Annotated Checklist of Vascular Flora Dinosaur National Monument

Home , Achnatherum aridum, Agoseris heterophylla, Allium nevadense, Almutaster, Amauriopsis, Amauriopsis dissecta

National Park Service U.S. Department of the Interior

Natural Resource Program Center

Natural Resource Technical Report NPS/NCPN/NRTR—2009/225 ON THE COVER Paria evening-primrose (Oenothera caespitosa var. navajoensis). Photograph by Walter Fertig. Annotated Checklist of Vascular Flora Dinosaur National Monument

Natural Resource Technical Report NPS/NCPN/NRTR—2009/225

Author Walter Fertig Moenave Botanical Consulting 1117 W. Grand Canyon Dr. Kanab, UT 84741

Editing and Design Alice Wondrak Biel Northern Colorado Plateau Network P.O. Box 848 Moab, UT 84532

July 2009

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Please cite this publication as:

Fertig, W. 2009. Annotated checklist of vascular ﬂ ora: Dinosaur National Monument. Natural Resource Technical Report NPS/NCPN/NRTR—2009/225. National Park Service, Fort Col- lins, Colorado.

NPS 122/100142, July 2009 ii Annotated Checklist of Vascular Flora: Dinosaur National Monument Contents

Figures ...... v Tables ...... v Abstract ...... vii Acknowledgements ...... ix

1 Introduction ...... 1 1.1 Background and justiﬁ cation ...... 1 1.2 Objectives ...... 1 1.3 Study area ...... 1 1.3.1 History and setting ...... 1 1.3.2 Geology ...... 2 1.3.3 Vegetation ...... 4 1.3.4 Previous botanical studies ...... 5

2 Methods ...... 11

3 Results ...... 13 3.1 Summary of the ﬂ ora of Dinosaur National Monument ...... 13 3.2 Life form and biogeographic diversity ...... 15 3.3 Plants of special concern ...... 15 3.4 Non-native species and noxious weeds ...... 15 4 Discussion ...... 21

5 Literature Cited ...... 23

Appendices ...... 27 Appendix A. Confi rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family and scientifi c name with taxonomic and biogeographic notes...... 29 Appendix B. Potential (unconfi rmed) taxa of Dinosaur National Monument...... 95 Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument...... 119 Appendix D. Confi rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes...... 137

Contents iii

Figures

Figure 1.3.1. Location of Dinosaur National Monument, Uintah County, Utah, and Moffat County, Colorado...... 3 Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Dinosaur National Monument by decade, 1910–2007...... 9

Tables

Table 1.3.4. Chronology of plant-collecting efforts in Dinosaur National Monument...... 7 Table 3.1-1. Statistical summary of the ﬂ ora of Dinosaur National Monument...... 13 Table 3.1-2. Statistical summary of the ﬂ ora of Dinosaur National Monument, by state...... 14 Table 3.3. Plant species of conservation concern from Dinosaur National Monument...... 16 Table 3.4. Noxious weed species of Dinosaur National Monument...... 20

Contents v

Abstract

In 2005–2006, the National Park Service’s Northern Colorado Plateau Network received funding to develop a revised checklist of the vascular plant fl ora of Dinosaur National Monument. This entailed a thorough review of existing literature, re-examination of specimens from the Dinosaur National Monument herbarium and other state and national museum collections, and additional fi eld work to corroborate unverifi ed species reports or locate new species. Based on this study, 756 vascular plant taxa have been confi rmed or reliably reported as occuring in the monument (705 are known from validated herbarium specimens and 51 from unvouchered literature reports). Another 123 species formerly considered part of the monument fl ora have now been shown to be falsely reported. The Utah portion of Dinosaur National Monument contains 484 species (13.5% of the state fl ora), and the Colorado segment contains 592 species (18.8% of the entire Colorado fl ora). Only one plant species from Dinosaur National Monument is listed as threatened or endangered under the U.S. Endangered Species Act: Ute ladies’-tresses (Spiranthes diluvialis), but at least 79 other species are either former candidates for listing, identifi ed as sensitive by the U.S. Forest Service or Bureau of Land Management, or tracked as species of concern by state natural heritage programs. Introduced plants make up 10.3% of the monument fl ora (78 species), of which 22 species are considered noxious weeds by the states of Utah and Colorado. Further surveys in Dinosaur National Monument are likely to fi nd additional new species (359 are known from similar habitats outside the monument boundary), especially among late- season fl owering, weedy, or wetland taxa.

Contents vii

Acknowledgements

I would like to thank the following individuals for their assistance with this project: Dr. Angela Ev- enden, botanist and former leader of the Northern Colorado Plateau Network (NCPN) program in Moab, Utah, was an active proponent of this project and helped secure funding for ﬁ eld work and herbarium studies in 2006. Tamara Naumann, Dinosaur National Monument (NM) botanist, provided digital data pertaining to her 1990 ﬂ oristic report and herbarium and literature review, granted research permits, facilitated housing, and assisted this project in numerous other ways. Ann Elder, former curator of museum collections for Dinosaur NM, provided access to all mounted and unmounted herbarium specimens from Dinosaur NM. Melissa Powell, Emily Spencer, and Doug Perry recorded specimen-label data from all mounted collections in the Dinosaur NM herbarium. Libby Nance, former database specialist with the NCPN, assisted with development and quality control of the monument’s NPSpecies database. Libby Nance, Janet Coles, and Helen Thomas also provided data from recent vegetation mapping and wetland inventories on the monument. Drs. Stanley Welsh and Duane Atwood provided access to herbarium collections at Brigham Young University. Dr. Michael Piep provided access to collections at the Utah State University (Intermountain) herbarium. Dr. Ron Hart- man and Ernie Nelson provided access and specimen loans from the Rocky Mountain Herbarium at the University of Wyoming. Thanks to the technical reviewers (Dr. Angela Evenden, Dr. Dusty Perkins, and Libby Nance) for their comments and suggestions. To any others whom I may have inadvertently forgotten, please accept my apologies (and thanks).

Research was conducted under permit #DINO-2006-SCI-0009 for study number DINO-00306. Funding was provided by the National Park Service through the University of Wyoming.

Contents ix

vascular plants for each park unit in the network 1 Introduction (Evenden et al. 2002). This eff ort has entailed reviewing existing species checklists, corroborating In 2006, the National Park Service (NPS) estab- the identifi cation of specimens in park museum lished a cooperative agreement with the Univer- and herbarium collections, locating additional sity of Wyoming, through the Rocky Mountains specimen records from other museums, mining Cooperative Ecosystem Studies Unit, to produce relevant literature and datasets, and conducting a series of technical reports summarizing the vas- fi eld surveys to locate new taxa or confi rm un- cular plant fl ora of each of the 16 national park substantiated reports.† All of these data have been units of the Northern Colorado Plateau Network synthesized in NPSpecies, the park service’s stan- (NCPN). These reports are based on existing in- dardized database for documenting park status formation in the park service’s NPSpecies data- (present, probably present, unconfi rmed, falsely base system and augmented by further data min- reported), abundance, residency, nativity, and ing, reviews of state and regional herbaria, and rarity. With the production of a technical series of new fi eld surveys to resolve the status of many reports on the fl ora of each park, the NPSpecies reported and unconfi rmed species. The objective data will now be available to a broader audience. of these reports is to summarize the current state of knowledge of the fl ora of each park for the benefi t of NPS biologists and interpretive staff , 1.2 Objectives outside researchers, and the general public. The The objectives of this project were to: following report addresses the vascular fl ora of Dinosaur National Monument, in northeast Utah 1. Document the current status (present, and northwest Colorado. Other NCPN park units reported, potential, or falsely reported) of all are addressed in separate reports. vascular plant species that have been cited as occurring in Dinosaur National Monument based on a review of herbarium collections 1.1 Background and justifi cation and past literature. One of the primary missions of the NPS is “to conserve unimpaired the natural and cultural 2. Develop an annotated checklist of the fl ora resources and values of the national park sys- of Dinosaur National Monument, with tem for the enjoyment of this and future genera- supplemental information on taxonomic tions” (NPS 2000). Due to their high degree of synonyms, distribution, within-park abun- protection and distribution across a wide variety dance, fl owering period, growth form, nativ- of habitat types, the nation’s national park units ity, habitat, and sources for each record. make a signifi cant contribution to the protection of biological diversity (Noss and Cooperrider 1.3 Study area 1994). Unfortunately, the full contribution of the 1.3.1 History and setting National Park System is not known, as many park units have traditionally lacked comprehensive Human occupation of Dinosaur National Monu- biological inventories or monitoring programs to ment (NM) dates back 9,000–10,000 years. The identify the composition and condition of their earliest inhabitants were hunter-gatherers, who biota. Knowing the status of park fl ora and fauna were ultimately replaced by the Fremont cul- is critical for eff ective park management and for ture around 400 A.D. Fremont peoples practiced systematically identifying and fi lling gaps in the farming of maize, beans, and squash, in addition nation’s protected area network (Margules and to hunting, and left behind an abundant archeo- Pressey 2000). logical record of petroglyphs, masonry, baskets, and fashioned projectile points. Following a pe- To rectify this shortcoming, Congress passed the riod of severe drought across the southwest in National Parks Omnibus Management Act in 1998 the thirteenth century, the Fremont culture gave to provide guidance and funding for an inventory way to nomadic Ute and Shoshone tribes prac- and monitoring program across the entire Na- ticing hunting and gathering. In the late 1800s, tional Park System. Since 2000, the NCPN Inven- Euro-American settlers moved into the Yampa tory and Monitoring Program has been develop- and Green River drainages to homestead and ing and verifying species lists of vertebrates and establish livestock ranches, though the area

†Taxa refer to all named species, subspecies, and varieties.

Chapter 1: Introduction 1 remained thinly populated (Harvey 1991). 1.3.2 Geology

Although John Wesley Powell’s Colorado River Although Dinosaur NM was created primarily to expedition fl oated through Dinosaur NM in 1869, protect its rich dinosaur fossil beds, the monu- little scientifi c exploration of the surrounding up- ment is equally noteworthy for having one of the lands occurred until the early twentieth century. most complete stratigraphic records of any NPS In 1909, Earl Douglass, a paleontologist with the unit. Faulting associated with the rise of the Uinta Carnegie Museum in Pittsburgh, Pennsylvania, Range and downcutting by the Green and Yampa discovered dinosaur fossils in the Morrison For- rivers has exposed 23 major geologic formations mation near Split Mountain, in Utah. Over the within the monument, dating back nearly 1.1 bil- next 15 years, Douglass and colleagues from the lion years (Chronic 1990; Gregson and Chure Carnegie Museum, U.S. National Museum, and 2003). University of Utah excavated nearly 300 partial- The oldest strata in Dinosaur NM are red quartz- to-complete skeletons of 14 dinosaur species, as ites and sandstones of the Precambrian Uinta well as fossils of crocodiles, freshwater turtles, Mountain Group. These outcrops are exposed bivalves, and silicifi ed wood (Gregson and Chure along the north fl ank of Douglas Mountain, the 2003). Concerned over growing vandalism of the Canyon of Lodore on the Green River, and in a fossil beds, Douglass lobbied the U.S. Department narrow band on the north side of Blue Moun- of the Interior and President Woodrow Wilson to tain (Gregson and Chure 2003). Sediments of the aff ord the quarry site federal protection. In Octo- Uinta Mountain Group were deposited in a large ber 1915, Wilson issued a presidential proclama- trough that represents the failed arm of a rift val- tion establishing Dinosaur National Monument ley system (analagous to the Rift Valley of east Af- (Elder 1999). Initially, the monument covered rica today). These sediments were later uplifted just 80 acres in the immediate area of the quarry. and eroded to a smooth, beveled edge, before President Franklin Roosevelt later expanded the ultimately being buried by marine sandstones monument to more than 82,000 acres, to include and shales of the Lodore Formation in the upper the confl uence of the Green and Yampa rivers. Cambrian, about 520 million years ago (Gregson Following the controversy surrounding the pro- and Chure 2003). posed Echo Park dam in the 1950s, the monument was expanded again in 1960, before attain- Rocks dating to the upper Paleozoic comprise the ing its current confi guation and size (210,276 most widely exposed strata within Dinosaur NM, acres/85,096 ha) in 1978 (Evenden et al. 2002). forming most of Split Mountain, the north slope of Blue Mountain, and the foothills at the base Dinosaur NM is located in northeastern Utah of Douglas Mountain. These sedimentary layers, (Uintah County) and northwestern Colorado including the Mississippian Madison Limestone (Moff at County), along the northern edge of the and Doughnut Shale, Pennsylvanian Round Val- Uinta Basin and foothills of the Uinta Range (Fig- ley Limestone and Morgan Formation, and Perm- ure 1.3.1). The monument is approximately 20 ian Park City Formation, are all of marine origin miles east of Vernal, Utah, and 50 miles west of and were deposited when northeastern Utah Craig, Colorado. Visitor centers are located at the and northwestern Colorado were located along quarry (north of Jensen, Utah) and at monument North America’s continental shelf and infl uenced headquarters in Dinosaur, Colorado. The Y-shape by sedimentation from ancestral mountain ranges of the monument accommodates 46 river miles of to the south (Gregson and Chure 2003). Only the the Yampa River (the last major undammed tribu- late Pennsylvanian–early Permian Weber Sand- tary of the Colorado River) and 45 river miles of stone was formed in a terrestrial setting, as part the Green River. Elevations within the monument of a massive sand dune or beach system. Weber range from 4,750 feet (1,448 m) near the quarry Sandstone forms the cliff walls and large mono- to 9,000 feet (2,743 m) on the summit of Zenobia liths of the canyon bottoms (such as Steamboat Peak, at the west end of Douglas Mountain. Av- and Jennie Lind rocks), and is noteworthy in the erage annual precipitation varies from 11 inches Uinta Basin as a reservoir of petrochemicals. (280 mm) at low-elevation sites to 20 inches (508 mm) at high elevations on the Colorado side (Ev- In the Mesozoic, Dinosaur NM was located along enden et al. 2002). Temperatures average 0–30°F the boundary of the mid-continental sea and was in January and 50–100°F in July. aff ected by repeated cycles of marine inundation

2 Annotated Checklist of Vascular Flora: Dinosaur National Monument Dinosaur National Monument Colorado and Utah

Colorado

Utah

318

Green River Yampa River Visitor Center Dinosaur Quarry ˘

Jensen

Visitor 40 Dinosaur Center

0246 Kilometers 0246 Map location Miles

Established October 4, 1915 Legend 1,448–2,743 m elevation (4,750–9,000 ft elevation) Intermittent streams Park boundary (NPS) 85,096 ha (210,276 ac) Perennial streams U.S. Bureau of Land Management Rivers U.S. Forest Service Paved roads Private Unpaved roads State Unimproved/4WD roads U.S. Fish and Wildlife Service

NCPN boundary Other state or local ownership

MAP CREATED BY ANETH WIGHT, NCPN (DERIVED FROM EVENDEN ET AL. 2002)

Figure 1.3.1. Location of Dinosaur National Monument, Uintah County, Utah, and Moffat County, Colorado.

Chapter 1: Introduction 3 and regression. Mesozoic sediments are exposed mostly dominated by desert shrublands of shad- primarily on Yampa Bench, Island Park, and in scale (Atriplex confertifolia), mat saltbush (Atri- the vicinity of the Dinosaur Quarry (Gregson plex corrugata), Wyoming big sagebrush (Artemi- and Chure 2003). The lower Triassic Moenkopi sia tridentata var. wyomingensis or Seriphidium Formation was deposited in a tidal fl at or shallow tridentatum ssp. wyomingense), budsage (Artemi- lagoon environment favorable to formation of sia spinescens or Picrothamnus desertorum), and evaporites (such as gypsum). In the upper Triassic, greenish rabbitbrush (Chrysothamnus nauseous). these beds were buried by Chinle siltstones, sand- Basin big sagebrush (Artemisia tridentata var. stones, and shales deposited by coastal streams or tridentata) becomes locally dominant on low- lakes. Thick, wind-blown sand dunes inundated elevation valley bottom sites with deeper soils or the area in the late Triassic–early Jurassic, forming a higher water table, while black sagebrush (Ar- the Glen Canyon Sandstone (equivalent to the ex- temisia nova) is common in areas with shallow tensive Nugget/Navajo dune system that covered soils over poorly drained bedrock (Goodrich and much of the Colorado Plateau). Marine trans- Neese 1986; Naumann 1990). gressions (Carmel and Stump formations) were interrupted by eolian Entrada Sandstone in the Woodlands of two-needle pinyon (Pinus edulis) mid-to-late Jurassic. Later in the Jurassic, parts of and Utah juniper (Juniperus osteosperma) pre- the monument were buried by alluvial and fl ood dominate over dry-to-mesic sandstone sites be- deposits laid down by streams meandering across tween 5,500 and 8,000 feet (Naumann 1990). The a steamy coastal fl at. These sediments became the understory of pinyon-juniper woodlands is usu- Morrison Formation, and contain the vast major- ally dominated by shrubs, such as basin big sage- ity of the dinosaur fossils that have given Dino- brush, antelope bitterbrush (Purshia tridentata), saur NM its fame (Chronic 1990). and alder-leaf mountain mahogany (Cercocarpus montanus), but may also have a high concentra- Cretaceous and younger strata are relatively poor- tion of perennial bunchgrasses, including Indian ly developed within Dinosaur NM. Marine sand- ricegrass (Stipa hymenoides or Achnatherum hy- stone and shales of the Dakota, Mowry, Frontier, menoides), Sandberg’s bluegrass (Poa fendleri- and Mancos formations are exposed in the vicin- ana), muttongrass (P. fendleriana), or bluebunch ity of the Dinosaur Quarry, Split Mountain, and wheatgrass (Elymus spicatus or Pseudoroegneria Deerlodge Park. Other Cretaceous formations spicata). Rocky sites exposed to frequent blowing are exposed elsewhere in the Uinta Basin, but not wind often support cushion-plant communities within the monument, itself. Regional uplift in of low forbs and bunchgrasses, rather than pin- the late Cretaceous resulted in formation of the yon-juniper stands. Such cushion communities Uinta Range, but was immediately followed by a support many of the monument’s rarer species, period of extensive erosion and sedimentation of especially on Blue Mountain and Zenobia Peak adjacent valleys. This reached its extreme in the (Naumann 1990). Oligocene, when much of the area was a beveled plane, locally preserved in areas of Blue Moun- At elevations above 8,000 feet, or on lower sites tain, the Yampa Plateau, and Ruple Ridge as the with deep soils, pinyon-juniper woodlands give Bishop Conglomerate. A repeated cycle of moun- way to montane shrublands. Mountain big sage- tain building in the late Tertiary resulted in exhu- brush (Artemisia tridentata var. vaseyana or Se- mation of much of this accumulated sediment by riphidium vaseyanum) shrublands, mixed with the modern Yampa and Green rivers (Gregson Idaho fescue (Festuca ovina var. ingrata or F. and Chure 2003). idahoensis); bluebunch wheatgrass; and a variety of perennial forbs typically occur on drier, south exposures. Alder-leaf mountain mahogany, 1.3.3 Vegetation Saskatoon serviceberry (Amelanchier alnifolia), The vegetation of Dinosaur NM and the Uinta western chokecherry (Prunus virginiana or Padus Basin has been described in detail by Goodrich virginiana), and squawbush (Rhus aromatica var. and Neese (1986), Graham (1937), Naumann trilobata) become dominant on shadier or more- (1990), and the NCPN (in ed.). In general, the mesic slopes with deeper soils (Goodrich and monument’s vegetation can be divided into 5–6 Neese 1986; Naumann 1990). main subdivisions based on elevation, dominant growth form, and proximity to water. Conifer and aspen forests occur at the highest elevations of Blue, Douglas, and Split mountains. Upland areas below 6,000 feet, with clay soils, are Quaking aspen (Populus tremuloides) woodlands

4 Annotated Checklist of Vascular Flora: Dinosaur National Monument occur in mesic sites with well-developed organic George Osterhout visited the quarry in 1925, soils and support a rich understory of peren- and collected the type specimen of Uinta Basin nial grasses, tall forbs, and mountain snowberry cryptanth (Cryptantha [Oreocarya] brevifl ora), (Symphoricarpos oreophilus or S. rotundifolius). the fi rst of more than one-half dozen holotypes Douglas-fi r (Pseudotsuga menziesii) forests occur from Dinosaur NM (Welsh 1982). As with sev- commonly on limestone or other non-quartzitic eral other type collections, however, the earliest substrates, especially on north-facing slopes. specimen of this Uinta Basin endemic was taken Mixed stands of Douglas-fi r and ponderosa pine by Douglass and Peterson in 1912. (Pinus ponderosa) occur in well-shaded side can- yons. Edward H. Graham, also of the Carnegie Muse- um, conducted an exhaustive fl oristic and ecolog- Riparian communities occur along the Green and ical survey of the Uinta Basin of Utah and Colo- Yampa rivers, or are associated with springs and rado from 1931 to 1935 (Graham 1937). During seeps. Fremont cottonwood (Populus fremontii), this period, Graham obtained at least 225 collec- box-elder (Acer negundo or Negundo aceroides), tions from Dinosaur NM, representing 70 new and willow species (Salix spp.) predominate along species for the fl ora. Over the next dozen years, alluvial terraces bordering the major rivers, while Bertrand Harrison, Bassett Maguire, Harold D. narrowleaf cottonwood (P. angustifolia) becomes Harrington, and Cedric L. Porter (all herbarium common along smaller streams draining the adja- curators from the leading herbaria of Utah, Col- cent mountains. Five-stamen tamarisk (Tamarix orado, and Wyoming) made several collecting chinensis or T. ramosissima) has become common trips to the area and added 40 more species to the along many reaches. Wet meadows at lower el- monument checklist (Table 1.3.4). evations are characterized by Baltic rush (Juncus arcticus) and three-square bulrush (Scirpus pun- Although Dinosaur NM had been greatly ex- gens or Schoenoplectus pungens), but often grade panded to include much of the Green and Yampa into big sagebrush–Great Basin wildrye (Elymus river drainage in 1938, the Colorado portion of cinereus or Leymus cinereus) stands on terraces the monument remained poorly known, bo- with a lower water table. Montane wet meadows tanically, until the late 1940s. In 1948, Ruth Ann tend to be locally dominated by Nebraska sedge Bradley (née Wolf) and Kathleen Dever, of the (Carex nebrascensis) or smallwing sedge (C. mi- University of Colorado, worked as fi re lookouts croptera). Hanging gardens occur infrequently on Roundtop Mountain and spent much of their in Dinosaur NM, and are associated with shady free time collecting plant specimens for the Di- seeps within sandstone ledges. Typical hanging- nosaur NM and University of Colorado herbaria. garden species include southern maidenhair fern Over a two-year period, Bradley and Dever col- (Adiantum capillus-veneris), alcove columbine lected more than 280 specimens and discovered (Aquilegia micrantha), giant helleborine (Epi- 139 new plant taxa for the monument as part of pactis gigantea), golden sedge (Carex aurea), and Bradley’s master’s-degree work on the fl ora of Jones’ reedgrass (Calamagrostis scopulorum). Moff at County, Colorado (Bradley 1950). From Many hanging-garden taxa are considered spe- 1948 to 1952, H. D. Harrington, William Weber cies of concern, and the community itself is rare (long-time curator of the University of Colorado (Fowler et al. 2007; Naumann 1990). herbarium), Hugo Rodeck, Alan Beetle, and C. L. Porter contributed 36 additional new species.

1.3.4 Previous botanical studies Stanley Welsh, future curator of the Brigham Western explorer and future presidential candi- Young University herbarium, spent two years sur- date John Charles Fremont led the ﬁ rst botanical veying the Utah side of Dinosaur NM, starting in expedition through the Uinta Basin of north- 1955, as part of his master’s thesis on the monu- western Utah in 1844 and 1845, but apparently ment’s vegetation. During this time, Welsh and did not collect specimens within Dinosaur NM, William Weber documented 65 new species for itself (Goodrich and Neese 1986). Paleontologist the monument ﬂ ora. The new species included Earl Douglass and O. A. Peterson, of the Carnegie the holotype of Ownbey’s thistle (Cirsium own- Museum, made the earliest collections from the beyi), taken by Welsh from the vicinity of Split future Dinosaur NM, taking 102 specimens (rep- Mountain in 1955. As part of his thesis, Welsh resenting 85 taxa) from the Dinosaur Quarry near (1957) produced a revised species list for the Utah Split Mountain in 1912 (Table 1.3.4, Figure 1.3.4). portion of Dinosaur NM, listing 360 taxa.

Chapter 1: Introduction 5 During the 1960s, local ranchers William Mac- The most exhaustive botanical survey of Dino- Leod and his parents, Marian and Bruce Mac- saur NM to date was conducted by Steve O’Kane, Leod, collaborated with William Weber to collect Betsey Neely, Tamara Naumann, and collabo- extensively in the Colorado portion of Dino- rators from 1987 to 1989. This team collected saur NM (Weber et al. 1982). Arthur Holmgren more than 1,400 specimens for the Dinosaur NM and colleagues from Utah State University con- herbarium (DINO) and other regional herbaria, ducted a botanical survey of the Green River in and documented 130 new vascular plant taxa, 1961–1962, and documented 40 new species for increasing the known fl ora of the monument by the monument fl ora. Holmgren (1962) published 18.5%. Naumann (1990) described the results of a revised checklist with 553 plant taxa. Additional this study, which included a detailed inventory of new taxa were collected by Jack Brotherson, Stan nearly 40 rare and endemic species. Welsh, Noel Holmgren, and James Reveal during the mid-to-late 1960s, including the types of Di- Since 1990, more than 50 additional species have nosaur wild buckwheat (Eriogonum lonchophyl- been recorded in Dinosaur NM. Lynn Reidel sur- lum var. saurinum) and orchard snakeweed (Guti- veyed the Hog Canyon area in 1991, and added errezia pomariensis). 19 new species to the fl ora, including the narrow endemic, Graham’s columbine (Aquilegia Elizabeth Neese, Joan Friedlander, Steve Wathen, grahamii). Jim Fowler and C. L. May surveyed Scott Peterson, Dieter Wilken, and others con- hanging gardens in the monument in 1993, and ducted fi eld work in Dinosaur NM in the early discovered one new species (Fowler et al. 2007). 1980s, and added nearly two dozen new species Tamara Naumann added three new early spring to the monument fl ora. Neese collected holotypes species in 1995–1996. Derek Tilley and colleagues of two narrow endemics from Blue Mountain in collected 18 new species from 2002 to 2004. Veg- 1982: rock hymenoxys (Hymenoxys lapidicola) etation mappers from the NCPN (in ed.) added and Blue Mountain penstemon (Penstemon scar- eight new species reports in 2002. Dewey and An- iosus var. cyanomontanus). Goodrich and Neese dersen (2006) conducted a weed inventory and (1986) published the Uinta Basin Flora in 1986, mapping study in 2004, and reported three new summarizing much of the fl oristic knowledge of invasive taxa. Rebecca Harms added four new the Dinosaur NM vicinity. wetland species to the fl ora during a survey of seeps and springs in 2005 (Springer et al. 2006).

6 Annotated Checklist of Vascular Flora: Dinosaur National Monument Table 1.3.4. Chronology of plant-collecting efforts in Dinosaur National Monument. # of new Year Collector(s) Comments taxa 1912 E. Douglass and O. A. 85 Earl Douglass and O. A. Peterson collected 102 plant specimens from the Peterson Dinosaur Quarry area (representing 85 species) for the Carnegie Museum. 1925 G. Osterhout 0 George Osterhout collected the type specimen of Uinta Basin cryptanth (Cryptantha [Oreocarya] brevifl ora) from the Dinosaur Quarry area. Douglass and Peterson had previously documented this species from the same site in 1912, though it remained misidentifi ed for several years. 1931– E. H. Graham, B. F. 71 + 4 Edward H. Graham collected at least 225 specimens in Dinosaur NM for the 1935 Harrison, Larsen, B. reported Carnegie Museum from 1931 to 1935, and discovered or reported 70 new Maguire, & G. Piranian species for the monument fl ora (Graham 1937). Bertrand Harrison, Bassett Maguire, and colleagues added 5 new species during fi eld work in 1934– 1935. 1945– H. D. Harrington, C. L. 33 + 1 Harold D. Harrington, of Colorado State University, and Cedric L. Porter, 1947 Porter, D. Ripley, & R. reported of the University of Wyoming, collected more than 40 specimens in the Barneby monument and discovered or reported 34 new species for the fl ora. Dwight Ripley and Rupert Barneby discovered the holotype of grass milkvetch (Astragalus chloodes) from a site southeast of Jensen, Utah (just outside the monument) in 1946. 1948– R. A. Bradley, K. S. 141 Ruth Ann Bradley (née Wolf) and Kathleen Dever, students from the 1949 Dever, H. Rodeck, H. University of Colorado, worked as fi re lookouts and collected more than 280 D. Harrington, A. A. plant specimens and 139 new species from the monument, focusing mostly Beetle, & C.L. Porter on the Colorado side (Bradley 1950). Harrington, Porter, Hugo Rodeck, and Alan Beetle added nearly 40 additional collections and 19 species for the fl ora in 1948. 1950– W. A. Weber 17 William Weber, longtime curator of the University of Colorado herbarium, 1952 collected 85 specimens from the monument over a three-year period and discovered 17 new species for the fl ora. 1953 A. H. Holmgren & S. S. 0 Arthur Holmgren and S. S. Tillett collected the type specimen of Dinosaur Tillett milkvetch (Astragalus saurinus) from the vicinity of the Dinosaur Quarry. This species had fi rst been collected (though misidentifi ed) by Douglass and Peterson in 1912. 1955– S. L. Welsh & W. A. 65 Stanley Welsh conducted a fl oristic and ecological survey of Dinosaur NM for 1956 Weber his master’s thesis (Welsh 1957). In 1955, Welsh documented 45 new species for the monument and collected the holotype of Ownbey’s thistle (Cirsium ownbeyi) from the Split Mountain area. Welsh and W. A. Weber combined to collect 20 new taxa in 1956. Welsh (1957) produced a checklist of 360 taxa from the Utah side of the monument. 1957– H. D. Harrington & W. 3 H. D. Harrington discovered one new species in 1957, and W. Barmore added 1959 Barmore 2 new taxa in 1959, on the Colorado side of the monument. 1960– W. A. MacLeod, 54 William A. MacLeod and his mother, Marian, collected extensively in the 1962 M. MacLeod, A. Colorado section of Dinosaur NM in 1960–1961, adding 12 new species to the Holmgren, N. monument fl ora. Arthur Holmgren, of Utah State University, Noel Holmgren, Holmgren, T. Jensen, J. Tom Jensen, and James Reveal made approximately 120 collections along Reveal, & S. Welsh the Green River in 1961–1962, and added 40 taxa to the fl ora. Holmgren (1962) published a revised monument checklist with 553 species. Stan Welsh discovered one new species in 1962. 1965– J. Brotherson, N. 12 Jack Brotherson discovered 6 new species for the monument in 1965–1966. 1966 Holmgren, J. Reveal, S. Noel Holmgren and James Reveal added 1 new species and collected the L. Welsh, M. MacLeod, holotype of Dinosaur wild buckwheat (Eriogonum lonchophyllum var. W. A. Weber, & P. J. saurinum) along the Island Park Road in 1966. Stanley Welsh and colleagues Salamun collected the type of orchard snakeweed (Gutierrezia pomariensis) from Orchard Draw in 1965. Marian MacLeod, William Weber, and P. J. Salamun contributed 5 new species. 1967– W. A. Weber & M. 5 William Weber found 4 new species and Marian MacLeod added 1 additional 1971 MacLeod species from the Colorado portion of the monument.

Chapter 1: Introduction 7 Table 1.3.4. Chronology of plant-collecting efforts in Dinosaur National Monument, cont.

# of new Year Collector(s) Comments taxa 1978– D. Wilken, E. Neese, 1 + 1 Dieter Wilken and colleagues collected pitseed goosefoot (Chenopodium 1979 S. L. Welsh, & W. A. reported album var. berlandieri) in the monument in 1978. Stanley Welsh and Weber Elizabeth Neese collected the holotype of park rockcress (Arabis vivariensis, sometimes included with A. fernaldiana) from the Jones Hole National Fish Hatchery, just outside Dinosaur NM, in 1979. Weber and others (1982) reported discovering western aster (Aster spathulatus) in the monument in 1979. Other collectors who worked in Dinosaur NM in the late 1970s, but did not document new species for the fl ora, include Larry England, Duane Atwood, Glen Moore, Barry Johnston, Janet Wingate, Leila Shultz, and Sherel Goodrich. 1982 E. Neese, K. Snyder, & 2 Elizabeth Neese and colleagues discovered two new species for Dinosaur NM, J. Trent and collected the type specimens of rock hymenoxys (Hymenoxys lapidicola) and Blue Mountain penstemon (Penstemon scariosus var. cyanomontanus) from Blue Mountain in 1982 (both had previously been collected in the monument by S. L. Welsh in 1955–1956). 1983– J. Friedlander, S. 22 + 2 Joan Friedlander and Steve Wathen collected 120 specimens for the DINO 1984 Wathen, S. Peterson, reported herbarium and documented 22 new taxa for the monument. Scott Peterson & D.Wilken and Dieter Wilken discovered two additional species, as reported by Goodrich and Neese (1986). 1986 B. Clarfi eld 3 Bonnie Clarfi eld collected 21 specimens for the DINO herbarium, including 3 new species for the monument. 1987– S. O’Kane, B. 130 Steve O’Kane, Betsey Neely, Tamara Naumann, and colleagues conducted 1989 Neely, L. Riedel, S. a major fl oristic and rare plant survey throughout Dinosaur NM (Naumann Galatowitsch, D. 1990). Over a three-year period, this group collected more than 1,400 Kuntz, A. Roszczewski, specimens for the DINO herbarium and documented 130 new taxa for the T. Naumann, E. L. monument, increasing the total fl ora by nearly 20%. In 1987, Steve O’Kane Painter, W. Kelley, discovered the type of needle-leaf fl eabane (Erigeron wilkenii) from Pool H. Campbell, & W. Canyon and named the species in 1990. Jennings 1991 L. Reidel 19 Lynn Reidel made more than 160 collections for the DINO herbarium and discovered 19 new taxa in the Hog Canyon area (including the fi rst record of Graham’s columbine, Aquilegia grahamii). 1993 J. Fowler & C. L. May 1 reported Jim Fowler and C. L. May conducted a survey of hanging gardens along the Colorado River drainage and reported one new species for the monument (Fowler et al. 2007). 1995– T. Naumann 3 Tamara Naumann discovered 3 new species for the monument, including 1996 under-collected early spring species Indian-potato (Orogenia linearifolia) and Jupiter buttercup (Ranunculus jovis). 2002– D. Tilley, Dewey and 22 + 11 Derek Tilley and colleagues collected 18 new species for Dinosaur NM 2005 Andersen, & NCPN reported in 2002–2004. Dewey and Andersen (2006) conducted a weed-mapping vegetation mapping survey and reported 3 new invasive species for the fl ora. The NCPN (in ed.) crew vegetation mapping crew reported 8 new species during fi eld surveys in 2002. Rebecca Harms added 4 new species to the fl ora during a survey of seeps and springs in 2005 (Springer et al. 2006).

8 Annotated Checklist of Vascular Flora: Dinosaur National Monument 800

700

600

500

400

300

Number of vascular taxa 200

100

0 1910 1920 1930 1940 1950 1960 1970 1980 1990 2000 2007

Year

Figure 1.3.4. Increase in the number of plant taxa conﬁ rmed as present in Dinosaur National Monument by decade, 1910–2007.

Chapter 1: Introduction 9

Each list was annotated with supplemental infor- 2 Methods mation on life form, geographic range (within the context of Utah and Colorado), population size In July 2005, I reviewed all mounted specimens in the monument (derived from NCPN in ed. and in the Dinosaur National Monument herbarium fi eld observations), fl owering period, distribu- to determine whether each was correctly identi- tion across general vegetation types, nativity, data fi ed to species, subspecies, or variety following source (collector and collection number from the nomenclature of Welsh and others (2003). As a sample specimen or literature citation), year necessary, I made corrections and annotations of earliest collection (from herbarium records), to each specimen and to the voucher records in and relevant synonymy and taxonomic problems the monument’s Automated National Cataloging (based on review of Barkworth et al. 2003, 2007; System (ANCS+) and NPSpecies database. Barneby 1989; Cronquist 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn 2001; Flora of North Based on this herbarium work and a review of America Editorial Committee 1993, 1997, 2000, monument references (Goodrich and Neese 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; 1986; Graham 1937; Holmgren 1962; Naumann Holmgren et al. 2005; and Weber and Wittmann 2002; Weber et al. 1982), I developed four lists: 2001). The annotations to these lists and label data from the DINO herbarium review were then 1. A revised checklist of species that were incorporated into the NPSpecies database for the present in the monument (confi rmed by an monument. authenticated voucher; included in Appen- dices A and D); Work in 2006–2007 focused on confi rmation of 2. A list of species that were reported in the lit- reported and potential species for the monument erature but not corroborated with a voucher fl ora through review of collections from state and (equivalent to “probably present” in NPSpe- regional herbaria not previously corroborated by cies terminology; included in Appendices A Naumann (1990, 2002). Herbarium studies were and D); undertaken at Brigham Young University (BRY), Utah State University (USU), and the University 3. A list of potential species (“unconfi rmed” of Wyoming (RM) in March, June, and October in NPSpecies) known from the vicinity of 2006, and February and March 2007. I also docu- Dinosaur NM, using an unpublished list mented monument specimens from the Virtual generated by Dr. N. Duane Atwood, of Herbarium website of the New York Botanical Brigham Young University, and Albee and Garden (http://sciweb.nybg.org/science2/ hcol/ others (1988); and intf/index.asp). Finally, I found additional new species reports by examining plot data from the 4. A list of species that are now known to be NCPN vegetation mapping project (NCPN in misidentifi ed or highly questionable (Ap- ed.) and reviewing recent literature (Dewey and pendix C). Andersen 2006; Fowler et al. 2007).

Chapter 2: Methods 11

New York Botanical Garden digital plant data- 3 Results base. As a result, the documented fl ora of Dino- saur NM remains at 705 taxa (Tables 3.1-1, 3.1-2, 3.1 Summary of the fl ora of Dinosaur Appendices A, D). National Monument Currently, 51 additional species have been report- Of the 2,235 vascular plant vouchers in the DINO ed for Dinosaur NM, but have yet to be corrobo- herbarium that I examined in 2005–2006, 280 rated with a voucher specimen (Table 3.1-1, Ap- were misidentifi ed (12.5%), 279 were revised to pendix A). Of these, 40 were reported by Fowler a more current name (12.5%), 522 had variety or and others (2007), Goodrich and Neese (1986), subspecies names added (23.4%), and 1,154 were Graham (1937), Holmgren (1962), Naumann found to be correctly determined (51.6%). Based (2002), and Weber and others (1982), and 11 oth- solely on these collections, the confi rmed fl ora of ers came from observations by recent weed and Dinosaur National Monument stood at 705 taxa vegetation mapping crews (Dewey and Andersen (Fertig 2006). No additional new species were 2005; NCPN in ed.). Taken together, the number discovered during a review of herbarium speci- of confi rmed and reported vascular plant taxa in mens at Brigham Young University, Utah State Dinosaur NM stands at 756 taxa (Table 3.1-1, Ap- University, the University of Wyoming, and the pendix A).

Table 3.1-1. Statistical summary of the ﬂ ora of Dinosaur National Monument. Number Number Category conﬁ rmed additionally Total in park reported for park Taxonomic diversity Total taxa 705 51 756 (including varieties and subspecies) Full species 668 49 717 (excluding varieties and subspecies) Families 78 2 80 Life form diversity Tree taxa 13 2 15 Shrub taxa 78 6 84 Perennial forb taxa 365 30 395 Annual forb taxa 124 7 131 Perennial graminoid taxa 101 5 106 Annual graminoid taxa 10 1 11 Fern taxa 14 0 14 Biogeographic diversity Introduced taxa 63 15 78 Native taxa Locally endemic taxa 18 1 19 Regionally endemic taxa 55 1 56 Disjunct taxa 0 0 0 Peripheral taxa 52 7 59 Sparse taxa 29 1 30 Widespread taxa 488 26 514 Total native taxa 642 36 678 The number of taxa and families is based on taxonomic concepts of Welsh and others (2003). See appendices for an explanation of the various categories. Species are listed as sparse or peripheral in the entire monument if they are considered sparse or peripheral in either Colorado or Utah.

Chapter 3: Results 13 Table 3.1-2. Statistical summary of the ﬂ ora of Dinosaur National Monument, by state. Present and Reported for park Total Category historical in park UT CO UT CO NA* UT CO Taxonomic diversity Total taxa 457 567 27 25 2 484 592 (including varieties and subspecies) Full species 445 556 26 24 2 471 580 (excluding varieties and subspecies) Families 71 81** 2 1 1 73 82 Life form diversity Tree taxa 12 13 2 0 0 14 13 Shrub taxa 64 64 5 3 0 69 67 Perennial forb taxa 231 302 13 16 2 244 318 Annual forb taxa 88 77 5 2 0 93 79 Perennial graminoid taxa 51 89 2 3 0 53 92 Annual graminoid taxa 7 8 0 1 0 7 9 Fern taxa 4 14 0 0 0 4 14 Biogeographic diversity Introduced taxa 41 45 12 1 2 53 46 Native taxa Locally endemic taxa 14 9 1 0 0 15 9 Regionally endemic 42 31 1 0 0 43 31 taxa Disjunct taxa 0 1 0 0 0 0 1 Peripheral taxa 4 34 0 6 0 4 40 Sparse taxa 5 18 0 0 0 5 18 Widespread taxa 351 429 13 18 0 364 447 Total native taxa 416 522 15 24 0 431 546 The number of taxa and families is based on taxonomic concepts of Welsh and others (2003) for Utah species and Weber and Wittmann (2001) for Colorado taxa. See appendices for an explanation of the various categories.

*Not Available. Reported for Dinosaur NM by Dewey and Andersen (2006) without the state given (these are indicated by a ? in the state distribution column in Appendix A).

**The number of families recognized in Colorado is higher than that in Utah (and the park as a whole) in part because of the recognition of several segregate families within Caryophyllaceae, Liliaceae, Polypodiaceae, and Ranunculaceae by Weber and Wittmann (2001).

Entering the 2006 ﬁ eld season, I recognized Monument is 359 (Appendix B). 374 taxa as potentially occurring in Dinosaur NM, based on the proximity of the monument Of the original pool of species documented or re- to known populations (Albee et al. 1988; Fertig ported for Dinosaur NM, I found that 119 taxa 2006; Weber et al. 1982). Fifteen of these species were falsely reported based on misidentiﬁ ed were eliminated from the potential list based on voucher specimens in the DINO herbarium and their low probability of occurring in the monu- other collections. I consider an additional four ment. Most of these were species of montane species cited (but not vouchered) by the NCPN coniferous forest or wetland habitats in the Uinta vegetation mapping team (NCPN in ed.) and Mountains. The current number of potential spe- Holmgren (1962) to be questionable based on the cies that I am recognizing for Dinosaur National absence of suitable habitat or the distance of Di-

14 Annotated Checklist of Vascular Flora: Dinosaur National Monument nosaur NM to the known or expected range of (see Table 3.1-1). Species at the margin of their each species (Appendix C). global range (peripheral), or that occur sparsely over their range in Utah or Colorado, contribute The confi rmed and reported fl ora of the Utah 13.1% of the native fl ora. portion of Dinosaur NM consists of 484 taxa. These species represent 13.5% of the 3,594 native and naturalized taxa documented for the state of 3.3 Plants of special concern Utah by Welsh and others (2003), and 48.3% of Ute ladies’-tresses (Spiranthes diluvialis) is the the state’s 151 plant families (Table 3.1-2). With- only plant species listed as threatened or endan- in Uintah County, the monument fl ora captures gered under the U.S. Endangered Species Act that 34.4% of the 1,406 taxa reported by Welsh and is known to occur in Dinosaur NM (Table 3.3). others (2003). At the ecoregional scale, the Utah At least 21 other monument taxa were formerly portion of Dinosaur NM contains 28.6% of the candidates for potential listing (Category 2 or 3C) 1,694 species reported for the Colorado Plateau under the act (Table 3.3) (CNHP 2007; UDWR in Utah (Shultz et al. 2006). 1998). Presently, two species from Dinosaur NM are listed as sensitive by the U.S. Forest Service, Species richness is higher in the Colorado sec- and 13 are considered sensitive by the Colorado tion of Dinosaur NM. In all, 592 plant taxa are or Utah state offi ces of the Bureau of Land Man- confi rmed or reported from the monument, rep- agement (Table 3.3). The Utah Division of Wildlife resenting 18.8% of the 3,145 species in the fl ora Resources Conservation Data Center (UTCDC) of Colorado (Weber and Wittmann 2001), and lists 50 species from Dinosaur NM as species of 51.3% of the 160 recognized families (Table 3.1- concern due to their global or state rarity (UDWR 2). The monument contains 71.6% of the 826 1998). The Colorado Natural Heritage Program plant species known for Moff at County (Kartesz tracks 47 taxa from the monument as species of 2003). The Colorado segment of the monu- concern (CNHP 2007). In all, 80 plant taxa from ment contains 31.2% of the 1,898 species in the the monument are considered rare species by the state’s portion of the Colorado Plateau ecoregion state natural heritage programs. Naumann (1990) (Kartesz 2003). conducted a multi-year survey of rare species from Dinosaur NM, and provided detailed infor- 3.2 Life form and biogeographic mation on the identifi cation, distribution, habitat, diversity and management needs of nearly 40 species. Although ecologically dominant, tree and shrub species make up only 13.1% of the confi rmed and 3.4 Non-native species and noxious reported plant taxa of Dinosaur NM (see Table weeds 3.1-1). Perennial forbs are the most species-rich Introduced plant species (not historically native life form in the monument fl ora, with 395 known to Utah, Colorado, or North America) make up and reported taxa (52.2% of the total fl ora). An- 10.3% of the confi rmed and reported fl ora of nual forbs and grass-like plants comprise approx- Dinosaur NM (see Table 3.1-1). This fi gure is imately 19% of the fl ora. Like trees and shrubs, lower than the statewide average of 13.5% non- perennial graminoids are ecologically important native species in Utah (Fertig 2007; Welsh et al. but relatively species-poor, representing just 14% 2003) and 15.6% for Colorado (Rejmanek and of the total fl ora. The diversity of ferns and fern- Randall 1994). Of the monument’s 78 introduced allies in Dinosaur NM is especially low (1.9%) taxa, 22 are listed as noxious by the states of Utah (see Table 3.1-1). (SUDAF 2008) or Colorado (CDA 2007) (Table 3.4). Plants on the Colorado state noxious weed Of the monument’s 678 native taxa, 514 (75.8%) list fall into three categories with diff ering man- are widespread across Utah, Colorado, and west- agement objectives and requirements (Colorado ern North America. Only 19 species (2.8% of the Noxious Weed Act of 1996 Title 35 Article 5.5). monument’s native fl ora) have extremely limited Weeds on List A are designated for full eradica- global distributions (local endemics found within tion wherever they occur, while containment of an area of less than 1° of latitude × 2° of longi- infestations is the goal for species on List B, and tude; see appendices for complete defi nitions of voluntary control, education, and research is the biogeographic categories). Regional endemics objective for species on List C. Currently, 15 spe- (largely restricted to the Colorado Plateau) com- cies from Dinosaur NM are on List B and 6 are on prise an additional 8.3% of the monument fl ora List C; none is on List A (Table 3.4).

Chapter 3: Results 15 Table 3.3. Plant species of conservation concern from Dinosaur National Monument.

