Great Basin Naturalist
Volume 40 Number 2 Article 10
6-30-1980
The genus Eriogonum Michx. (Polygonaceae) and Michel Gandoger
James L. Reveal University of Maryland, College Park, and Smithsonian Institution, Washington, D.C.
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Recommended Citation Reveal, James L. (1980) "The genus Eriogonum Michx. (Polygonaceae) and Michel Gandoger," Great Basin Naturalist: Vol. 40 : No. 2 , Article 10. Available at: https://scholarsarchive.byu.edu/gbn/vol40/iss2/10
This Article is brought to you for free and open access by the Western North American Naturalist Publications at BYU ScholarsArchive. It has been accepted for inclusion in Great Basin Naturalist by an authorized editor of BYU ScholarsArchive. For more information, please contact [email protected], [email protected]. THE GENUS ERIOGONUM MICHX. (POLYGONACEAE) AND MICHEL GANDOGER
James L. Reveal'
Abstract.— Michel Gandoger, a notorious "splitter," proposed several new entities in the plant genus Eriogonum (Polvgonaceae) in a 1906 paper published in Belgium. Because he used the term species at two different ranks, in violation of the International Code of Botanical Nomenclature, manv of his names are invalid. Unlike his papers published in France, this one was apparentl\- edited so that some names were validly published and some are invalid. A review of the 1906 Eriogonum paper shows that a majority of both specific and infraspecific entities proposed are valid, but some names, long in use and assinned to be valid, are, in fact, invalid. Even so, most of his names are sviionyms. Each name proposed by Gandoger is reviewed and a nomenclatural and taxonomic disposition made. Two
new combinations are made within E. hitcolum Greene, var. caninum (Greene) Reveal and var. pedunrulatum (S. Stokes) Reveal.
Michel Gandoger (1850-1926), in the ond order, in addition to terming them "spe- words of Keck (1958), was a "French abbe; ciebus secimdi ordinis," he also used the term author of 'Flora Europae' (27 vols.); volumi- siibspeciebus or subspecies to refer to entities nous writer; amasser of a huge herbarium he then proceeded to treat as binomials and now at Lyon; a 'splitter' who named thou- to designate by the expression sp. n.
sands of unacceptable species." Gandoger is The following year. Heller (1907) present- also mentioned in the International Code of ed a "compilation" of the Gandoger paper, Botanical Nomenclature (Stafleu et al. 1978) noting that he was presenting only "the new as an example of Art. 33.4. This article deals forms described in this paper." Heller himself with the problem of misplaced ranks such as had some difficulties with the names. He the use of the term species as a rank within a mentioned, after Eriogonum aspalathoides (as
species. As noted in the code, Gandoger ap- it was called bv Heller), that this name was plied the term species and used binary no- "perhaps intended as a variety of E. fasci-
menclature for two categories of taxa of con- ciilatiim, but the way the name is printed secutive rank, the higher rank being .should indicate a species." Only once does equivalent to that of species in contemporary Heller include Gandoger's terminology allud- literature, while he misapplied the same term ing to species of a lower rank, this being for to a lower rank. These latter terms are not the phrases published under E. sphoerocepha- validly published. In his 1906 paper, Gando- him, "Inter formas varias duae sequentes, ut ger used three sets of ranks, the species, spe- subspecies, praecipue distingui possunt." cies of the second order ("speciebus secundi It must be noted that Heller was merely ordinis"), and a rank which has come to be presenting Gandoger's results, and he cannot identified with the rank of variety (Heller be assumed to have validated any of the oth- 1907), although Gandoger causally refers to erwise invalid Gandoger names. this latter rank as forms ("formis," "formas Normally, Gandoger published his papers secernendas, " "formae memorabiles," "mem- in the Bulletin de la Societe Botaniqiie de orantur sequentes formae," "varieas formas," France, but for Eriogonum his choice was the "modo formae," etc.). He used the terms Bulletin de la Societe Roijale de Botanique de "varians" and "variabilis" to allude to the Belgique, and, because of this, some of the same category. As for the species of the sec- names may be validly described. Possibly
'Department of Botany, University ot Maryland. College Park. Maryland 20742, and National Museum of Natural History. Smitfisonian Institution, Wash- ington, D.C. 20560. Research supported by National Science Foundation Grant BMS75-13063. This is Scientific Article .\2714. Contribution No. 5761 of the Maryland .\gricultural Experiment Station, Department of Botany.
