FREDERICK H. RINDGE,

BULLETIN~ OF THE. AMERICAN MUSEUM. OF NATURAL HISTORY VOLUME -135: ARTICL 3 N,EW YORK.: 1967

A REVISION OF THE NEOTROPICAL SPECIES OF THE MOTH GENUS GLENA (LEPIDOPTERA, GEOMETRIDAE)

FREDERICK H. RINDGE Curator, Department of Entomology The American Museum of Natural History

BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY VOLUME 135 : ARTICLE 3 NEW YORK: 1967 BULLETIN OF THE AMERICAN MUSEUM OF NATURAL HISTORY Volume 135, article 3, pages 107-172, figures 1-50 Issued Feburary 27, 1967

Price: $1.50 a copy CONTENTS INTRODUCTION. 111 Materials and Methods . 111 Acknowledgments...... 112 SYSTEMATIC DESCRIPTIONS ...... 113 Genus Glena Hulst ...... 113 Key to Species Based on Male Genitalia and Secondary S5exu,al Characters . . . 115 Glena effusa, New Species . 116 Glena bipennaria (Guenee), New Combination . 119 Glena bipennaria bipennaria (Guen6e) 119 Glena bipennaria demissa, New Subspecies ...... 120

Glena dentata, New Species...... 121

...... 121 Glena grandillosa (Dognin), New Combination . . .

Glena sucula, New Species ...... 123

Glena vesana, New Species ...... 124

Glena bisulca, New Species...... 124 Glena juga, New Species ...... 125 Glena megale, New Species ...... 127 Glena unipennaria (Guenee), New Combination ...... 128 ...... 128 Glena unipennaria unipennaria (Guenee). .

...... 129 ...... Glena unipennaria cosmeta, New Subspecies. . Glena uncata, New Species ...... 130 Glena demissaria (Walker), New Combination ...... 131 Glena basalis, New Species ...... 133

Glena hima, New Species...... 133 Glena lora, New Species ...... 135 Glena agria, New Species...... 135 Glena trapezia, New Species...... 137

. . . . 138 Glena sacca, New Species......

Glena bulla, New Species. I ...... 139 Glena gemina, New Species...... 139 Glena laticolla, New Species...... 140

...... 141 Glena brachia, New Species...... Glena cretacea (Butler), New Combination ...... 143 Glena gampsa, New Species...... 144 Glena labecula, New Species ...... 144 Glena totana, New Species ...... 145 Glena quadrata, New Species ...... 146 Glena mopsaria (Schaus), New Combination ...... 147 Glena turba, New Species...... 148 Glena tyrbe, New Species...... 148 Glena asaccula, New Species ...... 149 LIST OF THE NEOTROPICAL SPECIES OF THE GENUS Glena, WITH THEIR...... KNOWN DISTRIBU- TION. 150 APPENDIX: SPECIMENS EXAMINED. 152 BIBLIOGRAPHY. 171

109

INTRODUCTION SEVERAL YEARS AGO I began working with the A.M.N.H., the American Museum of Natural geometrid moths of the genus Glena, paying History particular attention to the North American B.M.N.H., British Museum (Natural History), species. At that time preliminary work on the London Central and South American species showed U.S.N.M., United States National Museum, that much more time and labor were needed Smithsonian Institution, Washington, D. C. before the tropical taxa could be properly All the specimens studied by the author understood. The practical solution was to during the preparation of this paper have had publish separate papers on the two faunas; type or identification labels affixed. All too the one on the nearctic fauna was published often such labeling has not been done in the in 1965. The present paper is a revisional past, so that there is always the question of study of the neotropical species, in which I whether or not certain specimens were ex- have attempted to answer some of the ques- amined by a reviser. tions pertaining to their taxonomy, distribu- A total of 715 specimens were studied. A tion, and phylogeny, and to propose a more large number of genitalic dissections were pre- satisfactory systematic arrangement of the pared by the author, who also had at his dis- taxa. posal slides made by J. L. Sperry at the As now understood, Glena includes 31 American Museum of Natural History, and species from the neotropical area. Two of by H. W. Capps and E. L. Todd of the United these are polytypic, having two subspecies States National Museum. In all, 409 dissec- each; the remainder are monotypic. Six tions were examined. names are available for this group, all repre- senting valid species; none has heretofore LOCALITY DATA been published in Glena. Two of these names The locality data are briefly summarized were proposed by Guenee in 1857, and one under the heading of Distribution for every each was given by Walker (1868), Butler species. The complete information for the (1881), Dognin (1902), and Schaus (1913). In previously described species is contained in this revision 33 specific and subspecific names the Appendix, and that for the new species are recognized; 27 of them are proposed as and subspecies in the list of type material for new. that taxon in the text. Unfortunately many of the labels did not MATERIALS AND METHODS include adequate data and, in many cases, had very little information. The index to the MATERIALS STUDIED Map of Hispanic America of the American This revision is based primarily on the Geographical Society (1945) served as the specimens in the collections of the American main source of information in the obtaining of Museum of Natural History, the British additional facts. Whenever possible, the de- Museum (Natural History), and the United partment, state, or province is given for the States National Museum of the Smithsonian larger countries; if not indicated on the label, Institution. Types of three of the six known it is inserted in brackets in the text and Ap- species have been studied by the author. The pendix. This task was lightened immeasur- genitalia of the remaining three, in the collec- ably for Ecuadorian localities by the paper of tion of the British Museum (Natural His- Brown (1941), and for Central America by tory), have been dissected by Mr. D. S. the one of Selander and Vaurie (1962). Fletcher; photographs of these preparations In some instances it was possible to corre- have been examined by the author. late the distributional data of a species with The genitalic drawings were prepared from the climatic zones of South America. The specimens taken from the above-mentioned source of terms of these zones is the "Life collections. The following abbreviations have pictorial atlas of the world" (Time, Inc., been used: 1961). 111 112 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 PHYLOGENY abdominal segment, the presence of the The nearctic taxa of Glena were divided large, single spine of the vesica of the aedea- into three species groups, based largely on the gus, a broad ductus bursae, and a flat signum. secondary sexual characters of the male, wing When considered from a geographical point shape, pattern, and the nature of the arma- of view, the more primitive species of Glena ture of the aedeagus. As these criteria were are found in the United States and southern also followed when the neotropical species Canada. The species of group I are primarily were studied, the order in which they appear neotropical in distribution, although three in this paper is the same. When I completed species are found in the southwestern United my studies of the North American moths, a States and northwestern Mexico. Eight few tentative conclusions were proposed re- species are restricted to Mexico and Central garding evolutionary changes within this por- America, and one extends from Mexico into tion of the genus, using the above characters. Bolivia. In South America, 12 taxa are found Since the completion of the study of the along the Andes Mountains, usually at rather North American Glena, a revision of the low elevations, and seven are from the Humid species of the genus Anacamptodes Mc- Subtropical Zone. The remaining several Dunnough has been finished (Rindge, 1966). species are found in Trinidad (one), the The latter genus is closely allied to Glena, and Guianas (two), the Amazon Basin (one), and some of the characters within it show definite widespread throughout much of South and unmistakable evolutionary trends. These America east of the Andes (one). trends are the same as those found in the re- lated and very large genus Melanolophia ACKNOWLEDGMENTS Hulst (Rindge, 1964), and are just the op- posite of the ones given for the northern The author wishes to acknowledge with species of Glena. This fact, plus a knowledge thanks the cooperation and aid of the follow- of the tropical species of Glena, led to a re- ing colleagues who have allowed him to study evaluation of these characters, and I now the types and specimens in their charge: Dr. believe that my tentative conclusions were J. F. G. Clarke of the Smithsonian Institution; wrong. What I had considered as being primi- Mr. D. S. Fletcher of the British Museum tive characters are, in fact, the more highly (Natural History); Dr. E. L. Todd of the evolved ones. As a result, the species of Insect Identification and Parasite Introduc- groups II and III are actually the more primi- tion Research Branch, United States De- tive, whereas those of the much larger group partment of Agriculture; and Dr. P. E. L. I are the more advanced. Viette of the Museum National d'Histoire The following characters are the more Naturelle, Paris, France. highly evolved: moths of large size with The author wishes to thank Mr. Thomas whitish wings, the presence in the males of Hayden of the Graphic Arts Department of both the hair pencil on the hind tibia and the the American Museum of Natural History medioventral row of bristles on the third for preparing the drawings of the genitalia. SYSTEMATIC DESCRIPTIONS GENUS GLENA HULST those of the first group, and have broad, Boarmia auct. nec Treitschke: DRUCE, 1892 black, gray, or grayish brown wings, and lack (1891-1900), p. 72. the single, very large spine of the vesica. The Glena HULST, 1896, p. 358. RINDGE, 1965, p. definition of group III is unaffected. For the 269 (bibliography, synonymy, description). New World, the first group now contains a Catoria auct. nec Moore: PROUT, 1929, p. 132. total of 34 species, the second but three taxa, As the genus was completely described for and five species are included in group III. the revision of the nearctic species (Rindge, A study of the male genitalia of the neo- 1965), this information is not repeated here. tropical species necessitates but few changes However, the present study shows that my in the generic description for these organs. generic description must be slightly modi- The uncus ends in a single point in the ma- fied to include all the species now placed in jority of the species. Only four of the known Glena. species covered in this revision have two The males may or may not have a fovea at points on the uncus, as compared with all 11 the base of the forewing. All the nearctic of the nearctic taxa that have the latter con- species have this character; in the neotropical dition. The valves are basically similar in the region, 18 species possess the fovea, 11 do not, species of the two faunas, although more and and the males of the remaining two taxa are varied modifications occur in the more unknown. numerous tropical taxa. Seven species have In my revision of the nearctic fauna, the the apex of the costa sharply enlarged. There species were divided into three groups based is considerable variation in the combined on, among other characters, the presence or lengths of the sacculus and its process, rang- absence of both the hair pencil on the hind ing from 1.3 to 3.4 mm. In one of the shortest tibia and the bristle comb on the venter of of these the process is vestigial, and its func- the third abdominal segment in the male. tion is apparently assumed by what appears Continuing studies in this tribe of moths, to be a lateral extension of the transtilla. Two both on Anacamptodes (Rindge, 1966) and species are unique in the genus in that they the species included in this paper, show that have a group of heavy spines at the base of these probably sex-linked characters are of the costa. All the neotropical species have an more value on the specific level than as group elongate spine in the vesica of the aedeagus; characters. The males of 29 of the neotropical in the nearctic area, only three of the 11 taxa species are known; three of these lack both possess this character. the tibial hair pencil and bristle comb, and it The female genitalia of the neotropical is probable that one of the two unknown species show that there is a greater range of males will not have either of the characters. variability in these organs than in the ones These few taxa are so closely allied to the from the nearctic region, as would be ex- others in the genus that it does not seem pected, because of the larger number of reasonable to separate them into two groups, species involved. The variability manifests as was done for the nearctic species. This itself primarily in the sterigma, in the form change should also apply to the northern and position of origin of the ductus seminalis, species, so arcana Rindge should be trans- and in the form and location of the signum. ferred from group II to group I. The latter The sterigma shows variation in its compo- group includes all the neotropical species and nent parts, and there is also some variability three nearctic ones (grisearia Grote, fur- in the intersegmental area between A7 and furaria Hulst, and arcana). Group I is re- A8. The ductus seminalis may arise ventrally, defined as containing the larger species, all laterally on the left side, or dorsally, and it with the upper surface of the wings white or comes off the corpus bursae either as a large whitish gray, and having the very large spine sac, a swollen tube, or as a slender tube. The in the aedeagus. Group II is now restricted to signum is absent from one species; in the moths that are usually smaller in size than other taxa it may be found ventrally, 113 114 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 laterally on the left side, or dorsally. It is fortunately most of the species are known either rounded in shape, or else it is a more or from so few examples that figures on sex less elongate, transverse ellipse, with the ratios would be meaningless. Only 12 of the outer margin being either spinose or smooth. 31 species are represented by 10 or more The upper surface of the wings of all the specimens. Of this dozen, the sex ratio varies neotropical species is of an even white or from one male to three females (uncata) to grayish white color, variably overlain with the other extreme of nine males to one female brown or gray scales. The cross lines are rep- (unipennaria cosmeta). The ratios of males to resented by black or very dark dots, and females vary: 1/1 (agria), 2.5/1 (mopsaria), these may be either unconnected or con- 3/1 (quadrata), 4/1 (juga, totana, demissaria, nected. The hind wings are concolorous with and unipennaria unipennaria), 5/1 (grandil- the forewings, and they have the same type of losa and brachia), 6/1 (effusa), and 7/1 maculation as is found on the primaries. The (bipennaria bipennaria). Improved or modi- under surface of the wings may be white or fied collecting techniques are called for to gray, and in some species the wings are find the females of some of the species. heavily suffused with dark brown or grayish References in the literature to species of black scales; in some of the taxa the fore- this genus are usually completely unreliable, wings are darker in color than the hind wings. owing to the great similarity in size, color, The under surface is usually without macula- and maculation of all the taxa. None of the tion except for a broad, dark terminal band of earlier workers used the genitalia as an aid in varying extent. classification. An example is "Boarmia demis- The early stages are completely unknown saria Walker" as used by Druce [1892 (1891- for all the species included in this paper. 1900), p. 75]. Fortunately his material was The type species for Glena is cognataria labeled, and most of it has been studied by Hiibner; this moth is found in the eastern the present author. The above category con- United States. tained eight different species, none of them The range of the species covered in this re- being demissaria. Seven are included in this vision is from southern Mexico, through paper, with six being described as new species. Central America, and throughout most of The last taxon is represented by the single South America, with the exception of Chile female from Pinos Altos, Chihuahua and southern Argentina. (Buchan-Hepburn), which is referable to fur- The tropical species of Glena form a dis- furariafurfuraria (Grote), and represents the tinct and easily recognized group within the first known Mexican record for this species. Cleorini (McDunnough, 1920) of the sub- Unless the specific specimen has been person- family Ennominae. The adults can usually be ally examined, the literature records are not distinguished by the more or less uniform included in this paper. color and pattern that are present in every In nearly every case a dissection of the species. Specific determinations are com- genitalia is necessary for making specific de- plicated by both geographical and seasonal terminations. These structures are large and variability. Many of the species are repre- easy to work with, so few difficulties are to be sented by inadequate material, so our encountered when dissecting them. The knowledge of the variation is rather sketchy aedeagus is large, so it is strongly recom- at the present time. mended that the vesica be exserted during the There is some sexual dimorphism in this dissection, a procedure that will show several genus, as the females tend to have more dark characters that are otherwise difficult to see scaling on both surfaces of the wings than is and that are of specific importance. found in the males. The pattern of the upper Because the color and pattern of the wings surface is usually less strongly indicated, are so similar in all of the species, it is not whereas the terminal band of the lower sur- feasible to make a key to the adults based on faces is more strongly represented in the these characters. In addition, the females of females. one-third of the species are unknown, so a key Females are usually much fewer in number based upon the genitalia of this sex is not very in the collections studied than are males. Un- practical at this time. However, all the known 1967 RINDGE: GLENA 115 structures of this sex are described and fig- of length of costa; combined lengths of ured. The male genitalia offer good characters sacculus and process, 2.8 mm. for the recognition of the various species, and .l...... labecula a key to these organs has been prepared. Process of sacculus extending to middle of The species described as Boarmia nepia costa; combined lengths of sacculus and process, 1.5 to 1.9 mm...... 12 Druce [1892 (1891-1900), p. 76] was sus- 12(11). Apical section of process of sacculus pected of belonging to Glena (McDunnough, globular, with posteriorly directed 1920, p.24; Rindge, 1965, p. 283).An examina- spines in single or double row on poste- tion of photographs of the genitalia of the fe- rior margin of process . . . . . totana male type of nepia proves it to be a member of Apical section of process of sacculus the genus Anavitrinella McDunnough. elongate, slender, and curved, with multiple rows of outwardly curved KEY TO SPECIES BASED ON MALE spines arising on posterior one-half of GENITALIA AND SECONDARY process.... quadrata SEXUAL CHARACTERS1 13(7). Spine of vesica with recurved base . .14 1. Venter of third abdominal segment with Spine of vesica straight, without recurved median row of bristles ...... 2 base.... trapezia Venter of third abdominal segment with- 14(13). Anellus very broad, with width almost out median row of bristles . .. 27 equal to length ...... 15 2(1). Uncus terminating in single point . . 3 Anellus slender, with length at least twice Uncus terminating in two widely sepa- as great as width ...... 18 rated points ...... 25 15(14). Exserted vesica with lateral, bifurcate 3(2). Valves with dense patch of heavy spines caecum... sacca at base of costa ...... 4 Exserted vesica without caecum . . . 16 Valves without basal patch of spines . 5 16(15). Process of sacculus with spines on distal 4(3). Anellus with length twice as great as one-half only ...... 17 width ...... turba Process of sacculus with two widely sepa- Anellus with length equal to width . tyrbe rated spine patches, one basally and 5(3). Valves with elongate process of sacculus other terminally . . . . . grandillosa two ...... 6 17(16). Process of sacculus with spine patches Valves with rudimentary process of sac- ...... bisulca culus; middle of valve with lateral, Process of sacculus with single, terminal apically spinose extension of transtilla spine patch ...... juga Recurved base of of vesica ...... asaccula 18(14). spine very 6(5). Valves with costal arm attached at base short, being one-fifth to one-seventh of only ...... brachia length of spine ...... effusa Costa an integral part of valve. . . . 7 Recurved base longer, being from one- 7(6). Uncus triangular; exserted vesica tubular fourth to one-half of length of spine and usually having sclerotized, shortly ...... 19 . spinose dorsal strip ...... 8 19(18). Vesica with narrow, elongate, dentate Uncus elongate; exserted vesica not tubu- patch on dorsal surface, and with larger lar and without dorsal strip . . . .13 dentate area ventrally . . . . agria 8(7). Process of sacculus as long as, or longer Vesica not as above ...... 20 than, costa ...... 9 20(19). Recurved base of spine in vesica one- Process of sacculus shorter than costa fourth to one-third of length of spine

...... 10 ...... 21. 9(8). Process of sacculus of same length as costa Recurved base of spine one-third to one- ...... gampsahalf of length of spine ...... 22 Process of sacculus longer than costa . 21(20). Process of sacculus with apex not swollen

...... cretacea and with terminal two-fifths to one-half 10(9). Spines on terminal portion of process of spinose, the spines either in one group sacculus situated laterally . mopsaria or with median interruption .

Process of sacculus with terminal spines .bi...... bipennaria

...... 11 Process of sacculus with apical one-third 11(10). Process of sacculus extending two-thirds swollen and covered with spines .

...... dentata. 1 The males of basalis and lora are unknown. 22(20). Anellus strongly bilobed; uncus with ven- 116 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

tral margin concave between extended grayish white, with variable amount of pale

ventrolateral margins . . . . megale brown scaling and with tarsus and tibia of Anellus not bilobed although with small forelegs of many specimens dark brown or median incision on posterior margin; brownish black, with narrow pale gray bands uncus not as above ...... 23 at ends of segments; tibia of hind leg with 23(22). Anellus with prominent constriction an- teriorly and with raised lateral margins hair pencil. Abdomen white above, with vari- ...... unipennariaable amount of pale brown scaling and with Anellus without above characters .24 paired dark brown or brownish black spots 24(23). Length of spine in aedeagus, 1.4 to 2.0 posteriorly on segments As and A4; below dull

.... . demissaria grayish brown. Length of spine in aedeagus, 1.0 to 1.1 UPPER SURFACE OF WINGS: Forewings with mm...... uncata small fovea at base; ground color white, over- 25(2). Length of sacculus and its process, 1.3 lain with variable number of pale brown or mm...... gemina grayish brown scales and strigations; cross Length of sacculus and its process, at least lines appearing as brownish black spots on 2.0 mm...... 26 on t. 26(25). Process of sacculus spinose for most of costa and veins, with a., median, and length ...... bulla t. p. lines represented; discal spot absent; t. p. Process of sacculus with spines on outer line shaded distally by pale grayish brown in one-third or one-fourth only cells M2, M3, and above anal margin; s. t.

