Arachnida: Opiliones)
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Topic Paper Chilterns Beechwoods
. O O o . 0 O . 0 . O Shoping growth in Docorum Appendices for Topic Paper for the Chilterns Beechwoods SAC A summary/overview of available evidence BOROUGH Dacorum Local Plan (2020-2038) Emerging Strategy for Growth COUNCIL November 2020 Appendices Natural England reports 5 Chilterns Beechwoods Special Area of Conservation 6 Appendix 1: Citation for Chilterns Beechwoods Special Area of Conservation (SAC) 7 Appendix 2: Chilterns Beechwoods SAC Features Matrix 9 Appendix 3: European Site Conservation Objectives for Chilterns Beechwoods Special Area of Conservation Site Code: UK0012724 11 Appendix 4: Site Improvement Plan for Chilterns Beechwoods SAC, 2015 13 Ashridge Commons and Woods SSSI 27 Appendix 5: Ashridge Commons and Woods SSSI citation 28 Appendix 6: Condition summary from Natural England’s website for Ashridge Commons and Woods SSSI 31 Appendix 7: Condition Assessment from Natural England’s website for Ashridge Commons and Woods SSSI 33 Appendix 8: Operations likely to damage the special interest features at Ashridge Commons and Woods, SSSI, Hertfordshire/Buckinghamshire 38 Appendix 9: Views About Management: A statement of English Nature’s views about the management of Ashridge Commons and Woods Site of Special Scientific Interest (SSSI), 2003 40 Tring Woodlands SSSI 44 Appendix 10: Tring Woodlands SSSI citation 45 Appendix 11: Condition summary from Natural England’s website for Tring Woodlands SSSI 48 Appendix 12: Condition Assessment from Natural England’s website for Tring Woodlands SSSI 51 Appendix 13: Operations likely to damage the special interest features at Tring Woodlands SSSI 53 Appendix 14: Views About Management: A statement of English Nature’s views about the management of Tring Woodlands Site of Special Scientific Interest (SSSI), 2003. -
Towards a DNA Barcode Referencedatabasefor Spiders And
RESEARCH ARTICLE Towards a DNA Barcode Reference Database for Spiders and Harvestmen of Germany Jonas J. Astrin1*, Hubert Höfer2*, Jörg Spelda3*, Joachim Holstein4*, Steffen Bayer2, Lars Hendrich3, Bernhard A. Huber1, Karl-Hinrich Kielhorn5, Hans-Joachim Krammer1, Martin Lemke6, Juan Carlos Monje4, Jérôme Morinière3, Björn Rulik1, Malte Petersen1, Hannah Janssen1, Christoph Muster7 1 ZFMK: Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany, 2 SMNK: Staatliches Museum für Naturkunde Karlsruhe, Karlsruhe, Germany, 3 ZSM: Zoologische Staatssammlung München, München, Germany, 4 SMNS: Staatliches Museum für Naturkunde Stuttgart, Stuttgart,Germany, 5 Karl- a11111 Hinrich Kielhorn, Berlin, Germany, 6 Martin Lemke, Lübeck, Germany, 7 Zoologisches Institut und Museum, Universität Greifswald, Greifswald, Germany * [email protected] (JJA); [email protected] (HH); [email protected] (JS); joachim. [email protected] (JH) OPEN ACCESS Abstract Citation: Astrin JJ, Höfer H, Spelda J, Holstein J, As part of the German Barcode of Life campaign, over 3500 arachnid specimens have been Bayer S, Hendrich L, et al. (2016) Towards a DNA collected and analyzed: ca. 3300 Araneae and 200 Opiliones, belonging to almost 600 spe- Barcode Reference Database for Spiders and Harvestmen of Germany. PLoS ONE 11(9): cies (median: 4 individuals/species). This covers about 60% of the spider fauna and more e0162624. doi:10.1371/journal.pone.0162624 than 70% of the harvestmen fauna recorded for Germany. The overwhelming majority of Editor: Matja Kuntner, Scientific Research Centre of species could be readily identified through DNA barcoding: median distances between clos- the Slovenian Academy of Sciences and Art, est species lay around 9% in spiders and 13% in harvestmen, while in 95% of the cases, SLOVENIA intraspecific distances were below 2.5% and 8% respectively, with intraspecific medians at Received: June 3, 2016 0.3% and 0.2%. -
Comparative Functional Morphology of Attachment Devices in Arachnida
