. Moore J. T. Howell and G. John Howell and Gordon True collected 3 new species for Arches National 1968 3 True Monument for the New York Botanical Garden. Stan Welsh collected the fi rst specimen of spike pappusgrass (Enneapogon 1970 S. L. Welsh 1 desvauxii) from Arches National Monument. John Allan, a BYU graduate student, documented at least 7 new species for 1972– J. Allan 7 Arches National Park in the course of describing and mapping vegetation types 1973 (Allan 1977). K. Forsythe, K. 1979– Kay Forsythe, Kathy Kitchell, and Steve Swanke made 19 collections for the Kitchell, and S. 9 1981 ARCH herbarium and added 9 new species to the park fl ora. Swanke Damian Fagan, Jayne Belnap, and Sonja Paspal collected more than 130 D. Fagan, J. 1984– specimens for the ARCH herbarium and added 37 new species to the park Belnap, and S. 37 1988 checklist while establishing permanent vegetation monitoring plots across Paspal Arches National Park. Ken Heil, Rich Fleming, and Leo Vermilion, of San Juan College, collected more 1989– K. Heil, R. Fleming, 19 than 60 specimens for the ARCH herbarium and documented 19 new species for 1992 and L. Vermilion the park. Fowler and others (2007) conducted a fl oristic survey of hanging gardens in J. Fowler and C. 2 (+ 2 1993 the Colorado River drainage and documented 4 new riparian species in Arches May reported) National Park. 1997– Charlie Schelz, biologist with the Southeast Utah Group national parks, added 4 C. Schelz 4 1998 new species to the Arches NP fl ora.

Chapter 1: Introduction 5 Table 1.3.4. Chronology of plant-collecting efforts in Arches National Park, cont. # of new Year Collector(s) Comments taxa Dewey and Andersen (2005) documented two new invasive weed species and K. Andersen, L. reported two additional taxa without vouchers. Lisa Hahn and Sarah Topp added 2002– Hahn, C. Schelz, R. 15 (+ 2 6 new species during vegetation mapping ﬁ eld work (NCPN in ed.). Charlie 2005 Harms, M. Moran, reported) Schelz and Mary Moran collected 5 new species. Rebecca Harms added two new and S. Topp taxa while conducting an inventory of seeps and springs (Springer et al. 2006). W. Fertig, S. Topp, 53 (+ 5 Walter Fertig, Sarah Topp, and Mary Moran added 53 new species and reported 2006 M. Moran reported) 5 additional new taxa (without vouchers) for the park (Topp 2007).

600

500

400

300

200

Number of vascular taxa 100

0 1940 1950 1960 1970 1980 1990 2000 2006

Year

Figure 1.3.4. Increase in the number of plant taxa confi rmed as present in Arches National Park, 1940–2006. Wittmann 2001). The annotations to these lists 2 Methods and label data from the ARCH herbarium review were then incorporated into the NPSpecies data- In July 2005, we reviewed all mounted specimens base for the park. in the Arches NP herbarium (ARCH), housed in the SEUG offi ce in Moab, Utah, to determine Work in 2006–2007 focused on confi rmation of whether each was correctly identifi ed to species, reported and potential species for the park fl ora subspecies, or variety following the nomencla- through fi eld surveys and review of collections ture of Welsh and others (2003). As necessary, we from state and regional herbaria. We made col- made corrections and annotations to each speci- lecting trips to Arches NP in the summer and fall men and to the voucher records in the park’s Au- of 2006 to target missing species, emphasizing tomated National Cataloging System (ANCS+) undersampled wetland, weedy, and late-fl ow- and NPSpecies database. ering taxa. Herbarium studies were undertaken at Brigham Young University (BRY), Utah State Based on this herbarium work and a review of University (USU), and the University of Wyoming park references (Harrison et al. 1964; Schelz and (RM) in March, June, and October 2006, and Feb- Moran 2005; Springer et al. 2006), we developed ruary and March 2007. We also documented park four lists: specimens from the Virtual Herbarium website of the New York Botanical Garden (http://sciweb. 1. A revised checklist of species that were pres- nybg.org/science2/hcol/intf/index.asp). Finally, ent in the park (confi rmed by an authenti- we found additional new species reports by ex- cated voucher; included in Appendices A amining unmounted collections by SEUG and and D); NCPN researchers and reviewing plot data from 2. A list of species that were reported in the lit- recent vegetation mapping and wetland invento- erature but not corroborated with a voucher ries (NCPN in ed.; Springer et al. 2006). (equivalent to “probably present” in NPSpe- cies terminology; included in Appendices A and D); 3. A list of potential species (“unconfi rmed” in NPSpecies) known from the vicinity of Arches NP using an unpublished list gener- ated by Dr. N. Duane Atwood of Brigham Young University and Albee and others (1988) (included in Appendix B); and 4. A list of species that are now known to be misidentifi ed or highly questionable (Ap- pendix C). Each list was annotated with supplemental information on life form, geographic range (within the context of Utah), population size in the park (derived from Harrison et al. 1964; NCPN in ed., and fi eld observations), fl owering period, distribution across general vegetation types, nativity, data source (collector and collection number from a sample specimen or literature citation), year of earliest collection (from herbarium records), and relevant synonymy and taxonomic problems (based on review of Barkworth et al. 2003, 2007; Barneby 1989; Cronquist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn 2001; Flora of North America Editorial Committee 1993, 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; Holmgren et al. 2005; and Weber and

Chapter 2: Methods 7

barium), Utah State University, the University of 3 Results Wyoming’s Rocky Mountain Herbarium, and the New York Botanical Garden Virtual Herbarium. 3.1 Summary of the fl ora of Arches Field surveys in 2005–2006 netted another 56 new species: 15 from SEUG botanist Mary Moran, 21 National Park by NCPN vegetation mapper Sarah Topp, 17 by Of the 623 vascular plant vouchers in the ARCH Walter Fertig, and 3 by other NPS staff (Topp herbarium that we examined in 2005 (excluding 2007). Overall, the documented fl ora of Arches 240 specimens on loan), 32 were misidentifi ed NP has increased by 120 species (24.7%), bring- (5.1%), 28 were revised to a more current name ing the total number of taxa to 485 (Table 3.1, Ap- (4.5%), 122 had variety or subspecies names add- pendices A, D). ed (19.6%), and 441 were found to be correctly determined (70.8%). Based solely on these col- Currently, 37 additional taxa have been reported lections, the confi rmed fl ora of Arches NP stood for Arches NP but have yet to be corroborated at 365 taxa (Fertig 2006). From 2005 to 2007, we with a voucher specimen (Table 3.1, Appendix located an additional 64 taxa from collections A). Of these, 28 were reported by Schelz and oth- at Brigham Young University (mostly species ers (2006) and 9 others came from observations reported in Harrison et al. 1964, but without a by recent vegetation-mapping (NCPN in ed.), duplicate voucher deposited in the ARCH her- wetland-survey (Springer et al. 2006), and weed-

Table 3.1. Statistical summary of the ﬂ ora of Arches National Park. Number Number Category conﬁ rmed additionally Total in park reported for park Taxonomic diversity Total taxa 485 37 522 (including varieties and subspecies) Full species 461 33 494 (excluding varieties and subspecies) Families 68 2 70 Life form diversity Tree taxa 16 0 16 Shrub taxa 72 5 77 Perennial forb taxa 203 21 224 Annual forb taxa 95 7 102 Perennial graminoid taxa 70 2 72 Annual graminoid taxa 21 1 22 Fern taxa 8 1 9 Biogeographic diversity Introduced taxa 68 7 75 Native taxa Locally endemic taxa 9 0 9 Regionally endemic taxa 65 7 72 Disjunct taxa 0 0 0 Peripheral taxa 8 0 8 Sparse taxa 7 0 7 Widespread taxa 328 23 351 Total native taxa 417 30 447 The number of taxa and families is based on taxonomic concepts of Welsh and others (2003). See appendices for an explanation of the various categories.

Chapter 3: Results 9 mapping crews (Dewey and Andersen 2005). but relatively species-poor, representing less than Taken together, the number of conﬁ rmed and 14% of the total ﬂ ora. The diversity of ferns and reported vascular plant taxa in Arches National fern-allies in Arches NP is especially low (1.7%) Park stands at 522 taxa (Table 3.1, Appendix A). (Table 3.1).

Entering the 2006 fi eld season, we recognized 229 Of the park’s 447 native taxa, 351 (63.4%) are taxa as potentially occurring in Arches NP, based widespread across Utah and western North on the proximity of the park to known popula- America. Only nine species (2.0% of the total na- tions (Albee et al. 1988; Fertig 2006). At least 10 tive fl ora) have extremely limited global distribu- of these species were found to occur in the park tions (local endemics found within an area of less based on fi eld collections or relocated herbarium than 1° of latitude × 2° of longitude; see appen- specimens in 2006. Another 73 potential species dices for complete defi nitions of biogeographic were eliminated based on their low probability of categories). Regional endemics (largely restricted occurring in the park. Most of these were species to the Colorado Plateau) comprise an additional of montane coniferous forest or wetland habitats 16.1% of the park’s native fl ora (Table 3.1). An- of the La Sal Range. The current number of po- other 15 taxa (3.4%) are sparsely distributed tential species that we are recognizing for Arches across the state or occur in the Arches area at the National Park is 146 (Appendix B). periphery of their continuous range.

Of the original pool of species documented or reported for Arches NP, we found that 37 taxa were 3.3 Plants of special concern falsely reported based on misidentifi ed voucher Arches NP has no plants listed as threatened or specimens in the ARCH and BRY herbaria or lit- endangered under the U.S. Endangered Species erature reports based on collections made outside Act. At least nine park taxa were formerly candi- of the park. We consider an additional 14 species dates for potential listing (Category 2 or 3C) un- cited (but not vouchered) by Schelz and others der the act (Table 3.3) (UDWR 1998). Schelz and (2006), Fowler and others (2007), and Springer Moran (2007) cited 11 species from Arches NP as and others (2006) to be questionable based on NPS “species of concern,” but three of these are the absence of suitable habitat or the distance of not presently known to occur within the park: Ise- Arches NP to the known or expected range of ly’s milkvetch (Astragalus isleyi), Cisco milkvetch each species (Appendix C). (A. sabulosus), and Trotter oreoxis (Oreoxis trotteri). The Utah Conservation Data Center lists The confi rmed and reported fl ora of Arches Na- 30 species from Arches NP as species of concern tional Park represents 14.5% of the 3,594 native (UDWR 1998). Although not currently cited on and naturalized taxa documented for the state of any agency list, the recently described Moab Utah by Welsh and others (2003) and 46.4% of pepperplant (Lepidium moabense) is a narrow the state’s 151 plant families (Table 3.1). Within endemic of southeastern Utah that may warrant Grand County, the park fl ora captures 44.2% attention as a species of concern due to its appar- of the 1,180 taxa reported by Welsh and others ently limited range and low numbers (Welsh et al. (2003). At the ecoregional scale, Arches NP con- 2003). It should be noted, however, that Moab tains 30.8% of the 1,694 species reported for the pepperplant is considered a synonym of Lepidi- Utah portion of the Colorado Plateau (Shultz et um eastwoodiae (or L. alyssoides var. eastwoodiae) al. 2006). by Holmgren and others (2005).

3.2 Life form and biogeographic 3.4 Non-native and noxious weeds diversity Introduced plant species (not historically native Although ecologically dominant, tree and shrub to Utah or North America) make up 14.4% of the species make up only 17.8% of the confi rmed confi rmed and reported fl ora of Arches NP (Table and reported plant taxa of Arches National Park 3.1). This fi gure is slightly higher than the state- (Table 3.1). Perennial forbs are the most species- wide average of 13.5% non-native species (Fertig rich life form in the park’s fl ora, with 224 known 2007; Welsh et al. 2003). Of the 75 introduced and reported taxa (42.9% of the total fl ora). An- plant species confi rmed or reported for Arches nual forbs and grass-like plants comprise less NP (Table 3.4), only 6 are listed as noxious by the than 24% of the fl ora. Like trees and shrubs, State of Utah (SUDAF 2008). Dewey and Ander- perennial graminoids are ecologically important sen (2005) mapped the distribution of 11 noxious

10 Annotated Checklist of Vascular Flora: Arches National Park and other invasive weed species within the park logeton glomeratus). Weed species are especially from 2003 to 2004, and found extensive popula- well-developed in upper Salt Wash and Salt Val- tions of ﬁ ve-stamen tamarisk (Tamarix chinensis), ley. Recent tamarisk control eﬀ orts have cleared Russian knapweed (Centaurea repens), Russian infestations from much of Courthouse Wash. olive (Elaeagnus angustifolia), and halogeton (Ha-

Table 3.3. Plant species of conservation concern from Arches National Park. TNC UTCDC Family Species Common name global Legal status status rank Boraginaceae Cryptantha long-fl ower G3 Watch, S2 longifl ora cryptanth Boraginaceae Cryptantha Wetherill’s cryptanth G3? Watch, S3? wetherillii (R) Cactaceae Opuntia large pricklypear G5T5 Peripheral, phaeacantha var. S2S3 major Cactaceae Opuntia polyacantha slenderspine G5 Peripheral, S2 var. trichophora pricklypear T3T5Q Chenopodiaceae Krascheninnikovia Ruin Park winterfat G5 Rare, S1? lanata var. ruinina T1?Q Compositae Cirsium rydbergii Rydberg’s thistle G3 USFWS: former 3C, SEUG: Watch, S2S3 (Asteraceae) Species of Concern Compositae Flaveria campestris marshweed G4 Peripheral, S1 (Asteraceae) Compositae Grindelia fastigiata erect gumweed G3 Watch, S2 (Asteraceae) Compositae Lygodesmia Entrada rush-pink G1Q UT BLM: Sensitive, USFWS: Rare, S1 (Asteraceae) grandifl ora var. former C2 candidate, SEUG: entrada Species of Concern Compositae Perityle specuicola alcove rock-daisy G1 UT BLM: Sensitive, USFWS: Rare, S1 (Asteraceae) former C2 candidate, SEUG: Species of Concern Compositae Xylorhiza Moab woody-aster G4 Rare, S1S2 (Asteraceae) glabriuscula var. T1T2 linearifolia Cruciferae Lepidium alyssoides Eastwood’s tall G5? Add. Data, SR (Brassicaceae) var. eastwoodiae pepperplant T2T4 Cruciferae Lepidium moabense Moab pepperplant GNR (Brassicaceae) Euphorbiaceae Tragia ramosa noseburn G5 Peripheral, S1 Gramineae Muhlenbergia Mexican muhly G5 Infrequent, (Poaceae) mexicana S1 Gramineae Spartina pectinata prairie cordgrass G5 Add. Data, S? (Poaceae) Hydrophyllaceae Nama retrorsum Howell’s nama G3 USFWS: former 3C Watch, S2 Hydrophyllaceae Phacelia howelliana Howell’s phacelia G2 USFWS: former 3C, SEUG: Watch, S2 Species of Concern Leguminosae Caesalpinia repens creeping rush-pea G2 Add. Data, S2 (Fabaceae or Caesalpiniaceae)

Chapter 3: Results 11 Table 3.3. Plant species of conservation concern from Arches National Park, cont.

TNC UTCDC Family Species Common name global Legal status status rank Leguminosae Psoralidium slenderleaf scurfpea G5T3?Q Tax. Prob., (Fabaceae) lanceolatum var. S3? stenophyllum (H) Liliaceae Zigadenus vaginatus alcove death-camas G2 USFWS: former 3C, SEUG: Rare, S2 Species of Concern Loasaceae Mentzelia Thompson’s stickleaf G4 Watch, S2 thompsonii Orchidaceae Epipactis gigantea giant helleborine G3G4 SEUG: Species of Concern Orchidaceae Habenaria zothecina alcove bog-orchid G2 UT BLM: Sensitive, USFWS: Rare, S2 (Platanthera former C2 candidate, SEUG: zothecina) Species of Concern Polypodiaceae Pellaea glabella var. western cliffbrake G5T4 Peripheral, S2 occidentalis Polypodiaceae Pellaea limitanea border cloak-fern G4G5 Peripheral, S1 Primulaceae Primula specuicola cave primrose G4Q USFWS: former 3C Watch, S3 Rubiaceae Galium multifl orum Colorado bedstraw G3G4 Watch, S3? var. coloradense Saxifragaceae Fendlera rupicola fendlerbush G5 Peripheral, (Hydrangeaceae) S2S3 Umbelliferae Lomatium latilobum Canyonlands G1 USFS R4: Sensitive, UT BLM: Rare, S1 (Apiaceae) lomatium Sensitive, USFWS: former C2 candidate, SEUG: Species of Concern Zygophyllaceae Kallstroemia yellow kallstroemia G5 Add. Data, S1 californica Zygophyllaceae Kallstroemia orange kallstroemia G5 Peripheral, S1 parvifl ora This list is derived from Schelz and Moran (2007) and Utah Division of Wildlife Resources (1998) for Utah Conservation Data Center (UTCDC) status. All species are confi rmed as occurring in Arches NP, except those marked with an (R) (reported). Species known only from historic records are marked with an (H).

Codes: TNC global rank assesses abundance and conservation priority on a scale of 1–5 (1 being extremely vulnerable and 5 being secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank, Q = taxonomic questions, U = unknown, and NR = not ranked. A comparable scoring system is used to measure state abundance (S) in Utah under the column UTCDC status. Under legal status, USFWS = U.S. Fish and Wildlife Service. C2 = Category 2 candidate (a former category used for taxa that might warrant being proposed for threatened or endangered status following additional research). 3C = Category 3 candidates (species dropped from consideration for listing). USFS = U.S. Forest Service. UT BLM = U.S. Bureau of Land Management, Utah ofﬁ ce. UTCDC status includes conservation categories adopted by the state natural heritage program to prioritize endemic and rare plant taxa (UDWR 1998). These categories include: Historical (presumed extinct or only known historically through their range), Rare (plants with rangewide viability concerns), Watch (regional endemics without rangewide viability concerns), Peripheral (rare or uncommon in Utah, but more common rangewide), Infrequent (plants occur infrequently over western U.S.), Taxonomic Problems (validity of species, subspecies, or variety has been questioned), Additional Data (species appear rare, but more information needed before being assigned to a different category).

12 Annotated Checklist of Vascular Flora: Arches National Park Table 3.4. Noxious weed species of Arches National Park.

Family Species Common name Park status

Compositae (Asteraceae) Centaurea maculosa spotted knapweed Present Compositae (Asteraceae) Centaurea repens Russian knapweed Present Compositae (Asteraceae) Onopordum acanthium Scotch thistle Reported Convolvulaceae Convolvulus arvensis ﬁ eld bindweed Present Cruciferae (Brassicaceae) Lepidium latifolium broad-leaf pepperplant Present Lythraceae Lythrum salicaria purple loosestrife Present This list is derived from the State of Utah Department of Agriculture and Food (2008) and Dewey and Andersen (2005).

Chapter 3: Results 13

of the park would increase by nearly 22%. Many 4 Discussion of the “missing” potential or reported taxa consist of cryptic species that fl ower late in the grow- In the more than 40 years since the publication of ing season, or occur in undersampled wetland or Harrison and others (1964), the known fl ora of weedy habitats. Confi rmation of reported species Arches NP has increased by more than 40%, from should remain a high priority, as some of these 320 to 522 taxa (Tables 1.3.4, 3.1, Figure 1.3.4, Ap- taxa may ultimately prove to be erroneous. Addi- pendix A). Based on available habitats and prox- tional studies would also be useful to better refi ne imity, at least another 146 plant taxa may occur the abundance and distribution of species of high within the park (Appendix B). Should all of these management interest (especially rare plants and potential species be found, the documented fl ora invasive non-natives).

Chapter 4: Discussion 15

——. 1984. Volume 4, Subclass Asteridae (except 5 Literature Cited Asteraceae). Intermountain fl ora: Vascular plants of the Intermountain West, U.S.A. Albee, B. J., L. M. Shultz, and S. Goodrich. 1988. Bronx, N.Y.: New York Botanical Garden. Atlas of the vascular plants of Utah. +Occa- 573 pp. sional Publ. 7, Salt Lake City: Utah Museum of Natural History. 670 pp. Cronquist, A., N. H. Holmgren, and P. K. Hol- mgren, eds. 1997. Volume 3, Part A, Subclass Allan, J. S. 1977. The plant communities of Rosidae (except Fabales). Intermountain Arches National Park. Provo, Ut.: Brigham fl ora: Vascular plants of the Intermountain Young University. 98 pp. West, U.S.A. Bronx, N.Y.: New York Botani- Barkworth, M. E., K. M. Capels, S. Long, and M. cal Garden. 446 pp. B. Piep, eds. 2003. Magnoliophyta: Com- Dewey, S., and K. Andersen. 2005. An inventory melinidae (in part): Poaceae, part 2. Flora of of invasive non-native plants in Arches Na- North America north of Mexico, Volume 25. tional Park (2003–2004): Final report. Utah New York: Oxford University Press. 783 pp. State University, Logan, Ut. Weed Science Barkworth, M. E., K. M. Capels, S. Long, L. K. Research Project Report # SD0510A. 31 pp. Anderton, and M. B. Piep, eds. 2007. Mag- Doelling, H. 2003. Geology of Arches National noliophyta: Commelinidae (in part): Poace- Park. Pages 11–36 in Sprinkel, D. A., T. ae, part 1. Flora of North America north C. Chidsey, Jr., and P. B. Anderson, eds., of Mexico, Volume 24. New York: Oxford Geology of Utah’s parks and monuments. University Press. 911 pp. Utah Geological Association Publication 28 Barneby, R. C. 1989. Volume 3, Part B. Fabales. (second edition). In Cronquist, A., A. H. Holmgren, N. H. Dorn, R. D. 2001. Vascular plants of Wyoming. Holmgren, J. L. Reveal, and P. K. Holmgren, Third edition. Cheyenne, Wy.: Mountain eds., Intermountain fl ora: Vascular plants of West Publishing. 412 pp. the Intermountain West, U.S.A. Bronx, N.Y.: New York Botanical Garden. 279 pp. Evenden, A., M. Miller, M. Beer, E. Nance, S. Daw, A. Wight, M. Estenson, and L. Cudlip. Chronic, H. 1988. Pages of stone: Geology of 2002. Northern Colorado Plateau Vital western national parks and monuments: Signs Network and Prototype Cluster plan Grand Canyon and the Plateau Country. for natural resources monitoring: Phase I Seattle, Wa.: The Mountaineers. 158 pp. report. October 1. Two volumes. National Cronquist, A. 1994. Volume 5, Asterales. In Park Service, Northern Colorado Plateau Cronquist, A., A. H. Holmgren, N. H. Network, Moab, Ut. 138 pp + app. Holmgren, J. L. Reveal, and P. K. Holmgren, Fertig, W. 2006. Herbarium and fi eld studies of eds., Intermountain fl ora: Vascular plants of vascular plant fl ora of ARCH for National the Intermountain West, U.S.A. Bronx, N.Y.: Park Service Inventory and Monitoring Pro- New York Botanical Garden. 496 pp. gram. Investigator’s Annual Report, National Cronquist, A., A. H. Holmgren, N. H. Holmgren, Park Service. 2 pp. and J. L. Reveal, eds. 1972. Volume 1: Geo- ——. 2007. Introduced and naturalized plants of logical and botanical history of the region, Utah. Sego Lily 30(5):7–11. its plant geography and a glossary. The vascular cryptogams and the gymnosperms. Flora of North America Editorial Committee. Intermountain fl ora: Vascular plants of the 1993. Pteridophytes and Gymnosperms. Intermountain West, U.S.A. Bronx, N.Y.: Flora of North America north of Mexico, New York Botanical Garden. 270 pp. Volume 2. New York: Oxford University Press. 475 pp. Cronquist, A., A. H. Holmgren, N. H. Holmgren, J. L. Reveal, and P. K. Holmgren, eds. 1977. ——. 1997. Magnoliophyta: Magnoliidae and Volume 6: The Monocotyledons. Inter- Hamamelidae. Flora of North America mountain fl ora: Vascular plants of the In- north of Mexico, Volume 3. New York: Ox- termountain West, U.S.A. Bronx, N.Y.: New ford University Press. 590 pp. York Botanical Garden. 584 pp.

Chapter 5: Literature Cited 17 ——. 2000. Magnoliophyta: Alismatidae, Are- National Park Service (NPS). 2000. Manage- cidae, Commelinidae (in part), and Zin- ment policies. U.S. Department of Interior, giberidae. Flora of North America north National Park Service, Washington, DC. of Mexico, Volume 22. New York: Oxford Publication # NPS D1416. 137 pp. University Press. 352 pp. Northern Colorado Plateau Network (NCPN). ——. 2002a. Magnoliophyta: Commelinidae (in in ed. Arches National Park vegetation part): Cyperaceae. Flora of North America classifi cation and mapping project. Natural north of Mexico, Volume 23. New York: Resource Technical Report. National Park Oxford University Press. 608 pp. Service, Denver, Co. ——. 2002b. Magnoliophyta: Liliidae: Liliales and Noss, R. F., and A. Y. Cooperrider. 1994. Saving Orchidales. Flora of North America north nature’s legacy: Protecting and restoring of Mexico, Volume 26. New York: Oxford biodiversity. Washington, D.C.: Island Press. University Press. 723 pp. 416 pp. ——. 2003. Magnoliophyta: Caryophyllidae, part Schelz, C., and M. Moran. 2005. Southeast Utah 1. Flora of North America north of Mexico. Group plants. National Park Service South- Volume 4. New York: Oxford University east Utah Group (Arches NP, Canyonlands Press. 559 pp. NP, Hovenweep NM, & Natural Bridges NM), Moab, Ut. 17 pp. ——. 2005. Magnoliophyta: Caryophyllidae, part 2. Flora of North America north of Mexico. ——. 2007. Southeast Utah Group threatened, Volume 5. New York: Oxford University endangered and species of concern. South- Press. 656 pp. east Utah Group (Arches NP, Canyonlands NP, Hovenweep NM, & Natural Bridges ——. 2006a. Magnoliophyta: Asteridae, part 6: NM), Moab, Ut. 3 pp. Asteraceae, part 1. Flora of North America north of Mexico. Volume 19. New York: Schelz, C., M. Moran, and A. Lafever. 2006. Oxford University Press. 579 pp. Southeast Utah Group plants. National Park Service Southeast Utah Group (Arches NP, ——. 2006b. Magnoliophyta: Asteridae, part 7: Canyonlands NP, Hovenweep NM, & Natu- Asteraceae, part 2. Flora of North America ral Bridges NM), Moab, Ut. 16 pp. north of Mexico. Volume 20. New York: Oxford University Press. 666 pp. Shultz, L. M., R. D. Ramsey, and W. Lindquist. 2006. Revised atlas of Utah plants. College ——. 2006c. Magnoliophyta: Asteridae, part 8: of Natural Resources, Utah State University, Asteraceae, part 3. Flora of North America Logan, Ut. http://earth.gis.usu.edu/plants. north of Mexico. Volume 21. New York: Oxford University Press. 616 pp. Springer, A. E., L. E. Stevens, and R. Harms. 2006. Inventory and classifi cation of selected Fowler, J. F., N. L. Stanton, and R. L. Hartman. National Park Service springs on the Colo- 2007. Distribution of hanging garden vegeta- rado Plateau. Northern Arizona University, tion associations on the Colorado Plateau, Flagstaff , Az. NPS Cooperative Agreement # USA. Journal of the Botanical Research CA 1200-99-009; Task # NAU-118. Institute of Texas 1(1):585–607. State of Utah Department of Agriculture and Harrison, B. F., S. L. Welsh, and G. Moore. 1964. Food (SUDAF). 2008. Utah noxious weed Plants of Arches National Monument. list. http://ag.utah.gov/divisions/plant/nox- Brigham Young University Science Bulletin ious/documents/noxUtah.pdf. 5(1):1–23. Topp, S. 2007. Veg mappers fi nd “new” plants Holmgren, N. H., P. K. Holmgren, and A. Cron- for Utah National Parks. Sego Lily 30(2):8–9. quist. 2005. Volume 2, Part B, Subclass Dille- niidae. Intermountain fl ora: Vascular plants Utah Division of Wildlife Resources (UDWR). of the Intermountain West, U.S.A. Bronx, 1998. Inventory of sensitive species and N.Y.: New York Botanical Garden. 488 pp. ecosystems in Utah. Endemic and rare plants of Utah: An overview of their distribution Margules, C. R., and R. L. Pressey. 2000. Sys- and status. Report prepared for the Utah tematic conservation planning. Nature Reclamation Mitigation and Conservation 405:243–253.

