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New Species and New Records of the Genus Anomotarus Chaudoir, Subgenus Anomotarus S

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New Species and New Records of the Genus Anomotarus Chaudoir, Subgenus Anomotarus S ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de SPIXIANA 34 1 21-32 München, September 2011 ISSN 0341-8391 New species and new records of the genus Anomotarus Chaudoir, subgenus Anomotarus s. str., from the Oriental and Australian regions. Supplement to the “Revisions of the species of the subgenus Anomotarus Chaudoir s. str.” (Coleoptera, Carabidae, Lebiinae) Martin Baehr Baehr, M. 2011. New species and new records of the genus Anomotarus Chaudoir, subgenus Anomotarus s. str., from the Oriental and Australian regions. Supplement to the “Revisions of the species of the subgenus Anomotarus Chaudoir s. str.” (Coleo- ptera, Carabidae, Lebiinae). Spixiana 34 (1): 21-32. As a supplement to the revisions of the species of the genus Anomotarus Chaudoir s. str. from the Oriental and Papuan regions, and from Australia, four new species are described: A. mourae and A. toombae from Queensland, Australia, A. jakli from Tanimbar Islands, and A. scriptus from Thailand. Also new records of some rare or little known species are communicated. The new species are inserted in the most recent keys to the Oriental and Australian Anomotarus. Martin Baehr, Zoologische Staatssammlung München, Münchhausenstr. 21, 81247 München, Germany; e-mail: [email protected] Introduction The genus Anomotarus Chaudoir belongs to the lebiine subtribe Calleidini and consists of more or less While visiting recently a number of Australian elongate, usually ground living carabid beetles which collections and checking the unidentified carabid are characterized by more or less securiform labial material, I sorted out specimens of a number of rare palpi, rather projected eyes, usually rather swollen or undescribed species of the subgenus Anomotarus orbits, and denticulate tarsal claws. A number of s. str. of the lebiine genus Anomotarus Chaudoir, Australian species, in particular those of the subgen- 1875. They are communicated or described herein era Nototarus Chaudoir, 1875, Lithostrotus Blackburn, to receive a better idea of the taxonomic diversity 1894, Amplitarus Baehr, 2009, and Rubritarus Baehr, and the distribution patterns within this subgenus. 2009 possess atrophied metathoracic wings, whereas During a recent collecting trip through northern most species of the subgenus Anomotarus s. str., parts of Northern Territory and Western Australia except some montane Australian species, have well I sampled material from certain species which are developed wings. also communicated. Additional material of new The subgenus Anomotarus s. str. is distributed or rare species was kindly submitted by S. Bilý from Pakistan in the north-west through southern (Praha), M. Häcker (Praha), E. Riley (College Station, and eastern mainland Asia to Japan, the Indonesian Texas), and D. Wrase (Berlin). So the present paper and Philippine insular belts including the Moluccas, is regarded as a supplement to my revisions of the New Guinea, the Bismarck Archipelago, Solomon Oriental-Papuan and the Australian species of the Islands, New Caledonia and adjacent islands (Baehr subgenus (Baehr 2004, 2006). 1996, 2004, 2005), and Australia (Baehr 2006), with 21 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de two Australian species introduced into New Zealand In species described by earlier authors only the (Larochelle & Larivière 2007). original citation and the citation in the revisions (Baehr Baehr has revised the Oriental-Papuan and the 2004, 2006) are given. For full literature records see the Australian species in two papers (Baehr 2004, 2006), revisions. Additional records of the common and wide- and the other subgenera which exclusively occur in spread Oriental species Anomotarus stigmula (Chaudoir, 1852) and the widely distributed Australian species Australia were revised recently (Baehr 2009). Anomotarus crudelis (Newman, 1840) are not communi- From Australia, including adjacent Lord Howe cated. Only if few specimens are recorded, their exact and Norfolk Islands, at present 48 taxa of the nomi- number is noted. nate subgenus were recorded (Baehr 2006) that cover the whole of mainland Australia, whereas from Abbreviations of collections Tasmania so far only a single species is known to AMS Australian Museum, Sydney occur. Species diversity is highest along the east coast CBM Working collection M. Baehr in Zoologische and in tropical northern Australia, but a couple of Staatssammlung, München CHP Collection M. Häcker, Praha species has managed to live even in semiarid areas CWB Collection D. Wrase, Berlin in the interior. IRSNB Institut Royal des Sciences Naturelles, Bruxel- So far 38 extra-Australian taxa have been re- les corded (Baehr 2004, 2005), of which 21 occur in the NTD Northern Territory Museum and Art Gallery, Oriental Region and 17 in New Guinea, the Bismarck Darwin Archipelago, Solomon Islands, and New Caledonia QDPIB