Comparative functional morphology of attachment devices in Arachnida Vergleichende Funktionsmorphologie der Haftstrukturen bei Spinnentieren (Arthropoda: Arachnida) DISSERTATION zur Erlangung des akademischen Grades doctor rerum naturalium (Dr. rer. nat.) an der Mathematisch-Naturwissenschaftlichen Fakultät der Christian-Albrechts-Universität zu Kiel vorgelegt von Jonas Otto Wolff geboren am 20. September 1986 in Bergen auf Rügen Kiel, den 2. Juni 2015 Erster Gutachter: Prof. Stanislav N. Gorb _ Zweiter Gutachter: Dr. Dirk Brandis _ Tag der mündlichen Prüfung: 17. Juli 2015 _ Zum Druck genehmigt: 17. Juli 2015 _ gez. Prof. Dr. Wolfgang J. Duschl, Dekan Acknowledgements I owe Prof. Stanislav Gorb a great debt of gratitude. He taught me all skills to get a researcher and gave me all freedom to follow my ideas. I am very thankful for the opportunity to work in an active, fruitful and friendly research environment, with an interdisciplinary team and excellent laboratory equipment. I like to express my gratitude to Esther Appel, Joachim Oesert and Dr. Jan Michels for their kind and enthusiastic support on microscopy techniques. I thank Dr. Thomas Kleinteich and Dr. Jana Willkommen for their guidance on the µCt. For the fruitful discussions and numerous information on physical questions I like to thank Dr. Lars Heepe. I thank Dr. Clemens Schaber for his collaboration and great ideas on how to measure the adhesive forces of the tiny glue droplets of harvestmen. I thank Angela Veenendaal and Bettina Sattler for their kind help on administration issues. Especially I thank my students Ingo Grawe, Fabienne Frost, Marina Wirth and André Karstedt for their commitment and input of ideas. -
Opiliones, Palpatores, Caddoidea)
Shear, W. A. 1975 . The opilionid family Caddidae in North America, with notes on species from othe r regions (Opiliones, Palpatores, Caddoidea) . J. Arachnol . 2:65-88 . THE OPILIONID FAMILY CADDIDAE IN NORTH AMERICA, WITH NOTES ON SPECIES FROM OTHER REGION S (OPILIONES, PALPATORES, CADDOIDEA ) William A . Shear Biology Departmen t Hampden-Sydney, College Hampden-Sydney, Virginia 23943 ABSTRACT Species belonging to the opilionid genera Caddo, Acropsopilio, Austropsopilio and Cadella are herein considered to constitute the family Caddidae . The subfamily Caddinae contains the genu s Caddo ; the other genera are placed in the subfamily Acropsopilioninae. It is suggested that the palpatorid Opiliones be grouped in three superfamilies : Caddoidea (including the family Caddidae) , Phalangioidea (including the families Phalangiidae, Liobunidae, Neopilionidae and Sclerosomatidae ) and Troguloidea (including the families Trogulidae, Nemostomatidae, Ischyropsalidae an d Sabaconidae). North American members of the Caddidae are discussed in detail, and a new species , Caddo pepperella, is described . The North American caddids appear to be mostly parthenogenetic, an d C. pepperella is very likely a neotenic isolate of C. agilis. Illustrations and taxonomic notes ar e provided for the majority of the exotic species of the family . INTRODUCTION Considerable confusion has surrounded the taxonomy of the order Opiliones in North America, since the early work of the prolific Nathan Banks, who described many of ou r species in the last decade of the 1800's and the first few years of this century. For many species, no additional descriptive material has been published following the original de- scriptions, most of which were brief and concentrated on such characters as color and body proportions . -
Harvest-Spiders 515
PROVISIONAL ATLAS OF THE REF HARVEST-SPIDERS 515. 41.3 (ARACHNIDA:OPILIONES) OF THE BRITISH ISLES J H P SANKEY art å • r yz( I is -..a .e_I • UI II I AL _ A L _ • cta • • .. az . • 4fe a stir- • BIOLOGICAL RECORDS CENTRE Natural Environment Research Council Printed in Great Britain by Henry Ling Ltd at the Dorset Press, Dorchester, Dorset ONERC Copyright 1988 Published in 1988 by Institute of Terrestrial Ecålogy Merlewood Research Station GRANGE-OVER-SANDS Cumbria LA1/ 6JU ISBN 1 870393 10 4 The institute of Terrestrial Ecology (ITE) was established in 1973, from the former Nature Conservancy's research stations and staff, joined later by the Institute of Tree Biology and the Culture Centre of Algae and Protozoa. ITO contribbtes to, and draws upon, the collective knowledge of the 14 sister institutes which make up the Natural Environment Research Council, spanning all the environmental sciences. The Institute studies the factors determining the structure, composition and processes of land and freshwater systems, and of individual plant and animal species. It is developing a sounder scientific basis for predicting and modelling environmental trends arising from natural or man-made change. The results of this research are available to those responsible for the protection, management and wise use of our natural resources. One quarter of ITE's work is research commissioned by customers, such as the Department of Environment, the European Economic Community, the Nature Conservancy Council and the Overseas Development Administration. The remainder is fundamental research supported by NERC. ITE's expertise is widely used by international organizations In overseas projects and programmes of research. -