TNC Family Species Common name global Legal status State heritage status rank Apocynaceae Amsonia jonesii Jones’ amsonia G4 CO BLM: Sensitive CNHP: SOC, S1 Boraginaceae Cryptantha ambiguaH obscure cryptanth G4 UTCDC: Periph, S2? Boraginaceae Cryptantha brevifl ora Uinta Basin G4 CNHP: SOC, S1 (Oreocarya brevifl ora) cryptanth Boraginaceae Cryptantha rollinsii Rollins’ cryptanth G3 CO BLM: Sensitive CNHP: SOC, S2 (Oreocarya rollinsii) USFWS: former C2 Boraginaceae Cryptantha stricta erect cryptanth G3 USFWS: 3C CNHP: SOC, S3 (Oreocarya stricta) UTCDC: Watch, S2S3 Cactaceae Opuntia polyacantha slenderspine G5 UTCDC: Periph, S2 var. trichophora pricklypear T3T5 Capparaceae Cleomella palmeriana Goodrich’s G3G4 UT BLM: Sensitive UTCDC: Rare, S1 (Cleomaceae) var. goodrichii R stinkweed T? Caprifoliaceae Symphoricarpos western snowberry G5 UTCDC: Periph, S2? occidentalisH Caryophyllaceae Arenaria nuttallii Nuttall’s sandwort G5 CNHP: SOC, S1 (Alsinaceae) (Minuopsis nuttallii)R Chenopodiaceae Monolepis pusillaH tiny poverty-weed G5 CNHP: SOC, SH UTCDC: Periph., S1 Compositae Aster perelegans elegant aster G5 CNHP: SOC, S1 (Asteraceae) (Eucephalus elegans) Compositae Cirsium ownbeyi Ownbey’s thistle G3 CO BLM: Sensitive CNHP: SOC, S2 (Asteraceae) UT BLM: Sensitive UTCDC: Watch, S1 USFWS: former C2 Compositae Erigeron wilkenii (E. needle-leaf G1 (G3) CNHP: SOC, S1 (Asteraceae) nematophyllus) fl eabane Compositae Enceliopsis nudicaulis nakedstem G5T5 CNHP: SOC, S1 (Asteraceae) Compositae Gailladia aristata common G5 UTCDC: Periph, SR (Asteraceae) blanketfl ower Compositae Gutierrezia pomariensis orchard GQ USFWS: 3C UTCDC: Watch, S2S3 (Asteraceae) snakeweed Compositae Hymenoxys lapidicola rock hymenoxys G1 UT BLM: Sensitive UTCDC: Rare, S1 (Asteraceae) USFWS: 3C Compositae Hymenoxys torreyana Torrey’s hymenoxys G4 UTCDC: Periph, S1 (Asteraceae) (Tetraneuris torreyana) Compositae Platyschkuhria Uintah bahia G5T3? UTCDC: Watch, S3? (Asteraceae) integrifolia var. ourolepis Compositae Sphaeromeria argenteaR silver chicken-sage G3G4 CNHP: SOC, S1 (Asteraceae) Compositae Thelesperma Hopi-tea G5 UTCDC: Periph, S1 (Asteraceae) megapotamicum Compositae Townsendia mensana plateau G3 USFWS: 3C UTCDC: Watch, S3 (Asteraceae) townsendia Compositae Townsendia strigosaH hairy townsendia G4 CO BLM: Sensitive CNHP: SOC, S1 (Asteraceae) Crossosomataceae Glossopetalon spinesens spiny greasebush G5T3 CNHP: SOC, S3 var. meionandrum (Forsellesia meionandra)

16 Annotated Checklist of Vascular Flora: Dinosaur National Monument Table 3.3. Plant species of conservation concern from Dinosaur National Monument, cont. TNC Family Species Common name global Legal status State heritage status rank Cruciferae Arabis demissa var. nodding rockcress G5 T3T4 USFWS: 3C UTCDC: Watch, S2? (Brassicaceae) demissa (Boechera demissa var. demissa) Cruciferae Arabis nuttallii Nuttall’s rockcress G5 UTCDC: Periph, S2? (Brassicaceae) (Boechera nuttallii) Cruciferae Arabis vivariensis park rockcress G3G4 CO BLM: Sensitive CNHP: SOC, S2 (Brassicaceae) (Boechera fernaldiana) UT BLM: Sensitive UTCDC: Watch, S1 USFWS: former C2 Cruciferae Draba oligosperma var. Uinta draba G5TQ USFWS: former C2 UTCDC: Watch, S2 (Brassicaceae) juniperina (D. juniperina) Cruciferae Draba oligosperma var. few-seed whitlow- G5 CNHP: SOC, S2 oligosperma grass Cruciferae Rorippa sinuataH spreading G5 UTCDC: Infreq, S2? (Brassicaceae) yellowcress Cyperaceae Carex curatorum Canyonlands sedge G2 USFWS: 3C UTCDC: Watch, S2 Cyperaceae Scirpus pallidus pale bulrush G5 UTCDC: Infreq, S1S2 Gentianaceae Centaurium exaltatumR Great Basin G5 CNHP: SOC, S1 centaury Gramineae Muhlenbergia thurberi Thurber’s muhly G3G4Q CNHP: SOC, SR (Poaceae) Hydrophyllaceae Nama densum var. leafy fi ddleleaf G5T5 CNHP: SOC, S1 parvifl orumH Hydrophyllaceae Phacelia demissa var. brittle phacelia G5T3? UTCDC: Watch, S3? minor Juncaceae Juncus bryoidesR minute rush G4 CNHP: SOC, S1 UTCDC: Periph, S1 Leguminosae Astragalus argophyllus Martin’s milkvetch G5T4 CNHP: SOC, S1 (Fabaceae) var. martinii Leguminosae Astragalus chloodes grass milkvetch G3 USFWS: 3C CNHP: SOC, SNR (Fabaceae) UTCDC: Watch, S3 Leguminosae Astragalus duchesnensis Duchesne G3 CO BLM: Sensitive CNHP: SOC, S1S2 (Fabaceae) milkvetch Leguminosae Astragalus hamiltonii Hamilton’s G1 USFWS: former C2 CNHP: SOC, S1 (Fabaceae) milkvetch UTCDC: Rare, S1 Leguminosae Astragalus saurinus Dinosaur milkvetch G3 USFWS: 3C CNHP: SOC, SNR (Fabaceae) UTCDC: Watch, S3 Leguminosae Hedysarum boreale var. Rollins’ sweet- G5T1 UTCDC: Tax Prob., S? (Fabaceae) gremialeH vetch Leguminosae Oxytropis besseyi var. Maybell locoweed G5T2 CNHP: SOC, S2 (Fabaceae) obnapiformis Leguminosae Pediomelum large-fl owered G5T5 CNHP: Watch, S3S4 (Fabaceae) megalanthumH breadroot Leguminosae Thermopsis rhombifolia prairie golden-pea G5 UTCDC: Periph, S2S3 (Fabaceae) Leguminosae Trifolium andinum Andean clover G3 CNHP: SOC, S1 (Fabaceae)

Chapter 3: Results 17 Table 3.3. Plant species of conservation concern from Dinosaur National Monument, cont. TNC Family Species Common name global Legal status State heritage status rank Leguminosae Vicia americana var. plains vetch G5T5 UTCDC: Periph, S2 (Fabaceae) minor Liliaceae Allium nevadense Nevada onion G4 CNHP: SOC, S2 (Alliaceae) Liliaceae Fritillaria pudica yellow-bell G5 CNHP: SOC, S1 Liliaceae Zigadenus vaginatus alcove death- G2 USFWS: 3C CNHP: SOC, S2 (Melanthiaceae) (Anticlea vaginatus) camas UTCDC: Rare, S2 Loasaceae Mentzelia multicaulis Uinta Basin G3 CNHP: SOC, S3 var. uintahensis stickleaf (Nuttallia multicaulis) Loasaceae Mentzelia thompsonii Thompson’s G4 UTCDC: Watch, S2 (Acrolasia humilis)H stickleaf

Onagraceae Epilobium palustreR marsh willow-herb G5 UTCDC: Periph, SR Onagraceae Oenothera ﬂ ava var. large yellow G2 CO BLM: Sensitive CNHP: SOC, S2 acutissima eveing-primrose USFWS: 3C UTCDC: Watch, S2 (O. acutissima) Onagraceae Oenothera pallida var. pale evening- G5 T3T5 UTCDC: Periph, S? trichocalyx primrose Orchidaceae Epipactis gigantea giant helleborine G4 USFS R2: Sensitive CNHP: SOC, S2 Orchidaceae Habenaria zothecina alcove bog-orchid G2 UT BLM: Sensitive CNHP: SOC, S1 (Limnorchis zothecina) USFWS: former C2 UTCDC: Rare, S2 Orchidaceae Spiranthes diluvialis Ute ladies’-tresses G2 USFWS: Threatened CNHP: SOC, S2 UTCDC: Rare, S1 Polemoniaceae Gymnosteris parvulaH gymnosteris G4 UTCDC: Periph, S1S2 Polemoniaceae Leptodactylon watsonii Watson’s prickly- G3G5 CNHP: SOC, S2 phlox Polygonaceae Eriogonum brevicaule Duchesne wild G4Q CO BLM: Sensitive CNHP: SOC, S1 var. viridulum (E. buckwheat USFWS: 3C viridulum) Polygonaceae Eriogonum Dinosaur wild G4T3 USFWS: 3C CNHP: SOC, S1 lonchophyllum var. buckwheat UTCDC: Watch, S3 saurinum (E. saurinum) Polygonaceae Eriogonum tumulosum Woodside wild G3Q CO BLM: Sensitive CNHP: SOC, S2 buckwheat USFWS: 3C UTCDC: Watch, S2 Polypodiaceae Adiantum capillus- southern G5 CNHP: SOC, S2 (Adiantaceae) veneris maidenhair fern Polypodiaceae Asplenium forked spleenwort G4G5 CNHP: SOC, S1S2 (Aspleniaceae) septentrionale UTCDC: Infreq, S1 Polypodiaceae Pellaea breweri Brewer’s cliff-brake G5 CNHP: SOC, S2 (Adiantaceae, Sinopteridaceae) Polypodiaceae Pellaea glabella var. western cliff-brake G5T4 UTCDC: Periph, S2 (Adiantaceae, occidentalis Sinopteridaceae) Portulacaceae Lewisia rediviva Oregon bitter-root G5 CNHP: SOC, S2 Primulaceae Androsace ﬁ liformis slender-stem rock- G4 UTCDC: Periph, S1 jasmine

18 Annotated Checklist of Vascular Flora: Dinosaur National Monument Table 3.3. Plant species of conservation concern from Dinosaur National Monument, cont. TNC Family Species Common name global Legal status State heritage status rank Primulaceae Androsace occidentalis western rock- G5 UTCDC: Periph, S1 jasmine Ranunculaceae Aquilegia grahamii Graham’s G1 USFS R4: Sensitive UTCDC: Rare, S1 (Helleboraceae) columbine Ranunculaceae Ranunculus gmeliniiR lesser yellow water G5 UTCDC: Periph, S1 buttercup Rubiaceae Galium multiﬂ orum var. Colorado bedstraw G4 UTCDC: Watch, S3? coloradoense (G. coloradoense) Scrophulariaceae Penstemon angustifolius Vernal penstemon G5T3 USFWS: 3C UTCDC: Watch, S3 var. vernalensisH

Scrophulariaceae Penstemon radicosus mat-root G5 CNHP: SOC, S1 penstemon Scrophulariaceae Penstemon scariosus var. Blue Mountain G4T2 CNHP: SOC, S2 cyanomontanus penstemon UTCDC: Rare, S2 Umbelliferae Cymopterus acaulis plains spring- G5 UTCDC: Periph, S2? (Apiaceae) parsley Umbelliferae Cymopterus Duchesne spring- G3 CO BLM: Sensitive CNHP: SOC, S1 (Apiaceae) duchesnensis parsley USFWS: 3C UTCDC: Watch, S3 Urticaceae Urtica gracilis ssp. stinging nettle G5T5? CNHP: SOC, S1 holosericea This list was derived from Colorado Natural Heritage Program (2007) and Utah Division of Wildlife Resources (1998). Species preceded by an “H” are known only from historical records. Taxa marked with an “R” are reported from the monument. All other species are conﬁ rmed as present with a voucher specimen.

Codes: TNC global rank assesses abundance and conservation priority on a scale of 1–5 (1 = extremely vulnerable; 5 = secure) for full species (G) and varieties or subspecies (T) across their entire range. A “?” indicates uncertainty in the rank, Q = taxonomic questions, U = unknown, and NR = not ranked. A comparable scoring system is used to measure state abundance (S) in Colorado and Utah under the column State Heritage Status. TNC = The Nature Conservancy. Under legal status, USFWS = U.S. Fish and Wildlife Service. C2 = Category 2 candidate (a former category used for taxa that might warrant being proposed for threatened or endangered status following additional research). 3C = Category 3 candidates (species dropped from consideration for listing). USFS = U.S. Forest Service. R2 = USFWS, Region 2. UT BLM = U.S. Bureau of Land Management, Utah ofﬁ ce. State heritage status/UTCDC (Utah Conservation Data Center) includes conservation categories adopted by the state natural heritage program to prioritize endemic and rare plant taxa (UDWR 1998). These categories include: Historical (presumed extinct or only known historically through their range), Rare (plants with rangewide viability concerns), Watch (regional endemics without rangewide viability concerns), Peripheral (rare or uncommon in Utah, but more common rangewide), Infrequent (plants occur infrequently over western U.S.), Taxonomic Problems (validity of species, subspecies, or variety has been questioned), Additional Data (species ap- pear rare, but more information needed before being assigned to a different category). Under State heritage status/CNHP (Colorado Natural Heritage Program), SOC denotes plants on the state “Species of Concern” list and Watch indicates species on the watch list that may ultimately warrant listing as species of concern (CNHP 2007).

Chapter 3: Results 19 Table 3.4. Noxious weed species of Dinosaur National Monument.

State weed Family Species Common name Park status list Boraginaceae Cynoglossum offi cinale common hound’s-tongue Reported in UT CO List B Chenopodiaceae Halogeton glomeratus halogeton Present in UT CO List C Compositae Arctium minus burdock Reported in UT CO List C (Asteraceae) Compositae Carduus nutans var. musk thistle Present in CO CO List B, UT (Asteraceae) macrolepis Noxious Compositae Centaurea diffusa (Acosta diffuse knapweed Reported in UT CO List B, UT (Asteraceae) diffusa) Noxious Compositae Centaurea maculosa spotted kanpweed Reported in UT CO List B, UT (Asteraceae) (Acosta maculosa) Noxious Compositae Centaurea repens Russian knapweed Present in UT CO List B, UT (Asteraceae) (Acroptilon repens) Noxious Compositae Cichorium intybus chicory Present in UT CO List C (Asteraceae) Compositae Cirsium arvense Canada thistle Present in UT CO List B, UT (Asteraceae) (Breea arvense) Noxious Compositae Cirsium vulgare bull thistle Present in CO CO List B (Asteraceae) Present in UT Compositae Onopordum acanthium Scotch thistle Reported in UT CO List B, UT (Asteraceae) Noxious Convolvulaceae Convolvulus arvensis fi eld bindweed Present in UT CO List C, UT Noxious Cruciferae Cardaria chalepensis orbicular whitetop Present in CO UT Noxious (Brassicaceae) Cruciferae Lepidium latifolium broad-leaf pepperwort Present in CO CO List B, UT (Brassicaceae) (Cardaria latifolia) Noxious Elaeagnaceae Elaeagnus angustifolia Russian olive Reported in UT CO List B Euphorbiaceae Euphorbia esula leafy spurge Reported in UT CO List B, UT (Tithymalus esula) Noxious Gramineae (Poaceae) Elymus repens (Elytrigia quackgrass Present in UT CO List B, UT repens) Noxious Poaceae Bromus tectorum cheatgrass Present in CO CO List C (Anisantha tectorum) Present in UT Scrophulariaceae Linaria dalmatica Dalmatian toadfl ax Reported in UT CO List B (L. genistifolia ssp. dalmatica) Scrophulariaceae Linaria vulgaris butter-and-eggs Present in UT CO List B Scrophulariaceae Verbascum thapsus woolly mullein Reported in UT CO List C Tamaricaceae Tamarix chinensis fi ve-stamen tamarisk Present in CO CO List B (T. ramosissima) Present in UT List derived from the Colorado Department of Agriculture (2007) and State of Utah Department of Agriculture and Food (2008).

20 Annotated Checklist of Vascular Flora: Dinosaur National Monument cryptic species that fl ower late in the growing sea- 4 Discussion son, or occur in undersampled wetland or weedy habitats. Confi rmation of reported species should Since the fi rst plant collections of Earl Douglass remain a high priority, as some of these taxa may and O. A. Peterson at the Dinosaur Quarry in 1912, ultimately prove to be erroneous. Nearly 18% of the confi rmed and reported fl ora of Dinosaur the monument fl ora (138 taxa) consists of historic NM has increased from 85 to 756 taxa (see Tables species that have not been relocated since 1970. 1.3.4, 3.1-1, Figure 1.3.4, Appendix A). Based on Determining whether historic taxa are extirpated available habitats and proximity, at least another or still extant should be a high management pri- 359 plant taxa may occur within the monument ority. Additional studies would also be useful to (Appendix B). Should all of these potential spe- better refi ne the abundance and distribution of cies be found, the documented fl ora of the monu- species of high management interest (especially ment would increase by nearly 32%. Many of the rare plants and invasive non-natives). “missing” potential or reported taxa consist of

Chapter 4: Discussion 21

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24 Annotated Checklist of Vascular Flora: Dinosaur National Monument Rejmanek, M., and J. M. Randall. 1994. Invasive Weber, W. A., J. Wingate, B. MacLeod, and M. alien plants in California: 1993 summary MacLeod. 1982. Checklist of vascular plants and comparison with other areas in North of Moff at County, Colorado. University of America. Madrono 41(3):161–177. Colorado Museum, Boulder, Co. 66 pp. Shultz, L. M., R. D. Ramsey, and W. Lindquist. Weber, W. A., and R. C. Wittmann. 2001. Colo- 2006. Revised atlas of Utah plants. College rado fl ora: Western Slope. Third edition. of Natural Resources, Utah State University, Boulder: University Press of Colorado. 488 Logan, Ut. http://earth.gis.usu.edu/plants. pp. Springer, A. E., L. E. Stevens, and R. Harms. Welsh, S. L. 1957. An ecological survey of the 2006. Inventory and classifi cation of selected vegetation of the Dinosaur National Monu- National Park Service springs on the Colo- ment, Utah. Master’s thesis, Brigham Young rado Plateau. Northern Arizona University, University, Provo, Ut. Flagstaff , Az. NPS Cooperative Agreement ——. 1982. Utah plant types - historical per- #CA 1200-99-009; Task # NAU-118. spective 1840 to 1981 - annotated list, State of Utah Department of Agriculture and and bibliography. Great Basin Naturalist Food (SUDAF). 2008. Utah noxious weed 42(2):129–195. list. http://ag.utah.gov/divisions/plant/nox- Welsh, S. L., N. D. Atwood, S. Goodrich, and L. ious/documents/noxUtah.pdf. C. Higgins. 2003. A Utah fl ora. Third edition, Utah Division of Wildlife Resources (UDWR). revised. Provo, Ut: Brigham Young Univer- 1998. Inventory of sensitive species and sity. 912 pp. ecosystems in Utah. Endemic and rare plants of Utah: An overview of their distribution and status. Report prepared for the Utah Reclamation Mitigation and Conservation Commission and U.S. Department of the Interior. 566 pp + app.

Chapter 5: Literature Cited 25

Appendices Synonyms/Taxonomic notes (Appendices A, B, and C) Pertinent synonyms are included for species with alternative scientifi c names in other recent state and regional fl or- The following appendices contain annotated checklists of as (Barkworth et al. 2003, 2007; Barneby 1989; Cronquist the vascular fl ora of Dinosaur National Monument. 1994; Cronquist et al. 1972, 1977, 1984, 1997; Dorn 2001; Appendix A lists all species that have been confi rmed for Flora of North America Editorial Committee 1993, 1997, the fl ora with an authenticated voucher specimen (includ- 2000, 2002a, 2002b, 2003, 2005, 2006a, 2006b, 2006c; Hol- ing extant and historical records) or that have been reli- mgren et al. 2005). Only species not otherwise listed in the ably reported for the monument in the literature (Dewey Utah or Colorado species name columns are included. No and Andersen 2005; Fowler et al. 2007; Goodrich and eff ort has been made to identify additional synonyms from Neese 1986; Graham 1937; Holmgren 1962; Naumann monographs and fl oras preceding the early 1970s. 1990, 2002; NCPN in ed.; Springer et al. 2006; Weber et Common name (Appendices A, B, C, and D) al. 1982). Vouchers were examined from the DINO herbarium and collections from Brigham Young University Common names are derived primarily from Welsh and (BRY), Utah State University (USU), the University of Wy- others (2003) and the Intermountain Flora (Barneby 1989; oming (RM), and the digital museum database of the New Cronquist 1994; Cronquist et al. 1972, 1977, 1984, 1997; York Botanical Garden (NY). Species entries in Appendix Holmgren et al. 2005). A are organized alphabetically by family and genus. Life form (Appendices A, B, C, and D) Appendix B includes species that may potentially oc- Life form is based on the stature and growth habit of typi- cur within Dinosaur NM, based on the proximity of the cal members of a species at reproductive maturity. Trees monument to other populations or the presence of suit- are woody perennials with a single stem or trunk and are able habitat. typically over 3.5 m tall. Shrubs are woody perennials with one to many trunks arising from near the base of the Appendix C lists species that have been previously re- plant and are usually less than 3.5 m tall. Perennial Forbs ported for Dinosaur NM, but are now considered errone- (PerF) are broad-leaved dicots or monocots with non- ous because they are based on misidentifi ed specimens or woody stems (at least above ground level) that live for mul- questionable literature records. tiple years. Annual Forbs (AnnF) are broad-leaved dicots or monocots with non-woody stems that complete their Appendix D contains the same confi rmed, historical, and life cycle (mature, fl ower, and die) in one year. Perennial reported taxa as in Appendix A, but organized by life form Graminoids (PerG) are linear-leaved, grass-like monocots rather than by family in order to better meet the needs of that live for multiple years. Annual Graminoids (AnnG) NPS interpretive specialists. are linear-leaved, grass-like monocots that complete their life cycle (mature, fl ower, and die) in one year. Ferns and The codes and references for each fi eld in the appropriate fern-allies (Fern) are non-fl owering vascular plants that appendices are summarized below. reproduce by spores. Family (Appendices A, B, C, and D) Range UT/CO (Appendices A, B, and C) Family concepts and nomenclature follow A Utah Flora, Range represents the distribution of a species within Utah third edition (Welsh et al. 2003). Alternative family names or Colorado and each state’s contribution to the plant’s and treatments from other regional fl oras (Dorn 2001; overall global range. Usually these values are the same for Flora of North America Editorial Committee 1993, 2002b; both states, but in some cases a species might be wide- Holmgren et al. 2005; Weber and Wittman 2001) are listed spread in one state, but considered sparse, peripheral, or in parentheses. introduced in the other. Disjunct (Disj) taxa have their Species name (Appendices A, B, C, and D) Utah or Colorado distribution separated from the main, contiguous portion of their range by a gap of more than Because Dinosaur NM straddles Utah and Colorado, two 800 km (ca 500 miles). Introduced (Intro) taxa are those sets of species names are provided using the scientifi c that are not native to Utah, Colorado, or North Amer- names and taxonomic concepts of the most recent fl ora ica but have become naturalized (breeding on their own of each state (Weber and Wittmann 2001 and Welsh et al. without human assistance). Local Endemics (LocEn) are 2003). Scientifi c names are listed in alphabetical rather taxa whose entire global range is restricted to an area of than phylogenetic order and according to the Utah names less than 16,500 km2 (ca 6,370 mi2, or 1° of latitude × 2° of Welsh and others (2003). of longitude). Peripheral (Periph) taxa have a widespread

Chapter 6: Appendices 27 global distribution but occur at the margin of their contig- CO (Appendices A, C) uous range in Utah or Colorado and occupy less than 5% This column indicates whether a given species has been of the state’s area (usually only within a few miles of the documented in the Colorado portion of Dinosaur NM state border). Regional Endemics (RegEn) are taxa with a with a voucher (X). Species that are reported for the mon- 2 global range of 16,500–250,000 km (an area about the size ument without the state indicated are marked by a ?. of the state of Wyoming). Sparse taxa occur widely across Utah, Colorado, or North America but their range within Source (Appendix A) a state is small and patchy, with populations restricted to Sources are the evidence for the assigned park status. In specialized or uncommon habitats. Widespread (Wide) the case of Present and Historical taxa, sources are her- taxa have global ranges exceeding 250,000 km2 and occur barium records (with collector, collection number, and over at least 10% of each state. repository) that were corroborated during this study. For Park status (Appendices A, C, and D) reported taxa, the source is a published or unpublished reference citing the species within the monument. Speci- Park status identifi es whether a species is Present (Pres), men repositories include the Dinosaur NM herbarium Historical (Hist), or Reported (Rep) for Dinosaur NM. (DINO), Brigham Young University (BRY), Utah State Species are considered Present if a confi rmed voucher or University (USU), New York Botanical Garden (NY), and documented observation has been made within monu- the University of Wyoming’s Rocky Mountain Herbarium ment boundaries since 1970. Historical species have been (RM). confi rmed with a voucher from the monument but have not been relocated since before 1970. Reported species Year doc. (Appendix A) are listed for the monument in the literature (NCPN in ed.; Year documented indicates the fi rst year in which a spe- Springer et al. 2006), but have not been corroborated with cies was confi rmed or reported to occur in Dinosaur NM. a voucher specimen. Species that may potentially occur This year does not necessarily correspond with the date of in Dinosaur NM are listed in Appendix B. Rejected taxa the voucher specimen cited under Source. “NA” indicates (listed in Appendix C) have either been falsely reported that the fi rst year is not known (used mostly for “report- (FalsRep) in the literature based on misidentifi ed herbar- ed” taxa). ium specimens or are considered Questionable (Ques?) because park reports are from well outside their known Comments (Appendices A, B, C, and D) or expected range. This column includes additional comments on the status Pop. size (Appendix A) of a particular species in the monument, the place of origin for introduced species, legal status, and other data. Population size refers to the abundance and frequency with which a species occurs within Dinosaur NM. Abundance Nativity (Appendix D) data are derived from NCPN (in ed.) and personal obser- Nativity indicates whether a species is native to Utah or vations. Abundant (Abun) taxa have large populations, Colorado, or Introduced (Intro) based on Fertig (2007). have a broad ecological amplitude (often being dominant in one to several vegetation types), or occur extensively Flower time (Appendix D) across the monument. Common (Com) taxa have large local populations but may be restricted to a single vegeta- Flower time refers to the months in which a species is most tion type or do not occur across the entire monument. likely to be in fl ower. NA indicates a species that is a non- Uncommon (Unc) taxa have low to medium-sized popu- fl owering fern or gymnosperm. lations, typically have a narrow ecological amplitude (or at Habitat (Appendix D) least contribute relatively little to total vegetative cover), or have a small range across the monument. Rare taxa usu- The distribution of species across generalized habitats in ally have low population numbers and are restricted to a Dinosaur NM is indicated by an “x” in the columns Pin- single location within the monument. A “?” indicates that Jun through Wet, and is derived from the Dinosaur NM population size is unknown. vegetation classifi cation (NCPN in ed.) and Naumann (1990). DesShr = Desert shrub communities of saltbush UT (Appendices A, C) and sagebrush. PJ = Pinyon-juniper woodlands. MtCon This column indicates whether a given species has been = Montane conifer forests and shrublands of Douglas-fi r, documented in the Utah portion of Dinosaur NM with a aspen, and ponderosa pine. Wet = Wetlands (including voucher (X). Species that are reported for the monument hanging gardens). Additional comments are included for without the state indicated are marked by a ?. species that are restricted to a particular soil/geologic type (derived from Naumann 1990) or are found mostly in disturbed sites.

28 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments is Helper, is Helper, Carbon Co., UT s.n. (Trelease MO) Sensitive c name with 2002 1961 2002 1946 1945 1912 1991 Year Year doc. 1948 1933 1950 locality Type 1912 1935 1933 CO BLM: 1946 (DINO) et al. 413 (DINO) (DINO) 1955 (CSU) & Jensen 13961 (DINO) 2759 (DINO) (DINO) Source (DINO) 2823 (DINO) (DINO) Dever 5046 (DINO) (BRY) (DINO) 3134 (DINO) Pop. size UT CO Pres X Unc 2874 Tilley Hist X ? Holmgren Pres X Unc 2897 Tilley Pres X X Unc Harrington Pres X X Rare Holmgren Pres X X Unc O’Kane Pres X Unc Reidel 111 Pres X X Unc Neely 4000 Park stat. Pres X X Com O’Kane Hist X ? 5441 Weber Pres X X Com & Wolf Hist X Unc 367 Welsh Pres X Rare 5333 Weber Pres X X Unc O’Kane Wide Wide Wide Wide Wide Wide Wide Wide Range UT/CO Wide Wide Wide Wide RegEn Wide PerF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Shrub Wide/ Life form Shrub Wide/ Shrub Wide/ PerF RegEn/ PerF Wide/ water- plantain northern arrowhead prostrate pigweed common dogbane spider milkweed pallid milkweed swamp milkweed showy Rocky Mountain maple Common name box elder Tree Wide/ Harriman’s Harriman’s yucca squawbush Shrub Wide/ western poison ivy Jones’ amsonia milkweed

var.

var. & americanum var. A. graecizans Includes vars. cannabinum, glaberrimum, angustifolium Includes vars. neomexicanum, douglasii, tripartitum Synonyms/ notes Taxonomic A. negundo interius R. trilobata trilobata Rhus radicans, T. Rhus radicans, T. radicans

ssp. asperula interius Alisma trivialeplantago-aquatica A. Amaranthus blitoides Apocynum cannabinum Asclepias asperula ssp. Asclepias cryptoceras Acer glabrum ssp. Yucca harrimaniae Yucca trilobata Rhus aromatica Toxicodendron Toxicodendron rydbergii

var.

Species name var. var. asperula harrimaniae Alisma plantago- aquatica Sagittaria cuneata Sagittaria cuneata Amaranthus blitoides Apocynum cannabinum Asclepias asperula var. cryptoceras Asclepias cryptoceras Asclepias incarnata Asclepias incarnata Asclepias speciosa Asclepias speciosa glabrum UtahAcer glabrum Colorado Acer negundo Negundo aceroides Yucca harrimaniae Yucca var. Rhus aromatica trilobata Toxicodendron Toxicodendron rydbergii Amsonia jonesii Amsonia jonesii vascular plant taxa of Dinosaur National Monument, organized by family and scientiﬁ rmed, historical, and reported Alismataceae Alismataceae Amaranthaceae Anacardiaceae Apocynaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Asclepiadaceae Aceraceae Aceraceae Family Agavaceae Anacardiaceae Apocynaceae Appendix A. Conﬁ taxonomic and biogeographic notes.

Chapter 6: Appendices 29 Comments is 6.5 miles N of Jensen, Dinosaur NM (Osterhout 6414 RM) Year Year doc. 1933 1989 1933 1946 1912 locality Type 1991 1956 1933 1933 1935 1987 1912 1952 1950 Source Dever 5137 (DINO) 371 (DINO) 2818 (DINO) 1987 (CSU) 2963 (DINO) 15119 (BRY) (BRY) (DINO) (DINO) (DINO) Roszczewski 3162 (DINO) (DINO) (COLO) (DINO) c name with taxonomic and Pop. size UT CO Park stat. Pres X XPres Unc & Wolf X Unc Naumann Pres X X Unc O’Kane Hist X ? Harrington Pres X X Rare O’Kane Pres X X ? Atwood Hist X ? 493 Welsh Pres X X Com Neely 4151 Pres X X Com 5342 Weber Pres X Unc Reidel 36 Pres X Unc O’Kane & Pres X X Unc 5423 Weber Hist X ? 7405 Weber Hist X ? 5445 Weber Range UT/CO Wide Wide Wide Wide RegEn Wide Wide Wide Wide Wide Wide Sparse Periph Periph Life form AnnF Sparse/ PerF RegEn/ AnnF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ Common name Oregon-grape Shrub Wide/ thinleaf alder Shrub Wide/ water birch Tree Wide/ obscure cryptanth Uinta Basin cryptanth thick-sepaled cryptanth sand dune cryptanth plateau yellow cryptanth yellow-eye cryptanth slender cryptanth dwarf cryptanth Kelsey’s Kelsey’s cryptanth wing-nut cryptanth recurved cryptanth Synonyms/ notes Taxonomic A. tenuifolia

ora

ssp. var. ava nana avoculata tenuifolia Alnus incana Cryptantha ambigua Oreocarya breviﬂ elachantha Cryptantha crassisepala Oreocarya ﬂ ssp. Cryptantha kelseyana Cryptantha pterocarya Cryptantha recurvata

Species name var. avafl Oreocarya var. var. ora avoculata Cryptantha crassisepala UtahMahonia repens Mahonia repens Berberis repens Colorado Cryptantha fl Alnus incana occidentalis Betula occidentalis Betula fontinalis Cryptantha ambigua Cryptantha brevifl elachantha Cryptantha fendleri Cryptantha fendleri C. pattersonii Cryptantha fl Cryptantha gracilis Cryptantha gracilis Cryptantha humilis Oreocarya humilis Cryptantha kelseyana Cryptantha pterocarya pterocarya Cryptantha recurvata rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Boraginaceae Berberidaceae Family Boraginaceae Betulaceae Betulaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae biogeographic notes, cont. Appendix A. Confi

30 Annotated Checklist of Vascular Flora: Dinosaur National Monument noxious weed (List B). Native to Eurasia Comments 22 miles S of Uintah Ouray, Co., UT (Rollins 1715 GH). CO BLM: Sensitive, USFWS: former C2 candidate 1987 1960 USFWS: 3C NA CO state 1912 Year Year doc. 1950 locality is Type 1933 1912 1956 1986 1948 1955 2002 Roszczewski 3209 (DINO) Reidel 2780 (DINO) (2002) (DINO) Source DINO V-4 DINO V-4 (DINO) (DINO) (DINO) 2940 (DINO) (DINO) (DINO) 2735 (DINO) (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc O’Kane & Pres X Unc O’Kane & Rep X ? Naumann Pres X X Unc 5375 Weber Park stat. Pres X Rare Simonsen Pres X X Unc Painter 454 Pres X X Unc Neely 3968 Pres X X Unc O’Kane Pres X Unc Neely 4162 Pres X Unc Neely 4005 Pres X X Unc O’Kane Pres X Unc s.n. Tilley Wide RegEn Intro Wide Range UT/CO RegEn Wide Wide Wide RegEn RegEn Periph Wide PerF Wide/ PerF RegEn/ PerF Intro/ AnnF Wide/ Life form PerF RegEn/ AnnF Wide/ PerF Wide/ PerF Wide/ PerF RegEn/ PerF RegEn/ PerF Wide/ AnnF Wide/ ower showy stickseed erect cryptanth common hound’s- tongue cupseed stickseed Common name Rollins’ cryptanth western stickseed showy stoneseed western stoneseed spindle-rooted bluebell lanceleaf bluebell western bluebell meadow popcorn- ﬂ

var. M. M. viridis var. var. redowskii cupulata Synonyms/ notes Taxonomic redowskii Included in oblongifolia Included in in Cronquist et al. (1984) P. scouleri P. hispidulus, P. nelsonii hispidulus, P.

oribunda ssp. cinale Cynoglossum ofﬁ Lappula marginata L. texana, Lappula redowskii L. redowskii Lithospermum incisum Lithospermum ruderale Mertensia lanceolata Mertensia oblongifolia penicillatus Plagiobothrys scouleri

Species name var. var.

var. oribundafl Hackelia var. cinale Hackelia fl Lappula occidentalis Cryptantha stricta Oreocarya stricta Cynoglossum offi cupulata Lappula occidentalis UtahCryptantha rollinsii Oreocarya rollinsii Colorado occidentalis Lithospermum incisum Lithospermum ruderale Mertensia fusiformis coriacea Mertensia lanceolata Mertensia oblongifolia Plagiobothrys scouleri penicillatus rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Family Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 31 Rainbow Draw, Rainbow Draw, Uintah Co., UT (Goodrich UT 12312 BRY). BLM: Sensitive Colorado? Comments 1945 1987 1989 1912 1948 1935 locality is Type 1955 1970 New record for Year Year doc. 1935 1935 1987 1948 (DINO) O’Kane 3927 (DINO) 345 (DINO) 4443 (DINO) et al. 483 (DINO) (1937) 3173 (DINO) (COLO) Source 2775 (DINO) (BRY) 3019 (DINO) Dever s.n. (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc Neely 4328 Pres X Unc Neely & Pres X Unc Naumann Pres X X Unc Rodeck Pres X X Unc Holmgren Rep X ? Graham Pres X X Unc O’Kane Pres X Unc s.n. Weber Park stat. Pres X X Unc O’Kane Hist X ? 363 Welsh Pres X Unc O’Kane Pres X Unc & Wolf Wide Wide Wide Wide Wide NA Periph Periph Range UT/CO Wide NA Wide Wide Shrub Wide/ PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF LocEn/ Shrub Wide/ PerF Periph/ Life form PerF Wide/ PerF Wide/ PerF Wide/ ower blue elderberry Simpson’s Simpson’s hedgehog cactus common hops PerF Wide/ yellow beeplant Rocky Mountain beeplant Goodrich’s Goodrich’s stinkweed long-ﬂ snowberry slenderspine pricklypear Common name claretcup PerF Wide/ Utah pricklypear brittle pricklypear plains pricklypear

var. Flora

Flora & var. in

inermis. var.

Var. not Var. var. (2003).

ssp. spina ruﬁ Flora of North caerulea, S. cerulea Includes var. simpsonii C. hillmanii goodrichii of North America (2003) Includes not recognized Vars. in America Synonyms/ notes Taxonomic O. polyacantha hystricina. recognized in of North America (2003). O. rhodantha. Flora of North America (2003) includes vars. trichophora Included with O. polyacantha polyacantha

var. minor var. orus americanus Pediocactus simpsonii Humulus lupulus ssp. Symphoricarpos longiﬂ Echinocereus triglochidiatus brachyartha Opuntia polyacantha

var. Species name var. var. var. orus neomexicanus utahensis Sambucus caerulea Sambucus coerulea S. nigra Pediocactus simpsonii Humulus lupulus var. Cleome lutea Cleome lutea Cleome serrulata Cleome serrulata Cleomella palmeriana goodrichii UtahEchinocereus triglochidiatus melanacanthus Colorado Symphoricarpos longifl Opuntia erinacea var. Opuntia fragilis Opuntia fragilis Opuntia polyacantha polyacantha Opuntia polyacantha trichophora rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Caprifoliaceae (Adoxaceae) Cactaceae Cannabaceae Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) Cactaceae Family Caprifoliaceae Cactaceae Cactaceae Cactaceae Cactaceae biogeographic notes, cont. Appendix A. Confi

32 Annotated Checklist of Vascular Flora: Dinosaur National Monument Europe Europe Europe Sensitive Comments is Logan, Cache Co., UT (Rydberg s.n. US). near Duchesne, Duchesne Co., UT (Ripley & Barneby 8701 NY). 1948 1935 1988 Native to 1987 NA Native to 1991 Native to 1948 1912 1983 CO BLM: Year Year doc. 1955 1935 locality Type 1965 1987 locality is Type 1912 Dever 5188 (DINO) 2682 (DINO) 300 (DINO) (DINO) (2002) (DINO) 2733 (DINO) 41a (DINO) and Neese (1986) Source (BRY) Dever 5145 (DINO) 374 (COLO) 2949 (DINO) et al. 454 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc & Wolf Pres X X Com O’Kane Pres X Unc Naumann Pres X Unc Neely 3980 Rep X ? Naumann Pres X Unc Reidel 116 Pres X X Unc O’Kane Pres X X Com MacLeod Rep X ? Goodrich Park stat. Hist X ? 315 Welsh Pres X X Com & Wolf Hist X ? MacLeod Pres X Unc O’Kane Pres X X Com Holmgren Wide Wide Intro Wide Intro Intro Wide Wide Periph Range UT/CO Wide Wide Wide RegEn Wide PerF Wide/ Shrub Wide/ AnnF Intro/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Life form Shrub Periph/ Shrub Wide/ PerF RegEn/ Shrub Wide/ eld James’ chickweed mountain- lover salt sand- spurrey ﬁ chickweed holosteum AnnF Intro/ bouncing-bet PerF Intro/ ball-head sandwort Eastwood’s Eastwood’s sandwort Nuttall’s Nuttall’s sandwort Common name western snowberry mountain snowberry silver orach AnnF Wide/ Hooker’s Hooker’s sandwort fourwing saltbush

var.

var. var. hookeri E. ssp. includes var. (2005) cephaloidea nuttallii Flora of North myrsinites strictum E. congesta congesta, var. E. eastwoodiae eastwoodiae Included in hookeri in America var. Synonyms/ notes Taxonomic occidentalis cinalis jamesiana Paxistima myrsinites Pachistima Atriplex argentea Saponaria ofﬁ Holosteum umbellatum Eremogone congesta Eremogone eastwoodiae Eremogone hookeri Symphoricarpos occidentalis Symphoricarpos rotundifolius Atriplex canescens A. canescens

Species name var. argentea congesta eastwoodiae desertorum canescens cinalis Stellaria jamesiana Pseudostellaria Pachystima myrsinites Atriplex argentea var. Saponaria offi Spergularia marina Spergularia marina S. salina Holosteum umbellatum Cerastium arvense Cerastium strictum C. arvense Arenaria congesta var. Arenaria fendleri var. Arenaria hookeri var. Arenaria nuttallii Minuopsis nuttallii Minuartia nuttallii UtahSymphoricarpos occidentalis Colorado utahensis Symphoricarpos oreophilus Atriplex canescens var. rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Caryophyllaceae (Alsinaceae) Celastraceae Chenopodiaceae Caryophyllaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Caprifoliaceae Family Caprifoliaceae Chenopodiaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 33 Eurasia Comments is near Emery, is near Emery, Emery Co., UT (Jones 5443 RM) Salt Lake City, Salt Lake City, Salt Lake Co., UT (Jones 1760 POM). is Gunnison, Eurasia Sanpete Co., UT (Jones 6525 POM) 1978 1983 1949 1935 1948 Native to Year Year doc. 1912 1912 1912 locality Type NA locality is Type 1948 locality Type 1955 1961 Native to al. 13445 (CSU) & Wathen & Wathen 97 (DINO) 5355 (DINO) (1937) Dever 5251 (DINO) Source (DINO) Dever 5077 (DINO) (DINO) and Neese (1986) Dever 5235 (DINO) (BRY) et al. 450 (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc Wilken et Pres X Unc Friedlander Hist X ? Bradley Rep X ? Graham Hist X ? & Wolf Park stat. Pres X X Com 5424 Weber Pres X Unc & Wolf Pres X Unc 5365 Weber Rep X ? Goodrich Hist X ? & Wolf Pres X Unc 452 Welsh Hist X ? Holmgren Wide Wide RegEn Wide Intro Range UT/CO Wide RegEn RegEn NA RegEn Wide Intro AnnF Wide/ AnnF Wide/ AnnF Intro/ Life form Shrub RegEn/ AnnF Intro/ ve-horn pitseed goosefoot mountain goosefoot stalked orach AnnF RegEn/ wedge orach AnnF Wide/ summer- cypress Common name shadscale Shrub Wide/ mat saltbush Shrub RegEn/ Castle Valley Castle Valley saltbush basin saltbush Shrub Wide/ slender orach AnnF RegEn/ greenmolly PerF Wide/ ﬁ smotherweed

var. var. var. var. A. C. berlandieri zschackei C. fremontii atrovirens A. tridentata, includes tridentata caput-medusae Synonyms/ notes Taxonomic robusta itenuissima A. Chenopodium berlandieri Chenopodium atrovirens Atriplex saccaria A. saccaria Atriplex confertifolia Atriplex gardneri A. cuneata Atriplex wolﬁ

var. Species name i

var. utahensis saccaria cuneata Chenopodium album berlandieri Chenopodium atrovirens Atriplex gardneri var. Atriplex saccaria var. Atriplex truncata Atriplex truncata Atriplex wolfi Bassia scoparia Bassia sieversiana Kochia scoparia UtahAtriplex confertifolia Colorado Atriplex corrugata Atriplex corrugata Atriplex gardneri var. tenuissima Bassia americana Kochia americana Bassia hyssopifolia Bassia hyssopifolia rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Family Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae biogeographic notes, cont. Appendix A. Confi

34 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Indian Creek, Mtns., Tushar Beaver Co., UT (Martin & 17148 Welsh BRY) noxious weed (List C). Native to Eurasia 1949 1950 1948 1984 1935 1950 1933 Year Year doc. 1960 locality is Type 1961 1961 1933 1983 CO state 793 (BRY) (DINO) 283 (DINO) & Wathen & Wathen 128 (DINO) (DINO) (DINO) 7742 (CM) Source 49a (DINO) et al. 458 (DINO) et al. 438 (DINO) 30b (DINO) & Wathen & Wathen 89 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Brotherson Pres X Unc 5369 Weber Pres X X Unc Naumann Pres X Unc Friedlander Pres X X Unc Neely 4315 Pres X X Unc 5388 Weber Hist X ? Graham Pres X Unc MacLeod Park stat. Pres X Unc Holmgren Hist X ? Holmgren Pres X X Unc MacLeod Pres X Unc Friedlander Wide Wide Wide Wide Wide Wide Periph Wide Range UT/CO Wide Sparse Wide Intro AnnF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ AnnF Periph/ AnnF Wide/ Life form AnnF Wide/ AnnF Wide/ Fremont’s Fremont’s goosefoot silvery goosefoot oakleaf goosefoot narrow-leaf goosefoot winterfat Shrub Wide/ Nuttall’s Nuttall’s poverty-weed tiny poverty- weed smallhead goosefoot Common name maple-leaf goosefoot American bugseed spiny hopsage Shrub Wide/ halogeton AnnF Intro/ Vars. Vars.

& var. (2003). occidentale Flora of North C. incanum incanum, C. incanum var. C. salinum Ceratoides lanata, Eurotia lanata. not recognized in America pusilla C. overi Synonyms/ notes Taxonomic C. hybridum, gigantospermum, C. standleyanum Includes vars. americanum rydbergii Chenopodium fremontii Chenopodium incanum Chenopodium glaucum Chenopodium leptophyllum Krascheninnikovia lanata Monolepis nuttalliana Chenopodium foliosum Chenopodium simplex Corispermum americanum Halogeton glomeratus Species name

var. lanata var. var. var. var. Chenopodium fremontii fremontii Chenopodium fremontii incanum salinum Chenopodium glaucum Chenopodium leptophyllum Krascheninnikovia lanata Monolepis nuttalliana Monolepis pusilla Monolepis pusilla Micromonolepis UtahChenopodium capitatum parvicapitatum Colorado Chenopodium simplex Corispermum americanum Grayia spinosa Atriplex grayi Atriplex spinosa Halogeton glomeratus rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Family Chenopodi aceae Chenopodiaceae Chenopodiaceae biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 35 Comments is 15 miles E of Roosevelt, Uintah Co., UT (Stutz 9325 BRY) Year Year doc. 1948 Native to Asia 1948 1955 1983 1948 1948 1987 1987 1948 1955 1961 1948 1934 locality Type Source Dever 5225 (DINO) 2764 (DINO) (DINO) & Wathen & Wathen 127 (DINO) Dever 5055 (DINO) (DINO) 3056 (DINO) 3200 (DINO) Dever 5113 (DINO) et al. 485 (DINO) et al. 446 (DINO) & Wathen & Wathen 42 (DINO) (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc & Wolf Park stat. Pres X X Com O’Kane Pres X X Com Neely 3920 Pres X Unc Friedlander Hist X ? & Wolf Pres X X Com MacLeod 39 Pres X Unc O’Kane Pres X X Com O’Kane Pres X X Com & Wolf Pres X X Unc Holmgren Pres X Unc Holmgren Pres X Unc Friedlander Hist X ? 5430 Weber Intro Range UT/CO Wide Wide NA Wide Wide Periph Wide Wide Wide Wide Wide RegEn Life form AnnF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Shrub RegEn/ Russian thistle AnnF Intro/ Common name greasewood Shrub Wide/ low pussytoes PerF Wide/ western seepweed Torrey’s Torrey’s seepweed common yarrow orange agoseris pale agoseris PerF Wide/ cutleaf agoseris bur ragweed AnnF Wide/ skeleton-leaf bursage tall pussytoes PerF Wide/ Plummer’s Plummer’s siltbush

var. var.

by Vars. Vars. S. var. ora (2006) Flora of North pestifer Synonyms/ notes Taxonomic Included in calceoliformis some authors plummeri, Grayia brandegei occidentalis. not recognized in America A. parviﬂ Franseria acanthicarpa Franseria tomentosa plummeri

var. var. ssp. aurantiaca Sarcobatus vermiculatus Antennaria dimorpha Suaeda moquinii S. nigra, fruticosa Atriplex brandegei Z. brandegeei Achillea lanulosa A. millefolium Agoseris aurantiaca var. Agoseris glauca glauca Agoseris glauca laciniata Ambrosia acanthicarpa Ambrosia tomentosa Antennaria pulcherrima anaphaloides

var. var.

Species name ssp. torreyana plummeri aurantiaca UtahSalsola tragus Salsola australis Colorado S. kali, iberica, Sarcobatus vermiculatus Antennaria dimorpha Suaeda occidentalis Suaeda torreyana var. Zuckia brandegei var. lanulosa Achillea millefolium Agoseris aurantiaca var. Agoseris glauca glauca Agoseris glauca laciniata Ambrosia acanthicarpa Ambrosia tomentosa Antennaria anaphaloides rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Chenopodiaceae Chenopodiaceae (Sarcobataceae) Family (Asteraceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae biogeographic notes, cont. Appendix A. Conﬁ

36 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Year Year doc. 1945 1987 NA 1987 1960 2002 1961 1991 2002 1955 1949 1949 1948 Source (DINO) 2689 (DINO) (2002) 3185 (DINO) 14a (DINO) ed.) et al. 405 (DINO) (DINO) ed.) (DINO) et al. 480 (DINO) 263 (DINO) Dever 5061 (DINO) c name with taxonomic and Pop. size UT CO Park stat. Pres X X Com Neely 4187 Pres X X Unc O’Kane Rep X ? Naumann Pres X Unc O’Kane Hist X ? MacLeod Rep X ? NCPN (in Pres X Unc Holmgren Pres X Unc Reidel 156 Rep X Unc NCPN (in Pres X Unc Reidel 127 Pres X Unc Holmgren Pres X X Com Naumann Pres X X Com & Wolf Range UT/CO Wide Wide Intro Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Shrub Wide/ Shrub Wide/ Shrub Wide/ Shrub Wide/ PerF Wide/ Common name small-leaf pussytoes common pussytoes burdock PerF Intro/ heartleaf arnica orange arnica PerF Periph/ low sagebrush Shrub Wide/ biennial wormwood Bigelow’s Bigelow’s sagebrush silver sagebrush tarragon PerF Wide/ fringed sagebrush black sagebrush Louisiana wormwood

Flora

in var. var. var. Includes var. (2006). (2006) A. rosea in Welsh et in Welsh Vars. not Vars. Flora of North Flora of North Includes Synonyms/ notes Taxonomic Vars. not recognized Vars. in America Misspelled var. cordata al. (2003). A. arbuscula arbuscula. A. longiloba Cronquist (1994) biennis Vars. not recognized Vars. in Cronquist (1994) and America nova. recognized in of North America (2006)

. ssp. var ssp. viscidulum Antennaria microphylla Antennaria parvifolia Arnica cordifolia arbusculum arbusculum Seriphidium canum ssp. Oligosporus dracunculus glaucus ludoviciana Artemisia ludoviciana Seriphidium novum A. arbuscula

var.

var. Species name ssp. var. cordifolia UtahAntennaria microphylla Colorado Antennaria parvifolia Arctium minus Arctium minus Arnica cordifolia var. Arnica fulgens Arnica fulgens Artemisia arbuscula Seriphidium Artemisia biennis Artemisia biennis A. biennis Artemisia bigelovii Artemisia bigelovii Artemisia cana viscidula glauca Artemisia dracunculus Artemisia frigida Artemisia frigida Artemisia ludoviciana ludoviciana Artemisia nova nova rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 37

ora pauciﬂ Comments is Left Fork of Hobble Creek, Utah Co., UT (Goodrich et al. 21492 BRY) var. var. 1948 Year Year doc. 1912 1983 1955 2002 1988 1991 1988 locality of Type 1989 1960 1979 Dever 5118 (DINO) Source Dever 5067 (DINO) & Wathen 71 (DINO) Riedel 296 (DINO) ed.) 383 (DINO) (DINO) 229 (DINO) 380 (DINO) 392 (DINO) (1982) c name with taxonomic and Pop. size UT CO Pres X X Com & Wolf Park stat. Pres X X Unc & Wolf Pres X X Unc Friedlander Pres X X Abun Naumann & Rep XPres X Unc X NCPN (in Unc Naumann Pres X Unc Reidel 145 Pres X Unc Naumann Pres X X Unc Naumann Pres X Rare Naumann Rep X ? et al. Weber Wide Range UT/CO Wide Wide Wide Wide Wide Wide Wide Wide Periph Wide Shrub Wide/ Life form Shrub Wide/ AnnF Wide/ Shrub Wide/ PerF Wide/ c aster c PerF Wide/ Wyoming big sagebrush Common name budsage Shrub Wide/ Paciﬁ basin big sagebrush rayless alkali aster blueleaf aster PerF Wide/ Siskiyou aster PerF Wide/ mountain big sagebrush white heath aster elegant aster PerF Wide/ western aster PerF Wide/

var.

ssp. var. var. A. ora pansus, Synonyms/ notes Taxonomic A. chilensis adscendens, A. adscendens, Symphyotrichum ascendens Symphyotrichum ciliatum glauca Includes tridentata Herrickia glauca Symphyotrichum lanceolatum hesperium var. Symphyotrichum ericoides pauciﬂ pansum Symphyotrichum spathulatum

ssp. ssp. angusta hesperius Seriphidium tridentatum wyomingense Picrothamnus desertorum ascendens Seriphidium tridentatum tridentatum ssp. Seriphidium vaseyanum ssp. Species name

var. var. var. Artemisia tridentata wyomingensis UtahArtemisia spinescens Colorado Aster ascendens Virgulaster Artemisia tridentata tridentata Aster brachyactis Brachyactis ciliata Artemisia tridentata vaseyana Aster glaucodes Eucephalus glaucusAster hesperius Eurybia glauca, Aster lanceolatus Aster pansus Virgulus ericoides A. ericoides Aster perelegans Eucephalus elegans Aster spathulatus Aster spathulatus A. occidentalis, rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Compositae (Asteraceae) Compositae Compositae (Asteraceae) Family (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ

38 Annotated Checklist of Vascular Flora: Dinosaur National Monument noxious weed, CO state noxious weed (List B). Native to Europe noxious weed, CO state noxious weed (List B). Native to Mediterranean noxious weed, CO state noxious weed (List B). Native to Eurasia. Comments is Lake Point, Co., UT Tooele (Jones 1727 UC) American Fork, Jordan Valley, Utah Co., UT 493 (Watson YU) NA UT state NA UT state 1948 1946 2004 UT state 1948 2002 Year Year doc. 1987 locality Type 1962 locality is Type (2002) (2002) et al. 466 (DINO) & Wathen & Wathen 26 (DINO) DINO V-2 DINO V-2 (DINO) Dever 5105 (DINO) ed.) Source (DINO) (BRY) c name with taxonomic and Pop. size UT CO Rep X ? Naumann Rep X ? Naumann Pres X X Unc Holmgren Pres X X Unc Friedlander Pres X Unc Richards Pres X X Unc & Wolf Rep X Unc NCPN (in Park stat. Pres X Unc Neely 4257 Hist X ? 1779 Welsh Intro Intro Wide Wide Intro Wide Wide Range UT/CO Periph Wide PerF Intro/ PerF Intro/ PerF Wide/ Shrub Wide/ PerF Wide/ PerF Wide/ Life form PerF Wide/ Shrub Wide/ ower spotted knapweed diffuse diffuse knapweed tassel-ﬂ brickellbush rough brickellbush musk thistle PerF Intro/ arrowleaf balsamroot devil’s- devil’s- beggarsticks Common name hairy balsamroot Mohave brickellbush

ssp. micranthos, C. biebersteinii Includes ssp. leiophyllus Synonyms/ notes Taxonomic

var.

var. ora Acosta maculosa C. stoebe Brickellia grandiﬂ Brickellia microphylla scabra macrolepis Balsamorhiza sagittata Balsamorhiza hispidula Brickellia oblongifolia

Species name var. var.

var. ora Centaurea maculosa Centaurea diffusa Acosta diffusa Brickellia microphylla scabra Brickellia oblongifolia Balsamorhiza sagittata Bidens frondosa Bidens frondosa Brickellia grandifl Carduus nutans Carduus nutans UtahBalsamorhiza hookeri hispidula Colorado linifolia rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 39 Comments noxious weed, CO state noxious weed (List B). Native to Eurasia is St. George, Washington Co., UT (Palmer s.n. NY) 1988 1965 Year Year doc. 1989 UT state 1912 1912 1912 1949 1948 1961 1949 1991 locality Type 1935 301 (DINO) 766 (BRY) Source 388 (DINO) Dever 5117 (DINO) (DINO) (DINO) 5369 384 (DINO) et al. 462 (DINO) 5368 (DINO) (DINO) et al. 481 (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc Naumann Pres X Unc Brotherson Park stat. Pres X Unc Naumann Pres X X Com & Wolf Pres X Unc 5367 Weber Pres X X Com Neely 3987 Pres X Unc Bradley Pres X X Unc Naumann Pres X Unc Holmgren Pres X X Unc Bradley Pres X Unc Reidel 157 Pres X X Com Holmgren Wide NA Range UT/CO Intro Wide Wide Wide Wide Wide Wide RegEn Wide Wide Shrub Wide/ Shrub Wide/ Life form PerF Intro/ PerF Wide/ AnnF Wide / Shrub Wide/ Shrub Wide/ Shrub Wide/ Shrub RegEn/ PerF Wide/ Shrub Wide/ longbract rabbitbrush Nevada rabbitbrush Common name Russian knapweed hoary dusty- maiden stevia dusty- maiden rose-heath PerF Wide/ spear-leaf spear-leaf rabbitbrush greenish rabbitbrush glabrate rabbitbrush smoothseed rabbitbrush hispid golden- aster dwarf rabbitbrush

var. var.

minor

var. var. montana var. hispida, var. var. & var. attenuatus consimilis, graveolens, leiosperma Includes vars. douglasii Ericameria parryi howardii Ericameria parryi nevadensis, C. parryi var. Synonyms/ notes Taxonomic Lorandersonia linifolia Ericameria nauseosa var. E. nauseosa oreophila, C. nauseosus oreophilus Ericameria nauseosa var. E. nauseosa glabrata Ericameria nauseosa var. hispida, Heterotheca villosa H. villosa

ssp. ssp. ssp. howardii ssp. Chaenactis douglasii Chrysothamnus parryi Chaenactis stevioides Leucelene ericoides Aster arenosus Heterotheca villosa C. villosa Chrysothamnus linifolius Chrysothamnus nauseosus consimilis Chrysothamnus nauseosus graveolens Chrysothamnus nauseosus leiospermus Chrysothamnus depressus Species name

var. var. var.

howardii var. var. minor Chaenactis douglasii Chrysothamnus parryi nevadensis Chrysothamnus parryi UtahCentaurea repens Acroptilon repens Colorado Chaenactis stevioides Chaetopappa ericoides Chrysopsis villosa var. Chrysothamnus linifolius Chrysothamnus nauseosus consimilis Chrysothamnus nauseosus graveolens Chrysothamnus nauseosus leiospermus Chrysothamnus depressus rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ

40 Annotated Checklist of Vascular Flora: Dinosaur National Monument noxious weed (List B). Native to Europe Comments noxious weed, CO state noxious weed (List B). Native to Eurasia is Horse Trail is Horse Trail Canyon, Dinosaur NM, Uintah Co., 343 UT (Welsh UT BLM: BRY). Sensitive, CO BLM: Sensitive, USFWS: former C2 candidate noxious weed (List C). Native to Eurasia 1961 NA 1912 1988 CO state Year Year doc. 1961 UT state 1955 locality Type 1988 1955 1989 CO state & Wathen & Wathen 82 (DINO) (2002) Dever 5285 (DINO) 378 (DINO) Source 213 (DINO) 3168 (DINO) & Campbell 309 (DINO) et al. 478 (DINO) 389 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Friedlander Rep X ? Naumann Pres X X Unc & Wolf Pres X X Unc Naumann Park stat. Pres X Unc MacLeod Pres X X Rare O’Kane Pres X Com Naumann Pres X X Com Holmgren Pres X Unc Naumann Wide Wide Wide Intro Range UT/CO Intro LocEn Wide Wide Intro AnnF Wide/ PerF Wide/ Life form PerF LocEn/ Shrub Wide/ Shrub Wide/ Canadian horseweed Ownbey’s Ownbey’s thistle elk thistle PerF Wide/ wavy-leaved thistle bull thistle PerF Intro/ Common name Canada thistle PerF Intro/ slenderleaf rabbitbrush green rabbitbrush chicory PerF Intro/

Flora by some

& (2006) orus Flora of North Vars. not recognized Vars. in America Synonyms/ notes Taxonomic Includes vars. horridum integrifolium Included in var. Included in var. viscidiﬂ not authors. Var. recognized in of North America (2006).

ssp. ssp. orus orus orus Conyza canadensis Cirsium scariosum Cirsium undulatum Chrysothamnus viscidifl stenophyllus Chrysothamnus viscidifl viscidifl

Species name var. var. orus orus orus glabrata scariosum undulatum Conyza canadensis var. Cirsium ownbeyi Cirsium ownbeyi Cirsium scariosum var. Cirsium undulatum var. Cirsium vulgare Cirsium vulgare UtahChrysothamnus viscidifl Colorado Cirsium arvense Breea arvensis stenophyllus Chrysothamnus viscidifl Cichorium intybus Cichorium intybus viscidifl rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 41 Comments is ridge above Bear River Canyon, Uinta Mtns., Watson 546 GH) Year Year doc. 1948 1912 1935 1947 1948 1987 1912 1983 1991 1912 1987 1948 1948 locality Type Source 3220 (DINO) (BRY) (DINO) (1962) & et al. Weber (1982) (DINO) 3032 (DINO) 341 (DINO) (DINO) (DINO) (DINO) (DINO) (DINO) Dever 5183 (DINO) c name with taxonomic and Pop. size UT CO Park stat. Pres X Unc O’Kane Hist X ? 170 Welsh Pres X X Unc Neely 4399 Rep X ? Holmgren Pres X Unc Neely 4306b Pres X Unc O’Kane Pres X Unc Naumann PresPres X X Unc Neely 3984 Unc Reidel 47 Pres X X Rare Neely 4131 Pres X Unc Neely 4460 Pres XPres X Com 5289 Wolf X Unc & Wolf Range UT/CO Wide RegEn Wide Periph Wide Wide Periph Wide Wide Periph Wide Wide Wide Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ eabane Common name mountain hawksbeard noddinghead PerF RegEn/ gray hawksbeard tufted ﬂ Modoc hawksbeard western hawksbeard glaucous hawksbeard Engelmann’s daisy smooth daisy PerF Wide/ nakedstem PerF Wide/ cut-leaved daisy spreading daisy Eaton’s daisyEaton’s PerF Wide/

ssp. var.

var. var. Synonyms/ notes Taxonomic C. modocensis modocensis glabellus discoideus eatonii runcinata acuminata intermedia Erigeron caespitosus modocensis Erigeron compositus E. compositus Psilochenia occidentalis Psilochenia glauca ssp. Erigeron engelmannii Enceliopsis nudicaulis Erigeron divergens

Species name

var. var. occidentalis glauca divergens UtahCrepis acuminata Psilochenia Colorado Enceliopsis nutans Enceliopsis nutans Encelia nutans Crepis intermedia Psilochenia Erigeron caespitosus Crepis modocensis Psilochenia Erigeron compositus glabratus Crepis occidentalis var. Crepis runcinata var. Erigeron engelmannii Enceliopsis nudicaulis nudicaulis Erigeron glabellus Erigeron glabellus E. glabellus Erigeron divergens var. Erigeron eatonii Erigeron eatonii E. eatonii rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Compositae (Asteraceae) Family (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ

42 Annotated Checklist of Vascular Flora: Dinosaur National Monument is E. wilkenii is Mill Creek, Summit Co., UT (Payson 4984 RM) near mouth of Pool Canyon, Dinosaur NM, Co., CO Moffatt (O’Kane 3014 NY) Comments of 1950 1912 1948 locality Type Year Year doc. 1956 locality Type 1989 1955 1948 1988 1961 1961 1948 (DINO) 2650 (DINO) Dever 5162 (DINO) Source (DINO) 386 (DINO) et al. 423 (DINO) 4429 (DINO) & Campbell 298 (DINO) Reidel 285 (DINO) et al. 484 (DINO) Dever 5092 (DINO) c name with taxonomic and Pop. size UT CO Hist X ? 5425 Weber Pres X X Com O’Kane Pres X Unc & Wolf Park stat. Pres X X Com Neely 3934 Pres X Unc Naumann Pres X X Unc Holmgren Pres X X Unc Rodeck Pres X Unc Naumann Pres X Unc Naumann & Hist X ? Holmgren Pres X X Unc & Wolf Wide Wide RegEn Range UT/CO LocEn Wide Wide Wide Wide Wide Wide Wide PerF Wide/ Life form PerF RegEn/ PerF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ PerF Wide/ ower eabane eabane spotted joe- pye weed basin daisy PerF Wide/ Navajo fl Uinta daisy PerF RegEn/ Common name needle-leaf fl western goldenrod common blanketfl slender cudweed lowland cudweed cottonbatting cudweed curly-cup gumweed

var. var. includes var. (2006) subglaber Flora of North Eupatoriadelphus maculatus bruneri, Eutrochium maculatum bruneri concinnus, var. Synonyms/ notes Taxonomic Solidago occidentalis G. chilense Vars. not recognized Vars. in America serrulata Eupatorium maculatum Erigeron pulcherrimus Erigeron concinnus E. concinnus Erigeron uintahensis Erigeron wilkenii Euthamia occidentalis Gnaphalium uliginosum Gnaphalium palustre Pseudognaphalium stramineum Grindelia squarrosa var.