143 144 Great Basin Naturalist Vol. 40, No. 2 they may because of the way the editor had Eriogonum angulosum var. pauciflorum the paper set, thereby removing from the Gandoger, 187, valid, a synonym of E. ma- original manuscript some of Gandoger's ec- culatum Heller. centricities. In reviewing Gandoger's papers Eriogonum angulosum var. flabellatum in the French journal, it is clear that the ma- Gandoger, 187, valid, a synonym of E. ma- jority of his names are invalidly published. In culatum Heller. reviewing a few of his articles containing Eriogonum annuum Nutt. var. pauciflorum species from the United States, I failed to Gandoger, 187, valid, a synonym of E. an- note a single instance when a species name nuum. was validly published. This little known fact, Eriogonum hitchcockii Gandoger, 187, at least in the United States and perhaps else- valid, a synonym of E. annuum. where, means that all the Gandoger names Eriogonum juncinellum Gandoger, 187, must be carefully checked before they can be valid, a synonym of E. davidsonii Greene. accepted. This is particularly true for floristic Eriogonum salicorniodes Gandoger, 187, workers. valid, a good species restricted to clay slopes
Finally, it is sometimes difficult to know in southwestern Idaho and adjacent south- exactly where the various sections of Gando- western Oregon. It is most closely related to ger's paper start and end so that one can de- E. collinum Stokes ex Jones but most often termine if the name is validly published or, in confused with E. baileyi S. Wats. Synonyms fact, is not because of a sentence presented of this species include E. demissum S. Stokes several pages before. I have tried to follow (1936) and the var. romanum S. Stokes. the intent of Gandoger's paper, and hope I Eriogonum caespitosum Nutt. var. alijs- have interpreted each segment correctly. soides Gandoger, 188, valid, a synonym of E. The following treatment indicates which caespitosum. names are valid, which are not, and the tax- Eriogonum nevadense Gandoger, 188, onomic status of each name. The full author valid, a synonym of E. ochrocephalum S. citation of each name is given, and the num- Wats. var. ochrocephalum. ber following the name is the page on which Eriogonum elatum Dougl. ex Benth. var. it was proposed by Gandoger. limonifolium Gandoger, 188, valid, a syn- Erio^onum abertianum Torr. in Emory var. onym of £. elatum var. elatum. rubenimiim Gandoger, 185, valid, a synonym Eriogonum elatum var. erianthum Gando- of var. abertianum. ger, 188, valid, a synonym of var. elatum. Eriogonum aberiianum var. neomexicanum Eriogonum fasciculatum Benth. var. oleifo- Gandoger, 185, valid, a synonym of var. lium Gandoger, 189, valid, a synonym of E. abertianum. fasciculatum var. fasciculatum. Eriogonum arizonicum Gandoger, non Eriogonum fasciculatum Benth. var. as- Stokes ex Jones, 186, valid, a synonym of E. palathoides Gandoger, 189, valid, a synonym pharnaceoides Torr. in Emory var. pharna- of var. fasciculatum. As noted by Heller ceoides. (1907), there is some problem with this name;
Eriogonum alattim Torr. in Sitgr. var. mac- that is, there is an E. prior to the epithet. douglasii (Tandoger, 186, valid, a synonym of However, based upon the rest of the text, this var. mogollonense Stokes ex Jones. is clearly as printer's error and should be list- Eriogonum alatum var. brevifolium Gando- ed as a variety and not as a species, as was