De Hooiwagens 1St Revision14
Table of Contents INTRODUCTION ............................................................................................................................................................ 2 CHARACTERISTICS OF HARVESTMEN ............................................................................................................................ 2 GROUPS SIMILAR TO HARVESTMEN ............................................................................................................................. 3 PREVIOUS PUBLICATIONS ............................................................................................................................................. 3 BIOLOGY ......................................................................................................................................................................... 3 LIFE CYCLE ..................................................................................................................................................................... 3 MATING AND EGG-LAYING ........................................................................................................................................... 4 FOOD ............................................................................................................................................................................. 4 DEFENCE ........................................................................................................................................................................ 4 PHORESY, -
Biochemical Divergence Between Cavernicolous and Marine
The position of crustaceans within Arthropoda - Evidence from nine molecular loci and morphology GONZALO GIRIBET', STEFAN RICHTER2, GREGORY D. EDGECOMBE3 & WARD C. WHEELER4 Department of Organismic and Evolutionary- Biology, Museum of Comparative Zoology; Harvard University, Cambridge, Massachusetts, U.S.A. ' Friedrich-Schiller-UniversitdtJena, Instituifiir Spezielte Zoologie und Evolutionsbiologie, Jena, Germany 3Australian Museum, Sydney, NSW, Australia Division of Invertebrate Zoology, American Museum of Natural History, New York, U.S.A. ABSTRACT The monophyly of Crustacea, relationships of crustaceans to other arthropods, and internal phylogeny of Crustacea are appraised via parsimony analysis in a total evidence frame work. Data include sequences from three nuclear ribosomal genes, four nuclear coding genes, and two mitochondrial genes, together with 352 characters from external morphol ogy, internal anatomy, development, and mitochondrial gene order. Subjecting the com bined data set to 20 different parameter sets for variable gap and transversion costs, crusta ceans group with hexapods in Tetraconata across nearly all explored parameter space, and are members of a monophyletic Mandibulata across much of the parameter space. Crustacea is non-monophyletic at low indel costs, but monophyly is favored at higher indel costs, at which morphology exerts a greater influence. The most stable higher-level crusta cean groupings are Malacostraca, Branchiopoda, Branchiura + Pentastomida, and an ostracod-cirripede group. For combined data, the Thoracopoda and Maxillopoda concepts are unsupported, and Entomostraca is only retrieved under parameter sets of low congruence. Most of the current disagreement over deep divisions in Arthropoda (e.g., Mandibulata versus Paradoxopoda or Cormogonida versus Chelicerata) can be viewed as uncertainty regarding the position of the root in the arthropod cladogram rather than as fundamental topological disagreement as supported in earlier studies (e.g., Schizoramia versus Mandibulata or Atelocerata versus Tetraconata). -
A Multilocus Phylogeny of Podoctidae