...... laticolla line represented by large, dark brownish gray 27(1). Recurved base of spine in vesica very intravenular spots, with spots in cells R5 and short, not exceeding 0.2 mm. . .28 Ms absent from some specimens; terminal line Recurved base of spine one-third of length of prominent, blackish brown, intravenular of spine, 0.5 to 0.6 mm. in length spots; fringe concolorous with wing, with some ...... hima pale brownish gray scaling. Hind wings con- 28(27). Terminal portion of process of sacculus slender, bearing spines in two groups; colorous with forewings and with same type uncus 0.6 mm. in length; spine of of maculation, with intradiscal, extradiscal, and s. t. lines discal spot brown- aedeagus weakly curved . . . sucula represented; Terminal portion of process of sacculus ish black, elongate; terminal line and fringe capitate, bearing spines in one or two like those of forewings. groups; uncus 0.7 mm. in length; spine UNDER SURFACE OF WINGS: Pale gray, of aedeagus with strong curve . vesana heavily and evenly overlain with dark grayish brown on and with brown Glena effusa, new species forewings grayish on hind wings; discal dots absent from fore- Figures 1, 30 wings, weakly represented on hind wings; Boarmia demissaria auct.: DRUCE, 1892 (1891- without additional maculation except for 1900), p. 75 (partim). broad blackish gray terminal band on all Boarmia unipennaria auct.: OBERTHER, 1913, wings, with white apex on primary and with p. 278 (partim), p1. CLXIX, fig. 1656. narrow white band at wing margin; terminal This is the commonest and most widely line represented by dull black or grayish ranging species of the genus, as it occurs from black intravenular spots; fringe white. Mexico to Bolivia. It is one of the largest LENGTH OF FOREWING: 18 to 25 mm.; taxa, and the under side of the wings has a holotype, 22 mm. broad, dark terminal band. The genitalia FEMALE: Similar to male, but tending to should be used for identification. have upper surface of wings more irrorate MALE: Head with vertex white; front with dark scales, and to have t. p. and ex- brownish gray, whitish gray across top and tradiscal lines more completely represented bottom; palpi blackish brown, with terminal on upper surface of wings. segment becoming whitish gray or pale LENGTH OF FOREWING: 19 to 26 mm.; grayish brown. Thorax white above, with allotype, 23 mm. scattered light brown scales and with dark MALE GENITALIA: Uncus triangular, sides brown scaling at apex of collar and posteriorly more or less straight, with apical portion slen- on thorax; below dull grayish brown; legs der and laterally compressed; gnathos with 1967 RINDGE: GLENA 117 median enlargementvariable in shape, ranging swellings or denticulations; signum with flat from relatively broad, with apical region or weakly concave anterior margin, rounded rounded, to slender and tapering, with apical posteriorly, outer margin with numerous region bluntly pointed; valves with costa pointed projections, anterolateral ones larg- sclerotized, apical one-fourth slightly swollen est, those along anterior margin smallest and and with numerous anteromedially directed with flat median area recessed and bearing setae; valvula with poorly defined, lightly several small spinose processes. Posterior por- sclerotized strips extending length of valve; tion of ventral surface of A7 thickly covered sacculus swollen, sclerotized, tapering in with minute, sclerotized, globular swellings, width distally, extending about one-half of and having numerous short, longitudinal length of valve, with process arising from folds. distal portion; process long and slender, TYPES: Holotype, male, Avispas, Madre de slightly longer than costa, projecting short Dios, Peru, September 20-30, 1962 (L. E. distance beyond valvula, ribbon-like, without Pena); genitalia mounted on slide F.H.R. spines except for five or six at tip of process; No. 12337. Allotype, female, Quincemil, length of sacculus and process varying from Cuzco, Peru, September 1-6, 1962 (L. E. 2.7 to 3.4 mm.; cristae represented by large, Pefia); genitalia mounted on slide F.H.R. elongate patch of numerous slender setae No. 12685. Paratypes: Bolivia: Chapare, arising from area at base of sacculus; anellus [Beni], various dates in August and Decem- constricted medially, with large, smoothly ber, 1949 (L. E. Pefia), June 6, 1948, June 5, sclerotized base, very wide anteriorly, sharply 22 males and one female; Region Chapare, tapering medially, enlarging posteriorly into [Beni], July, 1951, one male; Yungas del large fanlike process, with prominent ridge on Palmar, elevation 1000 meters, October 15, each side extending from posterior margin to 1949, three females; San Ernesto, [La Paz], median constriction; vesica with very long elevation 1000 meters, August-September, spine, basal end sharply bent and very short, 1900 (Simons), three males, "saison seche," with terminal portion broadly curved and one male; Salampioni, elevation 800 meters, having numerous small teeth on inside of August, 1901, dry season (Simons), one male; curve; length of spine varying from 1.1 to R[io] Songo to R[io] Suapi, [La Paz], eleva- 1.4 mm., of recurved base about 0.2 mm.; tion 1100 meters, March-June, 1896 (Gar- vesica with weakly sclerotized, oval area lepp), one male; Songo (Gar[lepp]), one male; posteriad and away from base of spine on Rio Songo, elevation 750 meters (Fassl), one dorsal surface bearing short deciduous spines, male; Charaplaya, [Cochabamba], elevation and with large, narrow, lightly sclerotized 130Q meters, June, 1901 (Simons), one male; areas bearing numerous short spicules on Buenavista, elevation 750 meters, August, both ventral and dorsal surfaces of vesica 1906 to April, 1907 (Steinbach), one male; under terminal portion of spine. Ventral Yunga del Espiritu Santo, Cochabamba, surface of As with median row of bristles. 1888-1889 (P. Germain), five males; Yungas FEMALE GENITALIA: Sterigma with large, de la Paz, elevation 1000 meters, one male; no smoothly sclerotized, elliptical or oval further data, one male. Peru: Quincemil, lamella postvaginalis, and with large, sclero- Cuzco, elevation 2400 feet, August, Septem- tized lateral areas, their anteromedian mar- ber 1-6, October 20-30, 1962 (L. E. Pefna), gins extending over anterolateral portions of nine males, one female; Avispas, Madre de lamella postvaginalis; ductus bursae shorter Dios, September 10-30, 1962 (L. E. Pefia), than length of lamella postvaginalis; ductus two males; Coosnipata, Paucartambo, Cuzco, seminalis arising from large, rounded sac November 11 to December 4, 1951 (F. L. laterally on left side of corpus bursae, orig- Woytkowski), four males; Satipo, [Junin], inating between 1.1 and 1.4 mm. anteriad of June, 1948 (P. Paprzycki), one male; Chan- ductus bursae, with tubular portion coming chamayo, [Pasco], elevation 1000 meters, one off medially near corpus bursae; latter male; Chaquimayo, [Puno], elevation 2500- elongate, weakly swollen anteriorly, and with 3000 feet, September, 1910 (H. and C. posterior one-third weakly sclerotized and Watkins), one male; Cajon, Cuzco, Decem- striate, surface covered with very small ber, 1902 (Garlepp), one male; Huambo, 118 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 1889 (M. de Mathan), two males; Yahuar- females, with two males and two females mayo, [Puno], elevation 1200 feet, October- identified as "Boarmia demissaria Walk., November, 1910, March, 1911 (Watkin B. C. A. Lep. Het., Godman-Salvin Coll."; brothers), three males; San Gaban, elevation Coatepec, [Veracruz], two males; Paso San 2500 feet, March-April, 1913, two males, Juan, Veracruz, one female; Tamazunchale, two females; La Union, R[io] Huacamayo, San Luis Potosi, May 20, 1952 (Cazier, Carabaya, [Puno], elevation 2000 feet, No- Gertsch, Schrammel), one male. vember, 1904, wet season (G. Ockenden), The holotype and allotype are in the col- three males; Cushi, Huanuco, elevation lection of the American Museum of Natural 1820 meters, 1904 (W. Hoffmanns), one male; History; paratypes are in the collections of Pozuzu, Huanuco, elevation 800-1000 meters that institution, of the British Museum (W. Hoffmanns), two males. Ecuador: Dos (Natural History), and of the United States Puentes, Kilometer 99, [Chimborazo], ele- National Museum. vation 1700 feet, January, 1929 (W. J. DISTRIBUTION: From Mexico (the states Coxey), one male; Paramba, [Imbabura], of San Luis Potosi, Veracruz, and Chiapas) November-December, 1898, January-Au- through Central America to Venezuela, gust, 1899 (Flemming), two males; Canelos, Guyana, and Colombia (the departments of Rio Bobonaza, [Napo-Pastaza] (M. G. Pal- Norte de Santander, Santander, Caqueta, mer), one male. Colombia: "Ob. Rio Negro," and Magdalena), Ecuador (the departments [Norte de Santander], elevation 800 me- of Chimborazo, Imbabura, and Napo-Pas- ters (Fassl), three males, one female; Muzo, taza), Peru (the departments of Cuzco, Madre [Santander], elevation 400-800 meters (Fassl), de Dios, Pasco, Puno, and Huanuco), and two males, two females; Popoyan, [Caqueta] Bolivia (the departments of Beni, La Paz, (Lehmann), three males, 1896, one male; and Cochabamba). In South America this Minca, [Magdalena], elevation 2000 feet species has been taken at elevations ranging (H. H. Smith), one male, one female; from 1200 to 4300 feet. Rio Agnatal, elevation 1600 meters, October, The adults have been captured in the 1908 (Fassl), one male; no further data, four months of February, May, June, July, Aug- males. Guyana (former British Guiana): Rio ust, October, and December in Mexico and Demerara, one male. Venezuela: San Esteban, Central America, and in every month of the June, 1909 (S. M. Klages), one male; Suapure, year except February and May in Ecuador, April 20, 1899, one male; Las Quiguas, Este- Peru, and Bolivia. ban Valley, one female. Costa Rica: San REMARKS: A total of 138 specimens (117 Jose, [San Jose] (H. Schmidt), one male. Hon- males and 21 females) and 65 genitalic dis- duras: La Cambre, February 25-28, 1922 (J. sections (53 males and 12 females) have been Lienhart), three males. Guatemala: San Joa- studied. The adults of effusa usually can be quin, Vera Paz (Champion), one male, identi- recognized by their large size, and by the dark fied as "Boarmia demissaria Walk., B. C. A. coloration of the under surface of the wings, Lep. Het., Godman-Salvin Coll."; Cayuga, with the wide terminal band on all wings May (Schaus and Barnes), one male; Volcan with the sharply contrasting white fringe. Santa Maria, [Quezaltenango], July (Schaus Specimens from South America are rather and Barnes), one male. El Salvador: Santa uniform in size, with the exception of two Tecla, June 30, 1954 (M. Salazar V.), two males from Venezuela. Both of these are males. British Honduras: Columbia, Decem- smaller and have the upper surface of the ber, 1932 (J. J. White), one male. Mexico: wings darker in coloration than those of most Santa Anita, Chiapas, July 13-14, 1930 other examples. Specimens from Central (C. C. Hoffmann), two males; mountains America and Mexico tend to be slightly near Soconusco, Chiapas, August, 1931 (C. smaller than those from South America. Hoffmann), one male; Orizaba, [Veracruz], The male genitalia of this species can be 1937 (C. C. Hoffmann), one male; Presidio, recognized by the very long and slender pro- [Veracruz], July, 1951, one female; Cordoba, cess of the sacculus, which is unspined except Veracruz, October 5, 1924 (C. C. Hoffmann), for the very tip, and by the very short re- one female, (H. Rumeli), five males and four curved base of the long spine of the aedeagus. 1967 RINDGE: GLENA 119 There is some variation in the male geni- brownish black scales; forewings less elon- talia that seems to be associated with the gate; upper surface of wings with maculation geographical distribution of this species. darker and more strongly defined in most Specimens from Ecuador, Peru, and Bolivia specimens, with t. p. line more complete and tend to have the spine of the vesica with a having stronger, dark gray, distal shading; very short base, often appearing as if it were under surface with narrower terminal band, attached to a small sclerotized area, and to especially on secondaries, and with wings have a more sweeping curve to the posterior tending to be less heavily suffused with dark portion of the spine. Examples from Central gray scaling; hind tibia with hair pencil. America, Venezuela, and Colombia, on the LENGTH OF FOREWING: 18 to 24 mm. other hand, tend to have the spine with a FEMALE: Similar to male but tending to slightly longer base, without any trace of the have upper surface of wings more irrorate sclerotized attachment, and to have less of a with dark scales and to have less clearly de- curve posteriorly. Southern specimens tend to fined maculation. have the gnathos shorter and more truncate, LENGTH OF FOREWING: 21 to 23 mm. while many northern examples have a more MALE GENITALIA: Similar to those of elongate and pointed structure. Southern effusa, differing mainly as follows: gnathos specimens also tend to have a longer sacculus with more elongate median enlargement, and process than do those from Central slender and tapering in most specimens; America. valves with costa having outer one-third The females of effusa are unique in that the slightly enlarged; process of sacculus long posterior portion of the venter of the seventh and slender, slightly shorter than costa, abdominal segment is thickly set with extending short distance beyond valvula, minute, globular swellings. distal two-fifths to one-half spinose along inner side, spines directed posteriorly and Glena bipennaria (Guen6e), new combination tending to be curved around process, either Boarmia bipennaria GUENEE, 1857, p. 257. in one row or with median interruption; length of sacculus and process varying from This species is usually slightly smaller than 2.5 to 2.9 mm.; anellus with larger indenta- effusa, and the forewings are less elongate. tion posteriorly on midline, and appearing Most specimens have a slightly more promi- cleft anteriorly to median constriction; spine nent pattern on the upper side of the wings of vesica with longer recurved base, sharply and a narrower terminal band on the under pointed, and having terminal portion with surface of the hind wings. The genitalia more sinuous curve, posteriorly with small should be used for identification. teeth distributed around spine; length of This taxon occurs in two populations. The spine varying from 1.1 to 1.3 mm., of re- nominate one is found in southern South curved base from 0.3 to 0.4 mm.; vesica with America, and the second is known from weakly sclerotized oval area away from base French Guiana. of spine bearing short deciduous spines, with Glena bipennaria bipennaria (Guen6e) smoothly sclerotized area extending from re- curved base of spine, and with large, more or Figures 2, 31 less triangular area, decreasing in width Boarmia bipennaria GUENEE, 1857, p. 257. laterally, on dorsal surface, bearing very BOISDUVAL AND GUENEE, 1858, pl. 13, fig. 5. many short teeth, these increasing in size WALKER, 1860, p. 356. OBERTHYR, 1913, p. 278. anterolaterally; ventral surface of vesica with VIETTE, 1950, p. 203. weakly sclerotized area, its surface covered The nominate subspecies occurs in south- with extremely fine denticulation. Ventral ern South America, and it is larger and more surface of A3 with median row of bristles. heavily marked than the following popula- FEMALE GENITALIA: Sterigma with large, tion. sclerotized, elliptical or oval lamella post- MALE: Similar to that of effusa, differing vaginalis with more or less rough surface, mainly as follows: thorax and abdomen and with large, sclerotized lateral areas, their tending to be more heavily marked with anteromedian margins extending over an- 120 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 terolateral portions of lamella postvaginalis; long median invagination of the anellus, by intersegmental area between A7 and A8 reces- the fact that the spine of the aedeagus has a sed and convoluted ventrally; ductus bursae longer and more sharply pointed recurved about one-half of length of lamella post- base, and by the different ornamentation of vaginalis; ductus seminalis arising from large, the vesica. rounded sac ventrolaterally on left side of There is some variation in the spining on corpus bursae, originating between 0.6 and the process of the sacculus; this is apparently 0.7 mm. anteriad of ductus bursae, with tubu- correlated with the geographical distribution lar portion coming off medially near corpus of the specimens. Most specimens from Brazil bursae; latter elongate, weakly swollen an- tend to have both a slightly longer area of teriorly, and with posterior one-third to one- spining and to have it uninterrupted medi- fourth weakly sclerotized and striate, surface ally; examples from Paraguay, Bolivia, and covered with very small swellings or denticu- Peru tend to have a shorter area of spines lations; signum with flat or weakly concave that is usually interrupted to some degree in anterior margin, rounded posteriorly, outer or near the middle. margin with numerous pointed projections, The female genitalia of bipennaria are those along anterior margin smallest, and quite similar to those of effusa. These struc- with flat median area recessed and bearing tures in the present species can be recognized several small spinose processes. Posterior by the convoluted intersegmental membrane portion of ventral surface of A7 not modified. between A7 and A8 ventrally, and by the more TYPE: Guenee described bipennaria from posterior location of the ductus seminalis. one male and one female. The type male is in the Museum National d'Histoire Naturelle, Glena bipennaria demissa, new subspecies Paris; the female apparently has been lost. This species is smaller than nominate The genitalia of the type are mounted on bipennaria, and the wings are less strongly slide F.H.R. No. 13521. marked on the upper surface than are those of TYPE LOCALITY: Brazil. specimens of the nominate form. The present DISTRIBUTION: Southern Brazil (the states taxon is found in French Guiana. of Santa Catarina, Parang, Sao Paulo, MALE: Similar to that of nominate bipen- Guanabara, and Mato Grosso), Paraguay, naria but smaller, and with the upper surface the eastern portions of Bolivia (the depart- of the wings more suffused with pale grayish ments of Beni and Santa Cruz), Peru (the brown, and having less prominent macula- departments of Madre de Dios, Cuzco, Pasco, tion; under surface of all wings heavily suf- and Puno), and Ecuador (the province of fused with dark scales. Loja). (See Appendix for locality data of LENGTH oF FOREWING: 18 mm. (holotype). specimens examined.) This species occurs in FEMALE: Unknown. the Humid Subtropical Zone of South MALE GENITALIA: Similar to those of Bra- America. zilian examples but tending to have slightly The adults are probably on the wing during shorter area of uninterrupted spining on every month of the year, although no ma- process of sacculus. terial dated June has been examined. FEMALE GENITALIA: Unknown. REMARKS: A total of 65 specimens (57 TYPE: Holotype, male, French Guiana, ex males, including the type, and eight females) collection C. Bar, and in the collection of the and 40 genitalic dissections (33 males and British Museum (Natural History). The seven females) have been studied. This genitalia are on slide Geometridae No. 4805. species is very similar to effusa, but it has DISTRIBUTION: This subspecies is known shorter, less elongate forewings, and most only from French Guiana. specimens have a narrower terminal band on REMARKS: One specimen and one genitalic the secondaries below. dissection have been studied. This population The male genitalia of this species can be is more likely to be confused with demissaria separated from those of effusa by the more ex- (Walker), which flies in the same area, than tensive area of spining along the inner margin with nominate bipennaria. The genitalia of the process of the sacculus, by the very should be used for identification, as the two 1967 RINDGE: GLENA 121 species are easily distinguished by means of tion. Ventral surface of As with median row of these structures. bristles. FEMALE GENITALIA: Unknown. Glena dentata, new species TYPES: Holotype, male, Balzapamba, Bolivar, Ecuador, March-April, 1894 (M. de Figure 3 Mathan); genitalia mounted on slide Geomet- This species is similar to nominate bipen- ridae No. 4820; in the collection of the British naria in size and color, but the maculation of Museum (Natural History). Paratype, one the upper surface of the wings is less strongly male, Mesopotamia, Antioquia, Colombia, represented. The genitalia should be used for elevation 5000 feet; in the collection of the identification. This taxon occurs in Ecuador American Museum of Natural History. and Colombia. DISTRIBUTION: The western side of the MALE: Similar to that of bipennaria, dif- Andes Mountains in Ecuador (the depart- fering mainly as follows: upper surface of ment of Bolivar) and Colombia (the depart- wings more finely irrorate with light brown ment of Antioquia). The specimens were scales, and with cross lines more weakly in- captured at elevations of about 2100 to 5000 dicated; under surface more heavily suffused feet. The time of flight is unknown. with dark scales, with wider terminal band, REMARKS: Two specimens (both males) with larger white area at apex of forewing, and two genitalic dissections have been with small grayish white terminal spot in studied. cell Ms of forewings, and with wider white The male genitalia of this taxon are like terminal area on hind wings; hind tibia with those of bipennaria, but there are diagnostic hair pencil. differences in the shape and spining of the LENGTH OF FOREWING: 22 (holotype) to process of the sacculus, of the anellus, and of 24 mm. the armature of the vesica. FEMALE: Unknown. There are some differences between the two MALE GENITALIA: Similar to those of dissections of the male genitalia. The holo- bipennaria, differing mainly as follows: uncus type from Ecuador appears to have a longer with terminal portion tending to be more and more bluntly pointed gnathos, the pro- strongly compressed laterally and thicker; cess of the sacculus is shorter, and there are gnathos longer; valves with costa only weakly differences in the anellus, when compared enlarged apically; process of sacculus long with the Colombian specimen. The anellus of and slender, extending to edge of valvula, the latter appears flatter and narrower than apical one-third swollen and covered with does that of the former, and it does not have numerous, more or less appressed spines; the lateral ridges that are so prominent in the length of sacculus and process varying from Ecuadorian example. Perhaps these varia- 2.4 to 2.8 mm.; anellus with small anterior tions are individual, or perhaps they repre- constriction, gradually increasing in width sent geographical changes within this species; posteriorly as delimited by lateral ridges, but more material is needed to settle this ques- with actual sides of anellus more mem- tion. branous, convoluted, swollen, and with poste- rior margin rounded and with median in- Glena grandillosa (Dognin), new combination cision; spine of vesica S-shaped, with base Figures 4, 32 more sharply curved than apical region, latter with small teeth on inside of curve; length of Cymatophora ? grandillosa DOGNIN, 1902, p. spine varying from 1.3 to 1.4 mm., of re- 349. curved base 0.45 mm.; vesica with weakly Catoria unipennaria auct.: DYAR, 1913, p. 649. sclerotized area near recurved base of spine The adults of this species are similar to bearing short deciduous spines, and with those of bipennaria, but they tend to be elongate, tear-shaped area on dorsal surface smaller, to have slightly darker wings, and to bearing numerous teeth, these markedly in- have a more reduced terminal band on the creasing in size distally; ventral surface of under surface of the wings. The genitalia vesica covered with extremely fine denticula- should be used for identification. This taxon 122 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 occurs in Argentina, Bolivia, Peru, and Ecua- with extremely fine denticulation. Ventral dor. surface of A3 with median row of bristles. MALE: Similar to that of bipennaria, FEMALE GENITALIA: Sterigma with large, differing mainly as follows: head, thorax, and elliptical or oval lamella postvaginalis, abdomen tending to have more brown scales; smoothly sclerotized except for posterior front with wider white band across bottom; portion, and with large, sclerotized lateral upper surface of wings with more grayish areas, their anteromedian margins extending brown and brown scales, and with cross lines over anterolateral portions of lamella post- more weakly indicated; under surface less vaginalis, and having pleural areas outwardly heavily suffused with dark scales, especially swollen; intersegmental area between A7 and on hind wings, and with terminal band less As recessed and weakly convoluted ventrally; strongly represented, on secondaries greatly ductus bursae slightly less than one-half of reduced on or absent from some specimens; length of lamella postvaginalis; ductus semi- hind tibia with hair pencil. nalis arising from large, rounded sac ventro- LENGTH OF FOREWING: 18 to 23 mm. laterally on left side of corpus bursae, FEMALE: Similar to male but tending to originating between 0.7 and 0.9 mm. anteriad have upper surface of wings more irrorate of ductus bursae, with tubular portion coming with dark scales and to have less clearly de- off medially near corpus bursae; latter elon- fined maculation; under surface with terminal gate, weakly swollen anteriorly, and with band tending to be reduced farther, extending posterior one-third to one-fourth weakly only to middle of wing in some specimens. sclerotized and striate, surface covered with LENGTH OF FOREWING: 16 to 22 mm. very small swellings or denticulations; signum MALE GENITALIA: Similar to those of situated ventrally, relatively small, varying effusa, differing mainly as follows: uncus with in shape from an elongate ellipse to being sides tending to be slightly more concave, and widest anteriorly and having flat or weakly with laterally compressed apex more elon- concave anterior margin, rounded posteriorly, gate; gnathos with median enlargement outer margin with numerous pointed pro- longer, more pointed, and more heavily jections, those along anterior margin tending sclerotized; valves with terminal three-eighths to be widest in some specimens, and with flat of costa swollen; process of sacculus long and median area recessed and bearing several slender, extending to just beyond edge of small spinose processes. Posterior portion of valve; length of sacculus and process varying ventral surface of A7 not modified. from 2.5 to 3.1 mm.; process enlarged on in- TYPES: Dognin described grandillosa from ner side near base, this swelling covered with two females. One of these bears his type label, numerous heavy spines, medially slender and and has the genitalia mounted on slide naked, and with apex weakly capitate and E.L.T. No. 1370. This specimen is hereby covered with slender spines; anellus large, designated the lectotype; it is U.S.N.M. No. flat, more or less rectangular in outline but 32986. with weak anterior constriction, with lateral TYPE LOCALITY: Tucuman, Tucuman, and posterior margins more or less crenulate, Argentina. and having several small ridges extending DISTRIBUTION: Argentina (the provinces of basally; spine of vesica thick and of moderate Jujuy and Tucuman), Bolivia (the depart- length, with elongate, sharply angled base ments of Beni, Cochabamba, and La Paz), about one-half as long as spine, and with Peru (the departments of Cuzco, Pasco, terminal portion broadly curved anteriorly Puno, and Huancabamba), and Ecuador and without minute teeth; length of spine (the province of Loja). (See Appendix for varying from 1.0 to 1.2 mm., of recurved base locality data of specimens examined.) This from 0.4 to 0.6 mm.; vesica with weakly species is found along the eastern slopes of the sclerotized area away from base of spine Andes Mountains at elevations of from bearing numerous, short, deciduous spines, about 3000 to 6600 feet. with dorsal surface having subtriangular, Adults have been examined that have been sclerotized, dentate area; ventral surface with captured in every month except July and weakly sclerotized area, its surface covered November. 1967 RINDGE: GLENA 123 REMARKS: A total of 125 specimens (105 gray scaling, varying from being quite heavy males and 20 females, including the lecto- (holotype) to almost absent; most specimens type) and 44 genitalic dissections (35 males with large brown or brownish gray patch in and nine females) have been studied. Speci- cell M3 distad of t. p. line, and with s. t. line mens from Argentina, in the southernmost more heavily represented below costa and in part of the range of grandillosa, tend to be cells M1 and M2; under surface with forewings smaller and darker than examples from other heavily and evenly suffused with dark brown countries. scales, and with secondaries more lightly The male genitalia of this species are suffused with dark brown; discal spots and similar to those of the preceding species in broad, brownish black terminal band present the nature of the long process of the sacculus; on all wings. this process, in the present taxon, has two LENGTH OF FOREWING: 21 to 27 mm.; widely separated areas of spining, a condition holotype, 27 mm. that is diagnostic. Additional points of rec- FEMALE: Unknown. ognition are to be found in the broader MALE GENITALIA: Uncus elongate, sides anellus, and in the much longer base of the concave, apex curving ventrally and termina- spine in the vesica. ting in single spine; gnathos well sclerotized, The female genitalia of grandillosa can be narrowing ventrally and with large, termin- separated from those of the preceding species ally rounded median projection; costa of by the swollen pleural areas laterad of the valve extending short distance beyond val- sterigma. This taxon also has both a smaller vula; sacculus elongate, with process slightly lamella postvaginalis and signum than are to shorter than costa, not swollen apically, and be found in bipennaria, as well as a less with apical region bearing two groups of strongly crenulate intersegmental area be- spines, one at about three-fourths of length of tween A7 and A8. process, the other terminal; length of sacculus The genitalia of the lectotype do not agree and process varying from 2.1 to 2.3 mm.; exactly with those of the other specimens anellus large, length about one and one-half placed under this name. The genitalia of the times greater than width, with slight basal type lack the swollen pleural areas laterad of constriction, posterolateral areas with longi- the sterigma and appear to have a smaller tudinal ridges, with posterior margin rounded lamella postvaginalis and larger lateral areas and having median incision; spine of vesica of the sterigma. Among the dissections of sinuous or with apical portion curved, bearing seven Argentinian females (two from Tucu- small teeth along median concave surface, man), none was seen that exactly matched and with very short recurved base; length of Todd's dissection. Possibly two species are spine varying from 0.8 to 1.2 mm., of re- represented, but, if so, it is not obvious from a curved base 0.1 to 0.2 mm.; vesica with elon- study of the moths themselves. As far as we gate, lightly sclerotized patch extending know now, only one species of Glena is to be parallel with spine on both sides of base of found in northwestern Argentina. spine, bearing numerous short, deciduous Glena sucula, new species spines; dorsal surface of vesica with large, subtriangular, coarsely denticulate area, and Figure 5 ventral surface with finely denticulate area. This species resembles dentata but can be Ventral surface of As without row of bristles. distinguished from it by the lack of both the FEMALE GENITALIA: Unknown. tibial hair pencil and the row of bristles on TYPES: Holotype, male, Paso del Quirdiu, the under side of the third abdominal seg- Cent[ral] Cord[illera], Colombia, elevation ment of the male. The male genitalia also 3500 meters (Fassl); genitalia mounted on have good diagnostic characters. This taxon slide Geometridae No. 5806. Paratypes, all occurs in Colombia. from Colombia: Manizales (A. M. Patino), MALE: Similar to that of dentata, but dif- one male; San Antonio, [Valle del Cauca], fering mainly as follows: hind tibia without elevation 5800 feet, November, 1907 (M. G. hair pencil; upper surface of wings with Palmer), one male, and elevation 2000 variable amount of dark brown and brownish meters, February, 1909 (Fassl), one male; 124 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 Canon del Tolima, [Tolima], elevation 2700 LENGTH OF FOREWING: 21 to 22 mm.; meters, December, 1909 (A. H. Fassl), one holotype, 22 mm. male; Pacho, [Cundinamarca] (Chapman), FEMALE: Unknown. one male; no further data, one male. MALE GENITALIA: Similar to those of The holotype is in the collection of the sucula, but differing mainly as follows: uncus British Museum (Natural History); para- slightly longer; valvula with more heavily types are in the collections of that institution sclerotized and irregularly shaped area longi- and of the United States National Museum. tudinally; sacculus with process shorter, DISTRIBUTION: Colombia (the departments apically swollen and bearing either one group of Valle del Cauca, Tolima, and Cundina- of spines ventrally and medially, or two nar- marca). This is apparently a species that is to rowly separated groups of spines; length of be found in the mountains, as the elevations sacculus and process, 2.0 to 2.1 mm.; anellus range from 5800 to about 11,500 feet. large, slightly longer than wide, posterolateral The adults have been captured in Novem- areas rugose, posterior margin rounded and ber, December, and February. with wide median incision; spine of vesica REMARKS: A total of seven specimens (all with broad curve, bearing small teeth in males) and seven genitalic dissections have apical region, and with short recurved base; been studied. This species is one of the largest length of spine, 1.2 mm., of recurved base 0.1 in the genus, and it has one of the darkest to 0.2 mm. Ventral surface of A3 without row under surfaces of the wings of any taxon. In of bristles. addition it is one of the few species that does FEMALE GENITALIA: Unknown. not have a hair pencil on the hind tibia of the TYPES: Holotype, male, Incachaca, Cocha- male. bamba, Bolivia (J. Steinbach); genitalia This and the following species are unique mounted on slide F.H.R. No. 13546. Para- in having the very short recurved base of the types: Same data as holotype, one male; spine of the vesica. The present taxon can be Tinguri, Carabaya, [Puno], Peru, one male. recognized by the fact that the longer and All three specimens are in the collection of non-capitate process of the sacculus bears its the United States National Museum. spines in two groups, and by the more slender DISTRIBUTION: Bolivia (the department of anellus. The two groups of spines on the Cochabamba) and southern Peru (the de- process of the sacculus are very similar to partment of Puno). those in granillosa; these two species can be REMARKS: A total of three specimens (all separated by the nature of the spine of the males) and two genitalic dissections have vesica, and by the absence of the row of been studied. This species is closely related to bristles from the ventral surface of A3 in the sucuka but can be recognized by the paler present species. under surface of the hind wings, by the dis- tribution, and by the genitalia. Glena vesana, new species The male genitalia of vesana are quite simi- lar to those of the preceding taxon, and can Figure 6 be recognized by the shorter capitate process This species closely resembles sucula; the of the sacculus, and by the broader anellus. males of both species lack both the tibial hair pencil and the row of bristles on the under Glena bisulca, new species surface of the third abdominal segment. The male genitalia have good differences by which Figure 7 to separate the two taxa. The present species This large-sized species is very similar to occurs in Peru and Bolivia. dentata; both taxa occur in Ecuador and MALE: Similar to that of sucula, but differ- Colombia. The genitalia should be used for ing mainly as follows: under surface of wings identification. with less dark brown scaling, particularly on MALE: Similar to that of dentata, differing hind wings; hind wings with discal dot mainly as follows: upper surface of wings tending to be smaller, and with terminal band variably overlain with dark scales, tending to obsolescent. be paler in color than that of dentata; macula- 1967 RINDGE: GLENA 125 tion tending to be slightly more heavily repre- (Natural History) and the last two are in the sented; under surface of forewings tending to United States National Museum. have more sinuate inner border of terminal DISTRIBUTION: Ecuador (the province of band, represented by narrow, pale, more or Napo-Pastaza) and Colombia (the depart- less indistinct line; hind wings with incom- ments of Caldas and Cundinamarca). The plete terminal band and less dark scaling species apparently flies at higher altitudes medially; hind tibia with hair pencil. than does dentata, as the present taxon is LENGTH OF FOREWING: 22 (holotype) to known to occur from 4500 to 8000 feet. 26 mm. The adults have been taken in April, FEMALE: Unknown. September, and November. MALE GENITALIA: Similar to those of REMARKS: A total of five specimens (all grandillosa, differing mainly as follows: uncus males) and five genitalic dissections have with apical area more laterally compressed been studied. This species is similar to dentata and curved ventrally; valves with apex of in its distribution and variation. Both species costa only slightly swollen; process of sac- apparently tend to be slightly smaller and to culus elongate but not reaching edge of valve; have less strongly defined maculation in length of sacculus and process varying from Ecuador than in Colombia, factors that may 2.5 to 2.7 mm.; process without basal swelling prove to be of subspecific value when sufficient but with slight enlargement at about two- material comes to hand. thirds of length covered with elongate spines This species, like grandillosa, has two on inner side, and with apex bearing spines; patches of spines on the process of the sac- anellus with length equal to width, with culus. In the latter species these groups of moderate anterior constriction, posterior spines are separated by a distance of at least margin concave and in form of two cuplike 0.5 mm.; in the present species this is re- ovals, posterolateral angles with weak, poste- duced to about 0.2 mm. There are also differ- riorly extending ridges; spine of vesica with ences between the two taxa in the shape of elongate base, and with terminal portion the anellus and in both the spine and orna- curved dorsally and bearing minute teeth; mentation of the vesica. length of spine 1.2 to 1.3 mm., of sharply re- curved base 0.7 to 0.9 mm.; vesica with Glena juga, new species weakly sclerotized area posteriad of base of Figures 8, 33 spine bearing numerous short, deciduous The males usually have the under surface spines, and with dorsal surface having large, of the forewings dark and that of the hind sclerotized, dentate area; ventral surface wings white and immaculate, whereas the fe- with weakly sclerotized area, its surface males have a more or less complete terminal covered with extremely fine denticulation. band on the secondaries. The genitalia should Ventral surface of A3 with median row of be used for identification. This species occurs bristles. from Colombia south to Bolivia and Para- FEMALE GENITALIA: Unknown. guay. TYPES: Holotype, male, Hacienda La MALE: Similar to that of bipennaria, differ- Mascota, Rio Topo, [Napo-Pastaza], Ecua- ing mainly as follows: smaller in size; macula- dor, elevation 4500 feet, April 13, 1931 (W. J. tion less strongly indicated on upper surface; Coxey); genitalia mounted on slide F.H.R. under surface with forewings heavily suffused No. 13446. Paratypes, all from Colombia: with dark brownish gray scales, and with San Cajetano, elevation 8000 feet, Septem- broad, dark terminal band; under surface of ber, 1902, one male; Manizales, [Caldas] (A. hind wings varying from immaculate white to M. Patino), one male; Vergara, [Cundina- having partial representation of terminal marca], October 15 (Apollinaire), one male; band anteriorly; hind tibia with hair pencil. Rio Vitaco, elevation 2000 meters, Novem- LENGTH OF FOREWING: 18 to 27 mm.; ber, 1908 (Fassl), one male. The holotype is holotype, 19 mm. in the collection of the American Museum of FEMALE: Similar to male but tending to Natural History; the first two paratypes are have upper surface of wings more heavily ir- in the collection of the British Museum rorate with dark scales and to have less 126 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 clearly defined maculation, and to have more rior margin, rounded posteriorly, outer mar- strongly represented terminal band below, gin with numerous pointed projections, and with more dark scaling on hind wings. with flat median area recessed and bearing LENGTH OF FOREWING: 20 to 25 mm.; several small spinose processes. Posterior por- allotype, 22 mm. tion of ventral surface of A7 not modified. MALE GENITALIA: Similar to those of TYPES: Holotype, male, Carl[os] Pfannl, grandillosa, differing mainly as follows: Paraguay, July 6, 1951 (Schade); genitalia valves with apex of costa only slightly mounted on slide F.H.R. No. 13480. Allo- swollen; process of sacculus elongate but not type, female, Coosnipata, Paucartambo, reaching edge of valve; length of sacculus and Cuzco, Peru, November 29, 1951 (F. L. process varying from 2.1 to 2.4 mm.; process Woytkowski); genitalia mounted on slide with terminal one-third swollen and with F.H.R. No. 13377. Paratypes: Paraguay: numerous heavy spines; anellus without con- Carlos Pfannl, August 6, 1951 (Schade), three striction, increasing in width posteriorly, males; Paso Yobay, September, 1951 lateral margins and posterolateral areas with (Schade), two males. Bolivia: Rio Songo, several small ridges, and with posterior elevation 750 meters (Fassl), one female. margin convex and having median indenta- Peru: Coosnipata, Paucartambo, Cuzco, No- tion; spine of vesica relatively short and with vember 16, 19, 29, 1951 (F. L. Woytkowski), very long recurved base, terminal portion of three females; Oconeque, Carabaya, [Puno], spine somewhat variable in curvature, some elevation 6000-7000 feet, July, November, specimens with single curve, others with weak 1904, February, 1905 (G. Ockenden), five double curve; length of spine varying from males and one female; "Ob. Madre de Dios, 0.9 to 1.2 mm., of sharply recurved base, 0.5 Sud Peru," elevation 500-1200 meters (Fassl), to 0.8 mm.; vesica with weakly sclerotized one female; Limbani, Carabaya, [Puno], area posteriad of base of spine bearing numer- elevation 9500 feet, dry season, May, 1904 ous short, deciduous spines, and with dorsal (G. Ockenden), one male. Ecuador: Environs surface having large, sclerotized, dentate of Loja, [Loja], 1888-1893, 24 males, one fe- area; ventral surface with weakly sclerotized male; Loja, [Loja], one male. Colombia: Vil- area, its surface with extremely fine denticu- lavicencio, [Meta] (Apollinaire), one female. lation. Ventral surface of A3 with median row The holotype and allotype are in the collec- of bristles. tion of the American Museum of Natural FEMALE GENITALIA: Sterigma with large, History; paratypes are in the collections of elongate, smoothly sclerotized lamella post- that institution, of the British Museum (Nat- vaginalis, with width slightly more than one- ural History), and of the United States Na- half of length, narrowing posteriorly, with tional Museum. posterior end angled dorsally; lateral areas DISTRIBUTION: Paraguay, Bolivia, Peru of sterigma large, heavily sclerotized, with (the departments of Cuzco and Puno), Ecua- pleural area produced as strongly raised dor (the province of Loja), and Colombia (the ridge; intersegmental area between A7 and department of Meta). Apparently this spe- A8 sclerotized laterally, and weakly recessed cies occurs to the east of the Andes Moun- and convoluted ventrally; ductus bursae tains at elevations of from about 2500 to 9000 about one-fifth of length of lamella post- feet. vaginalis; ductus seminalis arising from large The adults have been captured in Feb- sac laterally or ventrolaterally on left side of ruary, May, July, August, September, and corpus bursae, originating between 0.9 and November. 