Queensland Department of Primary Industries, and surrounding islands. Brisbane Generally, very little has been recorded about QM Queensland Museum, Brisbane habits and life histories of any species of the genus, TAMU Texas A&M University Insect Collection, Col- because all species apparently are strictly nocturnal lege Station, Texas and thus, most specimens present in collections were WADAK Western Australian Department of Agricul- ture, Kununurra sampled at light. The nocturnal habits are corrobo- rated by the common observation that specimens Abbreviations of states of Anomotarus s. str. arrive at the lamp only in total NSW New South Wales darkness and rather late in the night. Therefore, any NT Northern Territory additional observations about habits and habitats are QLD Queensland important for better understanding the biology of VIC Victoria these beetles, and thus, even collecting circumstances WA Western Australia of apparent fairly common species are communicated in the present paper. The species Anomotarus humeralis Sloane Methods Sloane, 1917: 435. – Baehr 2006: 22. Examinations and descriptions follow the methods used in the revisions of the Oriental-Papuan and Australian Distribution. A fairly common species which is Anomotarus s. str. (Baehr 2004, 2006). It should be men- distributed through tropical northern Australia from tioned again that body length was measured from apex north Queensland to north Western Australia. of labrum to apex of elytra; length of orbit was taken from the posterior margin of the eye to the position Collecting circumstances. Most recently sampled where the orbital curvature meets the neck; length of specimens were collected at light near rivers or pronotum was measured along midline, and width of billabongs. base of pronotum at the position of the posterior lateral New records: NT: “NT9, Mary River NP, Mary River setae. Billabong, 5 km nnw. Mary River Crossing, 12.53.49S, The habitus photographs were obtained with a di- 131.38.33E, 28 m, 1.-2.11.2007, M. Baehr” (CBM); “NT19, gital camera using ProgRes CapturePro 2.6 and Auto- 3 km e. Elsey Creek, Roper Bar Rd. 15.00.23S, 133.16.72E, Montage and subsequently were edited with Corel 148 m, 5.-6.11.2007, M. Baehr” (CBM); “NT22, Roper Photo Paint 11. River Crossing 3 km e. Roper Bar, 14.42.83S, 134.30.51E, The types of the new species are stored in Queens- 10 m, 6.-7.11.2007, M. Baehr” (CBM); “NT25, 5 km s. land Museum, Brisbane (QM), the private collection of Larrimah, Stuart Hwy. 15.32.07S, 133.23.82E, 184 m, D. Wrase, Berlin (CWB), and the working collection of 7.-8.11.2007, M. Baehr” (CBM). – WA: “Kununurra, the author in the Zoologische Staatssammlung, Mün- 22.12.1991-6.1.1992, R. I. Storey” (WADAK); “WA84, chen (CBM), additional material is stored in the collec- Ord River Crossing, 102 km nne. Halls Creek, 17.28.74S, tions mentioned below under “Abbreviations”. 127.57.11E, 298 m, 26.-27.11.2007, M. Baehr” (CBM). 22 ©Zoologische Staatssammlung München/Verlag Friedrich Pfeil; download www.pfeil-verlag.de Anomotarus r. ruficornis Sloane Anomotarus micans Baehr Sloane, 1917: 436. – Baehr 2006: 25. Baehr 2006: 30. Distribution. The nominate subspecies of this rather Distribution. Apparently a fairly common species common species is distributed through tropical which is distributed through the tropical and sub- northern Australia from north Queensland to north tropical northern half of Australia from mid eastern Western Australia. Queensland to north Western Australia. Collecting circumstances. The recently sampled Collecting circumstances. All specimens were col- specimens were collected at light near rivers or bil- lected at light, mostly near rivers or billabongs. labongs, the one from Howard Springs was sampled New records: QLD: “Ayr, 8.XI.1990, W. F. Chamber- “ ” in litter . lain” (CBM, TAMU); “Mossman, 11.XI.1990, W. F. Cham- New records: NT: “Howard Springs, 11.11.1979, M. B. berlain” (TAMU). – NT: “NT19, 3 km e. Elsey Creek, Malipatil” (NTD); “NT13, Kakadu NP, South Alligator Roper Bar Rd. 15.00.23S, 133.16.72E, 148 m, 5.-6.11.2007, River Crossing at Old Jim Jim Rd. 13.02.95S, 132.19.13E, M. Baehr” (CBM); “NT22, Roper River Crossing, 3 km 43 m, 3.-4.11.2007, M. Baehr” (CBM); “NT29, 70 km n. e. Roper Bar, 14.42.83S, 134.30.51E, 10 m, 6.-7.11.2007, Top Springs, Buntine Hwy. 16.00.94S, 131.56.33E, 253 m, M. Baehr” (CBM); “NT25, 5 km s. Larrimah, Stuart Hwy. 9.-10.11.2007, M. Baehr” (CBM). 15.32.07S, 133.23.82E, 184 m, 7.-8.11.2007, M. Baehr” (CBM); “NT109, Kakadu NP, Jim Jim Billabong, 12 km sse. Cooinda, 12.56.26S, 132.33.10E, 16 m, 2.-3.12.2007, Anomotarus ruficornis plagiatus Baehr M. Baehr” (CBM); “NT114, Fogg Dam, 8 km ne. Hump- ty Doo, 12.33.44S, 131.17.86E, 23 m, 5.-6.12.2007, M. Baehr, 2006: 27. Baehr” (CBM). Distribution. This southern subspecies is distrib- uted in the southern half of eastern Queensland and Anomotarus rufescens Baehr in north-eastern New South Wales. Baehr 2006: 32. Collecting circumstances. Most species were col- lected in pitfall traps on the ground, one in “dry Distribution.
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