18 Annotated Checklist of Vascular Flora: Arches National Park Commission and U.S. Department of the Welsh, S. L., N. D. Atwood, S. Goodrich, and L. Interior. 566 pp + app. C. Higgins. 2003. A Utah ﬂ ora. Third edition, revised. Provo, Ut.: Brigham Young Univer- Weber, W. A., and R. C. Wittmann. 2001. Colo- sity. 912 pp. rado ﬂ ora: Western Slope. Third edition. Boulder: University Press of Colorado. 488 pp. Welsh, S. L. 1982. Utah plant types: Histori- cal perspective, 1840–1981, annotated list, and bibliography. Great Basin Naturalist 42(2):129–195.

Chapter 5: Literature Cited 19

Holmgren et al. 2005; Weber and Wittmann 2001). No ef- Appendices fort has been made to identify additional synonyms from monographs and fl oras preceding the early 1970s. The following appendices contain annotated checklists of the vascular fl ora of Arches National Park. Common name (Appendices A, B, C, and D) Common names are derived primarily from Welsh and Appendix A lists all species that have been confi rmed for others (2003) and the Intermountain Flora (Barneby 1989; the fl ora with an authenticated voucher specimen (includ- Cronquist 1994; Cronquist et al. 1972, 1977, 1984, 1997; ing extant and historical records) or that have been reli- Holmgren et al. 2005). ably reported for the park in the literature (Harrison et al. 1964; NCPN in ed.; Schelz et al. 2006; Springer et al. 2006). Life form (Appendices A, B, C, and D) Vouchers were examined from the ARCH herbarium and collections from Brigham Young University (BRY), Utah Life form is based on the stature and growth habit of typi- State University (USU), the University of Wyoming (RM), cal members of a species at reproductive maturity. Trees and the digital museum database of the New York Botani- are woody perennials with a single stem or trunk and are cal Garden (NY). Species entries in Appendix A are orga- typically over 3.5 m tall. Shrubs are woody perennials nized alphabetically by family and genus. with one to many trunks arising from near the base of the plant and are usually less than 3.5 m tall. Perennial Forbs Appendix B includes species that may potentially occur (PerF) are broad-leaved dicots or monocots with non- within Arches NP based on the proximity of the park to woody stems (at least above ground level) that live for mul- other populations or the presence of suitable habitat. tiple years. Annual Forbs (AnnF) are broad-leaved dicots or monocots with non-woody stems that complete their Appendix C lists species that have been previously report- life cycle (mature, fl ower, and die) in one year. Perennial ed for Arches NP but are now considered erroneous be- Graminoids (PerG) are linear-leaved, grass-like monocots cause they are based on misidentifi ed specimens or ques- that live for multiple years. Annual Graminoids (AnnG) tionable literature records. are linear-leaved, grass-like monocots that complete their life cycle (mature, fl ower, and die) in one year. Ferns and Appendix D contains the same confi rmed, historical, and fern-allies (Fern) are non-fl owering vascular plants that reported taxa as in Appendix A, but organized by life form reproduce by spores. rather than by family in order to better meet the needs of NPS interpretive specialists. Range (Appendices A, B, and C) Range represents the distribution of a species within Utah The codes and references for each fi eld in the appropriate and the state’s contribution to the plant’s overall global appendices are summarized below. range. Disjunct (Disj) taxa have their Utah distribution Family (Appendices A, B, C, and D) separated from the main, contiguous portion of their range by a gap of more than 800 km (ca 500 miles). Introduced Family concepts and nomenclature follow A Utah Flora, (Intro) taxa are those that are not native to Utah or North third edition (Welsh et al. 2003). Alternative family names America but which have become naturalized (breeding and treatments from other regional fl oras (Dorn 2001; on their own without human assistance). Local Endemics Flora of North America Editorial Committee 1993, 2002b; (LocEn) are taxa whose entire global range is restricted Holmgren et al. 2005; Weber and Wittman 2001) are listed to an area of less than 16,500 km2 (ca 6,370 mi2, or 1° of in parentheses. latitude × 2° of longitude). Peripheral (Periph) taxa have a widespread global distribution but occur at the margin of Species name (Appendices A, B, C, and D) their contiguous range in Utah and occupy less than 5% Nomenclature for scientifi c names and taxonomic con- of the state’s area (usually only within a few miles of the cepts are derived from Welsh and others (2003). Scientifi c state border). Regional Endemics (RegEn) are taxa with names are listed in alphabetical rather than phylogenetic a global range of 16,500–250,000 km2 (an area about the order. size of the state of Wyoming). Sparse taxa occur widely across Utah or North America but their range within Utah Synonyms/Taxonomic notes (Appendices A, B, and C) is small and patchy, with populations restricted to special- Pertinent synonyms are included for species with alter- ized or uncommon habitats. Widespread (Wide) taxa have native scientifi c names in other recent state and regional global ranges exceeding 250,000 km2 and occur over at fl oras (Barkworth et al. 2003, 2007; Barneby 1989; Cron- least 10% of the state. quist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn 2001; Flora of North America Editorial Committee 1993, Park status (Appendices A, C, and D) 1997, 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; Park status identifi es whether a species is Present (Pres),

Chapter 6: Appendices 21 Historical (Hist), or Reported (Rep) for Arches NP. Spe- Year doc. (Appendix A) cies are considered Present if a confi rmed voucher or doc- Year documented indicates the fi rst year in which a spe- umented observation has been made within park bound- cies was confi rmed or reported to occur in Arches NP. aries since 1970. Historical species have been confi rmed This year does not necessarily correspond with the date with a voucher from the park but have not been relocated of the voucher specimen cited under Source. “NA” indi- since before 1970. Reported species are listed for the park cates that the fi rst year is not known (used mostly for “re- in the literature (Dewey and Andersen 2005; Harrison et al. ported” taxa). 1964; NCPN in ed.; Schelz et al. 2006; Springer et al. 2006), but have not been corroborated with a voucher specimen. Comments (Appendices A, B, C, and D) Species that may potentially occur in Arches NP are listed This column includes additional comments on the status in Appendix B. Rejected taxa (listed in Appendix C) have of a particular species in the park, the place of origin for either been falsely reported (FalsRep) in the literature introduced species, legal status, and other data. based on misidentifi ed herbarium specimens or are considered Questionable (Ques?) because park reports are Nativity (Appendix D) from well outside their known or expected range. Nativity indicates whether a species is native to Utah or Pop. size (Appendix A) Introduced (Intro) based on Fertig (2007). Population size refers to the abundance and frequency Flower time (Appendix D) with which a species occurs within Arches NP. Abundance Flower time refers to the months in which a species is most data are derived from Dewey and Andersen (2005), Har- likely to be in fl ower. NA indicates a species that is a non- rison and others (1964), NCPN (in ed.), and personal ob- fl owering fern or gymnosperm. servations. Abundant (Abun) taxa have large populations, have a broad ecological amplitude (often being dominant Habitat (Appendix D) in one to several vegetation types), or occur extensively across the park. Common (Com) taxa have large local The distribution of species across generalized habitats in populations but may be restricted to a single vegetation Arches NP is indicated by an “x” in the columns BlckBr type or do not occur across the entire park. Uncommon through Wet and is derived from the Arches NP vegetation (Unc) taxa have low to medium-sized populations, typi- classifi cation (NCPN in ed.), Harrison and others (1964), cally have a narrow ecological amplitude (or at least con- and Welsh and others (2003). BlckBr = Blackbrush shrub tribute relatively little to total vegetative cover), or have a (including sagebrush communities). SaltDes = Salt des- small range across the park. Rare taxa usually have low ert shrub dominated by Atriplex spp. PinJun = Pinyon- population numbers and are restricted to a single location juniper woodlands. Wet = Wetlands (including hanging within the park. A “?” indicates that population size is un- gardens). Additional comments are included for species known. that are restricted to a particular soil/geologic type or are found mostly in disturbed sites. Source (Appendix A) Sources are the evidence for the assigned park status. In the case of Present and Historical taxa, sources are herbarium records (with collector, collection number, and repository) that were corroborated during this study. For reported taxa, the source is a published or unpublished reference citing the species within the park. Specimen repositories include the Arches National Park herbarium (ARCH), Brigham Young University (BRY), Utah State University (USU), New York Botanical Garden (NY), and the University of Wyoming’s Rocky Mountain Herbarium (RM).

22 Annotated Checklist of Vascular Flora: Arches National Park

A. eastwoodiana is Moab vicinity, Grand Co., UT (Rydberg & Garrett 8468 NY) America Helper, Carbon Helper, Co., UT (Trelease s.n. MO) America Comments c name with 1963 1985 locality of Type 1963 1963 1963 2006 Native to Central 1987 1949 locality is Type 1987 Native to tropical NA 1963 Year Year doc. (ARCH) (ARCH) Harrison, & Moore 2338b (ARCH) Harrison, & Moore 2308 (ARCH) Moore 2019 (ARCH) (ARCH) (ARCH) Moore 1920 (ARCH) (ARCH) (2006) Moore 2750 (ARCH) Source Pop. size Park status Range Life form western poison ivy Shrub Wide Pres Unc 1913 Welsh tomentose amsonia PerF Wide Pres Unc Paspal 59 squawbush Shrubsmooth sumac Wide Pres Shrub Wide Com Welsh, Pres Rare Welsh, squawbush Shrub Wide Pres Unc & Welsh redroot pigweed AnnF Intro Pres Rare Fertig 22998 prostrate pigweed AnnF Wide Pres Unc Belnap 444 tumble pigweed AnnF Intro Pres Unc Belnap 443 Harriman’s yuccaHarriman’s Shrub Wide Pres Com & Welsh narrowleaved yucca Shrub RegEn Rep ? Schelz et al. Common name box elder Tree Wide Pres Rare & Welsh . var .

. . var var var. ssp Rhus radicans, T. Rhus radicans, T. radicans A. eastwoodiana R. trilobata trilobata R. trilobata simplicifolia A. graecizans Y. harrimaniae Y. angustissima Synonyms/ notes Taxonomic interius, Negundo aceroides interius . . var var exus stenophylla . harrimaniae Amsonia tomentosa var. Rhus glabra Toxicodendron rydbergii Rhus aromatica trilobata Rhus aromatica simplicifolia Amaranthus retrofl Amaranthus albus Amaranthus blitoides Yucca harrimaniae Yucca var Yucca angustissima Yucca angustissima Y. Acer negundo A. negundo National Park, organized by family and scientifi vascular plant taxa of Arches rmed, historical, and reported Apocynaceae Anacardiaceae Anacardiaceae Anacardiaceae Anacardiaceae Amaranthaceae Amaranthaceae Amaranthaceae Agavaceae Agavaceae FamilyAceraceae Species name Appendix A. Confi taxonomic and biogeographic notes.

Chapter 6: Appendices 23 24

Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientiﬁ c name with taxonomic and Annotated Checklist ofVascular Flora: Arches NationalPark biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Apocynaceae Apocynum Includes vars. common dogbane PerF Wide Pres Unc Fagan 28 1963 cannabinum cannabinum, (ARCH) glaberrimum, & angustifolium Asclepiadaceae Asclepias asperula spider milkweed PerF Wide Rep Rare Northern 2006 var. asperula Colorado Plateau Network (in ed.) Asclepiadaceae Asclepias A. cryptoceras pallid milkweed PerF Wide Pres Unc Barnett 55 1947 cryptoceras var. (ARCH) cryptoceras Asclepiadaceae Asclepias latifolia broadleaf milkweed PerF Wide Pres Unc Fleming 824 1964 (ARCH) Asclepiadaceae Asclepias Included in A. bigseed milkweed PerF RegEn Pres Unc Pitts 76 1947 macrosperma involucrata by (ARCH) some authors Asclepiadaceae Asclepias ruthiae A. uncialis ssp. Ruth’s milkweed PerF RegEn Pres Rare Topp 2006 Type locality ruthiae ST05180601 is Calf Spring (ARCH) Canyon SE of Castledale, Emery Co., UT (Maguire 18310 UTC) Asclepiadaceae Asclepias speciosa showy milkweed PerF Wide Pres Rare Fagan 14 1963 (ARCH) Asclepiadaceae Asclepias A. galioides whorled milkweed PerF Wide Pres Unc Topp 2006 subverticillata ST08220606 (ARCH) Berberidaceae Mahonia fremontii Berberis fremontii Fremont’s mahonia Shrub Wide Pres Unc Barnett 60 1963 (ARCH) Berberidaceae Mahonia repens Berberis repens Oregon-grape Shrub Wide Pres Unc Forsythe & 1981 Swanke 16 (ARCH) Bignoniaceae Catalpa common catalpa Tree Intro Pres Rare Welsh & 1963 Native to E North bignonioides Moore 1998 America (BRY) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Boraginaceae Cryptantha thick-sepaled AnnF Wide Pres Com Pyrah 64 1947 crassisepala var. cryptanth (ARCH) elachantha Boraginaceae Cryptantha fl ava Oreocarya fl ava plateau yellow PerF Wide Pres Com Barnett 44 1947 cryptanth (ARCH) Boraginaceae Cryptantha Oreocarya yellow-eye PerF Wide Pres Unc Fleming 762 1989 fl avoculata fl avoculata cryptanth (ARCH) Boraginaceae Cryptantha Oreocarya plateau cryptanth PerF RegEn Pres Unc Pyrah 48 1947 fulvocanescens fulvocanescens, (ARCH) includes vars. fulvocanescens & echinoides Boraginaceae Cryptantha Kelsey’s cryptanth AnnF Wide Pres Unc Allan 477 1972 kelseyana (BRY) Boraginaceae Cryptantha Oreocarya long-fl ower PerF RegEn Pres Unc Barnett 41 1963 Type locality is longifl ora longifl ora cryptanth (ARCH) Palisades, Mesa Co., CO (Crandall s.n. RM) Boraginaceae Cryptantha Oreocarya Osterhout’s PerF RegEn Pres Rare Fleming 972 1990 Type locality osterhoutii osterhoutii cryptanth (ARCH) is Monument Park [Colorado National Monument], near Grand Junction, Mesa Co., CO (Osterhout 6138 RM) Boraginaceae Cryptantha wing-nut cryptanth AnnF Wide Pres Unc Fleming 779 1989 pterocarya var. (ARCH)

Chapter 6:Appendices pterocarya Boraginaceae Cryptantha recurved cryptanth AnnF Wide Pres Unc Allan 33 1972 recurvata (BRY) Boraginaceae Cryptantha tenuis slender cryptanth PerF RegEn Pres Rare Welsh & 1964 Type locality Moore 2801 is Courthouse (ARCH) Wash, Grand Co., UT (Eastwood s.n. CAS) 25 26

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Boraginaceae Cryptantha Wetherill’s PerF RegEn Rep ? Schelz et al. 1892 Type locality wetherillii cryptanth (2006) ? is Courthouse Wash, Grand Co., UT (Eastwood s.n. CAS) Boraginaceae Cynoglossum common hound’s- PerF Intro Rep ? Schelz et al. NA Native to Eurasia offi cinale tongue (2006) Boraginaceae Heliotropium Euploca bindweed AnnF Wide Pres Unc Welsh & 1952 convolvulaceum convolvulacea ssp. heliotrope Moore 2771 californica (ARCH) Boraginaceae Heliotropium salt heliotrope PerF Wide Rep ? Schelz et al. NA curassavicum var. (2006) obovatum Boraginaceae Lappula occidentalis L. texana, L. cupseed stickseed AnnF Wide Pres Com Pyrah 54 1963 var. cupulata redowskii var. (ARCH) cupulata, L. marginata Boraginaceae Lithospermum showy stoneseed PerF Wide Pres Rare Fagan 13 1963 incisum (ARCH) Boraginaceae Tiquilia latior Coldenia matted tiquilia PerF RegEn Pres Unc Welsh & 1963 (Ehretiaceae) hispidissima Moore 1989 (ARCH) Cactaceae Echinocereus Includes var. claretcup PerF Wide Pres Unc Welsh & 1963 triglochidiatus var. inermis. Vars. not Moore 2013 melanacanthus recognized in Flora (BRY) of North America (2003). Cactaceae Opuntia erinacea O. polyacantha var. common pricklypear PerF Wide Pres Unc Welsh & 1963 var. erinacea erinacea Moore 2022 (ARCH) Cactaceae Opuntia erinacea O. polyacantha var var. utahensis . Utah pricklypear PerF Wide Pres Com Welsh & 1963 hystricina. Var. not Moore 2020 recognized in Flora (ARCH) of North America (2003) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cactaceae Opuntia Vars. not large pricklypear PerF Wide Pres Unc Belnap 415 1987 phaeacantha var. recognized in Flora (ARCH) major of North America (2003) Cactaceae Opuntia Flora of North plains pricklypear PerF Wide Rep ? Schelz et al. NA polyacantha var. America (2003) (2006) polyacantha includes vars. rufi spina & trichophora Cactaceae Opuntia Included with O. rufous-spine PerF Wide Pres Unc Welsh & 1963 polyacantha var. polyacantha var. pricklypear Moore 2002 rufi spina polyacantha in (ARCH) Flora of North America (2003) Cactaceae Opuntia Included with O. slenderspine PerF Periph Pres Com Welsh & 1963 polyacantha var. polyacantha var. pricklypear Moore 2001 trichophora polyacantha in (ARCH) Flora of North America (2003) Cactaceae Opuntia whipplei Cylindropuntia Whipple’s cholla Shrub RegEn Rep ? Schelz et al. NA whipplei (2006) Cactaceae Sclerocactus S. parvifl orus, small-fl ower PerF Wide Pres Unc Fleming 744 1963 Type locality is whipplei var. roseus S. whipplei var. fi shhook cactus (ARCH) Glen Canyon, intermedius. San Juan Co., UT Includes S. (Clover & Jotter contortus & S. 2398 US) terrae-canyonae Capparaceae Cleome lutea yellow beeplant AnnF Wide Pres Unc Barnett 45 1963 (Cleomaceae) (ARCH) Chapter 6:Appendices Caprifoliaceae Symphoricarpos long-fl ower Shrub Wide Pres Unc Welsh & 1949 longifl orus snowberry Moore 2018 (ARCH) Caryophyllaceae Arenaria fendleri Eremogone Eastwood’s PerF Wide Pres Unc Fleming 968 1963 (Alsinaceae) var. eastwoodiae eastwoodiae var. sandwort (ARCH) eastwoodiae, A. eastwoodiae 27 28

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Caryophyllaceae Silene antirrhina sleepy catchfl y AnnF Wide Pres Unc Welsh & 1963 Moore 2015 (ARCH) Chenopodiaceae Allenrolfea iodine bush Shrub Wide Pres Unc Welsh & 1963 occidentalis Moore 2714 (ARCH) Chenopodiaceae Atriplex canescens A. canescens var. four-wing saltbush Shrub Wide Pres Com Welsh & 1963 var. canescens occidentalis Moore 1985 (ARCH) Chenopodiaceae Atriplex shadscale Shrub Wide Pres Com Welsh & 1963 confertifolia Moore 2047 (ARCH) Chenopodiaceae Atriplex corrugata mat saltbush Shrub RegEn Pres Unc Belnap 505 1987 (ARCH) Chenopodiaceae Atriplex gardneri A. cuneata Castle Valley Shrub RegEn Pres Com Welsh & 1964 Type locality var. cuneata saltbush Moore 3017 is near Emery, (ARCH) Emery Co., UT (Jones 5443 RM) Chenopodiaceae Atriplex garrettii A. garrettii Garrett’s saltbush Shrub LocEn Pres Unc Hanson 343 1961 Type locality is var. garrettii (BRY) vicinity of Moab, Grand Co., UT (Rydberg & Garrett 8465 NY) Chenopodiaceae Bassia americana Kochia americana greenmolly PerF Wide Rep Unc Schelz et 2006 al. (2006), Northern Colorado Plateau Network ( in ed.) Chenopodiaceae Bassia hyssopifolia Echinopsilon fi ve-horn AnnF Intro Pres Unc Welsh, 1963 Native to Eurasia hyssopifolium smotherweed Harrison, & Moore 2274 (ARCH) Chenopodiaceae Bassia scoparia Kochia scoparia, B. summer-cypress AnnF Intro Pres Unc Moran s.n. 2006 Native to Eurasia sieversiana (ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Chenopodiaceae Chenopodium C. album lambsquarter AnnF Intro Pres Unc Welsh, 1963 Native to Eurasia album var. album Harrison, & Moore 2328 (BRY) Chenopodiaceae Chenopodium C. berlandieri var. pitseed goosefoot AnnF Wide Pres Unc Welsh & 1963 album var. zschackei Moore 2824 berlandieri (ARCH) Chenopodiaceae Chenopodium C. pratericola desert goosefoot AnnF Sparse Pres Unc Howell & True 1968 dessicatum 44908 (NY) Chenopodiaceae Chenopodium C. fremontii Fremont’s goosefoot AnnF Wide Pres Unc Welsh & 1963 fremontii var. Moore 2723 fremontii (ARCH) Chenopodiaceae Chenopodium C. salinum oakleaf goosefoot AnnF Wide Rep ? Schelz et al. NA glaucum var. (2006) salinum Chenopodiaceae Chenopodium narrow-leaf AnnF Wide Rep ? Schelz et al. NA leptophyllum goosefoot (2006) Chenopodiaceae Corispermum Includes vars. American bugseed AnnF Wide Pres Rare Allan 251 1972 americanum americanum & (BRY) rydbergii Chenopodiaceae Cycloloma C. atriplicifolium winged pigweed AnnF Wide Pres Unc Belnap 452 1987 atriplicifolia (ARCH) Chenopodiaceae Grayia spinosa Atriplex spinosa, A. spiny hopsage Shrub Wide Pres Unc Pitts 66 1963 grayi (ARCH) Chenopodiaceae Halogeton halogeton AnnF Intro Pres Unc Welsh & 1963 Native to Eurasia glomeratus Moore 1986 (BRY) Chenopodiaceae Krascheninnikovia Ceratoides winterfat Shrub Wide Pres Unc Pyrah 33

Chapter 6:Appendices lanata var. lanata lanata, Eurotia (ARCH) 1963 lanata. Vars. not recognized in Flora of North America (2003). 29 30

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Chenopodiaceae Krascheninnikovia Ceratoides lanata Ruin Park winterfat Shrub LocEn Pres Unc Welsh & 1963 Type locality is lanata var. ruinina var. ruinina. Vars. Moore 2763 Ruin Park, Beef not recognized - paratype Basin, San Juan in Flora of North (ARCH) Co., UT (Welsh America (2003). & Welsh 22377 BRY) Chenopodiaceae Monolepis Nuttall’s poverty- AnnF Wide Pres Unc Belnap 522 1988 nuttalliana weed (ARCH) Chenopodiaceae Salsola tragus S. kali, S. iberica, S. Russian thistle AnnF Intro Pres Unc Welsh & 1963 Native to Asia pestifer, S. australis Moore 1978 (ARCH) Chenopodiaceae Sarcobatus greasewood Shrub Wide Pres Unc Welsh & 1963 (Sarcobataceae) vermiculatus Moore 2036 (ARCH) Chenopodiaceae Suaeda S. depressa broom seepweed AnnF Wide Pres Unc Welsh & 1963 calceoliformis Moore 2715 (ARCH) Chenopodiaceae Suaeda occidentalis Included in S. western seepweed AnnF Wide Pres Unc Welsh & 1963 calceoliformis by Moore 2722 some authors (ARCH) Chenopodiaceae Suaeda torreyana S. moquinii, S. Torrey’s seepweed AnnF Wide Pres Unc Hahn 2003 var. torreyana nigra, S. fruticosa LAH05210303 (ARCH) Chenopodiaceae Zuckia brandegei Z. brandegeei Brandegee’s siltbush Shrub RegEn Pres Unc Welsh & 1964 Type locality is var. brandegei var. brandegeei, Moore 3018 San Juan River, Atriplex brandegei (ARCH) San Juan Co., UT (Brandegee s.n. GH) Compositae Ambrosia Franseria bur ragweed AnnF Wide Pres Com Welsh & 1963 (Asteraceae) acanthicarpa acanthicarpa Moore 2712 (ARCH) Compositae Arctium minus burdock PerF Intro Pres Unc Moran s.n. 2006 Native to Eurasia (Asteraceae) (ARCH) Compositae Artemisia bigelovii Bigelow’s sagebrush Shrub Wide Pres Unc Welsh & 1963 (Asteraceae) Moore 2732 (ARCH) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Artemisia Oligosporus sand wormwood PerF Wide Pres Com Welsh & 1963 (Asteraceae) campestris var. campestris ssp. Moore 2739 scouleriana pacifi cus, O. (ARCH) pacifi cus, A. campestris ssp. pacifi ca Compositae Artemisia Oligosporus tarragon PerF Wide Pres Com Welsh & 1963 (Asteraceae) dracunculus ssp. dracunculus. Vars. Moore 2772 glauca not recognized in (ARCH) Cronquist (1994) and Flora of North America (2006) Compositae Artemisia fi lifolia Oligosporus sand sagebrush Shrub Wide Pres Com Welsh & 1963 (Asteraceae) fi lifolius Moore 2719 (BRY) Compositae Artemisia Louisiana PerF Wide Pres Unc Welsh & 1963 (Asteraceae) ludoviciana var. wormwood Moore 2728 albula (ARCH) Compositae Artemisia Louisiana PerF Wide Pres Unc Fowler & May 1993 (Asteraceae) ludoviciana var. wormwood 1671 (RM) ludoviciana Compositae Artemisia Louisiana PerF Wide Pres Unc Hahn 1963 (Asteraceae) ludoviciana var. wormwood LAH05220305 mexicana (ARCH) Compositae Artemisia nova var. A. arbuscula var. (Asteraceae) nova nova, Seriphidium black sagebrush Shrub Wide Rep ? Schelz et al. NA novum. Vars. not (2006) recognized in Flora of North America (2006). Chapter 6:Appendices

Compositae Artemisia spinescens Picrothamnus budsage Shrub Wide Pres Unc Fagan 7 1963 (Asteraceae) desertorum (ARCH) Compositae Artemisia tridentata Seriphidium basin big sagebrush Shrub Wide Pres Com Welsh & 1963 (Asteraceae) var. tridentata tridentatum ssp. Moore 2737 tridentatum (ARCH) 31 32

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Aster ascendens A. chilensis ssp. Pacifi c aster PerF Wide Pres Unc Welsh, 1963 (Asteraceae) adscendens, A. Harrison, & adscendens, Moore 2299 Symphyotrichum (BRY) ascendens, Virgulaster ascendens Compositae Aster brachyactis Symphyotrichum rayless alkali aster AnnF Wide Pres Unc Allan 443 1973 (Asteraceae) ciliatum, (BRY) Brachyactis ciliata ssp. angusta Compositae Aster hesperius A. lanceolatus Siskiyou aster PerF Wide Pres Unc Fertig 22992 2006 (Asteraceae) var. hesperius, (ARCH) Symphyotrichum lanceolatum var. hesperium Compositae Baccharis salicina Includes B. emoryi Rio Grande Shrub Wide Pres Rare Welsh & 1963 (Asteraceae) seepwillow Moore 2762 (ARCH) Compositae Bidens frondosa devil’s- beggarsticks PerF Wide Pres Unc Moran s.n. 2006 (Asteraceae) (ARCH) Compositae Brickellia longifolia B. longifolia var. longleaf Shrub Wide Pres Unc Welsh & 1963 (Asteraceae) longifolia brickellbush Moore 2731 (ARCH) Compositae Brickellia rough brickellbush Shrub Wide Pres Unc Welsh, 1963 (Asteraceae) microphylla var. Harrison, & scabra Moore 2298 (ARCH) Compositae Brickellia Mohave Shrub Wide Pres Rare Hahn 2003 (Asteraceae) oblongifolia var. brickellbush LAH05140310 linifolia (ARCH) Compositae Centaurea maculosa Centauera spotted knapweed PerF Intro Pres Unc Torrence s.n. 2002 UT state noxious (Asteraceae) biebersteinii, (ARCH) weed. Native to Acosta maculosa Europe Compositae Centaurea repens Acroptilon repens Russian knapweed PerF Intro Pres Unc Andersen 2003 UT state noxious (Asteraceae) ARCH V-1 weed. Native to (ARCH) Eurasia Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Chaenactis stevia dusty-maiden AnnF Wide Pres Unc Forsythe 4 1979 (Asteraceae) stevioides (ARCH) Compositae Chaetopappa Leucelene rose-heath PerF Wide Pres Unc Welsh & 1963 (Asteraceae) ericoides ericoides, Aster Moore 2010 arenosus (ARCH) Compositae Chrysopsis villosa C. villosa hispid golden-aster PerF Wide Pres Com Welsh & 1947 (Asteraceae) var. minor var. hispida, Moore 2740 Heterotheca villosa (ARCH) var. hispida, H. villosa var. minor Compositae Chrysothamnus Lorandersonia spear-leaf Shrub Wide Pres Unc Hahn 1963 (Asteraceae) linifolius linifolia rabbitbrush LAH05230307 (ARCH) Compositae Chrysothamnus Ericameria naus- greenish Shrub Wide Pres Com Fertig 22993 2006 (Asteraceae) nauseosus var. eosa var. consimilis, rabbitbrush (ARCH) consimilis E. nauseosa var. oreophila. Included in C. nauseosus var. oreophilus by some authors Compositae Chrysothamnus C. nauseosus graystem Shrub Wide Pres Com Welsh & 1963 (Asteraceae) nauseosus var. var. hololeucus, rabbitbrush Moore 2711 gnaphalodes Ericameria (ARCH) nauseosa var. hololeuca Compositae Chrysothamnus Ericameria (Asteraceae) nauseosus var. nauseosa var. glabrate Shrub Wide Pres Com Welsh & 1963 graveolens rabbitbrush Moore 2765 graveolens, E. (BRY) nauseosa var. Chapter 6:Appendices glabrata, includes C. nauseosus var. oreophilus, C. nauseosus ssp. pinifolius Compositae Chrysothamnus Ericameria rush rabbitbrush Shrub RegEn Pres Unc Belnap 455 1987 (Asteraceae) nauseosus var. nauseosa var. (ARCH) junceus juncea 33 34