1.0 mm. anteriad of ductus bursae, with REMARKS: A total of 46 specimens (37 tubular portion coming off medially near males and nine females) and 33 genitalic corpus bursae; latter elongate, weakly swol- dissections (26 males and seven females) have len anteriorly, and with posterior one-third to been studied. Examples from Paraguay tend two-fifths weakly sclerotized and striate, to be slightly smaller and paler in coloration surface covered with very many small swell- than specimens from farther north. Males ings or denticulations; signum situated from Paraguay have an immaculate white ventrally, with flat or weakly concave ante- under surface to the hind wings, whereas 1967 RINDGE: GLENA 127 Ecuadorian specimens have a partial terminal spines; length of sacculus and process vary- band on those wings. ing from 2.9 to 3.1 mm.; anellus about one This species has the apex of the process of and one-half times longer than broad, with the sacculus swollen and spinose, something wide anterior end, sharply constricted pos- like that of bipennaria. The present species teriad thereof, then broadly swollen into two can be separated from the latter by the large, flat lobes with minutely denticulate broader anellus and by the much longer base surface, each lobe rounded posteriorly, and of the spine in the vesica. with large median indentation; spine of vesica The female genitalia of juga can be recog- sinuously curved, terminal portion with nized by the very long, tapering lamella small teeth, and with elongate, sharply re- postvaginalis with the modified posterior end, curved base; length of spine varying from 1.4 and by the prominent ridge on each side of to 1.5 mm., of recurved base from 0.7 to 0.8 the heavily sclerotized lateral areas of the mm.; vesica with weakly sclerotized area sterigma. posteriad of base of spine bearing numerous short, deciduous spines, and with more Glena megale, new species heavily sclerotized area extending to apex of Figure 34 base of spine, latter area tending to have cross This is a large species with elongate fore- striations at right angle to spine; dorsal sur- wings and with the upper surface of the wings face of vesica with large, sclerotized, dentate more or less heavily dusted with dark scales. area, with teeth becoming larger anteriad of The genitalia should be used for identifica- apex of spine; ventral surface with weakly tion. This species occurs from Venezuela to sclerotized area, its surface with extremely Peru. fine denticulation. Ventral surface of A3 with MALE: Similar to that of effusa, differing median row of bristles. mainly as follows: front tending to have FEMALE GENITALIA: Sterigma with large, wider white band across bottom; upper sur- very weakly sclerotized, oval or elliptical face of wings more heavily suffused with pale lamella postvaginalis, becoming enlarged brownish gray scales, and with maculation and more heavily sclerotized posteriorly, and tending to be more strongly represented; with large, sclerotized lateral areas, their ir- under surface with more prominent white regularly shaped anteromedian margins ex- apical spot on forewings, with terminal band tending up to anterolateral portions of lamella tending to fade out before reaching anal angle postvaginalis; intersegmental membrane be- of hind wings, and with discal dots tending tween A7 and A8 recessed and weakly con- to be more prominent; hind tibia with hair voluted ventrally; ductus bursae very short, pencil. scarcely differentiated from lamella post- LENGTH OF FOREWING: 22 to 24 mm.; holo- vaginalis; ductus seminalis arising from large type, 22 mm. sac on dorsal or dorsolateral surface, origi- FEMALE: Similar to male, but with upper nating about 2.0 mm. anteriad of ductus surface of wings heavily and evenly suffused bursae, with tubular portion coming off pos- with grayish brown scaling, and with macu- terolaterally; corpus bursae elongate, weakly lation more weakly defined; under surface swollen anteriorly, and with posterior two- more heavily suffused with brown scales. fifths sclerotized, surface covered with LENGTH OF FOREWING: 24 mm. (allotype). very small swellings or denticulations, and MALE GENITALIA: Similar to those of juga, with median area of corpus bursae having differing mainly as follows: uncus more numerous striations; signum on left side of strongly developed, ventral surface tending corpus bursae, anterior margin flat, rounded to be concave, with ventrolateral margins posteriorly, outer margin with numerous slightly extended; gnathos very attenuate, pointed projections, and with flat median curving posteriorly, with long and slender area recessed and bearing several small spi- median projection; process of sacculus ex- nose processes. Posterior portion of ventral tending beyond edge of valve, very long and surface of A7 not modified. slender, scarcely swollen apically, with ter- TYPES: Holotype, male, Pedregosa, Merida, minal two-fifths having numerous heavy Venezuela, elevation 3000 meters, October, 128 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 1897 (Briceno). Allotype, female, same data, occurs in Panama and in much of South June 8, 1898; genitalia mounted on slide America. The genitalia should be used for Geometridae No. 4801. Paratypes: Venezuela: determination. Merida, November, 1898 (Briceno), one This species is divided into two popula- male. Colombia: "Nouv. Grenada," Honda, tions. In one, the under side of the hind wings 1899 (M. de Mathan), one male. Ecuador: of the male is usually without maculation; it Environs of Loja, [Loja], 1887, one male. occurs in Brazil and Paraguay. The second Peru: Santo Domingo, [Carabaya, Puno], population occurs from Panama to Bolivia, elevation 6000 feet (G. Ockenden), one male; and the under side of the hind wings of the Chanchamayo, [Pasco], 1898 (O. Schuncke), male has a dark terminal band. one male; no further data, one male. The holotype and allotype are in the col- Glena unipennaria unipennaria (Guenee) lection of the British Museum (Natural His- tory); paratypes are in the collections of that Figures 10, 35 institution and of the United States National Boarmia unipennaria GUENEE, 1857, p. 257. Museum. WALKER, 1860, p. 356. OBERTHUR, 1913, p. 278 DISTRIBUTION: Venezuela (the state of (partim), pl. CLXIX, fig. 1655 (lectotype). Catoria sp. (? Boarmia bipennaria): COSTA Merida), Colombia, Ecuador (the province of LIMA, "1949" [1950], p. 129, figs. 97 (adult), 98 Loja), and Peru (the departments of Puno (male genitalia). and Pasco). This species may be found at higher elevations; the two specimens with This is the paler population, in which the altitude data indicate that they were taken under side of the secondaries of the male is at 6000 and 9840 feet. usually immaculate. The adults were captured in June, October, MALE: Similar to that of bipennaria, dif- and November. fering mainly as follows: smaller; upper sur- REMARKS: Eight specimens (seven males face of wings paler and with weaker macula- and one female) and seven genitalic dissec- tion; under surface of wings considerably tions (six males and one female) have been paler and with terminal band tending to be studied. The genitalia of the holotype protrude absent, particularly from hind wings; hind from the end of the abdomen, with valves tibia with hair pencil. spread, and they have been studied without LENGTH OF FOREWING: 18 to 22 mm. dissection. FEMALE: Similar to male, but with heavier The adults are quite uniform in size, mac- brown and brownish gray scaling above and ulation, and color. The relatively dark color with more weakly defined pattern; under of the upper surface of the wings might be surface more heavily suffused with dark scales due to the age of the specimens, as all the and with terminal band more strongly rep- material was apparently taken between 1887 resented. and 1899. LENGTH OF FOREWING: 16 to 24 mm. The male genitalia of this species are easily MALE GENITALIA: Similar to those of juga, recognized by their large size, by the very differing mainly as follows: gnathos tending elongate gnathos, by the very long process of to be less attenuate medially; process of sac- the sacculus, and by the unique shape of the culus shorter, with terminal spinose swelling; anellus. length of sacculus and process varying from The female genitalia of megale can be iden- 1.8 to 2.3 mm.; anellus elongate and slender, tified by the very weakly sclerotized lamella its length approximately 2.5 times as great as postvaginalis and its more heavily sclerotized width, anterior portion broad, with prominent posterior portion, by the almost dorsal origin median constriction, lateral margins thick- of the ductus seminalis from the large sac, ened, expanding in width posteriorly but and by the fact that the posterior part of the with posterior one-fourth narrowed, with- corpus bursae is sclerotized but not striated. out thickened margins, posteriorly rounded Glena unipennaria (Guenee), new combination and with small median indentation, latter apparently continued anteriorly to near mid- Boarmia unipennaria GUENEE, 1857, p. 257. dle of anellus; spine of vesica more or less This rather pale species of moderate size S-shaped, with elongate, sharply recurved 1967 RINDGE: GLENA 129 base, and with small teeth on anterior surface were caught in every month except Febru- of spine near apex; length of spine varying ary. from 1.2 to 1.5 mm., of recurved base from REMARKS: A total of 65 specimens (51 0.5 to 0.8 mm.; vesica with weakly sclerotized males and 14 females) and 38 genitalic dis- area posteriad and away from base of spine sections (28 males and 10 females) have been bearing numerous short, deciduous spines, studied. This population can usually be dis- and with dorsal surface having large, sub- tinguished, at least in the males, by the al- triangular, dentate area; ventral surface with most immaculate under surface of the hind weakly sclerotized area, its surface with ex- wings. tremely fine denticulation. Ventral surface This species can be distinguished from of A8 with median row of bristles. juga in the male genitalia by the shorter FEMALE GENITALIA: Sterigma with large, process of the sacculus and by the much nar- sclerotized, oval lamella postvaginalis, and rower anellus. with large, sclerotized lateral areas, their The female genitalia of unipennaria can anteromedian margins extending to antero- be recognized by the very large ventral sac lateral portions of lamella postvaginalis; in- on the corpus bursae, with the ductus semi- tersegmental area between A7 and A8 recessed nalis arising posteroventrally therefrom, and and convoluted ventrally; ductus bursae very by the fact that the sclerotized area of the short; ductus seminalis arising from very corpus bursae is primarily on the dorsal sur- large, rounded sac ventrally on corpus bursae, face. originating between 0.8 and 1.0 mm. anteriad of ductus bursae, with tubular portion com- Glena unipennaria cosmeta, new subspecies ing off sac posteroventrally; corpus bursae Boarmia demissaria auct.: DRUCE, 1892 (1891- elongate, scarcely swollen anteriorly, and 1900), p. 75 (partim). with posterior one-third to two-fifths of dorsal This population differs from the nominate surface weakly sclerotized and striate, sur- one by the presence of a dark terminal band face covered with very small swellings or on the under side of the hind wings in the denticulations; signum on ventral surface of males. It occurs from Panama and the corpus bursae, with flat or weakly concave Guianas to Bolivia. The genitalia should be anterior margin, rounded posteriorly, outer used for determination. margin with numerous pointed projections, MALE: Similar to that of nominate uni- and with flat median area recessed and bear- pennaria, but differing mainly as follows: ing several small spinose processes. Posterior upper surface of wings tending to have slightly portion of ventral surface of A7 not modified. less dark scaling, and with maculation slightly TYPES: Guenee described unipennaria from less strongly indicated; under surface with a series of one male and three female speci- dark, wide terminal band on all wings. mens. Oberthiir (1913) selected one of the LENGTH OF FOREWING: 18 to 23 mm.; females as the "specimina typica," because holotype, 19 mm. what was presumably the original male was FEMALE: Similar to male but with more incomplete and in a very poor state of pres- obscure maculation and with upper surface ervation. This female becomes the lectotype; of wings more heavily suffused with dark it was figured by Oberthur, and it is in the scales; under surface with heavier terminal collection of the British Museum (Natural band and more suffused with dark scales. History). LENGTH OF FOREWING: 21 to 25 mm. TYPE LOCALITY: Brazil. MALE GENITALIA: Similar to those of nom- DISTRIBUTION: Southern Brazil (the states inate subspecies. of Santa Catarina, Parana, Sao Paulo, Gua- FEMALE GENITALIA: Similar to those of nabara, Rio de Janeiro, Minas Gerais, and nominate subspecies. Mato Grosso) and Paraguay. (See Appendix TYPES: Holotype, male, Fort Kobbe, Canal for locality data of specimens examined.) Zone, Panama, September 30, 1956 (W. E. This subspecies occurs from about sea level Lundy); genitalia mounted on slide F.H.R. up to about 3100 feet. No. 13492. Paratypes: Panama: Same data The adults apparently fly throughout the as holotype, one male; Tole, [Chiriqui] year, as specimens have been examined that (Champion), one male, identified as "Boarmia 130 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 demissaria Walk., B. C. A. Lep. Het., God- Glena uncata, new species man-Salvin Coll." French Guiana: Cayenne, Figures 11, 36 one male. Guyana (former British Guiana): Boarmia demissaria auct.: DRUCE, 1892 (1891- Christianburg, one male. Venezuela: Aroa, 75 [Yaracuy], two males. Colombia: "Ob. Rio 1900), p. (partim). Negro," [Norte de Santander], elevation 800 This rather pale species of moderate size meters (Fassl), two males; Minca, [Magda- occurs from Mexico to Costa Rica. The gen- lena], elevation 2000 feet (H. H. Smith), one italia should be used for determination. male; Florida, R[io] Putamayo, [Amazonas], MALE: Similar to that of unipennaria March, 1932 (G. Klug), one male; no further cosmeta, differing mainly as follows: upper data, one male. Ecuador: Zamora, Santiago- surface of wings with only sparse dusting of Zamora, March 27-30, 1965 (L. E. Pefia), one brownish gray scales; t. p. and extradiscal male; Sarayacu, [Napo-Pastaza] (C. Buckley), lines outwardly dentate on veins; under sur- two males; environs of Loja, [Loja], 1889, one face pale, with brownish black terminal band female. Bolivia: Chapare, [Beni], elevation on all wings, decreasing in width and inten- 400 meters, November 2, 1948, May, 1951, sity posteriorly on hind wings; hind tibia with two males; R[io] Songo to R[io] Suapi, [La hair pencil. Paz], elevation 1100 meters, March-June, LENGTH OF FOREWING: 18 to 20 mm.; holo- 1896 (Garlepp), one male; Prov[ince] del type, 20 mm. Sara, Santa Cruz, elevation 450 meters, FEMALE: Similar to male, but tending to be January (J. Steinbach), one female. more heavily dusted with dark scales on both The holotype is in the collection of the upper and under surfaces of wings; terminal American Museum of Natural History; para- band on hind wings varying from being like types are in the collections of that institution, that of male (allotype) to being broad and of the British Museum (Natural History), complete. and of the United States National Museum. LENGTH OF FOREWING: 20 to 23 mm.; allo- DISTRIBUTION: Panama (the province of type, 20 mm. Chiriqui; the Canal Zone), Guyana, French MALE GENITALIA: Similar to those of uni- Guiana, Venezuela (the state of Yaracuy), pennaria, differing mainly as follows: uncus Colombia (the departments of Norte de smaller, with flatter sides; process of sacculus Santander, Magdalena, and Amazonas), Ec- extending almost to edge of valve, very long uador (the provinces of Santiago-Zamora, and slender, scarcely swollen apically, with Napo-Pastaza, and Loja), and Bolivia (the terminal one-fourth having numerous short departments of Beni, La Paz, and Santa spines; length of sacculus and process vary- Cruz). The moths have been taken from ing from 1.7 to 2.0 mm.; anellus long and near sea level up to about 3600 feet. slender, broadly concave medially, swollen The adults have been captured in January, distally, with posterior end rounded and with March, May, June, September, and Novem- median indentation; spine of vesica curved, ber. terminal portion with a few small teeth, and REMARKS: A total of 20 specimens (18 with elongate, sharply recurved base, termi- males and two females) and 16 genitalic dis- nating in anteriorly directed point; length of sections (14 males and two females) have spine 1.0 to 1.1 mm., of recurved base, 0.45 been studied. Specimens of this population mm.; vesica with weakly sclerotized area are easily separated from examples of the posteriad of base of spine bearing numerous nominate taxon by the more heavily marked short, deciduous spines, and with dorsolateral under surface of the wings. Some individuals surface having large, weakly sclerotized, den- from Bolivia tend to be more lightly marked tate area; ventral surface membranous. Ven- than more northern examples. Greater diffi- tral surface of A3 with median row of bristles. culty will be found in separating members of FEMALE GENITALIA: Sterigma with rela- this population from specimens of demis- tively short, posteriorly rounded, smoothly saria; the genitalia should be studied. sclerotized lamella postvaginalis, with width 1967 RINDGE: GLENA 131 greater than length; lateral areas of sterigma be separated from unipennaria cosmeta by the weakly sclerotized, narrow, not attaining fact that the terminal band on the under side sterigma ventrally; intersegmental area be- of the wings is less heavily represented, by its tween A7 and A8 recessed and convoluted lat- more northern distribution, and by its gen- erally; ductus bursae less than one-half of italia. length of lamella postvaginalis; ductus semi- The male genitalia are very much like those nalis arising as large tube ventrolaterally of unipennaria, but can be distinguished by from swollen area on corpus bursae, originat- the different shape of the anellus, by the ing between 0.4 and 0.6 mm. anteriad of cor- different ornamentation of the vesica, and by pus bursae; corpus bursae elongate, weakly the shorter spine in the aedeagus. swollen anteriorly, and with posterior one- The female genitalia of uncata can be sep- fourth weakly sclerotized and striate, surface arated from those of any of the preceding covered with very small swellings or denticu- species by the fact that the ductus seminalis lations; signum situated laterally or dorsally arises as a thick tube, rather than as a large on corpus bursae, elliptical, outer margin with sac. numerous pointed projections of approxi- mately equal length, and with flat median Glena demissaria (Walker), new combination area recessed and bearing small spinose pro- Figures 12, 37 cesses. Posterior portion of ventral surface of Boarmia demissaria WALKER, 1860, p. 357. A7 not modified. Catoria unipennaria auct.: OBERTHUR, 1913, p. TYPES: Holotype, male, Juan Vinas, [Car- 278 (partim), pi. CLXIX, fig. 1657. DYAR, 1914, tago], Costa Rica, January (Schaus and p. 243. Barnes); genitalia mounted on slide F.H.R. This species is similar to unipennaria cos- No. 13599. Allotype, same data but collected meta, but the cross lines are less dentate, and in June, and with genitalia mounted on slide there is usually more dark scaling on the under F.H.R. No. 13610. Paratypes: Costa Rica: surface of the wings. The genitalia should be Same data as types, February and June, used for identification. This species occurs two females; Guapiles, [Limon], November in much of South America. (Schaus and Barnes), one female; no further MALE: Similar to that of unipennaria cos- data (P. Biolley), one male. Guatemala: Ca- meta, differing mainly as follows: upper sur- yuga, May (Schaus and Barnes), one male. face of wings with cross lines tending to be Mexico: Cordoba, [Veracruz], April 28, 1908, represented by spots on veins; under surface May 16, 1908 (F. Knab), two females; Ja- with broad terminal band on all wings, and lapa, [Veracruz] (Hoege), one female, identi- with wings more or less heavily suffused with fied as "Boarmia demissaria Walk., B. C. A. dark brownish gray scales; hind tibia with Lep. Het., Godman-Salvin Coll." hair pencil. The holotype and allotype are in the col- LENGTH OF FOREWING: 17 to 22 mm. lection of the United States National Mu- FEMALE: Similar to male, but both surfaces seum; paratypes are in the collections of that of wings more heavily suffused with dark institution and of the British Museum (Nat- scales, and with cross lines on upper surface ural History). more weakly represented. DISTRIBUTION: Costa Rica (the provinces LENGTH OF FOREWING: 18 to 22 mm. of Cartago and Limon), Guatemala, and MALE GENITALIA: Similar to those of uni- Mexico (the state of Veracruz). pennaria, differing mainly as follows: uncus The moths have been collected in Janu- with terminal area not laterally compressed; ary, February, April, May, June, and No- gnathos with median process tending to be vember. slightly shorter and less attenuate; process REMARKS: A total of 10 specimens (three of sacculus shorter and with terminal spinose males and seven females) and nine genitalic swelling; length of sacculus and process vary- dissections (three males and six females) ing from 1.4 to 1.9 mm.; anellus elongate and have been studied. This species can usually slender, its length approximately three times 132 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 as great as width, sides varying from being Bolivia, eastern Peru (the departments of subparallel anteriorly to having constriction Junin, Madre de Dios, and Puno), Colombia about one-third of distance from base, and (the department of Santander), Venezuela with prominent, rounded, rugose lobe on each (the states of Bolivar and Yaracuy), Guyana, margin at two-thirds of distance frombase; French Guiana, and Trinidad. This widely posterior margin of anellus bilobed, with ranging species is usually found at low ele- prominent median indentation extending vations, although one locality in Peru is given anteriorly as membranous strip to basal con- as 6000 feet. striction; spine of vesica very long, relatively The adults have been captured in every straight, terminal portion with weak bend month of the year. only and apex slightly hooked, and with REMARKS: A total of 94 specimens (76 area of slender teeth near apex; base of spine males and 18 females) and 55 genitalic dis- sharply recurved, its end pointed; length of sections (39 males and 16 females) have been spine varying from 1.4 to 2.0 mm., of recurved studied. Glena demissaria is the most com- base from 0.6 to 1.0 mm.; vesica with weakly mon and most widespread of the smaller sclerotized area posteriad of base of spine taxa in South America. It is a rather nonde- bearing numerous short, deciduous spines, script species that is difficult to characterize, and with dorsal, posterior and ventral sur- although most specimens have the wide ter- faces having extensive, sclerotized, dentate minal band on the under side of the wings, area, the teeth thereof becoming larger along and the same surface is usually suffused with margin of vesica. Ventral surface of A8 with dark scaling, particularly on the forewings. median row of bristles. The genitalia should be used for identifica- FEMALE GENITALIA: Sterigma with large, tion. sclerotized, rounded lamella postvaginalis, The male genitalia of this species are with width slightly greater than length, and similar to those of unipennaria but can be with large lateral areas partially sclerotized, recognized the their irregularly shaped median margins ex- by shorter process of the sac- tending to lateral margins of lamella post- culus, by the different configuration of the vaginalis; intersegmental area between A7 anellus, and by the longer and straighter and A8 weakly recessed and convoluted ven- spine in the vesica. trally; ductus bursae very short; ductus semi- There is some variation in the size of the nalis arising in form of large tube dorsally or parts of the male genitalia of demissaria. dorsolaterally on corpus bursae, originating Specimens from the southern Andes Moun- between 1.1 and 1.5 mm. anteriad of ductus tains tend to have both a longer sacculus and bursae; corpus bursae elongate, weakly process in the valve, and a longer spine in the swollen anteriorly, and with posterior one- aedeagus, than do examples from northern third to two-fifths weakly sclerotized and or eastern South America. striate, surface covered with very small swell- The female genitalia of this species can be ings or denticulations; signum on ventral sur- separated from those of all the preceding face of corpus bursae, elliptical, outer margin taxa, except uncata, by the entirely tubular with numerous pointed projections, those ductus seminalis. Additional diagnostic fea- along anterior margin tending to be longer tures include the short and broad lamella than posterior ones, and with flat median area postvaginalis and the spinose margins of the recessed and bearing many small spinose proc- signum; the latter will separate the genitalia esses. Posterior portion of ventral surface of of this species from those that follow. From A7 not modified. those of uncata, the genitalia of demissaria TYPE: Walker described demissaria from a can be recognized by the dorsal origin of the single male specimen; it is in the collection ductus seminalis. of the British Museum (Natural History). Several years after the appearance of the TYPE LOCALITY: Para, Brazil. description of demissaria, Walker (1868, pp. DISTRIBUTION: Brazil (the states of Para, 184, 197) redescribed this species. The second Amazonas, Mato Grosso, Minas Gerais, and description was based on a male from Bogota, Santa Catarina), Uruguay, Paraguay, eastern Colombia; this specimen has not been ex- 1967 RINDGE: GLENA 133 amined, so it is not known whether or not it is Costa Rica, elevation 5800 feet, August 31, conspecific with true demissaria. 1908; genitalia mounted on slide F.H.R. No. 13614. This specimen is in the collection of the Glena basalis, new species United States National Museum. Figure 38 DISTRIBUTION: This species is known only This species is similar to demissaria, but from the province of Cartago in central both surfaces of the wings are darker, and the Costa Rica. t. p. and extradiscal lines are situated nearer REMARKS: This taxon is represented by the the base of the wings. The genitalia should unique type only. The dark coloration of both be used for identification. This species is surfaces of the wings, the relatively basal known only from Costa Rica. position of the cross lines, and the distinctive MALE: Unknown. female genitalia will serve to distinguish this FEMALE: Similar to that of demissaria but species. with more white scales on front, and with Glena hima, new species third segment of palpi white; upper surface of wings more heavily dusted with grayish Figures 13, 39 brown scales, and with maculation tending to Boarmia-?: DRUCE, 1892 (1891-1900), p. 74. be less well t. p. and extra- Boarmia demissaria auct.: DRUCE, 1892 (1891- slightly defined; 75 discal lines more dentate and situated more 1900), p. (partim). basally on the wings; discal spot of hind wings This species is similar to demissaria, but touching extradiscal line; under surface the under surface of the wings is paler, and brownish gray, with very wide but rather both the tibial hair pencil and the bristle poorly defined terminal band, with prominent comb on the third abdominal segment are white apical spot on each forewing, and with lacking. The genitalia of both sexes are dis- prominent discal dots on all wings. tinctive. The present taxon occurs in Costa LENGTH OF FOREWING: 19 mm. (holotype). Rica and Panama. MALE GENITALIA: Unknown. MALE: Similar to that of demissaria, but FEMALE GENITALIA: Sterigma with large, differing mainly as follows: hind tibia with- elongate lamella postvaginalis, smoothly out hair pencil; upper surface of wings tend- sclerotized except for rugulose posterior one- ing to have slightly more brownish scaling, third, and with large, sclerotized lateral areas, and to have more prominent brown band their median margins not extending over outside t. p. and extradiscal lines; under sur- lamella postvaginalis, and with posterior por- face much paler, forewings lightly suffused tion having vertical ridge just anteriad of with pale brownish gray scales, and with pale outwardly swollen posterior margin; inter- brown terminal band weakly represented on segmental area between A7 and A8 recessed forewings only. but not convoluted; ductus bursae very short; LENGTH OF FOREWING: 18 to 24 (holotype) ductus seminalis arising ventrally as large, mm. swollen tube, extending posteriorly as far as FEMALE: Similar to male, but with upper ductus bursae, sharply narrowed at posterior surface of wings more heavily suffused with margin and ductus continued anteriorly and brown scales, and with maculation less then laterally, originating about 0.75 mm. strongly represented; under surface of wings anteriad of ductus bursae; corpus bursae rel- varying from lightly suffused with brown atively short and broad, weakly swollen an- scales and with prominent terminal band on teriorly, and with posterior one-fourth weakly all wings (Panamanian specimens) to more sclerotized and striate, surface covered with heavily suffused with brown scales and with very small swellings or denticulations; sig- much less prominent terminal band (Costa num a transverse ellipse, anterior margin Rican specimens). enlarged and smoothly rounded, and with LENGTH OF FOREWING: 17 to 24 mm.; allo- flat median area recessed and projecting an- type, 17 mm. teriorly at angle into ventral wall of corpus MALE GENITALIA: Uncus triangular, apex bursae, surface minutely ridged. curving ventrally and terminating in two TYPE: Holotype, female, Tuis, [Cartago], weak points; gnathos with moderate median 134 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 enlargement, apically wedge-shaped; costa of demissaria Walk. B. C. A. Lep. Het. God- valves concave, and with outer one-half man-Salvin Coll." Paratypes: Panama: Chiri- covered with spines; sacculus elongate, con- qui, one female. Costa Rica: Orosi, [Cartago], tinued posteriorly by long process extending elevation 1200 meters (Fassl), one male; R[io] to just beyond edge of valvula, with process Sucio, [San Jos6] (H. Rogers), and identified gradually increasing in width distally and as "Boarmia sp. (?), No. 10, Druce, B. C. A. bearing numerous short spines along inner Lep. Het. Godman-Salvin Coll.," one female; margin of outer one-half and continuing San Jos6, [San Jose], November, 1906, one around apex of process; length of sacculus female. and process varying from 1.9 to 2.6 mm.; The holotype and allotype are in the col- anellus elongate, slightly increasing in width lection of the British Museum (Natural His- posteriorly, with raised lateral margins, and tory); paratypes are in the collections of that with posterior margin strongly bilobed; spine institution and of the United States National of vesica sinuate, bearing small teeth distally Museum. and with large, strongly recurved base; DISTRIBUTION: Panama (the province of length of spine varying from 1.5 to 1.8 mm., Chiriqui) and Costa Rica (the provinces of of recurved base from 0.5 to 0.6 mm.; vesica Cartago and San Jose). with elongate, weakly sclerotized area along The only known date of capture is in No- base of spine bearing numerous short, decid- vember. uous spines; vesica with two lateral, elongate, REMARKS: A total of six specimens (two longitudinal, denticulate strips on dorsal males and four females) and six genitalic dis- surface, and with undifferentiated ventral sections have been studied. More material is surface. Ventral surface of A3 without needed before questions pertaining to the median row of bristles. variability of this species can be answered. FEMALE GENITALIA: Sterigma with lamella There is considerable variation in size among postvaginalis long and narrow, its length the few specimens that are in the type series, three times greater than width, smoothly and there is a marked difference in macula- sclerotized, with ends rounded and with tion on the under surface of the wings be- large, subtriangular, smoothly sclerotized tween the Panamanian and Costa Rican fe- lateral areas, slightly shorter than lamella males. Unfortunately five of the specimens postvaginalis; intersegmental area between do not have their dates of capture, so it is A7 and A8 deeply recessed; ductus bursae not possible to tell whether or not seasonal short, tapering; ductus seminalis arising ven- variation is present in this species. trally as large, rounded sac 0.6 mm. anteriad The male genitalia of this species can be of ductus bursae, with posterior portion curv- recognized by the nature of the process of the ing to right and producing narrow tube; cor- sacculus, by the elongate recurved base of the pus bursae weakly swollen anteriorly, with spine of the vesica, and by the ornamentation small swelling on dorsal surface opposite or- of the vesica. igin of ductus seminalis, and with posterior There are certain differences between the portion weakly sclerotized and longitudinally two male dissections of this species. Part of striate, surface