Species name var. concinnus uintahensis serrulata Eupatorium maculatum bruneri Erigeron pulcherrimus Erigeron pumilus var. Erigeron speciosus var. UtahErigeron nematophyllus Colorado Euthamia occidentalis Gaillardia aristata Gaillardia aristata Gnaphalium exilifolium Gnaphalium palustre Gnaphalium stramineum Grindelia squarrosa var. rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 43 Comments Orchard Creek Dinosaur Draw, NM, Duchesne Co., UT Shaw, (Welsh, Matthews, & Moore 9471 USFWS: BRY). 3C Abajo Mtns., San Juan Co., UT (Newberry s.n. GH) 1948 1983 1935 1989 1955 Year Year doc. 1965 locality is Type 1948 1987 1948 1989 locality is Type et al. 419 (DINO) & Wathen & Wathen 100 (DINO) et al. 421 (DINO) 387 (DINO) 207 (DINO) Source 9471 (BRY) 11a (DINO) 2697 (DINO) 4436 (DINO) & Jennings 395 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Holmgren Pres X Unc Friedlander Pres X X Unc Holmgren Pres X Unc Naumann Pres X Unc MacLeod Pres X Rare et al. Welsh Park stat. Pres X X Com MacLeod Pres X X Unc O’Kane Pres X X Unc Rodeck Pres X Unc Naumann Wide Wide Wide Wide Wide NA Range UT/CO Wide Wide Wide RegEn Shrub Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ AnnF Wide/ Shrub LocEn/ Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF RegEn/ ower ower ower ower ower broom snakeweed little sunfl common sunfl Nuttall’s Nuttall’s sunfl prairie sunfl orchard snakeweed Common name stemless goldenweed thrift goldenweed fall sneezeweed purpledisk sunfl

var.

acaulis var. ssp. var. var. (2006). (2006). Flora of North Flora of North Ssp. not recognized in America nuttallii Includes var. Includes var. petiolaris pomariensis, Xanthocephalum sarothrae pomariensis G. sarothrae Synonyms/ notes Taxonomic S. armerioides armerioides Vars. not recognized Vars. in America

var. ora Gutierrezia sarothrae Stenotus acaulis S. acaulis Helianthella uniﬂ Helianthus annuus Helianthus petiolaris Stenotus armerioides Helenium autumnale montanum Helianthella microcephala

Species name fallax var. var. acaulis ssp. var. ora lenticularis Gutierrezia sarothrae Haplopappus acaulis Helianthella unifl Helianthus annuus ssp. Helianthus nuttallii Helianthus nuttallii H. nuttallii Helianthus petiolaris UtahGutierrezia pomariensis Colorado armerioides Haplopappus armerioides Helenium autumnale montanum Helianthella microcephala rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi

44 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Blue Mountain, Uintah Co., UT (Neese & Fulmer 11734 UT BLM: BRY). Sensitive, USFWS: 3C Europe Year Year doc. 1961 1987 1955 locality is Type 1955 1971 1948 1983 Native to 1912 1948 1955 Source 215 (DINO) 2977 (DINO) 353 (DINO) (DINO) 2661 (DINO) (DINO) & Wathen & Wathen 108 (DINO) 3024 (DINO) Dever 5023 (DINO) (DINO) c name with taxonomic and Pop. size UT CO Park stat. Pres X X Unc MacLeod Pres X Unc O’Kane Pres X Rare Naumann Pres X X Unc Neely 4252 Pres X Unc O’Kane Pres X X Unc Neely 4265b Pres X Unc Friedlander Pres X X Com O’Kane Pres X X Unc & Wolf Pres X X Unc Neely 3983 Range UT/CO Wide Wide NA Wide RegEn Wide Intro Wide RegEn Wide Life form PerF Wide/ PerF LocEn/ PerF Wide/ PerF RegEn/ PerF Wide/ PerF RegEn/ PerF Wide/ Common name blue lettuce PerF Wide/ lanate hymenoxys rock hymenoxys Colorado rubber-plant Torrey’s Torrey’s hymenoxys poverty-weed PerF Wide/ prickly lettuce AnnF Intro/ common hyalineherb Wyoming hyalineherb Arizona hymenoxys

Flora lanata,

Flora H. var. var. var. lifolius by orus, var. in var. var. var. vars. not Tetraneuris Tetraneuris caespitosa paucifl lifolius lifolius lifolius Synonyms/ notes Taxonomic L. pulchella, oblongifolia, Mulgedium pulchellum H. acaulis Tetraneuris acaulis Tetraneuris var. Included in torreyana Cronquist (1994) and torreyana of North America (2006) axillaris, recognized in of North America (2006). H. fi lugens, H. fi var. H. fi megacephalus, H. fi tomentosus arizonica

var.

luteus var. var. lifolius lifolius Lactuca tatarica pulchella Hymenopappus ﬁ Hymenoxys lapidicola richardsonii Picradenia richardsonii Tetraneuris Tetraneuris torreyana cinereus Hymenopappus ﬁ Tetraneuris acaulis Tetraneuris acaulis T. Tetraneuris Tetraneuris brevifolia

var.

Species name var.

luteus var. var. arizonica caespitosa lifolius lifolius UtahHymenopappus fi Colorado Hymenoxys richardsonii Lactuca tatarica pulchella cinereus Hymenoxys lapidicola richardsonii Hymenoxys torreyana Iva axillaris Iva axillaris I. axillaris Lactuca serriola Lactuca serriola Hymenopappus fi Hymenoxys acaulis var. Hymenoxys acaulis var. rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 45 is Green River, is Green River, Emery Co., UT (Jones 54821 POM) Comments noxious weed. CO state noxious weed (List B). Native to Eurasia shores of Great Salt Lake, UT (Stansbury s.n. GH) 1987 1948 1912 1931 1948 1912 locality Type Year Year doc. 1948 NA Utah state 1948 1935 1912 locality is Type 2760 (DINO) (DINO) (DINO) 5361 (DINO) (DINO) 5302 (1949) Source Wolf 5295 Wolf (DINO) (2002) Neely 2967 (DINO) (DINO) 7762 (CM) c name with taxonomic and Pop. size UT CO Pres X Unc O’Kane Pres X X Unc Neely 3962 Pres X X Unc Neely 4141 Pres X X Unc Bradley Pres X X Unc Kelley 88-80 Pres X Unc Bradley Park stat. Hist X ? Dever & Rep X ? Naumann Pres X X Com O’Kane & Hist X ? 5377 Weber Hist X ? Graham Wide Wide Wide Wide Wide NA Range UT/CO Wide Intro Wide Wide Wide PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Life form PerF Wide/ PerF Wide/ AnnF Wide/ AnnF Wide/ woolly groundsel western groundsel showy rush- pink hoary aster PerF Wide/ rock- goldenrod Uintah bahia PerF LocEn/ Common name nodding microseris Scotch thistle PerF Intro/ gumweed aster sowthistle desert- dandelion Torrey’s Torrey’s desert- dandelion

var. var. canescens Aster canescens, Dieteria canescens var. Bahia nudicaulis ourolepis Synonyms/ notes Taxonomic Haplopappus nuttallii, Xanthisma grindelioides grindelioides ora Senecio integerrimus Lygodesmia Lygodesmia grandiﬂ Machaeranthera canescens Petradoria pumila Solidago petradoria Machaeranthera grindelioides Onopordum acanthium Malacothrix sonchoides

Species name

var. var. var. var. var. ora ora pumila Senecio canus Packera cana Senecio integerrimus exaltatus Machaeranthera canescens canescens Machaeranthera grindelioides Petradoria pumila var. ourolepis Platyschkuhria integrifolia Utah Lygodesmia grandifl grandifl Colorado grindelioides Microseris nutans Microseris nutans Onopordum acanthium Malacothrix sonchoides Malacothrix torreyi Malacothrix torreyi rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi

46 Annotated Checklist of Vascular Flora: Dinosaur National Monument . ed Comments is Uinta River, is Uinta River, Uintah Co., UT (Fremont 549 GH) Europe. Report from Holmgren (1962) based on misidentiﬁ specimen of S.uliginosus 1983 1948 1987 Year Year doc. 1956 locality Type 1948 1955 1955 1961 2005 Native to & Wathen & Wathen 107 (DINO) Dever 5123 (DINO) (DINO) Source Neely 2657 (DINO) 376 (DINO) 45b (DINO) 289 (DINO) et al. 440 (DINO) Andersen (2006) c name with taxonomic and Pop. size UT CO Pres X Unc Friedlander Pres X Unc & Wolf Pres X Unc Neely 4385 Park stat. Pres X X Com O’Kane & Pres X X Unc Naumann Pres X X Unc MacLeod Pres X X Unc Naumann Pres X X Unc Holmgren Rep ? ? Rare Dewey & Wide Wide Wide Range UT/CO Wide Wide Wide Wide Wide Intro PerF Wide/ PerF Wide/ PerF Wide/ Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Intro/ eld sow- eld broom groundsel Rocky Mountain groundsel montane groundsel Common name basin groundsel Canadian goldenrod Missouri goldenrod baby goldenrod three-nerve goldenrod ﬁ thistle

var.

Flora Vars. Vars. var. var. Vars. not Vars. ora, (2006) (2006) (2006). ora streptanthifola. salebrosa missouriensis, Flora of North Flora of North Flora of North Vars. not recognized Vars. in America P. streptanthifolia P. var. not recognized Vars. in America P. werneriifolia P. werneriifolia. not recognized in America Synonyms/ notes Taxonomic var. S. missouriensis var. includes var. extraria. recognized in of North America (2006). S. velutina sparsiﬂ arvensis Senecio spartioides Packera streptanthifolia Packera werneriifolia Packera multilobata Solidago canadensis S. altissima, lepida Solidago missouriensis

var. Species name var.

spartioides Senecio spartioides var. Senecio streptanthifolius var. streptanthifolius Senecio werneriifolius werneriifolius Solidago UtahSenecio multilobatus Colorado canadensis salebrosa Solidago missouriensis Solidago nana Solidago nana Solidago velutina Solidago velutina S. sparsifl Sonchus arvensis Sonchus arvensis S. arvensis rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 47 Eurasia Comments Europe Eurasia 1948 1948 1983 1955 1933 Native to 1948 1912 2002 1948 1912 Year Year doc. 1955 Native to 1987 Native to 1956 1945 Dever 5155 (DINO) Dever 5031 (DINO) and Neese (1986) (BRY) 2751 (DINO) Dever 5013 (DINO) (DINO) (DINO) Dever 5193 (DINO) Neely 2659 (DINO) Source 379 (DINO) (DINO) 9625 Welsh (COLO) & Jensen 13958 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc & Wolf Hist X Unc & Wolf Rep X ? Goodrich Hist X Unc 448 Welsh Pres X X Unc O’Kane Pres X Unc & Wolf Pres X X Unc Neely 4157 Pres X Unc s.n. Tilley Hist X ? & Wolf Pres X X Com O’Kane & Pres X X Unc Naumann Park stat. Pres X Unc Neely 3978 Hist X Unc & Weber Pres X X Unc Holmgren Wide RegEn Periph Wide Intro Sparse Wide Wide Wide Wide Intro Range UT/CO Intro Wide Wide Shrub Wide/ Shrub RegEn/ PerF NA/ AnnF Wide/ PerF Intro/ PerF Wide/ Shrub Wide/ PerF Wide/ PerF Wide/ PerF Intro/ Life form PerF Intro/ PerF Wide/ PerF Wide/ spineless horsebrush Nuttall’s Nuttall’s horsebrush silver chicken- sage white-plume wire-lettuce common dandelion desert wirelettuce shortspine horsebrush Hopi-tea PerF Periph/ Hooker’s Hooker’s townsendia hoary townsendia marsh sowthistle Common name red-seed dandelion common townsendia slender wirelettuce

exigua minor. minor. var. ssp. var. ssp. var. (2006) Flora of North Tanacetum nuttallii Tanacetum S. exigua spinosa glabrescens, S. arvensis uliginosus Synonyms/ notes Taxonomic erythrospermum T. S. minor not recognized Vars. in America cinale Tetradymia Tetradymia canescens Sphaeromeria argentea Stephanomeria exigua Taraxacum ofﬁ Stephanomeria runcinata Thelesperma megapotamicum Townsendia hookeri Townsendia Taraxacum Taraxacum laevigatum leptotes Townsendia Stephanomeria tenuifolia Species name

var. cinale Tetradymia Tetradymia canescens Tetradymia nuttalliiTetradymia nuttallii Tetradymia Tetradymia spinosaTetradymia spinosa Tetradymia spinosa T. Stephanomeria exigua Taraxacum offi Stephanomeria runcinata Stephanomeria tenuifolia Thelesperma megapotamicum Townsendia Townsendia hookeri Townsendia incanaTownsendia incana Townsendia UtahSonchus uliginosus Sonchus uliginosus S. arvensis Colorado Taraxacum Taraxacum laevigatum Townsendia leptotes tenuifolia rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi

48 Annotated Checklist of Vascular Flora: Dinosaur National Monument Europe noxious weed, CO state noxious weed (List C). Native to Eurasia Comments is ‘near Duchesne’, Duchesne Co., UT (Jones s.n. POM). USFWS: 3C. New to Colorado? Cisco, Grand Co., UT (Jones s.n. POM) 1948 1949 1935 1948 1950 1948 Native to 1991 Utah state Year Year doc. 1987 locality Type 1949 1933 locality is Type & Wathen & Wathen 101 (DINO) Kuntz 3050 (DINO) 275 (DINO) Dever 5120 (DINO) (DINO) (DINO) (DINO) Source 2742 (DINO) et al. 431 (DINO) (BRY) c name with taxonomic and Pop. size UT CO Pres X Unc Friedlander Pres X X Unc O’Kane & Pres X X Unc Naumann Pres X X Com & Wolf Hist X ? 5334 Weber Pres X X Unc 5291 Wolf Pres X Unc Reidel 147 Park stat. Pres X Unc O’Kane Pres X X Unc Holmgren Pres X Unc 178 Welsh Wide Periph Wide Wide RegEn Intro Intro Range UT/CO LocEn Wide RegEn PerF Wide/ PerF Wide/ Shrub Wide/ PerF Wide/ PerF RegEn/ PerF Intro/ Life form PerF LocEn/ AnnF Wide/ PerF RegEn/ eld showy goldeneye rough mule’s- ears red-osier dogwood lance-leaved stonecrop hairy townsendia yellow salsify PerF Intro/ ﬁ bindweed Common name plateau townsendia rough cocklebur Cisco woodyaster

& (2006). scabra Flora of North ssp. Includes ssp. lanceolatum subalpinum Synonyms/ notes Taxonomic X. pensylvanicum, X. saccharatum, vars. not recognized in America venusta

var. ora ora major Heliomeris multiﬂ multiﬂ Xanthium strumarium Swida sericea C. stolonifera lanceolatum Townsendia strigosa Townsendia var. Convolvulus arvensis Townsendia Townsendia mensana

ora var. Species name var. ora var. var. multifl Viguiera multifl var. Wyethia scabra Wyethia scabra Scabrethia scabra Xanthium strumarium canadense Cornus sericea sericea Sedum lanceolatum Amerosedum Townsendia Townsendia strigosa strigosa Tragopogon dubiusTragopogon dubius Tragopogon Convolvulus arvensis Utah Townsendia mensana Colorado Xylorhiza venusta Xylorhiza venusta Machaeranthera rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cornaceae Crassulaceae Compositae (Asteraceae) Compositae (Asteraceae) Convolvulaceae Compositae (Asteraceae) Family Compositae (Asteraceae) biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 49 A.

is . , & A. ora, A. ora, A. russeola 18 miles N of Uintah Vernal, Co., UT (Rollins 1757 GH). USFWS: 3C var. var. Comments Europe perennans considered a hybrid swarm involving drummondii with holboellii, A. sparsiﬂ pendulina 1948 1956 1948 locality of Type Year Year doc. 1950 1983 Native to 1987 1987 Often 1948 1933 Dever 5153 (DINO) (DINO) (DINO) Source 2675b (DINO) Neely 2672 (DINO) (DINO) (DINO) Dever 5195 (DINO) Reidel 2791 (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc & Wolf Pres X X Unc Neely 4371 Pres X Unc Neely 3955 Park stat. Pres X X Unc O’Kane Pres X Unc O’Kane & Pres X Unc Neely 4322 Pres X Unc Neely 4104 Pres X Unc & Wolf Pres X X Unc O’Kane & Wide Wide Wide Range UT/CO RegEn Intro Wide Wide Wide Wide PerF Wide/ PerF Wide/ PerF Wide/ Life form Shrub RegEn/ AnnF Intro/ PerF Wide/ PerF Wide/ PerF Wide/ Drummond’s Drummond’s rockcress tower- mustard nodding rock cress Common name spiny greasebush desert madwort hairy rockcress PerF Wide/ spreadingpod rockcress Fendler’s Fendler’s rockcress Holboell’s Holboell’s rockcress

var. var. includes var.

var. & Vars. Vars. nis, A. nis, russeola Boechera angustifolia, B. stricta Vars. not recognized Vars. in Intermountain Flora (2005) Includes vars. glabrata B. demissa languida, B. oxylobula, var. Synonyms/ notes Taxonomic pycnocarpa A. conﬁ divaricarpa, B. brachycarpa spatifolia, A. fendleri. not recognized in Holmgren et al. (2005) A. holboellii retrofracta, B. holboellii secunda demissa drummondii var. Forsellesia meionandra Alyssum desertorum Boechera divaricarpa Boechera fendleri B. fendleri Boechera retrofracta

Species name var. brachycarpa fendleri secunda Arabis drummondii Boechera Arabis glabra glabra Turritis Arabis hirsuta Arabis hirsuta Arabis demissa Boechera demissa UtahGlossopetalon spinescens meionandrum Colorado Alyssum desertorum Arabis holboellii var. Arabis holboellii var. Arabis holboellii var. rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Crossosomataceae Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) biogeographic notes, cont. Appendix A. Conﬁ

50 Annotated Checklist of Vascular Flora: Dinosaur National Monument is Jones Hole National Fish Hatchery, Uintah Co., & UT (Welsh Neese 18341 UT BLM: BRY). Sensitive, CO BLM: Sensitive, USFWS: former C2 candidate W shore Great Salt Lake, UT (Stansbury s.n. NY) noxious weed. Native to Europe Comments Colorado? is Westwater, is Westwater, Grand Co., UT (Jones s.n. US) Europe 1955 1948 locality Type 2002 Native to Asia 1952 locality is Type 1933 2003 UT state Year Year doc. 1983 1987 First report for 1912 locality Type 1987 Native to Asia 1946 Native to (DINO) 163 (DINO) (DINO) (DINO) (DINO) DINO V-3 DINO V-3 (DINO) Source & Wathen 5 & Wathen (DINO) (DINO) (DINO) (DINO) 2038 (CSU) c name with taxonomic and Pop. size UT CO Pres X X Unc Neely 4036 Pres X X Unc Naumann Pres X Unc s.n. Tilley Pres X X Unc Neely 4042 Pres X X Unc Neely 4343 Pres X Unc Richards Park stat. Pres X Unc Friedlander Pres X Unc Neely 4103 Pres X X Unc 5335 Weber Pres X Unc Neely 4066 Hist X X ? Harrington Wide Disj Intro Periph Wide Intro Range UT/CO Wide Periph Wide Intro Intro PerF Wide/ PerF Intro/ PerF Wide/ PerF Intro/ Life form PerF Sparse/ PerF Wide/ PerF Wide/ AnnF Intro/ AnnF Intro/ ax little-pod false Selby’s Selby’s rockcress park rockcress PerF LocEn/ hairy whitetop spindlestem PerF Wide/ perennial rockcress orbicular whitetop Common name Sweetwater rockcress Nuttall’s Nuttall’s rockcress desert rockcress ﬂ shepherd’s- purse

selbyi ssp. var. repens A. fernaldiana chalepensis, C. draba var. Synonyms/ notes Taxonomic pallens Camelina microcarpa fernaldiana Caulanthus crassicaulis Boechera perennans Cardaria chalepensis C. draba Boechera pulchra ssp. Capsella bursa- pastoris

var.

Species name var. perennans Camelina microcarpa Arabis vivariensis Boechera Cardaria pubescens Cardaria pubescens Caulanthus crassicaulis crassicaulis Arabis perennans var. Arabis pulchra Arabis selbyi Boechera selbyi B. selbyi Cardaria chalepensis UtahArabis lignifera Boechera lignifera Colorado Arabis nuttallii Boechera nuttallii pallens Capsella bursa- pastoris rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 51 C2 candidate Eurasia Comments Europe 1987 1955 1961 1956 USFWS: former 1948 Native to Year Year doc. 1987 Native to Asia 1948 Native to 1988 1912 1948 1955 1912 2747 (DINO) Neely 2678 (DINO) et al. 457 (DINO) 231 (DINO) 2703 (DINO) Source (DINO) Dever 5226 (DINO) (DINO) Dever 5074 (DINO) Roszczewski 3197 (DINO) (BRY) (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc O’Kane Pres X X Unc O’Kane & Pres X Unc Holmgren Pres X X Rare Naumann Pres X X Unc O’Kane Park stat. Pres X X Unc Neely 4065 Pres X Unc & Wolf Pres X Unc Painter 413 Pres X X Com & Wolf Pres X X Unc O’Kane & Hist X ? 221 Welsh Pres X X Com Painter 455 Sparse Wide Wide RegEn Intro Range UT/CO Intro Intro Wide Wide Wide Periph Sparse PerF Wide/ PerF Wide/ Life form PerF Wide/ PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ ower ower ixweed AnnF Intro/ few-seed whitlow-grass dwarf draba AnnF Wide/ western wallfl sand dune wallfl Uinta draba PerF RegEn/ woods draba AnnF Intro/ Common name blue mustard AnnF Intro/ fl California tansy-mustard mountain tansy-mustard western tansy- mustard western tansy- mustard

var. E. or

var. var. D. D. by some in var. var. var. Included capitatum incisa, D. incana capitatum, incana var. E. asperum arkansanum arkansanum, capitatum purshii. in var. in Holmgren et al. (2005) Included in oligosperma pectinipila authors Synonyms/ notes Taxonomic D. richardsonii sonnei, D. incana ssp. var. Included in pinnata osmiarum Holmgren et al. (2005) incisa intermedia halictorum Draba oligosperma Erysimum capitatum E. capitatum Erysimum asperum E. asperum Draba juniperina Descurainia californica Descurainia incisa ssp. Descurainia pinnata ssp. Descurainia pinnata ssp. Species name oligosperma elatum purshii juniperina sonnei intermedia osmiarum Draba oligosperma var. Draba reptans Draba reptans Erysimum asperum var. Erysimum asperum var. Draba oligosperma var. Draba nemorosa Draba nemorosa UtahChorispora tenella Chorispora tenella Colorado Descurainia sophia Descurainia sophia Descurainia californica Descurainia incana var. Descurainia pinnata var. Descurainia pinnata var. rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) biogeographic notes, cont. Appendix A. Conﬁ

52 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments noxious weed, CO state noxious weed (List B). Native to Eurasia Europe 18 miles N of Uintah Vernal, Co., UT (Rollins 1758 GH) Europe 1912 1912 Year Year doc. 1948 1950 1912 1965 1983 UT state 1948 1956 Native to Africa 1949 Native to 1948 locality is Type 1912 1987 Native to Neely 2666 (DINO) 2771 (DINO) Source Dever 5138 (DINO) (DINO) 7688 (CM) 809 (BRY) & Campbell 306 (DINO) (DINO) (DINO) 189 (DINO) 2746 (DINO) et al. 464 (DINO) 2755 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Com O’Kane & Pres X X Unc O’Kane Park stat. Pres X X Unc & Wolf Hist X ? 5437 Weber Hist X ? Graham Hist X Unc Brotherson Pres X Unc Naumann Pres X Unc Neely 4050 Pres X X Unc Neely 4216 Pres X X Unc Naumann Pres X X Unc O’Kane Pres X X Com Holmgren Pres X Unc O’Kane Wide Wide Range UT/CO Wide Sparse Wide NA Intro Periph Intro Intro RegEn Wide Intro PerF Wide/ PerF Wide/ PerF RegEn/ Life form AnnF Wide/ AnnF Wide/ PerF Intro/ PerF NA/ AnnF Intro/ PerF Wide/ AnnF Intro/ ower PerF Wide/ sharp-leaf twinpod silver bladderpod Rollins’ bladderpod Common name shy wallﬂ ovalpurse AnnF Wide/ prairie pepperwort St. George pepperwort broad-leaf pepperwort branched pepperwort African mustard watercress PerF Intro/ Jones’ pepperwort clasping pepperwort Synonyms/ notes Taxonomic Hornungia procumbens Includes var. Includes var. macrocarpum L. bourgeauanum Rorippa nasturtium- aquaticum

var. var. var. orum cinale Physaria acutifolia australis P. Lesquerella ludoviciana Lesquerella subumbellata Erysimum inconspicuum Hymenolobus procumbens Lepidium densiﬂ ramosum Lepidium montanum bourgeauanum Lepidium ramosissimum Nasturtium ofﬁ jonesii Lepidium perfoliatum

Species name var. var. var. orum acutifolia cinale Lepidium montanum Physaria acutifolia var. Physaria ludoviciana Physaria subumbellata UtahErysimum inconspicuum Colorado Hutchinsia procumbens Lepidium densifl ramosum Lepidium lasiocarpum georginum Lepidium latifolium Cardaria latifolia jonesii Malcolmia africana Malcolmia africana Nasturtium offi Lepidium perfoliatum rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 53 is Westwater, is Westwater, Grand Co., UT (Jones s.n. POM) Comments Europe is Stansbury Island, Tooele Co., UT (Watson 72 NY) 1948 1987 1987 1933 locality Type Year Year doc. 1961 1948 1948 1948 1961 Native to 1948 1912 1950 locality Type 1948 1933 375 (DINO) (DINO) (DINO) 3027 (DINO) Source et al. 401 (DINO) Dever 5231 (DINO) Dever 5224 (DINO) Dever 5196 (DINO) 210 (DINO) 770 (BRY) (BRY) (DINO) Dever 5167 (DINO) 4430 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Naumann Pres X Unc Neely 4093 Pres X Unc Neely 4041 Pres X X Unc O’Kane Hist X ? Holmgren Park stat. Hist X ? & Wolf Hst X X ? & Wolf Pres X X Unc & Wolf Pres X X Unc MacLeod Pres X Unc Brotherson Pres X Unc 376 Welsh Pres X Unc 5317 Weber Pres X Unc & Wolf Pres X X Abun Rodeck RegEn Wide Wide RegEn Sparse Range UT/CO Wide Wide Wide Intro Wide Wide Periph Wide Wide PerF RegEn/ PerF Wide/ PerF Wide/ PerF RegEn/ PerF Wide/ Life form PerF Wide/ PerF Sparse/ PerF Wide/ AnnF Intro/ PerF Wide/ AnnF Wide/ Shrub Wide/ ower ax-leaved ddle- Moab thelypody alpine pennycress heart-leaf jewelfl elegant thelypody smooth yellowcress Common name hairy yellowcress spreading yellowcress fl plainsmustard tumble mustard smoothleaf prince’s-plume prince’s-plume PerF Wide/ long-beak fi mustard common juniper Utah juniper Shrub Wide/

var. var. var. montana S. cordatus cordatus R. palustris fernaldiana Synonyms/ notes Taxonomic Sisymbrium linifolium alpina integrifolia glabra hispida pinnata Thelypodium integrifolium Noccaea montana N. montana Stanleya pinnata var. Streptanthus cordatus Thelypodiopsis elegans Rorippa palustris var. Rorippa palustris var. Schoenocrambe linifolia Sisymbrium altissimum var. Streptanthella longirostris Juniperus communis ssp. Sabina osteosperma

Species name

var. var. montanum glabra hispida Thelypodium integrifolium gracilipes Thlaspi montanum var. Juniperus communis Stanleya integrifolia Streptanthus cordatus Thelypodiopsis elegans UtahRorippa islandica var. Colorado Rorippa islandica var. Rorippa sinuata Rorippa sinuata Schoenocrambe linifolia Sisymbrium altissimum Stanleya pinnata Stanleya pinnata Streptanthella longirostris depressa Juniperus osteosperma rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cupressaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cupressaceae biogeographic notes, cont. Appendix A. Conﬁ

54 Annotated Checklist of Vascular Flora: Dinosaur National Monument C. . ed from DINO herbarium are based on misidentiﬁ specimens of nebrascensis Comments 1993 Other reports 1987 1961 1987 1987 1989 USFWS: 3C 1948 1988 1987 1948 Year Year doc. 1948 1961 (2007) (DINO) et al. 451 (DINO) Neely 3080 (DINO) 2817 (DINO) 340 (DINO) (DINO) (DINO) 3189 (DINO) Dever 5213 (DINO) Source 5341 (DINO) & Jensen 13980 (DINO) c name with taxonomic and Pop. size UT CO Rep X Rare Fowler et al. Pres X X Com Neely 4383 Hist X ? Holmgren Pres X X Com O’Kane & Pres XPres X X Unc O’Kane Rare Naumann Pres X Unc Neely 4368 Pres X Unc Painter 486 Pres X Unc O’Kane Pres X Unc & Wolf Park stat. Pres X X Unc Bradley Pres X Unc Holmgren Pres X Unc Harms s.n. 2005 Wide Wide Wide Wide Wide Wide NA Wide Wide Wide Range UT/CO Wide Wide Wide PerG Wide/ PerG Wide/ PerG Wide/ PerG RegEn/ PerG Wide/ Life form Tree Wide/ PerG Wide/ water sedge PerG Wide/ small-wing sedge slender- beaked sedge Nebraska sedge golden sedge PerG Wide/ western sedge PerG Wide/ Canyonlands sedge narrowleaf sedge elk sedge PerG Wide/ Hood’s sedgeHood’s PerG Wide/ Common name Rocky Mountain juniper bottlebrush sedge sheep sedge PerG Wide/

var. includes var. var. var. microptera, aquatilis C. limnophila curatorum C. eleocharis, stenophylla Synonyms/ notes Taxonomic var. C. festivella, macloviana microptera, eleocharis ssp. Sabina scopulorum Species name Carex athrostachya Carex athrostachya Carex nebrascensis Carex nebrascensis C. nebraskensis Carex aurea Carex aurea C. garberi Carex occidentalis Carex occidentalis Carex curatorumCarex duriuscula Carex stenophylla C. scirpoidea Carex geyeri Carex geyeri Carex hoodii Carex hoodii UtahJuniperus scopulorum Colorado Carex hystericina Carex hystericina C. hystricina Carex aquatilis Carex aquatilis C. aquatilis Carex illota Carex illota Carex microptera Carex microptera C. microptera rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cupressaceae Family Cyperaceae Cyperaceae Cyperaceae Cyperaceae biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 55

Comments C. specuicola is listed as Threatened by USFWS 1987 1988 1987 1987 1948 1961 1987 1955 1961 Year Year doc. 1988 1946 1934 2754 (DINO) (DINO) 2694 (DINO) (DINO) (DINO) et al. 436 (DINO) Neely 3081 (DINO) (DINO) et al. 449 (DINO) Source 323 (DINO) Wolf 5266 Wolf (DINO) & Jensen 13977 (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc O’Kane Pres XPres X Unc Painter 407 X Com O’Kane Pres X Unc Neely 4369 Pres X Unc Neely 4085 Pres X Unc Holmgren Pres X Unc O’Kane & Pres X Unc 348 Welsh Pres X Unc Holmgren Pres X Unc Naumann Park stat. Pres X X Com Dever & Pres X X Com Holmgren Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide Range UT/CO Wide Wide PerG Wide/ AnnG Wide/ PerG Wide/ PerG Wide/ Life form PerG Wide/ at- ower woolly sedge PerG Wide/ blackcreeper sedge Ross’ sedge PerG Wide/ dryland sedge PerG Wide/ valley sedge PerG Wide/ bearded ﬂ sedge few-ﬂ spikerush hardstem bulrush alkali bulrush PerG Wide/ Parry’s sedgeParry’s PerG Wide/ Common name common spikerush pale bulrush PerG Sparse/

& var. var.

acutus var. includes ora, ana E. E. var. var. exa paludosus, C. specuicola boottii, C. brevipes, C. pityophila Includes erythropoda E. pauciﬂ includes suksdorﬁ Schoenoplectus acutus occidentalis, Schoenoplectus acutus Scirpus maritimus var. Schoenoplectus maritimus parryana, UT specimens previously assigned to Synonyms/ notes Taxonomic uniglumis pallidus

acutus ssp. ora ssp. macrostachya Eleocharis quinquefl lacustris maritimus paludosus Species name ora Carex pellita Carex lanuginosa Carex praegracilis Carex praegracilis Carex rossiiCarex tahoensis Carex rossii Carex xerantica C. defl Carex vallicola Carex vallicola Cyperus squarrosus Cyperus aristatus Eleocharis palustris Eleocharis Scirpus acutus Schoenoplectus Scirpus maritimus Bolboschoenus Scirpus pallidus Scirpus pallidus S. atrovirens UtahCarex parryana Carex parryana C. parryana Colorado Eleocharis quinquefl rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Family Cyperaceae biogeographic notes, cont. Appendix A. Confi

56 Annotated Checklist of Vascular Flora: Dinosaur National Monument noxious weed (List B). Native to Europe. Comments Europe 1931 NA CO state Year Year doc. 1912 1987 2005 Native to 1948 1912 1948 1948 1948 1912 1948 (BRY) (2002) Source Kuntz 3040 (DINO) Kuntz 3041 (DINO) Andersen (2006) Dever 5045 (DINO) 5309 (DINO) Dever 5006 (DINO) (DINO) 141 (DINO) 3155 (DINO) (DINO) c name with taxonomic and Pop. size UT CO Hist X X ? 371 Welsh Rep X ? Naumann Park stat. Pres X X Com O’Kane & Pres X Unc O’Kane & Rep ? ? Unc Dewey & Pres X Unc & Wolf Pres X X Com Bradley Hist X ? & Wolf Pres X X Unc Painter 409 Pres X Unc Naumann Pres X X Unc O’Kane Hist X ? 5278 Wolf Wide Intro Range UT/CO Wide Wide Intro Wide Wide RegEn Wide Wide Wide Sparse Shrub Wide/ Life form PerG Wide/ PerG Wide/ Shrub Wide/ Shrub Wide/ Shrub RegEn/ Fern Wide/ Fern Wide/ Fern Sparse/ eld horsetail eld Fern Wide/ silver buffaloberry Torrey’s Torrey’s ephedra Russian olive Tree Intro/ Common name three-square bulrush soft-stem bulrush teasel PerF Intro/ green ephedra Cutler’s Cutler’s ephedra ﬁ tall scouring- rush smooth scouring-rush Ferris’ scouring-rush

var. validus E. hyemale var. var. var. ssp. polyphyllus, E. laevigatum Lepargyrea argentea Scirpus pungens var. Schoenoplectus pungens Synonyms/ notes Taxonomic longispicatus, Schoenoplectus pungens polyphyllus, Schoenoplectus pungens pungens Schoenoplectus tabernaemontani, Schoenoplectus lacustris sylvestris E. praealtum E. kansanum Hybrid of x nis afﬁ x validus ssp. ssp. Shepherdia argentea Elaeagnus angustifolia Schoenoplectus pungens lacustris Ephedra viridis Ephedra cutleri Hippochaete hyemalis Hippochaete laevigata ferrissii

var. var. Species name ferrissii Hippochaete x ne longispicatus affi Shepherdia argentea Ephedra torreyana Ephedra torreyana Elaeagnus angustifolia UtahScirpus pungens var. Colorado Scirpus validus Schoenoplectus Dipsacus sylvestris Dipsacus fullonum D. fullonum Ephedra viridis viridis Ephedra viridis viscida Equisetum arvense Equisetum arvense Equisetum hyemale var. Equisetum laevigatum Equisetum rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Elaeagnaceae Ephedraceae Elaeagnaceae Cyperaceae Family Cyperaceae Dipsacaceae Ephedraceae Ephedraceae Equisetaceae Equisetaceae Equisetaceae Equisetaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 57 Comments Colorado? noxious weed (List B), UT state noxious weed. Native to Europe 1956 1912 1966 2002 NA 1961 NA 1948 Year Year doc. 1961 First report for 1935 NA CO state (DINO) Dever 5002 (DINO) 482 (COLO) (DINO) (2002) Jensen 425 (DINO) (1962) Dever 5209 (DINO) Source & Jensen 13968 (DINO) Dever 5284 (DINO) (2002) c name with taxonomic and Pop. size UT CO Pres X X Unc Neely 4239 Pres X X Unc & Wolf Pres X Unc MacLeod Pres X Unc 2868 Tilley Rep X ? Naumann Pres X Unc Holmgren & Rep X ? Holmgren Pres X X Unc & Wolf Hist X ? Holmgren Pres X X Unc & Wolf Park stat. Rep X NA Naumann Wide Wide Wide Wide Periph Wide Wide Wide Periph Wide Range UT/CO Intro Shrub Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ Fern Sparse/ PerF Wide/ Life form greenleaf manzanita Fendler’s Fendler’s spurge golden corydalis ridge-seeded spurge Great Basin centaury thyme-leaved spurge Rocky Mountain gentian sticky crane’s- bill hybrid scouring-rush shorthorn spurge Common name leafy spurge PerF Intro/

Hybrid esula var. E. hyemale var. mackaii. x E. variegatum Euphorbia glyptosperma Euphorbia serpyllifolia E. between x robusta Synonyms/ notes Taxonomic E. robusta, brachycera

var. x ssp.

ssp. nis Arctostaphylos patula platyphylla Chamaesyce fendleri Euphorbia fendleri Corydalis curvisiliqua occidentalis Chamaesyce glyptosperma Centaurium exaltatum Pneumonanthe afﬁ Chamaesyce serpyllifolia Geranium viscosissimum nervosum Hippochaete mackaii Tithymalus Tithymalus brachyceras

var. Species name var. nis x nis affi Arctostaphylos patula Chamaesyce fendleri Corydalis aurea occidentalis Chamaesyce glyptosperma Centaurium exaltatum Chamaesyce serpyllifolia Gentiana affi var. Geranium viscosissimum incisum UtahEquisetum trachyodon Colorado Euphorbia brachycera Euphorbia esula esula Tithymalus E. esula rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Ericaceae Euphorbiaceae Fumariaceae Euphorbiaceae Gentianaceae Euphorbiaceae Gentianaceae Geraniaceae Equisetaceae Euphorbiaceae Family Euphorbiaceae biogeographic notes, cont. Appendix A. Confi

58 Annotated Checklist of Vascular Flora: Dinosaur National Monument Eurasia Eurasia and Africa Eurasia Comments 2002 1946 2002 Native to 1961 2002 1955 Native to 1912 Native to Year Year doc. 1988 1948 1955 2002 Colorado Plateau Network (in ed.) 1952 (CSU) (DINO) et al. 455 (DINO) Colorado Plateau Network (in ed.) & Jensen 13970 (DINO) (DINO) Source 287 (DINO) Dever 5032 (DINO) & Jensen 13973 (DINO) ed.) c name with taxonomic and Pop. size UT CO Rep X RarePres Northern X Unc Harrington Pres X Unc s.n. Tilley Hist X ? Holmgren Rep Rare Northern Pres X X Unc Holmgren Pres X X Unc Reidel 92 Park stat. Pres X Unc Naumann Pres X X Unc & Wolf Pres X X Unc Holmgren Rep X X Unc NCPN (in Wide Wide Intro Wide Wide Intro Intro Range UT/CO Wide Wide Wide Wide PerG Wide/ PerG Wide/ PerG Intro/ PerG Wide/ PerG Intro/ Life form PerG Wide/ AnnG Wide/ mountain bentgrass shortawn foxtail meadow foxtail spike bentgrass ticklegrass PerG Wide/ redtop PerG Intro/ crested wheatgrass Common name big bluestem PerG Wide/ purple three- awn American sloughgrass blue grama PerG Wide/

ssp. var. var. aequalis geminata A. cristatum pectinatum, A. pectinatum, A. sibiricum, A. desertorum, A. fragile, mongolicum Synonyms/ notes Taxonomic A. glauca, purpurea fendleriana, A. fendleriana

ssp. longiseta Alopecurus aequalis A. aequalis Alopecurus pratensis cristatum Agropyron cristatum Andropogon gerardii var. Beckmannia syzigachne Species name

var. Agrostis variabilis Agrostis variabilis Alopecurus aequalis Alopecurus pratensis Agrostis exarata Agrostis exarata Agrostis scabra Agrostis scabra A. hiemalis, Agrostis stolonifera Agrostis stolonifera A. alba, gigantea UtahAgropyron cristatum Colorado Andropogon gerardii gerardii Aristida purpurea Aristida purpurea Beckmannia syzigachne Bouteloua gracilis Chondrosum gracile rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 59 is Springdale, Washington Co., UT (Jones 6075 POM) noxious weed (List C). Native to Eurasia Comments Eurasia Eurasia and Africa Eurasia 1961 locality Type 1935 CO state 1982 1987 1946 Year Year doc. 1991 Native to 1961 1988 1989 Native to 1987 1987 1956 Native to et al. 468 (DINO) Wolf 5267 Wolf (DINO) Snyder 11894a (BRY) 3086 (DINO) Dever 5255 (DINO) Source (DINO) & Jensen 13965 (DINO) 236 (DINO) 337 (DINO) (DINO) (DINO) Kuntz 3042 (DINO) c name with taxonomic and Pop. size UT CO Hist X ? Holmgren Pres X X Unc Holmgren Pres X X Unc Dever & Pres X Unc Neese & Pres X X Unc O’Kane Pres X X Unc & Wolf Park stat. Pres X Unc Reidel 103 Pres X Unc Naumann Pres X Unc Naumann Pres X Unc Neely 4423 Pres X Unc Neely 4365 Pres X X Unc O’Kane & Wide Wide Intro Sparse Periph Wide Range UT/CO Intro Wide Intro Wide Wide Intro PerG Wide/ PerG Wide/ PerG Wide/ PerG Wide/ PerG Wide/ Life form AnnG Intro/ PerG Wide/ PerG Wide/ slimstem reedgrass Jones’ reedgrass cheatgrass AnnG Intro/ California oatgrass oneside oatgrass desert saltgrass Common name Japanese chess brookgrass PerG Wide/ orchard grass PerG Intro/ nodding brome mountain brome smooth brome PerG Intro/

or in stricta includes var. oras is oras B. porteri B. anomalus Material ascribed to Synonyms/ notes Taxonomic B. commutatus regional fl mostly B. lanatipes. B. marginatus, polyanthus Calamagrostis stricta C. neglecta Calamagrostis scopulorum Danthonia californica Danthonia unispicata carinata Species name Calamagrostis stricta Calamagrostis scopulorum Bromus tectorum Anisantha tectorum Danthonia californica Danthonia unispicata Distichlis spicata Distichlis stricta D. spicata UtahBromus anomalus Bromopsis porteri Bromus frondosus. Colorado Catabrosa aquatica Catabrosa aquatica Dactylis glomerata Dactylis glomerata Bromus japonicus Bromus japonicus B. arvensis, Bromus carinatus Ceratochloa Bromus inermis Bromopsis inermis rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Confi

60 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments noxious weed, CO state noxious weed (List B). Native to Eurasia top of Salina Pass, Sevier Co., UT (Jones 5447 POM) Year Year doc. 1948 1935 1935 1957 1948 1991 UT state 1987 locality is Type 1948 1955 Source et al. 463 (DINO) Kuntz 3039 (DINO) (DINO) 8727 (CSU) 2767 (DINO) Dever 5127 (DINO) (DINO) 209 (DINO) Neely 2970 (DINO) c name with taxonomic and Pop. size UT CO Park stat. Pres X Unc Holmgren Pres X XPres Com X O’Kane & X Com Neely 4428 Pres X Unc Harrington Pres X Unc O’Kane Pres X Unc & Wolf Pres X Unc Reidel 100 Pres X X Unc MacLeod Pres X X Com O’Kane & Range UT/CO Wide Wide Wide Wide Wide Wide Intro Wide Wide Life form PerG Wide/ PerG Wide/ PerG Wide/ PerG Wide/ PerG Wide/ Common name Canada wildrye Great Basin wildrye squirreltail PerG Wide/ blue wildrye PerG Wide/ Salina wildrye PerG Wide/ thickspike wheatgrass quackgrass PerG Intro/ western wheatgrass bluebunch wheatgrass

var.

var. smithii var. inermis . var. var. thsii albicans, lanceolatus, palmeri spicata Synonyms/ notes Taxonomic canadensis elymoides, Sitanion hystrix glaucus dasystachyum, includes var. riparium, A. albicans var. A. albicans var. Agropyron dasystachyum salinus, L. ambiguus, includes ssp salmonis includes vars. griffi & Agropyron spicatum, includes ssp. & spicata Species name UtahElymus canadensis Elymus canadensis E. canadensis Colorado Elymus cinereus Leymus cinereus Elymus elymoides Elymus elymoides E. elymoides Elymus glaucus Elymus glaucus E. glaucus Elymus lanceolatus Elymus lanceolatus E. lanceolatus Elymus salinus Leymus salina E. salina, Leymus Elymus smithii Pascopyrum smithii Agropyron smithii, Elymus repens Elytrigia repens Agropyron repens Elymus spicatus Pseudoroegneria rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Gramineae (Poaceae) Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 61 Comments Europe Eurasia 1955 1967 1912 1987 Year Year doc. 1934 1961 1982 Native to Asia 1988 1991 Native to 1945 1948 1989 Native to O’Kane 3919 (DINO) (DINO) Dever 5098 (DINO) (DINO) Source 3144 (DINO) et al. 434 (DINO) Trent 11804 Trent (BRY) 185 (DINO) (DINO) Roszczewski 3214 (DINO) (DINO) 336 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Neely & Pres X Unc Neely 4213 Pres X X Com & Wolf Pres X Unc Neely 4378 Pres X X Unc O’Kane Park stat. Hist X ? Holmgren Pres X ? Neese & Pres X X Unc Naumann Pres X Unc Reidel 62 Pres X Com O’Kane & Pres X Unc Neely 4167 Pres X Unc Naumann Wide Wide Wide Wide Wide Range UT/CO Wide Intro Wide Intro Wide Wide Intro AnnG Wide/ PerG Wide/ PerG Wide/ PerG Wide/ Life form AnnG Wide/ AnnG Intro/ PerG Wide/ PerG Intro/ six-weeks fescue fowl mannagrass galleta PerG Wide/ meadow barley slender wheatgrass Common name tufted lovegrass annual wheatgrass hairy tridens PerG Wide/ tall fescue PerG Intro/ Idaho fescue PerG Wide/ Rocky Mountain fescue meadow fescue ora glauca,

octoﬂ & G. elata var. latiglumis, trachycaulus, Includes vars. hirtella, Includes var. E. alaskanus ssp. Agropyron caninum, A. subsecundum, A. trachycaulum, latiglume Synonyms/ notes Taxonomic E. diffusa Agropyron triticeum arundinaceum, Schedonorus phoenix, S. arundinaceus F. ovina F. rydbergii Schedonorus pratensis ora Festuca idahoensis Critesion brachyantherum Eragrostis pectinacea Eremopyrum triticeum Festuca arundinacea Lolium Festuca saximontana

Species name var. var. oraoctofl Vulpia Festuca ovina ingrata Festuca octofl Glyceria striata Glyceria striata Hilaria jamesii Hilaria jamesii Pleuraphis jamesii Hordeum brachyantherum Utahtrachycaulus Elymus Elymus trachycaulus E. trachycaulus Colorado Eragrostis pectinacea Eremopyrum triticeum Erioneuron pilosum Erioneuron pilosum Festuca arundinacea Festuca ovina saximontana Festuca pratensis Festuca pratensis Lolium pratense, rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Confi

62 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Year Year doc. 1948 1948 1948 1946 1991 1961 1965 1961 1946 1961 1959 1961 Source Dever 5269 (DINO) Dever 5181 (DINO) (DINO) (DINO) (DINO) & Jensen 13967 (DINO) Salamun 12652 (COLO) et al. 467 (DINO) 1980 (CSU) ey al. 441 (DINO) & Jensen 13962 (DINO) et al. 473 (DINO) c name with taxonomic and Pop. size UT CO Park stat. Pres X X Unc & Wolf Pres X Com & Wolf Pres X Unc Neely 4087 Pres X Unc Neely 4084 Pres X X Unc Reidel 120 Pres X X Unc Holmgren Pres X Unc & Weber Hist X ? Holmgren Hist X ? Harrington Pres X X Unc Holmgren Pres X X Unc Holmgren Pres X Unc Holmgren Range UT/CO Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide Sparse Life form PerG Wide/ PerG Wide/ PerG Wide/ Common name foxtail barley PerG Wide/ Junegrass PerG Wide/ spike fescue PerG Wide/ oniongrass PerG Wide/ foxtail muhly PerG Wide/ scratchgrass PerG Wide/ sandhill muhly PerG Wide/ green muhly PerG Wide/ mat muhly PerG Wide/ Thurber’s Thurber’s muhly littleseed ricegrass bundle panicgrass

var. ssp. M. curtifolia Synonyms/ notes Taxonomic pyramidata, K. nitida Includes P. occidentale, P. tennessense, P. huachucae, D. P. acuminatum fasciculatum, D. lanuginosum fasciculatum Muhlenbergia andina Muhlenbergia asperifolia Muhlenbergia pungens Muhlenbergia racemosa Muhlenbergia richardsonis Muhlenbergia thurberi Piptatherum micranthum Dichanthelium acuminatum Species name UtahHordeum jubatum Critesion jubatum Colorado Koeleria macrantha Koeleria macrantha K. cristata, Leucopoa kingii Leucopoa kingii Hesperochloa kingii Melica bulbosa Bromelica bulbosa Muhlenbergia andina Muhlenbergia asperifolia Muhlenbergia pungens Muhlenbergia racemosa Muhlenbergia richardsonis Muhlenbergia thurberi Oryzopsis micrantha Panicum acuminatum rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Gramineae (Poaceae) Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 63

Eurasia Eurasia and Africa Eurasia and Africa Europe. Comments Eurasia P. pratensis P. have been documented (Naumann 143 DINO) 1933 1955 1987 Native to 1945 1912 1948 Native to 1933 1987 Native to 1987 Native to Year Year doc. 1961 2005 Native to 1987 Hybrids with Neely 2660 (DINO) (BRY) (DINO) 1454 (CSU) Neely 2670 (DINO) Dever 5254 (DINO) Wolf 5081 Wolf (DINO) 2983 (DINO) 3000 (DINO) Source et al. 410 (DINO) (DINO) Roszczewski 3194 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Com O’Kane & Hist X ? 311 Welsh Pres X Unc Neely 4125 Hist X ? Harrington Pres X X Com O’Kane & Pres X X Unc & Wolf Pres X X Unc Dever & Pres X Unc O’Kane Pres X X Com O’Kane Pres X Unc Holmgren Park stat. Pres X Unc Harms 193 Pres X Unc O’Kane & Wide Wide Intro Wide Wide Intro Wide Intro Intro Wide Range UT/CO Intro Wide PerG Wide/ PerG Intro/ PerG Wide/ PerG Wide/ AnnG Intro/ PerG Intro/ PerG Intro/ Life form PerG Wide/ muttongrass PerG Wide/ Nuttall’s Nuttall’s alkaligrass Canada bluegrass interior bluegrass Sandberg’s Sandberg’s bluegrass rabbitfoot- grass common reed PerG Wide/ bulbous bluegrass Kentucky bluegrass witchgrass AnnG Wide/ Common name timothy PerG Intro/ Wheeler bluegrass P.