" ger, 186, valid, a synonym of var. alatum. done by Heller in his "compilation. Eriogonum anemophiUim (as anemophijl- Eriogonum flavum Nutt. in Fras. var. folia- luni) Greene var. cusickii Gandoger, 186, tum Gandoger, 189, valid, a synonym of E. valid, a synonym of E. cusickii M. E. Jones. jamesii Benth. in DC. va.r. flavescens S. Wats. Eriogonum angulosum Benth. var. rectipes Eriogoni4m flavum var. linguifolium Gan- Gandoger, 186, valid, a synonym of E. ma- doger, 189, valid, a synonym of var. flavum. cukitum Heller. Eriogonum leucocladum Gandoger, 189, Eriogonum angulosum var. patens Gando- valid, a synonym of E. baileyi S. Wats. var. ger, 187, valid, a synonvm of E. maculatum divaricatum (Gandoger) Reveal. Heller. ErioEonum heracleoides Nutt. var. micran- June 1980 Reveal: Eriogonum Nomenclature 145
thum Gandoger, 189, valid, a synonym of £. Eriogonum myrianthum Gandoger, 191, in-
herocleoides var. angustifolium (Niitt.) Torr. valid, a synonym of E. cffusum Nutt. var. cf- & Gray. fusum. Eriogonum hcracleoides var. viride Gando- Eriogonum sputhulare Gandoger, 191, in- ger, 190, valid, a synonym of £. umhellatum valid, a synonym of E. microthecum Nutt. Torr. var. nevadense Gandoger. var. laxiflorum Hook. The name was used by Eriogonum hcracleoides var. multiccps Stokes (1936) at the variatal rank within E. Gandoger, 190, valid, a synonym of var. hcra- microthecum, where she validated the name cleoides. as E. microthecum var. spathulure S. Stokes. Eriogonum hcracleoides var. utahensis Eriogonum intricatum Gandoger, 191, in- Gandoger, 190, valid, a synonvm of var. hcra- valid, a synonym of E. microthecum Nutt. cleoides. var. laxiflorum Hook. Eriogonum hcracleoides var. rydbergii Gan- Eriogonum Iwlichrysoides Gandoger, 192, doger, 190, valid, a synonym of var. hcra- invalid, a synonym of E. cffusum Nutt. var. cleoides. rosmarinioides Benth. in DC. The name was Eriogonum jamesii Benth. in DC. var. sim- validly published by Rydberg (1931) and the plex Gandoger, 190, valid, a good variety of- citation of the name should be E. helichry- ten included within var. jamesii; this phase of soides Rydb., Brittonia 1: 87. 1931, without
the species is restricted to southwestern Kan- reference to Gandoger. Rydberg was in- sas. correct that Gandoger had proposed a varie- Eriogonum jamesii var. neomexicanum ty, so the combination attributed to Gando- Gandoger, 190, valid, a synonym of var. ger by Rydberg in synonymy, E. nucrothecum jamesii. var. helichrysoides, must be attributed to Eriogonum longifolium Nutt. var. long- Rydberg as well. idcns Gandoger, 190, valid, a synonym of E. Eriogonum sarothriforme Gandoger, 192, longifolium var. gnaphalifolium Gandoger. invalid. The type of this name, collected by Eriogonum longifolium var. gnaphalifo- Osterhout at Glenwood Springs, Garfield Co., lium Gandoger, 190, valid, that phase of the Colorado, may represent a distinct taxon. It species restricted to Florida; often called E. belongs to the E. brevicaule Nutt. complex floridanum Small. and is seemingly a part of the polymorphic Eriogonum longifolium var. floridanum (as species E. lonchopln/llum Torr. & Gray, a floridana) Gandoger, 190, valid, a synonym taxon that morphologically bridges the E. of £. longifolium var. gnaphalifolium Gando- brevicaule complex, a group of herbaceous ger. perennials, with those species typified bv E. Eriogonum longifolium var. lindheimeri corymbosum Benth. in DC, a series of shrubs Gandoger, 190, valid, the western phase of or subshnibs. The Garfield Co. plants tend to the species now best considered a synonvm of be more slender than typical E. lonchophyl- var. longifolium. lum of southern Colorado and adjacent Eriogonum longifolium var. caput-fclis northern New Mexico. These plants also tend Gandoger, 190, valid, a synonym of var. lon- to resemble some of the more robust, but yel- gifolium. low-flowered, forms of E. brevicaule found in Under the heading, Eriogonum micro- Rio Blanco Co., Colorado. Although well thccum Nutt., Gandoger states "inter quas se- known to me, I am still uncertain what to do quentes altem pro subspeciebus habueris" with the Glenwood Springs plants.