covered with very fine swell- these may be due to the way they are ings or denticulations; signum situated ven- mounted but cannot account for all the varia- trally, large, transverse, elliptical, with broad tion. The genitalia of the Panamanian speci- outer margin of more or less equal width, men are larger, and the extent of spining outer edge minutely serrate, and with median along the process of the sacculus is greater. section flat. The anellus of this specimen appears to be TYPES: Holotype, male, V[olcan] de Chiri- less rugose than that of the example from qui, [Chiriqui, Panama], below 4000 feet Costa Rica. (Champion); genitalia mounted on slide The female genitalia are very distinctive Geometridae No. 5812. Allotype, female, in having the very long and slender lamella David, [Chiriqui], Panama (Champion); gen- postvaginalis. There does not appear to be italia mounted on slide Geometridae No. very much difference between the dissections 5811. Both specimens are labeled "Boarmia of the specimens from the two countries, 1967 RINDGE: GLENA 135 although the Panamanian examples tend to DISTRIBUTION: Nicaragua. have a slightly stronger dorsal protuberance REMARKS: One specimen (a female) and on the posterior portion of the corpus bursae. one genitalic dissection have been studied. The color of the under surface of the type Glena lora, new species specimen is somewhat intermediate between Figure 40 the Panamanian and Costa Rican examples This species is similar to hima, but the t. p. of hima, while the pattern is more like the and extradiscal cross lines are slightly nearer specimens from the former country. We do the base of the wing in the present taxon. not begin to have enough material of either Good differences are present in the female of these species to understand their variabil- genitalia by which to separate the two species. ity. This one is known from Nicaragua. The female genitalia of lora are similar to MALE: Unknown. those of hima. These structures in the present FEMALE: Similar to that of hima, but dif- species can be recognized by the different fering mainly as follows: upper surface of shape of the lamella postvaginalis, by the wings more evenly suffused with brownish difference in the basal portion of the ductus gray scales and with cross lines tending to be seminalis, and by the different shape and posi- more clearly represented; t. p. and ex- tion of the signum. tradiscal cross lines slightly nearer base of Glena agria, new species wing; terminal area grayish brown, quite wide, with complete, crenulate s. t. line; under Figures 14, 41 surface pale grayish brown, with forewings Boarmia demissaria auct.: DRUCE, 1892 (1891- darker than hind wings; terminal band dark 1900), p. 75 (partim). brown, completely represented on all wings; This species resembles nominate unipen- discal dots present on all wings. naria, but the upper surface of the wings LENGTH OF FOREWING: 18 mm. (holotype). tends to be slightly darker in color. The gen- MALE GENITALIA: Unknown. italia should be used for identification. The FEMALE GENITALIA: Sterigma with lamella present taxon occurs from Mexico to Pan- postvaginalis long and narrow, more heavily ama. sclerotized medially, increasing in width an- MALE: Similar to that of unipennaria, dif- teriorly to become three to four times as fering mainly as follows: upper surface of wide as elongate posterior strip, and with wings tending to be slightly grayer; t. p. and large, subtriangular, smoothly sclerotized extradiscal cross lines situated slightly more lateral areas, shorter than lamella postva- basad on wings, with latter line tending to ginalis; intersegmental area between A7 and be more angled in middle of hind wing; under A8 deeply recessed; ductus bursae very short, surface varying in color from white to dull extending more or less vertically from swollen gray, forewings darker than hind wings, and end of lamella postvaginalis; ductus semi- with terminal band incomplete in most speci- nalis arising ventrally as slender tube about mens but completely represented in a few 0.3 mm. anteriad of ductus bursae; corpus examples; hind tibia with hair pencil. bursae swollen anteriorly, posterior portion LENGTH OF FOREWING: 16 to 22 mm.; not sclerotized, and with longitudinal stria- holotype, 16 mm. tions extending to anterior swelling; signum FEMALE: Similar to male but tending to be situated dorsally, large, transverse, narrowly more heavily suffused with dark scales above elliptical in outline, with narrow, stellate and below; maculation of upper surface tend- outer margin, and with flat median area hav- ing to be less clearly represented, and with ing several dentate ridges. terminal band more prominent on under sur- TYPE: Camoapa, w[estern] Nicaragua, ele- face. vation 2000 feet, wet season, 1906 (G. M. LENGTH OF FOREWING: 18 to 22 mm.; Palmer); genitalia mounted on slide Geomet- allotype, 21 mm. ridae No. 5829. This specimen is in the col- MALE GENITALIA: Similar to those of uni- lection of the British Museum (Natural His- pennaria, differing mainly as follows: ap- tory). proximately one-third or one-fourth shorter 136 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 in length; valves with costa concave, slightly ling); genitalia mounted on slide F.H.R. No. widened medially and not enlarged posteri- 13456. Allotype, female, same data; genitalia orly; process of sacculus with terminal spinose mounted on slide F.H.R. No. 13644. Both swelling; length of sacculus and process vary- types are from the author's collection. Para- ing from 1.6 to 1.8 mm.; anellus elongate and types: Mexico: Same data as holotype, var- slender, with length about twice as long as ious dates between September 22 and Octo- width, broadly concave anteriorly, enlarged ber 16, 1962, three males and nine females; posteriorly with convoluted lateral areas, and Mo Cuou, Cerro Pelon, Municipio Yolox, with posterior margins rounded and having Oaxaca, elevation 7050 feet, September 17, prominent median indentation; spine of 1962 (E. C. Welling), one female; Coatepec, vesica very long, somewhat curved and hav- [Veracruz], one male; Orizaba, [Veracruz], ing small hook at apex, and with area of October (F. Knab), one female, March, 1896 slender teeth near end; base of spine broadly (W. Schaus), two males and one female; recurved, its end bluntly pointed; length of Jalapa, [Veracruz], one female, July, 1897 spine about 1.5 mm., of recurved base 0.6 to (Schaus), one male, (M. Trujillo), one male 0.7 mm.; vesica with elongate, slender, and one female, (Hoege), identified as weakly sclerotized area along base of spine "Boarmia demissaria Walk., B. C. A. Lep. bearing numerous short, deciduous spines, Het., Godman-Salvin Coll.," one female; no and with elongate, slender, spinose strip dor- further data, one male. Guatemala: Quirigua, sally and longer and wider denticulate band [Izabal], May (Schaus and Barnes), one male; ventrally. Ventral surface of A3 with median Volcan Santa Maria, [Quezaltenango], Octo- row of bristles. ber (Schaus and Barnes), one female. Costa FEMALE GENITALIA: Sterigma with large, Rica: Puntarenas, Monteverde, elevation sclerotized, elliptical or oval lamella post- 1400 meters, 4600 feet, various dates in Jan- vaginalis, length one and one-half times uary, February, March, and October, 1960- greater than width, and with large, sclero- 1962 (C. W. Palmer), four males and one tized lateral areas, each of their anteromedian female; Sixola River, [Limon], March (Schaus margins extending over one-third of antero- and Barnes), one male. Panama: Chiriqui, lateral portions of lamella postvaginalis, and two males and one female; V[olcan de] Chiri- with posterolateral margins of lateral areas qui, below 4000 feet (Champion), one female, sharply curved outward; intersegmental area 3000-4000 feet (Champion), identified as between A7 and A8 recessed and convoluted "Boarmia demissaria Walk., B. C. A. Lep. ventrally; ductus bursae with length about Het., Godman-Salvin Coll.," one male. one-third of that of lamella postvaginalis; The holotype and the allotype are in the ductus seminalis arising laterally as large collection of the American Museum of Nat- tube on left side of corpus bursae, originating ural History; paratypes are in the collections between 0.7 and 0.8 mm. anteriad of ductus of that institution, of the British Museum bursae; corpus bursae elongate, weakly swol- (Natural History), and of the United States len anteriorly, and with posterior one-fourth National Museum. weakly sclerotized and striate, surface DISTRIBUTION: Mexico (the states of Vera- covered with very small swellings or dentic- cruz and Oaxaca), Guatemala (the provinces ulations; signum in form of long, narrow, of Izabal and Quezaltenango), Costa Rica transverse ellipse, posterior margin weakly (the provinces of Monteverde and Limon), concave, outer margin anteriorly and later- and Panama (the province of Chiriqui). This ally extended and weakly spinose, and with is apparently a montane species, as it occurs flat median area recessed and projecting at elevations of from about 4000 to 7050 anteriorly at angle into ventral wall of corpus feet. bursae, surface flat or weakly ridged. Pos- The adults have been captured in January, terior portion of ventral surface of A7 not February, March, May, July, September, and modified. October. TYPES: Holotype, male, Vista Hermosa, REMAR1KS: A total of 39 specimens (19 Municipio Comaltepec, Oaxaca, Mexico, ele- males and 20 females) and 20 genitalic dis- vation 4650 feet, October 1, 1962 (E. C. Wel- sections (10 males and 10 females) have been 1967 RINDGE: GLENA 137 studied. There is more variation of the under rounded; costa of valves with outer two- surface of the wings in this species than in the thirds slightly swollen and bearing slender preceding taxa. This applies not only to the spines; sacculus sclerotized, extending color of the wings but to the extent of the slightly more than one-half of length of valve, terminal band. As is the usual case with the distal one-third with slender, sclerotized species in this genus, the genitalia should be basal extension of process along outer edge of studied. valve; process of sacculus slender, curving The male genitalia of this species are into middle of valve, not attaining posterior shorter than those of both unipennaria and margin of valvula, and bearing row of heavy, demissaria, but the width of the spread valves outwardly directed spines, consisting of six or is about as great. Additional diagnostic char- seven basal ones in single line and six to eight acters are to be found in the form of the anel- smaller terminal spines in double row; length lus, in the shape of the spine, and in the ar- of sacculus and process 1.9 mm.; anellus with mature of the vesica. wide anterior margin, constricted posteriorly, The female genitalia of agria are like those then widening distally, posterolateral mar- of demissaria in that the ductus seminalis gins extended ventrally and being finely arises as a tube instead of from a large sac. spiculate, and with posterior margin be- The genitalia of the present taxon can be coming membranous, apparently bilobed and separated from those of the preceding species with prominent median incision; spine of by the lateral origin of this tube, and by the vesica without recurved base, gently curved different shape of the signum. medially, and tapering to long, slender point; length of spine 1.0 to 1.1 mm.; vesica ex- Glena trapezia, new species tending at right angle to aedeagus when Figures 15, 42 exserted, with sclerotized area near base of This small species is found in southern spine bearing numerous short, deciduous Brazil; it can be separated from all the pre- spines; posterior surface of vesica with elon- ceding taxa by the absence of the fovea at gate, slender, sclerotized, denticulate strip. the base of the forewing of the male. The Ventral surface of A8 with median row of genitalia should be used for identification. bristles. MALE: Similar to that of demissaria, dif- FEMALE GENITALIA: Sterigma roughly fering mainly as follows: base of forewing trapezoidal in outline, with elongate lamella without fovea; upper surface of wings tending postvaginalis, lightly sclerotized antero- to have more dark scaling in subterminal and medially, increasing in width posteriorly, terminal areas, with distinct s. t. line for most with two elongate, raised, padlike processes; of length of forewing; extradiscal line on sec- lateral areas of sterigma lightly and smoothly ondaries evenly rounded; under surface of all sclerotized, their median margins extending wings white, with forewings more or less out ventrally over lateral margins of lamella shaded with brownish gray; terminal band postvaginalis, and with rear margin be- dark brown, broad on forewings, greatly re- coming more heavily sclerotized and pro- duced and extending only to middle of jecting posteriorly in triangular fashion; secondaries; hind tibia with hair pencil. intersegmental area between A7 and A8 re- LENGTH OF FOREWING: 18 (holotype) to cessed and slightly convoluted; ductus bursae 19 mm. short; ductus seminalis arising on left side of FEMALE: Similar to male, but with both corpus bursae as slightly swollen tube surfaces of wings more heavily suffused with 0.6 mm. anteriad of ductus bursae; corpus grayish brown scales; beneath with terminal bursae with anterior portion swollen, poste- band complete on hind wings but becoming rior one-fifth lightly sclerotized and striate, less intense posteriorly. surface covered with minute denticulations LENGTH OF FOREWING: 20 mm. (allotype). extending farther down left side than on MALE GENITALIA: Uncus triangular, sides right; signum ventrolateral, elongate, trans- weakly concave, apex sharply pointed and verse, anterior margin weakly concave, outer not strongly curved ventrally; gnathos rel- margin weakly denticulate, and with flat atively small, with median enlargement median area recessed and projecting an- 138 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 teriorly at angle into wall of corpus bursae, and with apex more produced; upper surface its surface with minute teeth. Posterior por- of wings pale brownish gray, with very large tion of ventral surface of A7 not modified. costal spots at origin of cross lines; t. p. and TYPES: Holotype, male, Faz[enda] dos extradiscal lines situated nearer base of wing; Campos, Passa Quatro, Minas [Gerais], terminal intravenular dots small; under side Brazil, November 24, 1916 (J. F. Zikan); with nebulous dark brown terminal band in genitalia mounted on slide F.H.R. No. 13540. upper one-half of forewings only, with wide Allotype, female, Alto Serra, Sao Paulo, area of ground color distad thereto, and with Brazil, December 10 (Pohl); genitalia partial t. p. line present in upper portion of mounted on slide F.H.R. No. 13423. Para- wing; discal dots present on all wings; hind type, Petropolis, [Rio de Janeiro, Brazil], one tibia with hair pencil. male. LENGTH OF FOREWING: 20.5 mm. (holo- The holotype and allotype are in the collec- type). tion of the United States National Museum; FEMALE: Unknown. the paratype is in the collection of the MALE GENITALIA: Similar to those of British Museum (Natural History). trapezia, but differing mainly as follows: DISTRIBUTION: Southern Brazil (the states larger, with combined lengths of uncus, of Minas Gerais, Sao Paulo, and Rio de tegumen, and saccus almost one-half again Janeiro). as long; uncus with apex angled ventrally; The moths have been captured in Novem- valves with costa neither swollen nor spinose, ber and December. and extending well posteriad of valvula; sac- REMARKS: Three specimens (two males culus simple, with process not so strongly and one female) and three genitalic dissec- angled basally, thicker, tubular, and bearing tions have been studied. In addition to the band of small spines on posteromedian end of lack of the fovea at the base of the forewing process; length of sacculus and process 1.8 in the male, the adults can be recognized by mm.; anellus with posterior end large, the presence of the broad terminal band on rounded, and with lateral areas weakly the under side of the forewings. On the hind crenulate; spine parallel with aedeagus when wings, the males are without almost any vesica exserted, evenly curved dorsally, and trace of the terminal band, but the females with very short recurved base; length of spine have a wide band, decreasing in intensity 1.15 mm., of recurved base 0.25 mm.; vesica posteriorly. with weakly sclerotized area by base of spine, The male genitalia of this species are very presumably bearing numerous short, decidu- distinctive in that the spine of the vesica does ous spines; vesica with posterior end on dorsal not have a recurved base, and the spining of surface finely granulose, and with large, the process of the sacculus is directed out- apically bifurcate, exsertile caecum on left ward instead of inward and posteriorly. side just posteriad of aedeagus. Ventral sur- The female genitalia are unique in the face of A3 with median row of bristles. genus in the trapezoidal outline of the FEMALE GENITALIA: Unknown. sterigma. TYPE: Holotype, male, Chanchamayo, Glena sacca, new species [Pasco, Peru], elevation 2100-7500 feet; genitalia mounted on slide Geometridae No. Figure 16 5775. This specimen is in the collection of the This species, known from Peru, can be British Museum (Natural History). separated from trapezia by the absence of the DISTRIBUTION: Peru (the department of heavy black terminal band from the under Pasco). side of the forewing. The male genitalia are REMARKS: One specimen and one genitalic distinctive. dissection have been studied. The upper sur- MALE: Similar to that of trapezia, differing face of the unique type is somewhat worn, so mainly as follows: larger; front with narrower all the details of the maculation cannot be white band across bottom; palpi longer; fore- made out. It is also possible that the speci- wing without noticeable fovea but specialized men may be somewhat discolored. scaling present on under side at base of wing, The male genitalia of this species are very 1967 RINDGE: GLENA 139 distinctive in the type of spine and vesica of with apex hooked; spine with elongate, re- the aedeagus. The process of the sacculus is curved base, tending to become slightly mem- also characteristic. branous and flattened apically; length of spine varying from 2.0 to 2.1 mm., of re- Glena bulla, new species curved base from 0.8 to 0.9 mm.; vesica with Figure 17 elongate, narrow, sclerotized strip along base This is a relatively small species with nar- of spine bearing numerous, short, deciduous row, complete, dentate t. p. and extradiscal spines, and with lightly sclerotized, dentate lines, with a strong fovea at the base of the area continued from sclerotized strip, parallel forewing of the male, and with the terminal with spine, with denticulations extending band represented at the apex of the forewing posteriorly and decreasing in size; ventral only. The female is unknown. The taxon oc- surface without denticulate area. Ventral curs in Paraguay, Argentina, and Bolivia. surface of A3 with median row of bristles. The genitalia have good specific characters. FEMALE GENITALIA: Unknown. MALE: Similar to that of trapezia, but dif- TYPES: Holotype, male, Puerto Casado, fering mainly as follows: base of forewing [Boqueron], Paraguay, July 4, 1950; genitalia with small, strongly raised fovea; upper sur- mounted on slide F.H.R. No. 9800. Para- face of wings whitish gray, with very few dark types: P[uerto] Casado, Independencia, Para- scales; t. p. and extradiscal lines complete, guay, November 21, 1949, one male; Vincente narrow, outwardly dentate on veins; terminal Lopez, Buenos Aires, Argentina, August 5, area concolorous with, or only slightly darker 1951, one male; Yunga del Espiritu Santo, than, remainder of wing, with s. t. line Cochabamba, Bolivia, 1888-1889 (P. Ger- tending to appear more prominent as result. main), one male. Under surface of wings white; forewings with The holotype is in the collection of the broad blackish brown terminal band ex- American Museum of Natural History; the tending to vein M3 only, some specimens paratypes are in the collections of that in- with faint, narrow trace of band continued stitution and of the British Museum down primaries and onto hind wings; costa (Natural History). with three spots marking inception of cross DISTRIBUTION: Argentina (the province of lines of upper surface. Hind tibia with hair Buenos Aires), Paraguay (the department of pencil. Boquer6n), and Bolivia (the department of LENGTH OF FOREWING: 16 to 18 mm.; Cochabamba). This apparently is a southern holotype, 16 mm. species that is found at low elevations in the FEMALE: Unknown. Humid Subtropical Zone. MALE GENITALIA: Uncus triangular, with The adults have been taken in July, apical region extending posteriorly, apex with August, and November. parallel sides and terminally bifurcate; REMARKS: A total of four specimens (all gnathos relatively small, well sclerotized, males) and four genitalic dissections have median enlargement not prominent; valves been studied. The fovea at the base of the elongate, with distal two-thirds to three- forewing is of a different shape than the fourths of costa weakly swollen and bearing foveae of any of the preceding species. In numerous setae; sacculus rather short and bulla this modification of the wing is rel- wide; process of sacculus extending inward, atively short but quite high; in the other then angled to subparallel costa, latter part species the structure is more elongate, lower, elongate, slender, and bearing numerous and less prominent. short, heavy setae; length of sacculus and process varying from 2.0 to 2.2 mm.; anellus Glena gemina, new species long and slender, appearing pedicelate from Figures 18, 43 wide base, expanding in width posteriorly by Boarmia demissaria auct.: DRUCE, 1892 (1891- means of several convoluted ridges, and with 1900), p. 75 (partim). membranous posterior margin rounded; spine The adults of this species appear to be very of vesica weakly sinuous, inner surface of similar to those of agria, but the genitalia in- apical region having numerous fine teeth, and dicate a close relationship to bulla. This 140 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 species is known from Central America, and most 1.0 mm. anteriad of ductus bursae; the genitalic structures should be used for corpus bursae with anterior end rotated identifying it. about one-half turn to left, posterior one- MALE: Similar to that of agria, differing eighth lightly sclerotized and striate, surface mainly as follows: upper surface of wings covered with minute denticulations, and with with slightly heavier dusting of brownish striations extending almost entire length of gray scales, particularly in outer portion of corpus bursae; signum situated dorsally or wings; under surface with forewings pale dorsolaterally, elongate, transverse, outer brownish gray, with darker, broad terminal margin stellate, and with flat median area band separated from more basal portion of recessed and having minute teeth. Posterior wing by nebulous pale gray stripe, and with portion of ventral surface of A7 not modified. hind wings similar to forewings but paler; TYPES: Holotype, male, Cerro Zunil, hind tibia with hair pencil. [Quezaltenango], Guatemala, elevation 4000- LENGTH OF FOREWING: 20 mm. (holotype). 5000 feet (Champion), identified as "Boarmia FEMALE: Similar to male, but with both demissaria Walk., B. C. A. Lep. Het., God- surfaces of wings more heavily covered with man-Salvin Coll."; genitalia mounted on grayish brown scales, and with under surface slide Geometridae No. 5826. Allotype, female, having both more prominent terminal band Orosi, [Cartago], Costa Rica, elevation 1200 and white apical patch on forewings. meters (Fassl); genitalia mounted on slide LENGTH OF FOREWING: 23 mm. (allotype). Geometridae No. 5824. Both specimens are in MALE GENITALIA: Similar to those of the collection of the British Museum (Na- bulla, but differing mainly as follows: uncus tural History). with apical region wider and with posterior DISTRIBUTION: This species is known only margin truncate; gnathos with median en- from the mountains of Guatemala and Costa largement slightly longer and more slender; Rica. valves with costa neither enlarged nor REMARKS: Two specimens and two genitalic spinose; sacculus with much shorter process; dissections have been studied. The only safe length of sacculus and process 1.3 mm.; way to distinguish gemina from agria is by anellus almost twice as long as wide, flat studying the genitalia. except for lateral ridges posteriorly, with con- The male genitalia of this taxon show a striction near base, and with posterior margin close relationship to those of bulla, but are to bluntly wedge-shaped and having median in- be distinguished by the much shorter process cision; spine of aedeagus elongate, bisinuate, of the sacculus and by the different configura- and with apex curved; base of spine sharply tion of the anellus. recurved, elongate; length of spine 1.8 mm., of recurved base 0.65 mm.; vesica with Glena laticolla, new species slender sclerotized area along base of spine Figures 19, 44 bearing numerous short, deciduous spines; Boarmia demissaria auct.: DRUCE, 1892 (1891- vesica with two slender, longitudinal, den- 1900), p. 75 (partim). ticulate strips, one each on right and left This small species with variable maculation sides, both near aedeagus. Ventral surface of is found in Mexico and Nicaragua. The geni- A8 with median row of bristles. talia should be used for identification. FEMALE GENITALIA: Sterigma with large, MALE: Similar to that of gemina, differing elongate, ovate lamella postvaginalis, and mainly as follows: upper surface of wings with large lateral areas, more or less sub- more (holotype) or less (paratype) suffused triangular in outline, their anteromedian with grayish brown scales; maculation tend- margins extending out ventrally over antero- ing to be less clearly defined, with large lateral margins of lamella postvaginalis; in- grayish brown patch on forewings in cells M2 tersegmental area between A7 and A8 re- and M3, and with rather nebulous markings cessed, truncate anteriorly; ductus bursae in outer portion of wing; under surface vary- short, narrower than width of lamella post- ing from being dull gray, with faint, broad vaginalis; ductus seminalis arising on left side terminal band (holotype), to whitish gray, of corpus bursae as slightly swollen tube al- with distinct terminal band (paratype), both 1967 RINDGE: GLENA 141 specimens with hind wings paler than fore- The single male from Mexico tends to have a wings; hind tibia with hair pencil. suffused appearance on both surfaces of the LENGTH OF FOREWING: 18 (paratype) to wings, whereas the male from Nicaragua is 19 (holotype) mm. lighter in color and has the maculation on FEMALE: Similar to male, but upper sur- both surfaces of the wings much more con- face more heavily suffused with grayish trasting. brown scales; under surface gray, with dis- The genitalia of this species are very similar tinct, complete terminal band on all wings. to those of gemina. The male can be recog- LENGTH OF FOREWING: 23 mm. (allotype). nized by the much longer and thinner process MALE GENITALIA: Similar to those of of the sacculus, and the female can be known gemina, but differing mainly as follows: sac- by the much wider ductus bursae. culus with process straight, very long, and Glena brachia, new species slender, bearing spines on outer one-fourth or one-third; length of sacculus and process Figures 20, 45 varying from 2.0 to 2.1 mm.; spine of vesica This small to moderate-sized species has less curved and without curved apex; length the outer portion of the upper surface of the of spine 1.45 mm., of recurved base 0.45 mm. wings darkened, strongly dentate t. p. and Ventral surface of A8 with median row of extradiscal lines, and lacks the fovea at the bristles. base of the forewing in the male. It occurs FEMALE GENITALIA: Similar to those of from southern Brazil to Peru. The genitalia gemina, but differing mainly as follows: should be used for identification. sterigma less heavily sclerotized and with MALE: Similar to that of bulla, differing smaller lateral areas; intersegmental area be- mainly as follows: forewing without fovea at tween A7 and A8 not truncate anteriorly and base; upper surface suffused with grayish more convoluted; ductus bursae very broad, brown in terminal area; t. p. and extradiscal at least as wide as lamella postvaginalis; lines strongly dentate; s. t. line whitish gray, ductus seminalis arising about 0.8 mm. outwardly scalloped, more or less complete; anteriad of ductus bursae; corpus bursae under surface whitish gray, with faint pearly shorter, with narrowed area, owing to rota- luster, with blackish brown terminal band tion of anterior portion, situated more poste- complete on forewings, varying from being riorly. absent to being partially represented on hind TYPES: Holotype, male, Puerto Elegio, wings; hind tibia with hair pencil. Municipio Comaltepec, Oaxaca, Mexico, ele- LENGTH OF FOREWING: 17 to 20 mm.; holo- vation 2296 feet, September 26, 1961 (E. C. type, 20 mm. Welling), and from the author's collection; FEMALE: Similar to male but more heavily genitalia mounted on slide F.H.R. No. dusted with dark scales on both surfaces of 10554. Allotype, female, Cordova, Veracruz, wings; terminal band below tending to be [Mexico] (Rumeli), and identified as more heavily represented. "Boarmia demissaria Walk., B. C. A. Lep. LENGTH OF FOREWING: 18 to 21 (allotype) Het., Godman-Salvin Coll."; genitalia mm. mounted on slide Geometridae No. 5834. MALE GENITALIA: Uncus heavily sclero- Paratype, Eden, [Zelaya], Nicaragua, May tized, triangular, with apex wedge-shaped 13, 1922 (Wharton Huber), one male. and terminating in single point; gnathos The holotype and paratype are in the col- heavily sclerotized, lateral areas straplike, lection of the American Museum of Natural with elongate, slender, median projection; History; the allotype is in the collection of valves with costa in form of slender, S- the British Museum (Natural History). shaped, free arm, attached at base only; DISTRIBUTION: Southern Mexico (the states valvula with rounded posterior margin ex- of Veracruz and Oaxaca) and Nicaragua. tending to, or slightly beyond, tip of costa; The adults have been captured in the sacculus short, continued distally as very months of May and September. long, slender process extending as far as tip of REMARKS: A total of three specimens and costa; apex of process in form of slender, three genitalic dissections have been studied. diagonal enlargement, about four times 142 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 longer than width of process, and bearing (F. Plaumann), and from the Sperry collec- numerous spines along posterior margin; tion; genitalia mounted on slide J.L.S. No. length of sacculus and process varying from 1710. Allotype, female, Sapucay, [Paraguari], 2.4 to 3.0 mm.; anellus elongate, with length Paraguay, July 6, 1902 (W. Foster); genitalia twice that of width, with small, smoothly mounted on slide Geometridae No. 4838. sclerotized anterior portion with median in- Paratypes: Brazil: Same data as holotype, dentation and constriction, posteriorly in March, 1953, one male. Paraguay: Same data form of two elongate, slender lobes, separated as allotype (Collins), one male, October 27, by less than own width by more membranous 1903, one male. Bolivia: Prov[ince] del Sara, area; cristae prominent, situated at base of Santa Cruz, elevation 450 meters, October valve near anterior margin of anellus; base of (J. Steinbach), one male; Mapiri R[iver], east valve, on outer surface, with strong patch of Bolivia, one female; latitude 170 46' 55" S., very long, hairlike scales; vesica with well- longitude 630 5' 34" W. [about 6 miles south- developed, S-shaped spine, with hooked apex west of Santa Cruz, Santa Cruz], one male. and with several stout teeth nearby, and with Peru: Satipo, [Junin], December, 1941, strongly recurved base; length of spine August, 1948, October (P. Paprzycki), three varying from 1.3 to 1.5 mm., of recurved base males; Chanchamayo, [Pasco] (Schuncke), from 0.7 to 0.8 mm.; vesica with elongate, one male; Huambo, "IV Trin.," 1889 (M. de slender, lightly sclerotized strip along base of Mathan), one male. spine bearing numerous small, deciduous The holotype is in the collection of the spines; exserted vesica with finely denticulate American Museum of Natural History, and area dorsally and on right side, terminating the allotype is in that of the British Museum in heavily sclerotized strip; left side with (Natural History); paratypes are in the col- much smaller sclerotized and denticulate lections of those two institutions and of the area, in some specimens appearing as several United States National Museum. teeth only; ventral surface striate. Ventral DISTRIBUTION: Southern Brazil (the state surface of A8 with median row of bristles. of Santa Catarina), Paraguay (the depart- FEMALE GENITALIA: Sterigma with heavily ment of Paraguari), Bolivia (the department sclerotized, short, rhomboidal lamella post- of Santa Cruz), and Peru (the departments of vaginalis, with posterior end curved dorsally, Junin and Pasco). and with very large, sclerotized, lateral areas The adults have been taken in March, situated posteriad of lamella postvaginalis July, August, October, and December. and having irregularly curved margins; inter- REMARKS: A total of 13 specimens (11 segmental area between A7 and A8 shallowly males and two females) and 13 genitalic dis- recessed, weakly crenulate and membranous sections have been studied. Specimens from ventrally, becoming sclerotized laterally; duc- Brazil and Paraguay tend to have less of a tus bursae very short; ductus seminalis terminal band on the under surface of the arising from dorsal surface of corpus bursae wings in males than do those from Bolivia as swollen tube, originating about 0.6 mm. and Peru. In addition to this variability, anteriad of ductus bursae; corpus bursae rel- there seem to be some seasonal differences atively short and broad, with posterior one- within this taxon, as far as can be determined fourth to one-third weakly sclerotized, sur- from the limited amount of dated material. face covered with very small swellings or Specimens from Peru, dated May, have a denticulations, and having several small much heavier terminal band than do the ex- ridges; signum on dorsal surface of corpus amples dated October and December, as do bursae, long and slender, transverse, outer two Brazilian specimens dated March and margin smooth and tending to be slightly October. Much more material of this species wider on anterior side, and with flat median is needed before the entire picture of vari- area recessed and projecting anteriorly a-t ability can be satisfactorily explained. angle into wall of corpus bursae. This species can be recognized by the TYPES: Holotype, male, Nuevo Teutonia, unique, free costal arm of the costa in the [Santa Catarina], Brazil, October 28, 1948 male genitalia. The form of the uncus is dif- 1967 RINDGE: GLENA 143 ferent from that of all the preceding species, several ridges and with two ridges running although it is to be found in the succeeding length of anellus; base of valve on outer taxa. surface with strong hair pencil; spine of vesica The female genitalia of brachia are dis- slightly sinuous, with slightly curved apex tinctive in that the sterigma is heavily having thornlike projections apically, and sclerotized, with the lamella postvaginalis with sharply recurved base; length of spine anteriad of the irregularly shaped lateral 2.2 to 2.3 mm., of recurved base 0.85 mm.; areas. vesica with sclerotized strip near base of spine presumably bearing numerous short, decidu- ous spines; exserted vesica with slender, den- Glena cretacea (Butler), new combination ticulate strip extending to sclerotized, pro- Figure 21 jecting tip of aedeagus, in length about one- Tephrosia ? cretacea BUTLER, 1881, 327. half of that of spine, and with shorter, dentic- p. ulate area on vesica under outer portion of This species is similar to brachia, but the spine. Ventral surface of A3 with median row upper surface of the wings tends to be of bristles. slightly more brownish. The genitalia should FEMALE GENITALIA: Not examined. A be used for determination. This taxon occurs photograph of the genitalia of the type in- in Brazil and Peru. dicates that these structures are similar to MALE: Similar to that of brachia, differing those of totana, quadrata, and mopsaria. In mainly as follows: upper surface of wings the present species the sterigma is narrower slightly more suffused with brownish scaling; than the ductus bursae, and ends posteriorly cross lines tending to be more strongly repre- in two widely divergent points. sented; under surface of wings with terminal TYPE: Butler described cretacea from a band present on forewings, varying from single female; this specimen is in the collec- prominent to absent on hind wings; hind tibia tion of the British Museum (Natural His- with hair pencil. tory). LENGTH OF FOREWING: 18 mm. TYPE LOCALITY: "Prainha, Amazons," FEMALE: Not examined (known only from Brazil. unique type). MALE GENITALIA: Uncus heavily sclero- DISTRIBUTION: The Amazon basin, in both tized, triangular, with apex slightly attenuate Brazil and Peru (the department of Madre de and curved ventrally; gnathos heavily sclero- Dios). (See Appendix for locality data of tized, with large median projection termi- specimens examined.) nally rounded; valves with slender costa en- The adults have been captured in the larged distally into hirsute, triangular pro- months of September and November (the tuberance jutting inward toward center of type specimen). valve, not attaining end of valvula, and with REMARKS: Two specimens (both males) inner margin of costa irregular in outline and and two genitalic dissections have been bearing small spines; sacculus short but with studied. The under surface of the wings of very long and slender process extending be- these two males show the same type of varia- yond end of costa, outer one-half of process tion that is found in brachia. The Peruvian slightly wider than basal portion, and with specimen, caught in September, has but a apex somewhat enlarged and bearing curved very faint terminal band on the hind wings, band of spines extending from surface of whereas the undated Brazilian example has a valvula to ventral surface of process; length heavy terminal band on the secondaries. of sacculus and process varying from 2.9 to The male genitalia of this species are 3.0 mm.; anellus long and slender, with length basically similar to those of brachia. The about twice as great as width, strongly con- present taxon can be distinguished by the stricted above base, enlarging posteriorly, differences in the valves, particularly in the with median incision separating wedge- costa and process of the sacculus, and by the shaped posterior segments, lateral areas with nature of the anellus. 144 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