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Chrysothamnus Ericameria smoothseed Shrub RegEn Pres Rare Welsh & 1963 (Asteraceae) nauseosus var. nauseosa var. rabbitbrush Moore 2768 leiospermus leiosperma (BRY) Compositae Chrysothamnus C. parryi var. longbract Shrub Wide Pres Rare Welsh & 1963 (Asteraceae) parryi var. howardii attenuatus, rabbitbrush Moore 2764 Ericameria parryi (ARCH) var. howardii Compositae Chrysothamnus lanceleaf Shrub Wide Rep ? Schelz et al. NA (Asteraceae) viscidifl orus var. rabbitbrush (2006) lanceolatus Compositae Chrysothamnus Included in var. slenderleaf Shrub Wide Pres Com Welsh, 1963 (Asteraceae) viscidifl orus var. viscidifl orus by rabbitbrush Harrison, & stenophyllus some authors. Vars. Moore 2294 not recognized (ARCH) in Flora of North America (2006). Compositae Chrysothamnus green rabbitbrush Shrub Wide Pres Unc Welsh, 1963 (Asteraceae) viscidifl orus var. Harrison, & viscidifl orus Moore 2285 (ARCH) Compositae Cirsium calcareum Var. not recognized pretty thistle PerF Wide Pres Unc Fagan 41 1985 (Asteraceae) var. pulchellum in Cronquist (1994), (ARCH) included in C. arizonicum var. bipinnatum in Flora of North America 2006) Compositae Cirsium (Asteraceae) Vars. not New Mexico thistle PerF Wide Pres Unc Fleming 852 1989 neomexicanum var. recognized in Flora (ARCH) neomexicanum of North America (2006) Compositae Cirsium rydbergii Rydberg’s thistle PerF RegEn Pres Rare Fowler & May 1989 Type locality is (Asteraceae) 1732 (ARCH) along San Juan River near Bluff, San Juan Co., UT (Rydberg & Garrett 10001 NY). USFWS: 3C Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Cirsium undulatum C. undulatum wavy-leaved thistle PerF Wide Pres Unc Welsh, 1963 (Asteraceae) var. undulatum Harrison, & Moore 2281 (ARCH) Compositae Cirsium vulgare bull thistle PerF Intro Rep ? Schelz et al. NA Native to Europe (Asteraceae) (2006) Compositae Conyza canadensis Vars. not Canadian AnnF Wide Pres Unc Welsh, 1963 (Asteraceae) var. glabrata recognized in Flora horseweed Harrison, & of North America Moore 2261 (2006) (BRY) Compositae Crepis runcinata var. glaucous PerF Wide Pres Rare Welsh & 1963 (Asteraceae) glauca hawksbeard Moore 2044 (ARCH) Compositae Dicoria brandegei D. canescens ssp. Brandegee’s AnnF RegEn Pres Com Welsh & 1963 (Asteraceae) brandegei. Includes sandplant Moore 2713 var. wetherillii. (ARCH) Vars. not recognized in Flora of North America (2006) Compositae Encelia frutescens E. frutescens var. bush encelia Shrub Wide Pres Unc Welsh & 1963 (Asteraceae) var. resinosa frutescens, E. Moore 1994 resinifera (ARCH) Compositae Enceliopsis nakedstem PerF Wide Rep Rare Northern 2006 Type locality is (Asteraceae) nudicaulis var. Colorado “Southern Utah” nudicaulis Plateau (Bishop s.n. GH) Network (in ed.) Compositae Enceliopsis nutans Encelia nutans noddinghead PerF RegEn Pres Unc Barnett 54 1947

Chapter 6:Appendices (Asteraceae) (ARCH) Compositae Erigeron aphanactis E. concinnus var. hairy daisy PerF Wide Pres Rare Fleming 974 (Asteraceae) var. aphanactis aphanactis (ARCH) 1990

Compositae Erigeron E. bellidiastrum var. pretty daisy AnnF Wide Pres Unc Pyrah 65 1963 (Asteraceae) bellidiastrum bellidiastrum (ARCH) Compositae Erigeron divergens E. colomexicanus, spreading daisy PerF Wide Pres Unc Welsh & 1963 (Asteraceae) var. cinereus E. tracyi Moore 2043 (ARCH) 35 36

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Erigeron divergens spreading daisy PerF Wide Pres Unc Belnap 458 1987 (Asteraceae) var. divergens (ARCH) Compositae Erigeron basin daisy PerF Wide Pres Unc Belnap 77 1987 (Asteraceae) pulcherrimus (ARCH) Compositae Erigeron pumilus E. concinnus var. Navajo fl eabane PerF Wide Pres Unc Schelz s.n. 2003 (Asteraceae) var. concinnus concinnus, includes (ARCH) var. subglaber according to some authors Compositae Erigeron utahensis E. sparsifolius slenderleaf daisy PerF RegEn Pres Rare Moore 246 1964 Type locality is (Asteraceae) var. sparsifolius (BRY) Willow Creek, San Juan Co., UT (Eastwood 48 CAS) Compositae Erigeron utahensis E. utahensis Utah daisy PerF RegEn Pres Unc Welsh & 1963 Type locality is (Asteraceae) var. utahensis Moore 1992 Kanab, Kane Co., (ARCH) UT (Thompson s.n. GH) Compositae Euthamia Solidago western goldenrod PerF Wide Pres Unc Welsh & 1963 (Asteraceae) occidentalis occidentalis Moore 2720 (ARCH) Compositae Flaveria campestris marshweed AnnF Sparse Pres Rare Welsh & 1963 (Asteraceae) Moore 2751 (BRY) Compositae Gaillardia Hopi blanketfl ower PerF Wide Pres Com Fagan 23 1949 (Asteraceae) pinnatifi da (ARCH) Compositae Gaillardia pulchella Includes reports of pretty AnnF Intro Pres Unc Fertig 22996 2006 Probably escaped (Asteraceae) G. aristata blanketfl ower (ARCH) from highway planting. Compositae Gaillardia basin blanketfl ower PerF RegEn Pres Unc Welsh & 1963 Type locality is (Asteraceae) spathulata Moore 2049 Rabbit Valley, (ARCH) Wayne Co., UT (Ward 401 GH) Compositae Grindelia fastigiata Included in (Asteraceae) G. erect gumweed PerF RegEn Pres Unc Welsh & 1963 hirsutula in Flora Moore 2730 of North America (ARCH) (2006) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Gutierrezia thread snakeweed Shrub Wide Pres Com Welsh, 1963 (Asteraceae) microcephala Harrison, & Moore 2259 (ARCH) Compositae Gutierrezia broom snakeweed Shrub Wide Pres Abun Welsh & 1963 (Asteraceae) sarothrae Moore 2716 (ARCH) Compositae Haplopappus Stenotus thrift goldenweed PerF Wide Pres Unc Belnap 83 1949 (Asteraceae) armerioides var. armerioides var. (ARCH) armerioides armerioides Compositae Haplopappus rusbyi H. drummondii, Rusby’s goldenbush Shrub Wide Pres Unc Belnap 440 1963 Cited as (Asteraceae) Isocoma rusbyi, I. (ARCH) “Haplopappus drummondii sp.” in Harrison et al. (1964) Compositae Helianthus annuus Ssp. not recognized common sunfl ower AnnF Wide Pres Unc Welsh & 1963 (Asteraceae) ssp. lenticularis in Flora of North Moore 2725 America (2006). (ARCH) Compositae Helianthus Includes H. sand sunfl ower AnnF RegEn Pres Unc Welsh & 1963 Type locality (Asteraceae) anomalus deserticola Moore 2759 is desert S of (ARCH) Hanksville, Wayne Co., UT (Stanton 328 US). Compositae Helianthus Includes var. prairie sunfl ower AnnF Wide Pres Unc Welsh & 1963 (Asteraceae) petiolaris ssp. fallax petiolaris Moore 2050 (ARCH) Compositae Heterotheca H. psammophila, sand goldenaster PerF Periph Pres Rare Welsh & 1963 May be (Asteraceae) subaxillaris H. latifolia, H. Moore 2744 introduced from subaxillaris var. (ARCH) SW USA latifolia

Chapter 6:Appendices Compositae Hymenopappus H. fi lifolius (Asteraceae) fi lifolius var. var. common PerF Wide Pres Com Welsh & 1963 cinereus lugens, H. fi lifolius hyalineherb Moore 2008 var. paucifl orus, (ARCH) H. fi lifolius var. megacephalus, H. fi lifolius var. tomentosus Compositae Hymenoxys acaulis Tetraneuris acaulis Arizona hymenoxys PerF Wide Pres Unc Fleming 959 1987

37 (Asteraceae) var. arizonica var. arizonica (ARCH) 38

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Hymenoxys acaulis H. argentea Canyonlands PerF Wide Pres Com Pyrah 63 1947 (Asteraceae) var. ivesiana var. ivesiana, hymenoxys (ARCH) Tetraneuris ivesiana Compositae Lactuca serriola prickly lettuce AnnF Intro Pres Unc Topp 2006 Native to Europe (Asteraceae) ST05200602 (ARCH) Compositae Lactuca tatarica var. L. pulchella, L. blue lettuce PerF Wide Pres Unc Paspal 65 1988 (Asteraceae) pulchella oblongifolia, (ARCH) Mulgedium pulchellum Compositae Lygodesmia L. arizonica Arizona rush-pink PerF RegEn Rep ? Schelz et al. NA (Asteraceae) grandifl ora var. (2006) arizonica Compositae Lygodesmia L. entrada, Entrada rush-pink PerF LocEn Pres Rare Welsh & 1963 Type locality is (Asteraceae) grandifl ora var. considered a Moore 2006 Tusher Canyon, entrada synonym of var. (BRY) 15 mi WNW of dianthopsis by Moab, Grand some authors Co., UT (Welsh & Welsh 16725 BRY). UT BLM: Sensitive, USFWS: former C2 candidate. Compositae Lygodesmia showy rush-pink PerF Wide Pres Com Fagan 38 1985 (Asteraceae) grandifl ora var. (ARCH) grandifl ora Compositae Machaeranthera Aster canescens, hoary aster PerF Wide Pres Unc Welsh & 1963 (Asteraceae) canescens var. Dieteria canescens Moore 2718 canescens var. canescens (ARCH) Compositae Machaeranthera Haplopappus gumweed aster PerF Wide Pres Unc Paspal 58 1963 (Asteraceae) grindelioides var. nuttallii, Xanthisma (ARCH) grindelioides grindelioides var. grindelioides Compositae Machaeranthera tansyleaf aster AnnF Wide Pres Unc Welsh & 1963 (Asteraceae) tanacetifolia Moore 1976 (ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Malacothrix sowthistle desert- AnnF Wide Pres Unc Welsh & 1948 (Asteraceae) sonchoides dandelion Moore 1916 (ARCH) Compositae Onopordum Scotch thistle PerF Intro Rep Rare Dewey & 2004 Utah state (Asteraceae) acanthium Anderson noxious weed. (2005) Native to Eurasia Compositae Oxytenia acerosa Iva acerosa copperweed Shrub Wide Pres Unc Fertig 23000 2006 (Asteraceae) (ARCH) Compositae Perityle specuicola alcove rock-daisy PerF LocEn Pres Rare Schelz s.n. 1998 Type locality (Asteraceae) (ARCH) is 2 mi NE of Moab, Grand Co., UT (Welsh 16283 BRY). UT BLM: Sensitive, USFWS: former C2 candidate. Compositae Petradoria pumila Solidago petradoria rock-goldenrod PerF Wide Pres Unc Paspal 56 1963 (Asteraceae) var. pumila (ARCH) Compositae Platyschkuhria Bahia nudicaulis desert bahia PerF RegEn Pres Unc Welsh & 1949 Type locality is (Asteraceae) integrifolia var. var. desertorum Moore 2007 Cisco, Grand Co., desertorum (ARCH) UT (Jones s.n. US) Compositae Senecio douglasii S. longilobus, Douglas’ groundsel PerF Wide Rep ? Schelz et al. NA (Asteraceae) var. longilobus S. fl accidus var. (2006) douglasii, S. fl accidus var. fl accidus Compositae Senecio Packera basin groundsel PerF Wide Pres Unc Forsythe 15 1963 (Asteraceae) multilobatus multilobata (ARCH) Compositae Senecio spartioides S. multicapitatus; many-headed PerF Wide Pres Unc Welsh & 1963 Chapter 6:Appendices (Asteraceae) var. multicapitatus vars. not groundsel Moore 2724 recognized in Flora (ARCH) of North America (2006) Compositae Senecio spartioides (Asteraceae) var Vars. not broom groundsel PerF Wide Pres Unc Welsh & 1963 . spartioides recognized in Flora Moore 2754 of North America (ARCH) (2006) 39 40

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Senecio Packera Rocky Mountain PerF Wide Pres Rare Allan 66 1972 (Asteraceae) streptanthifolius streptanthifolia var. groundsel (BRY) var. streptanthifola. streptanthifolius Vars. not recognized in Flora of North America (2006) Compositae Solidago canadensis S. altissima, Canadian goldenrod PerF Wide Pres Unc Fleming 839 1963 (Asteraceae) var. salebrosa S. lepida var. (ARCH) salebrosa Compositae Solidago velutina S. sparsifl ora, three-nerve PerF Wide Pres Unc Fowler & May 1989 (Asteraceae) S. velutina var. goldenrod 1694 (ARCH) sparsifl ora Compositae Sonchus asper spiny-leaf sow- AnnF Intro Pres Unc Welsh & 1963 Native to Europe (Asteraceae) thistle Moore 2035 (ARCH) Compositae Sonchus oleraceus common sow-thistle AnnF Intro Pres Unc Moran s.n. 2003 Native to Eurasia (Asteraceae) (ARCH) Compositae Stephanomeria S. exigua ssp. white-plume wire- AnnF Wide Pres Com Welsh & 1963 (Asteraceae) exigua exigua lettuce Moore 1981 (ARCH) Compositae Stephanomeria brown-plume wire- PerF Wide Pres Unc Welsh, 1963 (Asteraceae) paucifl ora lettuce Harrison, & Moore 2263 (ARCH) Compositae Stephanomeria S. minor var. (Asteraceae) tenuifolia var. minor. Vars. not slender wirelettuce PerF Wide Pres Unc Welsh & 1963 tenuifolia recognized in Flora Moore 1991 of North America (ARCH) (2006)

Compositae Taraxacum common dandelion PerF Intro Pres Unc Fagan 34 1964 Native to Eurasia (Asteraceae) offi cinale (ARCH) Compositae Tetradymia spinosa T. spinosa var. shortspine Shrub Wide Pres Unc Fleming 969 1947 (Asteraceae) spinosa horsebrush (ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Thelesperma Vars. not scapose PerF Wide Pres Unc Paspal 55 1987 Type locality is (Asteraceae) subnudum var. recognized in Flora greenthread (ARCH) “Southern Utah” subnudum of North America (Parry 109 GH) (2006). Compositae Townsendia annua annual townsendia AnnF Wide Pres Unc Schelz s.n. 2005 Type locality is (Asteraceae) (ARCH) Cottonwood Wash, N of Bluff, San Juan Co., UT (Maguire 13509 GH) Compositae Townsendia incana hoary townsendia PerF Wide Pres Unc Pitts 74 1949 (Asteraceae) (ARCH) Compositae Tragopogon dubius T. dubius var. major yellow salsify PerF Intro Pres Unc Welsh & 1964 Native to Europe (Asteraceae) Moore 2813 (ARCH) Compositae Vanclevea stylosa Chrysothamnus resinbush Shrub RegEn Pres Com Welsh & 1963 Type locality (Asteraceae) stylosa Moore 2717 is near Epsom (ARCH) Creek [between Bluff and Mexican Hat], San Juan Co., UT (Eastwood 36 CAS) Compositae Wyethia scabra Scabrethia scabra rough mule’s-ears PerF Wide Pres Com Barnett 71 1947 (Asteraceae) ssp. scabra (ARCH) Compositae Xanthium X. pensylvanicum, rough cocklebur AnnF Wide Pres Unc Welsh, 1963 (Asteraceae) strumarium var. X. saccharatum; Harrison, & canadense vars. not Moore 2286 recognized in Flora (ARCH) of North America Chapter 6:Appendices (2006). Compositae Xylorhiza Machaeranthera Moab woodyaster PerF LocEn Pres Rare Belnap 94 1987 Type locality is (Asteraceae) glabriuscula var. linearifolia, X. (ARCH) 6 miles NW of linearifolia linearifolia Moab, Grand Co., UT (Watson 679 TEX) 41 42

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Compositae Xylorhiza tortifolia Machaeranthera hurtleaf woodyaster PerF RegEn Pres Rare Welsh & 1964 Type locality is E (Asteraceae) var. imberbis tortifolia var. Moore 2803 bank of Green imberbis (ARCH) River W of Moab (Cronquist 8994 NY) Compositae Xylorhiza venusta Machaeranthera Cisco woodyaster PerF RegEn Pres Unc Pyrah 50 1947 Type locality is (Asteraceae) venusta (ARCH) Cisco, Grand Co., UT (Jones s.n. POM) Convolvulaceae Convolvulus arvensis fi eld bindweed PerF Intro Pres Unc Fagan 33 1985 Utah state (ARCH) noxious weed. Native to Eurasia Convolvulaceae Cuscuta pentagona fi eld dodder AnnF Wide Pres Rare Welsh, 1963 (Cuscutaceae) Harrison, & Moore 2345 (BRY) Convolvulaceae Evolvulus evolvulus PerF Wide Pres Unc Fleming 755 1989 nuttallianus (ARCH) Convolvulaceae Ipomoea purpurea morning-glory AnnF Intro Pres Rare Moran s.n. 2006 Native to tropical (ARCH) America Cornaceae Cornus sericea var. C. stolonifera, red-osier dogwood Shrub Wide Pres Rare Fagan 39 1985 sericea Swida sericea (ARCH) Cruciferae Arabis perennans Boechera perennial rockcress PerF Wide Pres Unc Fagan 26 1947 (Brassicaceae) var. perennans perennans (ARCH) Cruciferae Arabis pulchra var. Boechera pulchra desert rockcress PerF Wide Pres Com Pyrah 56 1947 Type locality (Brassicaceae) pallens var. pallens (ARCH) is Westwater, Grand Co., UT (Jones s.n. US) Cruciferae Chorispora tenella blue mustard AnnF Intro Pres Unc Topp 2006 Native to Asia (Brassicaceae) ST05070603 (ARCH) Cruciferae Conringia orientalis hare’s-ear mustard AnnF Intro Rep ? Schelz et al. NA Native to Europe (Brassicaceae) (2006) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cruciferae Descurainia California tansy- PerF Wide Rep Rare Schelz et 2006 (Brassicaceae) californica mustard al. (2006), Northern Colorado Plateau Network (in ed.) Cruciferae Descurainia incana D. richardsonii var. mountain tansy- AnnF Wide Pres Com Pyrah 52 1963 (Brassicaceae) var. sonnei sonnei, D. incisa mustard (ARCH) var. incisa, D. incana ssp. incisa, D. incana var. incana Cruciferae Descurainia sophia ﬂ ixweed AnnF Intro Pres Unc Topp 2006 Native to Europe (Brassicaceae) ST05070601 (ARCH) Cruciferae Dithyrea wislizeni Dimorphocarpa spectacle-pod AnnF Wide Pres Com Pitts 65 1947 (Brassicaceae) wislizenii (ARCH) Cruciferae Draba cuneifolia wedgeleaf draba AnnF Wide Pres Unc Welsh & 1964 (Brassicaceae) var. cuneifolia Moore 2774 (ARCH) Cruciferae Draba reptans dwarf draba AnnF Wide Rep ? Schelz et al. NA (Brassicaceae) (2006) Cruciferae Erysimum asperum E. asperum var. sand dune PerF Wide Pres Unc Pitts 40 1963 Type locality is (Brassicaceae) var. purshii arkansanum, E. wallﬂ ower (ARCH) near the Great capitatum var. Salt Lake, Salt purshii, included in Lake Co., UT var. capitatum by (Carrington s.n.) some authors

Chapter 6:Appendices Cruciferae Lepidium alyssoides L. montanum var. Eastwood’s tall PerF RegEn Pres Rare Harms s.n. 2005 (Brassicaceae) var. eastwoodiae spathulatum, L. pepperwort (ARCH) eastwoodiae Cruciferae Lepidium (Brassicaceae) densiﬂ Includes var. prairie pepperwort AnnF Wide Pres Unc Welsh & 1964 orum var. macrocarpum Moore 2773 ramosum (ARCH) Cruciferae Lepidium latifolium Cardaria latifolia broad-leaf PerF Intro Pres Unc Moran s.n. 2006 Utah state (Brassicaceae) pepperwort (ARCH) noxious weed.

43 Native to Eurasia 44

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cruciferae Lepidium moabense Included in L. Moab pepperplant PerF LocEn Pres Rare Moran s.n. 2006 Type locality is (Brassicaceae) eastwoodiae by (ARCH) 2 miles NNE of some authors Moab, Grand Co., UT (Welsh 16351 BRY) Cruciferae Lepidium Jones’ pepperwort PerF Wide Pres Com Pyrah 55 1948 (Brassicaceae) montanum var. (ARCH) jonesii Cruciferae Lepidium clasping pepperwort AnnF Intro Pres Rare Fagan 17 1985 Native to Europe (Brassicaceae) perfoliatum (ARCH) Cruciferae Malcolmia africana African mustard AnnF Intro Pres Unc Welsh & 1964 Native to Africa (Brassicaceae) Moore 2808 (ARCH) Cruciferae Nasturtium Rorippa watercress PerF Intro Pres Rare Welsh, 1963 Native to Europe (Brassicaceae) offi cinale nasturtium- Harrison, & aquaticum Moore 2279 (ARCH) Cruciferae Physaria acutifolia P. australis sharp-leaf twinpod PerF Wide Pres Unc Barnett 42 1947 (Brassicaceae) var. acutifolia (ARCH) Cruciferae Sisymbrium tumble mustard AnnF Intro Pres Unc Belnap 49 1979 Native to Europe (Brassicaceae) altissimum (ARCH) Cruciferae Stanleya pinnata S. pinnata var. prince’s-plume PerF Wide Pres Com Barnett 65 1949 (Brassicaceae) pinnata (ARCH) Cruciferae Streptanthella long-beak fi ddle- AnnF Wide Pres Com Pitts 62 1949 (Brassicaceae) longirostris mustard (ARCH) Cruciferae Streptanthus S. cordatus var. heart-leaf PerF Wide Pres Unc Forsythe 13 1947 (Brassicaceae) cordatus cordatus jewelfl ower (ARCH) Cruciferae Thelypodiopsis elegant thelypody PerF RegEn Pres Rare Fagan 10 1985 Type locality (Brassicaceae) elegans (ARCH) is Westwater, Grand Co., UT (Jones s.n. POM) Cruciferae Thelypodium Moab thelypody PerF RegEn Pres Unc Welsh, 1963 (Brassicaceae) integrifolium var. Harrison, & gracilipes Moore 2337 (ARCH) Cupressaceae Juniperus Sabina Utah juniper Shrub Wide Pres Abun Pitts 73 1963 osteosperma osteosperma (ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cupressaceae Juniperus Sabina scopulorum Rocky Mountain Tree Wide Pres Rare Fertig 22646 2006 scopulorum juniper (ARCH) Cyperaceae Carex aurea C. garberi golden sedge PerG Wide Pres Unc Fleming 767 1947 Cited as C. (ARCH) garberi in Harrison et al. (1964) Cyperaceae Carex hystericina C. hystricina bottlebrush sedge PerG Wide Pres Unc Welsh, 1963 Harrison, & Moore 2300 (ARCH) Cyperaceae Carex nebrascensis Nebraska sedge PerG Wide Pres Unc Pitts 52 1963 (ARCH) Cyperaceae Carex praegracilis blackcreeper sedge PerG Wide Pres Unc Pyrah 42 1963 Cited as C. foena (ARCH) in Harrison et al. (1964) Cyperaceae Cyperus redroot ﬂ at-sedge AnnG Wide Pres Unc Schelz s.n. 1997 erythrorhizos (ARCH) Cyperaceae Cyperus esculentus Includes vars. chufa ﬂ at-sedge PerG Periph Pres Rare Fertig 23008 2006 Native to North leptostachys, & (ARCH) America, UT heermannii material may be introduced Cyperaceae Eleocharis palustris Includes E. common spikerush PerG Wide Pres Unc Barnett 49 1963 macrostachya, E. (ARCH) erythropoda, & E. uniglumis Cyperaceae Scirpus acutus Schoenoplectus hardstem bulrush PerG Wide Pres Unc Welsh, 1963 Cited as S. validus acutus var. Harrison, & in Harrison et al. occidentalis, Moore 2276 (1964). Schoenoplectus (ARCH) Chapter 6:Appendices lacustris ssp. acutus Cyperaceae Scirpus maritimus Scirpus maritimus alkali bulrush PerG Wide Pres Unc Topp 2006 var. paludosus, ST08220603 Schoenoplectus (ARCH) maritimus, Bolboschoenus maritimus var. paludosus 45 46

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Cyperaceae Scirpus pungens var. Scirpus pungens three-square PerG Wide Pres Unc Barnett 72 1963 longispicatus var. polyphyllus, bulrush (ARCH) Schoenoplectus pungens var. longispicatus, Schoenoplectus pungens var. polyphyllus Elaegnaceae Elaeagnus Russian-olive Tree Intro Pres Unc Welsh & 1963 Native to Europe angustifolia Moore 2748 (ARCH) Ephedraceae Ephedra torreyana Torrey’s ephedra Shrub Wide Pres Com Pyrah 59 1948 (ARCH) Ephedraceae Ephedra viridis var. E. viridis green ephedra Shrub Wide Pres Com Pyrah 32 1949 viridis (ARCH) Ephedraceae Ephedra viridis var. E. cutleri Cutler’s ephedra Shrub RegEn Pres Unc Welsh & 1952 viscida Welsh 21166 (BRY) Equisetaceae Equisetum arvense fi eld horsetail Fern Wide Pres Unc Welsh, 1963 Harrison, & Moore 2282 (ARCH) Equisetaceae Equisetum hyemale Hippochaete tall scouring-rush Fern Wide Pres Unc Welsh, 1963 var. affi ne hyemalis ssp. Harrison, & affi nis, E. prealtum Moore 2293 (ARCH) Equisetaceae Equisetum Hippochaete smooth scouring- Fern Wide Pres Unc Barnett 51 1963 laevigatum laevigata, E. rush (BRY) kansanum Euphorbiaceae Chamaesyce Euphorbia fendleri Fendler’s spurge PerF Wide Pres Unc Pyrah 61 1963 fendleri (ARCH) Euphorbiaceae Chamaesyce Euphorbia ridge-seeded spurge AnnF Wide Pres Unc Fertig 22991 2006 glyptosperma glyptosperma (ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Euphorbiaceae Chamaesyce parryi Euphorbia parryi Parry’s spurge AnnF Wide Pres Com Welsh & 1963 Type locality Moore 2710 is St. George, (ARCH) Washington Co., UT (Parry 247 MO) Euphorbiaceae Euphorbia E. robusta, E. shorthorn spurge PerF Wide Pres Com Fleming 766 1947 brachycera brachycera var. (ARCH) robusta, Tithymalus brachyceras Euphorbiaceae Tragia ramosa T. stylaris noseburn PerF Sparse Pres Unc Fleming 854 1989 (ARCH) Fagaceae Quercus gambelii Q. gambelii Gambel oak Tree Wide Pres Com Barnett 48 1947 var. gambelii (ARCH) Fagaceae Quercus turbinella turbinella live-oak Shrub Wide Pres Rare Forsythe 17 1981 (ARCH) Fagaceae Quercus welshii Q. havardii shinnery oak Shrub RegEn Pres Com Pyrah 66 1947 Type locality of var. tuckeri, Q. (ARCH) Q. havardii var. undulata tuckeri is Low Pass E of Kens Lake, San Juan Co., UT (Welsh & Higgins 23630 BRY) Fumariaceae Corydalis aurea var. C. curvisiliqua ssp. golden corydalis PerF Wide Pres Unc Belnap 74 1987 occidentalis occidentalis (ARCH) Gentianaceae Centaurium Great Basin PerF Wide Pres Unc Paspal 77 1991 exaltatum centaury (ARCH) Chapter 6:Appendices 47 48