and refers to the following six entities all Eriogonum niveum Dougl. ex Benth. var. published with binary names. suksdorfii Gandoger, 192, valid, a synonym of Eriogonum macdougalii Gandoger, 191, in- E. niveum. valid, a synonym of E. microthecum Nutt. Eriogonum niveuni var. candelabrum Gan- var. foliosum (Torr. & Gray) Reveal. The doger, 192, valid, a svnonvm of E. niveum. name was used by Stokes (1936) at the varia- Eriogonum ochrolcucum Small var. macro- tal rank within E. microthecum, where she podum Gandoger, 192, valid, the basionym validated the name as E. microthecum var. for E. ovalifolium Nutt. var. macropodum macdougalii S. Stokes. (Gandoger) Reveal. 146 Great Basin Naturalist Vol. 40. No. 2
Eriogonum ochroleucum var. decahans Cokeville, Uinta Co., Wyoming, upon which Gandoger, 192, valid a synonym of E. ocali- he also based the name E. dicJiroanthum. An folium Nutt. var. macropodum (Gandoger) examination of this collection b\ me in 1966, Reveal. and then by my brother, Jon A. Reveal, in Under the heading of Eriogonum ovalifo- 1973, was inconclusive in that it could not be lium Nutt., Gandoger proposed a series of fully determined if the flowers of var. multi- names in two ranks, all of which he refers to scapum were truly yellow. Gandoger divided by "formas quarum non paucas saltern pro the collection and moimted each on a sepa- speciebus secimdi ordinis haberi possimt," rate sheet, with E. dichroanthum having
which I believe he wished to apply only to "flores fructusque flavissimi." and the flowers those names he indicated by the designation of var. midtiscapum, to him. were "'ochro- oi sp. n. leuci." The same collection at KSC is strictly Eriogonum flavissitnum Gandoger, 193, in- yellow flowered, but the specimen at RM is a valid, a svTionym of E. ovalifolium Nutt. var. mixture of a bright-yellowed specimens Davis. matching E. dichroanthum and a whitish or. anserinum (Greene) R. J. The name was used by Stokes (1936) as a subspecies of at best, pale yellow-flowered specimen that E. ovalifoliwn, where the name was validated matches var. multiscapuryi. I now believe that as E. ovalifolium Nutt. ssp. anserinum S. the yellow-flowered, early spring flowering Stokes. phase of the species should be called var. Eriogonum cusickii Gandoger, non M. E. nevadense and the type of var. lyiultiscapum Jones (1903), 193, invalid, a synonym of E. assigned to var. ovalifolium. strictum var. proliferum (Torr. & Gray) Re- Eriogonum ovalifolium Nutt. var. cijclo- veal. The name was used at the variatal rank phijllum Gandoger, 194, valid, a synonym of by Stokes (1936) within E. strictum, where var. macropodum (Gandoger) Reveal. A reex- the name was validated as E. strictum Benth. amination of the types of var. cyclophyllum var. cusickii S. Stokes. and var. cerastoides (see below), as part of Eriogonum cusickii Gandoger var. califor- this study, clearly shows that these names nicum Gandoger, 193, invalid, a synonvm of must be referred to what I (Reveal 1968) had E. strictum var. proliferum (Torr. 6c Grav) earlier called var. macropodum. making a Reveal. new combination for this name by transfer-
Eriogonum ovalifolium Nutt. var. neva- ring it from E. ochroleucum Small to E. ovali- dense Gandoger, 193, valid, a good varietv of folium. Previously, these two names have the species, this being the yellow-flowered, been referred to var. ovalifolium (Hitchcock
early flowering expression which I have 1964). The var. cijclophyUum is close to var. called var. multiscapum Gandoger (see be- ovalifolium and may represent one of the low). many intermediate expressions between the
Eriogonum ovalifolium var. deltoideum two varieties. I retain the usage of var. Gandoger, valid, 193, a synonym of £. ovali- macropodum. folium var. nevadense Gandoger. Eriogonum ovalifolium Nutt. var. ceras- Eriogonum dichroanthum Gandoger, 193, toides Gandoger, 194, valid, a synonym of invalid, a synonym of E. ovalifolium Nutt. var. macropodum (Gandoger) Reveal (see the var. nevadense Gandoger (see discussion un- discussion above). der var. multiscapum below). Eriogonum ruhidum Gandoger. 194, in- Eriogonum ovalifolium Nutt. var. utahense valid, a synonym of E. ovalifolium Nutt. var. Gandoger, 194, valid, a synonym of var. ova- depressimi Blankinship. lifolium. Eriogonum ruhidum var. frigidum Gando- Eriogonum ovalifolium Nutt. var. multi- ger, 194, invalid, a synonym of E. ovalifolium scapum Gandoger, 194, valid, a synonym of Nutt. var. depressum Blankinship.