Glena gampsa, new species maculation these two species are very similar, Figure 22 but good differences are present in both the distribution and in the male genitalia to This species is very similar to brachia; it separate them. The male structures of the occurs in Venezuela. The genitalia should be present taxon can be recognized by the more used for identification. pointed uncus, the longer and more slender MALE: Similar to that of brachia, differing gnathos, and by the fact that the process of mainly as follows: upper surface of wings the sacculus does not extend beyond the apex with outer portion slightly darker and with of the costa. s. t. line complete; t. p. line tending to be more S-shaped; under surface with terminal Glena labecula, new species band broadly represented on forewings and Figure 23 on anterior portion of hind wings; hind tibia This species is very similar to brachia and with hair pencil. cretacea; it occurs in Bolivia. The genitalia LENGTH OF FOREWING: 17 mm. (holotype). should be used for identification. FEMALE: Unknown. MALE: Similar to that of brachia, differing MALE GENITALIA: Similar to those of mainly as follows; upper surface of wings cretacea, differing mainly as follows: smaller slightly more suffused with brownish scaling; by approximately one-fifth; uncus tapering cross lines tending to be more strongly repre- to single point; gnathos with elongate, sented; nebulous brownish gray spot present slender, median enlargement; valves with in cell M2 distad of t. p. line; under surface costa having inner margin without spines and without pearly luster, with dull brownish with apex rounded and with small median black terminal band on all wings; hind tibia cleft; sacculus with very long and slender pro- with hair pencil. cess extending as far as end of costa, with LENGTH OF FOREWING: 18 mm. (holotype). apical region curving medially, becoming FEMALE: Unknown. swollen and bearing thick group of short MALE GENITALIA: Similar to those of spines on posterior surface; length of sacculus cretacea, but differing mainly as follows: and process 2.1 mm.; anellus shorter and uncus shorter, more triangular; gnathos with broader, with posterior margin more or less much more slender median enlargement; truncate; spine of vesica slender, straight, costa of valves somewhat S-shaped, extend- with apex tapering and finely hooked at end ing slightly beyond valvula, with apex some- and with sharply recurved base; length of what attenuate and with slender, finger-like spine 1.0 mm., of recurved base 0.3 mm.; projection arising from inner margin, length vesica with sclerotized strip near base of of projection equal to width of costa; sac- spine bearing numerous short, deciduous culus with process shorter than costa, apex spines; exserted vesica with small swelling swollen and bearing spines posteriorly; length near middle of spine, having apical portion of sacculus and process 2.8 mm.; anellus sclerotized, and with elongate, slender den- slender, length two and one-half times width, tate strip on opposite side of vesica. Ventral posterolateral areas with several longitudinal surface of As with median row of bristles. ridges and with posterior margin deeply FEMALE GENITALIA: Unknown. bilobed; spine of vesica shallowly S-shaped, TYPE: Holotype, male, San Esteban, with several teeth posteriorly and sharply Venezuela, June, 1909 (S. M. Klages); geni- recurved base; length of spine 1.5 mm., of talia mounted on slide Geometridae No. recurved base 0.5 mm.; vesica with sclero- 5840. The type is in the collection of the tized strip near base of spine presumably British Museum (Natural History). bearing numerous short, deciduous spines; DISTRIBUTION: Venezuela. vesica with elongate, broad, denticulate strip, REMARKS: One specimen and one genitalic almost as long as spine. Ventral surface of As dissection have been studied. Although the with median row of bristles. type of this species is about the same size as FEMALE GENITALIA: Unknown. the adults of brachia, the genitalia of the TYPE: Holotype, male, Chapare, [Beni], male are considerably smaller. In color and Bolivia, April, 1951; genitalia mounted on 1967 RINDGE: GLENA 145 slide F.H.R. No. 9860. This specimen is vesica with additional small, denticulate from the Sperry collection and is in the Amer- patch near end of aedeagus under spine. ican Museum of Natural History. Ventral surface of A3 with median row of DISTRIBUTION: Bolivia (the department of bristles. Beni). FEMALE GENITALIA: Sterigma with lamella REMARKS: One specimen and one genitalic postvaginalis bilobate, in width two-thirds dissection have been studied. The only safe as wide as posterior portion of corpus bursae, way to separate this species from the preced- and with large, sclerotized, winglike, lateral ing one is by a study of the genitalia. areas united medially, twice as wide as pos- The male genitalia of labecula are like terior portion of corpus bursae and becoming those of brachia but can be distinguished by narrowed medially; intersegmental area be- the fact that the costa of the valves extends tween A7 and A8 weakly sclerotized and beyond the process of the sacculus. deeply recessed ventrally; ductus bursae not Glena new differentiated; ductus seminalis arising dor- totana, species sally from corpus bursae as slender tube 1.1 Figures 24, 46 mm. anteriad of anterior margin of interseg- Boarmia bipennaria auct.: KAYE AND LAMONT, mental area; corpus bursae gradually increas- 1927, p. 99 (partim). ing in width anteriorly, with posterior two- This species is very similar to the preced- fifths sclerotized, surface covered with very ing taxa; it occurs in Trinidad. The genitalia small swellings or denticulations; signum on should be used for identification. dorsal surface near anterior end of corpus MALE: Similar to that of brachia, differing bursae, outer margin smooth and wider on mainly as follows: upper surface of wings anterior side, and with flat median area re- slightly more suffused with gray scales; s. t. cessed and projecting anteriorly at angle into line tending to be slightly more strongly rep- wall of corpus bursae. resented, emphasized by dark gray or grayish TYPES: Holotype, male, Arima Valley, black scales along inner margin of line; under Trinidad, elevation 800-1200 feet, February surface without pearly luster and tending to 10-22, 1964 (J. G. Rozen and P. Wygodzin- be more strongly suffused with dark scales, sky); genitalia mounted on slide F.H.R. No. particularly on forewings; dark terminal band 12544. Allotype, female, Palmiste, Trinidad, present on all wings; hind tibia with hair March 7, 1917 (Sir N. Lamont); genitalia pencil. mounted on slide Geometridae No. 5825. LENGTH OF FOREWING: 16 to 18 mm.; holo- Paratypes, all from Trinidad: Same data as type, 16 mm. holotype, eight males, February 20, 24, 1966 FEMALE: Similar to male, but with more (J. G. Rozen), two males; no further data, light brown scaling on upper surface of wings two females. and with brown band distad of t. p. line; under The holotype and the male paratypes are surface dark brownish gray, with wider and in the collection of the American Museum of darker terminal band on all wings. Natural History; the allotype is in the collec- LENGTH OF FOREWING: 17 to 19 mm.; allo- tion of the British Museum (Natural His- type, 17 mm. tory), and the female paratypes are in the MALE GENITALIA: Similar to those of United States National Museum. labecula, but differing mainly as follows: DISTRIBUTION: Trinidad. The adults have costa straight, with slender, finger-like pro- been taken in February and March. jection arising from near middle of distal REMARKS: A total of 14 specimens (11 margin and with length of projection slightly males and three females) and five genitalic shorter than width of costa; sacculus with dissections (three males and two females) have shorter capitate process bearing fewer and been studied. Kaye and Lamont (1927, p. 99) longer spines; length of sacculus and process reported two specimens of "bipennaria" from 1.5 to 1.6 mm.; spine of vesica with weakly Trinidad. One of these is the allotype of curved apex, several thornlike teeth poste- totana. The second specimen (Guaico, April riorly, and sharply recurved base; length of 18, 1915) has not been examined, so it is not spine 1.4 mm., of recurved base 0.45 mm.; known whether it belongs to this species or 146 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 to demissaria, which also occurs on Trinidad. FEMALE GENITALIA: Sterigma with lamella The male genitalia of this species are to be postvaginalis more or less square in outline, separated from those of labecukl by the much almost as wide as posterior portion of corpus shorter sacculus and process. In the present bursae, increasing in width anteriorly to taxon they are slightly less than one-half of ductus bursae, with posterior margin weakly the length of these structures in the previous concave, and with large, sclerotized, winglike species. lateral areas, united medially by small, more The female genitalia of totana are quite membranous area, and with posterior margin distinct from any of the previously described outwardly oblique; intersegmental area be- structures in that the sterigma has the lateral tween A7 and A8 deeply recessed ventrally; areas united medially and much wider than ductus bursae not differentiated, perhaps the lamella postvaginalis. represented by anterior enlargement of lamella postvaginalis; ductus seminalis aris- Glena quadrata, new species ing dorsally from corpus bursae as slender Figures 25, 47 tube 1.0 mm. anteriad of anterior margin of This species closely resembles totana, but it intersegmental area; corpus bursae gradually is browner; it is known from French Guiana. increasing in width anteriorly, with posterior The genitalia should be used for identifica- two-fifths sclerotized, surface covered with tion. very small swellings or denticulations; sig- MALE: Similar to that of brachia, differing num on dorsal surface near anterior end of mainly as follows: upper surface of wings corpus bursae, outer margin smooth and more suffused with brown scales; cross lines wider on anterior side, and with flat median more strongly represented, with s. t. line area recessed and projecting anteriorly at emphasized by dark gray or grayish black angle into wall of corpus bursae. scales along inner margin of line; under sur- TYPES: Holotype, male, allotype, female, face without pearly luster and more strongly plus nine male and two female paratypes, all suffused with dark scales; dark terminal band from Cayenne, F[rench] Guiana. The geni- present on all wings; hind tibia with hair talia of the holotype are on slide F.H.R. No. pencil. 13561, of the allotype, F.H.R. No. 13584. All LENGTH OF FOREWING: 16 to 18 mm.; the type specimens are in the collection of the holotype, 17 mm. United States National Museum. FEMALE: Similar to male, but with heavier DISTRIBUTION: French Guiana. suffusion of brown and brownish gray scales REMARKS: A total of 13 specimens (10 on both upper and under surfaces of wings; males and three females) and four genitalic maculation less clearly represented above dissections (two males and two females) have and with heavier terminal band below. been studied. This species closely resembles LENGTH OF FOREWING: 18 to 19 mm.; the several preceding taxa. The upper sur- allotype, 19 mm. face of the wings has more brown scaling than MALE GENITALIA: Similar to those of do the others. Other differences are to be totana, differing mainly as follows: process of found in its distribution, and in the geni- sacculus slightly longer, apical process elon- talia. gate, bearing numerous outwardly curving The genitalia are of the same type as is spines on dorsal surface; length of sacculus found in the preceding taxa. In quadrata the and process 1.8 to 1.9 mm.; spine of vesica process of the sacculus is shorter than that with curved tip, with several thornlike teeth of cretacea and slightly longer than that of posteriorly, and with sharply recurved base, totana. The configuration of the apical section apical region of which becomes flattened and of the sacculus process, and the position and curves posteriorly; length of spine 1.2 to 1.3 length of the spines thereupon, are also dis- mm., of recurved base, 0.5 mm.; vesica with tinctive. dorsal, denticulate strip tending to be shorter The female genitalia of this species can be and broader. Ventral surface of As with me- recognized by the square lamella postva- dian row of bristles. ginalis. 1967 RINDGE: GLENA 147 Glena mopsaria (Schaus), new combination with narrow slit on midline; intersegmental Figures 26, 48 area between A7 and A8 deeply recessed ven- Boarmia plenaria auct.: DRUCE, 1892 (1891- trally; ductus bursae very short, poorly de- 1900), p. 75 (partim). fined; ductus seminalis arising dorsally from Alcis mopsaria SCHAUS, 1913, p. 347. corpus bursae as slender tube between 1.0 and 1.5 mm. anteriad of ductus bursae; corpus This species is very similar to the preceding bursae relatively thick, with posterior ones, but it tends to have less of a terminal one-fourth to one-third sclerotized, surface band on the under side of the wings of the covered with very small swellings or denticu- male. It occurs from Costa Rica to Colombia lations; signum on dorsal surface near ante- and Venezuela. The genitalia should be used rior end of corpus bursae, outer margin vary- for identification. ing from smooth to finely denticulate, wider MALE: Similar to that of brachia, differing on anterior side, and with flat median area mainly as follows: upper surface of wings recessed and projecting anteriorly at angle slightly more irrorate with brown scales, and into wall of corpus bursae. tending to have t. p. and extradiscal lines TYPE: Schaus described mopsaria from a more dentate and more heavily represented; single male specimen; it is U.S.N.M. No. under surface with dark terminal band on 17974. The genitalia are mounted on slide forewings, varying from being absent from, F.H.R. No. 13531. to being partially represented on, hind wings; TYPE LOCALITY: Juan Vinas, Avangarez, hind tibia with hair pencil. Costa Rica. LENGTH OF FOREWING: 16 to 19 mm. DISTRIBUTION: Costa Rica, Panama, Co- FEMALE: Similar to male, but with both lombia, and Venezuela. (See Appendix for upper and under surfaces of wings tending to locality data of specimens examined.) be more heavily dusted with dark scaling; The adults have been captured in March, cross lines on upper surface less clearly rep- April, June, and October. resented and with terminal band on under REMARKS: A total of 17 specimens (12 surface broader. males and five females) and 13 genitalic dis- LENGTH OF FOREWING: 17 to 20 mm. sections (eight males and five females) have MALE GENITALIA: Similar to those of been studied. labecula, but differing mainly as follows: The male genitalia of this species are sim- gnathos with median enlargement shorter, ilar to those of the three preceding species. broader, and with truncate posterior margin; The present taxon can be distinguished, in costa straight, with very short projection these structures, by the shorter and thicker arising from outer angle; sacculus with process of the sacculus and by the much shorter, wider capitate process bearing spines shorter projection from the end of the costa. along outer margin; length of sacculus and The female genitalia are also similar to process varying from 1.4 to 1.5 mm.; anellus those of the preceding species. These organs more slender; spine of vesica weakly sinuous, of mopsaria can be separated from those of with hooked apex and a few projecting teeth totana by the wider lamella postvaginalis medially, and with sharply recurved base; which projects farther posteriorly than does length of spine varying from 1.3 to 1.5 mm., that structure in the former taxon. of recurved base from 0.35 to 0.45 mm.; Five females have been available for dis- vesica without small denticulate patch under section. The one from Costa Rica has an ir- spine. Ventral surface of A3 with median row regular posterior margin to the lateral areas of bristles. of the sterigma, and the lamella postvaginalis FEMALE GENITALIA: Sterigma with lamella has the two swellings near the midline. Both postvaginalis slightly narrower than ductus Colombian examples have a heavily sclero- bursae, posterior margin weakly convex lat- tized and smooth posterior margin to the erally and concave medially, and with large, lateral areas, and the swellings of the lamella winglike lateral areas not quite twice as wide postvaginalis are situated farther from the as ductus bursae, narrowing medially and midline. The two Panamanian specimens are 148 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 somewhat intermediate between the above. very weakly sclerotized area on dorsal surface The remainder of the genitalia are very simi- near end of aedeagus. Ventral surface of A3 lar in all the specimens. with median row of bristles. FEMALE GENITALIA: Sterigma with lamella Glena turba, new species postvaginalis elongate, weakly sclerotized, Figures 27, 49 and with large, more heavily sclerotized lat- This species is very similar to the preced- eral areas, their median margins extending ing ones, so the genitalia should be used for partially over lateral margins of lamella post- identification. It occurs in southern Brazil. vaginalis; intersegmental area between A7 MALE: Similar to that of brachia, differing and A8 recessed, produced anteriorly as two mainly as follows: upper surface of wings rounded points, concave between; ductus more heavily suffused with grayish brown bursae small; ductus seminalis arising on scales, and tending to have t. p. and extra- left side of corpus bursae as slender tube, discal lines represented by heavier venular starting about 0.4 mm. anteriad of interseg- dots; under surface more suffused with dark mental area; corpus bursae widening poste- brownish scales, and with broad terminal riorly from narrow ductus bursae, and with band present on forewings and on anterior posterior end lightly sclerotized, surface portion of hind wings; hind tibia with hair covered with minute denticulations; signum pencil. elongate, transverse, lateral margins strongly LENGTH OF FOREWING: 17 mm. (holotype). stellate, anterior and posterior margins FEMALE: Similar to male, but with both weakly stellate. Posterior portion of ventral upper and lower surfaces of wings tending to surface of A7 not modified. be more heavily dusted with dark scaling; TYPES: Holotype, male, Rio Laeiss, Blu- cross lines on upper surface less clearly rep- menau, Santa Catarina, [Brazil], October, resented, and with terminal band on under 1934 (F. H. Hoffman); genitalia mounted on surface broader. slide Geometridae No. 5736. Allotype, fe- LENGTH OF FOREWING: 17 mm. (allotype). male, Santa Catarina, Brazil; genitalia MALE GENITALIA: Uncus with relatively mounted on slide Geometridae No. 5735. narrow base, elongate, slightly tapering, apex Both specimens are in the collection of the broad and bluntly rounded, terminating in British Museum (Natural History). single point; gnathos with small, rounded, DISTRIBUTION: Southern Brazil (the state median projection; valves with outer three- of Santa Catarina). fourths of costa straight, with parallel sides REMARKS: Two specimens (one male and and extending beyond valvula; valvula one female) and two genitalic dissections broad, posterior margin truncate, with have been studied. The male genitalia of this weakly sclerotized trough with irregular mar- species are very different from any of these gins extending from base of valve toward structures in the preceding species. This capitate enlargement of sacculus process, taxon, and the following taxa, can be recog- and with dense group of heavy, ventrally nized by the prominent patch of spines at the projecting spines at base near costa; sacculus base of the costa. Glena turba is characterized elongate, with short, capitate, spinose proc- both by the elongate process of the sacculus ess; length of sacculus and process 1.6 mm.; and by the elongate anellus. anellus elongate, with wide base and sharply The corpus bursae of the allotype is not constricted median area, increasing in width fully inflated, as the specimen was not mated. posteriorly, lateral surfaces rugose, posterior Additional details concerning this portion of margin with well-defined median incision; the genitalia will have to be supplied when spine of vesica weakly S-shaped, with more material comes to hand. strongly recurved base; length of spine 0.8 mm., of recurved base 0.3 mm.; vesica with Glena tyrbe, new species large, elongate, lightly sclerotized area near Figure 28 base of spine bearing numerous small, de- As far as can be determined from the worn ciduous spines; vesica with narrow, longi- condition of the unique type, tyrbe can be tudinal, denticulate strip ventrally and with distinguished by the well-developed fovea at 1967 RINDGE: GLENA 149 the base of the forewing of the male and by the type is badly rubbed, so that most of the the large discal spots on the upper surface of scales on the forewings are missing. As far as all wings. The male genitalia are very dis- can be determined from what scaling is left, tinctive. This taxon occurs in Peru. tyrbe has large discal spots on all wings above. MALE: Similar to that of turba, differing This is reminiscent of species of Iridopsis mainly as follows: base of forewing with Warren, but the genitalia do not agree with strongly developed fovea; upper surface of those of that genus. wings more heavily dusted with pale brown The male genitalia are similar to those of and brownish black scales; maculation mostly turba, but can be recognized by the differently lacking because of rubbed condition of shaped uncus, by the different ornamentation specimen, with large, dark discal spots having of the valves, and by the much shorter anel- centers of whitish gray on all wings; s. t. lus. line represented by larger spots; under sur- face with broad terminal band represented Glena asaccula, new species on primaries and becoming obsolescent pos- Figures 29, 50 teriorly on secondaries; large discal spot on This is a species with pale wings, having each forewing, absent from hind wing; hind the cross lines represented by large dots; the tibia with hair pencil. male does not have a fovea at the base of the LENGTH OF FOREWING: 17 mm. (holotype). forewing. The genitalia are very distinctive FEMALE: Unknown. in both sexes. This taxon occurs in southern MALE GENITALIA: Similar to those of Brazil. turba, differing mainly as follows: uncus very MALE: Similar to that of brachia, but dif- long, with posterior portion extending ven- fering mainly as follows: upper surface of trally at nearly right angle from anterior por- wings whitish gray, with very few dark scales tion and ending in slender point; valves with basad of t. p. line; cross lines represented by costa more curved, weakly enlarged apically, much larger, unconnected spots; t. p. and ex- and with larger group of spines at base; sac- tradiscal lines shaded distally by brown; s. t. culus and process wider and shorter, apex line complete on all wings; under surface capitate, with numerous spines arising from whitish gray, forewings with very faint brown dorsal and posterior surfaces; length of sac- suffusion; discal spots present on all wings culus and process 1.5 mm.; anellus short, beneath; terminal band dark brownish black, widely expanding from narrow base, posterior complete on forewing, partially represented margin apparently deeply concave; spine of on hind wings; hind tibia with hair pencil. vesica with apical portion angled, weakly LENGTH OF FOREWING: 20 mm. (holotype). enlarged, bearing small spines along inner FEMALE: Similar to male, but with both margin, and having strongly recurved base; surfaces of wings much more heavily suffused length of spine 0.9 mm., of recurved base with dark scaling; upper surface with cross 0.45 mm.; vesica with elongate, slender, lines much less strongly represented; under sclerotized strip near recurved base of spine, surface with broader and more complete ter- with elongate, slender, dentate strip near apex minal band. of spine, and with ventral surface slightly LENGTH OF FOREWING: 22 mm. (allotype). swollen and very finely setose. Ventral sur- MALE GENITALIA: Uncus short, triangular, face of AB with median row of bristles. with apex projecting ventrally into single FEMALE GENITALIA: Unknown. point; gnathos with elongate median projec- TYPE: Holotype, male, Satipo, [Junin], tion; valves with weakly biconcave costa ex- Peru (P. Paprzycki), from the John L. Sperry tending beyond valvula, apex bent at right collection; genitalia mounted on slide F.H.R. angle and projecting as long, pointed process; No. 13693. This specimen is in the collection sacculus short, with weakly developed pro- of the American Museum of Natural History. cess extending only to middle of outer mar- DISTRIBUTION: Peru (the department of gin of valvula; length of sacculus and process Junin). 1.35 mm.; transtilla membranous, but ap- REMARKS: One specimen and one genitalic parentlyprojectinglaterallyas strongly sclero- dissection have been studied. Unfortunately tized arm into middle of valve, terminating ISO BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135 in four or five elongate, ventrally projecting swellings or denticulations; signum absent. spines; anellus smoothly sclerotized, one and TYPES: Holotype, male, Rio Laeiss, Blu- one-half times longer than wide, with weak menau, Santa Catarina, [Brazil], December, basal constriction, raised lateral margins, 1933 (F. H. Hoffman); genitalia mounted on and with posterior margin rounded; spine of slide Geometridae No. 5728. Allotype, fe- vesica weakly S-shaped, and with strongly male, Jaragua do Sul, Santa Catarina, [Bra- recurved base; length of spine 1.4 mm., of zil], September, 1932 (F. H. Hoffman); recurved base 0.4 mm.; vesica with elongate, genitalia mounted on slide Geometridae no. weakly sclerotized area near base of spine 5726. Both type specimens are in the collec- bearing numerous short, deciduous setae; tion of the British Museum (Natural His- vesica with dorsal surface bearing two widely tory). separated, spinose patches near end of ae- DISTRIBUTION: Southern Brazil (the state deagus, and with elongate, slender denticu- of Santa Catarina). late area on ventral surface. Ventral surface REMARKS: A total of two specimens (one of A3 with median row of bristles. male and one female) and two genitalic dis- FEMALE GENITALIA: Sterigma with very sections have been studied. large, subtriangular, sclerotized lamella post- The genitalia of asaccula are distinctive vaginalis and with membranous lateral areas; and somewhat atypical of Glena. The males intersegmental area between A7 and A8 do not have an elongate process of the sac- weakly sclerotized and recessed ventrally; culus, and it would appear that this structure ductus bursae very short; ductus seminalis has been replaced by the lateral extension of arising as slender tube ventrolaterally on the transtilla. The female genitalia are left side of corpus bursae about 0.4 mm. an- unique, insofar as these structures are now teriad of ductus bursae; corpus bursae elon- known, in having neither a signum nor the gate, with posterior one-eighth sclerotized sclerotized lateral areas of the sterigma. and striate, surface covered with very small