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gentianaceae Swertia utahensis Frasera paniculata Utah swertia PerF RegEn Pres Unc Welsh, 1963 Cited as S. Harrison, & albomarginata Moore 2343 in Harrison (ARCH) et al. (1964). Type locality is Buckskin Mountains, Kaibab Plateau “on the S edge of Utah”, Kane Co., UT (Jones s.n. POM) Geraniaceae Erodium cicutarium stork’s-bill AnnF Intro Pres Com Welsh & 1963 Native to Europe Moore 2016 (ARCH) Gramineae Aegilops cylindrica Cylindropyrum jointed goatgrass AnnG Intro Pres Rare Fertig 22651 2006 Native to Eurasia (Poaceae) cylindricum (ARCH) Gramineae Agropyron A. cristatum var. crested wheatgrass PerG Intro Pres Rare Welsh & 1963 Native to Eurasia. (Poaceae) cristatum cristatum, A. Moore 2052 cristatum ssp. (ARCH) pectinatum, A. pectinatum, A. sibiricum, A. desertorum, A. fragile, A. mongolicum Gramineae Agrostis stolonifera A. alba, A. gigantea redtop PerG Intro Pres Unc Welsh, 1963 Native to Eurasia (Poaceae) Harrison, & and Africa Moore 2295 (ARCH) Gramineae Aristida purpurea A. glauca, A. purple three-awn PerG Wide Pres Com Welsh, 1948 (Poaceae) purpurea var. Harrison, & fendleriana, A. Moore 2330 purpurea var. (ARCH) longiseta Gramineae Bothriochloa B. ischaemum var. yellow bluestem PerG Intro Pres Rare Moran s.n. 2006 Native to Eurasia (Poaceae) ischaemum songarica (ARCH) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Bouteloua barbata Chondrosum sixweeks grama AnnG Wide Pres Com Welsh & 1963 (Poaceae) barbatum, Includes Moore 2531 vars. barbata & (ARCH) rothrockii Gramineae Bouteloua Includes vars. sideoats grama PerG Wide Pres Unc Welsh, 1949 (Poaceae) curtipendula curtipendula & Harrison, & caespitosa Moore 2313 (ARCH) Gramineae Bouteloua gracilis Chondrosum blue grama PerG Wide Pres Rare Topp 2003 (Poaceae) gracilie SET10130301 (ARCH) Gramineae Bromus carinatus B. marginatus, mountain brome PerG Wide Pres Unc Welsh & 1963 (Poaceae) B. polyanthus, Moore 2818 Ceratochloa (BRY) carinata Gramineae Bromus diandrus B. rigidus, ripgut brome AnnG Intro Pres Unc Moran s.n. 2006 Native to Eurasia (Poaceae) Anisantha diandra (ARCH) Gramineae Bromus japonicus B. arvensis, Includes Japanese chess AnnG Intro Pres Unc Topp 2006 Native to Eurasia (Poaceae) B. commutatus ST08080602 (ARCH) Gramineae Bromus tectorum Anisantha tectorum cheatgrass AnnG Intro Pres Com Pitts 44 1963 Native to Eurasia (Poaceae) (ARCH) Gramineae Calamagrostis Jones’ reedgrass PerG Wide Pres Rare Fertig 22994 2006 Type locality (Poaceae) scopulorum (ARCH) is Springdale, Washington Co., UT (Jones 6075 POM) Gramineae Cenchrus ﬁ eld sandbur AnnG Wide Pres Unc Welsh & 1963 (Poaceae) longispinus Moore 2756 Chapter 6:Appendices (ARCH) Gramineae Chloris virgata feather ﬁ nger grass AnnG Wide Pres Unc Moran s.n. 2006 (Poaceae) (ARCH) Gramineae Dactylis glomerata orchard grass PerG Intro Pres Rare Welsh, 1963 Native to Eurasia (Poaceae) Harrison, & and Africa Moore 2315 (ARCH) 49 50

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Digitaria sanguinalis hairy crabgrass AnnG Intro Pres Unc Fertig 22999 2006 Native to Eurasia (Poaceae) (ARCH) Gramineae Distichlis spicata D. spicata var. desert saltgrass PerG Wide Pres Unc Welsh & 1963 (Poaceae) stricta, D. stricta Moore 2031 (ARCH) Gramineae Echinochloa crus- Includes E. barnyard-grass AnnG Intro Pres Unc Welsh & 1963 Native to Eurasia (Poaceae) galli microstachya & Moore 2745 E. muricata var. (BRY) microstachya Gramineae Elymus canadensis E. canadensis var. Canada wildrye PerG Wide Pres Unc Fleming 830 1963 (Poaceae) canadensis (ARCH) Gramineae Elymus cinereus Leymus cinereus Great Basin wildrye PerG Wide Pres Unc Topp 2006 (Poaceae) ST08080606 (ARCH) Gramineae Elymus elymoides E. elymoides var. squirreltail PerG Wide Pres Unc Pitts 53 1949 (Poaceae) elymoides, Sitanion (ARCH) hystrix Gramineae Elymus salinus E. salina, E. Salina wildrye PerG Wide Pres Com Welsh, 1963 Type locality is (Poaceae) ambiguus, Leymus Harrison, & top of Salina salinus, L. salina, L. Moore 2297 Pass, Sevier Co., ambiguus (BRY) UT (Jones 5447 POM) Gramineae Elymus smithii Agropyron smithii, western wheatgrass PerG Wide Pres Unc Welsh & 1964 (Poaceae) Pascopyrum smithii, Moore 2809 includes vars. (ARCH) smithii & palmeri Gramineae Elymus trachycaulus E. trachycaulus slender wheatgrass PerG Wide Pres Com Welsh & 1963 (Poaceae) var. trachycaulus, Moore 2003 E. alaskanus (ARCH) ssp. latiglumis, Agropyron caninum, A. subsecundum, A. trachycaulum, A. latiglume Gramineae Enneapogon spike pappusgrass PerG Sparse Pres Rare Welsh 10933 1970 (Poaceae) desvauxii (BRY) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Eragrostis teal lovegrass AnnG Periph Pres Rare Hahn 2003 (Poaceae) hypnoides LAH08180301 (ARCH) Gramineae Eragrostis tufted lovegrass AnnG Wide Pres Unc Moran s.n. 2006 (Poaceae) pectinacea (ARCH) Gramineae Eremopyrum Agropyron annual wheatgrass AnnG Intro Pres Rare Topp 2006 Native to Asia (Poaceae) triticeum triticeum ST04210602 (ARCH) Gramineae Erioneuron pilosum hairy tridens PerG Wide Pres Unc Harrison 1947 (Poaceae) 11155 (BRY) Gramineae Erioneuron Dasyochloa fl uffgrass PerG Wide Pres Unc Harrison 1949 (Poaceae) pulchellum pulchella 11368 (BRY) Gramineae Festuca octofl ora Includes vars. six-weeks fescue AnnG Wide Pres Abun Barnett 62 1947 (Poaceae) glauca, hirtella, & (ARCH) octofl ora, Vulpia octofl ora Gramineae Festuca pratensis Lolium pratense, meadow fescue PerG Intro Pres Unc Fertig 22653 2006 Native to Eurasia (Poaceae) Schedonorus (ARCH) pratensis Gramineae Glyceria striata Includes G. elata fowl mannagrass PerG Wide Pres Unc Welsh, 1963 (Poaceae) Harrison, & Moore 2317 (ARCH) Gramineae Hilaria jamesii Pleuraphis jamesii galleta PerG Wide Pres Com Pyrah 60 1947 (Poaceae) (ARCH) Gramineae Hordeum jubatum Critesion jubatum foxtail barley PerG Wide Pres Rare Welsh, 1963 (Poaceae) Harrison, & Moore 2314 (ARCH) Chapter 6:Appendices Gramineae Hordeum murinum H. murinum rabbit barley AnnG Intro Pres Unc Belnap 521 1988 Native to Europe (Poaceae) ssp. leporinum, (ARCH) H. leporinum, Critesion glaucum Gramineae Muhlenbergia foxtail muhly PerG Wide Pres Unc Welsh, 1963 (Poaceae) andina Harrison, & Moore 2311 (ARCH) 51 52

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Muhlenbergia scratchgrass PerG Wide Pres Unc Welsh, 1952 (Poaceae) asperifolia Harrison, & Moore 2273 (ARCH) Gramineae Muhlenbergia Mexican muhly PerG Sparse Pres Unc Welsh, 1963 (Poaceae) mexicana Harrison, & Moore 2302 (ARCH) Gramineae Muhlenbergia sandhill muhly PerG Wide Pres Com Welsh, 1963 (Poaceae) pungens Harrison, & Moore 2329 (ARCH) Gramineae Muhlenbergia Includes M. Thurber’s muhly PerG Wide Pres Rare Welsh & 1963 (Poaceae) thurberi curtifolia Moore 2755 (ARCH) Gramineae Munroa squarrosa Monroa squarrosa false buffalograss AnnG Wide Pres Unc Fertig 22997 2006 (Poaceae) (ARCH) Gramineae Panicum P. occidentale, bundle panicgrass PerG Wide Pres Unc Welsh & 1952 (Poaceae) acuminatum P. tennessense, Moore 2042 P. huachucae, (ARCH) Dichanthelium acuminatum ssp. fasciculatum, D. lanuginosum var. fasciculatum Gramineae Panicum capillare P. capillare var. witchgrass AnnG Wide Pres Unc Moran s.n. 2006 (Poaceae) occidentale (ARCH) Gramineae Panicum obtusum vine mesquite PerG Wide Pres Rare Topp 2006 (Poaceae) ST08220607 (ARCH) Gramineae Panicum virgatum switchgrass PerG Wide Pres Unc Fertig 22648 2006 (Poaceae) (ARCH) Gramineae Phleum pratense timothy PerG Intro Pres Rare Welsh, 1963 Native to Eurasia (Poaceae) Harrison, & Moore 2316 (ARCH) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Phragmites australis P. communis common reed PerG Wide Pres Unc Welsh, 1963 (Poaceae) Harrison, & Moore 2291 (ARCH) Gramineae Poa fendleriana P. longiligula, muttongrass PerG Wide Pres Com Pyrah 30 1949 (Poaceae) P. fendleriana (ARCH) ssp. longiligula, includes reports of P. cusickii & P. epilis Gramineae Poa pratensis P. agassizensis Kentucky bluegrass PerG Intro Pres Unc Welsh & 1964 Native to Europe (Poaceae) Moore 2816 (BRY) Gramineae Poa secunda Includes P. Sandberg’s PerG Wide Pres Unc Pitts 46 1963 (Poaceae) gracillima, P. bluegrass (ARCH) juncifolia, P. sandbergii, P. canbyi, P. scabrella, P. nevadensis, P. ampla and all vars. of P. secunda Gramineae Polypogon rabbitfoot-grass AnnG Intro Pres Unc Welsh & 1963 Native to Eurasia (Poaceae) monspeliensis Moore 2034 and Africa (ARCH) Gramineae Puccinellia distans weeping alkaligrass PerG Intro Pres Unc Welsh & 1963 Native to Eurasia (Poaceae) Moore 2030 (ARCH) Gramineae Schizachyrium Andropogon little bluestem PerG Wide Pres Unc Welsh & 1963 (Poaceae) scoparium scoparius, S. Moore 2736 scoparium var. (ARCH) neomexicanum Chapter 6:Appendices Gramineae Secale cereale cultivated rye AnnG Intro Rep ? Schelz et al. NA Native to Eurasia (Poaceae) (2006) Gramineae Setaria glauca S. pumila ssp. yellow bristlegrass AnnG Intro Pres Unc Moran s.n. 2006 Native to Eurasia (Poaceae) pumila, Pennisetum (ARCH) glaucum 53 54

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Setaria viridis green bristlegrass AnnG Intro Pres Rare Welsh, 1963 Native to Eurasia (Poaceae) Harrison, & Moore 2346 (ARCH) Gramineae Spartina gracilis alkali cordgrass PerG Wide Pres Unc Topp 2006 (Poaceae) ST08070601 (ARCH) Gramineae Spartina pectinata prairie cordgrass PerG Sparse Pres Rare Topp 2006 (Poaceae) ST08220605 (ARCH) Gramineae Sphenopholis S. obtusata var. prairie wedgegrass PerG Wide Pres Rare Topp 1963 (Poaceae) obtusata obtusata SET08040301 (ARCH) Gramineae Sporobolus airoides S. airoides alkali sacaton PerG Wide Pres Unc Welsh & 1963 (Poaceae) var. airoides Moore 2041 (ARCH) Gramineae Sporobolus spike dropseed PerG Wide Pres Unc Welsh & 1952 (Poaceae) contractus Moore 1984 (BRY) Gramineae Sporobolus sand dropseed PerG Wide Pres Com Fagan 20 1963 (Poaceae) cryptandrus (ARCH) Gramineae Sporobolus mesa dropseed PerG Wide Pres Com Belnap 437 1952 (Poaceae) fl exuosus (ARCH) Gramineae Sporobolus giant dropseed PerG Wide Pres Unc Welsh, 1963 (Poaceae) giganteus Harrison, & Moore 2270 (ARCH) Gramineae Stipa arida Achnatherum Mormon PerG Wide Hist NA Harrison, 1941 Type locality is (Poaceae) aridum needlegrass Marshal, & near Milford, Nielson s.n. Beaver Co., UT (BRY) (Jones 2106) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Gramineae Stipa arnowiae Achnatherum Arnow’s ricegrass PerG RegEn Pres Rare Harrison 1952 Cited as S. x (Poaceae) arnowiae. Hybrid 12001 - bloomeri in of S. hymenoides x paratype Harrison et al. S. comata. (BRY) (1964). Type locality is ca 9 miles E of Johnson Canyon, Kane Co., UT (Welsh & O’Dell 28062 BRY) Gramineae Stipa comata var. Hesperostipa needle-and-thread PerG Wide Pres Com Welsh & 1952 (Poaceae) comata comata var. comata Moore 2028a (ARCH) Gramineae Stipa hymenoides Oryzopsis Indian ricegrass PerG Wide Pres Abun Pitts 77 1963 (Poaceae) hymenoides, (ARCH) Achnatherum hymenoides Gramineae Stipa speciosa Achnatherum desert needlegrass PerG Wide Pres Unc Hahn 1952 (Poaceae) speciosum, Jarava LAH05260301 speciosa (ARCH) Gramineae Tridens muticus T. muticus var. slim tridens PerG Wide Pres Rare Topp 2006 (Poaceae) elongatus ST08220608 (ARCH) Gramineae Triticum aestivum wheat AnnG Intro Pres Unc Welsh & 1963 Native to Eurasia (Poaceae) Moore 1979 (ARCH) Hydrophyllaceae Nama densum var. leafy ﬁ ddleleaf AnnF Wide Rep ? Schelz et al. NA parviﬂ orum (2006) Hydrophyllaceae Nama retrorsum Howell’s nama AnnF RegEn Pres Rare Welsh & 1963 USFWS: 3C

Chapter 6:Appendices Moore 2026 (ARCH) Hydrophyllaceae Phacelia crenulata P. corrugata corrugate phacelia AnnF Wide Pres Unc Fleming 955 1948 var. corrugata (ARCH) Hydrophyllaceae Phacelia demissa brittle phacelia AnnF RegEn Rep ? Schelz et al. NA var. demissa (2006) 55 56

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Hydrophyllaceae Phacelia howelliana Included in P. Howell’s phacelia AnnF RegEn Pres Rare Pitts 71 1963 Type locality is crenulata by some (ARCH) 0.4 miles N of authors Bluff, San Juan Co., UT (Atwood 2454 BRY). USFWS: 3C Hydrophyllaceae Phacelia ivesiana Ives’ phacelia AnnF Wide Pres Com Barnett 66 1963 (ARCH) Juncaceae Juncus arcticus J. balticus var. Baltic rush PerG Wide Pres Com Pyrah 67 1963 montanus, J. (ARCH) arcticus var. balticus & ssp. ater, includes var. mexicanus Juncaceae Juncus articulatus jointed rush PerG Sparse Pres Rare Schelz s.n. 1997 (ARCH) Juncaceae Juncus ensifolius J. saximontanus Tracy’s rush PerG Wide Pres Unc Welsh, 1963 var. brunnescens forma brunnescens. Harrison, & Included in J. Moore 2321 ensifolius var. (ARCH) montanus by some authors Juncaceae Juncus ensifolius J. saximontanus, mountain rush PerG Wide Rep Rare Fowler et al. 1993 var. montanus Includes J. tracyi, J. (2007) xiphoides Juncaceae Juncus longistylus longstyle rush PerG Wide Pres Unc Welsh, 1963 Harrison, & Moore 2301 (ARCH) Juncaceae Juncus nevadensis Includes var. badius Nevada rush PerG Wide Rep ? Schelz et al. NA & nevadensis (2006) Juncaceae Juncus tenuis Includes vars. poverty rush PerG Wide Pres Rare Welsh, 1963 dudleyi, congestus, Harrison, & & tenuis. J. dudleyi Moore 2305 (ARCH) Juncaceae Juncus torreyi Torrey’s rush PerG Wide Pres Unc Schelz s.n. 1997 (ARCH) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Juncaginaceae Triglochin maritima T. maritima var. seaside arrowgrass PerG Wide Pres Unc Welsh & 1963 elata, Includes T. Moore 2033 concinnum (ARCH) Labiatae Hedeoma mock pennyroyal PerF Wide Pres Unc Welsh & 1963 (Lamiaceae) drummondii Moore 2040 (ARCH) Labiatae Marrubium vulgare common horehound PerF Intro Rep ? Schelz et al. NA Native to Eurasia (Lamiaceae) (2006) Labiatae Poliomintha incana purple sage Shrub Wide Pres Com Welsh & 1952 (Lamiaceae) Moore 1980 (ARCH) Leguminosae Astragalus crescent milkvetch PerF Wide Pres Com Welsh & 1940 (Fabaceae) amphioxys var. Moore 1910 amphioxys (ARCH) Leguminosae Astragalus Sheldon’s milkvetch PerF RegEn Hist NA Harrison 1949 (Fabaceae) amphioxys var. 11399 (BRY) vespertinus Leguminosae Astragalus painted milkvetch PerF Wide Pres Rare Holmgren 1952 (Fabaceae) ceramicus var. 10926 (NY) ceramicus Leguminosae Astragalus coltonii Moab milkvetch PerF RegEn Pres Unc Welsh & 1947 Type locality is (Fabaceae) var. moabensis Moore 1902 near Monticello, (ARCH) San Juan Co., UT (Eastwood 9 POM) Leguminosae Astragalus canoe milkvetch PerF LocEn Pres Unc Belnap 55 1949 Type locality (Fabaceae) cymboides (ARCH) is Huntington, Emery Co., UT (Jones 5464j Chapter 6:Appendices POM) Leguminosae Astragalus rimrock milkvetch PerF RegEn Pres Unc Welsh & 1947 Type locality is (Fabaceae) desperatus var. Moore 1911 near Cisco, Grand desperatus (ARCH) Co., UT (Jones s.n. POM) Leguminosae Astragalus fl avus A. fl avus var. yellow milkvetch PerF Wide Pres Rare Welsh & (Fabaceae) var. fl avus candicans Moore 1905 1963

(ARCH) 57 58

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Leguminosae Astragalus geyeri A. geyeri var. Geyer’s milkvetch AnnF Wide Pres Unc Allan 277 1973 (Fabaceae) geyeri (BRY) Leguminosae Astragalus straggling milkvetch PerF RegEn Pres Com Welsh & 1947 Type locality (Fabaceae) lentiginosus var. Moore 1908 is Montezuma palans (ARCH) Canyon, San Juan Co., UT (Eastwood s.n. POM) Leguminosae Astragalus Moenkopi milkvetch PerF RegEn Rep ? Schelz et al. NA (Fabaceae) moencoppensis (2006) Leguminosae Astragalus woolly milkvetch PerF Wide Pres Unc Welsh & 1947 Type locality is (Fabaceae) mollissimus var. Moore 1909 Kanab, Kane Co., thompsoniae (ARCH) UT (Thompson s.n. GH) Leguminosae Astragalus Ellis’ milkvetch PerF Wide Pres Unc Welsh & 1949 (Fabaceae) praelongus var. Moore 1904 ellisiae (ARCH) Leguminosae Astragalus preussii Included in var. Preuss’ milkvetch PerF RegEn Pres Com Welsh & 1947 Type locality (Fabaceae) var. latus preussii by some Moore 1912 is Green River, authors (ARCH) Emery Co., UT (Jones s.n. POM) Leguminosae Astragalus gravel milkvetch AnnF Wide Pres Rare Allan 402 1973 (Fabaceae) sabulonum (BRY) Leguminosae Caesalpinia repens Hoffmanseggia creeping rush-pea PerF RegEn Pres Rare Paspal 61 1988 Type locality (Fabaceae or repens (ARCH) is Courthouse Caesalpinaceae) Wash, Grand Co., UT (Eastwood s.n. CAS) Leguminosae Dalea fl avescens var. Kanab prairie-clover PerF RegEn Pres Unc Welsh & 1963 Type locality (Fabaceae) fl avescens Moore 2004 is “Kanab” (ARCH) (Thompson s.n. GH) Leguminosae Dalea oligophylla D. candida var. white prairie-clover PerF Wide Pres Unc Fowler & May 1952 (Fabaceae) oligophylla, 1695 (ARCH) Petalostemon candidum, P. occidentale, Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Leguminosae Glycyrrhiza lepidota G. lepidota var. Nuttall’s licorice PerF Wide Pres Unc Welsh, 1963 (Fabaceae) lepidota Harrison, & Moore 2344 (ARCH) Leguminosae Hedysarum boreale Includes var. northern PerF Wide Pres Unc Welsh & 1963 (Fabaceae) var. boreale pabulare sweetvetch Moore 1903 (ARCH) Leguminosae Lathyrus Zion sweetpea PerF RegEn Pres Unc Fagan s.n. 1988 Type locality is 10 (Fabaceae) brachycalyx var. (ARCH) miles E of Zion zionis NP, Kane Co., UT (Hitchcock 19013 isotype NY) Leguminosae Lupinus argenteus silvery lupine PerF Wide Pres Rare Welsh & 1949 (Fabaceae) var. argenteus Moore 2028 (ARCH) Leguminosae Lupinus brevicaulis shortstem lupine AnnF Wide Pres Rare Fagan 32 1985 (Fabaceae) (ARCH) Leguminosae Lupinus pusillus var. dwarf lupine AnnF Wide Pres Abun Welsh & 1948 (Fabaceae) pusillus Moore 1906 (ARCH) Leguminosae Medicago sativa M. sativa var. sativa alfalfa PerF Intro Pres Unc Fagan s.n. 1988 Native to Europe (Fabaceae) (ARCH) Leguminosae Melilotus alba M. albus. Included white sweet-clover AnnF Intro Pres Unc Fleming 827 1963 Native to Europe (Fabaceae) in M. offi cinalis by (ARCH) some authors Leguminosae Melilotus offi cinalis yellow sweet-clover PerF Intro Pres Unc Welsh & 1964 Native to Europe (Fabaceae) Moore 3015 (ARCH) Leguminosae Pediomelum Pediomelum large-fl owered PerF RegEn Pres Rare Hahn 2003 Chapter 6:Appendices (Fabaceae) megalanthum megalanthum var. breadroot LAH05140305 megalanthum, (ARCH) Psoralea megalantha 59 60

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Leguminosae Psoralidium Vars. not slenderleaf scurfpea PerF RegEn Hist NA Harrison 1949 Type locality (Fabaceae) lanceolatum var. recognized in 11367 (BRY) is near Wilson stenophyllum Barneby (1989) Mesa, Grand Co., UT (Rydberg & Garrett 8367 NY) Leguminosae Robinia black locust Tree Intro Pres Unc Welsh & 1963 Native to E North (Fabaceae) pseudoacacia Moore 1996 America (ARCH) Leguminosae Sophora stenophylla silvery sophora PerF Wide Pres Rare Fleming 781 1963 (Fabaceae) (ARCH) Liliaceae Allium San Juan onion PerF Wide Pres Unc Barnett 53 1963 (Alliaceae) macropetalum (ARCH) Liliaceae Allium nevadense Includes A. Nevada onion PerF Wide Pres Rare Fagan s.n. 1988 (Alliaceae) atrorubens var. (ARCH) cristatum by Welsh et al. (2003) Liliaceae Allium textile textile onion PerF Wide Pres Unc Fleming 967 1990 (Alliaceae) (ARCH) Liliaceae Androstephium pink funnel-lily PerF Wide Pres Rare Pyrah 31 1963 Type locality (Alliaceae, brevifl orum (ARCH) is near Kanab, Themidaceae) Kane Co., UT (Thompson s.n.) Liliaceae Asparagus asparagus PerF Intro Pres Unc Welsh & 1964 Native to Eurasia (Asparagaceae) offi cinalis Moore 2828 (BRY) Liliaceae Calochortus nuttallii sego-lily PerF Wide Pres Unc Welsh & 1949 (Calochortaceae) Moore 2051 (ARCH) Liliaceae Eremocrinum sand lily PerF RegEn Pres Rare Welsh 7025 1968 Type locality is (Anthericaceae) albomarginatum (BRY) near Green River, Emery Co., UT (Jones s.n. POM) Liliaceae Smilacina stellata Maianthemum starry false PerF Wide Pres Unc Pyrah 44 1963 (Convallariaceae) stellatum Solomon’s-seal (ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Liliaceae Zigadenus vaginatus Anticlea vaginatus alcove death camas PerF LocEn Pres Unc Welsh, 1963 Type locality is (Melanthiaceae) Harrison, & Natural Bridges Moore 2335 NM, San Juan (ARCH) Co., UT (Rydberg & Garrett 9407 NY). USFWS: 3C Linaceae Linum aristatum Mesynium broom-fl ax AnnF Wide Pres Unc Welsh & 1963 aristatum Moore 2767 (ARCH) Linaceae Linum perenne ssp. L. lewisii var. blue fl ax PerF Wide Pres Unc Fagan 24 1985 lewisii lewisii, Adenolinum (ARCH) lewisii Loasaceae Mentzelia albicaulis M. obscura, white-stem AnnF Wide Pres Unc Fagan 16 1955 Acrolasia albicaulis, blazingstar (ARCH) A. gracilis Loasaceae Mentzelia M. multifl ora var. desert stickleaf PerF Wide Pres Unc Welsh & 1949 multifl ora multifl ora, Nuttallia Moore 1982 multifl ora (ARCH) Loasaceae Mentzelia Nuttallia wing-seed stickleaf PerF Wide Pres Unc Pitts 75 1963 Type locality is pterosperma pterosperma (ARCH) Willow Creek, San Juan Co., UT (Eastwood 31 CAS) Loasaceae Mentzelia Acrolasia humilis, Thompson’s stickleaf AnnF RegEn Pres Rare Heil & Heil 1992 Type locality thompsonii Mentzelia humilis, 7028 (ARCH) is 7 miles E Nuttallia humilis of Montrose, Montrose Co., CO (Weber 7470) Lythraceae Lythrum salicaria purple loosestrife PerF Intro Pres Rare Topp 2006 Utah state

Chapter 6:Appendices ST08220601 noxious weed. (ARCH) Native to Europe Malvaceae Hibiscus trionum ﬂ ower-of-an-hour AnnF Intro Pres Rare Fertig 23006 2006 Native to Africa (ARCH) Malvaceae Sphaeralcea Includes ssp. scarlet globe- PerF Wide Pres Unc Pitts 58 1947 coccinea coccinea, elata, & mallow (ARCH) dissecta 61 62