Molecular Phylogenetics and Evolution 106 (2017) 164–173 Contents lists available at ScienceDirect Molecular Phylogenetics and Evolution journal homepage: www.elsevier.com/locate/ympev A multilocus phylogeny of Podoctidae (Arachnida, Opiliones, Laniatores) and parametric shape analysis reveal the disutility of subfamilial nomenclature in armored harvestman systematics ⇑ Prashant P. Sharma a, , Marc A. Santiago b, Ricardo Kriebel c, Savana M. Lipps a, Perry A.C. Buenavente d, Arvin C. Diesmos d, Milan Janda e,f, Sarah L. Boyer g, Ronald M. Clouse b, Ward C. Wheeler b a Department of Zoology, University of Wisconsin-Madison, 430 Lincoln Drive, Madison, WI 53706, USA b Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79th Street, New York, NY, 10024, USA c Department of Botany, University of Wisconsin-Madison, 430 Lincoln Drive, Madison, WI 53706, USA d Zoology Division, National Museum of the Philippines, Padre Burgos Avenue, Ermita 1000, Manila, Philippines e Laboratorio Nacional de Análisis y Síntesis Ecológica, ENES, UNAM, Antigua Carretera a Pátzcuaro, 8701 Morelia, Mexico f Biology Centre, Czech Academy of Sciences, Branisovska 31, 370 05 Ceske Budejovice, Czech Republic g Biology Department, Macalester College, 1600 Grand Avenue, St. Paul, MN 55105, USA article info abstract Article history: The taxonomy and systematics of the armored harvestmen (suborder Laniatores) are based on various Received 9 August 2016 sets of morphological characters pertaining to shape, armature, pedipalpal setation, and the number of Accepted 20 September 2016 articles of the walking leg tarsi. Few studies have tested the validity of these historical character systems Available online 21 September 2016 in a comprehensive way, with reference to an independent data class, i.e., molecular sequence data. -
Behavioral Roles of the Sexually Dimorphic Structures in the Male Harvestman, Phalangium Opilio (Opiliones, Phalangiidae)
1763 Behavioral roles of the sexually dimorphic structures in the male harvestman, Phalangium opilio (Opiliones, Phalangiidae) Rodrigo H. Willemart, Jean-Pierre Farine, Alfredo V. Peretti, and Pedro Gnaspini Abstract: In various animal species, male sexual dimorphic characters may be used during intrasexual contests as orna- ments to attract females, or to hold them before, during, or after copulation. In the well-known harvestman, Phalangium opilio L., 1758, the behavioral functions of these male sexually dimorphic structures have never been studied in detail. Therefore, in addition to a morphometric study, 21 male contests and 43 sexual interactions were analyzed. Our observa- tions revealed that during contests, the male cheliceral horns form a surface by which the contestants use to push each other face-to-face while rapidly tapping their long pedipalps against the pedipalps of the opponent, occasionally twisting the opponent’s pedipalp. Scanning electron micrographs revealed contact mechanoreceptors on the pedipalp that would de- tect the intensity–frequency of contact with the contender’s pedipalp. Larger males won almost all contests, whereas the loser rapidly fled. During sexual interactions, the longer pedipalps of the male held legs IV of the female, whereas males with shorter pedipalps held the female by legs III. No contact with the male pedipalps and chelicerae by the females was visible before, during, or after copulation. Soon after copulating, males typically bent over the female, positioning their cheliceral horns against the females’s dorsum. Consequently, our data show that the cheliceral horns and the longer pedi- palps of the male seem to play an important role, during both intersexual and intrasexual encountering. -
The Phylogeny of Fossil Whip Spiders Russell J
Garwood et al. BMC Evolutionary Biology (2017) 17:105 DOI 10.1186/s12862-017-0931-1 RESEARCH ARTICLE Open Access The phylogeny of fossil whip spiders Russell J. Garwood1,2*, Jason A. Dunlop3, Brian J. Knecht4 and Thomas A. Hegna4 Abstract Background: Arachnids are a highly successful group of land-dwelling arthropods. They are major contributors to modern terrestrial ecosystems, and have a deep evolutionary history. Whip spiders (Arachnida, Amblypygi), are one of the smaller arachnid orders with ca. 190 living species. Here we restudy one of the oldest fossil representatives of the group, Graeophonus anglicus Pocock, 1911 from the Late Carboniferous (Duckmantian, ca. 315 Ma) British Middle Coal Measures of the West Midlands, UK. Using X-ray microtomography, our principal aim was to resolve details of the limbs and mouthparts which would