P. ssp. some P. epilis P. P. gracillima, P. & fendleriana longiligula, authors include cusickii P. interior P. Includes juncifolia, P. P. canbyi, sandbergii, P. scabrella, P. P. ampla nevadensis, P. and all vars. of secunda Synonyms/ notes Taxonomic

ssp. occidentale Puccinellia airoides interior Poa nemoralis Polypogon monspeliensis Phragmites australis communis P. var. Poa nervosa wheeleri P.

Species name var. var. Poa fendleriana Poa fendleriana longiligula, P. P. Poa glauca Puccinellia nuttalliana Poa compressa Poa compressa glauca Poa secunda Poa secunda Polypogon monspeliensis Phragmites australis Poa bulbosa Poa bulbosa Poa pratensis Poa pratensis agassizensis P. UtahPanicum capillare Panicum capillare Colorado Phleum pratense Phleum pratense Poa nervosa wheeleri rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family biogeographic notes, cont. Appendix A. Conﬁ

64 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments could not be substantiated by Goodrich and Neese (1986). Type locality is near Milford, Beaver Co., UT (Jones 2106) Year Year doc. 2005 1961 1912 1948 1955 NA DINO report 1935 1912 1959 1946 1945? Source (DINO) et al. 447 (DINO) Wolf 5096 Wolf (DINO) & Jensen 13971 (DINO) et al. 459 (DINO) (1962) 2955 (DINO) Dever 5003 (DINO) 3102 (DINO) (DINO) 176 (CSU) c name with taxonomic and Pop. size UT CO Pres X Unc Harms s.n. Park stat. Pres X X Unc Holmgren Hist X Com Dever & Pres X X Unc Holmgren Pres X X Com Holmgren Rep X ? Holmgren Pres X X Com O’Kane Pres X X Com & Wolf Pres X Unc O’Kane Pres X Unc Neely 4350 Pres X Unc Harrington Wide Range UT/CO Wide Wide Sparse Wide Periph Wide Wide Wide Wide Wide Life form PerG Sparse/ PerG Wide/ PerG Wide/ PerG Wide/ PerG Wide/ PerG Wide/ PerG Wide/ PerG Sparse/ little bluestem PerG Wide/ Common name prairie cordgrass alkali sacaton PerG Wide/ spike dropseed sand dropseed PerG Wide/ Mormon needlegrass needle-and- thread Indian ricegrass Letterman’s Letterman’s needlegrass Nelson’s Nelson’s needlegrass green needlegrass

var. var. var. var. H. comata comata neomexicanum Andropogon scoparius, S. scoparium Synonyms/ notes Taxonomic Oryzopsis hymenoides A. nelsonii nelsonii, S. columbiana nelsonii Hesperostipa comata Schizachyrium scoparium Sporobolus airoides Sporobolus contractus Sporobolus cryptandrus aridum hymenoides lettermanii Achnatherum nelsonii

Species name var. var. airoides Stipa comata comata UtahSchizachyrium scoparium Colorado Spartina pectinata Spartina pectinata Sporobolus airoides var. Sporobolus contractus Sporobolus cryptandrus Stipa arida Achnatherum Stipa hymenoides Achnatherum Stipa lettermanii Achnatherum Stipa nelsonii nelsonii Stipa viridula Nassella viridula rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 65 Comments is 1 mile E of Red Fleet Dam, Uintah Co., UT (Atwood 7247 BRY) may be from just outside the park (Johnston 2065, COLO) Year Year doc. 1987 1955 1987 1933 1948 1991 1935 1912 1912 locality Type 1991 1948 NA DINO report Source Roszczewski 3196 (DINO) (BRY) 3202 (DINO) (DINO) 204 (DINO) (DINO) (COLO) (DINO) 6612 (BRY) (DINO) Wolf 5237 Wolf (DINO) and Neese (1986) c name with taxonomic and Pop. size UT CO Pres X Unc O’Kane & Park stat. Hist X ? 397 Welsh Pres X Com O’Kane Pres X Unc Reidel 29 Pres X X Unc Naumann Pres X Unc Reidel 38 Hist X X ? 5434 Weber Pres X X Rare Neely 4221 Pres X Unc Atwood Pres X Unc Reidel 64 Pres X X Com Dever & Rep X ? Goodrich Wide Range UT/CO Wide Wide Wide Wide Wide Periph Wide NA Wide Wide Periph Life form PerF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF LocEn/ PerF Wide/ ddleleaf spike trisetum PerG Wide/ Common name common mare’s-tail lanceleaf phacelia Ives’ phacelia AnnF Wide/ silky phacelia PerF Wide/ Rocky Mountain iris leafy ﬁ corrugate phacelia brittle phacelia Idaho blue- eyed grass Baltic rush PerG Wide/ minute rush AnnG Sparse/

var.

& var. montanus, alpina Includes ssp. alaskanum congdonii, T. montanum Synonyms/ notes Taxonomic Includes vars. hastata, leucophylla, & Var. not recognized Var. by Cronquist et al. (1984) S. idahoense occidentale J. arcticus balticus, J. balticus var. includes var. mexicanus

ssp. ssp.

var. Phacelia sericea ciliosa Sisyrinchium idahoense Nama densum Phacelia crenulata occidentale ater

var. var. Species name orum corrugata minor Utah spicatumTrisetum spicatum Trisetum Colorado Hippuris vulgaris Hippuris vulgaris Nama densum Phacelia hastata Phacelia hastata Phacelia ivesiana Phacelia ivesiana Phacelia sericea ciliosa Iris missouriensis Iris missouriensis Sisyrinchium idahoense parvifl Phacelia crenulata var. Phacelia demissa var. Juncus arcticus Juncus arcticus Juncus bryoides Juncus bryoides J. kelloggii rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Gramineae (Poaceae) Hippuridaceae Family Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Iridaceae Iridaceae Hydrophyllaceae Hydrophyllaceae Juncaceae Juncaceae biogeographic notes, cont. Appendix A. Confi

66 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Europe Europe Year Year doc. 1946 2005 Native to 1987 1912 1946 1961 1955 1935 1991 1989 1989 1948 1991 Native to Source et al. 427 (DINO) (DINO) 3109 (DINO) 276 (DINO) 324 (DINO) 252 (DINO) & Jensen 13975 (DINO) & Jensen 13966 (DINO) (DINO) & Campbell 354 (DINO) 331 (DINO) (DINO) (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Holmgren Park stat. Pres X Unc Harms 195 Pres X Unc O’Kane Pres X X Unc Naumann Pres X X Unc Naumann Pres X Unc Naumann Hist X XHist ? Holmgren X ? Holmgren Pres X Unc Reidel 102 Pres X Unc Naumann Pres X Unc Naumann Pres X X Unc 5273 Wolf Pres X Unc Reidel 87 Wide Range UT/CO Intro Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide Intro Life form PerG Intro/ PerG Wide/ PerF Wide/ eld mint eld PerF Wide/ toad rush AnnG Wide/ Common name compressed rush Tracy’s rushTracy’s PerG Wide/ mountain rush PerG Wide/ longstyle rush PerG Wide/ poverty rush PerG Wide/ Torrey’s rushTorrey’s PerG Wide/ seaside arrowgrass American bugleweed self-heal PerF Wide/ little skullcap PerF Wide/ ﬁ catnip PerF Intro/

var. forma T. J.

var. var. & J. by var. J. tracyi, Includes tenuis. J. dudleyi Included in ensifolius Includes vars. bufonius, occidentalis, halophilus, ambiguus Synonyms/ notes Taxonomic brunnescens montanus some authors, saximontanus Includes xiphoides Includes vars. dudleyi, congestus, & elata. concinnum S. galericulata epilobifolia canadensis

var. Juncus ensifolius Juncus saximontanus Mentha arvensis M. arvensis Prunella vulgaris Scutellaria galericulata epilobifolia Species name brunnescens montanus glabrata lanceolata Juncus ensifolius var. UtahJuncus bufonius Juncus bufonius Colorado Juncus compressus Juncus compressus Juncus ensifolius var. Juncus longistylis Juncus longistylis Juncus tenuis Juncus tenuis Juncus torreyi Juncus torreyi Triglochin maritimaTriglochin maritima Triglochin maritima T. americanusLycopus americanus Lycopus Mentha arvensis var. Prunella vulgaris var. Scutellaria galericulata Nepeta cataria Nepeta cataria rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Juncaceae Juncaceae Juncaceae Family Juncaceae Juncaceae Juncaceae Juncaceae Juncaginaceae Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 67 Cisco, Grand Co., UT (Jones s.n. POM) Comments Holmgren (1962) is based on a collection from outside the monument. 13 miles below Theodore, Duchesne Co., UT (Jones s.n. POM). CO BLM: Sensitive 6 miles SE of Jensen, Uintah Co., UT (Ripley & Barneby 7797 CAS). USFWS: 3C 1912 1949 locality is Type 1956 Year Year doc. NA Report from 1912 1950 1933 locality is Type 1912 1912 locality is Type (DINO) 5358 (DINO) (BRY) 5328 (DINO) Source (2002) Dever 5115 (DINO) 5307 (DINO) (DINO) (DINO) c name with taxonomic and Pop. size UT CO Pres X X Rare Neely 4316 Hist X ? Bradley Pres X X Unc 320 Welsh Hist X ? Bradley Park stat. Rep X X ? Naumann Pres X X Unc & Wolf Pres X X Rare Bradley Pres X X Unc Neely 4021 Pres X Rare 5312 Weber Periph RegEn Wide Periph Range UT/CO Wide Wide RegEn RegEn NA PerF Wide/ PerF RegEn/ PerF Wide/ PerF Wide/ Life form PerF Wide/ PerF Wide/ PerF RegEn/ PerF RegEn/ PerF LocEn/ eld Martin’s Martin’s milkvetch milkweed milkvetch Hayden’s Hayden’s milkvetch painted milkvetch Common name ﬁ milkvetch lesser rushy milkvetch Duchesne milkvetch cicada milkvetch grass milkvetch

var. var. Synonyms/ notes Taxonomic A. chamaeleuce A. diversifolius campestris chamaeleuce

var. var. Astragalus argophyllus martinii Astragalus asclepiadoides Astragalus haydenianus Astragalus ceramicus ceramicus Astragalus chamaeleuce Astragalus convallarius Astragalus duchesnensis

Species name

var. var. var. var. Astragalus bisulcatus Astragalus argophyllus martinii Astragalus asclepiadoides haydenianus Astragalus ceramicus ceramicus UtahAstragalus agrestis Astragalus agrestis A. dasyglottis Colorado Astragalus chamaeleuce Astragalus convallarius convallarius Astragalus duchesnensis Astragalus chloodes rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Family Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Conﬁ

68 Annotated Checklist of Vascular Flora: Dinosaur National Monument is eld, candicans Comments is 5 miles S of Vernal, Uintah Co., UT (Hamilton & Beath s.n. RM). UT BLM: Sensitive, USFWS: former C2 candidate is Kanab, Kane Co., UT (Thompson s.n. GH) var. var. near Richﬁ Sevier Co., UT 246 GH) (Ward 1965 1950 2002 Year Year doc. 1949 2002 1987 locality Type 1987 1933 locality Type 1912 locality of Type Salamun 12646 (COLO) (DINO) ed.) Source 5336 (DINO) (DINO) 2768 (DINO) 213 (DINO) 317 (DINO) (DINO) c name with taxonomic and Pop. size UT CO Hist X ? & Weber Pres X X Unc 5346 Weber Rep X Rare NCPN (in Park stat. Hist X ? Bradley Pres X Unc 2781 Tilley Pres X Rare O’Kane Pres X Unc Naumann Pres X X Unc Naumann Hist X ? 5446 Weber Wide Wide NA Range UT/CO Periph Wide LocEn Wide Wide Wide PerF Wide/ PerF Wide/ PerF RegEn/ Life form AnnF Wide/ PerF Wide/ PerF LocEn/ PerF Wide/ PerF Wide/ PerF Wide/ tall kentrophyta Pursh’s Pursh’s milkvetch Trelease’s Trelease’s alkali milkvetch Common name Geyer’s Geyer’s milkvetch standing milkvetch Hamilton’s Hamilton’s milkvetch Rydberg’s weedy milkvetch woolly milkvetch yellow milkvetch

var. var. geyeri

var. var. exus avus Synonyms/ notes Taxonomic A. adsurgens robustior A. lonchocarpus hamiltonii candicans

var.

var. var. avus fl A. Astragalus kentrophyta elatus Astragalus purshii A. infl Astragalus fl Astragalus laxmannii Astragalus hamiltonii Astragalus miser oblongifolius Astragalus mollissimus thompsoniae

Species name

var. var. avus var. var. avus purshii fl oblongifolius Astragalus kentrophyta elatus Astragalus purshii var. Astragalus racemosus treleasei UtahAstragalus fl var. Colorado Astragalus geyeri Astragalus geyeri A. geyeri Astragalus laxmannii robustior Astragalus hamiltonii Astragalus miser var. Astragalus mollissimus thompsoniae rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Family Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 69 Wasatch Mtns. Wasatch or Antelope Island, Salt Lake Co., UT (Watson s.n. GH) Comments is 6 miles N of Jensen, Dinosaur NM, Uintah Co., UT (Holmgren & 9527 NY). Tillett USFWS: 3C 14 miles N of Uintah Vernal, Co., UT (Rollins 1733 RM) 1987 1948 1965 1948 locality is Type Year Year doc. 1912 locality Type 1983 1933 1988 1946 1987 1949 1965 locality is Type Neely 3083 (DINO) Dever 5106 (DINO) 379 (COLO) (DINO) Source 2653 (DINO) Roszczewski 4451 (DINO) 2774 (DINO) (DINO) Dever 5070 (DINO) (DINO) 5300 (DINO) 787 (BRY) c name with taxonomic and Pop. size UT CO Pres X Unc O’Kane & Pres X X Com & Wolf Pres X Unc MacLeod Pres X X Unc 5307 Weber Park stat. Pres X X Rare O’Kane Pres X Unc Neely & Pres X X Com O’Kane Pres X Unc Painter 469 Pres X X Unc & Wolf Pres X X Unc Neely 4039 Pres X X Unc Bradley Hist X ? Brotherson Periph Wide Wide Wide Range UT/CO LocEn Sparse Wide Sparse Wide Wide Periph NA PerF Wide/ PerF Wide/ PerF Wide/ Life form PerF LocEn/ PerF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ PerF LocEn/ white-leaved lupine spoonleaf milkvetch pulse milkvetch Utah lupine PerF Wide/ Common name Dinosaur milkvetch silvery lupine PerF Wide/ silvery lupine PerF Wide/ subalpine lupine Nuttall’s Nuttall’s licorice northern sweetvetch Great Basin lupine Rollins’ sweetvetch

ssp. ssp. var. pabulare var. caudatus, L. argenteus utahensis Synonyms/ notes Taxonomic L. argenteus spathulatus lepidota L. caudatus Some authors include var.

var. argentatus argenteus rubricaulis utahensis Lupinus leucophyllus Astragalus spatulatus Glycyrrhiza lepidota G. lepidota Lupinus pusillus Lupinus argenteus var. Lupinus argenteus var. Lupinus argenteus var. Lupinus caudatus var. Hedysarum boreale intermontanus

var. Species name argentatus argenteus rubricaulis utahensis boreale gremiale Lupinus leucophyllus Lupinus pusillus Astragalus spatulatus Astragalus tenellus Astragalus tenellus Glycyrrhiza lepidota intermontanus UtahAstragalus saurinus Astragalus saurinus Colorado Lupinus argenteus var. Lupinus argenteus var. Lupinus argenteus var. Lupinus caudatus var. Hedysarum boreale var. Hedysarum boreale var. rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae Leguminosae (Fabaceae) Family (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Conﬁ

70 Annotated Checklist of Vascular Flora: Dinosaur National Monument Europe Comments Europe Sensitive Europe Europe 8-9 miles W of Maybell, Moffat Co., CO (Porter 3864 RM) Europe 1948 Native to Year Year doc. 1948 Native to 1912 1946 1933 1987 CO BLM: 1955 1987 Native to 1988 1967 NA Native to 1987 locality is Type 1912 1987 Native to Wolf 5089 Wolf (DINO) Source Dever 5012 (DINO) Wolf 5086 Wolf (DINO) 348 (DINO) (DINO) 2656 (DINO) O’Kane 3925 (DINO) (DINO) (DINO) 13051 (COLO) (1962) (DINO) (DINO) (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Dever & Park stat. Pres X X Unc & Wolf Hist X ? Dever & Pres X X Unc Naumann Pres XPres X Unc Neely 4238 X Rare O’Kane Pres X X Unc Neely & Pres X Unc Neely 4382 Pres X Unc Painter 498 Hist X ? Weber Rep X ? Holmgren Pres X Rare Neely 4273 Hist X ? 5428 Weber Pres X X Unc Neely 4381 Intro Range UT/CO Intro RegEn Wide Periph Periph Wide Intro Wide Wide Intro RegEn Wide Intro PerF RegEn/ PerF Intro/ Life form AnnF Intro/ PerF RegEn/ PerF Wide/ PerF Periph/ PerF Wide/ owered Maybell locoweed yellow sweet- clover Common name white sweet- clover large- ﬂ breadroot lemon scurf- pea prairie golden pea Andean clover PerF Sparse/ dwarf clover PerF Wide/ white clover PerF Intro/ plains vetch PerF Periph/ lesser duckweed alfalfa PerF Intro/ dwarf lupine AnnF Wide/ black medick AnnF Intro/

var.

var. var. sativa

var. M. by some var. var. cinalis O. obnapiformis, O. nana obnapiformis T. gymnocarpon T. Synonyms/ notes Taxonomic Included in ofﬁ authors Psoralea megalantha, Pediomelum megalanthum megalanthum Psoralea lanceolata. not recognized Vars. in Barneby (1989). T. rhombifolia T. rhombifolia andinum gymnocarpon

var. var. cinale obnapiformis Oxytropis besseyi var. Trifolium Trifolium gymnocarpon Pediomelum megalanthum Psoralidium lanceolatum Thermopsis rhombifolia Vicia americana minor Lupinus pusillus pusillus

var. var. Species name var. cinalisoffi Melilotus obnapiformis Oxytropis besseyi var. Melilotus offi Trifolium Trifolium gymnocarpon Vicia americana UtahLupinus pusillus Colorado Melilotus alba Melilotus albus Pediomelum megalanthum Psoralidium lanceolatum lanceolatum Thermopsis rhombifolia Trifolium andinumTrifolium andinum Trifolium andinum T. Trifolium repensTrifolium repens Trifolium minor Lemna minor Lemna turionifera pusillus Medicago sativa Medicago sativa M. sativa Medicago lupulina Medicago lupulina rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Family Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Lemnaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 71 ower of ower Comments is near Kanab, Kane Co., UT (Thompson s.n.) Eurasia. Utah 1986 1987 1948 1912 1948 Year Year doc. 1955 1946 1988 Native to 1912 State ﬂ 2002 1935 1912 1933 locality Type Roszczewski 3191 (DINO) Neely 2673 (DINO) 421 (DINO) Dever 5241 (DINO) 2716 (DINO) Source (DINO) 2731 (DINO) 113 (DINO) Dever 5150 (DINO) (DINO) 154 (DINO) (DINO) (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc O’Kane & Pres X Unc O’Kane & Pres X X Rare Naumann Pres X X Unc & Wolf Pres X X Com O’Kane Park stat. Pres X X Unc Neely 4329 Pres X Unc O’Kane Pres X Unc Naumann Pres X X Unc & Wolf Pres X Unc 2777 Tilley Pres X X Rare Naumann Pres X X Unc 5382 Weber Pres X X Unc Neely 3948 Wide Wide Periph Wide Wide Range UT/CO Wide Wide Intro Wide Wide Periph Wide Wide PerF Wide/ PerF Wide/ Life form PerF Wide/ PerF Wide/ PerF Wide/ glacier-lily PerF Wide/ leopard-lily PerF Wide/ yellow-bell PerF Wide/ starry false Solomon’s-seal foothills death camas Common name taper-tip onion Brandegee’s Brandegee’s onion asparagus PerF Intro/ sego-lily PerF Wide/ Geyer’s onionGeyer’s PerF Wide/ Nevada onion PerF Wide/ textile onion PerF Wide/ pink funnel- lily

var. var. var. orum

A. orum grandifl E. grandifl Synonyms/ notes Taxonomic Some authors include atrorubens cristatum Brodiaea paysonii cinalis orum orum Erythronium grandifl Fritillaria atropurpurea stellatum Toxicoscordion Toxicoscordion paniculatum Asparagus offi Calochortus nuttallii Allium geyeri Androstephium brevifl

Species name var. orum orum cinalis Fritillaria atropurpurea Fritillaria pudica Fritillaria pudica Smilacina stellata Maianthemum Zigadenus paniculatus UtahAllium acuminatum Allium acuminatum Colorado Allium brandegei Allium brandegei A. brandegeei Allium geyeri Asparagus offi Calochortus nuttallii Erythronium grandifl geyeri Allium nevadense Allium nevadense Allium textile Allium textile Androstephium brevifl rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Liliaceae Liliaceae Liliaceae (Convallariaceae, Ruscaceae) Liliaceae (Melanthiaceae) Liliaceae (Alliaceae) Family Liliaceae (Alliaceae) Liliaceae (Alliaceae) Liliaceae (Asparagaceae) Liliaceae (Calochortaceae) Liliaceae Liliaceae (Alliaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae, Themidaceae) biogeographic notes, cont. Appendix A. Confi

72 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments is Natural Bridges NM, San Juan Co., UT (Rydberg & Garrett 9407 NY). USFWS: 3C is 7 miles E of Montrose, Montrose Co., CO (Weber 7470) is Watson- Rainbow Road, E of Evacuation Creek bridge, Uintah Co., UT (Holmgren & Holmgren 14465 NY) 1912 1935 1912 1987 Year Year doc. 1961 locality Type 1960 1948 1912 1950 locality Type 1948 locality Type (DINO) Trent 11800 Trent (BRY) Dever 5040 (DINO) 88 (DINO) Source et al. 470 (DINO) 12a (DINO) Dever 5100 (DINO) (DINO) (DINO) et al. 469 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Neely 4418 Pres X X Unc Neese & Pres X X Unc & Wolf Pres X Unc Kellley 88- Pres X Rare Holmgren Park stat. Pres X Unc MacLeod Pres X X Unc & Wolf Pres X X Unc Painter 464 Hist X ? 5324 Weber Pres X X Unc Holmgren Wide Wide Wide Wide LocEn Range UT/CO Wide Wide Wide RegEn RegEn PerF Wide/ PerF Wide/ PerF Wide/ PerF LocEn/ Life form PerF Wide/ AnnF Wide/ AnnF RegEn/ PerF RegEn/ ax PerF Wide/ scarlet globemallow small-leaf globemallow fragrant sand- verbena wild hollyhock PerF Wide/ alcove death camas Common name meadow death camas blue ﬂ white-stem blazingstar Thompson’s Thompson’s stickleaf Uinta basin stickleaf

var.

& diversa var.

lewisii,

Z. var. var. by some rivularis Includes ssps. coccinea, elata, dissecta fragrans elliptica, A. nana harrisii Z. venenosus Includes vars. & Included in elegans authors Synonyms/ notes Taxonomic venenosus Adenolinum lewisii A. gracilis, M. obscura Nuttallia humilis, M. humilis

ssp. dissecta Sphaeralcea parvifolia Toxicoscordion Toxicoscordion venenosum Sphaeralcea coccinea Anticlea vaginatus Adenolinum lewisii L. lewisii Acrolasia humilis Acrolasia humilis, Nuttallia multicaulis

ssp.

Species name var. Sphaeralcea parvifolia Abronia fragrans Abronia elliptica A. elliptica, Zigadenus venenosus Iliamna rivularis Iliamna rivularis Sphaeralcea coccinea UtahZigadenus vaginatus Colorado Linum perenne lewisii Mentzelia albicaulis Acrolasia albicaulis Acrolasia albicaulis, Mentzelia multicaulis Mentzelia thompsonii uintahensis rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Malvaceae Nyctaginaceae Liliaceae (Melanthiaceae) Malvaceae Malvaceae Liliaceae (Melanthiaceae) Family Linaceae Loasaceae Loasaceae Loasaceae biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 73 Comments is Labyrinth Canyon, San Juan Co., UT (Newberry s.n. NY) Paradox Valley, Paradox Valley, Montrose Co., CO 1988 1933 1933 1983 2002 NA NA 1988 Year Year doc. 1945 1935 1912 locality Type 1912 1965 locality is Type 183 (DINO) 7741 (CM) (DINO) (DINO) (DINO) (1962) (1962) 223 (DINO) Source 5283 Wolf (DINO) 316 (DINO) Dever 5065 (DINO) (DINO) 807 (BRY) c name with taxonomic and Pop. size UT CO Pres X Unc Naumann Hist X ? Graham Hist X ? 5435 Weber Pres X X Unc 2924a Tilley Pres X Unc 2928 Tilley Rep X ? Holmgren Rep X ? Holmgren Pres X Unc Naumann Pres X X Unc Dever & Park stat. Pres X X Unc Naumann Pres X X Com & Wolf Pres X Unc 5371 Weber Hist X ? Brotherson Wide Periph Wide Wide Wide Wide Periph Wide Wide Range UT/CO Wide Wide Wide Periph PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ PerF Periph/ AnnF Wide/ PerF Wide/ Life form AnnF Wide/ AnnF Wide/ ower lavender-leaf lavender-leaf sundrops lesser camissonia tiny camissonia autumn willow-herb northern willow-herb Hall’s willow- Hall’s herb marsh willow- herb deceptive groundsmoke narrow-leaf four-o’clock Common name small-ﬂ sandpuffs singleleaf ash Shrub Wide/ Paiute suncup AnnF Wide/ Walker’s Walker’s camissonia

var. M. Some

ssp. var. var. exuosa ﬂ grammado- C. hartwegii lavandulifolius var. E. paniculatum paniculatum ciliatum, E. watsonii, E. adenocaulon E. palustre palustre, E. palustre var. phyllum O. decumbens. authors include glabra Synonyms/ notes Taxonomic Abronia micrantha, pedunculatus T. Oenothera scapoidea grammado- Calylophus lavandulifolius Epilobium brachycarpum Epilobium halleanum var. phyllum Gayophytum decipiens Oxybaphus linearis Oxybaphus linearis, Tripterocalyx Tripterocalyx micranthus Camissonia scapoidea Species name

var. linearis Calylophus lavandulifolius Camissonia minor Camissonia minor Oenothera minor Camissonia parvula Camissonia parvula Oenothera contorta Camissonia scapoidea Epilobium brachycarpum Epilobium ciliatum Epilobium ciliatum E. ciliatum Epilobium halleanum Epilobium palustre Epilobium palustre Gayophytum decipiens UtahMirabilis linearis var. Colorado Tripterocalyx Tripterocalyx micranthus Fraxinus anomala Fraxinus anomala scapoidea Camissonia walkeri Camissonia walkeri rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Nyctaginaceae Nyctaginaceae Family Oleaceae Onagraceae biogeographic notes, cont. Appendix A. Conﬁ

74 Annotated Checklist of Vascular Flora: Dinosaur National Monument O. var. var. is Greendale Campground, Flaming Gorge NRA, Daggett Co., UT (Neese & Peterson 5428 MO). USFWS: 3C, CO BLM: Sensitive Comments may be annual forms of romat trichocalyx. is Paria River, is Paria River, Kane Co., UT (Cronquist 10202 MO) 1988 1955 1912 1983 1987 1955 1988 locality Type Year Year doc. 1948 1948 DINO reports 1950 locality Type (DINO) (BRY) Dever 5024 (DINO) (DINO) (DINO) & Jennings 394 (DINO) 157 (DINO) Source Dever 5216 (DINO) Porter 5517 (RM) 2704 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc O’Kane Pres X Unc Painter 438 Hist X X ? 231 Welsh Pres X X Unc & Wolf Pres X Unc 2935 Tilley Pres X Unc Neely 4209 Pres X Unc Naumann Pres X Rare Naumann Park stat. Hist X X ? & Wolf Hist X ? Beetle & RegEn Wide Wide RegEn Wide Wide Wide LocEn Range UT/CO Wide Wide PerF RegEn/ PerF Wide/ PerF Wide/ PerF RegEn/ PerF Wide/ PerF Wide/ PerF Wide/ PerF LocEn/ Life form AnnF Wide/ AnnF Wide/ Paria evening- primrose long-tube evening- primrose tufted evening- primrose pale evening- primrose biennial evening-primrose hairy-throat evening- primrose western evening- primrose large yellow evening- primrose Common name branching groundsmoke white-stem evening- primrose O. strigosa, biennis Synonyms/ notes Taxonomic

ssp. ssp. ssp. trichocalyx strigosa hirsutissima navajoensis marginata Oenothera caespitosa caespitosa Oenothera caespitosa Oenothera pallida ssp. Oenothera villosa ssp. Oenothera caespitosa Oenothera coronopifolia ssp. Oenothera acutissima Gayophytum ramosissimum Oenothera albicaulis Species name

ava var. var. var. trichocalyx strigosa acutissima navajoensis Oenothera caespitosa marginata Oenothera caespitosa Oenothera pallida var. Oenothera villosa var. caespitosa Oenothera caespitosa Oenothera coronopifolia Oenothera elata Oenothera elata Oenothera fl var. UtahGayophytum ramosissimum Colorado Oenothera albicaulis rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Family Onagraceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 75 Colorado? Threatened ca 1 mile N of Moab, Grand Co., UT (Welsh & Higgins UT 23269 BRY). BLM: Sensitive, USFWS: former C2 candidate Comments Sensitive 1949 First report for 1989 USFWS: 1987 1948 locality is Type Year Year doc. 1955 1912 USFS Region 2: 1983 1933 1912 1955 1935 5339 (DINO) & Jennings 393 (DINO) (DINO) 4442 (DINO) Source (BRY) 278 (DINO) & Wathen & Wathen 64 (DINO) 7556 (CM) (DINO) (DINO) 2748 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Bradley Pres X X Unc Naumann Pres X Unc Neely 4454 Pres X Rare Rodeck Park stat. Hist X ? 421 Welsh Pres X X Rare Naumann Pres X Unc Friedlander Hist X ? Graham Pres X X Com Neely 4113 Pres X X Com Neely 4260 Pres X X Com O’Kane Periph Sparse Wide RegEn Range UT/CO Wide Sparse Wide Wide Wide Wide Wide PerF Wide/ PerF Sparse/ PerF Wide/ PerF RegEn/ Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Tree Wide/ Tree Wide/ r Tree Wide/ at-topped ﬂ broomrape Ute ladies’- tresses Alaska rein orchid alcove bog- orchid Common name striped coral- root giant helleborine clustered broomrape Cooper’s Cooper’s broomrape two-needle pinyon ponderosa pine Douglas-ﬁ

striata var. var. O. ora, O. ora, Platanthera zothecina Synonyms/ notes Taxonomic Includes multiﬂ cooperi, O. ludoviciana ludoviciana

var. Aphyllon Orobanche corymbosa Spiranthes diluvialis Piperia unalascensis Limnorchis zothecina fasciculatum Orobanche ludoviciana Pinus ponderosa scopulorum Pseudotsuga menziesii

var. Species name

var. ana var. scopulorum Orobanche fasciculata Orobanche corymbosa Spiranthes romanzoffi diluvialis Habenaria unalascensis Habenaria zothecina UtahCorallorhiza striata Corallorhiza striata C. striata Colorado Epipactis gigantea Epipactis gigantea Amesia gigantea Orobanche ludoviciana cooperi Pinus edulis Pinus edulis Pinus ponderosa var. Pseudotsuga menziesii glauca rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Orobanchaceae Orobanchaceae Orchidaceae Orchidaceae Orchidaceae Orchidaceae Family Orchidaceae Orobanchaceae Pinaceae Pinaceae Pinaceae biogeographic notes, cont. Appendix A. Confi

76 Annotated Checklist of Vascular Flora: Dinosaur National Monument is Comments to Europe. Type locality native phase var. pachyphylla Salt Lake City, Salt Lake Co., UT (Jones 1030 POM) 1912 1946 1912 1948 1956 Year Year doc. 1961 Mostly native 1948 1912 1949 1946 1987 5330 (DINO) 2722 (DINO) Neely 2680 (DINO) (DINO) (BRY) Source et al. 402 (DINO) Dever 5119 (DINO) (DINO) 15129 (BRY) Wolf 5234 Wolf (DINO) 3103 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Bradley Pres X X Unc O’Kane Pres X X Unc O’Kane & Pres X X Unc Neely 3966 Hist X ? 497 Welsh Pres X X Unc Holmgren Park stat. Pres X X Unc & Wolf Pres X X Com Neely 4009 Pres X Rare Atwood Pres X X Unc Dever & Pres X Unc O’Kane Wide Wide Periph Wide Wide Intro Range UT/CO Wide Sparse Wide Wide Sparse AnnF Wide/ PerF Intro/ Life form AnnF Wide/- PerF Sparse/ woolly plantain small collomia AnnF Wide/ shy gilia AnnF Wide/ sticky gilia PerF Wide/ gymnosteris AnnF Sparse/ common plantain Common name scarlet gilia PerF Wide/- ball-head gilia PerF Wide/- spreading gilia dwarf gilia AnnF Wide/ matted prickly-phlox

da var. var. includes & patagonica, sinuata, G. P. purshii, P. vars. gnaphaloides, spinulosa, breviscapa Includes vars. inconspicua & ophthalmoides, G. sinuata, G. clokeyi, G. transmontana Includes var. Includes var. pachyphylla Synonyms/ notes Taxonomic Gilia aggregata aggregata Gilia congesta congesta Gilia polycladon Linanthus caespitosus

. dapinnatiﬁ Aliciella congesta Plantago patagonica Gymnosteris parvula Ipomopsis aggregata ssp aggregata Ipomopsis congesta var. Ipomopsis polycladon Leptodactylon caespitosum

Species name dapinnatifi Gilia var. congesta Plantago patagonica Collomia linearis Collomia linearis Gilia inconspicua Gilia sinuata Gilia pinnatifi Gymnosteris parvula aggregata Ipomopsis aggregata UtahPlantago major Plantago major Colorado Ipomopsis congesta var. Ipomopsis polycladon Ipomopsis pumila Ipomopsis pumila Gilia pumila Leptodactylon caespitosum rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Plantaginaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Plantaginaceae Family Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 77

, P. by canescens P. grahamii P. Comments Cottonwood Canyon, Salt Lake Co., UT s.n. (Watson GH) a specimen infested by rust. Price, Carbon Co., UT (Jones s.n. POM) var. var. is S of Great Salt Lake, Co., UT Tooele (Beckwith 4 GH) speciosa Graham (1937). Includes type of 1956 1946 1984 1912 locality of Type 1946 Year Year doc. 1912 1948 locality is Type 1912 Cited as 1949 locality is Type (DINO) (BRY) & Wathen 4 & Wathen (DINO) Dever 5163 (DINO) 2010 (CSU) Source 2959 (DINO) (DINO) 2715 (DINO) 5304 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Unc Neely 3951 Hist X X ? 501 Welsh Pres X Unc Friedlander Pres X X Com & Wolf Hist X ? Harrington Park stat. Pres X X Unc O’Kane Pres X X Unc Neely 4395 Pres X X Unc O’Kane Pres X Unc Bradley Wide Wide Wide Wide Wide Range UT/CO Wide Wide Wide RegEn AnnF Wide/ PerF Periph/ Life form PerF Wide/ PerF Wide/ PerF RegEn/ slender phlox AnnF Wide/ yellow navarretia desert phlox PerF Wide/ carpet phlox PerF Wide/ Rocky Mountain phlox Common name prickly-phlox Shrub Wide/ Watson’s Watson’s prickly-phlox long-leaf phlox Bateman’s Bateman’s wild buckwheat

& ora, ssp.

& var. ssp.

patula humilis Var. not recognized Var. in Cronquist et al. (1984) Includes ssp. glabrata, hoodii, viscidula Includes depressa, multiﬂ & Leptodactylon pungens Synonyms/ notes Taxonomic pungens, Linanthus pungens Linanthus watsonii Includes vars. longifolia stansburyi E. batemanii ora Microsteris gracilis Phlox gracilis Phlox austromontana Phlox hoodii Leptodactylon pungens Leptodactylon watsonii Eriogonum batemanii

var. Species name

var. oramultifl Phlox var. humilior Microsteris gracilis var. Navarretia breweri Navarretia breweri Phlox austromontana austromontana canescens Phlox hoodii Phlox multifl Eriogonum batemanii UtahLeptodactylon pungens Colorado Leptodactylon watsonii Phlox longifolia Phlox longifolia batemanii rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polygonaceae Polemoniaceae Family Polemoniaceae Polemoniaceae biogeographic notes, cont. Appendix A. Confi

78 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments is 8 miles E of Duchesne, Duchesne Co., UT (Reveal 675 UTC). USFWS: 3C is Aquarius Plateau, Wayne Co., UT (Ward 722 GH) is American Fork Canyon, Utah Co., 1033 (Watson US) 1948 Year Year doc. 1967 locality Type 1948 1965 1948 1912 1960 locality Type 1956 1965 locality Type Wolf 5084 Wolf (DINO) Source & Wathen 28 (DINO) 5371 (DINO) Reveal 1853 (BRY) Dever 5258 (DINO) (BRY) 44a (DINO) 3088 (DINO) 954 (BRY) c name with taxonomic and Pop. size UT CO Pres X X Unc Dever & Pres X Rare Friedlander Park stat. Pres X X Unc Bradley Hist X ? Holmgren & Pres X Unc & Wolf Pres X Unc 411 Welsh Hist X ? MacLeod Pres X X Unc O’Kane Hist X ? Brotherson Wide LocEn Range UT/CO Wide RegEn Wide NA Wide Wide Wide AnnF Wide/ PerF LocEn/ Life form Shrub Wide/ AnnF RegEn/ AnnF Wide/ AnnF RegEn/ PerF Wide/ PerF Wide/ AnnF Wide/ nodding wild buckwheat Duchesne wild buckwheat Common name crisp-leaf wild buckwheat bent wild buckwheat Gordon’s wild Gordon’s buckwheat Grand desert Valley trumpet Baker’s wild Baker’s buckwheat whorled wild buckwheat Hooker’s wild Hooker’s buckwheat

var. E. var. & (2005) Flora of North Vars. not recognized Vars. in America Includes var. Includes var. velutinum E. viridulum, duchesnense Synonyms/ notes Taxonomic lancifolium E. heracleoides E. fusiforme arcuatum heracleoides

exum var. var. Eriogonum cernuum Eriogonum corymbosum corymbosum Eriogonum brevicaule viridulum Eriogonum gordonii Eriogonum heracleoides Eriogonum jamesii E. arcuatum

Species name

var.

exumfl Stenogonum var. var. var. avescens fl atum Eriogonum cernuum cernuum corymbosum Eriogonum corymbosum viridulum UtahEriogonum brevicaule Colorado Eriogonum fl Eriogonum gordonii Eriogonum heracleoides Eriogonum infl fusiforme Eriogonum jamesii var. Eriogonum hookeri Eriogonum hookeri rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Polygonaceae Polygonaceae Polygonaceae Family Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 79

longilobum Red Plateau SW of Woodside, Emery Co., UT (Ripley & Barneby 8678 CAS). USFWS: 3C, CO BLM: Sensitive Comments Island Park Rd NE of Vernal, Dinosaur NM, Uintah Co., UT (Holmgren & Reveal 3019 UTC). USFWS: 3C var. var. is near Price, Carbon Co., UT (Jones 5590j POM) 1987 1948 1961 locality is Type Year Year doc. 1948 locality is Type 1948 1983 1912 1933 1961 locality of Type (DINO) Dever 5103 (DINO) (DINO) Source 3036 (DINO) Dever 5107 (DINO) 303 (DINO) Dever 5011 (DINO) (DINO) 24b (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc Neely 4253 Pres X Com & Wolf Pres X Rare Neely 4019 Park stat. Pres X X Rare O’Kane Pres X XPres Unc & Wolf X Unc Naumann Pres X X Com & Wolf Hist X ? 5330 Weber Hist X ? MacLeod RegEn Wide RegEn Range UT/CO LocEn Wide Wide Wide Wide Wide PerF RegEn/ PerF Wide/ PerF RegEn/ Life form Shrub LocEn/ Shrub Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ golden sulphur wild buckwheat sulfur wild buckwheat Woodside Woodside wild buckwheat Common name Dinosaur wild buckwheat slender wild buckwheat slender wild buckwheat cushion wild buckwheat smooth wild buckwheat Shockley’s Shockley’s wild buckwheat

var. var.

by some aureum , vars. includes (2005) Flora of North Included in var. Included in var. umbellatum authors Some authors include var. Vars. not recognized Vars. in America Synonyms/ notes Taxonomic phoeniceum E. ovalifolium purpureum not recognized in et al. (2003) Welsh Includes var. shockleyi E. microthecum foliosum,

var. var.

var. var. var. var. orum Eriogonum umbellatum aureum Eriogonum umbellatum umbellatum Eriogonum microthecum Eriogonum tumulosum Eriogonum saurinum laxiﬂ Eriogonum microthecum Eriogonum ovalifolium ovalifolium Stenogonum salsuginosum longilobum Eriogonum shockleyi simpsonii

var. Species name var. var. var. var. orum Eriogonum umbellatum aureum Eriogonum umbellatum umbellatum Eriogonum microthecum laxifl Eriogonum tumulosum UtahEriogonum lonchophyllum saurinum Colorado Eriogonum microthecum simpsonii Eriogonum ovalifolium Eriogonum salsuginosum Eriogonum shockleyi rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Family Polygonaceae Polygonaceae Polygonaceae Polygonaceae biogeographic notes, cont. Appendix A. Confi

80 Annotated Checklist of Vascular Flora: Dinosaur National Monument Eurasia. Eurasia Comments Eurasia 1984 1933 1948 1961 1988 1948 Native to 1988 1955 Native to Year Year doc. 1948 NA 1961 Native to & Wathen & Wathen 121 (DINO) (DINO) et al. 418 (DINO) et al. 428 (DINO) 281 (DINO) 3112 (DINO) 222 (DINO) (DINO) Source (DINO) (2002) et al. 403 (DINO) c name with taxonomic and Pop. size UT CO Pres X Unc Friedlander Hist X ? 5328 Weber Hist X ? Holmgren Hist X X ? Holmgren Pres X Rare Naumann Pres X X Unc O’Kane Pres X Unc Naumann Pres X X Unc Neely 4219 Hist X ? 5288 Wolf Park stat. Rep X ? Naumann Pres X Unc Holmgren Wide Wide Wide Wide Sparse Intro Wide Intro Wide Range UT/CO Wide Intro AnnF Wide/ Fern Wide/ AnnF Intro/ AnnF Wide/ PerF Wide/ Life form AnnF Wide/ Douglas’ knotweed canaigre dock PerF Wide/ golden dock AnnF Wide/ willow dock PerF Wide/ southern maidenhair fern yard knotweed Sawatch knotweed Kellogg’s Kellogg’s knotweed curly dock PerF Intro/ water smartweed Common name willow-weed AnnF Intro/

var.

var. var. Var. ssp. var. mexicanus, emersum, P. sawatchense P. R. fueginus R. mexicanus, R. salicifolius var. R. utahensis, triangulivalvis mexicanus, Rumex triangulivalvis montigenitus aviculare sawatchense. not recognized and by Weber Wittmann (2001). P. kelloggii P. kelloggii amphibium var. Polygonum coccineum, Persicaria coccinea Synonyms/ notes Taxonomic aviculare P.

ssp. fueginus Polygonum douglasii Rumex hymenosepalus Rumex maritimus var. Rumex triangulivalvis Adiantum capillus- veneris kelloggii Polygonum polygaloides Persicaria amphibia Polygonum Polygonum arenastrum Persicaria lapathifolia Species name

var. var. triangulivalvis fueginus douglasii Polygonum douglasii johnstonii Polygonum douglasii Rumex hymenosepalus Rumex maritimus var. Rumex salicifolius ssp. Adiantum capillus- veneris Polygonum kelloggii Rumex crispus Rumex crispus UtahPolygonum amphibium Colorado Polygonum aviculare Polygonum lapathifolium rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polypodiaceae (Adiantaceae) Polygonaceae Polygonaceae Polygonaceae Polygonaceae Family Polygonaceae biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 81

C. utahensis Comments of is Negro Bill Canyon, Grand Co., UT (Windham & Windham 90- 282) North America Year Year doc. 1987 1987 1948 1948 1948 1987 1987 2002 1961 1983 locality Type 1995 1961 Native to E Source (DINO) 3119 (DINO) Dever 5206 (DINO) 2685 (DINO) 4424 (DINO) 2736 (DINO) (DINO) (DINO) 13979 (DINO) 2737 (DINO) 414 (DINO) et al. 424 (DINO) c name with taxonomic and Pop. size UT CO Park stat. Pres X Rare Neely 4455 Pres X Unc O’Kane Pres X Rare & Wolf Pres X X Unc O’Kane Pres X Unc Rodeck Pres X X Unc O’Kane Pres X Rare Neely 4340 Pres X Unc s.n. Tilley Hist X ? Tanner Pres X Unc O’Kane Pres X Unc Naumann Hist X ? Holmgren Range UT/CO Sparse Wide Periph Periph Wide Wide Periph Wide Wide Wide Periph Intro Life form Fern Sparse/ Fern Wide/ Fern Wide/ Fern Periph/ Fern Periph/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Fern Wide/ AnnF Intro/ ower owered Common name forked spleenwort slender lip- fern Brewer’s cliff- Brewer’s brake western cliffbrake lanceleaf springbeauty Oregon bitter- root small- ﬂ fame-ﬂ northern pondweed brittle bladder fern Oregon woodsia miner’s lettuceminer’s AnnF Wide/ common purslane

C. & ssp. minor var. C. ssp. & Some scopulina C. orus & orus, P. P. orus, rediviva Synonyms/ notes Taxonomic Includes utahensis, C. reevesiana, P. occidentalis P. Includes ssps. oregana lanceolata. authors include rosea Includes vars. & intermontana parviﬂ confertiﬂ tenuis perfoliata ssp. orum Phemeranthus

var. cathcartiana Asplenium septentrionale simplex Pellaea glabella ssp. Claytonia lanceolata C. lanceolata Claytonia rubra C. perfoliata Talinum parviﬂ Talinum tenuifolius Potamogeton alpinus

var. Species name orum perfoliata UtahAsplenium septentrionale Colorado Cheilanthes feei Cheilanthes feei Cystopteris fragilis Cystopteris fragilis Pellaea breweri Pellaea breweri Pellaea glabella occidentalis Woodsia oregana Woodsia oregana Woodsia Claytonia lanceolata Lewisia rediviva Lewisia rediviva Montia perfoliata var. Talinum Talinum parvifl Potamogeton alpinus Portulaca oleracea Portulaca oleracea rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Polypodiaceae (Aspleniaceae) Family Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Aspleniaceae) Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Aspleniaceae, Woodsiaceae) Portulacaceae Portulacaceae Portulacaceae Portulacaceae Potamogetonaceae Portulacaceae biogeographic notes, cont. Appendix A. Confi

82 Annotated Checklist of Vascular Flora: Dinosaur National Monument ). (var. (var. Comments (DINO) is atypically robust and may be subvar. huberi zionense Year Year doc. 1961 1961 NA 2002 1948 1961 1948 1948 1933 O’Kane #3171 Source et al. 444 (DINO) et al. 448 (DINO) (2002) and et al. Weber (1982) (DINO) 3010 (DINO) & Jensen 13969 (DINO) Dever 5184 (DINO) (DINO) 4441 (DINO) c name with taxonomic and Pop. size UT CO Park stat. Hist X ? Holmgren Hist X ? Holmgren Rep X ? Naumann Pres X Unc s.n. Tilley Pres X Unc O’Kane Hist X ? Holmgren Hist X ? & Wolf Pres X Unc Neely 4004 Pres X X Unc Rodeck Range UT/CO Wide Wide Wide Wide Wide Wide Wide Wide Wide Life form PerF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ ower Common name longleaf pondweed fennel-leaf pondweed dwarf pondweed slender-stem slender-stem rock-jasmine western rock- jasmine pygmy-ﬂ rock-jasmine sea-milkwort PerF Wide/ sidebells pyrola dark-throat shooting-star

. & var

var. orum. huberi Synonyms/ notes Taxonomic Stuckenia pectinata, Coleogeton pectinatus P. pusillus P. pusillus Includes ssp. puberulenta, subumbellata, subilifera D. pauciﬂ Includes “subvars.” from UT Flora: coriaceum, atwoodii, & liformis Potamogeton nodosus Potamogeton pectinatus Potamogeton pusillus Androsace ﬁ Androsace occidentalis Androsace septentrionalis Dodecatheon pulchellum

Species name var. liformis UtahPotamogeton nodosus Colorado Potamogeton pectinatus Potamogeton pusillus Androsace fi Androsace occidentalis Androsace septentrionalis Glaux maritima Glaux maritima Pyrola secunda Orthilia secunda Dodecatheon pulchellum pulchellum rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Potamogetonaceae Family Potamogetonaceae Potamogetonaceae Primulaceae Primulaceae Primulaceae Primulaceae (Myrsinaceae) Pyrolaceae (Ericaceae) Primulaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 83 is Bluff, San is Bluff, Juan Co., UT s.n. (Wetherill CAS) Comments is Hole in the Canyon, Wall Uinta Mtns., 10 miles N of Uintah Vernal, Co., UT (Huber & Plunkett 91 USFS BRY). Region 4: Sensitive 1948 1933 NA 1948 locality Type Year Year doc. 1991 locality Type 1933 1987 NA Dever 5072 (DINO) (DINO) (2002) 4428 (DINO) Source (DINO) (DINO) 2741 (DINO) (1962) c name with taxonomic and Pop. size UT CO Pres X X Unc & Wolf Pres X X Unc 5387 Weber Rep X ? Naumann Pres X X Unc Rodeck Park stat. Pres X Unc Reidel 42 Pres X X Unc 5272 Wolf Pres X Unc O’Kane Rep X NA Holmgren Wide Wide Wide RegEn Range UT/CO NA Wide Wide Wide PerF Wide/ PerF Wide/ PerF Wide/ PerF RegEn/ Life form PerF LocEn/ PerF Wide/ PerF Wide/ PerF Sparse/ white virgin’s- bower Nuttall’s Nuttall’s larkspur thread water- buttercup alcove columbine Common name Graham’s columbine marsh buttercup sagebrush buttercup lesser yellow water buttercup

var. ssp. R. eradicatus, trichophyllus, D. nelsonii R. circinatus, longirostris, R. trichophyllus var. R. trichophyllus var. Batrachium longirostre, B. circinatum subrigidum, B. trichophyllum Synonyms/ notes Taxonomic R. cymbalaria cymbalaria Includes oreogenes

var. ssp. hookeri Clematis ligusticifolia Delphinium nuttallianum Batrachium circinatum Halerpestes cymbalaria Aquilegia micrantha saximontana Ranunculus glaberrimus ellipticus Ranunculus gmelinii var.