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Malvaceae Sphaeralcea slenderleaf PerF Wide Pres Rare Welsh & 1963 leptophylla globemallow Moore 2011 (ARCH) Malvaceae Sphaeralcea small-leaf PerF Wide Pres Com Welsh & 1952 parvifolia globemallow Moore 1915 (ARCH) Moraceae Morus alba white mulberry Tree Intro Pres Unc Welsh & 1963 Native to China Moore 3014 (ARCH) Nyctaginaceae Abronia fragrans A. elliptica, A. fragrant sand- PerF Wide Pres Com Pyrah 27 1947 fragrans var. verbena (ARCH) elliptica, A. nana var. harrisii Nyctaginaceae Mirabilis linearis Oxybaphus linearis, narrow-leaf four- PerF Wide Pres Unc Welsh & 1963 var. linearis O. decumbens, o’clock Moore 1977 Includes M. glabra (ARCH) Nyctaginaceae Mirabilis multifl ora Includes vars. showy four-o’clock PerF Wide Pres Rare Welsh & 1964 multifl ora, Moore 3016 pubescens, & (BRY) glandulosa Nyctaginaceae Tripterocalyx Abronia micrantha, small-fl ower AnnF Wide Pres Unc Welsh & 1963 micranthus T. pedunculatus sandpuffs Moore 2819 (ARCH) Oleaceae Forestiera F. neomexicana desert olive Shrub Wide Pres Rare Hahn 2003 pubescens LAH08270302 (ARCH) Oleaceae Fraxinus anomala singleleaf ash Shrub Wide Pres Com Pitts 42 1947 Type locality (ARCH) is Labyrinth Canyon, San Juan Co., UT (Newberry s.n. NY) Onagraceae Camissonia Eastwood’s AnnF RegEn Pres Rare Welsh 7023 1968 eastwoodiae camissonia (BRY) Onagraceae Camissonia O. scapoidea Paiute suncup AnnF Wide Pres Unc Paspal 76 1991 scapoidea var. (ARCH) scapoidea Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Onagraceae Epilobium ciliatum E. ciliatum northern willow- PerF Wide Pres Unc Welsh, 1963 var. ciliatum, herb Harrison, & E. watsonii, E. Moore 2318 adenocaulon (ARCH) Onagraceae Gaura parviﬂ ora G. mollis small-ﬂ owered AnnF Wide Pres Unc Welsh & 1964 gaura Moore 2814 (ARCH) Onagraceae Oenothera long-tube evening- PerF Wide Pres Com Pitts 47 1949 caespitosa var. primrose (ARCH) marginata Onagraceae Oenothera Paria evening- PerF RegEn Pres Unc Pitts 59 1963 Type locality caespitosa var. primrose (ARCH) is Paria River, navajoensis Kane Co., UT (Cronquist 10202 MO) Onagraceae Oenothera Bridges evening- PerF RegEn Pres Unc Welsh, 1963 Type locality is longissima primrose Harrison, & Armstrong & Moore 2332 White canyons (BRY) [Natural Bridges NM], San Juan Co., UT (Rydberg & Garrett 9410 NY) Onagraceae Oenothera pallida Includes var. pale evening- PerF Wide Pres Com Fleming 846 1948 var. pallida runcinata primrose (ARCH) Orchidaceae Epipactis gigantea giant helleborine PerF Wide Pres Unc Welsh & 1948 Moore 2038 (ARCH) Orchidaceae Habenaria dilatata Platanthera Lindley’s white bog- PerF Wide Pres Rare Welsh & 1963 Cited as H. spar-

Chapter 6:Appendices var. leucostachys dilatata var. orchid Moore 2735 siﬂ ora in Harrison leucostachys, P. (ARCH) et al. (1964) leucostachys 63 64

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Orchidaceae Habenaria Platanthera alcove bog-orchid PerF RegEn Pres Unc Fleming 835 1989 Type locality is zothecina zothecina, (ARCH) ca 1 mile N of Limnorchis Moab, Grand zothecina Co., UT (Welsh & Higgins 23269 BRY). UT BLM: Sensitive, USFWS: former C2 candidate. Orobanchaceae Orobanche Aphyllon clustered PerF Wide Pres Rare Welsh & 1963 fasciculata fasciculatum broomrape Moore 2023 (BRY) Orobanchaceae Orobanche Includes O. Cooper’s broomrape PerF Wide Pres Rare Franklin 3821 1986 ludoviciana var. multifl ora, O. (BRY) cooperi cooperi Pinaceae Pinus edulis two-needle pinyon Tree Wide Pres Com Pitts 60 1963 (ARCH) Plantaginaceae Plantago major Includes var. common plantain PerF Intro Pres Unc Moran s.n. 2006 Mostly native pachyphylla (ARCH) to Europe. Type locality of the var. pachyphylla is Salt Lake City, Salt Lake Co., UT (Jones 1030 POM) Plantaginaceae Plantago Includes vars. woolly plantain AnnF Wide Pres Abun Pyrah 2 1963 patagonica patagonica, (ARCH) gnaphaloides, spinulosa, & breviscapa Polemoniaceae Eriastrum diffusum spreading eriastrum AnnF Wide Rep ? Schelz et al. NA (2006) Polemoniaceae Gilia inconspicua Includes var. shy gilia AnnF Wide Pres Unc Pitts 57 1963 If recognized, inconspica & (ARCH) park material is sinuata, G. all var. sinuata ophthalmoides, G. sinuata, G. clokeyi Polemoniaceae Gilia leptomeria var. Aliciella leptomeria common gilia AnnF Wide Pres Unc Fleming 951 1948 leptomeria (ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Polemoniaceae Gilia subnuda Aliciella subnuda carmine gilia PerF RegEn Rep ? Schelz et al. NA (2006) Polemoniaceae Ipomopsis Gilia aggregata var. scarlet gilia PerF Wide Pres Unc Fagan 37 1985 aggregata var. aggregata (ARCH) aggregata Polemoniaceae Ipomopsis Gilia gunnisonii Gunnison’s gilia AnnF RegEn Pres Com Welsh & 1963 Type locality is gunnisonii Moore 1983 “sand banks of (ARCH) the Green River”, UT (Cruetzfeldt s.n. NY) Polemoniaceae Ipomopsis Gilia polycladon spreading gilia AnnF Wide Pres Unc Fagan 15 1985 polycladon (ARCH) Polemoniaceae Ipomopsis roseata Gilia roseata San Rafael gilia PerF RegEn Pres Unc Welsh & 1963 Moore 1995 (ARCH) Polemoniaceae Leptodactylon Linanthus watsonii Watson’s prickly- PerF Wide Pres Unc Fagan 22 1949 Type locality is watsonii phlox (ARCH) Cottonwood Canyon, Salt Lake Co., UT (Watson s.n. GH) Polemoniaceae Phlox Var. not recognized desert phlox PerF Wide Rep ? Schelz et al. NA Type locality is austromontana var. in Cronquist et al. (2006) Beaver Dam Mts, austromontana (1984) Washington Co., UT (Fremont 447 US) Polygalaceae Polygala subspinosa P. subspinosa var. cushion milkwort PerF Wide Pres Rare Welsh, 1963 subspinosa Harrison, & Moore 2306 (BRY)

Chapter 6:Appendices Polygonaceae Eriogonum alatum E. alatum winged wild PerF Wide Pres Unc Welsh & 1952 var. alatum, buckwheat Moore 2046 Pterogonum (ARCH) alatum Polygonaceae Eriogonum bicolor pretty wild PerF RegEn Pres Unc Fleming 961 1947 Type locality buckwheat (ARCH) is Thompson’s Springs, Grand Co., UT (Jones s.n. POM) 65 66

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Polygonaceae Eriogonum Includes var. crisp-leaf wild Shrub Wide Pres Com Welsh & 1963 corymbosum var. velutinum and E. buckwheat Moore 2709 corymbosum lancifolium (ARCH) Polygonaceae Eriogonum rimrock wild Shrub RegEn Pres Unc Welsh, 1963 Type locality corymbosum var. buckwheat Harrison, & is Green River, orbiculatum Moore 2260 Emery Co., UT (ARCH) (Jones s.n. POM) Polygonaceae Eriogonum gordonii Gordon’s wild AnnF Wide Pres Unc Howell 44866 1963 buckwheat (NY) Polygonaceae Eriogonum infl atum E. fusiforme Grand Valley desert AnnF RegEn Pres Unc Fertig 22658 2006 var. fusiforme trumpet (ARCH) Polygonaceae Eriogonum infl atum E. infl atum desert trumpet PerF Wide Pres Unc Pyrah 47 1963 var. infl atum (ARCH) Polygonaceae Eriogonum sand wild Shrub RegEn Pres Unc Welsh, 1963 Reported as leptocladon var. buckwheat Harrison, & “Eriogonum sp.” leptocladon Moore 2265 in Harrison et (ARCH) al. (1964). Type locality is Green River, Emery Co., UT (Cruezfeldt s.n. NY) Polygonaceae Eriogonum slender wild Shrub Wide Pres Unc Welsh & 1963 microthecum var. buckwheat Moore 2733 laxifl orum (ARCH) Polygonaceae Eriogonum E. microthecum var. slender wild PerF Wide Pres Com Welsh & 1963 microthecum var. foliosum buckwheat Moore 1993 simpsonii (ARCH) Polygonaceae Eriogonum E. ovalifolium var. cushion wild PerF Wide Pres Unc Fleming 743 1989 ovalifolium ovalifolium, E. buckwheat (ARCH) ovalifolium var. purpureum; vars. not recognized by Welsh et al. (2003) Polygonaceae Eriogonum Stenogonum smooth wild AnnF Wide Pres Unc Schelz s.n. 2003 salsuginosum salsuginosum buckwheat (ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Polygonaceae Eriogonum Wetherill’s wild AnnF RegEn Pres Unc Forsythe 7 1964 Type locality is wetherillii buckwheat (ARCH) along the San Juan River, San Juan Co., UT (Eastwood 124 CAS) Polygonaceae Polygonum Persicaria willow-weed AnnF Intro Pres Rare Topp 2006 Native to Eurasia lapathifolium lapathifolia ST08080605 (ARCH) Polygonaceae Rumex crispus curly dock PerF Intro Rep Unc Dewey & 2003 Native to Eurasia Anderson (2005) Polygonaceae Rumex canaigre dock PerF Wide Pres Com Barnett 61 1948 hymenosepalus (ARCH) Polygonaceae Rumex venosus veiny dock PerF Wide Pres Unc Fagan 27 1979 (ARCH) Polypodiaceae Adiantum capillus- southern Fern Wide Pres Unc Pyrah 41 1947 (Adiantaceae) veneris maidenhair fern (ARCH) Polypodiaceae Cystopteris fragilis Includes C. brittle bladder fern Fern Wide Pres Unc Welsh, 1963 Schelz et al. (Aspleniaceae) utahensis, C. Harrison, & (2006) report C. reevesiana, & C. Moore 2320 utahensis, but tenuis (ARCH) ARCH specimen in currently on loan and could not be veriﬁ ed. Polypodiaceae Pellaea breweri Brewer’s cliff-brake Fern Wide Rep Rare Fowler et al. 1993 Report in (Adiantaceae, (2007) Harrison et al. Sinopteridaceae) (1964) is based on a misidentiﬁ ed

Chapter 6:Appendices specimen of P. glabella var. occidentalis. Polypodiaceae Pellaea glabella var. P. occidentalis western cliffbrake Fern Periph Pres Rare Welsh, 1963 (Adiantaceae, occidentalis Harrison, & Sinopteridaceae) Moore 2341 (ARCH) 67 68

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Polypodiaceae Pellaea limitanea Argyrochosma border cloak-fern Fern Periph Pres Rare Welsh & 1963 (Adiantaceae, limitanea ssp. Moore 2757 Sinopteridaceae) limitanea, (BRY) Notholanea limitanea Polypodiaceae Pteridium aquilinum P. aquilinum var. bracken fern Fern Wide Pres Unc Pitts 54 1963 (Dennstaedtiaceae, var. pubescens lanuginosum (ARCH) Hypolepidiaceae) Portulacaceae Portulaca oleracea common purslane AnnF Intro Pres Unc Moore 41 1963 Native to E North (BRY) America. Portulacaceae Talinum brevifolium Phemeranthus sausageleaf talinum PerF Wide Pres Rare Topp 2006 brevifolius ST05200601 (ARCH) Primulaceae Primula specuicola cave primrose PerF RegEn Pres Unc Barnett 50 1947 Type locality is (ARCH) along San Juan River near Bluff, San Juan Co., UT (Rydberg 9882 NY). USFWS: 3C Ranunculaceae Aquilegia micrantha alcove columbine PerF RegEn Pres Unc Pyrah 45 1947 Type locality is (Helleboraceae) (ARCH) Bluff, San Juan Co., UT (Wetherill s.n. CAS) Ranunculaceae Clematis white virgin’s-bower PerF Wide Pres Unc Welsh, 1963 ligusticifolia Harrison, & Moore 2322 (ARCH) Ranunculaceae Delphinium D. scaposum pale larkspur PerF Wide Pres Unc Welsh & 1949 (Helleboraceae) andersonii var. Moore 1924 scaposum (ARCH) Ranunculaceae Delphinium D. nelsonii Nuttall’s larkspur PerF Wide Pres Unc Forsythe 9 1979 (Helleboraceae) nuttallianum (ARCH) Appendix A. Conﬁ rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientiﬁ c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Ranunculaceae Ranunculus R. cymbalaria marsh buttercup PerF Wide Pres Unc Belnap 525 1979 cymbalaria var. cymbalaria, (ARCH) Halerpestes cymbalaria ssp. saximontana. Vars. not recognized in Flora of North America (1997) Ranunculaceae Ranunculus Ceratocephala bur buttercup AnnF Intro Pres Unc Paspal 75 1991 Native to Eurasia testiculatus orthoceras, C. (ARCH) testiculata Rhamnaceae Rhamnus betulifolia Frangula betulifolia birchleaf buckthorn Shrub Wide Pres Unc Welsh & 1963 Moore 2760 (ARCH) Rosaceae Amelanchier Utah serviceberry Shrub Wide Pres Unc Barnett 58 1947 Type locality utahensis (ARCH) is Toquerville, Washington Co., UT (Jones 1716 GH) Rosaceae Cercocarpus C. ledifolius var. dwarf mountain Shrub Wide Pres Com Pitts 39 1948 Type locality is intricatus intricatus mahogany (ARCH) American Fork Canyon, Utah Co., UT (Watson 314 US) Rosaceae Cercocarpus alder-leaf mountain Shrub Wide Pres Com Pyrah 58 1947 montanus mahogany (ARCH) Rosaceae Coleogyne blackbrush Shrub Wide Pres Abun Barnett 64 1947 ramosissima (ARCH) Rosaceae Holodiscus dumosus H. discolor bush oceanspray Shrub Wide Pres Unc Fleming 770 1988

Chapter 6:Appendices (ARCH) Rosaceae Petrophyton Spiraea caespitosa rock spiraea PerF Wide Pres Unc Welsh & 1964 caespitosum Moore 2829 (BRY) Rosaceae Physocarpus dwarf ninebark Shrub Wide Pres Unc Fleming 771 1979 alternans (ARCH) 69 70

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Rosaceae Purshia mexicana P. stansburiana, cliff-rose Shrub Wide Pres Com Barnett 39 1963 Type locality is var. stansburyana Cowania mexicana (ARCH) Great Salt Lake, Davis Co., UT (Stansbury s.n. NY) Rosaceae Purshia tridentata antelope Shrub Wide Rep Rare Schelz et 2006 bitterbrush al. (2006), Northern Colorado Plateau Network (in ed.) Rosaceae Rosa woodsii R. woodsii var. Woods’ rose Shrub Wide Pres Unc Welsh, 1963 ultramontana, Harrison, & Welsh et al. (2003) Moore 2309 include R. manca (ARCH) Rubiaceae Galium aparine var. G. spurium cleavers AnnF Wide Pres Rare Topp 2006 echinospermon ST05020602 (ARCH) Rubiaceae Galium multifl orum G. coloradoense Colorado bedstraw PerF RegEn Pres Unc Welsh & 1963 var. coloradoense Moore 2741 (BRY) Rubiaceae Galium multifl orum G. multifl orum shrubby bedstraw PerF Wide Pres Rare Topp 2006 var. multifl orum ST08080601 (ARCH) Rubiaceae Galium trifl orum sweet-scented PerF Wide Pres Unc Welsh, 1963 bedstraw Harrison, & Moore 2319 (BRY) Salicaceae Populus alba white poplar Tree Intro Pres Unc Welsh & 1963 Native to Eurasia Moore 1997 (ARCH) Salicaceae Populus fremontii P. deltoides var. Fremont Tree Wide Pres Com Barnett 68 1947 var. fremontii fremontii cottonwood (ARCH) Salicaceae Populus x Hybird of P. Carolina poplar Tree Intro Pres Unc Welsh & 1963 canadensis deltoides x P. nigra Moore 2747(ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Salicaceae Populus x P. acuminata, P. x lance-leaf Tree Wide Pres Unc Topp 2006 Type locality intercurrens acuminata. Hybrid cottonwood ST05020604 is Jesse Ewing of P. fremontii x P. (ARCH) Canyon, Daggett angustifolia Co., UT (Goodrich 24441 BRY) Salicaceae Salix amygdaloides peach-leaf willow Tree Wide Pres Unc Welsh & 1964 Moore 2820 (ARCH) Salicaceae Salix eriocephala S. lutea, S. rigida yellow willow Shrub Wide Pres Unc Harms s.n. 1963 var. watsonii var. watsonii (ARCH) Salicaceae Salix exigua var. S. exigua var. coyote willow Shrub Wide Pres Unc Harms s.n. 2005 stenophylla exigua (ARCH) Salicaceae Salix gooddingii S. nigra var. Goodding’s black Tree Wide Pres Unc Schelz s.n. 1963 vallicola willow (ARCH) Salicaceae Salix lucida ssp. S. lasiandra var. whiplash willow Shrub Wide Pres Unc Barnett 70 1963 caudata caudata (ARCH) Santalaceae Comandra C. pallida bastard toadﬂ ax PerF Wide Pres Unc Barnett 63 1947 umbellata var. (ARCH) pallida Saxifragaceae Fendlera rupicola F. falcata fendlerbush Shrub Wide Pres Unc Welsh & 1963 (Hydrangeaceae) Moore 2752 (ARCH) Saxifragaceae Philadelphus P. microphyllus littleleaf mock- Shrub Wide Pres Rare Topp 2006 (Hydrangeaceae) microphyllus var. occidentalis, orange ST05150602 P. occidentalis var. (ARCH) occidentalis

Scrophulariaceae Castilleja chromosa C. angustifolia desert paintbrush PerF Wide Hist NA Harrison 1947 var. dubia, C. 11131 (BRY) Chapter 6:Appendices applegatei ssp. martinii Scrophulariaceae Castilleja exilis C. minor ssp. minor annual paintbrush AnnF Wide Pres Unc Fowler & May 1993 1652 (ARCH) Scrophulariaceae Castilleja linariifolia C. linariaefolia Wyoming PerF Wide Pres Unc Welsh & 1947 paintbrush Moore 2039 (ARCH) Scrophulariaceae Castilleja scabrida Eastwood’s PerF RegEn Pres Unc Heil 5791 1963 var. scabrida paintbrush (ARCH) 71 72

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Scrophulariaceae Cordylanthus Wright’s bird’s-beak AnnF Wide Pres Unc Belnap 449 1963 wrightii (ARCH) Scrophulariaceae Linaria dalmatica L. genistifolia ssp. Dalmatian toadfl ax PerF Intro Pres Unc Welsh & 1964 Native to Europe dalmatica Moore 2804 (ARCH) Scrophulariaceae Mimulus Eastwood’s monkey- PerF RegEn Pres Unc Welsh & 1963 Type locality is eastwoodiae fl ower Moore 2037 San Juan River (ARCH) near Bluff, San Juan Co., UT (Rydberg 9883 NY) Scrophulariaceae Mimulus glabratus M. glabratus var. glabrous PerF Wide Pres Rare Fagan 36 1985 Type locality var. utahensis glabratus monkeyfl ower (ARCH) is Preuss Lake, Millard Co., UT (Tidestrom 11180) Scrophulariaceae Penstemon barbatus Welsh et al. (2003) beardlip penstemon PerF Wide Rep ? Schelz et al. NA var. barbatus include var. torreyi (2006) Scrophulariaceae Penstemon bluestem PerF RegEn Pres Rare Fleming 958 1952 cyanocaulis penstemon (ARCH) Scrophulariaceae Penstemon eatonii undulate PerF Wide Pres Unc Pitts 72 1963 Type locality var. undosus penstemon (ARCH) is St. George, Washington Co., UT (Jones 5110h US) Scrophulariaceae Penstemon Loa penstemon PerF RegEn Pres Unc Fleming 960 1979 Type locality is ophianthus (ARCH) Thurber, Wayne Co., UT (Jones 5708 US) Scrophulariaceae Penstemon Utah penstemon PerF Wide Pres Rare Pitts 50 1947 Type locality utahensis (ARCH) is between Hatch’s Wash & Monticello, San Juan Co., UT (East-wood s.n. CAS) Scrophulariaceae Verbascum thapsus woolly mullein PerF Intro Pres Unc Moran s.n. 2006 (ARCH) Appendix A. Confi rmed, historical, and reported vascular plant taxa of Arches National Park, organized by family and scientifi c name with taxonomic and biogeographic notes, cont.

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Scrophulariaceae Veronica anagallis- water speedwell PerF Intro Pres Rare Fleming 954b 1990 Native to Europe aquatica (ARCH) Solanaceae Datura wrightii D. meteloides angel’s trumpet AnnF Wide Pres Rare Topp 2006 ST05080601 (ARCH) Solanaceae Physalis hederifolia P. fendleri, P. Fendler’s ground- PerF Wide Pres Unc Welsh & 1964 var. fendleri hederifolia var. cherry Moore 2815 cordifolia, P. (ARCH) hederaefolia var. fendleri Solanaceae Physalis longifolia P. virginiana var. long-leaf ground- PerF Wide Pres Unc Fertig 23007 2006 sonorae cherry (ARCH) Tamaricaceae Tamarix chinensis T. pentandra, T. ﬁ ve-stamen Shrub Intro Pres Com Welsh & 1963 Native to Eurasia ramosissima tamarisk Moore 2729 (ARCH) Typhaceae Typha domingensis southern cattail PerG Wide Pres Unc Welsh, 1963 Cited as T. Harrison, & latifolia in Moore 2280 Harrison et al. (BRY) (1964). Typhaceae Typha latifolia broad-leaved cattail PerG Wide Pres Unc Fleming 840 1989 (ARCH) Ulmaceae Celtis reticulata C. laevigata var. netleaf hackberry Tree Wide Pres Unc Welsh & 1963 (Celtidaceae) reticulata Moore 2045 (ARCH) Ulmaceae Ulmus pumila Siberian elm Tree Intro Pres Unc Welsh & 1963 Native to Asia Moore 2746 (BRY) Umbelliferae Cymopterus acaulis C. fendleri, C. Fendler’s PerF Wide Pres Unc Fleming 975 1988 (Apiaceae) var. fendleri glomeratus cymopterus (ARCH) Chapter 6:Appendices Umbelliferae Cymopterus onion spring-parsley PerF Wide Pres Unc Fagan 21 1985 (Apiaceae) bulbosus (ARCH) Umbelliferae Cymopterus sticky spring-parsley PerF RegEn Pres Unc Fleming 957 1948 (Apiaceae) newberryi (ARCH)

Umbelliferae Cymopterus basin white-cup PerF Wide Pres Unc Belnap 504 1988 (Apiaceae) purpurascens spring-parsley (ARCH) 73 74

Synonyms/ Life Park Pop. Year Family Species name Common name Range Source Comments Taxonomic notes form status size doc. Umbelliferae Cymopterus C. purpureus Colorado Plateau PerF RegEn Pres Unc Hahn 1990 (Apiaceae) purpureus var. spring-parsley LAH05140309 purpureus (ARCH) Umbelliferae Lomatium latilobum Aletes latilobus Canyonlands PerF LocEn Pres Unc Heil 5794 1947 Type locality is (Apiaceae) lomatium (ARCH) Wilson Mesa, Grand Co., UT (Rydberg & Garrett 8371 NY). USFS: Region 4 Sensitive, UT BLM: Sensitive, USFWS: former C2 candidate Umbelliferae Lomatium parryi Parry’s lomatium PerF Wide Pres Unc Pyrah 61 1949 Type locality is (Apiaceae) (ARCH) “Southern Utah” (Parry 85) Verbenaceae Verbena bracteata prostrate vervain PerF Wide Rep ? Schelz et al. NA (2006) Violaceae Viola nephrophylla Viola sororia var. northern bog violet PerF Wide Pres Rare Belnap 524 1988 affi nis (ARCH) Viscaceae Arceuthobium pinyon dwarf- PerF Wide Rep ? Schelz et al. NA Typically parasitic divaricatum mistletoe (2006) on Pinus edulis and P. monophylla Viscaceae Phoradendron juniper mistletoe PerF Wide Rep ? Schelz et al. NA Typically parasitic juniperinum (2006) on Juniperus osteosperma Zygophyllaceae Kallstroemia yellow kallstroemia AnnF Periph Pres Rare Schelz s.n. 2002 californica (ARCH) Zygophyllaceae Kallstroemia orange kallstroemia AnnF Periph Pres Rare Moran s.n. 2006 parvifl ora (ARCH) Zygophyllaceae Tribulus terrestris puncture vine AnnF Intro Pres Unc Welsh & 1963 Native to Eurasia Moore 2532 (ARCH) Mesa Co., CO is San Rafael Swell, Emery Co., UT (Jones s.n. CAS) Co., UT (Jones s.n. POM) Range Comments Life form hempcommon hopsRocky Mountain beeplant AnnF Wide PerF Wide PerF Intro Native to Eurasia aspen bluebell PerF Wide dwarf cryptanthsouthwestern stoneseed PerF PerF Wide Wide slender cryptanth AnnF Wide Cutler’s milkweedCutler’s milkweedRusby’s barberryFendler’s PerFwestern hop-hornbeam PerF RegEn obscure cryptanth Tree RegEn cushion cryptanth Shrub RegEn tall cryptanth RegEn AnnF AnnF Sparse Wide PerF LocEn locality is Grand Junction, Type spreading dogbaneJones’ cycladenia PerF Wide PerF RegEn locality USFWS: Threatened. Type Common name datil yucca yuccaBailey’s sea purslanenorthern arrowheadJones’ amsonia Shrub PerF Shrub Wide Periph PerF Wide PerF Wide RegEn Palmer’s stinkweedPalmer’s AnnF RegEn Emery locality is Green River, Type . . var ssp . . baccata . var nana, ssp var H. lupulus americanus grahamii nana Includes var. Includes var. Oreocarya humilis rusbyi C. jonesii Synonyms/Taxonomic Synonyms/Taxonomic notes . . . var var var National Park. rmed) taxa of Arches orum . palmeriana Humulus lupulus Cleome serrulata Cleomella palmeriana var neomexicanus Cannabis sativa Lithospermum multifl Mertensia arizonica M. arizonica Cryptantha gracilis Cryptantha humilis Asclepias cutleri Asclepias rusbyiBerberis fendleri Ostrya knowltonii A. engelmanniana Cryptantha ambigua Cryptantha circumscissa Cryptantha elata Oreocarya elata Cycladenia humilis androsaemifolium jonesii Yucca baileyi Yucca Sesuvium verrucosum Sagittaria cuneata Amsonia jonesii Apocynum androsaemifolium Yucca baccataYucca baccata Y. Cannabaceae Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) Cannabaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Asclepiadaceae Asclepiadaceae Berberidaceae Betulaceae Boraginaceae Boraginaceae Boraginaceae Apocynaceae Agavaceae Aizoaceae Alismataceae Apocynaceae Apocynaceae FamilyAgavaceae Species name Appendix B. Potential (unconfi

Chapter 6: Appendices 75 River, Grand Co., UT (Stutz 9547 River, BRY) along UT Hwy 128, Grand Co., (Hanson 322 BRY) Range Comments Life form y PerF Wide Arizona siltbush Shrub RegEn barbwire Russian thistle AnnF Intro Native to Eurasia mountain goosefoot AnnF Wide Powell’s orachPowell’s thinleaf orach AnnF Wide AnnF Wide Green River orach AnnF LocEn locality is 3 miles E of Green Type Welsh’s saltbushWelsh’s Shrub LocEn locality is 4 miles S of Cisco Type Drummond’s catchﬂ Drummond’s mountain snowberry Shrub Wide clammy-weed AnnFball-head sandwort Wide PerF Wide Common name common yarrow PerF Wide & . .

ssp . (2006). var includes Flora of parishii . Vars. not Vars. var. var in Cronquist var includes vars. cephaloidea . congesta, . drummondii, drummondii, Z. brandegeei arizonica atrovirens A. powellii triangularis A. minuticarpa A. welshii var. var. striata, striata, kruckebergii, & Gastrolychnis drummondii S. rotundifolius. Includes vars. oreophilus et al. (1984) Eremogone congesta var P. trachysperma P. ssp S. drummondii Synonyms/Taxonomic Synonyms/Taxonomic notes A. lanulosa, millefolium occidentalis. recognized in North America . . . var . .