var. ovalifolium. For several years I have Eriogonum roseiflorum Gandoger. 194, in- misapplied this name to the yellow-flowered valid, a svnonvm of E. ovalifolium Nutt. var. phase of the species (Reveal & Munz 196S, ovalifolium. Reveal 1973, 1976). Gandoger based this Eriogonum piperi Greene var. ochrocepha- name on plants gathered by Nelson (4658) at lum Gandoger, 195, valid, a synonym of E. June 1980 Reveal: Eriogonum Nomenclature 14'
flaviim \utt. in Fras. var. piperi (Greene) M. mioides S. Stokes without reference to Gan- E. Jones. doger. Eriogonum piperi var. longifloruni Gando- Eriogonuin subalpinum Greene var. arach- ger, 195, valid, a synonym of E. flavum Xutt. noideum Gandoger, 197. valid, a synonym of in Fras. var. piperi (Greene) M. E. Jones. E. umbellatum Torr. var. dichrocephalum Under Eriogonum pohjanthum Benth., now Gandoger. better known as E. umbellatum Torr. var. Eriogonum subalpinum var. vulcanicum pohjanthum (Benth. in DC.) M. E. Jones, Gandoger, 197, valid, a synonymy of £. um- Gandoger introduces two validly described bellatuin Torr. var. majus Hook. species simply stating that "species duae se- Eriogonum subalpinum var. stenophyllum quentes huic sunt affines." Gandoger, 197, valid, a synonym of E. um- Eriogonum marginale Gandoger, 195, bellatum Torr. var. majus Hook. valid, a synonym of E. umbellatum Torr. var. Eriogonum subalpinum var. subnivale aureum (Gandoger) Reveal. Gandoger, 197, valid, a synonym of E. um- Eriogonum glaherrimum Gandoger, 195. bellatum Torr. var. majus Hook. valid, the basionym of E. umbellatum Torr. Eriogonum tenellum Torr. var. grandi- var. glaberrimum (Gandoger) Reveal. florum Gandoger, 197, valid, a synonym of E. Eriogonum glaberrimum var. aureum Gan- microthecum Nutt. var. laxiflorum Hook. doger, 195, valid, the basionym of E. um- Eriogonum tenelhw} var. sessiiflorum Gan- bellatum Torr. var. aureum (Gandoger) Re- doger, 198, valid, a synonym of E. micro- veal. thecum Xutt. var. laxiflorum Hook.
Eriogonum pohjcladon Benth. var. mexi- Eriogonum tenellum var. erianthum Gan- canum Gandoger, 196, valid, a synonym of E. doger, 198, valid, a synonym of E. micro- pohjcladon. thecum Xutt. var. ambiguum (M. E. Jones) Reveal in Mimz. Eriogonum polycladon Benth. var. crispum Gandoger, 196, valid, a synonym of E. poly- Eriogonum thurberi Torr. var. parishii Gan- cladon. doger. 198, valid, a svnonvm of £. thurberi. Eriogonum thurberi var. acutangulum Gan- Eriogonum racemosum Xutt. var. sagitta- doger, 198. valid, a synonvm of E. macula- tum Gandoger, 196, valid, a synonym of E. racemosum. turn Heller. Eriogonum racemosum Xutt. var. cordi- Eriogonum tlnjmoides Benth. in DC. var. folium Gandoger, 196, valid, a svnonvm of E. pallens Gandoger, 198, valid, a synonym of racemosum. £. thymoides. Eriogonum reniforme Torr. & Frem. var. Eriogonum umbellatum Torr. var. cran- asarifolium Gandoger, 196, valid, a svnonvm dallii Gandoger, 198, valid, a synonym of var. of E. pusillum Torr. 6f Gray. umbellatum. Eriogonum praebens Gandoger, 196, valid, Eriogonum umbellatum var. chrysanthum a s\nonym of E. bailcyi S. Wats. var. diva- Gandoger, 198, valid, a synonym of E. um- ricatum (Gandoger) Reveal. bellatum var. stellatum (Benth.) M. E. Jones. Eriogonum praebens var. diiaricatum Gan- Eriogonum umbellatum var. nevadense doger, 196, valid, the basionym of E. baileyi Gandoger, 198, valid, a good variety referr- S. Wats. var. divaricatum (Gandoger) Reveal. ing to that phase of the species found in the Under the heading Eriogonum sphaero- Sierra Xevada of California northward into cephalum Dougl. ex Benth.. Gandoger states Oregon and eastward into uestern Xevada "inter formas varias duae sequentes. ut sub- that has been routinelv called var. umbella- species, praecipue distingui possunt." tum (Reveal & Munz 1968; see Howell 1976). Eriogonum cupreum Gandoger, 196. in- Eriogonum umbellatum var. cladophorum valid, a synonym of E. umbellatum Torr. var. Gandoger, 198, valid, a synonym of var. um- umbellatum. bellatum. Eriogonum halimioides Gandoger, 197, in- Eriogonum umbellatum var. dichro- valid. This name was validated by Stokes cephalum Gandoger, 199, valid, a good varie- (1936) as E. sphaerocephalum var. hali- ty applied to that phase previously called E. 148 Great Basin Naturalist Vol. 40, No. 2
umbeUatum var. aridum (Greene) C. L. based on E. pedunculatum S. Stokes, Leafl. Hitchc. (Hitchcock 1964). W. Bot. 2: 48. 1937. Eriogonwn umhellatum var. californicuni Eriogonum vimineum var. oregonense Gan- Gandoger, 199, valid, a synonym of E. um- doger, 199, valid, a synonym of E. vimineum. beUatum var. nevadense Gandoger. Eriogonum restioioidcs Gandoger, 199, Eriogonum vimineiim Dougl. ex Benth. valid, a synonym of E. baileyi S. Wats. var.
var. rigescens Gandoger, 199, valid, a synon- bailey i. ym of E. vimeneum. Dougl. ex Benth. Eriogonum vimineum Literature Cited var. califomicum Gandoger, 199, valid, a syn- onym of E. luteohim Greene var. caninum Gandoger, M. 1906. Le genre Eriogonum (Poly- Bull. Soc. Belgique (Greene) Reveal, comb, nov., based on E. vi- gonaceae). Roy. Bot. 42: 18.3-200. mineum var. caninum Greene, Fl. Francisc. Heller, A. A. 1907. Compilations. Miihlenbergia 3: 150. 1891. The type of var. califomicum is a3-96. somewhat intermediate between var. luteo- Hitchcock, C. L. 1964. Eriogonum. Univ. Wash. Publ. lum and var. caninum, being closer to the Biol. 17(2): 104-138. Howell, T. 1970. Marin flora. 2d ed. Berkeley: Univ. latter than the former. I have long recog- J. California Press. 366 pp. nized the caninum phase (Reveal & Munz 1976. Eriogonum notes VII. Mentzelia 1: 17-22. E. T. 1968) as distinct from vimineum, but J. Keck, D. D. 1959. "Abbreviations of authors' names." Howell, who has considered the expression Pages 1551-1576 in P. A. Munz, & D. D. Keck, only as a variant, has pointed out in our con- eds. A California flora. Berkeley: Univ. California Press. versation that he had observed a large series L. in Reveal, J. 1968. Some nomenclatural changes of intermediate populations, as had I, which Eriogonum (Polygonaceae). Taxon 17: 531-533. held the Mt. Tamalpais plant, var. caninum, 1973. Eriogonum (Polvgonaceae) of Utah. Phvto- well within the boundaries of what he called logia 25: 169-217. 1976. Eriogonum (Polvgonaceae) of Arizona and E. vimineum. This latter expression, however, New Mexico. Phytologia 34: 409-484. proved to be E. luteohim rather than E. vimi- L., P. A. 1968. "Eriogonum." Reveal, J. and Munz. neum, and I am now following Howell's Pages 33-72 in P. A. Munz, ed. Supplement to A (1970) taxonomic disposition of this local en- California flora. Berkeley: Univ. California Press. demic. In addition to this variant of E. luteo- Rydberg, p. a. 19.31. Taxonomic notes on the flora of the prairies and plains of central North .\merica. lum, the Sierra Nevada plant I recognized as Brittonia 1: 79-104. a distinct species previously (Reveal 1970) Stafleu, F. a., et al. 1978. International code of bot- should be included within this species as well. anical nomenclatme. Regnum Veg. 97: 1-457. Thus, I propose E. luteohim var. peduncula- Stokes, S. G. 1936. The genus Eriogonum, a preliminary studv based on geographic distribution. San Fran- tum. (S. Stokes) Reveal, stat. & comb, nov., cisco: B. Nebiett. 124 J. pp.