LIST OF THE NEOTROPICAL SPECIES OF THE GENUS GLENA, WITH THEIR KNOWN DISTRIBUTION 1. effusa, new species Southern Mexico to Bolivia 2. bipennaria (Guenee) a. bipennaria (Guenee) Southern Brazil, Paraguay, Bolivia, Peru, and Ecuador b. demissa, new subspecies French Guiana 3. dentata, new species Ecuador and Colombia 4. grandillosa (Dognin) Argentina, Bolivia, Peru, and Ecuador 5. sucula, new species Colombia 6. vesana, new species Bolivia and Peru 7. bisulca, new species Ecuador and Colombia 8. juga, new species Paraguay, Bolivia, Peru, Ecuador, and Colombia 9. megale, new species Venezuela, Colombia, Ecuador, and Peru 10. unipennaria (Guen6e) a. unipennaria (Guenee) Southern Brazil and Paraguay b. cosmeta, new subspecies Panama and the Guianas to Bolivia 11. uncata, new species Southern Mexico to Costa Rica 12. demissaria (Walker) Brazil, Uruguay, Paraguay, Bolivia to Colombia, Venezuela, the Guianas, and Trinidad 13. basalis, new species Costa Rica 14. hima, new species Costa Rica and Panama 15. lora, new species Nicaragua 16. agria, new species Southern Mexico to Panama 17. trapezia, new species Southern Brazil 18. sacca, new species Peru 19. bulla, new species Argentina, Paraguay, and Bolivia 20. gemina, new species Guatemala and Costa Rica 1967 RINDGE: GLENA 151