var . var var var var var. . utahensis var rmed) taxa of Arches National Park, cont. Achillea millefolium arizonica Zuckia brandegei Salsola paulsenii Atriplex powellii Atriplex prostrataChenopodium atrovirens C. fremontii A. patula powellii Atriplex powellii minuticarpa Atriplex gardneri welshii Arenaria congesta congesta trachysperma Symphoricarpos oreophilus drummondiiLychnis Silene drummondii lanulosa Polanisia dodecandra Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Caryophyllaceae (Alsinaceae) Caprifoliaceae Caryophyllaceae Appendix B. Potential (unconﬁ FamilyCapparaceae (Cleomaceae) Species name

76 Annotated Checklist of Vascular Flora: Arches National Park Potash plant). Type locality is 1 mi Potash plant). Type San Juan Co., E of San Juan River, UT (Atwood et al. 22406 BRY). Range Comments Life form ower brickellbush ower PerF Wide western aster PerF Wide California brickellbush Shrub Wide white heath aster PerFalkali aster Wide PerF Wide Common name tassel-ﬂ Atwood’s mule-fatAtwood’s Shrub LocEn Known from vicinity of park (S skeleton-leaf bursagefringed sagebrush PerF Wide Wyoming big sagebrush Shrub Shrub asterEaton’s Wide Wide leafy asterblueleaf aster PerF Wide AnnF Wide PerF Wide cutleaf agoseris PerF Wide orange agoseris PerF Wide hoary dusty-maiden PerF Wide

& orus . pansus, douglasii . pansum, var B. salicifolia . glauca var ora (2006) var wyomingense montana Flora of North includes var. includes var. Symphyotrichum spathulatum Included in in America Symphyotrichum ericoides Virgulus ericoides Synonyms/Taxonomic Synonyms/Taxonomic notes Seriphidium tridentatum ssp. eatonii, A. bracteolatus frondosum, Brachyactis frondosa Eurybia glauca, Herrickia glauca A. parvifl var. . . . var var . var ora var oruspaucifl Almutaster rmed) taxa of Arches National Park, cont. Chaenactis douglasii Baccharis viminea Brickellia californica Brickellia grandifl atwoodii Aster paucifl Aster spathulatus A. occidentalis, Artemisia frigida Artemisia tridentata Aster eatoniiAster frondosusAster glaucodes Symphyotrichum Symphyotrichum Aster pansus Eucephalus glaucus, A. ericoides wyomingensis Ambrosia tomentosa Franseria tomentosa laciniata Agoseris glauca Agoseris aurantiaca aurantiaca Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi FamilyCompositae (Asteraceae) Species name

Chapter 6: Appendices 77 ed San Juan Co., UT (Newberry s.n. GH) verifi (Baker, Earle, & Tracy 90) Earle, & Tracy (Baker, Lake, UT (Stansbury s.n. GH) Juan Co., UT (Eastwood 45 CAS) Range Comments Life form ower PerF RegEn locality is Abajo Mountains, Type ower PerF Wide Specimen on loan - cannot be Clement’s goldenweedClement’s PerFpurpledisk sunfl Wide sunfl Nuttall’s curly-cup gumweed PerF Wide trailing daisywhite thoroughwort PerF PerF Wide Wide Eaton’s daisyEaton’s PerF Wide poverty-weed PerF Wide Tracy’s thistleTracy’s PerF RegEn locality is Mancos, Colorado Type Torrey’s desert-dandelionTorrey’s AnnF Wide locality is shores of Great Salt Type Common name Greene’s rabbitbrushGreene’s rabbitbrushVasey’s Shrub Wide puberulent rabbitbrush Shrub Shrub Wide Wide chicory PerF Intro Native to Eurasia aristate aster PerF RegEn locality is Willow Creek, San Type .

var ssp. var. axillaris. vacans, eatonii var. var. var. clementis nuttallii Vars. not recognized in Vars. Flora of North America (2006) M. linearis, rigida, canescens Dieteria canescens Vars. not recognized in Vars. Flora of North America (2006). C. tracyi Synonyms/Taxonomic Synonyms/Taxonomic notes aristata . . var var

. aristata var. var agellaris orus rmed) taxa of Arches National Park, cont. Helianthella microcephala Helianthus nuttallii Helianthus nuttallii Haplopappus clementis Pyrrocoma clementis serrulata Eupatorium herbaceum Ageratina herbacea Grindelia squarrosa Iva axillaris Iva axillaris Erigeron fl Erigeron eatonii E. eatonii Machaeranthera canescens tracyi Malacothrix torreyi Chrysothamnus vaseyi Chrysothamnus viscidifl Cichorium intybus Cirsium undulatum Chrysothamnus greenei puberulus Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi FamilyCompositae (Asteraceae) Species name

78 Annotated Checklist of Vascular Flora: Arches National Park collected outside the park. Type collected outside the park. Type locality is near Kanab, Kane Co., UT (Thompson s.n. G) Range Comments Life form ower PerF Sparse ax-leaved plainsmustard ax-leaved PerF Wide marsh sow-thistlegolden crownbeard PerFshowy goldeneye Intro AnnF Native to Europe Wide PerFArizona mulesears Wide Sweetwater rockcress PerFshepherd’s-purse PerF RegEn Sparse AnnF Intro Native to Europe showy goldeneye PerF Wide western tansy-mustard AnnF Wide western tansy-mustard drabaWoods AnnF Wide mountain draba AnnF AnnF Intro RegEn Native to Eurasia prairie conefl prairie pepperwortfl AnnF Wide Common name prenanthellapiliferous turtleback AnnF AnnF RegEn Wide Specimen at ARCH herbarium was ora ora by some . . in Welsh in Welsh D. pinnata ssp var ora . multifl . nevadensis . osmiarum . uliginosus glabrescens, S. arvensis ssp Ximenesia encelioides Heliomeris multifl var Heliomeris multifl var Included in var authors D. pinnata halictorum Includes var. Includes var. macrocarpum et al. (2003) Synonyms/Taxonomic Synonyms/Taxonomic notes

. . . .

var. var var var var var. orum ora ora ora ora orum rmed) taxa of Arches National Park, cont. Verbesina encelioides Verbesina Viguiera multifl Viguiera multifl nevadensis Arabis ligniferaCapsella bursa-pastoris Descurainia pinnata Boechera lignifera multifl intermedia exauriculata Wyethia arizonica Descurainia pinnata osmiarum Draba nemorosa Draba rectifructa Ratibida columnifera Sonchus uliginosus S. arvensis densifl Schoenocrambe linifolia Sisymbrium linifolium Psathyrotes pilifera Prenanthella exigua exigua Lygodesmia Lepidium densifl Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Appendix B. Potential (unconfi FamilyCompositae (Asteraceae) Species name (Brassicaceae)

Chapter 6: Appendices 79 is marginale eld Co., UT (Rydberg & Aquarius Plateau, head of Poison Creek, Garfi Carlton 7401 NY) Virgin, Kane Co., UT (Siler s.n. ISC) Co., UT (Jones s.n. POM) Co., UT (Jones s.n. POM) Range Comments Life form at-sedge AnnG Wide eld’s bluestem eld’s PerG Sparse ower dragonhead ower PerF Wide Hayden’s milkvetchHayden’s milkvetchEastwood’s PerF PerF Wide RegEn Grand locality is Westwater, Type American bugleweed PerF Wide stinkgrassbearded sprangletopgreen muhly AnnGwater polypogon Wide Indiangrass AnnG Introtall dropseed PerG PerG Native to Eurasia Intro Wide Native to Eurasia and Africa compressed rush PerGsmallfl PerG Wide Sparse PerG Intro Native to Europe toad rush AnnG Wide Springfi Nevada ephedrasmall-leaf geranium Shrub PerF Wide Wide locality of var. Type silver buffaloberryroundleaf buffaloberry Shrub Shrub RegEn Wide locality is upper valley of the Type Common name Diehl’s milkvetchDiehl’s PerF RegEn locality is Farnham, Carbon Type bearded fl smoothleaf prince’s-plume PerF Wide

. ora var. var bufonius, . var fascicularis var. ambiguus A. haydenianus fusca Agrostis semiverticillata, viridis P. Includes vars. occidentalis, halophilus, & Moldavica parviﬂ G. caespitosum atropurpureum, G. atropurpureum, G. atropurpureum cowenii Synonyms/Taxonomic Synonyms/Taxonomic notes integrifolia . .

var var eldii exuosus orum rmed) taxa of Arches National Park, cont. . marginale Astragalus eastwoodiae haydenianus Astragalus bisulcatus Lycopus americanus Lycopus Eragrostis cilianensis Leptochloa fascicularis Diplachne fascicularis, L. Muhlenbergia racemosa Polypogon semiverticillatus Sorghastrum nutansSporobolus asperJuncus bufonius S. avenaceum S. compositus Dracocephalum parvifl Juncus compressus Bothriochloa springfi Ephedra nevadensis Geranium caespitosum var Shepherdia rotundifolia Shepherdia argentea Astragalus fl diehlii Stanleya integrifolia S. pinnata Cyperus squarrosus C. aristatus Leguminosae (Fabaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Juncaceae Labiatae (Lamiaceae) Juncaceae Gramineae (Poaceae) Ephedraceae Geraniaceae Elaeagnaceae Elaeagnaceae Leguminosae (Fabaceae) Appendix B. Potential (unconfi FamilyCruciferae (Brassicaceae) Species name Cyperaceae

80 Annotated Checklist of Vascular Flora: Arches National Park miles NE of Moab, Grand Co., UT (Shultz & Shultz 2070 UTC). UT BLM: Sensitive, USFWS: former C2 candidate Blanding and 10 miles E of Hite, San Juan Co., UT (Cronquist 9030 NY) Wasatch Co., UT (Watson 234 GH) Co., UT (Watson Wasatch Ranch, Grand Co., UT (Welsh & Ranch, Grand Co., UT (Welsh 14632 BRY) Taylor UT (Jones s.n. POM). UTM BLM: Sensitive, USFWS: former C2 candidate The Jump, Canyonlands NP, San The Jump, Canyonlands NP, Moore, & Juan Co., UT (Welsh, Canter 2979 BRY) Range Comments PerF Wide Life form owered milkvetch owered AnnF RegEn ower sweetpea ower PerF RegEn gooseberry-leaf globemallow Shultz’ stickleaf PerF LocEn 11 locality is Fisher Valley, Type Colorado stickleaf PerF RegEn locality is 75 miles W of Type Gunnison’s mariposaGunnison’s PerF Wide American vetch PerF Wide King’s lupineKing’s AnnF Wide locality is Heber Valley, Type Moab lupine PerF RegEn locality is Professor Valley Type whiteﬂ Cisco milkvetch PerF LocEn locality is Cisco, Grand Co., Type Fisher milkvetch PerF LocEn locality is Salt Canyon above Type Common name pliant milkvetchsmall-ﬂ PerF Periph . var . . var var S. grossulariifolia grossulariifolia cronquistii, M. cronquistii, Nuttallia cronquistii gunnisonii L. leucanthus Synonyms/Taxonomic Synonyms/Taxonomic notes

. . . var var. var . var . kingii L. kingii var var exuosus rmed) taxa of Arches National Park, cont. . micranthiformis exuosus Sphaeralcea grossulariifolia Mentzelia shultziorum Mentzelia marginata M. marginata Calochortus gunnisonii C. gunnisonii americana Vicia americana Lupinus kingii moabensis Lupinus argenteus Lathyrus lanszwertii leucanthus Astragalus sabulosus sabulosus Astragalus piscator Astragalus nuttallianus var Astragalus fl fl Malvaceae Loasaceae Loasaceae Liliaceae (Calochortaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix B. Potential (unconfi FamilyLeguminosae (Fabaceae) Species name

Chapter 6: Appendices 81 Colorado River along Hwy 129, 6857 BRY) Grand Co., UT (Welsh of Green River, Emery Co., UT of Green River, (Ownbey & Ownbey 2146 ) Montrose Co., CO Grand Co., UT (Welsh, Welsh, & Welsh, Grand Co., UT (Welsh, 16689 BRY) Welsh miles WNW of Hanksville, Wayne miles WNW of Hanksville, Wayne UT 13348 BRY). Co., UT (Welsh BLM: Sensitive, USFWS: former C2 candidate Road, Canyonlands National Park, 7064 BRY). San Juan Co., UT (Welsh UT BLM: Sensitive, USFWS: former C2 candidate Range Comments PerF Wide AnnF Wide Life form dwarf gilia AnnF Wide long-leaf phlox PerF Wide San Rafael prickly-poppy PerF LocEn along locality is bluffs Type hairy-throat evening- primrose yellow evening-primroseSan Rafael prickly-poppy PerF PerF Wide RegEn locality is 37.9 miles SW Type branching groundsmoke AnnFwhite-stem evening- Wide primrose autumn willow-herb AnnF Wide Walker’s camissoniaWalker’s AnnF Wide locality is Paradox Valley, Type lavender-leaf sundropslavender-leaf PerF Wide clay sand-verbena PerF RegEn locality is 6 miles S of Cisco, Type scurfpea globemallow PerF LocEn ca 17 locality is Salt Wash, Type Common name Jane’s globemallowJane’s PerF LocEn locality is along White Rim Type

var. longifolia ssp. ava janeae stansburyi Includes vars. & paniculatum lavandulifolius Synonyms/Taxonomic Synonyms/Taxonomic notes var.

var. var. avafl O. fl var. ava rmed) taxa of Arches National Park, cont. Ipomopsis pumilaPhlox longifolia Gilia pumila Oenothera fl Argemone corymbosa parva Oenothera coronopifolia Argemone corymbosa arenicola Oenothera albicaulis Gayophytum ramosissimum Epilobium brachycarpum E. paniculatum Camissonia walkeri Calylophus lavandulifolius C. hartwegii Abronia argillosa Sphaeralcea psoraloides Sphaeralcea janeae Sphaeralcea leptophylla Polemoniaceae Polemoniaceae Onagraceae Papaveraceae Onagraceae Papaveraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Nyctaginaceae Malvaceae Appendix B. Potential (unconfi FamilyMalvaceae Species name

82 Annotated Checklist of Vascular Flora: Arches National Park E. gordonii. City, Millard Co., UT (Holmgren & City, Holmgren 11298 NY) Juan Co., UT (Welsh, Welsh, & Welsh, Juan Co., UT (Welsh, 22244 BRY) Welsh is Range Comments Shrub Wide locality is ca 10 km E of Oak Type Shrub RegEn Life form ower AnnF Wide ower rock-jasmine ower AnnF Wide Rio Grande cottonwood Tree Periph curl-leaf mountain mahogany blister buttercup desert-lilacFranklin’s AnnF Shrub Wide LocEn locality is Muley Point, San Type Oregon woodsiapygmy-ﬂ Fern Periph redroot wild buckwheat PerF Wide Palmer’s wild buckwheatPalmer’s AnnF Wide yard knotweedslender lip-fern AnnF Intro Fern Native to Eurasia Wide sulfur wild buckwheat PerF Wide Grand Valley wild Grand Valley buckwheat Common name nodding wild buckwheat AnnF Wide Report for ARCH from Barnett (67) blue-eyed Mary AnnF Wide seep monkeyﬂ coyote tobaccosilverleaf nightshade PerF AnnF Intro Wide Native to C North America blunt-leaf spike-moss Fern Wide

. . oregana . & var. mutica aviculare aureum var ssp var var. ssp. (2005) Flora of North P. deltoides P. wislizenii C. ledifolius intercedens Hecatonia scelerata Includes ssp. puberulenta, subumbellata, subilifera Includes var. Includes var. from America E. racemosum aviculare Synonyms/Taxonomic Synonyms/Taxonomic notes Vars. not recognized in Vars. Flora of North America (2005)

var. . var. var. var ora oribundus dus rmed) taxa of Arches National Park, cont. umbellatum racemosum . intermontanus wislizenii Collinsia parvifl Populus fremontii Cercocarpus ledifolius var Ranunculus sceleratus Ceanothus greggii multifi franklinii Androsace septentrionalis Eriogonum umbellatum var. Eriogonum racemosum var. Eriogonum palmerianum Cheilanthes feei oregana Woodsia oregana W. Polygonum aviculare arenastrum, P. P. Eriogonum contortum Eriogonum cernuum cernuum Mimulus fl Solanum elaeagnifolium Selaginella mutica Nicotiana attenuata S. mutica Scrophulariaceae Salicaceae Rosaceae Ranunculaceae Rhamnaceae Primulaceae Polygonaceae Polygonaceae Polygonaceae Polypodiaceae Polypodiaceae Polygonaceae Polygonaceae Appendix B. Potential (unconfi FamilyPolygonaceae Species name Scrophulariaceae Solanaceae Selaginellaceae Solanaceae

Chapter 6: Appendices 83 (Albee et al. 1988), and rmed for the park. Species on this Range Comments Atlas of the Vascular Flora of Utah Atlas of the Vascular Life form ity, in 2001, the ity, ave not previously been reported or conﬁ been reported ave not previously ed here. Common name rock spring-parsleysouthern ligusticum PerF Wide PerF Wide Synonyms/Taxonomic Synonyms/Taxonomic notes C. petraeus, Aletes petraea, Pteryxia petraea rmed) taxa of Arches National Park, cont. . petraeus Ligusticum porteri Cymopterus terebinthinus var Umbelliferae (Apiaceae) Appendix B. Potential (unconﬁ FamilyUmbelliferae (Apiaceae) Species name The vascular plant species shown in this table are known from the vicinity of Arches National Park in Grand County, Utah, but h National Park in Grand County, the vicinity of Arches known from The vascular plant species shown in this table are list are derived from a checklist of potential species for NCPN parks prepared by Dr. N. Duane Atwood, of Brigham Young Univers N. Duane Atwood, of Brigham Young by Dr. a checklist of potential species for NCPN parks prepared derived from list are not includ as potentially occurring in the park, but are Appendix C could be considered Schelz et al. (2006). Some species from

84 Annotated Checklist of Vascular Flora: Arches National Park . . var. var. A. Centaurea D. brandegei A. ludoviciana E. pulcherrimus cation may be ed specimen of ed specimen of Also reported (without voucher) ed specimen of ed specimen of . Utah state noxious weed. Native misidentifi Comments Colorado Plateau Network (in ed.), but park is well outside the known range of this species. Reports may be based on macrosperma specimen from ARCH herbarium collected from just outside the park. # 40) on loan. on a collection (Vermilion Specimen is from outside the known range of this species and identifi mexicana. by NCPN (in ed.) a misidentifi erroneous. voucher in ARCH herbarium), but suitable habitat is lacking in ARCH. maculosa to Mediterranean. a misidentifi misidentifi but ARCH is outside the known a voucher, locality is Kachina range of the species. Type Bridge, Natural Bridges NM, San Juan Co., USFS: Region & Moore 2398 BRY). UT (Welsh 4 Sensitive, UT BLM: USFWS: former C2 candidate Park status Range PerF Sparse FalsRep Report from ARCH herbarium is based on a Life form PerF Wide Ques? Report from ARCH herbarium is based Louisiana wormwood Common name dwarf milkweed PerF Periph Ques?water birch Reported by Schelz et al. (2006) & Northern basin yellow Treecryptanth Wide FalsRep Report from Schelz et al. (2006) based on knapweedDiffuse PerF Intro FalsRep Report from ARCH herbarium is based on Fendler’s sandwortFendler’s PerF Periph Ques? Reported by Schelz et al. 2006 (without a gray sandplant AnnFsilver daisy Periph FalsRepKachina daisy Report from Harrison et al. (1964) based on PerF PerF Wide LocEn FalsRep Ques? Report from ARCH herbarium is based on a Reported by Springer et al. (2006) without (2006). . candicans Flora of North Artemisia ludoviciana ssp Synonyms/ notes Taxonomic Vars. not recognized Vars. in America Eremogone fendleri . . var var ora . latiloba canescens Betula occidentalis B. fontinalis Cryptantha confertifl Centaurea diffusa Acosta diffusa Asclepias involucrata fendleri Arenaria fendleri Artemisia ludoviciana var Dicoria canescens Erigeron argentatus Erigeron kachinensis Betulaceae Boraginaceae Compositae (Asteraceae) FamilyAsclepiadaceae Species name Caryophyllaceae (Alsinaceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix C. Rejected (falsely reported or questionable) taxa of Arches National Park. or questionable) taxa of Arches Appendix C. Rejected (falsely reported

Chapter 6: Appendices 85

var. var. ed var. var. . C. ed

M. canescens C. aurea. acutifolia X. glabriuscula P. integrifolia P. P. pilifera. P. subnudum T. var. var. eld Co., UT (Jones ed specimen of eld Co., UT (Cronquist Type locality is 11 miles NE Type Type locality is Bromide Pass, Type ed specimen of G. fastigiata. P. acutifolia P. ed specimens of Flora of North America . ed specimen of . ed specimen of ed specimen of ed specimen of canescens. linearifolia. var. var. misidentifi on a misidentifi Comments Henry Mountains, Garfi 5695y US) (2007) this species is endemic to southern California. misidentifi desertorum. misidentifi according to var. var. on misidentifi ARCH herbarium based on misidentifi specimens of subnudum a misidentifi based on a misidentifi of Henrieville, Garfi 9164 NY). USFWS: 3C. voucher in ARCH herbarium), but is well outside the known range of this species in Utah. nebrascensis specimens of Park status Range Life form PerF LocEn Ques? Reported by Fowler et al. (2007), but PerF LocEn FalsRep Report from ARCH herbarium is based on PerF Periph Ques? Reported by Schelz et al. 2006 (without a San Juan bahia PerF RegEn FalsRep Report from ARCH herbarium is based on a Bigelow’s asterBigelow’s PerF RegEn FalsRep Report from Harrison et al. (1964) is based Common name Hopi-tea PerF Periph FalsRep Report from ARCH herbarium is based on a mealy rosettesSan Diego rabbit- AnnFtobacco Wide FalsRep Report from ARCH herbarium is based on a Garber’s sedgeGarber’s PerG Wide FalsRep Reports from Harrison et al. (1964) are based Rydberg’s gumweedRydberg’s PerF Periph FalsRep Reports from Harrison et al. (1964) and Henrieville woodyaster water sedge PerG Wide FalsRep Report from Harrison et al. (1964) is Fremont’s Fremont’s pepperwort twinpodNewberry’s PerF RegEn FalsRep Reports from ARCH based on misidentifi . . var . var .

var

var Included C. aurea var var. var. (2006) G. squarrosa Flora of North oblongifolia commixta, Dieteria bigelovii commixta Synonyms/ notes Taxonomic Bahia nudicaulis aphanactis. in M. bigelovii Gnaphalium canescens microcephalum in America Included in et al. (2003). by Welsh confertifolia aquatilis racemosa . var Platyschkuhria integrifolia Thelesperma megapotamicum oblongifolia Machaeranthera commixta Psathyrotes annua Pseudognaphalium microcephalum Grindelia aphanactis Grindelia nuda Carex garberi Xylorhiza confertifolia Machaeranthera Carex aquatilis C. aquatilis Lepidium fremontii Physaria newberryi newberryi P. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyCompositae (Asteraceae) Species name Cyperaceae Compositae (Asteraceae) Cyperaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Arches National Park, cont.

86 Annotated Checklist of Vascular Flora: Arches National Park

var. ed . in BRY in BRY S. utahensis S. pungens . S. acutus N. retrorsum N. retrorsum. Stipa arnowiae ed specimen of is based on a misidentifi . C. praegracilis ed specimen of ed specimen of ed specimen of ed specimen of C. foena voucher in ARCH herbarium), but suitable habitat is lacking in ARCH. a misidentifi misidentifi on a misidentifi voucher in ARCH herbarium), but is well outside the known range of this species in Utah. on a paratype of herbarium. misidentifi misidentifi Comments name specimen of longispicatus specimen from ARCH herbarium collected locality is Brumley outside the park. Type 10970 BRY). Ridge, San Juan Co., UT (Welsh USFWS: Category 3C. Park status Range PerF WidePerG FalsRep RegEn Report from Harrison et al. (1964) is based Ques? Reported by Schelz et al. 2006 (without a Life form rusty sedgeAmerican bulrush PerG PerG Sparse Wide Ques? FalsRep Reported by Schelz et al. 2006 (without a Report in Harrison et al. (1964) is based on Bloomer’s ricegrassBloomer’s PerG Wide FalsRep Report from Harrison et al. (1964) is based soft-stem bulrush PerG Widewhite-margined FalsRepswertia Report in Harrison et al. (1964) is based on a depauperate needlegrass purple-mat AnnF Wide FalsRep herbarium is based on a Report from BRY hairy nama milkvetchIsely’s AnnF PerF Periph LocEn FalsRep Report from ARCH herbarium is based on a FalsRep Report from Schelz et al. (2006) based on Common name silvertop sedge PerG Wide FalsRep Report from Harrison et al. (1964) under the x

and . validus species Hybrids S. ssp S. comata. Stipa depauperatum bloomeri, Achnatherum bloomeri. between hymenoides other except x Schoenoplectus americanus tabernaemontani, Schoenoplectus lacustris Frasera albomarginata Achnatherum coronatum, A. parishii ssp. Synonyms/ notes Taxonomic . var bloomeri Stiporyzopsis x Carex subfusca Scirpus americanus Scirpus olneyi, Scirpus validus Schoenoplectus Swertia albomarginata Stipa coronata depauperata Stipa Nama demissum Nama hispidum Astragalus iselyi Carex siccata C. foenea Cyperaceae Cyperaceae Cyperaceae Gentianaceae Gramineae (Poaceae) Gramineae (Poaceae) Hydrophyllaceae Hydrophyllaceae Leguminosae (Fabaceae) FamilyCyperaceae Species name Appendix C. Rejected (falsely reported or questionable) taxa of Arches National Park, cont.

Chapter 6: Appendices 87 .

var var. var. ed M. . Z. vaginatus. zionis P. lanceolatum P. var. var. A. praelongus A. praelongus . Type locality is . Type ed specimen of cation may be erroneous. avescens fl ed specimen of L. brachycalyx ed specimen of ed specimen of ed specimen of . ora. stenophyllum Comments collection (Heil # 1614) on loan. Specimen is from outside the known range of this species and identifi locality is White Canyon, San Juan Co., Type UT (Harrison 11595 US). USFWS: 3C misidentifi a misidentifi on a misidentifi based on a misidentifi multifl ellisiae misidentifi var. specimen from ARCH herbarium collected from outside the park. ellisiae. specimens from ARCH herbarium collected locality is from outside the park. Type Grand Co., UT (Welsh Courthouse Wash, UT BLM: Sensitive. 22709 BRY). & Moore # 2004, which based on Welsh represents var. 10 mi E of Halls Crossing, San Juan Co., UT BLM: Sensitive, 5205 BRY). UT (Welsh USFWS: former C2 candidate specimen of Park status Range Life form PerF LocEn Ques? Report from ARCH herbarium is based on a PerF Wide FalsRep Report from Harrison et al. (1964) is based Shrub RegEn FalsRep Report from Schelz et al. (2006) based on a PerF LocEn FalsRep Report from UT Conservation Data Center Common name Monument milkvetch longstipe milkvetch PerF RegEn FalsRep Report from ARCH herbarium is based on a lemon scurf-pea PerF Wide FalsRep Report from ARCH herbarium is based on mountain death camas lax stickleaf PerF RegEn FalsRep Report from Harrison et al. (1964) is stinking milkvetch PerF Wide FalsRep Report from ARCH herbarium is based on a narrow-leaf dunebroom stage milkvetch PerF LocEnHole-in-the-Rock FalsRepprairie-clover Report from Schelz et al. (2006) based on seemly sweetpea PerF Sparse FalsRep based on a misidentifi Report from BRY

. var by ora M. pumila Var. not Var. eucosmus M. multifl Synonyms/ notes Taxonomic A. monumentalis monumentalis Psoralea lanceolata. not recognized Vars. in Barneby (1989) Included in or some authors D. epica. recognized in Barneby (1989). var. . . var var . var lifolia avescens lonchopus . praelongus . vehiculus Astragalus praelongus var. Astragalus monumentalis lanceolatum Zigadenus elegans Anticlea elegans Mentzelia pumila lagarosa Astragalus praelongus var Parryella fi Psoralidium lanceolatum Astragalus sabulosus var Dalea fl epica Lathyrus eucosmus Lathyrus brachycalyx Leguminosae (Fabaceae) FamilyLeguminosae (Fabaceae) Species name Loasaceae Leguminosae (Fabaceae) Liliaceae (Melanthiaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Arches National Park, cont.