Species name

var. var. Clematis ligusticifolia Delphinium nuttallianum Ranunculus aquatilis diffusus Ranunculus cymbalaria Aquilegia micrantha UtahAquilegia grahamii Colorado Ranunculus glaberrimus ellipticus Ranunculus gmelinii rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Family Ranunculaceae Ranunculaceae biogeographic notes, cont. Appendix A. Conﬁ

84 Annotated Checklist of Vascular Flora: Dinosaur National Monument

at DINO herbarium are A. utahensis determined on DINO plants. locality Type is ca 10 km E of Oak City, Millard Co., UT (Holmgren & Holmgren 11298 NY) Comments Eurasia American Fork Canyon, Utah Co., UT (Watson 314 US) 1912 1948 1935 All specimens 1948 not Variety Year Year doc. 1996 NA 1987 1991 Native to 1912 1948 locality is Type 2726 (DINO) Dever 5222 (DINO) (1937) and Northern Colorado Plateau Network (in ed.). Dever 5157 (DINO) Source 417 (DINO) (1962) (DINO) (DINO) Dever 5062 (DINO) Neely 2671 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Com O’Kane Pres X X Unc & Wolf Rep X X Unc Graham Pres X X Com & Wolf Park stat. Pres X Unc Naumann Rep X ? Holmgren Pres X Unc Neely 4379 Pres X Unc Reidel 169 Pres X X Com & Wolf Pres X X Unc O’Kane & Wide Wide Wide Wide Range UT/CO RegEn Wide Wide Intro Wide Wide Shrub Wide/ Shrub Wide/ Shrub Wide/ Life form PerF RegEn/ PerF Wide/ AnnF Wide/ Shrub Wide/ Shrub Wide/ alder-leaf alder-leaf mountain mahogany deer-brush Shrub Wide/ Saskatoon serviceberry curl-leaf mountain mahogany Common name Jupiter buttercup Macoun’s Macoun’s buttercup blister buttercup bur buttercup AnnF Intro/ Utah serviceberry dwarf mountain mahogany

var. var. var. velutinus C. ledifolius intercedens Synonyms/ notes Taxonomic C. testiculata C. ledifolius intricatus Cercocarpus montanus Ceanothus velutinus C. velutinus Amelanchier alnifolia Amelanchier utahensis Cercocarpus ledifolius Ranunculus macounii Hecatonia scelerata Ceratocephala orthoceras Cercocarpus intricatus Species name

var. var. dus Cercocarpus montanus Ceanothus velutinus Amelanchier alnifolia Amelanchier utahensis intermontanus Cercocarpus ledifolius UtahRanunculus jovis Ranunculus jovis Colorado Ranunculus macounii Ranunculus sceleratus Ranunculus testiculatus Cercocarpus intricatus multifi rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Rosaceae Rhamnaceae Rosaceae Rosaceae Rosaceae Ranunculaceae Family Ranunculaceae Ranunculaceae Ranunculaceae Rosaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 85 Comments is Mt, Barrette near Marysvale, Piute Co., UT (Rydberg & Carlton 7261 NY) 1955 NA Year Year doc. NA 1948 1956 1948 1987 1948 1948 1945 1986 locality Type 1987 1956 1946 (BRY) (1962) Source (2002) & Jensen 13957 (DINO) 3100 (DINO) 4440 (DINO) (DINO) Dever 5230 (DINO) Dever 5214 (DINO) 3110 (DINO) (DINO) (DINO) (BRY) 70b (DINO) c name with taxonomic and Pop. size UT CO Hist X ? 425a Welsh Rep X ? Holmgren Park stat. Rep X ? Naumann Pres X X Unc Holmgren Pres X X Unc O’Kane Pres X X Unc Rodeck Pres X Unc Neely 4107 Pres X X Unc & Wolf Hist X ? & Wolf Pres X Unc O’Kane Pres X Unc Neely 4320 Pres X Unc Neely 4367 Hist X ? 474 Welsh Pres X Unc MacLeod Sparse Wide Range UT/CO Wide Wide Wide Wide Wide Wide Wide Wide RegEn Sparse Wide Wide PerF Wide/ PerF Wide/ Life form Shrub Wide/ Shrub Wide/ PerF Wide/ Shrub Wide/ AnnF Wide/ PerF RegEn/ PerF Wide/ PerF Wide/ PerF Wide/ glandular cinquefoil comb-leaf cinquefoil Common name river hawthorn bush oceanspray Gordon’s Gordon’s ivesia rock spiraea PerF Wide/ dwarf ninebark silverweed PerF Wide/ tall cinquefoil PerF Wide/ biennial cinquefoil modest cinquefoil glabrate cinquefoil beautiful cinquefoil sheep cinquefoil

in arguta P. gracilis P. by some var. elmeri Included in var. authors P. pectinisecta P. P. gracilis P. Synonyms/ notes Taxonomic Spiraea caespitosa Includes ssp. Combined with rubricaulis P. Holmgren et al. (1997) nuttallii brunnescens glabrata Potentilla gracilis var. Potentilla gracilis var. Crataegus rivularis Holodiscus discolor H. microphyllus Petrophyton caespitosum Physocarpus alternans Drymocallis arguta Potentilla concinna concinna. P. Potentilla pulcherrima Potentilla ovina

var. Species name brunnescens elmeri fastigiata rivularis convallaria modesta pulcherrima Potentilla gracilis var. Potentilla gracilis var. Potentilla gracilis var. UtahCrataegus douglasii var. Colorado Holodiscus dumosus Ivesia gordonii Ivesia gordonii Petrophyton caespitosum Physocarpus alternans Potentilla anserina Argentina anserina Potentilla arguta var. Potentilla biennis Potentilla biennis Potentilla concinna var. Potentilla gracilis var. Potentilla ovina ovina rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Rosaceae Rosaceae Rosaceae Rosaceae Family Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae biogeographic notes, cont. Appendix A. Conﬁ

86 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Eurasia Wasatch Mtns., Wasatch Salt Lake Co., 480 UT (Watson US) 1935 1912 1988 Year Year doc. 1935 1948 1948 1987 Native to 1987 1983 1948 locality is Type 1988 1933 1948 1945 1948 3025 (DINO) (DINO) 188 (DINO) Source Reidel 2804 (DINO) Dever 5132 (DINO) Dever 5219 (DINO) (DINO) (DINO) & Wathen & Wathen 117 (DINO) Dever 5217 (DINO) 179 (DINO) 4432 (DINO) 2752 (DINO) May 1453 (RM) Dever 5186 (DINO) c name with taxonomic and Pop. size UT CO Pres X X Com O’Kane Pres X X Unc Neely 4225 Pres X Unc Naumann Park stat. Pres X X Unc O’Kane & Pres X X Com & Wolf Pres X Unc & Wolf Pres X Unc Neely 4333 Pres X Unc Neely 4357 Pres X Unc Friedlander Pres X X Unc & Wolf Pres X Unc Naumann Pres X X Unc Rodeck Pres X X Com O’Kane Pres X Unc Fowler & Pres X X Com & Wolf Wide RegEn Periph Range UT/CO Wide Wide Wide Intro Wide Wide Wide Wide Wide Wide Wide Wide PerF RegEn/ PerF Wide/ Life form Shrub Wide/ Shrub Wide/ Shrub Wide/ AnnF Wide/ PerF Wide/ Tree Wide/ Tree Wide/ Tree Periph/ Woods’ roseWoods’ Shrub Wide/ Colorado bedstraw shrubby bedstraw Common name western chokecherry antelope bitterbrush Nootka rose Shrub Wide/ tea rose Shrub Intro/ common red raspberry cleavers AnnF Wide/ twin-leaf bedstraw sweet-scented bedstraw narrow-leaved cottonwood Fremont cottonwood Rio Grande cottonwood quaking aspen Tree Wide/ R.

ultra-

var. var. includes var. montana, manca Synonyms/ notes Taxonomic R. idaeus aculeatissimus fremontii

orum ssp. orum melanocarpa wislizenii Galium coloradoense Galium multiﬂ Padus virginiana ssp. Rosa nutkana Rubus idaeus melanolasius Populus angustifolia Populus fremontii deltoides P. Populus deltoides ssp. Populus tremuloides Galium spurium

var. Species name var. orum orum var. orumtrifl Galium orum coloradoense multifl melanocarpa fremontii wislizeni Rosa woodsii Rosa woodsii R. woodsii Rubus idaeus Galium multifl var. Galium multifl var. Galium trifl UtahPrunus virginiana var. Colorado Purshia tridentata Purshia tridentata Rosa nutkana hispida Rosa odorata Rosa odorata strigosus Galium aparine Galium bifolium Galium bifolium Populus angustifolia Populus fremontii var. Populus fremontii var. Populus tremuloides echinospermum rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Rosaceae Rosaceae Rubiaceae Rubiaceae Rubiaceae Rosaceae Family Rosaceae Rosaceae Rosaceae Rubiaceae Rubiaceae Salicaceae Salicaceae Salicaceae Salicaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 87

utahensis Comments is Jesse Ewing Canyon, Daggett Co., UT (Goodrich 24441 BRY) var. var. is City Creek Canyon, Salt Lake Co., UT (Jones 1458 NY) Year Year doc. 1987 1987 1948 locality Type 1933 1933 1933 1948 1987 1987 1955 1933 1950 1948 locality of Type Source (DINO) 2986 (DINO) 4433 (DINO) 167 (DINO) (DINO) (1937) & NCPN (in ed.) Dever 5187 (DINO) Riedel 2799 (DINO) (DINO) 2826 (DINO) Dever 5026 (DINO) (DINO) Dever 5136 (DINO) c name with taxonomic and Pop. size UT CO Park stat. Pres X X Unc Neely 4203 Pres X Unc O’Kane Pres X X Com Rodeck Pres X X Unc Naumann Pres X X Com 5306 Weber Rep X X Unc Graham Pres X Unc & Wolf Pres X X Unc O’Kane & Pres X Unc Neely 3949 Pres X X Unc O’Kane Pres X X Unc & Wolf Pres X X Unc 5338 Weber Pres X X Unc & Wolf Range UT/CO Wide Wide Wide Wide Wide Wide Wide Wide Sparse Wide Wide Periph Wide Life form Tree Wide/ PerF Wide/ Tree Wide/ Shrub Wide/ PerF Wide/ PerF Wide/ Shrub Wide/ Shrub Wide/ PerF Wide/ Shrub Wide/ PerF Wide/ ower ax Common name peach-leaf willow woodlandstar lance-leaf cottonwood yellow willow Shrub Wide/ coyote willow Shrub Wide/ Scouler’s Scouler’s willow fringecup woodlandstar smallﬂ woodlandstar slender whiplash willow mountain willow bastard toadﬂ Utah fendlerella littleleaf alumroot & P.

x Hybrid exigua

var. var. var. ora P. fremontii P. of Synonyms/ notes Taxonomic angustifolia watsonii S. lasiandra L. bulbifera, glabra L. parviﬂ L. tenella caudata C. pallida Includes vars. parvifolia, major, utahensis

ssp. acuminata acuminata. P. x orum Populus Salix lutea S. rigida Salix exigua S. exigua Salix lucida caudata Lithophragma glabrum Lithophragma parviﬂ Lithophragma tenellum Comandra umbellata Fendlerella utahensis

Species name

var. ssp. var. var. x orum watsonii UtahPopulus intercurrens Colorado Salix amygdaloides Salix amygdaloides Salix eriocephala var. stenophylla Salix exigua caudata Salix lucida Salix scouleriana Salix scouleriana Comandra umbellata Lithophragma glabrum ramulosum Lithophragma parvifl Lithophragma tenellum Salix monticola Salix monticola pallida Fendlerella utahensis Heuchera parvifolia Heuchera parvifolia rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Salicaceae Salicaceae Family Salicaceae Salicaceae Salicaceae Salicaceae Santalaceae Saxifragaceae Saxifragaceae Saxifragaceae Salicaceae Saxifragaceae (Hydrangeaceae) Saxifragaceae biogeographic notes, cont. Appendix A. Confi

88 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments noxious weed (List B). Native to Eurasia is Leroy, Uinta is Leroy, Co., WY (Nelson, A. 4577 RM). noxious weed (List B). Native to Europe Year Year doc. 1988 1948 1933 1949 1948 1955 2003 CO state 1948 1912 locality Type 1956 1983 1961 NA CO state Source 256 (DINO) Wolf 5281 Wolf (DINO) 114 (DINO) 5359 (DINO) Dever 5177 (DINO) Neely 2664 (DINO) DINO V-5 DINO V-5 (DINO) Dever 5176 (DINO) 2725 (DINO) (DINO) & Wathen & Wathen 120 (DINO) et al. 435 (DINO) (2002) c name with taxonomic and Pop. size UT CO Park stat. Pres X Unc Naumann Pres X X Unc Dever & Pres X X Unc Naumann Pres X X Unc Bradley Pres X X Unc & Wolf Pres X X Unc O’Kane & Pres X Unc Ballard Pres X X Com & Wolf Pres X X Com O’Kane Pres X X Unc Neely 4100 Pres X Unc Friedlander Pres X Unc Holmgren Rep X ? Naumann Range UT/CO Wide Wide Wide Wide Wide RegEn Intro Wide Wide Wide Wide Wide Intro Life form PerF Wide/ Shrub Wide/ Shrub Wide/ AnnF Wide/ PerF Wide/ PerF RegEn/ PerF Intro/ PerF Wide/ AnnF Wide/ AnnF Wide/ PerF Intro/ ax Common name marsh grass- of-Parnassus littleleaf mock-orange golden currant annual paintbrush Wyoming paintbrush Eastwood’s Eastwood’s paintbrush butter-and- eggs wax currant Shrub Wide/ desert paintbrush blue-eyed Mary bushy bird’s- beak mudwort PerF Wide/ Dalmatian toadﬂ

var. var.

ssp.

minor tenuis var. var. ssp. martinii occidentalis, Includes var. Synonyms/ notes Taxonomic P. microphyllus P. var. occidentalis P. occidentalis aureum pedicellare dubia, C. applegatei ssp. L. genistifolia dalmatica ora dalmatica Parnassia palustris Philadelphus microphyllus Ribes cereum R. cereum Castilleja scabrida C. scabrida Cordylanthus ramosus ssp.

oraparvifl Collinsia Species name var. montanensis scabrida UtahParnassia palustris var. Colorado Philadelphus microphyllus Ribes aureum Ribes aureum R. aureum inebrians Ribes cereum Castilleja exilis Castilleja minor C. minor Castilleja linariifolia Castilleja linariifolia C. linariaefolia Castilleja scabrida var. Collinsia parvifl Linaria vulgaris Linaria vulgaris Castilleja chromosa Castilleja chromosa C. angustifolia Cordylanthus ramosus Limosella aquatica Limosella aquatica Linaria dalmatica Linaria genistifolia rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Saxifragaceae (Parnassiaceae) Family Saxifragaceae (Hydrangeaceae) Saxifragaceae (Grossulariaceae) Saxifragaceae (Grossulariaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 89 Comments is 3 miles N of Maeser, Uintah Co., UT (Holmgren, Holmgren, & Barneby 8748 NY). USFWS: 3C is 6.5 km S of Manila, Daggett Co., UT (Holmgren, Holmgren, & Barneby 8747 NY) Year Year doc. NA 1987 1991 1956 2002 1912 locality Type 1987 1912 1948 1933 locality Type 1960 1987 Source (1982) 3108 (DINO) (DINO) (BRY) (DINO) (DINO) (DINO) 2762 (DINO) Dever 5101 (DINO) Neely 2668 (DINO) 13a (DINO) (DINO) c name with taxonomic and Pop. size UT CO Rep X ? et al. Weber Park stat. Pres X X Unc O’Kane Pres X Unc Reidel 15 Hist X ? 496 Welsh Pres X Unc s.n. Tilley Hist X ? 5366 Weber Pres X Unc Neely 4237 Pres X X Unc O’Kane Pres X Unc & Wolf Pres X X Unc O’Kane & Hist X ? MacLeod Pres X Unc Neely 4310a Wide Range UT/CO Wide Wide Wide Wide LocEn RegEn RegEn Periph RegEn Wide Periph AnnF Wide/ Life form PerF Wide/ AnnF Wide/ AnnF Wide/ AnnF Wide/ PerF LocEn/ PerF RegEn/ PerF RegEn/ PerF Wide/ PerF RegEn/ PerF Wide/ PerF Periph/ ower ower ower ower ower seep monkeyfl Common name yellow monkeyfl reddish monkeyfl Suksdorf’s Suksdorf’s monkeyfl yellow owl- clover Vernal Vernal penstemon mat penstemon Fremont’s Fremont’s penstemon low penstemon thick-leaf penstemon small-fl penstemon mat-root penstemon Synonyms/ notes Taxonomic P. mucronatus P. i

var. var. var. ssp. oribundus Mimulus ﬂ Penstemon angustifolius vernalensis Penstemon caespitosus Penstemon fremontii fremontii Penstemon humilis Penstemon pachyphyllus mucronatus procerus Penstemon confertus Penstemon radicosus isuksdorﬁ Mimulus

Species name var. var.

var. var. humilis oribundus Penstemon caespitosus caespitosus Penstemon procerus UtahMimulus fl Colorado Mimulus guttatus Mimulus guttatus Mimulus rubellus Mimulus rubellus Mimulus suksdorfi Orthocarpus luteus Orthocarpus luteus vernalensis Penstemon angustifolius Penstemon fremontii Penstemon humilis var. Penstemon pachyphyllus mucronatus procerus Penstemon radicosus rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Family Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae biogeographic notes, cont. Appendix A. Confi

90 Annotated Checklist of Vascular Flora: Dinosaur National Monument noxious weed (List C). Native to Eurasia. Comments Blue Mountain, Uintah Co., UT (Neese et al. 11808 BRY) North America, may be adventive in UT. Europe Europe 1961 1987 Native to 1987 1948 1946 2005 CO state Year Year doc. 1956 locality is Type 1956 1987 2002 2002 Native to C 1955 NA Native to et al. 429 (DINO) (DINO) (DINO) Dever 5189 (DINO) (BRY) Andersen (2006) Source (DINO) (BRY) 2948 (DINO) (DINO) (DINO) & Jensen 13956 (DINO) (2002) c name with taxonomic and Pop. size UT CO Pres X Unc Holmgren Pres X Unc Neely 4375 Pres X Unc Neely 4351 Pres X Unc & Wolf Hist X X ? 424 Welsh Rep X Unc Dewey & Pres X X Unc Neely 3982 Park stat. Hist X ? 498 Welsh Pres X Unc O’Kane Pres X Unc s.n. Tilley Pres X Unc s.n. Tilley Pres X X Unc Holmgren Rep X ? Naumann Wide Intro Wide Wide Wide Intro LocEn Range UT/CO Wide Wide Wide Wide Wide Intro PerF Wide/ PerF Intro/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Intro/ PerF LocEn/ Life form AnnF Wide/ Fern Wide/ AnnF Wide/ AnnF Wide/ Fern Wide/ gwort American brooklime water speedwell Rocky Mountain penstemon Watson’s Watson’s penstemon lance-leaf ﬁ woolly mullein Blue Mountain penstemon Common name purslane speedwell dense spike- moss coyote tobacco cut-leaf nightshade blunt-leaf spike-moss black henbane AnnF Intro/ mutica var. scopulorum, S. Synonyms/ notes Taxonomic Includes vars. standleyi, densa, & scopulorum orum cyanomontanus xalapensis Veronica anagallis- Veronica aquatica Penstemon watsonii Scrophularia lanceolata Penstemon scariosus var. Veronica peregrina Veronica var. Nicotiana attenuata Species name

orumtrifl Solanum var. xalapensis Veronica americanaVeronica americana Veronica Veronica anagallis- Veronica aquatica Penstemon strictus Penstemon strictus Penstemon watsonii Scrophularia lanceolata Verbascum thapsusVerbascum thapsus Verbascum UtahPenstemon scariosus cyanomontanus Colorado Veronica peregrina Veronica var. Selaginella densa Selaginella densa Selaginella mutica Selaginella mutica S. mutica Nicotiana attenuata Solanum trifl Hyoscyamus niger Hyoscyamus niger rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Family Scrophulariaceae Selaginellaceae Selaginellaceae Solanaceae Solanaceae Solanaceae biogeographic notes, cont. Appendix A. Confi

Chapter 6: Appendices 91 Comments noxious weed List B). Native to Eurasia is Myton, Duchesne Co., UT (Jones s.n. POM). CO BLM: Sensitive, USFWS: 3C 1948 1933 NA Native to Asia 1949 1912 1950 Year Year doc. 1948 CO state 1912 1987 1912 locality Type 1933 1912 1987 Dever 5054 (DINO) 267 (DINO) (2002) & Jensen 13978 (DINO) (DINO) (DINO) Source Wolf 5090 Wolf (DINO) (DINO) 2801 (DINO) (BRY) 259 (DINO) 3015 (DINO) (DINO) c name with taxonomic and Pop. size UT CO Pres X Com & Wolf Pres X X Com Naumann Rep X ? Naumann Pres X X Unc Holmgren Pres X Unc Neely 3924 Hist X ? 5341 Weber Park stat. Pres X X Com Dever & Hist X ? 5384 Weber Pres X Unc O’Kane Pres X Rare 180 Welsh Pres X X Unc Naumann Pres X X Unc O’Kane Pres X X Unc Neely 4255 Wide Wide Intro Wide Periph Wide Range UT/CO Intro Wide Wide RegEn RegEn Periph Periph PerG Wide/ Tree Wide/ PerF Wide/ PerF Periph/ PerF Wide/ Life form Shrub Intro/ PerF Wide/ PerF Periph/ PerF RegEn/ PerF RegEn/ PerF Wide/ PerF Wide/ ve-stamen broad-leaved cattail netleaf hackberry Siberian elm Tree Intro/ cutleaf water- parsnip plains spring- parsley Fendler’s Fendler’s cymopterus Common name ﬁ tamarisk onion spring- parsley Constance’s Constance’s spring-parsley Duchesne spring-parsley Colorado Plateau spring-parsley aromatic spring-parsley rock spring- parsley

var. ora albiﬂ reticulata, C. douglasii Synonyms/ notes Taxonomic C. purpureus Pteryxia terebinthina var. petraea

var. Berula erecta Cymopterus acaulis C. glomeratus Cymopterus fendleri C. glomeratus Cymopterus bulbosus Cymopterus constancei Cymopterus duchesnensis Cymopterus purpureus Cymopterus terebinthinus calcareus Aletes petraeus C. petraeus, Pteryxia

Species name

var. var. var. var. orus acaulis fendleri Typha latifoliaTypha latifolia Typha Celtis reticulata Celtis reticulata C. laevigata Ulmus pumila Ulmus pumila Cymopterus purpureus Berula erecta incisa Cymopterus acaulis var. Cymopterus acaulis var. Utah chinensisTamarix ramosissima Tamarix pentandra T. Colorado Cymopterus bulbosus Cymopterus constancei Cymopterus duchesnensis purpureus Cymopterus terebinthinus albifl Cymopterus terebinthinus petraeus rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientifi Typhaceae Ulmaceae (Celtidaceae) Ulmaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae Tamaricaceae Family (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) biogeographic notes, cont. Appendix A. Confi

92 Annotated Checklist of Vascular Flora: Dinosaur National Monument is N of Parley’s is N of Parley’s Park, Summit Co., UT (Watson 440 US) Comments Great Salt Lake (Stansbury s.n. NY) is Coalville, Summit Co., UT (Jones s.n. POM) 1987 1987 1989 1934 1948 1933 1996 locality Type Year Year doc. NA 1948 NA 1948 locality is Type 1955 1987 locality Type (DINO) 3190 (DINO) 343 (DINO) et al. 407 (DINO) Dever 5190 (DINO) (DINO) 420 (DINO) Source (1962) Dever 5208 (DINO) and Neese (1986) Dever 5212 (DINO) (DINO) 2781 (DINO) c name with taxonomic and Pop. size UT CO Pres X Rare Neely 4475 Pres X Unc O’Kane Pres X Unc Naumann Hist X X ? Holmgren Pres X Unc & Wolf Pres X Unc Reidel 31 Pres X Rare Naumann Park stat. Rep X ? Holmgren Pres X X Unc & Wolf Rep X ? Goodrich Pres X X Unc & Wolf Pres X X Unc Neely 3979 Pres X Unc O’Kane Wide Wide Periph Wide Wide Wide Wide Range UT/CO Wide Wide Wide Wide Wide RegEn PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF RegEn/ at-seed Chilean sweet- cicely blunt sweet- cicely stinging nettle PerF Wide/ prostrate vervain hook-spur violet northern bog violet Indian-potato PerF Wide/ Common name southern ligusticum giant lomatium big-seed lomatium ﬂ lomatium milfoil lomatium juniper lomatium nis

afﬁ

var. var. ssp. dum U. dioica holosericea Synonyms/ notes Taxonomic multiﬁ L. simplex

ssp. var. Osmorhiza depauperata Urtica gracilis holosericea Orogenia linearifolia Lomatium dissectum L. dissectum Lomatium macrocarpum Lomatium triternatum platycarpum Lomatium grayi Lomatium juniperinum

var.

Species name

var. ssp. var. Osmorhiza chilensis Osmorhiza chilensis O. berteroi Osmorhiza depauperata Urtica dioica occidentalis Verbena bracteataVerbena bracteata Verbena Viola adunca Viola adunca Viola nephrophylla Viola sororia sororia V. Orogenia linearifolia UtahLigusticum porteri Ligusticum porteri Colorado Lomatium dissectum eatonii Lomatium grayi Lomatium macrocarpum Lomatium triternatum platycarpum grayi Lomatium juniperinum rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Urticaceae Verbenaceae Violaceae Violaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Family Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) biogeographic notes, cont. Appendix A. Conﬁ

Chapter 6: Appendices 93 and Comments parasitic on Pinus edulis monophylla P. 1955 Year Year doc. 1948 1988 1946 1955 Typically (BRY) Source 56b (DINO) 180 (DINO) 2040 (CSU) (BRY) c name with taxonomic and Pop. size UT CO Hist X ? 395 Welsh Hist X X ? MacLeod Park stat. Pres X ? Naumann Hist X ? Harrington Hist X ? 243 Welsh Wide Wide Range UT/CO Wide Wide Wide PerF Wide/ PerF Wide/ Life form PerF Wide/ PerF Wide/ horned pondweed upland yellow violet Common name valley violet PerF Wide/ goose-foot violet pinyon dwarf- mistletoe

altior

ssp. var. var. praemorsa, V. praemorsa, V. praemorsa lingulifolia, V. praemorsa V. nuttallii V. Synonyms/ notes Taxonomic

ssp. ssp. major Zannichellia palustris Viola praemorsa Viola vallicola venosa Viola purpurea Arceuthobium divaricatum

var. Species name var. var. Zannichellia palustris UtahViola nuttallii Colorado major Viola nuttallii vallicola Viola purpurea venosa Arceuthobium divaricatum rmed, historical, and reported vascular plant taxa of Dinosaur National Monument, organized by family scientiﬁ Zannichelliaceae Violaceae Violaceae Family Violaceae Viscaceae biogeographic notes, cont. Appendix A. Conﬁ

94 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments ca 5 miles S of Jensen, Uintah Co., UT (Welsh 18461 BRY). tropical America Central America Eurasia Category 3C. Range (UT/CO) Periph Periph Wide NA Wide NA RegEn Periph Life form PerF Wide/ PerF Sparse/ Common name redroot pigweed AnnF Intro/Intro Native to creeping yucca Shrubsea purslane LocEn/ NA locality is Type narrow-leaf water- plantain echinodorus PerFtumble pigweed Wide/ AnnF AnnF Periph / Intro/Intro Native to clasping-leaved dogbane milkweedCircle Cliffs PerFcatchweed RegEn/ AnnF Intro/Intro Native to bigtooth maple Tree Wide/ ally cryptanth cryptanthBarneby’s PerF AnnF LocEn/ NA Wide/NA USFWS: tufted cryptanthcushion cryptanth PerF AnnF RegEn Wide/

var. Synonyms/ notes Taxonomic Var. not recognized by Var. some authors hypericifolium exus retroﬂ circumscissa

var. exus Amaranthus Species name nis rmed) taxa of Dinosaur National Monument. Utah Colorado Yucca harrimaniae Yucca sterilis Sesuvium verrucosum Sesuvium verrucosum Alisma gramineumEchinodorus berteroi Alisma gramineum Amaranthus albus Amaranthus albus Amaranthus retrofl Apocynum sibiricum E. rostratus Apocynum sibiricumAsclepias labriformis A. cannabinum Asperugo procumbensprocumbens Asperugo Cryptantha affi Acer grandidentatum Acer grandidentatum Cryptantha barnebyi Cryptantha caespitosa Oreocarya caespitosa Cryptantha circumscissa Cryptantha Agavaceae Aizoaceae Alismataceae Alismataceae Amaranthaceae Amaranthaceae Apocynaceae Asclepiadaceae Boraginaceae Boraginaceae Family Aceraceae Boraginaceae Boraginaceae Boraginaceae Appendix B. Potential (unconfi

Chapter 6: Appendices 95 2003 Comments is mouth of Sand Wash, Uintah Co., UT (Graham 7924 GH). USFWS: 3C USFWS: Category 3C Eurasia. Native to Europe Vars. not Vars. recognized in Flora of North America Range (UT/CO) RegEn Wide Periph RegEn Wide Wide Wide Periph Wide Life form PerF Wide/ shhook ower ﬁ ower Common name Graham’s cryptanthGraham’s PerF LocEn/ NA locality Type Paradox cryptanth PerF RegEn/ cactus desert cryptanthsilky cryptanth AnnFsalt heliotrope Periph/ PerF RegEn/ PerF Wide/NA European stickseed AnnF Intro/Introcomfrey Native to PerF Intro/Intro Cultivated. secret water-starwort PerF Sparse/ streamside bluebell PerF Wide/ tiquiliaNuttall’s pincushion cactus AnnF PerF Wide/ small-ﬂ Periph/ pretty heliotrope PerF Periph/

var.

var. orus Synonyms/ notes Taxonomic squarrosa, L. echinata vivipara S. parviﬂ

cinale var. intermedius Callitriche hermaphroditica Heliotropium curassavicum oculatum Sclerocactus whipplei var. Coryphantha vivipara Escobaria vivipara

Species name cinaleoffi Symphytum var. var. rmed) taxa of Dinosaur National Monument, cont. roseus vivipara Utah Colorado Cryptantha grahamii Cryptantha paradoxa Oreocarya paradoxa Callitriche hermaphroditica Cryptantha scoparia Cryptantha scoparia Cryptantha sericeaHeliotropium Oreocarya sericea curassavicum obovatum Heliotropium curassavicum oculatum Sclerocactus whipplei var. Lappula squarrosa Mertensia ciliata Lappula squarrosa Symphytum offi L. squarrosa Mertensia ciliata nuttalliiTiquilia Coryphantha vivipara var. nuttallii Tiquilia Coldenia nuttallii Family Boraginaceae Boraginaceae Callitrichaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Cactaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae (Ehretiaceae) Cactaceae Appendix B. Potential (unconfi

96 Annotated Checklist of Vascular Flora: Dinosaur National Monument C. in Weber in Weber Comments Cited as verna et al. (1982). Eurasia Eurasia Eurasia Range (UT/CO) Wide Wide Wide NA Wide Wide Wide Wide Wide Wide Wide Life form y PerF Wide/ Common name vernal water-starwort PerF Wide/ black twinberrynodding chickweed Shrub AnnFtall baby’s-breath Wide/ Sparse/ PerF Intro/Intro Native to clammy-weed AnnF Wide/ Drummond’s catchﬂ Drummond’s long-leaved starwort PerFthickleaf orach Sparse/ two-seed orach AnnFgarden orach Wide/ AnnF Intro/Intro Native to AnnF Intro/Intro Native to Menzies’ campion PerF Wide/ alpine pearlwort PerF Wide/ creeping nailwort PerF Wide/ thinleaf orach AnnF Wide/NA Powell’s orachPowell’s AnnF Wide/

& & ssp.

var. var. var. includes vars. drummondii menziesii Synonyms/ notes Taxonomic Silene drummondii ssp. striata, striata, kruckebergii, drummondi subspicata, A. subspicata menziesii, A. menziesii var. triangularis ora Polanisia dodecandra trachysperma P. drummondii Atriplex powellii

orasessilifl Paronychia var. Species name rmed) taxa of Dinosaur National Monument, cont. trachysperma Utah Colorado Callitriche palustris Callitriche palustris C. verna Polanisia dodecandra var. Lonicera involucrata Distegia involucrata Cerastium nutans Gypsophila paniculata Gypsophila paniculata drummondiiLychnis Gastrolychnis Paronychia sessilifl Stellaria longifoliaAtriplex dioica Stellaria longifolia Atriplex heterosperma Atriplex patulaheterosperma Atriplex A. micrantha Atriplex hortensis A. patula Atriplex powellii Atriplex hortensis powellii Sagina saginoidesSilene menziesii Sagina saginoides Anotites menziesii S. menziesii Atriplex prostrata A. patula Family Callitrichaceae Capparaceae (Cleomaceae) Caprifoliaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae Caryophyllaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae Chenopodiaceae Appendix B. Potential (unconfi

Chapter 6: Appendices 97 Comments Eurasia Eurasia Eurasia Range (UT/CO) Wide Wide Wide Wide Wide Wide Sparse Wide RegEn Periph Periph Wide Periph Wide Life form PerF Intro/NA Native to Common name tumbling orach AnnF Intro/Intro Native to sandstone pussytoes PerF Wide/ prostrate kochia, forage kochia Chamisso’s arnicaChamisso’s PerF Wide/ mountain pussytoes PerF Wide/ desert goosefoot AnnF Sparse/ red goosefoot AnnF Wide/ winged pigweed AnnF Wide/ red saltwortbarbwire Russian thistle AnnF Intro/Intro AnnF Native to Wide/ broom seepweedArizona siltbush AnnF Wide/ Shrub RegEn/ short agoseris PerF Wide/ annual agoseris AnnF Periph/ western ragweed PerF Wide/ rush pussytoes PerF Wide/ alpine pussytoes PerF Wide/

var. var. var. ssp. A. neglecta Synonyms/ notes Taxonomic Includes previous records of from UT A. chamissonis andina C. pratericola S. rubra arizonica heterophylla A. coronopifolia luzuloides

var. rubra foliosa coronopifolia umbrinella Chenopodium desiccatum Arnica chamissonis ssp. Chenopodium rubrum atriplicifolium ssp. Atriplex brandegei Z. brandegeei Agoseris glauca dasycephala var. Antennaria media

var. var. var. Species name var. rmed) taxa of Dinosaur National Monument, cont. rubrum Utah Colorado Antennaria umbrinella Antennaria Atriplex rosea Atriplex rosea Antennaria marginata Antennaria marginata Bassia prostrata Kochia prostrata Chenopodium desiccatum Arnica chamissonis foliosa Chenopodium rubrum var. Cycloloma atriplicifolia Cycloloma Salicornia europaea Salicornia europaea Salsola paulsenii Salsola paulsenii Suaeda calceoliformis Suaeda calceoliformisZuckia brandegei S. depressa arizonica Agoseris glauca dasycephala Agoseris heterophylla Agoseris heterophylla A. heterophylla Ambrosia psilostachya Ambrosia psilostachya Antennaria alpina media Antennaria luzuloides Antennaria luzuloides A. luzuloides Compositae (Asteraceae) Family Chenopodiaceae Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconﬁ

98 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments is 10 miles W of Vernal, Uintah Co., UT (Goodrich 23215 BRY) Eurasia. CO state noxious weed (List B) Periph Range (UT/CO) Wide Wide Wide Periph Wide Wide Wide Wide Wide Wide Periph Life form Common name graystem rabbitbrush Shrub Wide/ Carruth’s wormwoodCarruth’s PerFDuchesne sagebrush Wide/ Shrub LocEn/ NA locality Type hairy arnica PerF Wide/ whitestem rabbitbrush Shrub Wide/ pygmy sagebrush Shrub Wide/ leafy aster AnnF Wide/ Parry’s asterParry’s PerF Wide/ cutleafnodding bur-marigold PerF Wide/ PerF Wide/ alkali aster PerF Wide/ Eastwood-plant PerF Wide/ ox-eye daisy PerF Intro/Intro Native to Greene’s rabbitbrushGreene’s Shrub Wide/

parva speciosa var. var. var. A. ovata hololeuca Synonyms/ notes Taxonomic Includes C. nauseosus albicaulis, Ericameria nauseosa frondosum C. scaposa Ericameria nauseosa var.

parryi var. var. var. orus Chrysothamnus nauseosus speciosus pygmaeum Symphyotrichum foliaceum pauciﬂ Chamaechaenactis scaposa Leucanthemum vulgare greenei Chrysothamnus nauseosus hololeucus

Species name var. var.

speciosus orus Almutaster var. var. rmed) taxa of Dinosaur National Monument, cont. Utah Colorado Chrysothamnus nauseosus Artemisia carruthiiArtemisia nova Artemisia carruthii duchesnicola Arnica mollis Arnica mollis Artemisia pygmaea Seriphidium Aster foliaceus parryi Aster frondosusAster pauciﬂ Brachyactis frondosa Symphyotrichum Bahia dissectaBidens cernua Bahia dissectaChamaechaenactis scaposa Bidens cernua Amauriopsis dissecta Chrysanthemum leucanthemum Chrysothamnus greenei Chrysothamnus Chrysothamnus nauseosus gnaphalodes Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconﬁ

Chapter 6: Appendices 99 Comments Range (UT/CO) Wide Wide Wide RegEn Wide Wide Wide Wide RegEn Wide RegEn RegEn RegEn Wide RegEn Life form PerF Wide/ ower PerF RegEn/ eabane PerF Wide/ wandering ﬂ Common name trailing daisybeautiful daisylongleaf daisy PerF PerF Wide/ Wide/ PerF Wide/ dune rabbitbrush Shrub RegEn/ aspen daisybear daisybasin blanketﬂ PerF Wide/ PerF Wide/ San Rafael daisy PerF RegEn/ hispidulous hawksbeard Parry’s rabbitbrushParry’s thistleBarneby’s Shrub Wide/ Wyoming thistle PerF RegEn/ PerF RegEn/ Tracy’s thistleTracy’s PerF RegEn/ sticky cudweed AnnF Sparse/ crustweed AnnF Wide/NA

var.

var. turbinata nis E. peregrinus scaposus Synonyms/ notes Taxonomic Ericameria nauseosa var. Ericameria parryi afﬁ G. viscosum, Pseudognaphalium macounii

var. agellaris nis hispidulosa callianthemus afﬁ formosissimus lonchophylla Erigeron peregrinus Chrysothamnus nauseosus turbinatus ssp. Erigeron speciosus Erigeron ursinus Erigeron compactus E. consimilis Psilochenia runcinata ssp. Chrysothamnus parryi var. Cirsium tracyi viscosum

var. var.

var. var. Species name var. var.

var. agellarisfl Erigeron rmed) taxa of Dinosaur National Monument, cont. nis affi Utah Colorado Erigeron formosissimus Erigeron Erigeron lonchophyllus Trimorpha Erigeron peregrinus callianthemus Chrysothamnus parryi var. Chrysothamnus nauseosus turbinatus Erigeron speciosus macranthus Erigeron ursinus ursinus Gaillardia spathulata Gaillardia spathulata Erigeron compactus consimilis Erigeron fl Cirsium barnebyiCirsium pulcherrimum Cirsium barnebyi Cirsium pulcherrimum Cirsium undulatum Crepis runcinata hispidulosa tracyi Glyptopleura marginata Gnaphalium macounii Pseudognaphalium Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix B. Potential (unconfi Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

100 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Europe Europe Europe CO BLM: Sensitive Range (UT/CO) Wide Wide Periph Wide Wide NA Periph Wide Wide Periph RegEn Life form ower PerF Wide/ venerve sunﬂ venerve Common name ﬁ grass goldenweed PerF goldenweedClement’s PerF LocEn/ NA Wide/ hedge bindweed PerF Sparse/ common tansylittleleaf horsebrush PerF Shrub Wide/NA Intro/Intro Native to showy goldeneye PerF Wide/ marsh-elder AnnF Wide/ desert bahiaprenanthella PerFcommon groundsel RegEn/ AnnF AnnF Intro/Intro Wide/ Native to spiny-leaf sow-thistle AnnF Intro/Intro Native to mountain tarweeed AnnFcopperweed Wide/ low feverfew Shrub Wide/ PerF RegEn/ false boneset PerF Periph/

var.

var. var. var. gramineus corymbulosa, B. Synonyms/ notes Taxonomic Stenotus armerioides var. clementis Convolvulus sepium desertorum ligulatum K. eupatorioides var. eupatoriodies corymbulosa Bahia nudicaulis

var.

ora ssp. nevadensis Helianthella quinquenervis Calystegia sepium angulata Heliomeris multiﬂ var. xanthifolia Brickellia rosmarinifolia chlorolepis

var. var. Species name

ora var. var. rmed) taxa of Dinosaur National Monument, cont. chlorolepis Utah Colorado Helianthella quinquenervis Haplopappus armerioides gramineus Haplopappus clementis Pyrrocoma clementis clementis P. Tanacetum vulgareTanacetum glabrata Tetradymia vulgare Tanacetum Viguiera multiﬂ Calystegia sepium angulata nevadensis Iva xanthifoliaKuhnia eupatorioides var. Cyclachaena Prenanthella exigua Prenanthella exiguaSenecio vulgaris exigua Lygodesmia Senecio vulgaris Sonchus asper Sonchus asper Madia glomerataOxytenia acerosa Madia glomerata Parthenium ligulatum Oxytenia acerosa Bolophyta ligulata alpinum P. Iva acerosa Platyschkuhria integrifolia desertorum Compositae (Asteraceae) Appendix B. Potential (unconﬁ Family Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Convolvulaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 101 Comments Mediterranean New species, cited for UT and CO in Holmgren et al. (2005) Range (UT/CO) Periph NA Wide Wide Wide Wide LocEn NA Periph NA Wide Life form PerF LocEn/ NA ower dodder ower AnnF Periph/ exed rockcress exed PerF Periph/ eld dodder eld AnnF Wide/NA Common name Payson’s tansymustardPayson’s AnnF Periph/ western tansy-mustard AnnF Wide/ heartleaf bittercress PerFwestern tansy-mustard AnnF Wide/ Wide/ fi rockcressDorn’s PerF LocEn/ slender dodder PerF Intro/Intro Native to large-seed dodder AnnF Wide/ defl elegant rockcressnorthern wintercress PerF PerF Wide/NA Wide/ slenderfl smalltooth dodder AnnF Wide/ rockcressHolboell’s PerF Wide/ Thorne’s perennial Thorne’s rockcress

var. ora var.

paysonii var. ora var. sparsiﬂ Synonyms/ notes Taxonomic D. incisa neuropetala A. pendulocarpa pendulocarpa, A. exilis, Boechera exilis Boechera sparsiﬂ var. Name may be misapplied, according to Holmgren et al. (2005) Boechera selbyi thorneae

var. nelsonii approximata pinetorum lipes Descurainia incisa Descurainia pinnata ssp. Cardamine cordifolia ﬁ Cuscuta epithymum ssp. Boechera holboellii var.

var. var var.

var. var. Species name var. var. ora rmed) taxa of Dinosaur National Monument, cont. ora cordifolia urceolata lipes Utah Colorado Cardamine cordifolia var. Cuscuta approximata var. Descurainia pinnata paysonii Descurainia pinnata fi Descurainia pinnata nelsonii Cuscuta pentagona Arabis holboellii Boechera glareosa Boechera glareosa pendulocarpa Barbarea orthoceras Barbarea orthoceras Cuscuta cephalanthi Cuscuta denticulataCuscuta indecora Cuscuta denticulata Grammica denticulata Grammica indecora C. indecora Arabis holboellii Arabis perennans Arabis sparsifl sparsifl pinetorum thorneae Appendix B. Potential (unconfi Family Convolvulaceae (Cuscutaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Convolvulaceae (Cuscutaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Convolvulaceae (Cuscutaceae) Convolvulaceae (Cuscutaceae) Convolvulaceae (Cuscutaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

102 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Europe noxious weed. Native to Europe Periph Periph Wide Wide Range (UT/CO) RegEn Wide Wide NA Wide Wide Periph Sparse Life form PerF RegEn/ eld pepperwort AnnF Intro/Intro Native to Asia green prince’s-plume PerF Wide/ Common name Bebb’s sedgeBebb’s beautiful sedge PerG PerG Periph/ Wide/ arrowhead thelypody PerF Wide/ ﬁ Eastwood’s tall Eastwood’s pepperwort Dame’s rocketDame’s woadDyer’s PerF Intro/Intro Native to PerF Intro/Intro UT state wedgeleaf draba AnnF Wide/ Indian Canyon twinpod PerF LocEn/ NA Utah bladderpodalpine yellowcress PerFblunt-leaf yellowcress PerF RegEn/ PerFlow yellowcress Wide/ Wide/ AnnF Wide/ branched pepperwort PerF Wide/ poor-man’s pepperwortpoor-man’s AnnF Wide/

var. in included var. P. grahamii P. Synonyms/ notes Taxonomic Thelypodium sagittatum sagittatum L. montanum spathulatum, L. eastwoodiae Holmgren et al. (2005) P. repanda; P. in ora eastwoodiae Thelypodium sagittatum Lepidium alyssoides var. Draba cuneifolia Rorippa alpina Rorippa curvipes Lepidium ramosissimum Lepidium virginicum

var. var. var. var. var. Species name oraviridifl Stanleya rmed) taxa of Dinosaur National Monument, cont. sagittata ramosissimum pubescens Thelypodiopsis sagittata var. Utah Colorado Carex bebbiiCarex bella Carex bebbii Carex bella Stanleya viridifl Lepidium campestre Neolepia campestris Lepidium ramosissimum var. Hesperis matronalisIsatis tinctoria Hesperis matronalis Lepidium alyssoides Isatis tinctoria eastwoodiae Draba cuneifolia cuneifolia Lepidium virginicum var. Physaria utahensisRorippa curvipes alpina Rorippa curvipes curvipes Rorippa tenerrima Rorippa tenerrima Lesquerella utahensis Physaria acutifolia repanda Cruciferae (Brassicaceae) Cyperaceae Cyperaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix B. Potential (unconfi Family Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

Chapter 6: Appendices 103 Comments Range (UT/CO) Sparse Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide Sparse Sparse Periph Wide Wide Life form Common name dwarf spikerush PerG Wide/ Douglas’ sedgeebony sedge PerGblunt sedge Wide/ Liddon sedge PerG Wide/ dunhead sedge PerG PerGmeadow sedge Wide/ PerG Wide/ Raynolds’ sedge Wide/ PerGslender spikerush Wide/ PerG PerG Wide/ Wide/ soft-leaved sedge PerG Wide/ red-stem waterwort AnnFNevada ephedra Sparse/ Shrub Wide/NA Canada buffaloberry Shrub Wide/ sharpscale bulrush AnnG Sparse/ river bulrushpanicled bulrush PerG PerG Sparse/ Wide/ pipsissewa PerF Wide/ uviatilis Synonyms/ notes Taxonomic E. coloradensis rubella Lepargyrea canadensis Scirpus supinus Schoenoplectus ﬂ

var. occidentalis uviatilis Eleocharis parvula anachaeta canadensis Schoenoplectus saximontanus ﬂ Chimaphila umbellata ssp.

var. Species name var. uviatilis Bolboschoenus rmed) taxa of Dinosaur National Monument, cont. occidentalis Utah Colorado Scirpus ﬂ Carex douglasiiCarex ebenea Carex douglasii Carex obtusataCarex petasata Carex ebenea Carex obtusata Carex phaeocephala Carex petasata Carex phaeocephala Carex praticolaCarex raynoldsii Carex praticola Eleocharis acicularis Carex raynoldsii Eleocharis parvula Eleocharis acicularis anachaeta Carex disperma Carex disperma Elatine triandraEphedra nevadensis Chimaphila umbellata Elatine triandravar. E. brachysperma, Shepherdia canadensis Shepherdia Scirpus microcarpusScirpus supinus Scirpus microcarpus saximontanus Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Appendix B. Potential (unconﬁ Family Cyperaceae Elatinaceae Ephedraceae Ericaceae Elaeagnaceae Cyperaceae Cyperaceae

104 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Europe. CO state noxious weed (List A) Europe. CO state noxious weed (List B) Eurasia. CO state noxious weed (List C) Eurasia Range (UT/CO) Wide Wide Wide Wide Wide Wide Wide Wide Wide Wide Life form Common name blue spurge PerF Intro/Intro Native to Gambel’s oakGambel’s Tree Wide/ alpine foxtailtall oatgrass PerG Periph/ PerG Intro/Intro Native to Thurber bentgrass PerG Periph/ felwortelkweedstork’s-bill PerF PerF crane’s-billRichardson’s Wide/ PerF AnnF Wide/ jointed goatgrass Wide/ Intro/Intro Native to AnnGalpine bengrass Intro/Intro Native to PerG Wide/ pine dropseed PerG Wide/ Engelmann’s gentianEngelmann’s PerF Wide/ northern gentian PerF Wide/

amarella var. A. humilis var. acuta, Gentianella Synonyms/ notes Taxonomic Gentianella amarella ssp. amarella magellanicus Included in by some authors G. amarella heterosepala Quercus gambelii elatius cylindricum thurberiana Blepharoneuron tricholepis Gentianella heterosepala

var. Species name rmed) taxa of Dinosaur National Monument, cont. Utah Colorado Euphorbia myrsinites myrsinites Tithymalus Quercus gambelii Gentianella amarella Gentianella acuta Gentiana amarella, gambelii Alopecurus alpinusArrhenatherum elatius Alopecurus alpinus Arrhenatherum A. borealis, Swertia perennisSwertia radiata Swertia perennis Erodium cicutarium Frasera speciosa Erodium cicutarium Geranium richardsonii Geranium richardsonii Aegilops cylindrica Cylindropyrum Agrostis humilisAgrostis thurberiana Podagrostis humilis Podagrostis Blepharoneuron tricholepis Gentianella heterosepala Appendix B. Potential (unconﬁ Family Euphorbiaceae Fagaceae Gentianaceae Gramineae (Poaceae) Gramineae (Poaceae) Gentianaceae Gentianaceae Geraniaceae Geraniaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gentianaceae

Chapter 6: Appendices 105 Comments Eurasia Eurasia. Eurasia Russia Eurasia North America Range (UT/CO) Wide Wide Wide NA Wide Wide Wide Wide Wide Wide Life form PerG Intro/Intro Native to Common name rattlesnake chess AnnG Intro/Intro Native to drooping woodreed PerGtimber oatgrass Wide/ tufted hairgrass PerG Wide/ PerG Wide/ barnyard-grass AnnGintermediate wheatgrass Intro/Intro Native to Russian wildryealkali wildrye PerGstinkgrass Intro/Intro Native to Mexican lovegrass PerG RegEn/ fescueThurber’s AnnG AnnG Intro/Introbearded sprangletop Native to W Intro/Intro PerG Native to AnnG Wide/ Wide/ false buffalograssroughleaf ricegrass AnnG Wide/ reed canary grass PerG Periph/ PerG Wide/ pullup muhly AnnG Wide/

simplex, ) var. var. fascicularis var. var. var. E. microstachya E. muricata ( Synonyms/ notes Taxonomic Welsh et al. 2003 Welsh include microstachya E. hispidus hispidus, Agropyron intermedium, Elytrigia intermedia Leymus simplex liformis intermedium juncea arundinacea fi Species name liformis Muhlenbergia rmed) taxa of Dinosaur National Monument, cont. Utah Colorado Bromus briziformis Bromus briziformis B. brizaeformis Cinna latifoliaDanthonia intermedia Danthonia intermedia Cinna latifolia Deschampsia cespitosacespitosa Deschampsia Echinochloa crus-galli Echinochloa crus-galli Elymus hispidus Thinopyrum Elymus junceusElymus simplex Psathyrostachys Eragrostis cilianensis Eragrostis cilianensis Eragrostis mexicanaFestuca thurberi Eragrostis mexicana E. orcuttiana Leptochloa fascicularis Diplachne fascicularis Festuca thurberi Muhlenbergia fi L. fusca E. simplex Munroa squarrosaOryzopsis asperifolia Monroa squarrosa Oryzopsis asperifolia Phalaris arundinacea Phalaroides Appendix B. Potential (unconfi Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae)