21. laticolla, new species Southern Mexico and Nicaragua 22. brachia, new species Southern Brazil, Paraguay, Bolivia, and Peru 23. cretacea (Butler) Brazil and Peru 24. gampsa, new species Venezuela 25. labecula, new species Bolivia 26. totana, new species Trinidad 27. quadrata, new species French Guiana 28. mopsaria (Schaus) Costa Rica, Panama, Colombia, and Venezuela 29. turba, new species Southern Brazil 30. tyrbe, new species Peru 31. asaccula, new species Southern Brazil APPENDIX: SPECIMENS EXAMINED ALTHOUGH THE RANGE is summarized under Glena grandillosa (Dognin) the heading of Distribution for each taxon, ARGENTINA: Jujuy, Jujuy, Jan., May, June, the complete data are given below for all 1948, Jan., Feb., Mar., May, Dec., 1950 (J. specimens examined of previously described Foerster), 15 6, 6 9; La Almona, Jujuy, Feb. species. Whenever possible, the department, 16, 1949, 16'; Prov. Jujuy, Mar., Apr., May, province, or state is given for the larger coun- 1948, Mar., Apr., 1949, Jan., Feb., Mar., Dec., tries; if this was not indicated on the label it 1950, 166', 4 9; Tucuman, [Tucuman], 1 9, (R. Schreiter), 46', 2 9, Apr., 1903 (L. Monetti), 19, has been inserted in brackets. The spelling (F. Gerard), 19 lectotype; Siambon, Tucuman, of certain names has been altered from that 1600 meters, Jan., 1931 (R. Schreiter), 19; no indicated on the labels, unless given in quo- additional data (W. Rait-Smith), 16'. tation marks, to conform with the approved BOLIVIA: Chapare, [Beni], Aug. 23, 1949 (Pefla), present-day spelling. 1 6; Rio Songo, Prov. Yungas, Feb.-May, 1896, The arrangement of the species is the same 1 6, 750 meters (Fassl), 1 6; R[io] Tanampaya, as that in the text. 1894 (Garlepp), 16'; Chulumani, [La Paz], 2000 meters, Dec., 1900 (Simons), 1 6; La Paz, 1000 GENUS GLENA HULST meters (E. Garlepp), 26; Charaplaya, [Cocha- Glena bipennaria bipennaria (Guen6e) bamba], 1300 meters, June, 1901 (Simons), 1 6; Yunga del Espiritu Santo, Cochabamba, 1888- BRAZIL: Nuevo Teutonia, [Santa Catarina], 1889 (P. Germain), 56; Suapi (Garlepp), 16'. various dates in Jan., Mar., Apr., May, July, PERU: Coosnipata, Paucartambo, Cuzco, Nov. Aug., Sept., and Nov., 1943-1953 (F. Plaumann), 14 to Dec. 7, 1951 (F. L. Woytkowski), 24 d; Cal- 106', 3 9; Santa Catarina, Aug., 1882, 1 c; Jara- langa, Paucartambo, Cuzco, Feb. 14-17, 1953 gua do Sul, Santa Catarina, Jan. 30 (Hoffmann), (F. L. Woytkowski), 5 6; Marcapata, eastern 1 6; Castro, Parani, 950 meters (E. D. Jones), Peru, 4500 ft., 16'; Chanchamayo, [Pasco], 16'; 3 d; Iguassu, Parani, Oct. 15, 1921, 1 dc; Rio [de] Tinguri, Carabaya, [Puno], 26, 3400 ft., dry Janeiro, [Guanabara], 1 d, 1 9; Burity, 30 miles season, Aug., 1904 (G. Ockenden), 26; R[io] northeast of Cuyaba, Mato Grosso, 2250 ft., Sept. Inambari, La Oroya, [Puno], 3100 ft., dry season, 22-30, Oct. 1-15, 1927 (C. L. Collenette), 1 d, Sept., 1904, wet season, Oct., 1904 (G. Ockenden), 1 9; Santa Cruz, Bresil meridionale, 1 '; Sao 36; La Oroya, Carabaya, [Puno], 3000 ft., Mar., Paulo, [Sao Paulo], Apr., 1 9; no further data, 1905 (G. Ockenden), 1 d; Oxapampa, [Pasco], type c. 6400 ft., 26; El Porvenir, 900 meters, Oct., 1909, PARAGUAY: Caa-genz'u, Nov. 20, Dec. 6, 1950, 1 6; Cerro de Pasco, Huancabamba, 6000-10,000 2 c; Carlos Pfannl, Aug. 6, 1951 (Schade), 1 6; ft. (E. Boettger), 5 6; Huancabamba, 6000- Independencia, [Caazapa], various dates in Oct., 10,000 ft., 16 ; Santo Domingo, Carabaya, Nov., and Dec., 1950, 66'; Villarrica, Feb., 1924 [Puno], 46; Pampaconas R[iver], Aug., 1911 (P. Jorgensen), 1 c; Sapucay, [Paraguari], Nov. (Yale Peruvian Expedition), 1 6. 17, 1908 (W. Forster), 1 c; no further data, 1 c. ECUADOR: Loja, [Loja], Sept., 1896, 1 9. BOLIVIA: Chapare, and Region Chapare, [Beni], 400 meters, Nov. 26, Dec. 18, 1948, Apr. and Glena unipennaria unipennaria (Guenee) July, 1951, 46d; Rio Suruta, Santa Cruz, 400 BRAZIL: Nuevo Teutonia, [Santa Catarina], meters, Aug. (J. Steinbach), 2 e; Santiago del various dates in Jan., May, July, Aug., 1948-1953 Estero, 1905-1906 (J. Steinbach), 1 c; Buena- (F. Plaumann), 66'; Nova Bremen, Santa Cata- vista, July-Oct., 1906 (J. Steinbach), 1 ci. rina, 250 meters, Apr., 1933, Aug., 1937 (F. Hoff- PERU: Avispas, Madre de Dios, Sept. 10-30, man), 26; Santa Catarina, 1 d; Jaragua do Sul, 1962 (L. E. Pefna), 66'; Quincemil, Cuzco, 2400 Santa Catarina, Dec., 1927 (F. Hoffmann), 1 9; ft., Aug., 1962 (L. E. Pefia), 16 ; Chanchamayo, Iguassu, Parana, Nov. 20, 1921, 16 ; Alta Parana, [Pasco], 1000 meters, 1898, Oct. and Nov., 1906 June, 1927 (F. Wucherpfennig), 1di; Castro, (O. Schuncke), 66'; La Merced, Chanchamayo, Parani, 950 meters (E. D. Jones), 26d, 1 9; [Pasco] (C. 0. Schuncke), 36; Tinguri, Carabaya, Aracatuba, Sao Paulo, 150 meters, Apr. 1, 1913 [Puno], 3400 ft., wet season, Jan., 1905 (G. (E. D. Jones), 16; Alta da Serra, Santos, [Sao Ockenden), 1 6; Chanchamayo, [Pasco], 1 9. Paulo], 2600 ft., Mar. 5, 1910 (E. D. Jones), 1 6'; ECUADOR: "Environs de Loja," [Loja], 1890, Rio [de Janeiro, Guanabara], 26', (Derg.), 1 9, 1891, 26'. 1883, (P. Germain), 16; Prov[ince] Rio de Ja- 152 1967 RINDGE: GLENA 153 neiro, 1 9, (Zikan), 1 6; "Lagune de Sacuaresma," Chavez, Esperanza, east Bolivia, 2 6; Prov. del Prov[ince] Rio de Janeiro, Aug.-Sept., 1884 (P. Sara, Santa Cruz, 450 meters, May, Oct. (J. Stein- Germain), 1 9; Petropolis, [Rio de Janeiro], 1 c; bach), 1 6, 1 9; Santa Cruz de la Sierra to San N[ovo] Friburgo, [Rio de Janeiro], 1 9; Uberaba, Jose de Chiquitos, end of July, 1904 (Steinbach), Minas Gerais, 1 c, 1 9; Prov[ince] Mato Grosso, 19 . 1886 (P. Germain), 3 ci; Porto Alegre, 1 6; Vic- PERU: Satipo, [Junin], July, Aug., 1948 (P. toria, 1 6; "Casa Br., Bras. m., G," 1 e. Paprzycki), 26'; Avispas, Madre de Dios, Sept. PARAGUAY: Independencia, [Caazapa], various 20-30, 1962 (L. E. Pefna), 1 6; Clevelandia, Nov. dates in Sept., Oct., Nov., Dec., 1950-1951, 5 c; 28, 1937, 1 9; Rio Ucayali (Stuart), 16; Santo Paso Yobay, Sept., Nov., 1951 (Schade), 26, 1 9; Domingo, [Carabaya, Puno], 6000 ft., May, 1902, Carlos Pfannl, Sept. 30, Nov. 25, 1951 (Schade), Nov., 1904 (G. Ockenden), 26; Tarapoto, May- 2 ci, 1 9; no further data, Nov. 9, 1951, 1 ei; Pa- Aug., 1886 (de Mathan), 1 6. lino cue, also spelled Patino cu6 (Monforts), 1 c, COLOMBIA: Muzo, [Santander], 400-800 meters 2 9, Feb., 2 e, 2 9; Sapucay, various dates in (Fassl), 46. July, Sept., Oct., Dec., 1902-1903 (W. Forster), VENEZUELA: Suapure, [Bolivar], Apr. 20, 1899 4 d; Sapucay, near Villa Rica (Forster), 1 i; Vil- (Klages), 1 9; Aroa, [Yaracuy], 1 9. larica, Oct., Dec., 1922-1924 (F. Schade), 3 ci, GUYANA (FORMER BRITISH GUIANA): Rock- 1 9 ; Yhu, Sept.-Dec., 1896 (Andeer), 1 6; central stone, Essequebo, 1 6'; Essequebo River (Whib- Paraguay, 1885-1886 (P. Germain), 2 e. ley), 1 9 ; confluence of Oronoque and New River, July 20 to Sept. 20, 1937 (C. A. Hudson), 1 9; Glena demissaria (Walker) Potoro, Jan., Feb., 1908 (S. M. Klages), 2 6; BRAZIL: Nuevo Teutonia, [Santa Catarina], Tumatumari, Rio Potoro, 1 6'; no further data, Mar., 1953 (F. Plaumann), le; Uberaba, Minas 1 9. Gerais, 4ci, 1 9; Prov. Mato Grosso, 1886 (P. FRENCH GUIANA: St. Jean de Maroni, 146 Germain), 2 "; Parfa (A. M. Moss), 2 d, 2 9; Sao 3 9 ; no further data (C. Bar), 36, 3 9. Paulo de Olivenca, Amazonas, Nov.-Dec. (Fassl), TRINIDAD: Arima Valley, 800-1200 ft., Feb. 3 d; Taperinha, Amazonas, 1 c; Tonantins, 10-22, 1964 (Rozen and Wygodzinsky), 36, Feb. Amazonas, July-Sept., 1880 (de Mathan), 1 6; 24, 1966 (J. G. Rozen), 1 6'; no further data, 1 6. Codajas, upper Amazon [River, Amazonas], Apr., 1907 (S. M. Klages), 1 c; Santarem, Parfa, Jan., Glena cretacea (Butler) 1878 (de Mathan), 1 c; Fura, Amazonas (H. BRAZIL: Ponta Nova, Rio Xingu, Amazones, Fassl), 1 6; Rio Madeira, 380 meters up river 16. (A. M. Moss), 16'; Calama, Rio Madeira, below PERU: Avispas, Madre de Dios, Sept. 20-30, Rio Machados, [Amazonas], Aug.-Oct., 1907 (W. 1962 (L. E. Pefia), 16'. Hoffmanns), 1 6; Humayta, Rio Madeira, [Ama- zonas], July-Sept., 1906 (W. Hoffmanns), 16; Glena mopsaria (Schaus) lower Amazon [River] (A. M. Moss), 1 9; Ama- COSTA RICA: Juan Vinas, Avangarez, type 6; zons (Leech), 1 6; "Amaz." (Bates; Felder collec- San Jose, without date and Apr. 16, 1923 (H. tion), 16'. Schmidt), 66' and 1 9; San Carlos, June, 1899 URUGUAY: Santa Cruz, Sept., 16'. (Underwood), 1 6'. PARAGUAY: Villarica, 1924 (F. Jorgensen), 36; PANAMA: V[olcan] de Chiriqui, 2000-3000 ft. Trinidad, 1 9; "Paraguay central," 1885 (P. Ger- (Champion), identified as "Boarmia plenaria main; illustrated by Oberthur, 1913, pl. CLXIX, Walk., B. C. A. Lep. Het., Godman-Salvin Coll.," fig. 1657), 16'. 1 6; Chiriqui, 1 9; Lino, 800 meters (Fassl), 1 6'; BOLIVIA: Rio Grande, Prov. Cordillera, Dec., Corazal, Canal Zone, Mar. 24, 1911 (Aug. Busck), 1903 (J. Steinbach), 1 6; San Ernesto, [La Paz], 19. 1000 meters, Aug.-Sept., 1900 (Simons), 4 6'; COLOMBIA: Makasaka, Santa Marta (V. de Buenavista, east Bolivia, 750 meters, Aug., 1906, Andreis), 1 9; Don Ano, Magdalena, 4000 ft., to Apr., 1907 (Steinbach), 1 6'; Chimate, 760 June (H. H. Smith), 1 9. meters, Sept., 1900 (Simons), 2 6'; Nutio de VENEZUELA: Merida, 1 d, Oct., 1898, 16'. 154 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