allow us to test whether this fossil belongs in the extant, relict family Paracharontidae; represented today by a single, blind species Paracharon caecus Hansen, 1921. Results: Tomography reveals several novel and significant character states for G. anglicus; most notably in the chelicerae, pedipalps and walking legs. These allowed it to be scored into a phylogenetic analysis together with the recently described Paracharonopsis cambayensis Engel & Grimaldi, 2014 from the Eocene (ca. 52 Ma) Cambay amber, and Kronocharon prendinii Engel & Grimaldi, 2014 from Cretaceous (ca. 99 Ma) Burmese amber. We recovered relationships of the form ((Graeophonus (Paracharonopsis + Paracharon)) + (Charinus (Stygophrynus (Kronocharon (Charon (Musicodamon + Paraphrynus)))))). This tree largely reflects Peter Weygoldt’s 1996 classification with its basic split into Paleoamblypygi and Euamblypygi lineages; we were able to score several of his characters for the first time in fossils. -
A Stable Phylogenomic Classification of Travunioidea (Arachnida, Opiliones, Laniatores) Based on Sequence Capture of Ultraconserved Elements
A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements The Harvard community has made this article openly available. Please share how this access benefits you. Your story matters Citation Derkarabetian, Shahan, James Starrett, Nobuo Tsurusaki, Darrell Ubick, Stephanie Castillo, and Marshal Hedin. 2018. “A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements.” ZooKeys (760): 1-36. doi:10.3897/zookeys.760.24937. http://dx.doi.org/10.3897/zookeys.760.24937. Published Version doi:10.3897/zookeys.760.24937 Citable link http://nrs.harvard.edu/urn-3:HUL.InstRepos:37298544 Terms of Use This article was downloaded from Harvard University’s DASH repository, and is made available under the terms and conditions applicable to Other Posted Material, as set forth at http:// nrs.harvard.edu/urn-3:HUL.InstRepos:dash.current.terms-of- use#LAA A peer-reviewed open-access journal ZooKeys 760: 1–36 (2018) A stable phylogenomic classification of Travunioidea... 1 doi: 10.3897/zookeys.760.24937 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research A stable phylogenomic classification of Travunioidea (Arachnida, Opiliones, Laniatores) based on sequence capture of ultraconserved elements Shahan Derkarabetian1,2,7 , James Starrett3, Nobuo Tsurusaki4, Darrell Ubick5, Stephanie Castillo6, Marshal Hedin1 1 Department of Biology, San Diego State University, San -
Carinostoma Elegans New to the Slovakian Harvestmen Fauna (Opiliones, Dyspnoi, Nemastomatidae)
Arachnologische Mitteilungen 48: 16-23 Karlsruhe, Dezember 2014 Carinostoma elegans new to the Slovakian harvestmen fauna (Opiliones, Dyspnoi, Nemastomatidae) Anna Šestáková & Ivan Mihál doi: 10.5431/aramit4804 Abstract. A new genus and species of small harvestman was found for the first time in Slovakia – Carinostoma elegans (Sørensen, 1894). One male and two females were collected in the Mlyňany arboretum of the Slovak Academy of Science (western Slovakia). Descriptions and photographs of both sexes of C. elegans are provided. Additional com- ments, and a map of distribution of all species of this genus, are provided. Keywords: arboretum, faunistics, harvestmen, new record, western Slovakia Zusammenfassung. Carinostoma elegans neu für die Weberknechtfauna der Slowakei (Opiliones, Dyspnoi, Nemastomatidae). Eine neue Weberknechtgattung und –art wurde erstmals in der Slowakische Republik nachge- wiesen – Carinostoma elegans (Sørensen, 1894). Ein Männchen und zwei Weibchen wurden im Mlyňany Arboretum der Slovakischen Akademie der Wissenschaften nachgewiesen. Beide Geschlechter sowie die Verbreitung der Art werden beschrieben und abgebildet. Altogether five species in three genera from the and the number of genera increases to 25 (Bezděčka family Nemastomatidae are known to occur in Slo- & Bezděčková 2011, Mihál & Astaloš 2011). As the vakia. During a brief zoological investigation into species is new to the Slovakian harvestmen fauna, we the arachnid fauna in the arboretum Mlyňany of provide a description of its morphology and compare the Slovak Academy of Science three specimens of its distribution to other species of the genus. a harvestman so far not known as a member of the Slovakian opilionid fauna were found. The specimens Methods were identified asCarinostoma elegans Sørensen, 1894.