106 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Eurasia and W North America Eurasia Eurasia Eurasia noxious weed (List B). Native to Europe. Range (UT/CO) Wide Wide Wide Wide Wide Wide NA Wide Wide Sparse Life form Common name annual bluegrass AnnGfowl bluegrass Intro/Intro Native to nodding bluegrass PerG PerG Wide/ Wide/ alpine timothy PerG Wide/ yellow bristlegrass AnnG Intro/Intro Native to green bristlegrass AnnG Intro/Intro Native to tall dropseed PerG Sparse/ prairie wedgegrass PerG Wide/ grain sorghumalkali cordgrass PerG Intro/NA PerG Native to Wide/ European milfoil PerF Intro/Intro CO state western St. John’s-wort PerF Wide/ Siberian water-milfoil PerF Wide/ two-leaf waterweed PerF Periph/ Arnow’s ricegrassArnow’s PerG RegEn/

x pumila x var. var. S. ssp. by some Synonyms/ notes Taxonomic Included in S. obtusata obtusata M. exalbescens, M. spicatum exalbescens bloomeri authors. Hybrid of S. hymenoides comata exa obtusata sibiricum spicatum Species name exarefl Poa rmed) taxa of Dinosaur National Monument, cont. Utah Colorado Poa annuaPoa palustrisPoa refl Poa annua Setaria glauca Poa palustris Setaria glauca S. pumila Phleum alpinum Phleum commutatum Setaria viridis Setaria viridis Sporobolus asperStipa arnowiae Sporobolus asper S. compositus Sorghum bicolor Spartina gracilisSphenopholis obtusata Sphenopholis Spartina gracilis Hypericum formosum Hypericum formosum Myriophyllum sibiricum Myriophyllum Myriophyllum spicatum Myriophyllum Elodea bifoliata Elodea bifoliata Elodea longivaginata Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix B. Potential (unconfi Family Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Guttiferae (Hypericaceae, Clusiaceae) Haloragaceae Haloragaceae Hydrocharitaceae

Chapter 6: Appendices 107 Comments Wide Wide Range (UT/CO) Wide Wide Wide Sparse Periph Wide Periph Wide Wide Wide Wide Wide Wide Wide Wide PerF Wide/ Life form PerG Wide/ owered ower motherwortrough bugleweed PerF PerF Wide/ Intro/Intro Native to Asia smallfl dragonhead Common name spike woodrushmarsh arrowgrass PerG PerG Wide/ Wide/ small-fl woodrush dwarf hesperochiron PerFGreat Basin nemophila AnnF Wide/ hoary scorpion-weed Wide/ AnnFblue-eyed grass Wide/ PerF Wide/ Canadian waterweed PerF Sparse/ annual sage PerF Sparse/ beebalm PerF Periph/ Mertens’ rushNevada rush PerGknotted rush Wide/ rushParry’s PerG Wide/ PerG Wide/ PerG Wide/ Drummond’s rushDrummond’s PerG Wide/ & ora badius & orus Moldavica parvifl Synonyms/ notes Taxonomic Includes vars. drummondii subtrifl Includes vars. nevadensis

var. ora ora stulosa exa orum Monarda ﬁ Dracocephalum parviﬂ pumilus demissum menthifolia

var. orabrevifl Nemophila Species name oraparvifl Luzula stulosa exarefl Salvia rmed) taxa of Dinosaur National Monument, cont. orum Utah Colorado Leonurus cardiaca asperLycopus Leonurus cardiaca Monarda fi menthifolia asper Lycopus Luzula spicata palustrisTriglochin Luzula spicata Dracocephalum palustris Triglochin parvifl Hesperochiron pumilus Hesperochiron Nemophila brevifl Phacelia incanaSisyrinchium demissum Sisyrinchium Phacelia incana Juncus drummondii Juncus drummondii Elodea canadensis Elodea canadensis Salvia refl Juncus mertensianus Juncus mertensianus Juncus nevadensisJuncus nodosus Juncus nevadensis Juncus parryi Juncus nodosus Luzula parvifl Juncus parryi Labiatae (Lamiaceae) Labiatae (Lamiaceae) Labiatae (Lamiaceae) Juncaceae Juncaginaceae Labiatae (Lamiaceae) Hydrophyllaceae Hydrophyllaceae Hydrophyllaceae Iridaceae Juncaceae Appendix B. Potential (unconfi Family Hydrocharitaceae Labiatae (Lamiaceae) Juncaceae Juncaceae Juncaceae Juncaceae Juncaceae

108 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Europe Type locality Type is ‘SW of Theodore’ (Duchesne), Duchesne Co., UT (Jones s.n. POM). USFWS: Category 3C. CO BLM: Sensitive E of Horseshoe Bend, Uintah Co., UT (Neese & Welsh 7380 BRY). USFWS: former Category 1 Candidate Type locality Type is near Grand Canyon of the Grand River, Grand Co., CO (Patterson s.n. GH) Range (UT/CO) Wide RegEn Wide RegEn Wide RegEn Life form Common name hairy hedge-nettle PerF Wide/ Sheldon’s milkvetchSheldon’s PerF RegEn/ chickpea milkvetch PerF Intro/Intro Native to two-grooved milkvetch PerF Wide/ debris milkvetch PerF RegEn/ horseshoe milkvetch PerF LocEn/ NA locality is Type Patterson’s milkvetchPatterson’s PerF RegEn/ mountain kentrophyta PerF Wide/

var. pilosa var. S. pilosa Synonyms/ notes Taxonomic neeseae

ssp. . ssp vespertinus pilosa Stachys palustris Astragalus amphioxys var. Astragalus bisulcatus implexus Astragalus kentrophyta

Species name var. rmed) taxa of Dinosaur National Monument, cont. vespertinus bisulcatus tegetarius Utah Colorado Stachys palustris pilosa Astragalus amphioxys var. Astragalus bisulcatus var. Astragalus cicer Astragalus cicer Astragalus detritalis Astragalus detritalis Astragalus equisolensis A. desperatus Astragalus kentrophyta var. Astragalus pattersonii Astragalus pattersonii Appendix B. Potential (unconﬁ Family Labiatae (Lamiaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae)

Chapter 6: Appendices 109 Comments USFWS: 3C, USFS Region 2: Sensitive USFWS: Category 3C. Europe Europe Europe Wide Range (UT/CO) RegEn RegEn Wide NA Wide Wide Wide Wide NA Wide Life form ax PerF RegEn/ Utah yellow-ﬂ false Solomon’s-seal PerF Wide/ Common name Wetherill’s milkvetchWetherill’s PerF RegEn/ Jones’ locoweed PerF RegEn/ Green River milkvetch PerF RegEn/ silky lupine PerF Wide/ perennial sweetpea PerF Intro/Intro Native to silvery sophorared bladder-vetch PerFstrawberry clover PerF Wide/NA Intro/Intro PerF Native to Asia Intro/Intro Native to stemless lupine PerF Wide/ shortstem lupine AnnF Wide/ red cloverAmerican vetch PerF PerF Intro/Intro Wide/ Native to Gunnison’s mariposaGunnison’s PerF Wide/ wing-seed stickleaf PerF Wide/

var. var. var. amplexicaule Synonyms/ notes Taxonomic Vars. not recognized in Vars. Barneby (1989) L. caespitosus utahensis C. gunnisonii M. racemosum gunnisonii

var. ssp. Astragalus pubentissimus Lupinus sericeus Lupinus lepidus caespitosus Vicia americana americana gunnisonii amplexicaule

var.

Species name var. var. var. var. rmed) taxa of Dinosaur National Monument, cont. Utah Colorado Linum subteres Astragalus wetherillii Astragalus wetherillii Oxytropis oreophila jonesii Astragalus pubentissimus pubentissimus Lathyrus latifolius Lathyrus latifolius Lupinus sericeus sericeus Sophora stenophylla Sphaerophysa salsula Sphaerophysa salsula fragiferumTrifolium Swainsona salsula fragiferum Trifolium Lupinus brevicaulisLupinus lepidus Lupinus brevicaulis utahensis pratenseTrifolium Vicia americana pratense Trifolium americana Calochortus gunnisonii Calochortus Smilacina racemosa Maianthemum Mentzelia pterosperma Nuttallia pterosperma Linaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix B. Potential (unconﬁ Family Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Calochortaceae) Liliaceae (Convallariaceae, Ruscaceae) Loasaceae

110 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Eurasia Wide Range (UT/CO) Wide Sparse Periph Wide Wide Wide Wide Sparse Wide Periph NA Periph NA Wide Life form PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/NA ower willow- ower owered gaura owered AnnF Wide/ Common name Rocky Mountain willow-herb robust toothcupcommon mallow AnnFalkali mallow Sparse/ AnnF Intro/Intro Native to PerF Wide/ Rusby’s stickleafRusby’s PerF Periph/ small-ﬂ yellow evening- primrose white checker-mallow PerF Wide/ slender willow-herb PerF Periph/ white-ﬂ herb New Mexico checker- mallow gooseberry-leaf globemallow smooth spike-primrose AnnFAndean camissonia Periph/ wingseed camissonia AnnF AnnF Periph/ club willow-herb Periph/ PerF Wide/ pepperwort four-o’clockWatson’s PerF Wide/ Fern Wide/NA

var.

gracile vestita candida crenulata var. var. var. orum glabrata neomexicana Synonyms/ notes Taxonomic Includes vars. & & Includes vars. E. palustre lactifl S. grossulariifolia grossulariifolia orumalpinum E. ava Epilobium saximontanum Oenothera fl leptophyllum Epilobium clavatum Epilobium lactifl

var. Species name var. oramollis Gaura ava orum rmed) taxa of Dinosaur National Monument, cont. lactifl ava Utah Colorado Gaura parvifl Epilobium saximontanum Ammannia robustaMalva neglecta Ammannia robusta Malvella leprosa Malva neglecta Sidalcea candida Malvella leprosa Sidalcea candida Mentzelia rusbyi Nuttallia rusbyi Oenothera fl fl Sidalcea neomexicana Sidalcea neomexicana Epilobium leptophyllum Epilobium Sphaeralcea grossulariifolia Marsilea vestita M. vestita Boisduvalia glabellaCamissonia andinaCamissonia Camissonia andinapterosperma Epilobium alpinum Oenothera andina clavatum Epilobium hornemannii var. Epilobium pygmaeum Mirabilis alipes Mirabilis alipes Hermidium alipes Onagraceae Lythraceae Malvaceae Malvaceae Malvaceae Appendix B. Potential (unconfi Family Loasaceae Onagraceae Malvaceae Onagraceae Onagraceae Malvaceae Marsileaceae Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Nyctaginaceae

Chapter 6: Appendices 111 Eurasia Comments USFWS: 3C Wide Wide Sparse Wide Wide Periph RegEn Range (UT/CO) Wide Wide Wide Periph LocEn Wide RegEn RegEn Life form PerF Wide/ PerF Wide/ PerF LocEn/ PerF Wide/ Shrub RegEn/ PerF RegEn/ northern linanthus AnnF Wide/ longleaf plantainEnglish plantain AnnF Periph/ PerF Intro/Intro Native to limber pine Tree Wide/ linanthusNuttall’s PerF Wide/ common giliaUinta gilia AnnF Wide/ PerF RegEn/ naked broomrape PerF Sparse/ Common name blue leafy Jacob’s- ladder western Jacob’s-ladder PerF Wide/ bronze evening- primrose pale evening-primrose PerF Wide/ ephedra wild buckwheat longleaf wild buckwheat winged wild buckwheat Porter’s wild Porter’s buckwheat

alatum var. runcinata nuttallii, var. var. var. ora Leptosiphon septentrionalis Linanthastrum nuttallii Leptosiphon nuttallii occidentalis Synonyms/ notes Taxonomic P. occidentale P. Includes var. Includes var. E. brevicaule ephedroides

ssp.

orumuniﬂ O. var. ssp. exilis septentrionalis Linanthus harknessii var. Aliciella leptomeria nuttallii Polemonium foliosissimum Polemonium caeruleum amygdalinum Oenothera pallida pallida ephedroides Eriogonum lonchophyllum Eriogonum umbellatum porteri

var.

Species name var. ora unifl Aphyllon var. var. rmed) taxa of Dinosaur National Monument, cont. exilisfl Pinus pterospermum porteri Utah Colorado Polemonium caeruleum var. Plantago elongataPlantago lanceolata Plantago elongata Plantago lanceolata Linanthus septentrionalis Pinus fl Gilia leptomeria leptomeria Gilia stenothyrsaLinanthastrum nuttallii Linanthastrum Aliciella stenothyrsa Ipomopsis stenothyrsa Polemonium foliosissimum foliosissimum Eriogonum alatum Pterogonum alatum E. alatum Oenothera pallida pallida Oenothera howardii Oenothera howardii O. brachycarpa Orobanche unifl Eriogonum ephedroides Eriogonum Eriogonum lonchophyllum lonchophyllum Eriogonum umbellatum var. Polemoniaceae Plantaginaceae Plantaginaceae Polemoniaceae Pinaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polygonaceae Onagraceae Appendix B. Potential (unconfi Family Onagraceae Orobanchaceae Polygonaceae Polygonaceae Polygonaceae

112 Annotated Checklist of Vascular Flora: Dinosaur National Monument Comments Eurasia not Vars. recognized in et al. Welsh 2003 Eurasia Eurasia Wide Wide Wide NA Wide Wide Range (UT/CO) Wide Wide Wide Sparse Wide Wide Fern Wide/ Life form Rocky Mountain woodsia alpine bitter-rootslenderleaf pondweed PerF PerF Wide/ Wide/ woodland pinedrops PerF Wide/ Zion shooting-star PerF RegEn/ Richardson’s pondweedRichardson’s PerF Wide/ Common name lady’s-thumbbushy knotweed AnnF AnnF Intro/Intro Native to Wide/ sheep sorrelnarrow-leaved dock PerF PerF Intro/Intro Native to Intro/Intro Native to western dock PerF Wide/ veiny dockgreen spleenwort Fern PerF Periph/ Wide/ leafy pondweed PerF Wide/ grass pondweed PerF Wide/

var.

var. ssp. maximus var. liformis cum, P. P. cum, proliﬁ gramineus scopulina occidentalis Includes “subvar.” Includes “subvar.” huberi Synonyms/ notes Taxonomic P. ramosissimum P. var. ramosissimum ramosissimum R. aquaticus fenestratus Includes vars. & ramosum Stuckenia ﬁ

ssp.

ssp. liformis andromedea richardsonii Polygonum ramosissimum occidentalis Potamogeton gramineus Potamogeton ﬁ Potamogeton foliosus Potamogeton perfoliatus Species name

liformis var. rmed) taxa of Dinosaur National Monument, cont. occidentalis foliosus Utah Colorado Lewisia pygmaeaPotamogeton ﬁ var. Oreobroma pygmaea Potamogeton foliosus var. Pterospora andromedea Pterospora zionense Dodecatheon pulchellum Polygonum persicaria Persicaria maculosa Polygonum ramosissimum Rumex acetosella Acetosella vulgaris Rumex stenophyllus Rumex stenophyllus Rumex occidentalis Rumex aquaticus Rumex venosusAsplenium viride Rumex venosus scopulinaWoodsia Asplenium viride scopulina Woodsia A. trichomanes- scopulina W. Potamogeton gramineus Potamogeton richardsonii Portulacaceae Potamogetonaceae Potamogetonaceae Pyrolaceae (Ericaceae, Monotropaceae) Primulaceae Appendix B. Potential (unconﬁ Family Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polypodiaceae (Aspleniaceae) Polypodiaceae (Aspleniaceae, Woodsiaceae) Potamogetonaceae Potamogetonaceae

Chapter 6: Appendices 113 Comments Native to Eurasia Wide Wide Wide Wide Range (UT/CO) Intro Wide Wide Wide Wide Wide RegEn Wide Wide Wide Wide Wide PerF Wide/ Life form ower meadow- ower Virginia strawberryerect avens PerF Wide/ PerF Sparse/ Fendler’s mountain-lilacFendler’s Shrub Wide/ few-ﬂ rue Common name creeping buttercup PerF Intro/ sugar-bowls PerF Wide/ tiny mouse-tail AnnF Sparse/ western larkspurbristly mouse-tail PerF AnnF Wide/ Wide/ Geyer’s larkspurGeyer’s PerF RegEn/ purple virgin’s-bower PerF Wide/ alpine buttercup PerF Wide/ heart-leaf buttercup PerF Wide/ plantain buttercup PerF Wide/ creeping buttercup PerFdrab buttercup Wide/ PerF Wide/

var. var. D. ammula adoneus glauca var.

x orum reptans ammula alpinus liformis, R. fl platypetala Includes ssp. T. sparsifl T. saximontanum Synonyms/ notes Taxonomic hirsutissima Includes vars. & D. glaucum barbeyi, D. glaucum fi var. Includes var. alpeophilus

var. var. orum ora hirsutissima C. hirsutissima glauca ssp. Ranunculus repens sparsiﬂ Myosurus apetalus M. aristatus occidentale Atragene occidentalis Ranunculus cardiophyllus Ranunculus alismifolius montanus inamoenus Ranunculus reptans R. ﬂ inamoenus

var. var. orum Thalictrum Species name ammula rmed) taxa of Dinosaur National Monument, cont. grosseserrata montanus ovalis Utah Colorado Geum aleppicum Geum aleppicum Ceanothus fendleriFragaria virginiana Ceanothus fendleri Fragaria virginiana Clematis hirsutissima Corifl Thalictrum sparsifl Clematis occidentalis var. Myosurus apetalus Myosurus minimusRanunculus adoneus Myosurus minimus Ranunculus adoneus montanus Delphinium geyeriDelphinium occidentale Delphinium geyeri Delphinium Ranunculus alismifolius var. repens Ranunculus cardiophyllus Ranunculus fl var. Ranunculus inamoenus Ranunculus Ranunculus repens Rosaceae Rhamnaceae Rosaceae Appendix B. Potential (unconfi Family Ranunculaceae Ranunculaceae (Thalictraceae) Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae

114 Annotated Checklist of Vascular Flora: Dinosaur National Monument Eurasia Comments Eurasia Native to C North America Wide Wide NA Range (UT/CO) Wide Wide Wide Wide Wide Periph Wide Sparse Wide Wide Wide Wide Life form PerF RegEn/ Common name crack willowwavy-leaved Indian- Treepaintbrush Intro/Intro Native to Drummond’s willowDrummond’s Shrub Wide/ willowGeyer’s Shrubsaline bird’s-beak Wide/ AnnF LocEn/ NA white poplar Tree Intro/Intro Native to ditch-grass PerF Wide/ prairie smoke PerF Wide/ rough bedstraw PerF Periph/ woolly cinquefoil PerF Wide/ brook cinquefoilAmerican plum AnnF Sparse/ Shrub Intro/ sticky cinquefoil PerFsheep cinquefoil Wide/ bushy cinquefoil PerF Wide/ AnnF Wide/ rock mountain-ashnorthern bedstraw Shrub Wide/ PerF Wide/ western thimbleberry Shrub Wide/

ssp. var. var. var. var. Synonyms/ notes Taxonomic C. applegatei pinetorum geyeriana R. maritima occidentalis hippiana scopulina

ssp. ora var. ssp. orum asperrimum occidentalis Galium mexicanum ssp. Erythrocoma triﬂ Drymocallis glandulosa Potentilla ovina decurrens paradoxa Rubacer parviﬂ

var.

var. Species name var. var. orus orum rmed) taxa of Dinosaur National Monument, cont. orus orus viscida pseudorupestris Utah Colorado Salix fragilisCastilleja applegatei Salix fragilis var. Cordylanthus kingii Salix drummondiana Salix drummondiana densifl Salix geyeriana Salix geyeriana S. geyeriana Geum trifl Populus alba Populus alba Potentilla glandulosa var. Ruppia cirrhosa Ruppia cirrhosa asperulum ciliatum Potentilla hippiana Potentilla ovina Potentilla hippiana hippiana P. Potentilla paradoxaPotentilla rivalis Potentilla supina Prunus americana Potentilla rivalis Prunus americana decurrens Rubus parvifl parvifl Sorbus scopulina Galium boreale Sorbus scopulina Galium mexicanum S. scopulina Galium septentrionale Salicaceae Scrophulariaceae Scrophulariaceae Salicaceae Appendix B. Potential (unconfi Family Rosaceae Salicaceae Rosaceae Ruppiaceae Salicaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Rubiaceae Rubiaceae

Chapter 6: Appendices 115 Comments is 3.7 km E of Lapoint, Uintah Co., UT (Holmgren, Holmgren, & Barneby 8760 NY). UT BLM: Sensitive, USFWS: former C2 candidate Type locality Type is near Sand Uintah Wash, Co., UT (Graham 7883 CM). USFWS: formerly Proposed Threatened USFWS: Candidate Range (UT/CO) NA RegEn LocEn NA LocEn Wide Wide NA LocEn Life form ower PerF Periph/ eshy penstemon eshy PerF RegEn/ Common name dusty penstemon penstemonGoodrich’s PerF PerF RegEn/ LocEn/ NA locality Type shortleaf penstemon PerFﬂ RegEn/ penstemonGraham’s PerF LocEn/ Yampa penstemonYampa PerF LocEn/ dwarf lousewort PerF Wide/ thickleaf penstemon PerF RegEn/ glabrous monkeyﬂ White River penstemon PerF LocEn/

var. Synonyms/ notes Taxonomic P. yampaensis P. jamesii

var. uvis albiﬂ Penstemon comarrhenus centranthera Penstemon acaulis yampaensis Mimulus glabratus M. glabratus Penstemon scariosus var.

var. var. Species name

var. uvis rmed) taxa of Dinosaur National Monument, cont. perbrevis albiﬂ Utah Colorado Penstemon comarrhenus Penstemon goodrichii Penstemon caespitosus var. Penstemon carnosus Penstemon grahamii Penstemon grahamii Pedicularis centranthera Pedicularis Penstemon acaulis yampaensis Penstemon pachyphyllus pachyphyllus Penstemon scariosus var. Mimulus glabratus fremontii Appendix B. Potential (unconﬁ Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Family Scrophulariaceae

116 Annotated Checklist of Vascular Flora: Dinosaur National Monument noxious weed (List C). Native to Eurasia. Comments Eurasia Eurasia Native to C North America America CO state noxious weed (List B). Native to Eurasia. Wide Range (UT/CO) NA Wide Wide Wide Wide Intro PerF Wide/NA Life form PerF Wide/NA PerF Wide/ PerF Wide/NA AnnF Intro/Intro Native to South PerF Wide/ water-hemlockpoison-hemlock PerF PerF Wide/ Intro/Intro CO state long-stalk spring- parsley basin white-cup spring- Common name parsley two-lobe speedwellchain speedwell AnnF Intro/IntroAmerican alpine Native to Asia PerFspeedwell matrimony-vine Sparse/ Shrub Intro/Intro Native to plateau penstemon PerF RegEn/ European bittersweet PerFbuffalobur Intro/Intro Native to AnnF Intro/ long-leaf ground- cherry ground-cherry nightshade narrowleaved burr- reed caraway PerF Intro/

var. V. anagallis- V. var. by some longipes Synonyms/ notes Taxonomic Included in aquatica authors sonorae S. villosum

var. Cicuta douglasii C. occidentalis physalifolium nitidibaccatum Sparganium angustifolium

Species name var. rmed) taxa of Dinosaur National Monument, cont. scariosus Utah Colorado angustifolia Conium maculatum Conium maculatum Cymopterus longipesCymopterus purpurascens C. longipes Veronica bilobaVeronica catenataVeronica Pocilla biloba catenata Veronica wormskjoldiiVeronica nutans Veronica barbarumLycium barbarum Lycium L. halmifolium Penstemon scariosus var. Physalis longifoliaSolanum dulcamaraSolanum rostratum Solanum dulcamara Solanum rostratum virginiana P. Solanum sarrachoides Solanum Sparganium angustifolium Carum carvi Carum carvi Cicuta maculata Appendix B. Potential (unconﬁ Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Scrophulariaceae Scrophulariaceae Scrophulariaceae Solanaceae Family Scrophulariaceae Solanaceae Solanaceae Solanaceae Solanaceae Sparganiaceae Umbelliferae (Apiaceae) Umbelliferae (Apiaceae)

Chapter 6: Appendices 117

) Atlas of P. Comments Europe Eurasia North America noxious weed (List C). Native to Eurasia. Typically Typically parasitic on Pinus contorta (rarely on ponderosa Range (UT/CO) Wide Wide Wide Wide Wide Wide Wide Life form PerF Periph/ considered as potentially occurring in the monu- considered . Duane Atwood, of Brigham Young University, in 2001, the University, . Duane Atwood, of Brigham Young tah) and Moffat County, Colorado, but have not previously been reported been reported Colorado, but have not previously County, tah) and Moffat Common name common parsnipPennsylvania pellitory PerF AnnFstinging nettle Intro/Intro Wide/ Native to tobacco-root PerFwedge-leaf fog-fruit Periph/ PerFblue vervain PerF Sparse/ pansy Wide/ lodgepole pine dwarf- PerFmistletoe Wide/ AnnFVirginia creeper Intro/NApuncture vine Native to Shrub Intro/Intro Native to E AnnF Intro/Intro CO state cow parsnip PerF Wide/

var. gracilis ssp. Synonyms/ notes Taxonomic U. dioica H. maximum, sphondylium lanatum

ssp. ssp. pensylvanica Urtica gracilis gracilis Arceuthobium americanum Parthenocissus quinquefolia sphondylium montanum Species name

var. rmed) taxa of Dinosaur National Monument, cont. Utah Colorado Pastinaca sativaParietaria pensylvanica Parietaria Pastinaca sativa Urtica dioica procera edulisValeriana Phyla cuneifolia edulis Valeriana hastataVerbena Phyla cuneifoliaViola tricolor hastata Verbena Arceuthobium Lippia cuneifolia americanum Parthenocissus quinquefolia terrestrisTribulus terrestris Tribulus Heracleum lanatum Heracleum (Albee et al. 1988), and records collected just outside Dinosaur NM (Weber et al. 1982). Some species from Appendix C could be et al. 1982). Some species from collected just outside Dinosaur NM (Weber (Albee et al. 1988), and records rmed for the monument. Species on this list are derived from a checklist of potential species for NCPN parks prepared by Dr. N by Dr. a checklist of potential species for NCPN parks prepared derived from rmed for the monument. Species on this list are Umbelliferae (Apiaceae) Urticaceae Urticaceae Valerianaceae Verbenaceae Verbenaceae Violaceae Viscaceae Vitaceae Zygophyllaceae Appendix B. Potential (unconﬁ Family Umbelliferae (Apiaceae) he Vascular Flora of Utah he Vascular t not included here. ment, but are The vascular plant species listed here are known from the vicinity of Dinosaur NM, in Daggett, Duchesne, and Uintah counties (U known from are The vascular plant species listed here or conﬁ

118 Annotated Checklist of Vascular Flora: Dinosaur National Monument

. ed ed . . var. var. A. fendleri M. repens. ora

ed specimens ed ed specimens ed ed specimen ed ed specimen ed A. uintahensis eastwoodiae. O. erinacea M. fusiformis Lappula occidentalis C. brevifl utahensis. of herbarium (without variety) based on misidentifi of et al. (1982) based on misidentifi of (1962) based on misidentifi locality is Wasatch Type Mountains, Salt Lake 858 Co., UT (Watson GH) (1962) based on misidentifi of Museum herbarium based on misidentifi specimen of Native to NW North America. (1962) as based on misidentifi specimens of var. UT CO Comments FalsRep X Reported by Holmgren FalsRep X Report from DINO FalsRep X Reported by Weber FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Report from Carnegie Park status Wide Wide Wide Wide RegEn NA Range PerF Wide/ PerF Wide/ AnnF Wide/ PerF RegEn/ Shrub Intro/ Life form basin sandwort PerF Wide/ berry pricklypear short-styled bluebell Watson’s Watson’s cryptanth plateau cryptanth shining mahonia Common name

ssp. ssp. var. plateauensis, E. kingii glabrescens, A. kingii uintahensis, A. kingii compacta, A. kingii ssp. A. uintahensis Synonyms/ notes Taxonomic uintahensis Eremogone kingii ssp. Opuntia phaeacantha Mertensia brevistyla Cryptantha watsonii Oreocarya fulvocanescens Species name glabrescens Arenaria fendleri var. Opuntia phaeacantha Mertensia brevistyla Cryptantha watsonii Cryptantha fulvocanescens Mahonia aquilifolium Utah Colorado Caryophyllaceae (Alsinaceae) Cactaceae Boraginaceae Boraginaceae Boraginaceae Family Berberidaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument. Appendix C. Rejected (falsely reported

Chapter 6: Appendices 119 . var. var. lanata. . Native to Ambrosia . var. . ed specimen ed ed specimen ed ed specimen ed ed ed specimen ed ed specimen ed ed specimen ed A. perelegans. A. pansus K. lanata C. americanum C. simplex B. microphylla of (1962) based on misidentifi of (2002) based on misidentifi herbarium based on misidentifi of (1962) based on misidentifi specimen of tomentosa of (1962) based on misidentifi of Eurasia. (1962) based on misidentifi of scabra. (1962) based on misidentifi (in ed.), but DINO is outside the expected range of the species. UT CO Comments FalsRep X Reported by Holmgren FalsRep X Reported by Naumann FalsRep X Report from DINO FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren Ques? Reported by NCPN Park status Wide Wide NA LocEn NA Intro NA Periph Range PerF Wide/ Shrub Periph/ AnnF NA/ Shrub Wide/ Life form Shrub Wide/ Engelmann’s Engelmann’s aster Eaton’s asterEaton’s PerF Wide/ linearleaf ragweed shrubby winterfat hairy bugseed AnnF Wide/ lambsquarter AnnF Intro/ Watson’s Watson’s brickellbush Common name longleaf brickellbush

Flora var. var. subspinosa. B. longifolia eatonii Ceratoides lanata var. not Vars. recognized in of North America (2003). C. hyssopifolium, C. nitidum longifolia Synonyms/ notes Taxonomic Brickellia longifolia engelmannii Ambrosia linearis Franseria linearis Chenopodium album

Species name var.

album var. var. Brickellia longifolia Aster engelmannii Eucephalus Aster eatonii Aster bracteolatus Symphyotrichum Krascheninnikovia lanata subspinosa Corispermum villosum Chenopodium album Brickellia microphylla watsonii Utah Colorado Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Family Chenopodiaceae Compositae (Asteraceae) Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

120 Annotated Checklist of Vascular Flora: Dinosaur National Monument ed

var. var. var. var. from var. var. var. . E. . orus ed specimen ed ed specimen ed specimen ed ed specimen ed ed specimen ed . E. divergens C. undulatum C. scariosum C. viscidifl E. pumilus divergens. (1962) without variety based on misidentifi specimen of pulcherrimus of (2002) based on misidentifi (2002) without variety based on misidentifi of undulatum. (1962) based on misidentifi of outside DINO. Native to northern North America. Not native to UT or CO. of (2002) based on misidentifi stenophyllus concinnus (1962) and Weber et (1962) and Weber al. (1982) based on misidentifi of UT CO Comments FalsRep X Reported by Holmgren FalsRep X Reported by Naumann FalsRep X Reported by Naumann FalsRep X Reported by Naumann FalsRep X Reported by Holmgren Park status NA Wide Wide Wide RegEn Range PerF RegEn/ Shrub Wide/ PerF RegEn/ Life form eabane slenderleaf daisy pretty daisy AnnF Wide/ pretty thistle PerF Wide/ dwarf thistle PerF NA/NA FalsRep X Reported by Holmgren lanceleaf rabbitbrush La Veta Pass La Veta fl Common name in C. var. var. 2006) bellidiastrum E. sparsifolius E. bellidiastrum var. Var. not Var. recognized by Cronquist (1994), included in arizonicum bipinnatum Flora of North America Cirsium drummondii Synonyms/ notes Taxonomic

ssp. orus pulchellum Erigeron bellidiastrum Cirsium calcareum var. lanceolatus Chrysothamnus viscidiﬂ

Species name var. orus sparsifolius pulchellum Erigeron utahensis var. Erigeron bellidiastrum Cirsium calcareum var. lanceolatus Chrysothamnus viscidiﬂ Erigeron vetensis Erigeron vetensis Utah Colorado Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

Chapter 6: Appendices 121

M. ora

var. var. . USFWS: var. var. ora , but ora ora. ed specimens ed ed specimen ed ed specimen ed ed specimen ed specimen ed ). Aster lifolius Stephanomeria multifl grandifl L. grandifl H. lapidicola H. fi G. squarrosa Viguiera multifl and runcinata. var. (1962) based on misidentifi of cinereus. herbarium based on misidentifi of former C2 (1962) based on misidentifi of (1962) based on misidentifi of serrulata. of var. herbarium based on misidentifi (1962) as rubrotinctus monument is outside the expected range of this taxon (report Probably based on canescens UT CO Comments FalsRep X Reported by Holmgren FalsRep X Report from DINO FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Report from DINO Ques? X Reported by Holmgren Park status Wide NA NA Periph RegEn Wide Range PerF Periph/ PerF RegEn/ PerF Periph/ PerF Wide/ Life form ower PerF Intro/ rush skeleton- plant low hymenoxys PerF LocEn/ little sunfl bulb hyalineherb Rydberg’s Rydberg’s gumweed Bigelow’s tansy- Bigelow’s aster Common name

var. bigelovii, Aster nana H. depressa, acaulis Tetraneuris var. aphanactis G. nuda Dieteria bigelovii var. rubrotinctus Synonyms/ notes Taxonomic Lygodesmia Lygodesmia juncea Helianthus pumilus Grindelia aphanactis Machaeranthera bigelovii

Species name

var. var. nana lifolius Hymenoxys acaulis var. Lygodesmia Lygodesmia juncea Helianthus pumilus Hymenopappus ﬁ eriopodus Grindelia aphanactis Machaeranthera bigelovii bigelovii Utah Colorado Compositae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) (Asteraceae) Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

122 Annotated Checklist of Vascular Flora: Dinosaur National Monument

ed ed . var. var. var. var. var. var. var. T. S. nana var. var. ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed S. spathulata . S. canadensis S. canadensis P. integrifolia P. M. canescens S. multilobatus Platyschkuhria (2002) as (1962) based on a misidentifi of salebrosa et al. (1982) based on misidentifi of salebrosa. of (1962) based on misidentifi ourolepis. (2002) without variety based on misidentifi specimens of mensana of herbarium based on misidentifi canescens. (1962) based on misidentifi of herbarium based on misidentifi of integrifolia ourolepis. based on misidentifi specimens of UT CO Comments FalsRep X Reported by Naumann FalsRep X Reported by Holmgren FalsRep X Reported by Weber FalsRep X Reported by Holmgren FalsRep X Reported by Naumann FalsRep X Report from DINO FalsRep X Reported by Holmgren FalsRep X Report from DINO Park status Wide Wide Wide RegEn NA NA Wide Periph Range PerF Wide/ PerF Periph/ PerF NA/ PerF Wide/ Life form coast goldenrod PerF Wide/ Rocky Mountain goldenrod late goldenrod PerF Periph/ San Juan bahia PerF RegEn/ Wyoming townsendia thickleaf aster PerF RegEn/ Fendler’s Fendler’s groundsel scapose greenthread Common name

nana var. simplex oblongifolia, & var. includes vars. Bahia nudicaulis var. B. oblongifolia T. alpigena T. alpigena M. leucanthemifolia, Aster leucanthemifolius Synonyms/ notes Taxonomic

var. Solidago multiradiata Platyschkuhria integrifolia oblongifolia Packera fendleri Senecio fendleri Thelesperma subnudum

var. Species name var. var. var. var. Solidago simplex Solidago simplex S. spathulata, Solidago multiradiata scopulorum Solidago gigantea Solidago gigantea Platyschkuhria integrifolia oblongifolia montana Townsendia Townsendia montana Machaeranthera canescens leucanthemifolia Thelesperma subnudum subnudum Utah Colorado Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) Compositae (Asteraceae) Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

Chapter 6: Appendices 123 var. var. var. var. . . CO . Native to ed specimen ed ed specimen ed ed specimen ed ed specimen ed specimen ed Aster based on L. ramosissimum C. pubescens X. venusta. T. dubius T. L. montanum (2002) without variety based on misidentifi of bourgeauanum (1962) based on specimen collected from outside the monument. (2002) based on misidentifi of state noxious weed (List B). Native to Europe (1962) as xylorhiza misidentifi of (1962) based on specimen collected from outside the monument. of herbarium based on misidentifi of jonesii. (1962) based on misidentifi Europe. UT CO Comments FalsRep X Reported by Naumann FalsRep X Reported by Holmgren FalsRep X Reported by Naumann FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Report from DINO FalsRep X Reported by Holmgren Park status Wide Wide Intro Wide Wide Wide Intro Range AnnF Wide/ PerF Wide/ PerF Periph/ PerF Wide/ Life form prairie pepperwort slender whitlow-grass whitetop PerF Intro/ smooth woodyaster mule’s-ears PerF Wide/ salsify PerF Intro/ mountain pepperwort Common name draba var. draba Machaeranthera glabriuscula Synonyms/ notes Taxonomic

var. orum Lepidium densiﬂ Draba albertina Xylorhiza glabriuscula Wyethia amplexicaulis Tragopogon Tragopogon porrifolius Lepidium montanum montanum

var. var. Species name var. orum orum nana densiﬂ Lepidium densiﬂ Draba stenoloba var. Cardaria draba Cardaria draba Lepidium draba, C. Xylorhiza glabriuscula glabriuscula Wyethia amplexicaulis Tragopogon Tragopogon porrifolius Lepidium montanum montanum Utah Colorado Cruciferae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Compositae (Asteraceae) (Brassicaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

124 Annotated Checklist of Vascular Flora: Dinosaur National Monument

E. ed ed

. var. var. Not . R. islandica P. acutifolia P. based on ed specimens ed specimen ed ed specimen ed ed specimen ed R. obtusa Lesquerella based on glabra. acutifolia. P. subumbellata. P. C. nebrascensis P. acutifolia P. E. palustris (1962) as based on misidentifi specimen of var. of (2002) based on misidentifi of al. (1982) and Naumann (2002) as alpina misidentifi State University herbarium based on misidentifi of acutifolia. Museum herbarium based on misidentifi specimen of var. native to UT or CO. (1962) based on specimen collected from outside the monument. Holmgren (1962) as engelmannii misidentifi of UT CO Comments FalsRep X Reported by Holmgren FalsRep X Reported by Naumann FalsRep X X et Reported by Weber FalsRep X Report from Colorado FalsRep X Reported by Holmgren FalsRep X Reported by Park status Sparse Wide Wide LocEn Wide Wide Range AnnF Sparse/ PerF Wide/ Life form PerF NA/NA FalsRep X Report from Carnegie round-pod yellowcress lakeshore sedge PerG Wide/ alpine bladderpod mesa twinpod PerF LocEn/ Common name common twinpod beaked sedge PerG Wide/ ovate spikerush AnnG Periph/

var. (sensu includes spatulata, P. spatulata, P. ovata, E. R. obtusa C. lenticularis pallida Lesquerella alpina, var. reediana Physaria didymocarpa lato) E. obtusa var. engelmannii Synonyms/ notes Taxonomic detonsa alpina lipocarpa oribunda Rorippa sphaerocarpa Carex lenticularis var. Lesquerella alpina ssp. Physaria ﬂ var. Species name lipocarpa reediana oribunda Rorippa sphaerocarpa Carex lenticularis var. Physaria reediana var. Carex utriculata Carex utriculata C. rostrata Physaria ﬂ Eleocharis ovata Eleocharis obtusa Utah Colorado Cruciferae (Brassicaceae) Cyperaceae Cruciferae (Brassicaceae) Family Cruciferae (Brassicaceae) Cyperaceae Cruciferae (Brassicaceae) Cyperaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

Chapter 6: Appendices 125

. . var. Native to . romati. var. var. ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed B. carinatus E. laevigatum Limosella F. ovina F. S. pungens E. smithii. E. elymoides. of herbarium based on misidentifi (2002) based on misidentifi of (1962) based on specimen collected from outside the monument. of et al. (1982) based on misidentifi herbarium based on misidentifi of at al. (1982) based on misidentifi of saximontana. E and N North America (1962) based on misidentifi of longispicatus of herbarium based on misidentifi UT CO Comments FalsRep X Report from DINO FalsRep X Reported by Naumann FalsRep X Report from DINO FalsRep X Reported by Holmgren FalsRep X Reported by Weber FalsRep X Report from DINO FalsRep X Reported by Weber FalsRep X Reported by Holmgren Park status Periph Wide NA Wide Sparse Disj Intro NA Range Fern Disj/ PerG Wide/ Life form big squirreltail PerG Sparse/ fringed brome PerG Wide/ false quackgrass PerG Sparse/ bearberry Shrub Wide/ northern scouring-rush Texas bergiaTexas AnnF Sparse/ red fescue PerG Intro/ American bulrush Common name

var. E. x rubra x E. var. & Agropyron pseudorepens. Hybrid of trachycaulus Includes vars. ciliatus richardsonii lanceolatus Vars. not Vars. recognized in Holmgren et al. (2005) E. variegatum variegatum Schoenoplectus americanus Synonyms/ notes Taxonomic

ssp. canadensis coactilis Arctostaphylos uva-ursi Hippochaete variegata Species name x Elymus multisetus Elymus multisetus Sitanion jubatum Elymus pseudorepens Bromus ciliatus Bromopsis Arctostaphylos uva-ursi Equisetum variegatum Bergia texana Bergia texana Festuca rubra Festuca rubra rubra F. Scirpus americanus Scirpus olneyi, Utah Colorado Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Ericaceae Equisetaceae Elatinaceae Gramineae (Poaceae) Family Cyperaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

126 Annotated Checklist of Vascular Flora: Dinosaur National Monument

. ciliosa. var. var. . P. nervosa P. var. var. . var. var. ed specimen ed ed hybrid ed specimen ed ed specimen ed ed specimen ed ed specimens ed ed specimen ed P. crenulata P. S. comata P. acuminatum P. M. bulbosa. P. sericea P. P. hastata P. P. pratensis P. (2002) based on misidentifi of corrugata specimen of of comata. (2002) based on misidentifi x herbarium based on misidentifi of (1962) based on misidentifi of (1962), Weber et al. (1962), Weber (1982), and Naumann (2002) based on misidentifi and Neese (1986) based on Goodrich # 12317 which was collected just outside the monument. Endemic to Idaho. of (1962) based on misidentifi of (2002) based on misidentifi UT CO Comments FalsRep X Reported by Naumann FalsRep X Reported by Naumann FalsRep X Report from DINO FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reported by Goodrich FalsRep X Reported by Naumann Park status RegEn Wide Wide Wide Wide Wide Wide Range PerF RegEn/ PerG Wide/ PerG Sparse/ PerG Wide/ Life form glandular scorpion-weed bog bluegrass PerG Wide/ needle-and- thread Scribner panicgrass purple oniongrass little barley AnnG Wide/ Common name Idaho phacelia PerF NA/NA FalsRep X X Reported by Holmgren wand phacelia PerF Wide/

& var.

scribnerianum Includes ssp. D. oligosanthes var. heterophylla Phacelia idahoensis virgata Synonyms/ notes Taxonomic

var. Phacelia heterophylla Phacelia glandulosa Hesperostipa comata intermedia Dichanthelium oligosanthes spectabilis

var. var. Species name Phacelia heterophylla Phacelia glandulosa Poa leptocoma Poa leptocoma Stipa comata intermedia Panicum oligosanthes scribnerianum Melica spectabilis Bromelica Hordeum pusillum Critesion pusillum Utah Colorado Hydrophyllaceae Hydrophyllaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Gramineae (Poaceae) Hydrophyllaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

Chapter 6: Appendices 127 . A. . var or . . USFWS: USFS . ed specimen ed ed specimens ed ed specimen ed ed specimen ed ed specimen ed ed specimens ed ed specimen ed ed specimen ed . Endemic to A. saurinus A. chloodes A. duchesnensis A. chamaeleuce. A. saurinus. J. tenuis A. spatulatus A. convallarius of 3C (1962) based on misidentifi (1962) based on misidentifi of of USFWS: 3C herbarium based on misidentifi of convallarius. Region 4: Sensitive (1962) based on misidentifi (1962) based on misidentifi of herbarium based on misidentifi of of (1962) and Naumann (2002) based on misidentifi (1962) based on misidentifi of chloodes Wyoming. UT CO Comments FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Report from DINO FalsRep X X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reports from DINO FalsRep X Reported by Holmgren Park status RegEn NA NA NA NA NA Wide Range PerF RegEn/ PerF RegEn/ PerF RegEn/ PerF RegEn/ Life form PerF NA/NA FalsRep X Reported by Holmgren San Rafael milkvetch Moenkopi milkvetch lancer milkvetch PerF RegEn/ meadow milkvetch canoe milkvetch PerF LocEn/ Colton’s Colton’s milkvetch perplexing rush PerG Wide/ Common name little bun milkvetch

var. diversifolius A. diversifolius Astragalus simplicifolius Synonyms/ notes Taxonomic Astragalus rafaelensis Species name coltonii Astragalus rafaelensis Astragalus moencoppensis Astragalus lancearius Astragalus diversifolius Astragalus cymboides Astragalus coltonii var. Juncus confusus Juncus confusus Utah Colorado Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Family Juncaceae Leguminosae (Fabaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

128 Annotated Checklist of Vascular Flora: Dinosaur National Monument

ed . . var. var. .

var. O. besseyi ed specimen ed ed specimen ed ed specimens ed specimen ed ed specimen ed . obnapiformis. P. megalanthum P. T. gymnocarpon T. A. mollissimus L. pusillus T. rhombifolia T. herbarium based on misidentifi of (2002) without variety based on misidentifi specimen of var. (1962) based on specimen collected from outside the monument. pusillus Holmgren (1962) based on specimen collected from outside the monument. Report from DINO herbarium based on a misidentifi of of thompsoniae (1962) based on misidentifi of (2002) based on misidentifi (1962) based on misidentifi of UT CO Comments FalsRep X Report from DINO FalsRep X Reported by Naumann FalsRep X Reported by Holmgren FalsRep X Reported by FalsRep X Reported by Holmgren FalsRep X Reported by Naumann FalsRep X Reported by Holmgren Park status NA Wide Wide Wide NA NA Wide Range PerF RegEn/ PerF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ Life form skunk breadroot Lambert’s Lambert’s locoweed white locoweed PerF Wide/ Wind River clover Utah milkvetch PerF Wide/ blackbrush lupine montane golden pea Common name

var. by var.

includes included pusillus mephitica montana, rhombifolia T. T. pinetorum, T. pinetorum, T. T. rhombifolia Psoralea mephitica var. Includes var. lambertii L. rubens, in var. some authors ovata Synonyms/ notes Taxonomic Thermopsis montana Oxytropis lambertii Oxytropis sericea Trifolium longipes Trifolium

Species name var. exum sericea rubens reﬂ Thermopsis montana Pediomelum mephiticum Oxytropis lambertii bigelovii Oxytropis sericea var. Lupinus pusillus var. Trifolium longipes Trifolium var. Astragalus utahensis Utah Colorado Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Family Leguminosae (Fabaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

Chapter 6: Appendices 129 ...... ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed G. decipiens G. ramosissimum S. parvifolia. M. thompsonii Z. vaginatus Ipomopsis pumila A. acuminatum H. zothecina (2002) based on misidentifi of of et al. (1982) based on misidentifi of (2002) based on misidentifi (1962) based on misidentifi of (1962) based on misidentifi of of (1962) based on misidentifi of herbarium based on misidentifi (1962) based on misidentifi of UT CO Comments FalsRep X Reported by Naumann FalsRep X Reported by Weber FalsRep X Reported by Naumann FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Report from DINO FalsRep X Reported by Holmgren Park status Wide Wide NA Wide Wide Periph Wide Wide Range AnnF Wide/ AnnF Wide/ PerF Wide/ PerF Wide/ AnnF Wide/ PerF Wide/ Life form raceme groundsmoke diffuse diffuse groundsmoke white-stem globe-mallow Nevada stickleaf AnnF Wide/ mountain death camas spreading eriastrum nodding onion PerF Wide/ northern bog orchid Common name

latifolia & & &

G. lasiospermum Includes ssps. munroana subrhomboidea Includes vars. compacta, dispersa, Includes vars. cernuum obtusum Includes Platanthera aquilonis, P. huronensis Synonyms/ notes Taxonomic

ssp. orum Limnorchis Gayophytum racemosum Gayophytum diffusum parviﬂ Eriastrum diffusum hyperborea

Species name var. Habenaria hyperborea Gayophytum racemosum Gayophytum diffusum strictipes Sphaeralcea munroana Mentzelia dispersa Acrolasia dispersa Zigadenus elegans Anticlea elegans Eriastrum diffusum Allium cernuum Allium cernuum Utah Colorado Orchidaceae Onagraceae Onagraceae Malvaceae Loasaceae Liliaceae (Melianthaceae) Polemoniaceae Family Liliaceae (Alliaceae) Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

130 Annotated Checklist of Vascular Flora: Dinosaur National Monument

. Type . Type . CO var. var. da ed specimens ed ed specimen ed ed specimen ed . Type locality . Type E. tumulosum I. congesta I. aggregata G. pinnatifi herbarium based on misidentifyied specimen of BLM: Sensitive. (1962) based on specimen collected from outside the monument. (1962), but DINO is well outside the expected range of this species (Cronquist et al. 1984). Not native to UT or CO. of congesta is Grand Junction, Mesa Co., CO (Eastwood s.n. NY). (1962) based on misidentifi aggregata of herbarium based on misidentifi of locality is “plains of the San Juan River in S Colorado or adjacent Utah” (Brandegee s.n.). (1962) based on misidentifi UT CO Comments FalsRep X Report from DINO FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Report from DINO FalsRep X Reported by Holmgren Park status RegEn Wide RegEn Wide RegEn Range PerF RegEn/ AnnF Sparse/ Life form single-stem wild buckwheat Harkness’ linanthus spreading phlox PerF NA/NA Ques? X Reported by Holmgren San Rafael gilia PerF RegEn/ scarlet gilia PerF Wide/ San Juan gilia PerF RegEn/ Common name maculata Leptosiphon harknessii Phlox diffusa Gilia aggregata var. Synonyms/ notes Taxonomic

ssp. var. var. septentrionalis Linanthus harknessii Ipomopsis aggregata formosissima

Species name var. Eriogonum acaule Eriogonum acaule Linanthus harknessii Ipomopsis roseata Ipomopsis roseata Gilia roseata Ipomopsis aggregata maculata Gilia haydenii Gilia haydenii Aliciella haydenii Utah Colorado Polemoniaceae Polygonaceae Polemoniaceae Polemoniaceae Polemoniaceae Family Polemoniaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

Chapter 6: Appendices 131

P. and var. var. based from ed P. glabella P. (Goodrich ana ed specimen ed ed specimen ed ed specimen ed ed specimens ed occidentalis P. kelloggii P. E. tumulosum. E. tumulosum. E. batemanii Holmgren (1962) as suksdorfi P. on misidentifi specimens of var. breweri. outside the monument of herbarium based on misidentifi (1962) based on specimen collected from outside the monument. of herbarium based on misidentifi herbarium based on misidentifi of batemannii & Neese 1986). Type locality is Thompson’s Springs, Grand Co., UT (Jones s.n. POM). Holmgren (1962) probably based on misidentifi of UT CO Comments FalsRep X Reported by FalsRep X Report from DINO FalsRep X Reported by Holmgren FalsRep X Report from DINO FalsRep X Report from DINO FalsRep X Reported by Park status Periph Wide Wide NA NA RegEn Range Fern Wide/ AnnF Periph/ PerF Wide/ PerF RegEn/ PerF Wide/ PerF RegEn/ Life form Suksdorf’s Suksdorf’s cliffbrake broadleaf knotweed American bistort Gray’s wild Gray’s buckwheat matted wild buckwheat pretty wild buckwheat Common name ana P. suksdorfi P. Synonyms/ notes Taxonomic simplex Pellaea glabella ssp. Polygonum minimum Bistorta bistortoides Species name orum simplex Pellaea glabella ssp. Polygonum minimum Polygonum bistortoides Eriogonum villifl Eriogonum caespitosum Eriogonum bicolor Eriogonum bicolor Utah Colorado Polypodiaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Family Polygonaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

132 Annotated Checklist of Vascular Flora: Dinosaur National Monument c Native .Native to ed specimen ed ed specimen ed P. biennis P. P. anserina. P. ower of Colorado. (2002) based on misidentifi of Europe and possibly E North America. (1962) based on specimen collected from outside the monument. to northern North America and Pacifi Coast states. of (1962) based on specimen collected from outside the monument. herbarium based on misidentifi (1962) based on specimen collected from outside the monument. State fl (1962) based on specimen collected from outside the monument. Native to Asia. UT CO Comments FalsRep X Reported by Naumann FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren Park status Intro Wide Wide Wide Intro Range AnnF Intro/ Shrub Wide/ PerF Wide/ PerF NA/NA FalsRep X Report from DINO PerF Wide/ PerF Intro/ Life form c Norwegian cinquefoil shrubby cinquefoil Fendler’s Fendler’s meadow-rue Pacifi silverweed Colorado columbine Oriental clematis Common name

ssp. & ssp. oribunda, D. oribunda, oribunda monspeliensis P. norvegica P. fl fl Pentaphylloides fruticosa, Dasiphora fruticosa Potentilla egedii A. caerulea, includes vars. ochroleuca pinetorum Synonyms/ notes Taxonomic oribunda Potentilla norvegica Pentaphylloides fl Thalictrum fendleri Aquilegia coerulea Species name Potentilla norvegica Potentilla fruticosa Thalictrum fendleri Aquilegia coerulea Clematis orientalis Viticella orientalis Utah Colorado Rosaceae Rosaceae Rosaceae Ranunculaceae (Thalictraceae) Ranunculaceae (Helleboraceae) Family Ranunculaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

Chapter 6: Appendices 133 var. var. ed Fendlerella . . ed specimen ed ed specimen ed ed specimen ed . C. exilis P. ovina P. S. eriocephala (1962) based on specimen collected from outside the monument. (1962) and Naumann (2002) based on misidentifi of (1962) based on specimen collected from outside the monument. Type locality is upper Bear near the River Valley Uintas, Summit Co., UT 811 NY). (Watson herbarium based on misidentifi specimen of utahensis (1962) and Naumann (2002) based on misidentifi of watsonii. State University herbarium based on misidentifi of UT CO Comments FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Report from DINO FalsRep X Reported by Holmgren FalsRep X Report from Colorado Park status Wide Wide Wide Wide Wide Wide Range PerF Wide/ PerF Wide/ PerF Wide/ PerF Wide/ Life form rhexia-leaved paintbrush scarlet paintbrush yellow paintbrush Fendlerbush Shrub Wide/ Bebb’s willowBebb’s Shrub Wide/ redstem cinquefoil Common name ava fl var. ava Synonyms/ notes Taxonomic avafl C. Castilleja rhexifolia Potentilla rubricaulis

Species name avaﬂ Castilleja var. rhexifolia Castilleja rhexifolia Castilleja miniata Castilleja miniata Castilleja ﬂ Fendlera rupicola Fendlera falcata Salix bebbiana Salix bebbiana Potentilla rubricaulis Utah Colorado Scrophulariaceae Scrophulariaceae Scrophulariaceae Saxifragaceae (Hydrangeaceae) Salicaceae Family Rosaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

134 Annotated Checklist of Vascular Flora: Dinosaur National Monument

var.

var. var. var. . var. ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimen ed ed specimens ed . P. procerus P. P. humilis P. T. latifolia T. P. scariosus P. Zannichellia P. angustifolius P. (1962) based on specimen collected from outside the monument. humilis of procerus. (1962) based on misidentifi (1962) based on misidentifi of (1962), but DINO is well outside the expected range of this species (Cronquist et al. 1984) (1962) based on misidentifi of (1962) based on misidentifi of cyanomontanus. of palustris. Herbarium based on a misidentifi (1962) based on misidentifi of vernalensis. UT CO Comments FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren Ques? X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X Report from BYU FalsRep X Reported by Holmgren Park status Wide Wide RegEn RegEn Wide RegEn Wide RegEn Range PerF Wide/ PerF Periph/ PerF RegEn/ PerF NA/ PerG NA/ PerF RegEn/ PerF Wide/ PerF RegEn/ Life form thyme-leaved speedwell Rydberg’s Rydberg’s penstemon Moffat Moffat penstemon Osterhout’s Osterhout’s beardtongue narrowleaf cattail bluestem penstemon emersed burr- reed Red Desert penstemon Common name

var. ssp. Veronica Veronica serpyllifolia humifusa stoloniferum S. simplex, erectum Synonyms/ notes Taxonomic

ssp. Veronicastrum Veronicastrum serpyllifolium humifusum Penstemon rydbergii Penstemon moffatii Penstemon osterhoutii Penstemon cyanocaulis Penstemon arenicola Sparganium emersum

Species name var. Sparganium Veronica Veronica serpyllifolia Penstemon rydbergii rydbergii Penstemon moffatii Typha angustifoliaTypha angustifolia Typha Penstemon cyanocaulis Penstemon arenicola emersum Utah Colorado Sparganiaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Typhaceae Scrophulariaceae Family Scrophulariaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont.