I

2

FIGS. 1, 2. Male genitalia. 1. Glena effusa, new species, paratype, Popoyan, [Colombia] (Lehmann; B.M.N.H.). 2. G. bipennaria bipennaria (Guenee), Sapucay, Paraguay, November 17, 1903 (W. Forster; B.M.N.H.). 1967 RINDGE: GLENA 155

3

FIGS. 3, 4. Male genitalia. 3. Glena dentata, new species, holotype, Balzapamba, Bolivar, [Ecuador], March-April, 1894 (de Mathan; B.M.N.H.). 4. G. grandillosa (Dognin), Callanga, Cuzco, Peru, February 16, 1953 (F. L. Woytkowski; A.M.N.H.). 156 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

FIGS. 5, 6. Male genitalia. 5. Glena sucula, new species, paratype, Colombia (B.M.N.H.). 6. G. vesana, new species, holotype, Incachaca, Cochabamba, Bolivia (J. Steinbach; U.S.N.M.). 1967 RINDGE: GLENA 157

FIGS. 7, 8. Male genitalia. 7. Gkena bisulca, new species, holotype, Hacienda La Mascota, Rio Topo, Ecuador, April 13, 1931 (W. J. Coxey; A.M.N.H.). 8. G. juga, new species, paratype, environs of Loja, Ecuador, 1888 (U.S.N.M.). 158 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

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9

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FIGS. 9, 10. Male genitalia. 9. Glena megale, new species, paratype, Santo Domingo, Peru (G. Ocken- den; B.M.N.H.). 10. G. unipennaria unipennaria (Guen6e), Independencia, Paraguay, September 22, 1951 (A.M.N.H.). 1967 RINDGE: GLENA1159

12 FIGS. 11, 12. Male genitalia. 11. Glena uncata, new species, holotype, Juan Vinas, Costa Rica, January (Schaus and Barnes; U.S.N.M.). 12. G. demissaria (Walker), Chimate, Bolivia, September, 1900 (Si- mons; B.M.N.H.). 160 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

15

FIGS. 13-15. Male genitalia. 13. Glena hima, new species, holotype, V[olcan] de Chiriqui, [Panama] (Champion; B.M.N.H.). 14. G. agria, new species, paratype, Orizaba, [Mexico], March, 1896 (W. Schaus; B.M.N.H.). 15. G. trapezia, new species, holotype, Faz[enda] dos Campos, Passa Quatro, Minas [Gerais], Brazil, November 24, 1916 (J. F. Zikan; U.S.N.M.). 1967 RINDGE: GLENA 161

16

FIGS. 16, 17. Male genitalia. 16. Glena sacca, new species, holotype, Chanchamayo, [Peru] (B.M.N.H.). 17. G. bulla, new species, paratype, Independencia, Paraguay, November 21, 1949 (P. Casada; A.M.- N.H.). 162 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

18

19

20 iv

FIGS. 18-20. Male genitalia. 18. Glena gemina, new species, holotype, Cerro Zunil, [Guatemalal (Champion; B.M.N.H.). 19. G. laticolla, new species, holotype, Puerto Elegio, Oaxaca, Mexico, Septem- ber 26, 1961 (E. C. Welling; A.M.N.H.). 20. G. brachia, new species, paratype, Chanchamayo, Peru (Schuncke; B.M.N.H.). 1967 RINDGE: GLENA 163

21

22

FIGS. 21, 22. Male genitalia. 21. Glena cretacea (Butler), Avispas, Madre de Dios, Peru, September 20- 30, 1962 (L. E. Penia; A.M.N.H.). 22. G. gampsa, new species, holotype, San Esteban, Venezuela, June, 1909 (5. M. Klages; B.M.N.H.). 164 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

23

FIGS. 23, 24. Male genitalia. 23. Glena labecula, new species, holotype, Chapare, Bolivia, April, 1951 (A.M.N.H.). 24. G. totana, new species, paratype, Arima Valley, Trinidad, February 10-22, 1964 (Rozen and Wygodzinsky; A.M.N.H.). 1967 RINDGE: GLENA 165

25

v27 , <

FIGS. 25-27. Male genitalia. 25. Glena quadrata, new species, holotype, Cayenne, French Guiana (U.S.N.M.). 26. G. mopsaria (Schaus), San Jose, Costa Rica (H. Schmidt; B.M.N.H.). 27. G. turba, new species, holotype, R[io] Laeiss, Blumenau, Santa Catarina, [Brazil], October, 1934 (F. H. Hoffman; B.M.N.H.). 166 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

FIGS. 28, 29. Male genitalia. 28. Glena tyrbe, new species, holotype, Satipo, Peru (P. Paprzycki; A.M.N.H.). 29. G. asaccula, new species, holotype, Rio Laeiss, Blumenau, Santa Catarina, [Brazil], December, 1933 (F. H. Hoffman; B.M.N.H.). 1967 RINDGE: GLENA 167

31 32

30

33 34

FIGS. 30-34. Female genitalia. 30. Glena effusa, new species, paratype, Presidio, Mexico, July, 1951 (A.M.N.H.). 31. G. bipennaria bipennaria (Guenee), Nuevo Teutonia, Brazil, September 2, 1943 (F. Plaumann; A.M.N.H.). 32. G. grandillosa (Dognin), Tucuman, Argentina (R. Schneiter; B.M.N.H.). 33. G. juga, new species, paratype, Coosnipata, Paucartambo, Cuzco, Peru, November 29, 1951 (F. L. Woytkowski; A.M.N.H.). 34. G. megale, new species, allotype, Pedregosa, Merida, Venezuela, June 8, 1898 (Briceno; B.M.N.H.). 168 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

36 37

35

38 39

FIGS. 35-39. Female genitalia. 35. Glena unipennaria unipennaria (Guenee), Patino cue, Paraguay (Monforts; B.M.N.H.). 36. G. uncata, new species, allotype, Juan Vinas, Costa Rica, June (Schaus and Barnes; U.S.N.M.). 37. G. demissaria (Walker), Para, [Brazil] (A. M. Moss; B.M.N.H.). 38. G. basalis, new species, holotype, Tuis, Costa Rica, August 31, 1908 (U.S.N.M.). 39. G. hima, new species, paratype, Chiriqui, Panama (U.S.N.M.). 1967 RINDGE: GLENA 169

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40

FIGS. 40-44. Female genitalia. 40. Glena lora, new species, holotype, Camoapa, Nicaragua, 1906 (G. M. Palmer; B.M.N.H.). 41. G. agria, new species, allotype, Vista Hermosa, Oaxaca, Mexico, October 1, 1962 (E. C. Welling; A.M.N.H.). 42. G. trapezia, new species, allotype, Alto Serra, Sao Paulo, [Brazil], December 10 (Pohl; U.S.N.M.). 43. G. gemsna, new species, allotype, Orosi, Costa Rica (Fassl; B.M.N.H.). 44. G. latscolla, new species, allotype, Cordova, Veracruz, [Mexico] (Rumeli; B.M.N.H.). 170 BULLETIN AMERICAN MUSEUM OF NATURAL HISTORY VOL. 135

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45

49 48 50

FIGS. 45-50. Female genitalia. 45. Glena brachia, new species, paratype, Sapucay, Paraguay, July 6, 1902 (W. Foster; B.M.N.H.). 46. G. totana, new species, allotype, Palmiste, Trinidad, March 7, 1917 (N. Lamont; B.M.N.H.). 47. G. quadrata, new species, allotype, Cayenne, French Guiana (U.S.N.M.). 48. G. mopsaria (Schaus), Makasaka, Santa Marta, [Colombia] (V. de Andreis; B.M.- N.H.). 49. G. turba, new species, allotype, Santa Catarina, Brazil (B.M.N.H.). 50. G. asaccula, new species, allotype, Jaragua do Sul, Santa Catarina, [Brazil], September, 1932 (F. Hoffman; B.M.- N.H.). BIBLIOGRAPHY

AMERICAN GEOGRAPHICAL SOCIETY McDUNNOUGH, J. 1945. Map of Hispanic America, 1:1,000,000. 1920. Studies in North American Cleorini Washington, Government Printing Of- (Geometridae). Dominion of Canada, fice. Dept. Agr., Ent. Branch, Tech. Bull., BOISDUVAL [J. B. A. D. DE], AND [A.] GUENEE no. 18, pp. 1-64, pls. 1-11. 1858. Histoire naturelle des insectes. Species OBERTHUR, CHARLES gen6ral des lepidopteres. Uranides, pha- 1913. etudes de lepidopterologie comparee. lenites, siculides. Paris, atlas, pp. 1-6, Rennes, fasc. 7, 677 pp., pls. 1-111, 1-4, pls. 1, 1-22, 1. CLXI-CXCVII. BROWN, F. MARTIN PROUT, LOUIS B. 1941. A gazetteer of entomological stations in 1929. On the geometrid genus Catoria Moore. Ecuador. Ann. Ent. Soc. Amer., vol. 34, Novitates Zool., vol. 35, pp. 132-142. pp. 809-851, 10 figs. RINDGE, FREDERICK H. BUTLER, ARTHUR GARDINER 1964. A revision of the genera Melanolophia, 1881. On the Lepidoptera of the Amazons, col- Pherotesia, and Melanotesia (Lepidop- lected by Dr. James W. H. Trail during tera, Geometridae). Bull. Amer. Mus. the years 1873 to 1875. Trans. Ent. Soc. Nat. Hist., vol. 126, pp. 241-434, figs. London, pp. 315-349. 1-163. COSTA LIMA, A. DA 1965. A revision of the nearctic species of the "1949" [1950]. Insetos do Brasil. Lepidopteros, genus Glena (Lepidoptera, Geometri- 2^ parte. Esc. Nac. Agronomia, vol. 6, dae). Ibid., vol. 129, pp. 265-306, figs. 420 pp., 331 figs. 1-28. DOGNIN, PAUL 1966. A revision of the moth genus Anacamp- 1902. H6t6roceres nouveaux de l'Amerique du todes (Lepidoptera, Geometridae). Ibid., sud. Ann. Soc. Ent. Belgique, vol. 46, vol. 132, pp. 175-244, figs. 1-53, pls. 22- pp. 335-350. 25. DRUCE, HERBERT SCHAUS, W. 1891-1900. Biologia Centrali-Americana. In- 1913. New from secta. Lepidoptera-Heterocera. London, species of Heterocera Costa vol. 2, pp. 1-622. Rica. XXI. Ann. Mag. Nat. Hist., ser. DYAR, HARRISON G. 8, vol. 11, pp. 342-357. 1913. Results of the Yale Peruvian expedition SELANDER, RICHARD B., AND PATRICIA VAURIE of 1911. Lepidoptera. Proc. U. S. Natl. 1962. A gazetteer to accompany the "In- Mus., vol. 45, pp. 627-649. secta" volumes of the "Biologia Cen- 1914. Report on the Lepidoptera of the trali-Americana." Amer. Mus. Novi- Smithsonian biological survey of the tates, no. 2099, pp. 1-70, figs. 1-8. Panama Canal Zone. Ibid., vol. 47, pp. TIME, INC. 139-350. 1961. Life pictorial atlas of the world. New GUENEE, A. York, Time, Inc. 1857. Histoire naturelle des insectes. Species VIETTE, P. ge'nral des lepidopteres. Paris, vol. 9, 1950. Sur quelques especes de geometrides lvi+514 pp. d6scrites par Guen6e (1857). Bull. HULST, GEO. D. Mens. Soc. Linn6enne, Lyon, yr. 19, pp. 1896. A classification of the Geometrina of 201-206. North America, with descriptions of WALKER, FRANCIS new genera and species. Trans. Amer. 1860. List of the specimens of lepidopterous Ent. Soc., vol. 23, pp. 245-386, pls. 10, insects in the collection of the British 11. Museum. Geometrites. London, pt. 21, KAYE, WILLIAM JAMES, AND NORMAN LAMONT pp. 277-498. 1927. A catalogue of the Trinidad Lepidop- 1868. Characters of some undescribed hetero- tera Heterocera (moths). Mem. Dept. cerous Lepidoptera. Jour. Linnean Soc., Agr., Trinidad and Tobago, no. 3, pp. 1- London, Zool., vol. 9, pp. 181-199. 144.

171