88 Annotated Checklist of Vascular Flora: Arches National Park . var . I. roseata. Mimulus Physocarpus M. alba . cation may be erroneous. Habenaria dilatata ed specimen of ed specimen of utahensis ed ed specimen of ed specimen of var. var. . on a misidentifi Comments Harrison et al. (1964) is based on a misidentifi voucher in ARCH herbarium), but is well outside the known range of this species in Utah. a misidentifi leucostachys. Co., locality is Springdale, Washington Type UT (Jones 5247 NY). but ARCH is well outside the a voucher, locality is known range of this taxon. Type Emery Co., UT (Holmgren & The Big Flat Top, Reveal 3012 UTC). UT BLM: Sensitive, USFWS: former C2 candidate a misidentifi collection (Heil # 5792) on loan. Specimen is from outside the known range of this species and identifi locality is near Canaan Ranch, Rockville, Type Co., UT (Jones 5262 POM) Washington alternans on a misidentifi voucher in ARCH herbarium), but is well outside the known range of this species in Utah. glabratus Park status Range Life form PerF RegEn Ques? Reported by Schelz et al. 2006 (without a Shrub LocEn Ques? Reported by Schelz et al. (2006) without Shrub Wide FalsRep Report from ARCH herbarium is based on PerF Wide Ques? Report from ARCH herbarium is based on a PerF Wide FalsRepPerF Report from ARCH herbarium is based RegEn Ques? Reported by Schelz et al. 2006 (without a ower at top wild shrubby gilia PerF RegEn FalsRep Report from Harrison et al. (1964) is based Common name red mulberrypale evening- Treeprimrose Intro FalsRep Native to E North America. Report from slender bog orchid PerF Wide FalsRep Report in Harrison et al. 1964 is based on fl buckwheat whitestem gooseberry Rockville penstemon yellow monkeyfl Fremont’s penstemon . . (2002) var Flora of var Vars. not Vars. ora. Platanthera sparsifl recognized in North America Gilia congesta frutescens Synonyms/ notes Taxonomic E. smithii fremontii .

var ora . . var var ora . sparsiﬂ . frutescens trichocalyx Oenothera pallida Habenaria sparsiﬂ var Morus rubra Ipomopsis congesta var Eriogonum corymbosum smithii Ribes inerme congestus Mimulus guttatus Penstemon fremontii fremontii P. Penstemon pachyphyllus Onagraceae Orchidaceae FamilyMoraceae Species name Polemoniaceae Polygonaceae Saxifragaceae (Grossulariaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Appendix C. Rejected (falsely reported or questionable) taxa of Arches National Park, cont.

Chapter 6: Appendices 89 e or Ques?). cation may be erroneous. Comments collection (Fleming # 774) on loan. Specimen is from outside the known range of this species and identiﬁ locality is Mancos, Montezuma Co., CO Type 76 NY). Earle, & Tracy (Baker, collection in ARCH herbarium from outside locality is 20 km NW of the park. Type 22729 & Trotter Moab, Grand Co., UT (Welsh UT BLM: Sensitive, USFWS: former C2 BRY). candidate. Park status ature (Harrison et al. 1964; Schelz et al. 2006) or from herbarium specimens, (Harrison et al. 1964; Schelz 2006) or from ature Range Life form PerF RegEn Ques? Report from ARCH herbarium is based on a PerF LocEn FalsRep Report by Schelz et al. (2006) based on a Common name Natural Bridges penstemon spring- Trotter’s parsley cations (False Reports or FalsRep) because the park is well outside known expected range of species (Questionabl Synonyms/ notes Taxonomic Oreoxis trotteri Cymopterus trotteri Penstemon strictiformis Umbelliferae (Apiaceae) FamilyScrophulariaceae Species name Appendix C. Rejected (falsely reported or questionable) taxa of Arches National Park, cont. The vascular plant species shown in this table have been reported for Arches National Park from published and unpublished liter National Park from for Arches The vascular plant species shown in this table have been reported because of misidentiﬁ now believed to be erroneous but are

90 Annotated Checklist of Vascular Flora: Arches National Park Cultivated x x x xx PinJun Wet Comments Des Salt xx xx xx x Br Blck Flower time Park status white mulberrytwo-needle pinyon white poplarFremont cottonwood Native IntroCarolina poplar Pres Nativelance-leaf cottonwood Prespeach-leaf willow Pres Intro NA black willowGoodding’s Apr–Jun Native Mar–Jun netleaf hackberry Pres Intro Pres Native Native Mar–JunSiberian elm Pres Pres Apr–Jun Pres Native Apr–May Apr–Junnarrowleaved yucca Apr–Jun yuccaHarriman’s Pres x squawbush Apr–May Native Intro Repsquawbush Native Pres Disturbed May–Jul Pressmooth sumac Mar–Aprwestern poison ivy x Apr–May Native mahoniaFremont’s Cultivated x Pres x Native Native Cultivated Native May–Jul Pres Native Pres Pres x Pres x May–Jul May–Jun Jun–Jul Apr–Jul x Sandy soils Cultivated x x x x x Gambel oak Native Pres Apr–Jun black locust Intro Pres May–Jun Oregon-grape chollaWhipple’s Native Native Pres Rep Apr–Jun May–Jul x x x box eldercommon catalpaRocky Mountain juniperRussian-olive Native Intro Pres Native Pres NA Pres Intro Jun–Jul Apr–Jun Pres May–Jul x x x Cultivated x Disturbed .

. . . var var. var var var intercurrens canadensis x x Pinus edulis Populus alba Populus fremontii fremontii Populus Salix amygdaloides Salix gooddingii Celtis reticulata angustissima Yucca harrimaniae Yucca harrimaniae rydbergii Toxicodendron Mahonia fremontii Mahonia repens Morus alba Populus Ulmus pumila Rhus aromatica simplicifolia Rhus aromatica trilobata Rhus glabra gambelii Robinia pseudoacacia Opuntia whipplei Acer negundo Catalpa bignonioides Juniperus scopulorum Elaeagnus angustifolia Quercus gambelii National Park, organized by life form with ecological notes. taxa of Arches rmed, historical, and reported Pinaceae Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Ulmaceae (Celtidaceae) Shrubs Agavaceae Agavaceae Anacardiaceae Berberidaceae Berberidaceae Moraceae Salicaceae Ulmaceae Anacardiaceae Anacardiaceae Anacardiaceae Leguminosae (Fabaceae) Cactaceae FamilyTrees Aceraceae Bignoniaceae Cupressaceae Species nameElaegnaceae Fagaceae Common name Nativity Appendix D. Conﬁ

Chapter 6: Appendices 91 Saline or seleniferous soils Clayey to gravelly saline soils Sandy soils Rocky areas Saline clays Mancos & Morrison formations Clayey to gravelly soils Sandy soils x x x PinJun Wet Comments x x xx xx xx x xx Des Salt xx x xx xx xx x Br Blck Flower time Park status ower snowberry ower Native Pres May–Jul x x Bigelow’s sagebrushBigelow’s Native Pres Aug–Oct Ruin Park winterfat Native siltbushBrandegee’s Pres May–Jul Native Pres Jun–Aug greasewood Native Pres May–Aug budsage Native Pres Apr–Jun black sagebrush Native Rep Sep–Oct sand sagebrush Native Pres Aug–Oct winterfat Native Pres May–Jul Garrett’s saltbushGarrett’s spiny hopsage Native Pres Native May–Sep Pres Apr–Jul x x x Castle Valley saltbushCastle Valley Native Pres May–Sep mat saltbush Native Pres Apr–Sep shadscale Native Pres May–Sep four-wing saltbushfour-wing Native Pres May–Sep long-ﬂ iodine bush Native Pres Jul–Sep x x basin big sagebrush Native Pres Jul–Sep . orus .

var . . var . nova var. var var var lifolia . ruinina . lanata Artemisia bigelovii brandegei Zuckia brandegei Sarcobatus vermiculatus Artemisia spinescens Artemisia nova Artemisia fi Krascheninnikovia lanata var Krascheninnikovia lanata var Atriplex garrettii garrettii Grayia spinosa Atriplex gardneri cuneata Atriplex corrugata Atriplex confertifolia canescens Atriplex canescens Allenrolfea occidentalis Symphoricarpos longifl Artemisia tridentata tridentata rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae (Sarcobataceae) Compositae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae FamilyShrubs, cont. Caprifoliaceae Species name Common name Nativity (Asteraceae) Appendix D. Confi

92 Annotated Checklist of Vascular Flora: Arches National Park Sandy to saline soils Sandstone x x x x x PinJun Wet Comments xx x xx xx Des Salt xx x xx x xx x xx x xx xx x xx xx x xx Br Blck Flower time Park status slenderleaf rabbitbrush Native Pres Jul–Sep lanceleaf rabbitbrush Native Rep Jul–Sep smoothseed rabbitbrush Native Pres Aug–Oct longbract rabbitbrush Native Pres Jul–Sep rush rabbitbrush Native Pres Aug–Oct glabrate rabbitbrush Native Pres Aug–Oct graystem rabbitbrush Native Pres Aug–Oct greenish rabbitbrush Native Pres Aug–Oct thread snakeweedbroom snakeweed Native Pres Native Jul–Oct Pres Jul–Oct bush encelia Native Pres Apr–Jun green rabbitbrush Native Pres Jul–Sep Mohave brickellbush rabbitbrushspear-leaf Native Pres Native Pres Jun–Aug Jul–Sep longleaf brickellbush Native Pres Aug–Sep Rio Grande seepwillow Native Presrough brickellbush May–Nov Native Pres Aug–Oct . . . orus orus orus var var var . var orus gnaphalodes . viscidifl . stenophyllus . lanceolatus . leiospermus . junceus . graveolens . consimilis Chrysothamnus viscidifl var Chrysothamnus viscidifl var howardii Chrysothamnus parryi Chrysothamnus viscidifl var Chrysothamnus nauseosus var Chrysothamnus nauseosus var Chrysothamnus nauseosus var. Chrysothamnus nauseosus var Gutierrezia microcephala Gutierrezia sarothrae resinosa Encelia frutescens Chrysothamnus nauseosus var Brickellia oblongifolia linifolia Chrysothamnus linifolius Brickellia longifolia Brickellia microphylla Baccharis salicina scabra rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi FamilyShrubs, cont. Compositae (Asteraceae) Species name Common name Nativity

Chapter 6: Appendices 93 Sandstone Saline soils x x PinJun Wet Comments xx xx xx xx xx Des Salt xx x xx xx Br Blck Flower time Park status Native Pres May–Jun birchleaf buckthornUtah serviceberrydwarf mountain mahogany Native Native mountain alder-leaf mahogany Pres Pres Native Apr–Jun Pres May–Jun Apr–Jun x x x x Rock crevices Rock crevices x slender wild buckwheat Native Pres Jun–Oct rimrock wild buckwheat Nativesand wild buckwheat Pres Native Aug–Oct Pres Jun–Oct crisp-leaf wild buckwheat Native Pres Jul–Oct green ephedra ephedraCutler’s turbinella live-oakshinnery oakpurple sage Native Native Nativedesert olive Pres Pressingleleaf ash Pres NA Native NA Apr–Jun Native Pres Pres x Apr–Jun Native x Native May–Jun Pres x Pres x x Apr–May Apr–May x x x x x x Sandy soils x x resinbush Native Pres Jul–Sep shortspine horsebrush Native Pres Apr–Jun Rusby’s goldenbushRusby’s copperweed Native Pres Aug–Sep Nativered-osier dogwoodUtah juniper Pres ephedraTorrey’s Jun–Oct Native Pres Native Native May–Jul Pres Pres NA NA x x x x x x x .

var . viridis . viscida . sericea var var var orum . laxiﬂ . orbiculatum . corymbosum Amelanchier utahensis Cercocarpus intricatus Cercocarpus montanus Rhamnus betulifolia Eriogonum leptocladon Eriogonum microthecum var leptocladon Eriogonum corymbosum var Ephedra viridis Quercus turbinella Quercus welshii Poliomintha incana Fraxinus anomala Eriogonum corymbosum var Forestiera pubescens Vanclevea stylosa Vanclevea Tetradymia spinosa Tetradymia Cornus sericea Haplopappus rusbyi Juniperus osteosperma Ephedra torreyana Ephedra viridis Oxytenia acerosa rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Rosaceae Rosaceae Rosaceae Rhamnaceae Polygonaceae Polygonaceae Polygonaceae Ephedraceae Fagaceae Fagaceae Labiatae (Lamiaceae) Oleaceae Polygonaceae Oleaceae Compositae (Asteraceae) Compositae (Asteraceae) Cornaceae FamilyShrubs, cont. Compositae (Asteraceae) Species name Common nameCupressaceae Ephedraceae Ephedraceae Nativity Compositae (Asteraceae) Appendix D. Conﬁ

94 Annotated Checklist of Vascular Flora: Arches National Park Shallow sandy to clay soil x x PinJun Wet Comments Des Salt xx x xx xx xx x xx x Br Blck Flower time Park status ve-stamen tamarisk ve-stamen Intro Pres Apr–Sep x whiplash willowfendlerbushlittleleaf mock-orange Nativeﬁ Native Pres Native Prestomentose amsonia Apr–Jun Pres May–Sep common dogbane Apr–Jun spider milkweed Native Pres Native Apr–May Pres Native Rep Jun–Aug x May–Jul x pallid milkweed Native Pres Apr–Jun broadleaf milkweedbigseed milkweed milkweedRuth’s Nativeshowy milkweedwhorled milkweed Pres Nativeplateau yellow cryptanth Pres Native Jun–Aug Native Native Pres Native Apr–Jul Pres Pres Pres Apr–Jun Apr–Jun x x May–Aug May–Sep x x x x x x x x x x x x Disturbed Disturbed coyote willow Native Pres Apr–Aug x bush oceanspraydwarf ninebarkcliff-roseantelope bitterbrush Native roseWoods’ Native Presyellow willow Native Pres Jun–Aug Rep Native May–Aug Pres Apr–Jul Native Native May–Jun Pres Pres x May–Jul x Apr–Jun x x Rock crevices Rock crevices x blackbrush Native Pres Apr–Jun yellow-eye cryptanth Native Pres Apr–Jun x .

. var . var. . var var var . stenophylla . caudata ava avoculata var ssp Fendlera rupicola Amsonia tomentosa stenophylla Asclepias asperula asperula Philadelphus microphyllus chinensis Tamarix Apocynum cannabinum Asclepias cryptoceras cryptoceras Asclepias macrosperma Asclepias ruthiae Asclepias speciosa Asclepias subverticillata Cryptantha fl Asclepias latifolia Salix lucida Salix exigua Physocarpus alternans Purshia mexicana stansburyana Rosa woodsii Salix eriocephala watsonii Holodiscus dumosus Purshia tridentata Coleogyne ramosissima Cryptantha fl rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Saxifragaceae (Hydrangeaceae) Forbs Perennial Apocynaceae Asclepiadaceae Saxifragaceae (Hydrangeaceae) Tamaricaceae Apocynaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Boraginaceae Boraginaceae Asclepiadaceae Salicaceae Salicaceae Rosaceae Rosaceae Rosaceae Salicaceae Rosaceae Rosaceae Appendix D. Confi FamilyShrubs, cont. Rosaceae Species name Common name Nativity

Chapter 6: Appendices 95 Sandy soils Disturbed x PinJun Wet Comments xx xx xx x xx xx Des Salt xx x xx x xx xx xx xx Br Blck Flower time Park status Native Pres Apr–May shhook ower fi ower ower cryptanth ower Native Pres May–Jun x x cactus rufous-spine pricklypear Nativeslenderspine pricklypear Pres Nativesmall-fl May–Jun Pres May–Jun common pricklypearUtah pricklypear Native Pres Native May–Jun Pres May–Jun claretcuplarge pricklypear Nativeplains pricklypear Pres Apr–Jun Native Pres Native Rep Apr–Jul May–Jun showy stoneseedmatted tiquilia Native Pres Native Pres Apr–Jul May–Aug x salt heliotrope Native Rep May–Oct sand wormwood Native Pres Jul–Sep plateau cryptanthlong-fl cryptanthOsterhout’s slender cryptanth Native cryptanthWetherill’s Native Prescommon hound’s-tongue Pres Intro May–Jun Native Native May–Jun Rep Pres Rep x x May–Jun Apr–Jun May–Jul x x x x x x x x x Mancos Shale x x Disturbed greenmollyburdock Native Rep Intro Aug–Oct Pres Jul–Sep x x Eastwood’s sandwortEastwood’s Native Pres May–Sep ...... var var var var var var . . . var var ora var cinale . melanacanthus . obovatum spina Arenaria fendleri Opuntia polyacantha Sclerocactus whipplei trichophora roseus Opuntia erinacea erinacea Opuntia erinacea utahensis Opuntia phaeacantha Opuntia polyacantha polyacantha major Opuntia polyacantha rufi Echinocereus triglochidiatus var Tiquilia latior Tiquilia Lithospermum incisum scouleriana Cryptantha fulvocanescens Cryptantha longifl Cryptantha tenuis Cryptantha wetherillii Cynoglossum offi Heliotropium curassavicum var Cryptantha osterhoutii Artemisia campestris Arctium minus Bassia americana eastwoodiae rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Caryophyllaceae (Alsinaceae) Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Cactaceae Boraginaceae (Ehretiaceae) Boraginaceae FamilyPerennial Forbs, cont. Boraginaceae Boraginaceae Species nameBoraginaceae Boraginaceae Boraginaceae Boraginaceae Common name Nativity Boraginaceae Compositae (Asteraceae) Compositae (Asteraceae) Chenopodiaceae Appendix D. Confi

96 Annotated Checklist of Vascular Flora: Arches National Park Disturbed Disturbed x x x x x x x xx xx xx xx PinJun Wet Comments xx xx xx x Des Salt xx x xx x Br Blck Flower time Park status c aster c Native Pres Jul–Oct wavy-leaved thistle Native Pres Jun–Sep pretty thistle thistleRydberg’s Native Pres Native Jun–Sep Pres May–Aug New Mexico thistle Native Pres May–Jun hispid golden-aster Native Pres May–Sep devil’s- beggarsticksdevil’s- Spotted knapweed NativeRussian knapweed Pres Introrose-heath Jun–Aug Pres Intro Jul–Sep Pres Jun–Sep Native Pres May–Sep Disturbed Siskiyou aster Native Pres Jul–Sep Louisiana wormwoodPaciﬁ Native Pres Jul–Oct Louisiana wormwood Native Pres Jul–Oct Louisiana wormwood Native Pres Jul–Oct tarragon Native Pres Jul–Oct bull thistleglaucous hawksbeard Native Pres Intro May–Aug Rep Jul–Aug

. . . . var. . ssp var var var . . minor . glauca var var var var undulatum pulchellum Cirsium neomexicanum Cirsium rydbergii Cirsium undulatum neomexicanum Cirsium calcareum Centaurea repens Bidens frondosa Centaurea maculosa Chaetopappa ericoides Chrysopsis villosa Aster hesperius mexicana Aster ascendens ludoviciana Artemisia ludoviciana Artemisia ludoviciana albula Artemisia ludoviciana glauca Crepis runcinata Artemisia dracunculus Cirsium vulgare rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Compositae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity (Asteraceae)

Chapter 6: Appendices 97 Sandstone Disturbed x x xx PinJun Wet Comments xx xx x x xx xx xx x Des Salt xx x xx xx x xx xx x Br Blck Flower time Park status ower Native Pres May–Oct ower Native Pres Apr–Aug eabane Native Pres May–Aug thrift goldenweed Native Pres May–Jun western goldenrodHopi blanketfl Native Pres Jul–Oct Utah daisy Native Pres Apr–Jun slenderleaf daisy Native Pres Jun–Sep Navajo fl erect gumweed Native Pres Aug–Sep basin blanketfl spreading daisybasin daisy Native Pres May–Sep Native Pres May–Jun spreading daisy Native Pres May–Sep hairy daisy Native Pres May–Jul Arizona hymenoxys Native Pres Apr–Jun nakedstemnoddinghead Native Native Rep Pres May–Jun May–Jun sand goldenastercommon hyalineherb Native Native Pres Pres May–Oct May–Oct Disturbed . .

. . . var . var var var var var. var. da var lifolius cinereus . armerioides Erigeron utahensis Euthamia occidentalis Gaillardia pinnatifi utahensis Erigeron utahensis sparsifolius concinnus Haplopappus armerioides var Grindelia fastigiata Gaillardia spathulata Erigeron pumilus Erigeron divergens divergens Erigeron pulcherrimus Erigeron divergens cinereus aphanactis arizonica Erigeron aphanactis nudicaulis Enceliopsis nutans Hymenoxys acaulis Enceliopsis nudicaulis Heterotheca subaxillaris Hymenopappus fi var. rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Compositae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity (Asteraceae) Compositae (Asteraceae) Appendix D. Confi

98 Annotated Checklist of Vascular Flora: Arches National Park x x x x x x PinJun Wet Comments xx xx xx xx Des Salt xx x xx x xx x xxx xx xx Br Blck Flower time Park status Douglas’ groundselbasin groundsel Native Rep Native May–Oct Pres Apr–Aug desert bahia Native Pres May–Jun gumweed aster Native Pres May–Jul hoary asterScotch thistle Nativealcove rock-daisy Pres May–Sep Intro Native Rep Pres Jun–Sep Jul–Sep Disturbed rock-goldenrod Native Pres Jul–Sep showy rush-pink Native Pres May–Jun Canadian goldenrod Native Pres Aug–Oct Rocky Mountain groundsel Native Pres Jun–Aug canyonlands hymenoxys Native Pres Apr–Jun many-headed groundsel Nativebroom groundsel Pres Jun–Oct Native Pres Jun–Oct Arizona rush-pinkEntrada rush-pink Native Rep Native Pres May–Jun Jun blue lettuce Native Pres Jun–Sep . . .

var . var var . .

. . var

var var ora ora ora var. var var . var. var ora . desertorum . canescens . streptanthifolius longilobus Senecio multilobatus Senecio douglasii Machaeranthera grindelioides grindelioides Perityle specuicola Onopordum acanthium Petradoria pumila pumila Platyschkuhria integrifolia var var Machaeranthera canescens Solidago canadensis salebrosa grandifl Senecio spartioides Senecio streptanthifolius var Hymenoxys acaulis Senecio spartioides multicapitatus spartioides ivesiana Lactuca tatarica grandifl Lygodesmia arizonica grandifl Lygodesmia entrada grandifl Lygodesmia pulchella rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 99 Disturbed Disturbed Sandy soils Chinle, Shinarump, & Moenkopi formations Sandy soils x xx x x x PinJun Wet Comments xx xx x x xx Des Salt xx x xx xx x xx x xx xx Br Blck Flower time Park status ower Native Pres Apr–Aug eld bindweed eld Intro Pres Jun–Aug Disturbed California tansy-mustard Native Rep Jun–Sep desert rockcress Native Pres Mar–Jun perennial rockcress Native Pres Mar–Jul common dandelion Intro Pres Apr–Dec slender wirelettuce Native Pres Jun–Sep sand dune wallﬂ ﬁ evolvulus Native Pres May–Jun x hurtleaf woodyasterCisco woodyaster Native Pres Native Apr–Jun Pres Apr–Jun yellow salsifyrough mule’s-ears Intro Native Pres Pres May–Aug May–Jul hoary townsendia Native Pres Apr–Jul scapose greenthread Native Pres May–Aug three-nerve goldenrod Native Pres Aug–Oct brown-plume wire-lettuce Native Pres May–Oct Moab woodyaster Native Pres Apr–May . . var

var . ora . var var. . pallens var var cinale . tenuifolia Descurainia californica perennans Arabis pulchra Taraxacum offi Taraxacum Erysimum asperum purshii Evolvulus nuttallianus Arabis perennans Convolvulus arvensis imberbis Xylorhiza venusta Tragopogon dubius Tragopogon Wyethia scabra Townsendia incana Townsendia subnudum Thelesperma subnudum Stephanomeria tenuifolia var Solidago velutina Stephanomeria paucifl Xylorhiza tortifolia Xylorhiza glabriuscula linearifolia rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Convolvulaceae Cruciferae (Brassicaceae) Convolvulaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyPerennial Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi

100 Annotated Checklist of Vascular Flora: Arches National Park Disturbed Often on seleniferous soils x x x x x PinJun Wet Comments xx xx xx x xx xx Des Salt xx x xx xx x Br Blck Flower time Park status ower Native Pres Apr–Jul crescent milkvetch Native Pres Mar–Jun Fendler’s spurgeFendler’s shorthorn spurgenoseburngolden corydalis Native NativeGreat Basin centaury PresUtah swertia Presmock pennyroyal Native May–Sep Native May–Jun Native Prescommon horehound x Pres Pres Apr–Sep Native Jun–Sep x Native May–Aug Intro Pres Pres x Rep May–Aug Jun–Jul x x May–Oct x x x x x x Sandy soils Disturbed Moab thelypody Native Pres Jun–Sep elegant thelypody Native Pres Apr–Jul Moab pepperplant Native Pres Aug–Oct heart-leaf jewelﬂ Eastwood’s tall pepperwortEastwood’s Nativebroad-leaf pepperwort Pres Intro Jul–Oct Jones’ pepperwort Preswatercress Jun–Oct Nativesharp-leaf twinpod Presprince’s-plume Apr–Sep Native Intro Pres Pres Native May–Jul May–Sep Pres Apr–Sep .

. var . var. var

var var. cinale gracilipes amphioxys Euphorbia brachycera ramosa Tragia Corydalis aurea occidentalis Swertia utahensis Hedeoma drummondii Astragalus amphioxys Chamaesyce fendleri Centaurium exaltatum Marrubium vulgare Thelypodium integrifolium var. Thelypodiopsis elegans Lepidium moabense Lepidium montanum Nasturtium ofﬁ Physaria acutifolia Streptanthus cordatus Lepidium alyssoides eastwoodiae jonesii acutifolia Lepidium latifolium Stanleya pinnata rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Euphorbiaceae Euphorbiaceae Fumariaceae Gentianaceae Labiatae (Lamiaceae) Leguminosae (Fabaceae) Euphorbiaceae Gentianaceae Labiatae (Lamiaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix D. Conﬁ FamilyPerennial Forbs, cont. Cruciferae (Brassicaceae) Species name Common name Nativity Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

Chapter 6: Appendices 101 Sandy soils Sandy soils Sandy soils Sandy soils Sandy soils x x xx x PinJun Wet Comments xx xx xx xx xx xx xx xx x Des Salt xx x xx xx x xx x xx x x Br Blck Flower time Park status Kanab prairie-clover Native Pres May–Jun Ellis’ milkvetchPreuss’ milkvetchcreeping rush-pea Native Native Pres Pres Native Apr–Jul Pres Apr–Jun Apr–Jun woolly milkvetch Native Pres Feb–Jun Moenkopi milkvetch Native Rep May–Jul straggling milkvetch Native Pres Apr–Jun yellow milkvetch Native Pres Apr–Jul northern sweetvetch Native Pres May–Aug Nuttall’s licoriceNuttall’s Zion sweetpea Native Pres May–Jul Native Pres Apr–Jun Moab milkvetchcanoe milkvetchrimrock milkvetch Native Pres Native Native Apr–Jun Pres Pres Apr–Jun May–Jul Sheldon’s milkvetchSheldon’s Native Hist Mar–Jun white prairie-clover Native Pres May–Sep painted milkvetch Native Pres Apr–Jun

. . . . var. . var. var. latus var avus var . var var . ﬂ

var var. var var var. avus avescens avescens Dalea fl fl Astragalus preussii Astragalus praelongus ellisiae Caesalpinia repens Astragalus mollissimus thompsoniae Astragalus moencoppensis Astragalus lentiginosus palans Hedysarum boreale boreale Glycyrrhiza lepidota Lathyrus brachycalyx zionis Astragalus desperatus Astragalus fl desperatus Astragalus coltonii moabensis Astragalus cymboides Astragalus amphioxys vespertinus Dalea oligophylla Astragalus ceramicus ceramicus rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae or Caesalpinaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix D. Confi

102 Annotated Checklist of Vascular Flora: Arches National Park Saline to seleniferous soils Sandy soils x xx PinJun Wet Comments xx xx Des Salt xx xx x xx xx x xx Br Blck Flower time Park status owered breadroot owered Native Pres Apr–Jun ax Native Pres Apr–Aug x blue fl desert stickleafwing-seed stickleafpurple loosestrifescarlet globe-mallow Native Native Pres Intro Native Pres Pres Mar–Aug Pres Apr–Sep Apr–Oct x Jul–Aug x x x x x x x x starry false Solomon’s-seal Nativealcove death camas Pres Apr–Jun Native Pres Jul–Aug sand lily Native Pres Apr–Jun pink funnel-lilyasparagussego-lily Native Pres Apr–May Intro Pres Native Jun–Jul Pres Jun–Jul Disturbed silvery sophoraSan Juan onionNevada oniontextile onion Native Pres Native Pres Native Jul–Aug Native Pres May–Jun Pres Apr–Jul May–Jun x x x x x x x slenderleaf scurfpea Native Hist Apr–Aug silvery lupinealfalfayellow sweet-clover Nativelarge-fl Pres Intro May–Aug Pres Intro May–Oct Pres May–Sep Disturbed Disturbed slenderleaf globemallow Native Pres May–Sep x x x . orum var . lewisii ora ssp cinalis cinalis . stenophyllum Mentzelia multifl Mentzelia pterosperma salicaria Lythrum Sphaeralcea coccinea Sphaeralcea leptophylla Smilacina stellata Zigadenus vaginatus Linum perenne Eremocrinum albomarginatum Asparagus offi Calochortus nuttallii Allium nevadense Allium textile Androstephium brevifl Sophora stenophylla Allium macropetalum argenteus Melilotus offi Pediomelum megalanthum Medicago sativa Psoralidium lanceolatum var Lupinus argenteus rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Loasaceae Loasaceae Lythraceae Malvaceae Malvaceae Liliaceae (Convallariaceae) Liliaceae (Melanthiaceae) Linaceae Liliaceae (Anthericaceae) Liliaceae (Asparagaceae) Liliaceae (Calochortaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae, Themidaceae) Leguminosae (Fabaceae) Liliaceae (Alliaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix D. Confi FamilyPerennial Forbs, cont. Leguminosae (Fabaceae) Species name Common name Nativity

Chapter 6: Appendices 103 Sandy or gravelly soils x xx x PinJun Wet Comments xx xx x Des Salt xx x xx x xx x xx x Br Blck Flower time Park status cushion milkwortwinged wild buckwheat Native Native Pres Pres Jun–Oct May–Jun x x x Sandy soils alcove bog-orchidclustered broomrape broomrapeCooper’s Native Nativecommon plantain Nativecarmine gilia Pres Presscarlet gilia Pres Jul–Aug Jun–Jul IntroSan Rafael gilia May–Jul prickly-phloxWatson’s x Presdesert phlox Native Native x May–Aug Rep Native Native Pres Pres May–Jun x Pres May–Aug May–Aug x x Native May–Jun x x Rep x Apr–Jun x x x x Disturbed Paria evening-primroseBridges evening-primrose Nativepale evening-primrose Native Pres Presgiant helleborine Native Apr–Jun white bog-orchidLindley’s Jul–Sep Pres Native Pres May–Jul Native Pres Jun–Aug Apr–Aug x x long-tube evening-primrose Native Pres Apr–Jun desert trumpet Native Pres Mar–Oct showy four-o’clocknorthern willow-herb Native Native Pres Pres Apr–Jul Jun–Aug x x x narrow-leaf four-o’clock Native Pres May–Aug small-leaf globemallowfragrant sand-verbena Native Native Pres Pres Mar–Oct Apr–Sep x x x x x x pretty wild buckwheat Native Pres Apr–Jul x x

. . .