Chapter 6: Appendices 135 . Native . ed specimen ed ed specimen ed C. reticulata T. latifolia T. (1962) based on specimen collected from outside the monument. (1962) based on specimen collected from outside the monument. Type locality is Uinta Mountains, Summit Co., 458 GH) UT (Watson (1962) based on misidentifi of to E. North America. of (2002) based on misidentifi UT CO Comments FalsRep X Reported by Holmgren FalsRep X Reported by Holmgren FalsRep X X Reported by Holmgren FalsRep X Reported by Naumann Park status Wide Wide Wide Wide Range nd Neese 1986; Holmgren 1962; Naumann 2002; NCPN, in ed.; and Weber 1962; Naumann 2002; NCPN, in ed.; and Weber nd Neese 1986; Holmgren PerF Wide/ PerF Wide/ Tree Intro/ Life form cations (False Reports or FalsRep) because the monument is well outside known expected western valerian pinnate angelica common hackberry southern cattail PerG Wide/ Common name Synonyms/ notes Taxonomic Valeriana Valeriana occidentalis Typha Typha domingensis Species name Valeriana Valeriana occidentalis Angelica pinnata Angelica pinnata Celtis occidentalis Celtis occidentalis Typha Typha domingensis Utah Colorado Valerianaceae Umbelliferae (Apiaceae) Ulmaceae (Celtidaceae) Family Typhaceae Appendix C. Rejected (falsely reported or questionable) taxa of Dinosaur National Monument, cont. The vascular plant species listed here have been reported for Dinosaur NM from published and unpublished literature (Goodrich a published and unpublished literature for Dinosaur NM from have been reported The vascular plant species listed here because of misidentifi now believed to be erroneous herbarium specimens, but are et al. 1982) or from range of the species (Questionable or Ques?).

136 Annotated Checklist of Vascular Flora: Dinosaur National Monument X X X X X X X X Wet Comments X X Mt Con XX X PJ XXX XXX XXX XXX Shr Des Flower time Park status Native Pres Apr–May Native Pres Apr–Jun Native Pres Apr–Jun Native Pres Mar–Jun Native Pres Mar–Jun Native Pres Apr–Jun Native Hist Apr–May Native Pres NA Native Pres Apr–Jun Native Pres NA r Native Pres NA netleaf hackberry peach-leaf willow lance-leaf cottonwood quaking aspen Native Pres Mar–Jun X X Rio Grande cottonwood Fremont cottonwood squawbush Native Pres May–Jul ponderosa pine NativeDouglas-ﬁ Pres NA narrow-leaved cottonwood Harriman’s Harriman’s yucca Russian olive Introtwo-needle pinyon Rep May–Jul Rocky Mountain maple water birchRocky Mountain juniper Native Pres Apr–May X box elder Native Pres Apr–Jun Siberian elm Intro Rep Mar–Apr X X X Common name Nativity

ssp. var. ssp. acuminata x interius wislizenii Populus deltoides Populus fremontii Rhus aromatica trilobata Pinus ponderosa scopulorum Pseudotsuga menziesii Yucca harrimaniae Yucca angustifolia Acer glabrum ssp.

var. var. var.

var. Species name var. var. intercurrens Populus x glauca Celtis reticulata Celtis reticulata Salix amygdaloides Salix amygdaloides Populus Populus tremuloides Populus tremuloides Populus fremontii wislizeni Populus fremontii fremontii Rhus aromatica trilobata Pinus ponderosa scopulorum Pseudotsuga menziesii var. Populus angustifolia Populus angustifolia Yucca harrimaniae Yucca harrimaniae Elaeagnus angustifolia Elaeagnus Pinus edulis Pinus edulis glabrum Utah Colorado Juniperus scopulorum Sabina scopulorum Betula occidentalis Betula fontinalis Acer negundo Negundo aceroides Acer glabrum Ulmus pumila Ulmus pumila taxa of Dinosaur National Monument, organized by life form with ecological notes. rmed, historical, and reported Ulmaceae (Celtidaceae) Salicaceae Salicaceae Salicaceae Salicaceae Salicaceae Anacardiaceae Pinaceae Pinaceae Salicaceae Agavaceae Elaeagnaceae Pinaceae Cupressaceae Betulaceae Family Trees Aceraceae Shrubs Aceraceae Ulmaceae Appendix D. Conﬁ

Chapter 6: Appendices 137 Saline clays Sandy soils X X X Wet Comments Mt Con PJ X XX XXX XX XXX X XXX X X XX XX X Shr Des Flower time Park status Native Pres May–Sep Native Pres May–Sep Native Pres Jun–Aug Native Hist Jun–Aug Native Pres May–Jul Native Hist May–Jun Native Hist Jun–Aug ower basin saltbush Native Rep Jun–Sep Castle Valley Castle Valley saltbush shadscalemat saltbush Native Native Pres Pres May–Sep Apr–Sep X X X Clayey soils Morrison F four-wing four-wing saltbush mountain-lover Native Pres May–Aug X X X mountain snowberry western snowberry thinleaf alder Nativeblue elderberry Pres Nativelong-ﬂ Pres Apr–May snowberry May–Jul Oregon-grape Native Pres Apr–Jun X X X X low sagebrush Native Rep Jun–Sep western poison ivy spiny hopsagewinterfat Native Presgreasewood Native Apr–Jul Native Plummer’s Pressiltbush X Pres May–Jul May–Aug X Common name Nativity

. ssp. var orus Atriplex gardneri Atriplex canescens Symphoricarpos rotundifolius Symphoricarpos occidentalis tenuifolia Symphoricarpos longiﬂ Alnus incana arbusculum arbusculum Toxicodendron Toxicodendron rydbergii Krascheninnikovia lanata Sarcobatus vermiculatus Atriplex brandegei

var. var. var. var.

Species name

var. var. lanata var. orus Atriplex gardneri utahensis Atriplex corrugataAtriplex gardneri cuneata Atriplex corrugata Atriplex confertifolia Atriplex confertifolia Atriplex canescens canescens Pachystima myrsinites Paxistima myrsinites utahensis Symphoricarpos oreophilus Symphoricarpos occidentalis Sambucus caeruleaSymphoricarpos Sambucus coerulea longiﬂ Alnus incana occidentalis Mahonia repens Mahonia repens Artemisia arbuscula Seriphidium Utah Colorado Toxicodendron Toxicodendron rydbergii Grayia spinosaKrascheninnikovia lanata Atriplex grayi Sarcobatus vermiculatus Zuckia brandegei plummeri rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Celastraceae Caprifoliaceae Caprifoliaceae Caprifoliaceae (Adoxaceae) Caprifoliaceae Betulaceae Berberidaceae Compositae (Asteraceae) Family Shrubs, cont. Anacardiaceae Chenopodiaceae Chenopodiaceae (Sarcobataceae) Chenopodiaceae Appendix D. Conﬁ

138 Annotated Checklist of Vascular Flora: Dinosaur National Monument Sandstone Sandy to saline soils Clayey to gravelly saline soils Rocky areas X Wet Comments Mt Con PJ XX XXX XXX XXX XXX XX XXX X XXX XXX XX XXX XX XX XXX Shr Des Flower time Park status Native Pres Aug–Oct Native Pres Jul–Sep Native Pres Aug–Oct Native Pres Jul–Sep Native Pres Aug–Oct Native Pres Jul–Sep Native Pres Aug–Oct Native Pres Jul–Sep Native Pres Jul–Sep Native Pres Jul–Sep Native Pres Aug–Oct Native Hist Jun–Aug rough brickellbush Wyoming big sagebrush smoothseed rabbitbrush mountain big sagebrush glabrate rabbitbrush basin big sagebrush greenish rabbitbrush black sagebrush Native Presbudsage Sep–Oct Native Pres Apr–Jun spear-leaf spear-leaf rabbitbrush silver sagebrush Native Rep Aug–Sep fringed sagebrush dwarf rabbitbrush Bigelow’s Bigelow’s sagebrush Mohave brickellbush Common name Nativity

ssp. ssp. ssp. ssp. ssp. viscidulum scabra Brickellia microphylla var. wyomingense Seriphidium tridentatum Chrysothamnus nauseosus leiospermus Seriphidium vaseyanum Chrysothamnus nauseosus graveolens tridentatum Chrysothamnus nauseosus consimilis Seriphidium tridentatum Seriphidium novum desertorum Chrysothamnus linifolius Seriphidium canum ssp. Chrysothamnus depressus Brickellia oblongifolia

Species name var. var.

var. var. var. linifolia scabra wyomingensis vaseyana tridentata Brickellia oblongifolia var. Brickellia microphylla var. Artemisia tridentata var. Chrysothamnus nauseosus leiospermus Artemisia tridentata var. Chrysothamnus nauseosus graveolens Chrysothamnus nauseosus consimilis Artemisia tridentata var. Artemisia nova nova Artemisia spinescens Picrothamnus Chrysothamnus linifolius Artemisia cana viscidula Artemisia frigida Artemisia frigida Chrysothamnus depressus Utah Colorado Artemisia bigelovii Artemisia bigelovii rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ Family Shrubs, cont. Compositae (Asteraceae)

Chapter 6: Appendices 139 Glen Canyon, Moenkopi, & Mowry F. X X Wet Comments X Mt Con PJ XX XX XX XXX X XXX XXX XXX XX XX XXX XXX Shr Des Flower time Park status Native Pres Apr–Jun Native Hist May–Jun Native Pres Jun–Sep Native Pres Jul–Oct Native Pres Jul–Sep Native Pres Jul–Sep Native Pres NA Native Hist Apr–May Native Pres Jul–Sep Native Pres NA Native Pres Mar–Jun Native PresNative Jul–Sep Pres Jul–Sep Native Pres May–Jul shortspine horsebrush Nuttall’s Nuttall’s horsebrush spineless horsebrush broom snakeweed orchard snakeweed green ephedra Native Pres NA green rabbitbrush Torrey’s Torrey’s ephedra Utah junipersilver Nativebuffaloberry Pres NA X X X X slenderleaf rabbitbrush common juniper spiny greasebush longbract rabbitbrush Nevada rabbitbrush red-osier dogwood Common name Nativity

ssp. ssp. orus orus orus alpina howardii Ephedra viridis viscidifl Chrysothamnus viscidifl stenophyllus Juniperus communis ssp. Chrysothamnus viscidifl Forsellesia meionandra Chrysothamnus parryi ssp. Swida sericea

Species name

var. var.

var. var. var. orus orus orus depressa nevadensis howardii Cornus sericea Tetradymia spinosaTetradymia spinosa Tetradymia Tetradymia nuttalliiTetradymia nuttallii Tetradymia Tetradymia canescensTetradymia canescens Tetradymia Gutierrezia sarothrae Gutierrezia sarothrae Gutierrezia pomariensis Ephedra viridis viridis Ephedra torreyana Ephedra torreyana Chrysothamnus viscidifl viscidifl Shepherdia argentea Shepherdia argentea Juniperus osteosperma Sabina osteosperma Juniperus communis var. Chrysothamnus parryi var. Chrysothamnus viscidifl stenophyllus Glossopetalon spinescens meionandrum Utah Colorado Chrysothamnus parryi var. sericea rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Cornaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Ephedraceae Ephedraceae Compositae (Asteraceae) Elaeagnaceae Cupressaceae Cupressaceae Compositae (Asteraceae) Compositae (Asteraceae) Crossosomataceae Family Shrubs, cont. Compositae (Asteraceae) Appendix D. Confi

140 Annotated Checklist of Vascular Flora: Dinosaur National Monument Rock crevices Moenkopi & Mowry F. Rock crevices Rocky sites Sandy soil X Wet Comments Mt Con XX XX PJ XXX XXX X XXX XXX XXX XX XXX XXX XXX XX Shr Des Flower time Park status Native Pres Apr–Jun Native Pres Apr–Jun Native Rep May–Jul Native Pres Jun–Oct Native Pres Jun–Oct Native Pres Jun–Aug Native Pres Jul–Oct Native Pres May–Jun Native Pres Mar–Jul Native Pres May–Jun curl-leaf mountain dwarf mountain mahogany Utah serviceberry deer-brushSaskatoon serviceberry Native Pres Jun–Jul X X slender wild buckwheat Dinosaur wild buckwheat dwarf ninebark Native Pres May–Aug X X X Rock crevices bush oceanspray river hawthorn Native Rep May–Jun prickly-phlox Native Pres May–Jul singleleaf ash Native Pres Apr–May X X X crisp-leaf wild buckwheat alder-leaf alder-leaf mountain mahogany greenleaf manzanita Cutler’s ephedraCutler’s Native Hist NA mahogany Common name Nativity

var. var. orum . platyphylla Cercocarpus ledifolius utahensis Eriogonum microthecum laxiﬂ Eriogonum saurinum Crataegus rivularis pungens Eriogonum corymbosum corymbosum montanus ssp Ephedra cutleri

var.

Species name var. var. var. var. orum intermontanus Cercocarpus ledifolius var. Cercocarpus intricatus Cercocarpus intricatus Amelanchier utahensis Amelanchier Amelanchier alnifolia Amelanchier alnifolia Ceanothus velutinus Ceanothus velutinus Eriogonum microthecum laxiﬂ saurinum Physocarpus alternans Physocarpus alternans Eriogonum lonchophyllum Holodiscus dumosus Holodiscus discolor Crataegus douglasii rivularis Leptodactylon pungens Leptodactylon Fraxinus anomala Fraxinus anomala Eriogonum corymbosum corymbosum Cercocarpus montanus Cercocarpus Utah Colorado Arctostaphylos patula Arctostaphylos patula Ephedra viridis viscida rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Rosaceae Rosaceae Rosaceae Rosaceae Rhamnaceae Polygonaceae Rosaceae Polygonaceae Rosaceae Rosaceae Polemoniaceae Oleaceae Polygonaceae Rosaceae Ericaceae Appendix D. Conﬁ Family Shrubs, cont. Ephedraceae

Chapter 6: Appendices 141 X X X X X X X Wet Comments XX X Mt Con XX PJ XXX X XXX XXX XXX XXX Shr Des Flower time Park status Native Pres May–Sep Native Rep May–Jun Native Pres Jun–Aug Native Pres Apr–Jul Native Hist Jun–Sep Native Pres May–Jul Intro Pres Apr–Sep ve-stamen golden currant Native Pres Mar–Jul Scouler’s willowScouler’s NativeUtah fendlerella Native Pres Preslittleleaf mock- Apr–Junorange Apr–Sep X X whiplash willow Native Presmountain willow Apr–Jun coyote willow Native Pres Apr–Aug yellow willow Native Pres Apr–Jun common red raspberry Nootka rose Nativetea rose roseWoods’ Pres Jun–Jul Native Intro Pres Pres May–Jul May–Jul X X Cultivated antelope bitterbrush water-plantain Nativenorthern Presarrowhead Jun–Sep western chokecherry ﬁ tamarisk wax currant Native Pres Apr–Aug Common name Nativity

ssp.

ssp. ssp. Philadelphus microphyllus caudata Salix exigua Salix lutea melanolasius Rubus idaeus Rosa nutkana Alisma triviale melanocarpa Padus virginiana Ribes cereum

var.

Species name

caudata Salix lucida var. var. var. var. ssp. Ribes aureum Ribes aureum Fendlerella utahensis Fendlerella utahensis Salix scouleriana Salix scouleriana Philadelphus microphyllus Salix monticola Salix monticola Salix exigua stenophylla Salix lucida Salix eriocephala watsonii Rosa woodsiiRubus idaeus strigosus Rosa woodsii Rosa nutkana hispida Rosa odorata Rosa odorata Purshia tridentata Purshia tridentata Alisma plantago- aquatica Sagittaria cuneata Sagittaria cuneata Utah Colorado Tamarix chinensisTamarix ramosissima Tamarix inebrians Prunus virginiana melanocarpa Ribes cereum rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Saxifragaceae (Grossulariaceae) Saxifragaceae (Hydrangeaceae) Salicaceae Saxifragaceae (Hydrangeaceae) Salicaceae Salicaceae Salicaceae Salicaceae Rosaceae Rosaceae Rosaceae Rosaceae Rosaceae Perennial Forbs Perennial Alismataceae Alismataceae Tamaricaceae Family Shrubs, cont. Rosaceae Saxifragaceae (Grossulariaceae) Appendix D. Conﬁ

142 Annotated Checklist of Vascular Flora: Dinosaur National Monument Disturbed Moenkopi F. Bishop, Moenkopi, & Morrison Disturbed areas areas Bishop, Chinle Glen Canyon, & Moenkopi F. X X X Wet Comments Mt Con PJ XXX X XXX XX XXX XXX XXX XXX XXX XXX XX XXX XXX Shr Des Flower time Park status Intro Rep May–Jul Native Pres May–Jun Native PresNative Apr–Jun Pres Apr–Jun Native Pres May–Jun Native Pres May–Aug Native Pres May–Jun Native Pres Jun–Aug Native Pres Apr–Jun Native Pres May–Jul Native Pres Apr–Jul Native Pres Jun–Aug common hound’s-tongue erect cryptanth Native Pres May–Jun X X X dwarf cryptanth Native PresRollins’ cryptanth Apr–Jun plateau yellow cryptanth yellow-eye cryptanth Uinta Basin cryptanth showy milkweed spindle-rooted bluebell pallid milkweed Native Presswamp milkweed Apr–Jun western stoneseed spider milkweed showy stoneseed showy stickseed Native Pres Jun–Aug X X X Jones’ amsonia Native Prescommon Apr–Jun dogbane Common name Nativity

ssp. ssp. ora ava avoculata oribunda cinale ofﬁ nana Asclepias cryptoceras ruderale Asclepias asperula asperula incisum cannabinum

var. orabrevifl Oreocarya Species name cinale Cynoglossum ava avoculatafl fl Oreocarya Oreocarya oribundafl Hackelia cryptoceras Cynoglossum offi Cryptantha stricta Oreocarya stricta Cryptantha fl Cryptantha fl Cryptantha humilisCryptantha rollinsii Oreocarya humilis Oreocarya rollinsii Cryptantha brevifl Asclepias speciosa Asclepias speciosa Mertensia fusiformis Asclepias cryptoceras var. Asclepias incarnata Asclepias incarnata Lithospermum ruderale Lithospermum Asclepias asperula asperula Utah Colorado Lithospermum incisum Lithospermum Hackelia fl Amsonia jonesii Amsonia jonesii Apocynum cannabinum Apocynum rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Asclepiadaceae Boraginaceae Asclepiadaceae Asclepiadaceae Boraginaceae Asclepiadaceae Boraginaceae Boraginaceae Appendix D. Confi Family Perennial Forbs, cont. Apocynaceae Apocynaceae

Chapter 6: Appendices 143 scree areas X Wet Comments Mt Con PJ XXX XXX XXX XX XX XXX XXX X XXX XXX XXX XX XXX XXX Shr Des Flower time Park status Native Pres May–Aug Native Pres May–Jul Native Pres May–Jun Native Pres May–Jun Native Pres Jun–Aug Native Hist May–Jun Native Pres Jun–Jul Native Rep May–Aug and Talus Native Pres Apr–May Native Pres Apr–Aug Native Pres Jun–Aug Native Pres May–Sep eld chickweed eld Native Pres Apr–Jun Eastwood’s Eastwood’s ball-head sandwort common hops Native Pres Jul–Aug Simpson’s Simpson’s hedgehog cactus greenmolly Native Pres Aug–Oct X X plains pricklypear slenderspine pricklypear ﬁ James’ chickweed claretcupUtah Nativepricklypear Pres Apr–Jun brittle pricklypear bouncing-bet Intro Pres May–Sep Disturbed Nuttall’s Nuttall’s sandwort western bluebell Hooker’s Hooker’s sandwort lanceleaf bluebell sandwort Common name Nativity

ssp. var. cinalis minor Eremogone eastwoodiae Eremogone congesta Humulus lupulus americanus var. Opuntia polyacantha jamesiana Echinocereus triglochidiatus brachyartha oblongifolia Eremogone hookeri Mertensia lanceolata

var.

var. var. var. var. Species name var. cinalisoffi Saponaria trichophora polyacantha coriacea Arenaria fendleri eastwoodiae Arenaria congesta congesta Humulus lupulus neomexicanus Opuntia polyacantha var. Pediocactus simpsonii Pediocactus simpsonii Bassia americana Kochia americana Opuntia fragilisOpuntia polyacantha var. Opuntia fragilis Cerastium arvenseSaponaria offi Cerastium strictum Stellaria jamesiana Pseudostellaria Echinocereus triglochidiatus melanacanthus Opuntia erinacea utahensis Arenaria nuttallii Minuopsis nuttallii Mertensia oblongifolia Mertensia Arenaria hookeri desertorum Utah Colorado Mertensia lanceolata var. rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Cannabaceae Cactaceae Cactaceae Chenopodiaceae Cactaceae Cactaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae Caryophyllaceae (Alsinaceae) Cactaceae Cactaceae Caryophyllaceae (Alsinaceae) Boraginaceae Caryophyllaceae (Alsinaceae) Family Perennial Forbs, cont. Boraginaceae Appendix D. Confi

144 Annotated Checklist of Vascular Flora: Dinosaur National Monument Disturbed areas X X X Wet Comments X Mt Con PJ XX XXX XXX XXX X XXX XXX X XXX X XXX XX XXX XXX X XX Shr Des Flower time Park status Native Pres May–Jul Native Pres Jun–Aug Native Pres Jun–Oct Native Pres Jul–Oct Native Pres May–Sep Native Pres Aug–Oct c aster c Native Pres Jul–Oct heartleaf arnica Native Pres May–Aug burdock Intro Rep Jul–Sep common pussytoes low pussytoes Nativesmall-leaf Prespussytoes Apr–Jun tall pussytoes Native Pres Jun–Aug Paciﬁ orange agoseris Native Pres Jun–Aug skeleton-leaf bursage tarragonLouisiana Nativewormwood Pres Jul–Oct pale agoseris Nativecutleaf agoseris Native Pres Pres May–Sep May–Sep common yarrow orange arnica Nativebiennial Histwormwood May–Aug Common name Nativity

var. var.

ssp. ssp. ludoviciana aurantiaca Arnica cordifolia microphylla Antennaria pulcherrima anaphaloides Agoseris aurantiaca var. Agoseris glauca Agoseris glauca Oligosporus dracunculus glaucus Artemisia ludoviciana ssp. glauca laciniata Achillea lanulosa

var.

var. Species name var. var. glauca lanulosa ludoviciana Arnica cordifolia cordifolia Arctium minus Arctium minus Antennaria parvifolia Antennaria parvifolia Antennaria dimorpha Antennaria dimorpha Antennaria microphylla Antennaria Antennaria anaphaloides Aster ascendens Virgulaster ascendens Agoseris aurantiaca aurantiaca Agoseris glauca glauca Agoseris glauca laciniata Ambrosia tomentosa Ambrosia tomentosa Artemisia dracunculus ssp. Artemisia ludoviciana var. Utah Colorado Achillea millefolium ssp. Arnica fulgens Arnica fulgens Artemisia biennis Artemisia biennis rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ Family Perennial Forbs, cont. Compositae (Asteraceae) (Asteraceae) Compositae (Asteraceae)

Chapter 6: Appendices 145 Disturbed areas areas areas Disturbed areas Disturbed areas areas X X X X Wet Comments XX XX Mt Con XX PJ XXX XXX XXX XX XXX XXX XXX XXX X Shr Des Flower time Park status Intro Pres Jun–Sep Intro Rep Jul–Sep Disturbed Intro Rep Jul–Sep Disturbed Native Pres Jul–Oct Native Rep Jun–Aug Native Pres Apr–Jul Native Pres Apr–Jul Native Pres May–Sep Native Pres Aug–Oct Native Pres May–Sep ower Russian knapweed spotted knapweed diffuse diffuse knapweed musk thistle Intro Pres Jun–Sep tassel-ﬂ brickellbush Devil’s- Devil’s- beggarsticks Canada thistle Intro Pres Jul–Sep arrowleaf balsamroot elegant aster Nativewestern aster Pres Nativehairy Jul–Aug Repbalsamroot Jul–Sep chicory Intro Pres Jul–Oct Disturbed rose-heath Native Pres May–Sep hispid golden- aster Siskiyou aster Nativewhite heath Presaster Jul–Sep blueleaf aster Native Pres Jul–Sep hoary dusty- maiden Common name Nativity

ssp. ora var. macrolepis sagittata Balsamorhiza hispidula Heterotheca villosa hesperius

var. oragrandifl Brickellia Species name hispidula Centaurea repens Acroptilon repens Centaurea maculosa Acosta maculosa Centaurea diffusa Acosta diffusa Carduus nutans Carduus nutans Brickellia grandifl Bidens frondosa Bidens frondosa Cirsium arvense Breea arvensis Balsamorhiza sagittata Balsamorhiza Aster perelegansAster spathulatus Eucephalus elegans Balsamorhiza hookeri Aster spathulatus var. Cichorium intybus Cichorium intybus Chaetopappa ericoides Leucelene ericoides Chrysopsis villosa minor Aster pansus Virgulus ericoides Utah Colorado Aster glaucodes Eucephalus glaucus Chaenactis douglasii Chaenactis douglasii Aster hesperius Aster lanceolatus rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Compositae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Perennial Forbs, cont. Compositae (Asteraceae) (Asteraceae) Compositae (Asteraceae) Appendix D. Confi

146 Annotated Checklist of Vascular Flora: Dinosaur National Monument Moenkopi F. Disturbed areas Lodore & F. Weber X X X Wet Comments Mt Con PJ XX X XXX XXX XXX XX XXX XX XX XX X XXX X XXX X XXX Shr Des Flower time Park status Native Pres May–Aug Native Pres May–Jul Native Pres May–Jul Native Pres May–Jul Native Pres May–Jul Native Pres Jun–Sep Native Pres May–Jul Native Pres Jun–Aug eabane Native Rep Jun–Aug nakedstemnoddinghead Native Native Pres Hist May–Jun May–Jun glaucous hawksbeard western hawksbeard Modoc hawksbeard gray hawksbeard bull thistlemountain Introhawksbeard Pres Jul–Aug smooth daisy Native Pres Jun–Aug elk thistle Native Rep Jun–Aug wavy-leaved thistle cut-leaved daisy Native Presspreading daisy Native May–Aug daisyEaton’s Pres Native May–Sep Engelmann’s daisy Pres May–Jul Ownbey’s Ownbey’s thistle tufted ﬂ Common name Nativity runcinata Enceliopsis nudicaulis Psilochenia glauca ssp. Psilochenia occidentalis modocensis intermedia Cirsium scariosum Cirsium undulatum Erigeron compositus Erigeron divergens

var. var. var. var. var. Species name nudicaulis glabratus Enceliopsis nudicaulis var. Enceliopsis nutans Enceliopsis nutans Crepis runcinata glauca Crepis occidentalis occidentalis Crepis modocensis Psilochenia Crepis intermedia Psilochenia Cirsium vulgareCrepis acuminata Cirsium vulgare Psilochenia acuminata Erigeron glabellus Erigeron glabellus Cirsium scariosum scariosum Cirsium undulatum undulatum Erigeron compositus var. Erigeron divergens divergens Erigeron eatoniiErigeron engelmannii Erigeron eatonii Erigeron engelmannii Utah Colorado Cirsium ownbeyi Cirsium ownbeyi Erigeron caespitosus Erigeron caespitosus rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ Family Perennial Forbs, cont. Compositae (Asteraceae)

Chapter 6: Appendices 147 Green River Shale & Weber Sandstone X X X X Wet Comments Mt Con XX PJ XXX XXX XXX XXX XX XXX XX XXX XX XXX XXX Shr Des Flower time Park status Native Pres May–Jun Native Pres May–Jul Native Pres Jul–Sep Native Pres Jun–Sep Native Pres Jul–Oct Native Pres Jul–Sep Native Pres May–Oct Native Pres May–Oct Native Pres Jul–Sep Native Pres May–Jul Native Pres Jul–Sep ower Native Pres Jun–Aug ower eabane Native Pres May–Aug ower ower eabane thrift goldenweed stemless goldenweed curly-cup gumweed common blanketfl western goldenrod Uinta daisyspotted joe-pye Nativeweed Pres Jul–Aug Navajo fl basin daisy Native Hist May–Jun Wyoming hyalineherb common hyalineherb Nuttall’s Nuttall’s sunfl little sunfl needle-leaf fl purpledisk sunfl fall sneezeweed Native Pres Jul–Sep Common name Nativity ora luteus cinereus var. var. serrulata montanum lifolius lifolius Stenotus armerioides Stenotus acaulis Grindelia squarrosa var. Erigeron uintahensis Eupatorium maculatum Erigeron concinnus Hymenopappus fi Hymenopappus fi Erigeron wilkenii Helianthella microcephala Helenium autumnale var.

var. var. var. Species name orauniﬂ Helianthella

var. luteus cinereus var. var. acaulis bruneri montanum lifolius lifolius Helenium autumnale Haplopappus armerioides armerioides Haplopappus acaulis var. Grindelia squarrosa serrulata Gaillardia aristata Gaillardia aristata Euthamia occidentalis Euthamia occidentalis Erigeron speciosus uintahensis Eupatorium maculatum var. concinnus Hymenopappus fi Hymenopappus fi Erigeron pulcherrimus Erigeron pulcherrimus Erigeron pumilus Helianthella unifl Helianthus nuttallii Helianthus nuttallii Utah Colorado Erigeron nematophyllus Helianthella microcephala var. rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Perennial Forbs, cont. Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi

148 Annotated Checklist of Vascular Flora: Dinosaur National Monument Weber F. Weber areas X Wet Comments Mt Con XX PJ XXX XXX XXX XXX X XXX XXX XX XXX XX XXX XXX XXX Shr Des Flower time Park status Native Pres May–Jun Native Pres May–Jul Native Pres May–Aug Native Pres May–Aug Native Pres May–Jul Native Pres Apr–Jun Native Hist Apr–Aug hoary aster Native Pres May–Sep showy rush- pink blue lettuce Native Pres Jun–Sep poverty-weed Native Pres May–Sep Torrey’s Torrey’s hymenoxys rock hymenoxys Native PresColorado rubber-plant Jun rock-goldenrod NativeUintah bahia Pres Native Jul–Sep woolly Presgroundsel May–Jun lanate hymenoxys Scotch thistle Intro Rep Jun–Sep Disturbed Arizona hymenoxys nodding microseris gumweed aster Native Pres May–Jul Common name Nativity

var.

var. ora Machaeranthera grindelioides Machaeranthera canescens Lygodesmia Lygodesmia grandiﬂ Lactuca tatarica pulchella Picradenia richardsonii richardsonii Petradoria pumila Tetraneuris brevifolia Tetraneuris acanthium Tetraneuris acaulis Tetraneuris

ora var. var. var.

Species name var.

var. var. ora var. grandifl richardsonii Machaeranthera canescens Machaeranthera grindelioides grindelioides canescens Lygodesmia grandifl Lygodesmia var. pulchella Iva axillaris Lactuca tatarica Iva axillaris Hymenoxys torreyana torreyana Tetraneuris Hymenoxys lapidicola Hymenoxys lapidicola Hymenoxys richardsonii var. Petradoria pumila pumila Platyschkuhria integrifolia ourolepis Senecio canus Packera cana Hymenoxys acaulis caespitosa Onopordum acanthium Onopordum Utah Colorado Hymenoxys acaulis arizonica Microseris nutans Microseris nutans rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Perennial Forbs, cont. Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi

Chapter 6: Appendices 149 Disturbed areas Sandy soils X X X X Wet Comments X Mt Con PJ X XXX XXX X XXX X XX XXX X XXX XXX XXX XX Shr Des Flower time Park status Native Rep Jun–Jul Intro Pres Jul–Oct Native Pres Aug–Oct Native Pres Jul–Oct Native Pres Aug–Oct Native Pres Jul–Aug Intro PresIntro Mar–Dec Pres Apr–Dec Native Pres Jun–Aug Native Pres Jun–Sep Native Pres Jun–Oct Native Pres Jun–Aug Native Pres Jul–Aug eld sow-thistle eld Intro Rep Jul–Oct silver chicken- sage marsh sowthistle ﬁ baby goldenrod Native Presthree-nerve goldenrod Aug–Sep Missouri goldenrod Canadian goldenrod montane groundsel red-seed dandelion common dandelion Rocky Mountain groundsel slender wirelettuce basin groundsel Nativebroom Presgroundsel Apr–Aug western groundsel desert wirelettuce Common name Nativity cinale Sphaeromeria argentea Solidago canadensis Packera werneriifolia laevigatum Packera streptanthifolia Stephanomeria tenuifolia Senecio spartioides Senecio integerrimus Stephanomeria runcinata

var. var. Species name cinaleofﬁ Taraxacum

var. salebrosa werneriifolius exaltatus Sonchus uliginosus Sonchus uliginosus Sonchus arvensis Sonchus arvensis Solidago nanaSolidago velutina Solidago nana Solidago velutina Solidago missouriensis Solidago missouriensis Solidago canadensis var. Taraxacum ofﬁ Taraxacum Senecio werneriifolius var. Taraxacum laevigatumTaraxacum Taraxacum Senecio streptanthifolius streptanthifolius tenuifolia spartioides Senecio multilobatus Packera multilobata Senecio spartioides Stephanomeria tenuifolia Utah Colorado Senecio integerrimus var. Stephanomeria runcinata rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Family Perennial Forbs, cont. Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Conﬁ

150 Annotated Checklist of Vascular Flora: Dinosaur National Monument areas Saline & seleniferous soils Sandy soils Disturbed areas Wet Comments X Mt Con PJ X XXX XXX X XX XX XX XXX X XX XXX XXX XX XXX XX XXX XX Shr Des Flower time Park status Native Pres Apr–Jun Native Pres May–Aug Native Pres Jun–Sep Native Hist May–Jun Native Pres Apr–Jun Native Pres Jun–Jul Native Hist Jun–Jul Native Pres May–Sep Native Pres Apr–Jul Native Pres May–Aug Native Hist Apr–Jun Native Pres Jun–Aug eld bindweed eld Intro Pres Jun–Aug Disturbed lance-leaved stonecrop ﬁ Cisco woodyaster rough mule’s- ears showy goldeneye yellow salsify Intro Pres May–Jul hairy townsendia plateau townsendia tower-mustard Nativehairy rockcress Pres Native May–Sep Pres Jun–Aug spreadingpod rockcress common townsendia Drummond’s Drummond’s rockcress hoary townsendia nodding rock cress Hopi-tea Native Pres Jun–Aug Hooker’s Hooker’s townsendia Common name Nativity

var. ora ora multiﬂ major lanceolatum Heliomeris multiﬂ var. var. Townsendia strigosa Townsendia Boechera divaricarpa demissa Thelesperma megapotamicum

var. var. Species name ora ora strigosa Sedum lanceolatum Amerosedum Convolvulus arvensis Convolvulus arvensis Xylorhiza venusta Xylorhiza venusta Wyethia scabra Wyethia scabra Viguiera multifl multifl Townsendia strigosa Townsendia var. dubiusTragopogon dubius Tragopogon Townsendia mensanaTownsendia mensana Townsendia Arabis glabraArabis hirsuta glabra Arabis holboellii Turritis Arabis hirsuta brachycarpa Townsendia leptotesTownsendia leptotes Townsendia Arabis drummondii Boechera drummondii Townsendia incanaTownsendia incana Townsendia Utah Colorado Arabis demissa Boechera demissa Thelesperma megapotamicum Townsendia hookeriTownsendia hookeri Townsendia rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Crassulaceae Convolvulaceae Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Family Perennial Forbs, cont. Compositae (Asteraceae) Compositae (Asteraceae) Appendix D. Confi

Chapter 6: Appendices 151 areas areas Glen Canyon F. & Weber Weber F. Weber Wet Comments Mt Con XX PJ XXX XXX XXX XX XXX XX XX XX XXX XXX XXX XXX XX XXX Shr Des Flower time Park status Native Pres Jun–Sep Intro Pres May–Aug Disturbed Native Pres Mar–Jul Native Pres Apr–Jun Native Pres Apr–Aug Native Pres May–Jul Native Pres Apr–Aug Native Pres May–Jul Native Pres May–Aug Native Pres Apr–Aug ower Native Pres Apr–Jul ower ower spindlestemCalifornia tansy- Nativemustard Pres May–Jul hairy whitetop Intro Pres May–Sep Disturbed orbicular whitetop desert rockcress Native rockcressSelby’s Pres Native Mar–Jun Prespark rockcress Native May–Aug Pres May perennial rockcress Nuttall’s Nuttall’s rockcress shy wallfl sand dune wallfl Sweetwater rockcress western wallfl Holboell’s Holboell’s rockcress Uinta drabafew-seed Nativewhitlow-grass Pres May–Jun Fendler’s Fendler’s rockcress Common name Nativity

ssp. Caulanthus crassicaulis californica Boechera pulchra pallens Boechera perennans Erysimum inconspicuum Erysimum asperum Erysimum capitatum Boechera retrofracta Draba juniperina Draba oligosperma Boechera fendleri

var. var. var. var.

var. var. var. Species name var. crassicaulis Caulanthus crassicaulis var. Descurainia californica Descurainia Cardaria pubescens Cardaria pubescens Cardaria chalepensis Cardaria chalepensis Arabis pulchra pallens Arabis selbyiArabis vivariensis Boechera selbyi Boechera fernaldiana Arabis perennans perennans Arabis nuttallii Boechera nuttallii Erysimum inconspicuum Erysimum asperum purshii Arabis lignifera Boechera lignifera Erysimum asperum elatum Arabis holboellii secunda Utah Colorado juniperina Draba oligosperma oligosperma Arabis holboellii fendleri Draba oligosperma rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Family Perennial Forbs, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix D. Conﬁ

152 Annotated Checklist of Vascular Flora: Dinosaur National Monument Often on seleniferous soils Disturbed areas X X X X X X Wet Comments Mt Con PJ XX XXX XXX XXX X XXX XX XXX XXX XXX Shr Des Flower time Park status Native Hist Apr–Sep Native Hist Jun–Sep Native Hist Jun–Sep Native Pres May–Jul Native Pres May–Jun Native Pres Jun–Sep Native Pres May–Jul Native Pres Apr–Jul Native Pres Jun–Aug Native Pres Apr–Jul Native Pres Apr–Sep Native Pres Apr–Sep Intro Pres Jun–Oct Native Pres Apr–Jul ower ax-leaved spreading yellowcress hairy yellowcress smooth yellowcress Rollins’ bladderpod silver bladderpod Moab thelypody sharp-leaf twinpod watercress Intro Pres May–Sep elegant thelypody branched pepperwort heart-leaf jewelﬂ prince’s-plume Native Pres Apr–Sep Jones’ pepperwort smoothleaf prince’s-plume broad-leaf pepperwort ﬂ plainsmustard Common name Nativity

var. var. var. var. var. cinale jonesii Rorippa palustris hispida Rorippa palustris glabra subumbellata ludoviciana Thelypodium integrifolium Physaria acutifolia elegans Lepidium ramosissimum bourgeauanum pinnata Lepidium montanum var. integrifolia Schoenocrambe linifolia

var. var. var.

Species name cinaleoffi Nasturtium var. jonesii Rorippa sinuata Rorippa sinuata Rorippa islandica hispida Rorippa islandica glabra Physaria subumbellata Lesquerella Physaria ludoviciana Lesquerella Thelypodium integrifolium gracilipes Nasturtium offi Physaria acutifolia acutifolia Thelypodiopsis elegans Thelypodiopsis Streptanthus cordatus Streptanthus cordatus Stanleya pinnata Stanleya pinnata Lepidium montanum var. Stanleya integrifolia Stanleya pinnata Utah Colorado Lepidium latifolium Cardaria latifolia Schoenocrambe linifolia rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Family Perennial Forbs, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Appendix D. Confi

Chapter 6: Appendices 153 Disturbed areas X X X X X X X Wet Comments XX Mt Con PJ XX XX XX XXX XXX X XXX XX Shr Des Flower time Park status Native Pres May–Aug Native Hist Jun–Sep Native Pres Jun–Aug Native Rep Jul–Sep Native Rep Jun–Sep Native Pres Jul–Aug Native Pres May–Jun Native Pres May–Aug Native Pres Apr–Aug Native Pres May–Jul eld mint eld Native Pres Jun–Sep silky phacelia Native Pres Jun–Aug lanceleaf phacelia common mare’s-tail sticky crane’s- bill catnip Intro Pres Jun–Sep ﬁ golden corydalis Native Pres Apr–Sep Rocky Mountain gentian leafy spurge IntroGreat Basin Repcentaury May–Aug X American bugleweed shorthorn spurge teasel spurgeFendler’s Native Pres Intro May–Sep Rep X Jul–Sep X X Idaho blue-eyed grass alpine pennycress Rocky Mountain iris Common name Nativity

nis

ssp.

var. var. occidentalis Phacelia sericea ciliosa Geranium viscosissimum nervosum Mentha arvensis Corydalis curvisiliqua ssp. Pneumonanthe afﬁ exaltatum brachyceras idahoense occidentale Noccaea montana

var.

var. Species name var. var. var. var. nis nis Phacelia sericea ciliosa Iris missouriensis Iris missouriensis Phacelia hastata Phacelia hastata Hippuris vulgaris Hippuris vulgaris Geranium viscosissimum incisum Gentiana affi Nepeta cataria Nepeta cataria Mentha arvensis glabrata Corydalis aurea occidentalis affi Euphorbia esula esula Tithymalus Centaurium exaltatum Centaurium Lycopus americanusLycopus americanus Lycopus Euphorbia brachycera Tithymalus Dipsacus sylvestrisChamaesyce fendleri Dipsacus fullonum Chamaesyce fendleri Sisyrinchium idahoense Sisyrinchium Utah Colorado Thlaspi montanum montanum rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Hydrophyllaceae Iridaceae Hydrophyllaceae Hippuridaceae Geraniaceae Gentianaceae Labiatae (Lamiaceae) Labiatae (Lamiaceae) Fumariaceae Euphorbiaceae Gentianaceae Labiatae (Lamiaceae) Euphorbiaceae Dipsacaceae Euphorbiaceae Iridaceae Family Perennial Forbs, cont. Cruciferae (Brassicaceae) Appendix D. Confi

154 Annotated Checklist of Vascular Flora: Dinosaur National Monument Entrada, Glen Canyon, & Frontier, Dakota F. Sandy soils Morrison F. Weber F. Weber Morrison F. X Wet Comments XX Mt Con X PJ XX XX XX X XX XX XXX X XX XX XX XXX XXX Shr Des Flower time Park status Native Pres May–Aug Native Pres Apr–Jun Native Hist Apr–Jun Native Pres May–Aug Native Pres May–Aug Native Pres May–Jun Native Hist May–Jun Native Hist Apr–Jul Native Pres Apr–Jun Native Pres Apr–Jul eld milkvetch eld Native Rep May–Aug lesser rushy milkvetch grass milkvetch Native Pres May–Jul cicada milkvetch painted milkvetch tall kentrophyta Native Histstanding Jun–Sep milkvetch Hayden’s Hayden’s milkvetch Hamilton’s Hamilton’s milkvetch milkweed milkvetch yellow milkvetch little skullcap Native Presﬁ Jun–Aug Martin’s milkvetch self-heal Native Pres May–Sep Duchesne milkvetch Common name Nativity

var. var. var. avus ceramicus Astragalus convallarius Astragalus chamaeleuce Astragalus ceramicus var. Astragalus kentrophyta elatus Astragalus laxmannii Astragalus haydenianus Astragalus asclepiadoides Astragalus ﬂ galericulata epilobifolia Astragalus argophyllus martinii Prunella vulgaris Astragalus duchesnensis

var. var. Species name avus convallarius ceramicus elatus robustior haydenianus martinii avus Astragalus convallarius var. Astragalus chloodes Astragalus chamaeleuce Astragalus ceramicus var. Astragalus kentrophyta var. Astragalus laxmannii var. Astragalus bisulcatus var. Astragalus hamiltonii Astragalus hamiltonii Astragalus asclepiadoides Astragalus fl fl Utah Colorado Scutellaria galericulata Scutellaria Astragalus agrestisAstragalus argophyllus Astragalus agrestis var. Prunella vulgaris lanceolata Astragalus duchesnensis rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Labiatae (Lamiaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Family Perennial Forbs, cont. Labiatae (Lamiaceae) Leguminosae (Fabaceae) Appendix D. Confi

Chapter 6: Appendices 155 Moenkopi F. Moenkopi & Morrison F. areas Uinta F. areas X Wet Comments XX XX Mt Con XX PJ XXX XXX XXX XX XX XX XX XX XX XXX XXX Shr Des Flower time Park status Native Hist May–Aug Native Pres May–Aug Native Pres May–Jul Native Pres Apr–Jun Intro Pres May–Oct Disturbed Native Pres Apr–Jun Native Rep May–Jul Native Pres Feb–Jun Native Pres Jun–Aug Native Pres Jun–Aug Native Pres May–Aug Rollins’ sweetvetch northern sweetvetch pulse milkvetch Native licoriceNuttall’s Pres Native May–Aug Pres May–Jul spoonleaf milkvetch Dinosaur milkvetch yellow sweet- clover Pursh’s Pursh’s milkvetch Trelease’s alkali Trelease’s milkvetch alfalfa Intro Rep May–Sep Disturbed woolly milkvetch Utah lupine Native Pres May–Aug white-leaved lupine Rydberg’s Rydberg’s weedy milkvetch silvery lupine Nativesubalpine lupine Pres May–Aug silvery lupine Native Pres May–Aug Common name Nativity

var. var. cinale thompsoniae argenteus rubricaulis argentatus Hedysarum boreale Astragalus purshii Astragalus mollissimus var. Lupinus caudatus utahensis Astragalus miser oblongifolius Lupinus argenteus var. Lupinus argenteus var. Lupinus argenteus var.

var. var. var. var. var. var. var. var. Species name cinalisoffi Melilotus treleasei thompsoniae Lupinus argenteus Hedysarum boreale gremiale Hedysarum boreale boreale Astragalus tenellusGlycyrrhiza lepidota Astragalus tenellus Glycyrrhiza lepidota Astragalus spatulatus Astragalus spatulatus Astragalus saurinus Astragalus saurinus Astragalus racemosus var. Melilotus offi Astragalus purshii purshii Medicago sativa Medicago sativa Astragalus mollissimus var. Lupinus caudatus utahensis Lupinus leucophyllus Lupinus leucophyllus Utah Colorado Astragalus miser oblongifolius Lupinus argenteus argenteus Lupinus argenteus rubricaulis argentatus rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Family Perennial Forbs, cont. Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Appendix D. Confi

156 Annotated Checklist of Vascular Flora: Dinosaur National Monument Weber F. Weber Moenkopi F. Brown’s Park Brown’s F. Saline to seleniferous soils Disturbed areas Bishop & Moenkopi F. X X Wet Comments Mt Con PJ XX XXX XXX XXX XX XXX XX X XX XX XXX XX XX Shr Des Flower time Park status Native Pres Jun–Jul Native Pres May–Aug Native Hist Apr–Jun Native Pres May–Jul owered Geyer’s onionGeyer’s NativeNevada onion Pres Native May–Sep Pres Apr–Jul dwarf clover Nativewhite clover Pres Intro Apr–Aug Pres Apr–Sep plains vetch Native Pres May–Aug Andean clover Native Pres May–Jul lesser duckweed Native oniontaper-tip Hist Native Brandegee’s onion Pres Jun–Oct May–Jul X X X X lemon scurf-pea Native Presprairie golden Apr–Aug pea large-ﬂ breadroot asparagussego-lily Introglacier-lily Pres Native Jun–Jul Pres Native Pres Jun–Jul May–Jul textile onionpink funnel-lily Native Native Pres Pres May–Jun Apr–May X X Maybell locoweed Common name Nativity

var. var. cinalis orum orum Allium geyeri gymnocarpon Vicia americana minor Psoralidium lanceolatum rhombifolia Pediomelum megalanthum Erythronium grandiﬂ Androstephium breviﬂ Oxytropis besseyi obnapiformis

var.

Species name

var. var. cinalisoffi Asparagus var. orum orum Allium geyeri geyeri Allium nevadense Allium nevadense Trifolium gymnocarponTrifolium Trifolium repensTrifolium Vicia americana minor repens Trifolium Allium acuminatumAllium brandegei Allium acuminatum Allium brandegei Trifolium andinumTrifolium andinum Trifolium Lemna minor Lemna turionifera Psoralidium lanceolatum lanceolatum Thermopsis rhombifolia Thermopsis Pediomelum megalanthum Utah Colorado Asparagus offi Calochortus nuttallii Calochortus nuttallii Erythronium grandifl Androstephium brevifl Allium textile Allium textile Oxytropis besseyi obnapiformis rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Liliaceae (Alliaceae) Liliaceae (Alliaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Alliaceae) Liliaceae (Alliaceae) Leguminosae (Fabaceae) Lemnaceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Liliaceae (Asparagaceae) Liliaceae (Calochortaceae) Liliaceae Liliaceae (Alliaceae, Themidaceae) Liliaceae (Alliaceae) Family Perennial Forbs, cont. Leguminosae (Fabaceae) Appendix D. Confi

Chapter 6: Appendices 157 X X X X Wet Comments Mt Con PJ XXX XXX X XXX XXX XXX XX XXX XXX X XXX XXX X XXX Shr Des Flower time Park status Native Pres May–Aug Native Pres May–Aug Native Pres Apr–Sep Native Pres Mar–Oct Native Pres Apr–Oct Native Pres May–Sep Native Pres Apr–Jul Native Pres Jul–Aug Native Rep Jul–Aug Native Pres Mar–Jun Native Rep Jun–Aug Native Pres Apr–Jun Native Pres Jun–Aug ax Native Pres Apr–Aug lavender-leaf lavender-leaf sundrops narrow-leaf four-o’clock fragrant sand- verbena small-leaf globemallow scarlet globemallow wild hollyhock Native Pres Jun–Oct X Uinta Basin stickleaf blue ﬂ meadow death camas alcove death camas marsh willow- herb foothills death camas Hall’s willow- Hall’s herb leopard-lily Nativestarry false PresSolomon’s-seal Apr–Jul yellow-bell Native Pres Mar–Jun X X northern willow-herb Common name Nativity

var. dissecta Calylophus lavandulifolius Oxybaphus linearis ssp. Nuttallia multicaulis Adenolinum lewisii venenosum palustre grammadophyllum paniculatum atropurpurea stellatum

var. Species name ssp. uintahensis Calylophus lavandulifolius Mirabilis linearis linearis Abronia fragrans Abronia elliptica Sphaeralcea parvifolia Sphaeralcea parvifolia Sphaeralcea coccinea Sphaeralcea coccinea Iliamna rivularis Iliamna rivularis Mentzelia multicaulis var. Linum perenne lewisii Zigadenus venenosus Toxicoscordion Zigadenus vaginatus Anticlea vaginatus Epilobium palustre Epilobium Zigadenus paniculatus Toxicoscordion Epilobium halleanum Epilobium halleanum Utah Colorado Fritillaria atropurpurea Fritillaria Smilacina stellata Maianthemum Fritillaria pudica Fritillaria pudica Epilobium ciliatum Epilobium ciliatum rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Onagraceae Onagraceae Nyctaginaceae Nyctaginaceae Malvaceae Malvaceae Malvaceae Loasaceae Linaceae Liliaceae (Melanthiaceae) Liliaceae (Melanthiaceae) Onagraceae Liliaceae (Melanthiaceae) Onagraceae Family Perennial Forbs, cont. Liliaceae Liliaceae (Convallariaceae, Ruscaceae) Liliaceae Appendix D. Conﬁ

158 Annotated Checklist of Vascular Flora: Dinosaur National Monument Weber F. Weber X X X X Wet Comments X Mt Con PJ XXX XX XXX X XXX XXX XXX XXX XXX XXX XX Shr Des Flower time Park status Native Pres Apr–Aug Native Hist Jun–Jul Native Pres Jun–Aug Native Pres May–Jul Native Pres May–Jun Native Pres Jun–Aug Native Pres Jun–Jul Native Pres Jun–Aug Native Pres Jun–Aug Native Pres Apr–Jun Native Pres Jul–Sep Native Pres Apr–Jun Native Pres Jul–Aug Native Hist Apr–Jun Native Pres Jun–Aug at-topped giant helleborine striped coral- root biennial evening- primrose pale evening- primrose large yellow evening- primrose western evening- primrose clustered broomrape hairy-throat evening- primrose ﬂ broomrape Paria evening- primrose Ute ladies’- tresses long-tube evening- primrose alcove bog- orchid tufted evening- primrose Alaska rein orchid Common name Nativity

ssp. ssp. trichocalyx navajoensis marginata caespitosa Oenothera villosa strigosa Oenothera pallida ssp. Oenothera acutissima hirsutissima Oenothera coronopifolia Oenothera caespitosa ssp. Spiranthes diluvialis Oenothera caespitosa ssp. Oenothera caespitosa ssp.

var. var. var. Species name var. ava ana navajoensis marginata caespitosa Epipactis gigantea Epipactis gigantea Corallorhiza striata Corallorhiza striata Oenothera villosa strigosa Oenothera pallida trichocalyx Oenothera fl acutissima Oenothera elata Oenothera elata Orobanche fasciculata Aphyllon fasciculatum Oenothera coronopifolia Orobanche corymbosacorymbosa Orobanche Oenothera caespitosa var. Spiranthes romanzoffi diluvialis Oenothera caespitosa var. Habenaria zothecina Limnorchis zothecina Utah Colorado Oenothera caespitosa var. Habenaria unalascensis Piperia unalascensis rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Orchidaceae Orchidaceae Orchidaceae Onagraceae Onagraceae Onagraceae Onagraceae Orobanchaceae Onagraceae Orobanchaceae Onagraceae Orchidaceae Onagraceae Orchidaceae Family Perennial Forbs, cont. Onagraceae Appendix D. Confi

Chapter 6: Appendices 159 Disturbed areas Morrison F. Wet Comments Mt Con PJ XX XX XXX XXX XXX XXX XX XXX XXX X XXX XXX X XX XXX XX Shr Des Flower time Park status Native Pres Apr–Aug Native Pres Jun–Sep Native Pres Jun–Oct Native Pres May–Aug Native Pres Jun–Aug Native Hist Jun–Oct Native Pres May–Jul Intro Pres May–Aug Native Pres May–Sep Native Hist May–Jul Native Pres Jul–Oct cushion wild buckwheat desert phlox Nativecarpet phlox Pres Nativelong-leaf phlox Apr–Jun Native Pres Pres Apr–Jun wild Bateman’s Apr–Julbuckwheat X X X slender wild buckwheat Watson’s Watson’s prickly-phlox Rocky Mountain phlox Baker’s wild Baker’s buckwheat scarlet giliaball-head gilia Native Native Presmatted prickly- Presphlox May–Aug Jun–Aug sticky gilia Native Pres May–Sep X X common plantain whorled wild buckwheat Cooper’s Cooper’s broomrape Duchesne wild buckwheat Common name Nativity

var. da ora var. aggregata congesta viridulum . Eriogonum brevicaule var. ovalifolium ovalifolium Phlox austromontana Phlox hoodii Eriogonum batemanii Eriogonum microthecum simpsonii watsonii Eriogonum jamesii Ipomopsis aggregata ssp Ipomopsis congesta var. Leptodactylon caespitosum Eriogonum heracleoides Orobanche ludoviciana

var. var.

Species name

var. var. dapinnatifi Gilia oramultifl Phlox viridulum austromontana batemanii aggregata cooperi avescens Eriogonum brevicaule var. Eriogonum ovalifolium Eriogonum Phlox multifl Phlox austromontana var. Phlox hoodii canescens Phlox longifoliaEriogonum batemanii Phlox longifolia var. Eriogonum microthecum simpsonii Leptodactylon watsonii Leptodactylon Gilia pinnatifi Eriogonum jamesii fl Ipomopsis aggregata var. Ipomopsis congesta congesta Leptodactylon caespitosum Plantago major Plantago major Eriogonum heracleoides Utah Colorado Orobanche ludoviciana var. rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Polygonaceae Polygonaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polygonaceae Polygonaceae Polemoniaceae Polemoniaceae Polygonaceae Polemoniaceae Polemoniaceae Polemoniaceae Plantaginaceae Polygonaceae Family Perennial Forbs, cont. Orobanchaceae Appendix D. Confi

160 Annotated Checklist of Vascular Flora: Dinosaur National Monument Disturbed areas Disturbed areas Brown’s Park Brown’s F. & Weber X X X X X X X Wet Comments Mt Con XX PJ XXX XXX X XXX XXX XXX XX XX Shr Des Flower time Park status Native Hist Jun–Aug Native Pres Apr–Sep Native Pres May–Jun Native Pres Apr–Jul Native Hist Jun–Sep Native Pres Jun–Sep Native Pres Mar–Aug Native Pres Jun–Sep Native Rep Jul–Aug Native Pres May–Jul Native Hist Jul–Sep Native Hist May–Aug Native Hist Jun–Aug ower owered northern pondweed small-ﬂ fame-ﬂ Oregon bitter- root curly dockcanaigre dock Intro Nativewillow dock Pres Hist Native May–Sep lanceleaf Apr–Jun springbeauty Hist May–Sep water smartweed sulfur wild buckwheat dark-throat shooting-star golden sulphur wild buckwheat dwarf pondweed Woodside wild Woodside buckwheat fennel-leaf pondweed Shockley’s wild Shockley’s buckwheat longleaf pondweed Common name Nativity

orum var. var. tenuifolius longilobum var. hymenosepalus Rumex triangulivalvis Eriogonum umbellatum umbellatum Dodecatheon pulchellum Eriogonum umbellatum aureum tumulosum Potamogeton pectinatus var. nodosus

ssp. Species name

orumparvifl Talinum var. umbellatum aureum Potamogeton alpinus Potamogeton alpinus Talinum parvifl Talinum Lewisia rediviva Lewisia rediviva Rumex crispusRumex hymenosepalus Rumex Rumex crispus Rumex salicifolius triangulivalvis Claytonia lanceolata Claytonia lanceolata Polygonum amphibium Persicaria amphibia Eriogonum umbellatum var. pulchellum Dodecatheon pulchellum Eriogonum umbellatum var. Potamogeton pusillus Potamogeton pusillus Eriogonum tumulosum Eriogonum Potamogeton pectinatus Utah Colorado Eriogonum shockleyi Eriogonum shockleyi Potamogeton nodosus Potamogeton rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Potamogetonaceae Portulacaceae Portulacaceae Polygonaceae Polygonaceae Polygonaceae Portulacaceae Polygonaceae Polygonaceae Primulaceae Polygonaceae Potamogetonaceae Polygonaceae Potamogetonaceae Family Perennial Forbs, cont. Polygonaceae Potamogetonaceae Appendix D. Confi

Chapter 6: Appendices 161 Rock crevices X X X X X X X X Wet Comments X X Mt Con PJ XX XXX XXX XX XXX Shr Des Flower time Park status Native Rep May–Jul Native Pres Feb–Jun Native Rep Jun–Aug Native Pres Apr–Jun Native Pres May–Sep Native Rep May–Sep Native Pres Apr–Jul Native Pres Jul–Aug Native Pres May–Jun Native Pres Jun–Jul Native Pres Jun–Aug Gordon’s ivesiaGordon’s Native Pres Jun–Aug X Macoun’s Macoun’s buttercup Jupiter buttercup lesser yellow water buttercup sagebrush buttercup marsh buttercup thread water- buttercup Nuttall’s Nuttall’s larkspur white virgin’s- bower alcove columbine Graham’s Graham’s columbine sidebells pyrola Native Hist Jun–Aug sea-milkwort Native Hist May–Jul modest cinquefoil tall cinquefoil Native Hist Jun–Jul silverweed Native Pres May–Aug X rock spiraea Native Pres Aug–Sep Common name Nativity

var. ssp. hookeri var. Ranunculus glaberrimus ellipticus saximontana cymbalaria Batrachium circinatum Delphinium nuttallianum Potentilla concinna Drymocallis arguta Petrophyton caespitosum

var. var. Species name

var. diffusus Petrophyton Ivesia gordonii Ivesia gordonii Ranunculus macounii Ranunculus macounii Ranunculus jovis Ranunculus jovis Ranunculus gmelinii Ranunculus gmelinii Ranunculus glaberrimus ellipticus Ranunculus cymbalaria Halerpestes Ranunculus aquatilis var. Delphinium nuttallianum Clematis ligusticifolia Clematis ligusticifolia Aquilegia micrantha Aquilegia micrantha Utah Colorado Aquilegia grahamii Pyrola secunda Orthilia secunda Glaux maritima Glaux maritima Potentilla concinna modesta Potentilla arguta convallaria Potentilla anserina Argentina anserina caespitosum rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Rosaceae Rosaceae Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae Ranunculaceae (Helleboraceae) Ranunculaceae (Helleboraceae) Pyrolaceae (Ericaceae) Family Perennial Forbs, cont. Primulaceae (Myrsinaceae) Rosaceae Rosaceae Rosaceae Appendix D. Conﬁ

162 Annotated Checklist of Vascular Flora: Dinosaur National Monument Sandstone X X X Wet Comments X X Mt Con PJ XXX XXX X XXX XXX XX XXX XX XX XXX X XXX XXX X XXX Shr Des Flower time Park status Native Pres May–Sep Native Pres Apr–Aug Native Pres May–Aug Native Pres Jul–Aug Native Pres May–Aug Native Pres May–Jul Native Pres Jul–Aug Native Hist Jun–Aug Native Pres Jul–Aug Native Pres Jun–Aug Native Pres Apr–Jun Native Pres May–Oct Native Rep Jun–Aug Native Pres Apr–Aug Native Hist Jun–Aug Native Pres Apr–Aug Native Pres Mar–Aug ower ax littleleaf alumroot bastard toadﬂ shrubby bedstraw sweet-scented bedstraw Colorado bedstraw Eastwood’s Eastwood’s paintbrush sheep cinquefoil beautiful cinquefoil Wyoming paintbrush glabrate cinquefoil desert paintbrush marsh grass-of- Parnassus comb-leaf cinquefoil slender woodlandstar glandular cinquefoil smallﬂ woodlandstar fringecup woodlandstar Common name Nativity

var. var. orum orum orum Comandra umbellata Galium multiﬂ Galium coloradoense Castilleja scabrida Potentilla ovina Potentilla pulcherrima glabrata Potentilla gracilis Parnassia palustris tenellum brunnescens Lithophragma parviﬂ Potentilla gracilis Lithophragma glabrum

var. var. var. var. var. var. Species name var. orum orum orumtrifl Galium orum orum pallida multifl coloradoense ramulosum Heuchera parvifolia Heuchera parvifolia Galium trifl Comandra umbellata var. Galium multifl var. Galium multifl var. scabrida Potentilla ovina ovina Castilleja scabrida Potentilla gracilis pulcherrima Castilleja linariifolia Castilleja linariifolia Castilleja chromosa Castilleja chromosa Potentilla gracilis fastigiata Parnassia palustris montanensis Potentilla gracilis elmeri Lithophragma tenellum Lithophragma Lithophragma parvifl Utah Colorado brunnescens Potentilla gracilis Lithophragma glabrum var. rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Saxifragaceae Rubiaceae Santalaceae Rubiaceae Rubiaceae Rosaceae Scrophulariaceae Rosaceae Scrophulariaceae Scrophulariaceae Rosaceae Saxifragaceae (Parnassiaceae) Rosaceae Saxifragaceae Saxifragaceae Family Perennial Forbs, cont. Rosaceae Saxifragaceae Appendix D. Confi

Chapter 6: Appendices 163 Disturbed areas Disturbed areas Disturbed areas Lodore & F. Weber X X Wet Comments X Mt Con PJ XXX XX XXX XXX XX XXX X XXX XXX XXX XX XXX XX Shr Des Flower time Park status Native Pres May–Jul Native Hist Jun–Aug Native Pres May–Jun Native Pres Jul–Aug Native Pres May–Jun Native Hist Jun–Jul Native Hist May–Jun Native Pres Jun–Aug Native Pres Jun–Aug Native Pres Jun–Aug Intro Rep Jul–Aug Native Pres May–Jul ower ower ax gwort mat-root penstemon small-fl penstemon low penstemon Nativethick-leaf Prespenstemon May–Jun American brooklime woolly mullein Intro Rep Jun–Aug mat penstemon Native Fremont’s Prespenstemon May–Jul lance-leaf fi Vernal Vernal penstemon Watson’s Watson’s penstemon butter-and-eggs Introyellow Presmonkeyfl Jun–Aug Rocky Mountain penstemon Dalmatian toadfl mudwort Native Pres Jun–Aug X Blue Mountain penstemon Common name Nativity

var. var. procerus dalmatica fremontii cyanomontanus Penstemon confertus ssp. Penstemon humilis Penstemon pachyphyllus mucronatus Penstemon caespitosus var. lanceolata Penstemon angustifolius vernalensis ssp. Penstemon scariosus var.

var.

Species name var. var. procerus caespitosus cyanomontanus Penstemon radicosus Penstemon radicosus Penstemon procerus var. humilis Penstemon humilis Penstemon pachyphyllus mucronatus Veronica americanaVeronica americana Veronica Verbascum thapsusVerbascum thapsus Verbascum Penstemon caespitosus var. Penstemon fremontii Penstemon fremontii Scrophularia lanceolata Scrophularia Penstemon angustifolius vernalensis Penstemon watsonii Penstemon watsonii Linaria vulgarisMimulus guttatus Linaria vulgaris Mimulus guttatus Penstemon strictus Penstemon strictus Utah Colorado Limosella aquaticaLinaria dalmatica Limosella aquatica Linaria genistifolia Penstemon scariosus var. rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Family Perennial Forbs, cont. Scrophulariaceae Scrophulariaceae Appendix D. Conﬁ

164 Annotated Checklist of Vascular Flora: Dinosaur National Monument Sandy to heavy clay soils Morrison F. Sandy soils Sandy to clayey soils X X Wet Comments Mt Con PJ XXX XXX XX XXX XX XX XXX XX XXX XX XX XXX X XX Shr Des Flower time Park status Native Pres Apr–Jun Native Pres Apr–Jun Native Pres Apr–May Native Pres May–Jul Native Pres Apr–May Native Rep Apr–May Native Hist Apr–May Native Pres May–Jul Native Hist Apr–May Native Pres Apr–May Native Pres Apr–May Native Rep Jun–Aug Intro PresNative Pres Jun–Aug Jun–Sep Native Pres Apr–May at-seed aromatic spring- parsley Colorado Plateau spring- parsley Duchesne spring-parsley ﬂ lomatium Constance’s Constance’s spring-parsley big-seed lomatium onion spring- parsley juniper lomatium Fendler’s Fendler’s cymopterus giant lomatium Nativemilfoil Preslomatium Apr–Jun plains spring- parsley southern ligusticum water speedwell cutleaf water- parsnip rock spring- parsley Common name Nativity

var. platycarpum Cymopterus terebinthinus calcareus Cymopterus purpureus Cymopterus duchesnensis Lomatium triternatum var. constancei Lomatium macrocarpum juniperinum Cymopterus fendleri Lomatium dissectum Lomatium grayi Cymopterus acaulis Veronica anagallis- Veronica aquatica Aletes petraeus

var. var.

incisa Berula erecta Species name var. var. var. var. orus platycarpum purpureus eatonii Cymopterus terebinthinus Cymopterus terebinthinus albiﬂ Cymopterus purpureus var. Cymopterus duchesnensis Lomatium triternatum ssp. Cymopterus constancei Cymopterus Lomatium macrocarpum Cymopterus bulbosus Cymopterus bulbosus Lomatium juniperinum Lomatium Cymopterus acaulis fendleri Lomatium dissectum var. Lomatium grayi grayi Berula erecta Cymopterus acaulis acaulis Ligusticum porteri Ligusticum porteri Utah Colorado Veronica anagallis- Veronica aquatica petraeus rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Umbelliferae (Apiaceae) Appendix D. Conﬁ Family Perennial Forbs, cont. Scrophulariaceae

Chapter 6: Appendices 165 areas X X Wet Comments X XX XX Mt Con X PJ XXX XX XX XXX XXX XX XXX XXX XX Shr Des Flower time Park status Native Hist Mar–Nov Native Hist Aug–Sep Native Hist Mar–Aug Native Hist Apr–Aug Native Pres Apr–Jul Native Pres Mar–Aug Native Hist May–Jul Native Hist May–Sep Native Pres Apr–Jun Native Pres Jun–Jul Native Hist May–Jul Native Pres May–Jul Native Pres Jun–Sep Disturbed horned pondweed pinyon dwarf- mistletoe valley violetgoose-foot Nativeviolet Pres Mar–Jul upland yellow violet northern bog violet hook-spur violet sand dune cryptanth stinging nettle Nativeprostrate Presvervain Jul–Aug thick-sepaled cryptanth blunt sweet- cicely obscure cryptanth Indian-potato NativeChilean sweet- Prescicely Mar–May prostrate pigweed Common name Nativity

ssp.

ssp. ssp. var. Arceuthobium divaricatum Viola vallicola Viola purpurea venosa major Viola praemorsa Urtica gracilis holosericea Cryptantha crassisepala elachantha Osmorhiza depauperata

Species name

var. var. var. var. elachantha Zannichellia palustris Zannichellia palustris Arceuthobium divaricatum Viola nuttallii vallicola Viola purpurea venosa Viola nuttallii major Viola nephrophylla Viola sororia Viola adunca Viola adunca Cryptantha fendleri Cryptantha fendleri Urtica dioica occidentalis bracteataVerbena bracteata Verbena Cryptantha crassisepala var. Osmorhiza depauperata Cryptantha ambigua Cryptantha ambigua Utah Colorado Orogenia linearifolia Orogenia linearifolia Amaranthus blitoides Amaranthus blitoides Osmorhiza chilensis Osmorhiza chilensis rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Zannichelliaceae Viscaceae Violaceae Violaceae Violaceae Violaceae Violaceae Boraginaceae Urticaceae Verbenaceae Boraginaceae Umbelliferae (Apiaceae) Boraginaceae Family Perennial Forbs, cont. Umbelliferae (Apiaceae) Annual Forbs Amaranthaceae Umbelliferae (Apiaceae) Appendix D. Conﬁ

166 Annotated Checklist of Vascular Flora: Dinosaur National Monument Morrison F Disturbed areas Disturbed areas Disturbed areas Saline soils Disturbed areas Mancos & Morrison F X X Wet Comments Mt Con PJ XX X XXX X XXX XXX XXX XX XX XXX XXX XXX X XX X X XX Shr Des Flower time Park status Intro Pres Jun–Sep Native Rep Apr–Jun Native Pres May–Oct Native Pres May–Aug Native Pres Apr–Jun Native Pres Apr–Jun Native Hist Apr–Jun Native Hist Apr–Jul Native Pres Jun–Jul Native Pres Apr–Jul Intro Hist Aug–Oct ower ve-horn holosteumsalt sand- Introspurrey Rep Feb–May Goodrich’s Goodrich’s stinkweed meadow popcorn-fl yellow beeplant Native PresRocky Mountain beeplant Apr–Sep western stickseed cupseed stickseed recurved cryptanth wing-nut cryptanth Kelsey’s Kelsey’s cryptanth summer-cypress Intro Hist Aug–Oct slender cryptanth fi smotherweed slender orach Native Hist Jun–Sep wedge orach Native Rep Jun–Sep X X silver orach Native Hist Jun–Sep stalked orach Native Hist Aug–Sep Common name Nativity i penicillatus umbellatum Plagiobothrys scouleri ssp. Lappula redowskii Lappula marginata Cryptantha pterocarya Atriplex wolfi Atriplex argentea Atriplex saccaria

var. var. Species name var. i goodrichii penicillatus occidentalis cupulata pterocarya Holosteum umbellatum Holosteum Spergularia marina Spergularia marina Cleomella palmeriana var. Plagiobothrys scouleri var. Cleome luteaCleome serrulata Cleome lutea Cleome serrulata Lappula occidentalis var. Lappula occidentalis var. Cryptantha recurvata Cryptantha recurvata Cryptantha pterocarya var. Cryptantha kelseyana Cryptantha kelseyana Bassia scoparia Bassia sieversiana Utah Colorado Cryptantha gracilis Cryptantha gracilis Bassia hyssopifolia Bassia hyssopifolia Atriplex wolﬁ tenuissima Atriplex truncata Atriplex truncata Atriplex argentea argentea Atriplex saccaria saccaria rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Chenopodiaceae Caryophyllaceae (Alsinaceae) Caryophyllaceae (Alsinaceae) Capparaceae (Cleomaceae) Boraginaceae Capparaceae (Cleomaceae) Capparaceae (Cleomaceae) Boraginaceae Boraginaceae Boraginaceae Boraginaceae Boraginaceae Chenopodiaceae Appendix D. Conﬁ Family Annual Forbs, cont. Boraginaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae

Chapter 6: Appendices 167 Sandy soils Disturbed areas Sandy soils areas Disturbed areas X X X X Wet Comments X Mt Con PJ XX XX XXX X XXX XX X XXX XXX XXX X XX XXX Shr Des Flower time Park status Native Hist Aug–Oct Native Pres Jul–Oct Native Rep Aug–Sep Native Pres Jul–Sep Native Hist Jun–Sep Native Pres Aug–Sep Native Pres Jun–Sep Native PresNative Aug–Oct Pres Jun–Aug Native Pres Jun–Sep Native Pres Jul–Sep Native Hist Jun–Aug Native Pres Jul–Sep Native Pres Apr–Jul halogeton Nuttall’s Intro Pres Jul–Aug American bugseed maple-leaf goosefoot bur ragweed Native Pres Jul–Oct rayless alkali aster narrow-leaf goosefoot Torrey’s Torrey’s seepweed oakleaf goosefoot western seepweed Fremont’s Fremont’s goosefoot silvery goosefoot smallhead goosefoot Russian thistle Intro Pres Jun–Sep Disturbed mountain goosefoot tiny poverty- weed pitseed goosefoot poverty-weed Common name Nativity

ssp. glomeratus Corispermum americanum acanthicarpa angusta Chenopodium leptophyllum Suaeda moquinii Chenopodium glaucum Chenopodium fremontii Chenopodium incanum Chenopodium foliosum Chenopodium atrovirens Chenopodium berlandieri

var. Species name

fremontii incanum var. var. var. salinum berlandieri Halogeton glomeratus Halogeton Monolepis nuttalliana Monolepis nuttalliana Corispermum americanum Chenopodium simplex Chenopodium simplex Ambrosia acanthicarpa Ambrosia Aster brachyactis Brachyactis ciliata Chenopodium leptophyllum Suaeda torreyana torreyana Chenopodium fremontii Chenopodium fremontii Chenopodium glaucum var. Suaeda occidentalis Chenopodium capitatum parvicapitatum Salsola tragus Salsola australis Chenopodium atrovirens Monolepis pusilla Monolepis pusilla Utah Colorado Chenopodium album var. rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Compositae (Asteraceae) Compositae (Asteraceae) Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Chenopodiaceae Family Annual Forbs, cont. Chenopodiaceae Appendix D. Conﬁ

168 Annotated Checklist of Vascular Flora: Dinosaur National Monument Disturbed areas Disturbed areas areas areas areas X X X X Wet Comments XX Mt Con PJ XX X XX XXXX X XXX XX XX XXX Shr Des Flower time Park status Native Hist May–Sep Native Hist Apr–Jul Native Hist Apr–Jun Native Pres Jun–Sep Native Pres Jun–Sep Native Hist Jun–Oct Native Pres May–Oct Native Pres Apr–Jul Native Pres Jul–Oct Native Pres Jul–Sep Intro Hist Apr–Sep Disturbed Native Pres Apr–Jul Intro Pres Apr–Jul Disturbed ower ower ax white-plume wire-lettuce Torrey’s desert- Torrey’s dandelion sowthistle desert- dandelion prickly lettuce Intro Pres Jul–Sep prairie sunfl common sunfl cottonbatting cudweed lowland cudweed mountain tansy-mustard slender cudweed blue mustard Intro Pres Apr–Jun Disturbed Canadian horseweed shepherd’s- purse stevia dusty- maiden desert madwort Introlittle-pod false Presfl Apr–Jul rough cocklebur Native Pres Apr–Oct Common name Nativity

ssp. exigua sonchoides Helianthus petiolaris Helianthus annuus Pseudognaphalium stramineum Descurainia incisa incisa uliginosum Conyza canadensis pastoris Xanthium strumarium

var. ssp. var. Species name fallax canadense Stephanomeria exigua Stephanomeria Malacothrix torreyi Malacothrix torreyi Malacothrix sonchoides Malacothrix Lactuca serriola Lactuca serriola Helianthus petiolaris ssp. lenticularis Helianthus annuus Gnaphalium stramineum Gnaphalium palustre Gnaphalium palustre Descurainia incana sonnei Gnaphalium exilifolium Gnaphalium Chorispora tenella Chorispora tenella Conyza canadensis glabrata Capsella bursa-pastoris Capsella bursa- Utah Colorado Chaenactis stevioides Chaenactis stevioides Alyssum desertorum Alyssum desertorum Camelina microcarpa Camelina microcarpa Xanthium strumarium var. rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Compositae (Asteraceae) Cruciferae (Brassicaceae) Appendix D. Conﬁ Family Annual Forbs, cont. Compositae (Asteraceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae)

Chapter 6: Appendices 169 areas areas Morrison, Duchesne Mancos River, F. Disturbed areas Disturbed areas Sandy or clayey soils X Wet Comments Mt Con PJ XXX XX XX XX XXX XX XXX XX XXX XXX XX XXX XXX XX Shr Des Flower time Park status Native Pres May–Jun Intro Pres May–Jul Native Hist Mar–Jun Native Hist May–Jul Native Pres May–Sep Native Pres Mar–Aug Native Pres Jun–Aug Native Hist Mar–Aug Native Pres Mar–Jun ddleleaf Native Hist May–Aug ddle-mustard ixweed Intro Pres Mar–Aug long-beak fi tumble mustard Intro Pres May–Aug Disturbed brittle phacelia Native Pres May–Jun African mustard Intro Pres Apr–Jul Disturbed corrugate phacelia clasping pepperwort leafy fi St. George pepperwort woods draba Introdwarf draba Pres Nativeovalpurse Apr–Jun Presprairie Apr–Jul Nativepepperwort Hist Apr–Jun fl thyme-leaved spurge western tansy- mustard ridge-seeded spurge western tansy- mustard Common name Nativity orum halictorum intermedia ramosum Streptanthella longirostris altissimum Phacelia crenulata Nama densum procumbens Lepidium densifl var. Chamaesyce serpyllifolia Descurainia pinnata ssp. Chamaesyce glyptosperma Descurainia pinnata ssp.

var.

var. Species name orum var. orum georginum ramosum osmiarum intermedia Streptanthella longirostris Sisymbrium altissimum Sisymbrium Phacelia demissa minor Malcolmia africana Malcolmia africana Phacelia crenulata corrugata Lepidium lasiocarpum var. Lepidium perfoliatum Lepidium perfoliatum Nama densum parvifl Draba nemorosaDraba reptans Draba nemorosa Hutchinsia procumbens Hymenolobus Draba reptans Lepidium densifl var. Descurainia sophia Descurainia sophia Chamaesyce serpyllifolia Descurainia pinnata var. Utah Colorado Chamaesyce glyptosperma Descurainia pinnata var. rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Hydrophyllaceae Cruciferae (Brassicaceae) Hydrophyllaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Hydrophyllaceae Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Cruciferae (Brassicaceae) Euphorbiaceae Cruciferae (Brassicaceae) Euphorbiaceae Family Annual Forbs, cont. Cruciferae (Brassicaceae) Appendix D. Confi

170 Annotated Checklist of Vascular Flora: Dinosaur National Monument Mancos Shale Disturbed areas Sandy soils X X Wet Comments Mt Con PJ XX X XX X X XXX XX XXX XX XXX XXX X XXX Shr Des Flower time Park status Native Hist Apr–Jun Native Hist May–Jun Native Pres Apr–Sep Native Hist Apr–Jun Native Pres Apr–Aug Native Hist Mar–Jun Intro Pres May–Oct Native Hist May–Aug Native Pres Apr–Jul Native Pres Jun–Aug Native Hist Apr–Jul Native Pres Jul–Sep ower Paiute suncup Native Pres May–Jun tiny camissonia Native Hist Walker’s May–Juncamissonia X X lesser camissonia small-ﬂ sandpuffs Thompson’s Thompson’s stickleaf woolly plantain woolly plantain Native Pres Mar–Aug X X X white-stem blazingstar white-stem evening- primrose black medick Introwhite sweet- clover Pres May–Oct dwarf lupine Native Hist Apr–Jul branching groundsmoke Great Basin lupine deceptive groundsmoke Ives’ phacelia Geyer’s Nativemilkvetch Pres Apr–Jun X X X autumn willow- herb Common name Nativity

var. var. Epilobium Camissonia scapoidea Tripterocalyx Tripterocalyx micranthus Lupinus pusillus pusillus Gayophytum ramosissimum Lupinus pusillus intermontanus decipiens brachycarpum

Species name var. var. scapoidea Epilobium brachycarpum Camissonia scapoidea var. Camissonia parvula Camissonia parvula Camissonia walkeri Camissonia walkeri Camissonia minor Camissonia minor Tripterocalyx Tripterocalyx micranthus Mentzelia thompsonii Acrolasia humilis Plantago patagonica Plantago patagonica Mentzelia albicaulis Acrolasia albicaulis Oenothera albicaulis Oenothera albicaulis Medicago lupulinaMelilotus alba Medicago lupulina Melilotus albus Lupinus pusillus pusillus Gayophytum ramosissimum Lupinus pusillus intermontanus Gayophytum decipiens Gayophytum Utah Colorado Phacelia ivesianaAstragalus geyeri Phacelia ivesiana Astragalus geyeri rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Onagraceae Onagraceae Onagraceae Onagraceae Onagraceae Nyctaginaceae Loasaceae Plantaginaceae Loasaceae Onagraceae Leguminosae (Fabaceae) Leguminosae (Fabaceae) Leguminosae (Fabaceae) Onagraceae Leguminosae (Fabaceae) Onagraceae Appendix D. Conﬁ Family Annual Forbs, cont. Hydrophyllaceae Leguminosae (Fabaceae)

Chapter 6: Appendices 171 areas Clay soils X X Wet Comments Mt Con XX PJ XXX XXX XX XXX X XX XX XXX XX XXX Shr Des Flower time Park status Native Pres Jun–Aug Native Pres Jun–Oct Native Hist Apr–Sep Native Hist Jun–Oct Native Pres Apr–Jul Native Pres May–Oct Native Hist Apr–Sep Native Pres Apr–Oct Native Rep Jun–Sep Native Hist May–Jul Sawatch knotweed Douglas’ knotweed yard knotweed Intro Pres May–Oct Disturbed smooth wild buckwheat Hooker’s wild Hooker’s buckwheat Grand Valley Grand Valley desert trumpet Gordon’s wild Gordon’s buckwheat golden dock Native Hist May–Sep bent wild buckwheat willow-weed Intro Pres Jul–Nov nodding wild buckwheat Kellogg’s Kellogg’s knotweed small collomiashy gilia Nativegymnosteris Presspreading gilia May–Aug Native Native Nativedwarf gilia Hist Xslender Pres Pres phlox Native Native May–Aug Apr–Jul Apr–Junyellow Pres Presnavarretia X X Mar–Jun X X Apr–Jun X X X X X X Common name Nativity

var.

exum ssp. Polygonum douglasii arenastrum Stenogonum salsuginosum fueginus Persicaria lapathifolia Rumex maritimus Eriogonum cernuum kelloggii Microsteris gracilis polygaloides

var. var. Species name atum exumfl Stenogonum johnstonii douglasii fusiforme cernuum Polygonum douglasii var. Polygonum douglasii var. Polygonum aviculare Polygonum Eriogonum salsuginosum Eriogonum infl var. Eriogonum hookeri Eriogonum hookeri Eriogonum gordonii Eriogonum gordonii Eriogonum fl Polygonum lapathifolium Rumex maritimus fueginus Eriogonum cernuum var. Utah Colorado Collomia linearisGilia inconspicua Gymnosteris parvula Collomia linearis Ipomopsis polycladon Gilia sinuata Gymnosteris parvula Ipomopsis pumila Ipomopsis polycladon Microsteris gracilis Ipomopsis pumila humilior Navarretia breweri Navarretia breweri Polygonum kelloggii Polygonum rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Polygonaceae Family Annual Forbs, cont. Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Polemoniaceae Appendix D. Confi

172 Annotated Checklist of Vascular Flora: Dinosaur National Monument Disturbed areas areas X X X X X Wet Comments X Mt Con XX PJ XXX XX XXX X XX XXX XXX XXX XXX XXX X Shr Des Flower time Park status Native Pres Jun–Aug Native Pres Jun–Aug Native Pres May–Aug Native Pres May–Aug Native Pres May–Sep Native Hist Jun–Aug Native Pres May–Aug Native Pres Jul–Aug Native Pres Apr–Jun Native Hist May–Jun Native Pres Jun–Aug Native Pres Apr–Jun Intro Hist Jun–Sep Disturbed Native Rep May–Jul ower ower ower ower seep monkeyfl bushy bird’s- beak blue-eyed Mary Native Pres Apr–Jun X X X annual paintbrush cleaverstwin-leaf bedstraw Native Pres Apr–Jun bur buttercup Introbiennial cinquefoil Pres Mar–Jun blister buttercup purslane speedwell pygmy-fl rock-jasmine yellow owl- clover western rock- jasmine Suksdorf’s Suksdorf’s monkeyfl slender-stem slender-stem rock-jasmine reddish monkeyfl common purslane miner’s lettuceminer’s Native Pres Apr–Jun Common name Nativity i ora liformis oribundus xalapensis Galium spurium orthoceras Hecatonia scelerata Veronica peregrina Veronica var. Androsace septentrionalis occidentalis Claytonia rubra

var.

var. isuksdorfi Mimulus Species name var. oraparvifl Collinsia liformisfi Androsace dus oribundusfl Mimulus multifi Mimulus fl Collinsia parvifl Cordylanthus ramosus Cordylanthus ramosus Castilleja exilis Castilleja minor Galium aparine echinospermum Galium bifolium Galium bifolium Ranunculus testiculatus Ceratocephala Potentilla biennis Potentilla biennis Ranunculus sceleratus var. Veronica peregrina Veronica xalapensis Androsace septentrionalis Orthocarpus luteus Orthocarpus luteus Androsace occidentalis Androsace Mimulus suksdorfi Androsace fi Utah Colorado Mimulus rubellus Mimulus rubellus Portulaca oleracea Portulaca oleracea Montia perfoliata perfoliata rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Scrophulariaceae Scrophulariaceae Scrophulariaceae Scrophulariaceae Rubiaceae Rubiaceae Ranunculaceae Rosaceae Ranunculaceae Scrophulariaceae Primulaceae Scrophulariaceae Primulaceae Scrophulariaceae Primulaceae Scrophulariaceae Portulacaceae Appendix D. Confi Family Annual Forbs, cont. Portulacaceae

Chapter 6: Appendices 173 Disturbed areas Disturbed areas X X X X X Wet Comments XX Mt Con PJ XXX XX XX XXX Shr Des Flower time Park status Native Pres May–Aug Native Pres Jun–Aug Native Pres May–Jun Native Pres Jun–Jul Native Pres May–Aug Native Hist May–Aug Native Pres May–Aug Native Pres May–Sep common Ross’ sedgedryland sedge Nativevalley sedge Native Pres Pres Native May–Aug Jul–Aug Pres X May–Jul X X X X X Nebraska sedge Nativewestern sedge Pres sedgeParry’s Nativewoolly sedge Pres May–Jul Nativeblackcreeper Native Jun–Aug Pressedge Pres May–Jul May–Sep X X X X sheep sedgesmall-wing Nativesedge Pres Jun–Aug X elk sedge sedgeHood’s bottlebrush Native Nativesedge Pres Pres May–Jul Apr–Jul X X X narrowleaf sedge golden sedgeCanyonlands Nativesedge Pres Apr–Aug X black henbane Introcoyote tobacco Rep Native Pres Jun–Aug Jun–Sep water sedge slender-beaked Nativesedge Rep Jun–Aug X spikerush Common name Nativity cut-leaf nightshade

ssp. orum macrostachya eleocharis Species name orumtrifl Solanum Carex tahoensisCarex vallicolaEleocharis palustris Carex xerantica Eleocharis Carex vallicola Carex rossii Carex rossii Carex occidentalisCarex parryana Carex pellita Carex occidentalis Carex praegracilis Carex parryana Carex praegracilis Carex lanuginosa Carex nebrascensis Carex nebrascensis Carex microptera Carex microptera Carex illota Carex illota Carex hoodiiCarex hystericina Carex hystericina Carex hoodii Carex geyeri Carex geyeri Carex duriuscula Carex stenophylla Carex curatorum Carex aurea Carex aurea Utah Colorado Hyoscyamus niger Hyoscyamus niger Carex athrostachya Carex athrostachya Nicotiana attenuata Nicotiana attenuata Solanum trifl Carex aquatilis Carex aquatilis rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Cyperaceae Family Annual Forbs, cont. Solanaceae Cyperaceae Solanaceae Solanaceae Graminoids Perennial Cyperaceae Appendix D. Confi

174 Annotated Checklist of Vascular Flora: Dinosaur National Monument X X X X X X X X Wet Comments XX X XX Mt Con PJ XXX XXX XXX XXX XXX Shr Des Flower time Rep Jul–Aug Park status Native Pres Jun–Aug Intro Pres Jun–Aug Native Pres Jun–Aug Native Pres May–Aug Native Pres Jun–Aug Native Pres Mar–Sep Native Pres Jun–Aug ower big bluestem Native Pres Jul–Sep meadow foxtail Intro Pres Jun–Jul shortawn foxtail mountain bentgrass redtop Intro Pres Jun–Sep ticklegrass Native Rep Jul–Aug spike bentgrass Native Hist Jul–Aug crested wheatgrass soft-stem bulrush three-square bulrush pale bulrush Native Pres Jun–Aug X alkali bulrush Native Pres Jun–Sep blue gramanodding brome Native Native Rep Pres Jul–Sep Jun–Aug hardstem bulrush purple three- awn Common name Nativity few-ﬂ spikerush

var.

validus acutus ssp. ora ssp. ssp. cristatum Andropogon gerardii ssp. lacustris Schoenoplectus pungens maritimus paludosus quinqueﬂ lacustris longiseta

ora Eleocharis Species name var. gerardii Andropogon gerardii var. Alopecurus pratensis Alopecurus pratensis Alopecurus aequalis Alopecurus aequalis Agrostis variabilis Agrostis variabilis Agrostis stolonifera Agrostis stolonifera Agrostis scabra Agrostis scabra Agrostis exarata Agrostis exarata Agropyron cristatum Agropyron cristatum Scirpus validus Schoenoplectus Scirpus pungens longispicatus Scirpus maritimus Bolboschoenus Scirpus pallidus Scirpus pallidus Bouteloua gracilisBromus anomalus Chondrosum gracile Bromopsis porteri Utah Colorado Eleocharis quinqueﬂ Scirpus acutus Schoenoplectus Aristida purpurea Aristida purpurea rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Cyperaceae Cyperaceae Cyperaceae Cyperaceae Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Conﬁ Family Perennial Graminoids, cont. Cyperaceae Cyperaceae Gramineae (Poaceae)

Chapter 6: Appendices 175 Disturbed areas Disturbed areas X X X Wet Comments XX X X X XX Mt Con XX XX PJ XX XXX XXX X XXX X XX XX XXX Shr Des Flower time Park status Native Pres Jun–Aug Native Pres Jun–Aug Native Pres Jun–Aug Native Pres Jun–Aug Native Hist Jul–Aug Native Pres May–Aug Native Pres Jun–Aug Salina wildrye Native Pres Jun–Jul quackgrass Intro Pres Jun–Aug squirreltailblue wildrye Native Pres Nativethickspike wheatgrass Pres May–Aug Jul–Aug desert saltgrass NativeCanada wildrye Pres Native Jun–Sep Great Basin Preswildrye Jul–Aug oneside oatgrass brookgrassorchard grass Native Intro PresCalifornia oatgrass Jun–Sep Pres Jun–Aug smooth brome IntroJones’ reedgrass Native Pres Presslimstem Jun–Aug reedgrass Jul–Aug mountain brome western wheatgrass Common name Nativity Calamagrostis scopulorum Species name Elymus salinus Leymus salina Elymus repens Elytrigia repens Elymus elymoides Elymus glaucus Elymus elymoides Elymus lanceolatus Elymus glaucus Elymus lanceolatus Distichlis spicata Elymus canadensis Distichlis stricta Elymus cinereus Elymus canadensis Leymus cinereus Danthonia unispicata Danthonia unispicata Catabrosa aquaticaDactylis glomerata Catabrosa aquatica Danthonia californica Dactylis glomerata Danthonia californica Bromus inermisCalamagrostis scopulorum Bromopsis inermis Calamagrostis stricta Calamagrostis stricta Utah Colorado Bromus carinatus Ceratochloa carinata Elymus smithii Pascopyrum smithii rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Perennial Graminoids, cont. Gramineae (Poaceae) (Poaceae) Appendix D. Conﬁ

176 Annotated Checklist of Vascular Flora: Dinosaur National Monument X X X Wet Comments XX XX X Mt Con PJ XXX X XXX XXX XXX XX XXX XX X XX XXX XX Shr Des Flower time Park status Native Pres Jul–Aug Native Pres Jul–Aug Native Pres Jun–Aug Native Pres Jun–Aug foxtail barley NativeJunegrass Presspike fescue Jun–Aug Native Nativeoniongrass Pres Pres Jun–Aug foxtail muhly Native Jun–Aug Native Pres Pres May–Aug Jul–Sep galletameadow barley Native Native Pres Pres Jun–Aug May–Aug fowl mannagrass meadow fescue Intro Pres Jun–Aug hairy tridens Nativetall fescue PresIdaho fescue Intro May–Jul NativeRocky Mountain Presfescue Pres May–Jul May–Aug slender wheatgrass bluebunch wheatgrass sandhill muhly Native Pres Jul–Sep scratchgrass Native Pres Jul–Aug Common name Nativity Muhlenbergia Critesion brachyantherum Festuca idahoensis Festuca saximontana spicata pungens asperifolia

Species name var. var. Muhlenbergia asperifolia Hordeum jubatumKoeleria macrantha Critesion jubatum Koeleria macrantha Leucopoa kingiiMelica bulbosa Leucopoa kingii Muhlenbergia andina Muhlenbergia andina Bromelica bulbosa Hilaria jamesiiHordeum brachyantherum Hilaria jamesii Glyceria striata Glyceria striata Festuca pratensis Festuca pratensis Erioneuron pilosum Erioneuron pilosum Festuca arundinacea Festuca arundinacea Festuca ovina ingrata Festuca ovina saximontana Elymus trachycaulus Elymus trachycaulus Utah Colorado Elymus spicatus Pseudoroegneria Muhlenbergia pungens Muhlenbergia rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Conﬁ Family Perennial Graminoids, cont. Gramineae (Poaceae) Gramineae (Poaceae)

Chapter 6: Appendices 177 Sandy soils X X X X Wet Comments X X Mt Con PJ XXX X XXX X XXX XXX X XXX X XX XXX X XXX XXX X XXX XXX X Shr Des Flower time Park status Intro Pres May–Aug Native Pres Apr–Aug Native Hist Jun–Aug Intro Pres Jun–Aug Intro Pres Apr–Jul Native Pres Jun–Sep Native Pres Jul–Sep Native Hist May–Aug Native Pres Jun–Aug Native Pres Apr–Sep Kentucky bluegrass Wheeler bluegrass muttongrass Nativeinterior bluegrass Pres May–Aug Canada bluegrass timothycommon reed Intro Nativebulbous Presbluegrass Pres Jul–Sep Jun–Sep little bluestem Nativeprairie Prescordgrass Jul–Sep bundle panicgrass Nuttall’s Nuttall’s alkaligrass mat muhly muhlyThurber’s Native Native Hist Preslittleseed ricegrass Jul–Sep Jul–Sep green muhly Native Hist Jul–Sep Sandberg’s Sandberg’s bluegrass Common name Nativity

ssp. Poa nervosa interior Schizachyrium scoparium acuminatum Muhlenbergia richardsonis thurberi micranthum Muhlenbergia racemosa Species name

glauca Poa nemoralis var. var. Poa pratensis Poa pratensis Poa nervosa wheeleri Poa fendlerianaPoa glauca Poa fendleriana Poa compressa Poa compressa Phleum pratensePhragmites australis Phleum pratense Phragmites australis Poa bulbosa Poa bulbosa Schizachyrium scoparium Spartina pectinata Spartina pectinata Panicum acuminatum Dichanthelium Puccinellia nuttalliana Puccinellia airoides Utah Colorado Muhlenbergia richardsonis Muhlenbergia thurberi Muhlenbergia Oryzopsis micrantha Piptatherum Muhlenbergia racemosa Poa secunda Poa secunda rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Family Perennial Graminoids, cont. Gramineae (Poaceae) Appendix D. Conﬁ

178 Annotated Checklist of Vascular Flora: Dinosaur National Monument X X X X X Wet Comments XX Mt Con PJ X XXX XXX XXX XXX XX XXXX X XX Shr Des Flower time Park status Intro Pres Jun–Jul Native Pres Jun–Jul Native Pres Jun–Sep Native Pres Jul–Sep Native Pres May–Jul Native Rep May–Jul Native Hist May–Aug Tracy’s rushTracy’s Native Pres Jun–Aug compressed rush Baltic rush Native Pres May–Jul spike trisetum Native Pres Jul–Sep green needlegrass Nelson’s Nelson’s needlegrass Indian ricegrass Native Letterman’s Presneedlegrass May–Jul needle-and- thread spike dropseed Nativesand dropseed Pres Native Jun–Sep Mormon Presneedlegrass Jun–Aug alkali sacaton Native Hist Jun–Aug longstyle rushpoverty rush Native rushTorrey’s Pres Nativeseaside Native Jun–Aug Presarrowgrass Hist Jun–Aug Jun–Aug X X X mountain rush Native Pres Jun–Aug Common name Nativity

ssp. Juncus ensifolius ater Achnatherum nelsonii hymenoides lettermanii Hesperostipa comata cryptandrus Sporobolus airoides Juncus saximontanus

var.

var. var.

Species name var. var. Juncus ensifolius Juncus ensifolius brunnescens Juncus compressus Juncus compressus Trisetum spicatumTrisetum Juncus arcticus spicatum Trisetum Juncus arcticus Stipa viridula Nassella viridula Stipa nelsonii nelsonii Stipa hymenoidesStipa lettermanii Achnatherum Achnatherum Stipa comata comata Utah Colorado Sporobolus contractus Sporobolus contractus Sporobolus cryptandrus Sporobolus Stipa arida Achnatherum aridum Sporobolus airoides airoides Juncus tenuisJuncus torreyi maritimaTriglochin Juncus tenuis maritima Triglochin Juncus torreyi Juncus longistylis Juncus longistylis montanus rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Juncaceae Juncaceae Juncaceae Gramineae (Poaceae) Juncaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Appendix D. Conﬁ Family Perennial Graminoids, cont. Gramineae (Poaceae) Juncaceae Juncaceae Juncaginaceae Juncaceae

Chapter 6: Appendices 179 Disturbed areas Disturbed areas Disturbed areas X X X X X X X X Wet Comments Mt Con PJ XX XXX X XXX XX XXX Shr Des Flower time Park status Native Pres NA Intro Pres May–Jul Native Pres Jul–Aug Native Hist NA Native Pres Jun–Aug Native Hist NA Native Pres Jun–Aug Native Pres NA at- eld horsetail eld Native Pres NA X X X X tall scouring- rush minute rushtoad rush Native Repﬁ Native Jun–Jul Pres Jun–Aug X X X X six-weeks fescue Native Preswitchgrass Apr–Jun rabbitfoot-grass Native Intro Pres Pres Jun–Sep Jun–Aug annual wheatgrass cheatgrasstufted lovegrass Intro Native Hist Pres Jul–Aug May–Jul Japanese chess Intro Pres Jun–Aug American sloughgrass hybrid scouring- rush bearded ﬂ sedge Ferris’ scouring- rush broad-leaved cattail smooth scouring-rush Common name Nativity

var. ferrissii x x ora nis afﬁ laevigata Hippochaete hyemalis ssp. occidentale Polypogon monspeliensis triticeum syzigachne Hippochaete mackaii

var. Species name oraoctofl Vulpia ferrissii Hippochaete x x ne Equisetum laevigatum Hippochaete Festuca octofl affi Juncus bufoniusEquisetum arvense Juncus bufonius Equisetum hyemale Equisetum arvense Juncus bryoides Juncus bryoides Panicum capillarePolypogon Panicum capillare monspeliensis Eremopyrum triticeum Eremopyrum Bromus tectorumEragrostis pectinacea Anisantha tectorum Eragrostis pectinacea Bromus japonicus Bromus japonicus Beckmannia syzigachne Beckmannia Equisetum trachyodon Cyperus squarrosus Cyperus aristatus Equisetum Utah Colorado Typha latifoliaTypha latifolia Typha rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Equisetaceae Gramineae (Poaceae) Juncaceae Ferns and Allies Equisetaceae Equisetaceae Juncaceae Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Gramineae (Poaceae) Equisetaceae Annual Graminoids Cyperaceae Equisetaceae Family Perennial Graminoids, cont. Typhaceae Appendix D. Confi

180 Annotated Checklist of Vascular Flora: Dinosaur National Monument Rock crevices Rock crevices Rock crevices Rock crevices Weber F. Weber X X Wet Comments XX Mt Con PJ XXX XXX XXX XX XXX XXX Shr Des Flower time Park status Native Pres NA Native Pres NA Native Pres NA Native Pres NA Native Pres NA Native Pres NA Native Pres NA Native Pres NA blunt-leaf spike- moss dense spike- moss Oregon woodsia western cliffbrake Brewer’s cliff- Brewer’s brake slender lip-fern Native Presbrittle bladder fern NA forked spleenwort southern maidenhair fern Common name Nativity

ssp. ssp. cathcartiana Pellaea glabella simplex Asplenium septentrionale Adiantum capillus- veneris

Species name var. Selaginella mutica Selaginella mutica Selaginella densa Selaginella densa Woodsia oregana Woodsia oregana Woodsia Pellaea glabella occidentalis Pellaea breweri Pellaea breweri Cheilanthes feeiCystopteris fragilis Cheilanthes feei Cystopteris fragilis Asplenium septentrionale Utah Colorado Adiantum capillus- veneris rmed, historical, and reported taxa of Dinosaur National Monument, organized by life form with ecological notes, cont. Selaginellaceae Selaginellaceae Polypodiaceae (Aspleniaceae, Woodsiaceae) Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Adiantaceae, Sinopteridaceae) Polypodiaceae (Aspleniaceae) Polypodiaceae (Aspleniaceae) Appendix D. Conﬁ Family Ferns and Allies, cont. Polypodiaceae (Adiantaceae)

Chapter 6: Appendices 181

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