. var. .

var var var var. var . var var. var ora atum atum Eriogonum alatum Polygala subspinosa Orobanche fasciculata Orobanche ludoviciana Gilia subnuda Ipomopsis aggregata aggregata Leptodactylon watsonii Phlox austromontana austromontana Habenaria zothecina cooperi Plantago major Ipomopsis roseata navajoensis Oenothera pallida Habenaria dilatata leucostachys Oenothera longissima pallida Epipactis gigantea marginata Oenothera caespitosa infl Epilobium ciliatum Oenothera caespitosa Mirabilis multifl linearis Sphaeralcea parvifolia Abronia fragrans Mirabilis linearis Eriogonum infl Eriogonum bicolor rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Polygonaceae Polygonaceae Polygalaceae Orobanchaceae Orobanchaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Orchidaceae Plantaginaceae Polemoniaceae Onagraceae Orchidaceae Onagraceae Orchidaceae Onagraceae Onagraceae Onagraceae Nyctaginaceae FamilyPerennial Forbs, cont. Malvaceae Nyctaginaceae Nyctaginaceae Species name Common name Nativity Polygonaceae Appendix D. Confi

104 Annotated Checklist of Vascular Flora: Arches National Park Sandstone Navajo & Cedar Mesa sandstone x x x PinJun Wet Comments xx xx Des Salt xx x xx xx x xxx xx x Br Blck Flower time Park status ower Native Pres May–Sep x Sandstone ower Native Pres Jun–Aug ax Intro Pres Jul–Aug x Disturbed ax Native Pres Apr–Aug glabrous monkeyfl pale larkspur larkspurNuttall’s marsh buttercuprock spiraea Native NativeColorado bedstraw Pres Pres Native Apr–Jun Apr–Jul Native Pres Native Pres May–Sep Pres May–Aug Aug–SepDalmatian toadfl x x Rock crevices shrubby bedstrawsweet-scented bedstrawbastard toadfl Native Native Pres Pres paintbrushEastwood’s Jul–Aug May–Aug Native Pres May–Jul x cushion wild buckwheatcurly dock Nativecanaigre dockveiny dock Pressausageleaf talinum Apr–Aug Native Intro Native Pres Pres Rep Native x Apr–Jun Jun–Oct Pres May–Sep x x Apr–Jun x x x Disturbed slender wild buckwheat Native Pres Jun–Oct Eastwood’s monkey-fl Eastwood’s white virgin’s-bower Native Pres Jul–Aug x cave primrosealcove columbine Native Native Pres Pres May–Jun Feb–Juldesert paintbrushWyoming paintbrush x Native Native Pres Hist Jul–Aug Apr–Jun x x x x

. . . var. . var . var var var var orum orum orum orum . simpsonii utahensis scaposum Petrophyton caespitosum Galium multifl coloradoense Delphinium nuttallianum Ranunculus cymbalaria Galium multifl Galium trifl Linaria dalmatica Comandra umbellata scabrida pallida Rumex crispus Rumex hymenosepalus Rumex venosus brevifolium Talinum Eriogonum ovalifolium Mimulus eastwoodiae Mimulus glabratus Delphinium andersonii Clematis ligusticifolia Aquilegia micrantha multifl Castilleja linariifolia Castilleja scabrida Primula specuicola Castilleja chromosa Eriogonum microthecum var rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Rosaceae Rubiaceae Ranunculaceae (Helleboraceae) Ranunculaceae Rubiaceae Rubiaceae Scrophulariaceae Santalaceae Polygonaceae Polygonaceae Polygonaceae Portulacaceae Polygonaceae Scrophulariaceae Scrophulariaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae (Helleboraceae) Scrophulariaceae Scrophulariaceae Primulaceae Scrophulariaceae Appendix D. Confi FamilyPerennial Forbs, cont. Polygonaceae Species name Common name Nativity

Chapter 6: Appendices 105 Sandy soils Sandy soils Sandy to heavy clay soils Entrada Sandstone Sandy to heavy clay soils x x PinJun Wet Comments xx xx Des Salt xx x xx xx x xx x xx x xx x Br Blck Flower time Park status Native Pres Mar–May Native Pres Apr–Jun pinyon dwarf-mistletoejuniper mistletoe Native Rep Native Aug–Sep Rep Jul–Sep x x beardlip penstemon Native Rep Jun–Jul sticky spring-parsley Native Pres Apr–May onion spring-parsley Nativebasin white-cup spring- Presparsley Apr–May bluestem penstemonundulate penstemon NativeLoa penstemon NativeUtah penstemon Preswoolly mullein Pres May–Junwater speedwell ground-cherry May–Jul Fendler’s Native x Native Preslong-leaf ground-cherry Native Pres Intro cymopterusFendler’s Intro Pres May–Jun Native Apr–Jun Pres Pres May–Oct Pres x Native x Jun–Aug Jun–Aug Apr–Aug Pres Apr–May Sandy soils x x x x x x Disturbed Colorado Plateau spring- parsley Parry’s lomatiumParry’s prostrate vervainnorthern bog violet Native Pres Native Native Rep Apr–Jun Pres May–Sep Apr–Jul x x Disturbed Canyonlands lomatium Native Pres Apr–Jun . . var . . . var icatum var var var var Phoradendron juniperinum Penstemon barbatus barbatus Cymopterus newberryi Cymopterus bulbosus Cymopterus purpurascens Penstemon eatonii Penstemon utahensis thapsus Verbascum anagallis-aquatica Veronica Physalis hederifolia Cymopterus acaulis fendleri Penstemon cyanocaulis undosus Penstemon ophianthus fendleri Physalis longifolia Cymopterus purpureus purpureus Viola nephrophylla Arceuthobium di Lomatium parryi bracteata Verbena Lomatium latilobum rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Viscaceae FamilyPerennial Forbs, cont. Scrophulariaceae Species name Common name Nativity Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Solanaceae Umbelliferae (Apiaceae) Scrophulariaceae Scrophulariaceae Solanaceae Umbelliferae (Apiaceae) Violaceae Viscaceae Umbelliferae (Apiaceae) Verbenaceae Umbelliferae (Apiaceae) Appendix D. Conﬁ

106 Annotated Checklist of Vascular Flora: Arches National Park Disturbed Saline soils. Disturbed Sandy soils Disturbed PinJun Wet Comments xx xx xx Des Salt xx x xx x xx x xx xx xx Br Blck Flower time Park status y Native Pres May–Aug x x ve-horn smotherweed ve-horn Intro Pres Aug–Oct narrow-leaf goosefootAmerican bugseed Native Rep Native Jul–Sep Pres Aug–Oct x x Sandy soils oakleaf goosefoot Native Rep Aug–Sep desert goosefoot goosefootFremont’s Native Native Pres Pres Aug–Oct Jun–Sep x x x lambsquarterpitseed goosefoot Intro Native Pres Pres Jul–Sep Jun–Sep Disturbed summer-cypress Intro Pres Aug–Oct x x Disturbed yellow beeplantsleepy catchﬂ ﬁ Native Pres Apr–Sep wing-nut cryptanth Native Prescupseed stickseed Apr–Jul Native Pres Apr–Jun recurved cryptanthbindweed heliotrope Native Native Pres Pres Apr–Jun May–Sep x x x redroot pigweedthick-sepaled cryptanth cryptanthKelsey’s Native Intro Pres Pres Apr–Jun Native Jun–Sep Pres Jun–Jul x x Disturbed tumble pigweedprostrate pigweed Intro Native Pres Pres Jun–Sep Jun–Oct Disturbed Disturbed winged pigweedhalogeton poverty-weedNuttall’s Native Native Pres Pres Intro Aug–Oct Apr–Jul Pres x x Jul–Aug x x x x x Disturbed . . .

var var var var. var var. exus fremontii Corispermum americanum Chenopodium leptophyllum salinum Chenopodium fremontii var. Chenopodium dessicatum Chenopodium glaucum album Chenopodium album berlandieri Chenopodium album Bassia scoparia Lappula occidentalis Cleome lutea Silene antirrhina Bassia hyssopifolia cupulata pterocarya Heliotropium convolvulaceum Cryptantha recurvata Cryptantha crassisepala elachantha Cryptantha pterocarya Cryptantha kelseyana Amaranthus albus Amaranthus blitoides Amaranthus retroﬂ Halogeton glomeratus Monolepis nuttalliana Cycloloma atriplicifolia rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Boraginaceae Capparaceae (Cleomaceae) Caryophyllaceae Chenopodiaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Appendix D. Conﬁ FamilyAnnual Forbs Amaranthaceae Amaranthaceae Amaranthaceae Species name Common name Nativity Chenopodiaceae Chenopodiaceae

Chapter 6: Appendices 107 Disturbed Sandy soils Disturbed Sandy soils Disturbed x x x x x PinJun Wet Comments xx xx x xx xx Des Salt xx xx xx xx x xx x x xx x Br Blck Flower time Park status ower Intro Pres Jul–Sep Disturbed ower Native Pres Jun–Sep ower Native Pres Jun–Sep ower Native Pres May–Sep sand sunfl common sunfl prickly lettuce Intro Pres Jul–Sep prairie sunfl pretty daisymarshweedpretty blanketfl Native Pres Native May–Aug Pres Sep Brandegee’s sandplantBrandegee’s Native Pres Aug–Oct Canadian horseweed Native Pres Jul–Sep tansyleaf aster Native Pres May–Sep stevia dusty-maiden Native Pres Apr–Jul rayless alkali aster Native Pres Aug–Sep Russian thistlebroom seepweedwestern seepweed seepweedTorrey’s Intro Nativebur ragweed Native Pres Native Pres Pres Pres Jul–Sep Jun–Sep Jun–Sep Jun–Sep Native Pres Jul–Oct x x Disturbed sowthistle desert-dandelion Native Pres Apr–Jun spiny-leaf sow-thistle Intro Pres Jul–Oct

. .

ssp. ssp var var. Helianthus anomalus lenticularis Lactuca serriola Helianthus petiolaris fallax Erigeron bellidiastrum Flaveria campestris Gaillardia pulchella Helianthus annuus Dicoria brandegei glabrata Machaeranthera tanacetifolia Chaenactis stevioides Conyza canadensis Aster brachyactis Salsola tragus Suaeda calceoliformis Suaeda occidentalis Suaeda torreyana torreyana Ambrosia acanthicarpa Malacothrix sonchoides Sonchus asper rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) FamilyAnnual Forbs, cont. Chenopodiaceae Chenopodiaceae Chenopodiaceae Species nameChenopodiaceae Common name Nativity Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ

108 Annotated Checklist of Vascular Flora: Arches National Park Disturbed Sandy soils Disturbed Disturbed Disturbed Disturbed x x PinJun Wet Comments xx xx xx xx Des Salt xx x xx x xx x xx xx xx x xx xx x Br Blck Flower time Park status ddle-mustard Native Pres Mar–Jun ixweed Intro Pres Mar–Aug eld dodder eld Native Pres Jul–Sep tumble mustardlong-beak fi Intro Pres May–Aug Disturbed African mustard Intro Pres Apr–Jul Disturbed dwarf drabaprairie pepperwortclasping pepperwort Native Native Intro Pres Rep May–Jul Pres Apr–Jul May–Jul wedgeleaf draba Native Pres Feb–Jul spectacle-pod Native Pres Apr–Sep mountain tansy-mustard Nativefl Pres Apr–Jul morning-gloryblue mustard mustardhare’s-ear Intro Intro Intro Pres Pres Rep Jul–Sep Apr–Jun Apr–Aug Disturbed Disturbed white-plume wire-lettuce Nativeannual townsendia Pres May–Sep Nativefi Pres Apr–Jun common sow-thistle Intro Presrough cocklebur Jul–Oct Native Pres Apr–Oct . . var var .

var var. orum Streptanthella longirostris Sisymbrium altissimum Malcolmia africana Lepidium densiﬂ ramosum Draba cuneifolia Draba reptans Lepidium perfoliatum cuneifolia Dithyrea wislizeni Descurainia incana sonnei Descurainia sophia Chorispora tenella Conringia orientalis Ipomoea purpurea Stephanomeria exigua annua Townsendia Xanthium strumarium Cuscuta pentagona Sonchus oleraceus canadense rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Convolvulaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Convolvulaceae (Cuscutaceae) Appendix D. Conﬁ FamilyAnnual Forbs, cont. Compositae (Asteraceae) Species name Common name Nativity

Chapter 6: Appendices 109 Disturbed Sandy soils Mancos Shale x x PinJun Wet Comments xx xx x xx xx Des Salt xx x xx x Br Blck Flower time Park status ax Native Pres Apr–Jun x xsoils Sandy ower sandpuffs ower Native Pres Apr–Sep x owered gaura owered Native Pres May–Aug x ddleleaf Native Rep May–Aug ower-of-an-hour Intro Pres May–Oct Disturbed broom-fl Paiute suncup Native Pres May–Jun white sweet-clover Intro Pres May–Oct shortstem lupinedwarf lupine Native Pres Native May–Jul Pres Apr–Jul gravel milkvetch Native Pres Feb–Jul Howell’s phaceliaHowell’s Ives’ phacelia milkvetchGeyer’s Native Pres Native Native Apr–Jun Pres Pres x Apr–Jul Apr–Jun x x x x x brittle phacelia Native Rep May–Jun Howell’s namaHowell’s corrugate phacelia Native Native Pres Pres May–Jun May–Aug x white-stem blazingstar stickleafThompson’s fl Nativesmall-fl Pres Native camissoniaEastwood’s Pres Apr–Aug Native Apr–Jun x Pres x Apr–Jun x x x x x Mancos Shale leafy fi ridge-seeded spurge spurgeParry’s stork’s-bill Native Pres Native Jun–Aug Pres x Intro May–Sep Pres x x Feb–Oct x x x x Disturbed x Disturbed small-fl woolly plantain spreading eriastrum Native Native Pres Rep Mar–Aug May–Jun x x x x x x .

var . . pusillus var. . var var var ora orum Linum aristatum scapoidea Melilotus alba Lupinus pusillus Lupinus brevicaulis Astragalus sabulonum Phacelia ivesiana Astragalus geyeri Phacelia howelliana Phacelia demissa demissa Phacelia crenulata corrugata Nama retrorsum Mentzelia albicaulis Mentzelia thompsonii Hibiscus trionum micranthus Tripterocalyx Camissonia eastwoodiae Camissonia scapoidea parvifl Chamaesyce glyptosperma Chamaesyce parryi Erodium cicutarium Nama densum Gaura parvifl Plantago patagonica Eriastrum diffusum rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Linaceae Onagraceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Hydrophyllaceae Leguminosae (Fabaceae) Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Loasaceae Loasaceae Malvaceae Nyctaginaceae Onagraceae Onagraceae FamilyAnnual Forbs, cont. Euphorbiaceae Euphorbiaceae Geraniaceae Species nameHydrophyllaceae Common name Nativity Plantaginaceae Polemoniaceae Appendix D. Confi

110 Annotated Checklist of Vascular Flora: Arches National Park Clay soils x PinJun Wet Comments x Des Salt xxx xx x Br Blck Flower time Park status Native Pres Apr–Jul at-sedge Native Pres May–Sep Disturbed smooth wild buckwheat wild buckwheatWetherill’s Native Nativewillow-weedcommon purslane Pres Presbur buttercup Apr–Sep Apr–Octcleavers Intro xannual paintbrush Intro bird’s-beakWright’s Pres Intro x trumpetangel’s Pres xyellow kallstroemia Jun–Sep Native Pres Jul–Nov Nativeorange kallstroemia x x Native Prespuncture vine Pres Mar–Jun Pres Native Native Jun–Aug Native Jul–Sepgolden sedge Pres Pres Apr–Jun bottlebrush sedge PresNebraska sedge x Jun–Oct Apr–Oct x Intro Jul–Sepblackcreeper sedgechufa ﬂ x x Native Pres x Native Pres x x Native Native Pres May–Sep Disturbed Pres Pres May–Jun Apr–Aug x x Disturbed x May–Aug May–Jul Sandy soils Disturbed x x Disturbed x x Grand Valley desert Grand Valley trumpet common spikerushhardstem bulrushalkali bulrush Native Native Pres Pres May–Aug Native Jun–Aug Pres Jun–Sep x x x Gunnison’s giliaGunnison’s spreading gilia wild buckwheatGordon’s Native Native Pres Native Pres May–Oct Pres May–Jun Apr–Jun x x x x x x x Sandy soils shy giliacommon gilia Native Native Pres Pres May–Jun Apr–Jul x x x three-square bulrush Native Pres May–Aug . var

. . ora var. var var atum Eriogonum wetherillii Polygonum lapathifolium Portulaca oleracea Ranunculus testiculatus Galium aparine echinospermon Cordylanthus wrightii Datura wrightii Kallstroemia californica Kallstroemia parvifl terrestris Tribulus Carex aurea Carex hystericina Carex nebrascensis Carex praegracilis Cyperus esculentus Eriogonum salsuginosum Castilleja exilis fusiforme Eleocharis palustris Scirpus acutus Scirpus maritimus Scirpus pungens Ipomopsis polycladon Eriogonum gordonii Eriogonum infl Ipomopsis gunnisonii Gilia inconspicua Gilia leptomeria leptomeria longispicatus rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Polygonaceae Polygonaceae Portulacaceae Ranunculaceae Rubiaceae Scrophulariaceae Solanaceae Zygophyllaceae Zygophyllaceae Zygophyllaceae Graminoids Perennial Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Polygonaceae Scrophulariaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Polemoniaceae Polygonaceae Polygonaceae Polemoniaceae Appendix D. Confi FamilyAnnual Forbs, cont. Polemoniaceae Polemoniaceae Species name Common name Nativity

Chapter 6: Appendices 111 Slickrock x x x x x xx x PinJun Wet Comments xx xx xx xx Des Salt xx x x xx x xx x xx x Br Blck Flower time Park status spike pappusgrass Native Pres Aug–Sep slender wheatgrass Native Pres Jun–Aug Great Basin wildryesquirreltail NativeSalina wildrye Preswestern wheatgrass Jun–Aug Native Native Native Pres Pres Pres May–Aug Jun–Jul Jun–Aug blue grama Native Pres Jul–Sep Canada wildrye Native Pres Jul–Aug purple three-awnyellow bluestemsideoats grama Native Pres Intro Mar–Sep mountain brome Pres NativeJones’ reedgrass Pres Jun–Seporchard grass Native Jul–Sep Presdesert saltgrass Native May–Aug Pres Intro Jul–Aug Native Pres Pres Jun–Aug Jun–Sep Disturbed Disturbed crested wheatgrassredtop Intro Pres Jun–Aug Intro Pres Jun–Sep Enneapogon desvauxii Elymus trachycaulus Elymus cinereus Elymus elymoides Elymus salinus Elymus smithii Bouteloua gracilis Elymus canadensis Bouteloua curtipendula Distichlis spicata Aristida purpurea Bothriochloa ischaemum Bromus carinatus Calamagrostis scopulorum Dactylis glomerata Agropyron cristatum Agrostis stolonifera rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Gramineae (Poaceae) Appendix D. Conﬁ

112 Annotated Checklist of Vascular Flora: Arches National Park Sandy soils x x x x x x x x x xx xx xx PinJun Wet Comments xx xx x Des Salt xx x xx x xx x Br Blck Flower time Park status uffgrass Native Pres Apr–Aug Mexican muhlysandhill muhly muhlyThurber’s Nativebundle panicgrass Pres Nativevine mesquite Aug–Sep Native Pres Nativeswitchgrass Pres Jul–Sep Pres Jul–Sep timothy Native Jul–Sep common reed Presmuttongrass Native Aug–Sep Pres Intro Native Jun–Sep Pres Pres Native Jul–Sep Jun–Sep Pres May–Aug scratchgrass Native Pres Jul–Aug foxtail muhly Native Pres Jul–Sep ﬂ meadow fescuefowl mannagrassgalleta Introfoxtail barley Native Pres Pres Jun–Aug Jul–Aug Native Native Pres Pres Jun–Aug May–Aug hairy tridens Native Pres May–Jul Muhlenbergia mexicana Muhlenbergia pungens Muhlenbergia thurberi Panicum acuminatum Panicum obtusum Panicum virgatum Phleum pratense Phragmites australis Poa fendleriana Muhlenbergia asperifolia Muhlenbergia andina Erioneuron pulchellum Festuca pratensis Glyceria striata Hilaria jamesii Hordeum jubatum Erioneuron pilosum rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Conﬁ FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity

Chapter 6: Appendices 113 Disturbed x x x x x x xx PinJun Wet Comments xx xx xx xx xx x Des Salt xx x x xx x xx x xx x xx x x Br Blck Flower time Park status Mormon needlegrass ricegrassArnow’s Native Hist Native May–Jul Pres Jun–Aug giant dropseed Nativeneedle-and-thread Pres Jun–Sep Native Pres May–Jul prairie cordgrassprairie wedgegrass Native Native Presspike dropseed Pres Jun–Sep sand dropseed Jul–Sep mesa dropseed Native Pres Native Jun–Sep Pres Native Pres Jun–Aug Jun–Sep Indian ricegrassdesert needlegrass Native Native Pres Pres May–Jul Apr–Jun alkali sacaton Native Pres Jun–Aug alkali cordgrass Native Pres Jun–Sep little bluestem Native Pres Jul–Sep Sandberg’s bluegrassSandberg’s weeping alkaligrass Native Pres Intro Apr–Sep Pres Jun–Aug Kentucky bluegrass Intro Pres May–Aug . var comata var. exuosus Stipa arida Stipa arnowiae Stipa comata Sporobolus giganteus Spartina pectinata Sphenopholis obtusata Sporobolus airoides Sporobolus contractus Sporobolus cryptandrus Sporobolus ﬂ Stipa hymenoides Stipa speciosa airoides Spartina gracilis Schizachyrium scoparium Poa secunda Puccinellia distans Poa pratensis rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Appendix D. Conﬁ

114 Annotated Checklist of Vascular Flora: Arches National Park Disturbed Disturbed Disturbed x x x PinJun Wet Comments xx x Des Salt xx x xx xx xx x Br Blck Flower time Park status at-sedge Native Pres Jun–Sep x nger grass nger Native Pres Aug–Sep eld sandbur eld Native Pres Jul–Sep Disturbed longstyle rushNevada rushpoverty rush rushTorrey’s seaside arrowgrass Nativesouthern cattailbroad-leaved cattail Pres Native Native Native Rep Jun–Augredroot fl Native Pres Pres Native Native Jun–Aug Pres Jun–Aug Pres May–Aug Pres Jun–Aug Jun–Aug Jul–Aug x x x x x x x mountain rush Rep Jun–Aug hairy crabgrass Intro Pres Jul–Sep Disturbed fi feather fi cheatgrass Intro Pres May–Jul Japanese chess Intro Pres Jun–Aug jointed goatgrasssixweeks gramaripgut brome Intro Pres Native May–Jun Pres Intro Aug–Sep Pres Apr–Jun Disturbed Tracy’s rushTracy’s Native Pres Jun–Aug Baltic rushjointed rush Native Native Pres Pres May–Jul Jun–Aug x x slim tridens Native Pres May–Sep . . var var Juncus nevadensis Juncus tenuis Juncus torreyi maritima Triglochin domingensis Typha latifolia Typha Cyperus erythrorhizos Juncus longistylus montanus Digitaria sanguinalis Chloris virgata Cenchrus longispinus Bromus tectorum Bromus japonicus Bouteloua barbata Aegilops cylindrica Bromus diandrus Juncus ensifolius brunnescens Juncus articulatus Juncus ensifolius Juncus arcticus Tridens muticus Tridens rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Juncaceae Juncaceae Juncaceae Juncaginaceae Typhaceae Typhaceae Annual Graminoids Cyperaceae Juncaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Juncaceae Juncaceae Juncaceae Juncaceae Appendix D. Confi FamilyPerennial Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity

Chapter 6: Appendices 115 Disturbed Disturbed Disturbed Disturbed x x x x x PinJun Wet Comments x xx xx xx Des Salt xx x x Br Blck Flower time Park status eld horsetail eld Native Pres NA x smooth scouring-rushsouthern maidenhair fern Native Native Pres Pres NA NA x yellow bristlegrassgreen bristlegrass Introwheat Pres Introfi Jul–Sep Prestall scouring-rush Jun–Sep Intro Native Pres Pres Apr–Jul NA Disturbed Disturbed Disturbed rabbitfoot-grasscultivated rye Intro Pres Intro Jun–Aug Rep May–Jul witchgrass Native Pres Jun–Sep six-weeks fescuerabbit barleyfalse buffalograss Native Pres Intro Apr–Jun Native Pres Pres Jun–Sep May–Jul Disturbed teal lovegrasstufted lovegrassannual wheatgrass Native Native Pres Intro Pres Jul–Aug Pres Jul–Aug May–Jul Disturbed barnyard-grass Intro Pres Aug–Sep . var ora ne Adiantum capillus-veneris Equisetum laevigatum Setaria viridis aestivum Triticum Equisetum arvense Equisetum hyemale affi Setaria glauca Polypogon monspeliensis Secale cereale Panicum capillare Munroa squarrosa Festuca octofl Hordeum murinum Eremopyrum triticeum Eragrostis hypnoides Eragrostis pectinacea Echinochloa crus-galli rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Polypodiaceae (Adiantaceae) Equisetaceae Gramineae (Poaceae) Gramineae (Poaceae) Ferns and Allies Equisetaceae Equisetaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) FamilyAnnual Graminoids, cont. Gramineae (Poaceae) Species name Common name Nativity Appendix D. Confi

116 Annotated Checklist of Vascular Flora: Arches National Park Rock crevices x x x PinJun Wet Comments Des Salt Br Blck Flower time Park status western cliffbrakeborder cloak-fern Native Presbrittle bladder fern Native NA Pres Native NA Pres NA Rock crevices Rock crevices bracken fern Native Pres NA Brewer’s cliff-brakeBrewer’s Rep NA

var.

var. occidentalis Pellaea limitanea Cystopteris fragilis Pteridium aquilinum pubescens Pellaea glabella Pellaea breweri rmed, historical, and reported taxa of Arches National Park, organized by life form with ecological notes, cont. Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Aspleniaceae) Polypodiaceae (Dennstaedtiaceae, Hypolepidiaceae) Polypodiaceae (Adiantaceae, Sinopteridaceae) Appendix D. Conﬁ FamilyFerns and Allies, cont. Polypodiaceae (Adiantaceae, Sinopteridaceae) Species name Common name Nativity

Chapter 6: Appendices 117

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