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KUSC E G ee es i a su u a e i o me a ew ea a case s u y e i e i y a s a us o Co eo e a i e a u a a mo i ie a i a s o y ie Auck a (197 -19 9 / G Kusc e — Auck a SI 199 ( SI a o ec io e o ISS 11 - 1 ; o 3 IS - 77- 59 - I i e II Se ies U C 595 7 (931 11 © C ow Co y ig
u is e y SI a o ec io M A e esea c Ce e i a e ag Auck a ew ea a ecem e 199 i e y Ge e a i i g Se ices e so ew ea a Etiam pristina in a u a A sο i a a u a se i g summa securitas et futura sweet tranquility and nature
O IS IECE e o e - ee i g emo yc i ee e i o y c us u u us ( ou o i s a i e o oca os ka ika ea ( ac yca us ac y ioi es om e y ie ee e su ey a ea Auck a ew ea a e w i e g a u es o e ee e a e o e c us e s a ass e e s is ee e as a e ic s a us o u ike a o e ua a a (S e o o as ossi e i e ce s ows a e wee i g ou was i i g i u assic imes way ack i e e a o e i osau s a gym os e ms mo e a 1 mi io yea s ago O EWO
As a small boy in the 1930s I used to collect butterflies on the South Downs in southern England. On one occasion I met a elderly gentleman collecting beetles. He explained to me that most entomologists started with butterflies but later they graduated to more important groups such as beetles. I have been in awe of coleopterists ever since! This splendid publication by Dr Willy Kuschel has reinforced my awe. I have been asked to write this foreword as a conservation biologist. I can say with conviction that conservation cannot be effective unless it is based on good taxonomy and knowledge of the habitat requirements of species. The Lynfield Beetle Survey will not only be a valuable study for New Zealand entomologists, but will also be a very significant contribution to conservation biology. Movement of people and goods will increase in the future, and therefore the number of foreign species will increase in all biotopes. It will become more and more important to discover the extent to which autochthonous species — especially endemic ones — survive increased com- petition from foreign species. If we are to retain endemic species we must establish reserves of native habitat. As the years go by more of these will become habitat islands in a sea of crop and urban land, where foreign species predominate. This situation raises a host of difficult practical problems. How large should the reserves be? How close to each other? Are habitat corridors necessary between reserves? These problems can best be tackled by recording what happens in the field. Joseph Banks, who noted "a few Butterflies and Beetles" in New Zealand in the eighteenth century, would have been amazed by the number of beetle species recorded by Willy Kuschel. a ks ο e os i a i y o Wi y Kusc e a e e ey o oway I a e some i s - a e e ie ce o e y ie is ic is muc ess is i guis e isi o wou e e a e guesse ow ma y e emic s ecies emai i i Wi y Kusc e s ows a mo e a ee- qua e s o e ee e s ecies i e a ea a e e emic is is o i e es i g a e cou agi g As ea y a e e emic s ecies a e co i e o e emai i g agme s o us i is i a a suc aces e e ai e wi i y ie a simi a is ic s The conservation value of quite small areas of native habitat within urban and suburban areas has been demonstrated in several countries. It is particularly interesting that so many endemics survive in New Zealand where the proportion of foreign species is so great. Only monitoring will show the extent to which the endemic species will remain in their habitat islands. Wi y Kusc e s mas e y s u y o i es a i m ase ο w ic o e s ca ui e sce e which he has studied so thoroughly, will increase in significance; future generations will be deeply grateful for his pioneer work.
Norman W. Moore The Farm House Swavesey Cambridge [ o ma Moo e was o me y Se io i ci a Scie i ic O ice wi e a u e Co se a cy Cou ci o e U i e Ki g om] 18 May 1990
I A S AC
The Auckland suburb of Lynfield, on the northern side of the Manukau Harbour, presents a typical picture of urban development under topographical constraint. Contour and substrate along the coastline have kept urbanisation away, with the result that substantial areas of vegetation remained relatively unmodified. These are now set aside as reserves. There is also some farmland and recreational land in the area, apart from domestic gardens. The proximity of these varied environments to each other and to the author's home has made possible an intensive study of aspects of their natural history. Between 197 and 19 9 but especially 1975- 1 the area was subjected to regular and intensive sampling of its beetles, the insect group chosen for a survey. No accessible habitat type was left unshaken, unswept, u sei e or otherwise u sam e The several collecting methods employed to secure examples of the beetle fauna are outlined, and the habitats and diversity of life are discussed. Although the domestic gardens had a high content of indigenous plant species, only 9 of the native beetle species were found to be present also outside the native bush. To expose and partly explain the low adaptability of the indigenous fauna to modified environmental condi- tions, the geological background is briefly outlined; plant genera that thrive equally well in the bush and in gardens, and that are known to be highly attractive to many beetle species are scrutinised; and the ecological preferences of beetle genera represented by native and foreign
species are examined.
• At the conclusion of the field work 9 beetle species in 5 families had been obtained, 753 of them endemic. Some general statistics on their overall status are tabulated, and these are followed by a systematic tabulation, species by species, of such va riables as provenance, wing condition, relative abundance, and habitat preference. More expansive information is given in a annotated list of the species, which includes for each family a illustration of a representative member. Essential nomenclatural changes are formalised in a separate section; these include new synonymies, new combinations, new status, and a few new genera and species from among
the many awaiting desc ription in this material. The work is illustrated with habitat photographs, habitus and morphological drawings, a topographical map, and aerial photos. A brief account by A. E Es e of the botany of the survey area is appended, with a vegetation map. IS O CO E S C eck is o ami ies I o uc io 7 Ack ow e gme s 7 S uc u e a ese a io o su ey e o Co ec io me o s 9 a i a y es a ei ee e au a 11 e a i e io ic i e si y i e su ey a ea oes e i ige ous au a easi y a a o mo i ie e i o me a co i io s? I a u a summa ies o s ecies a a i c u i g a e o s a is ics 3 A o a e is o s ecies 39 a o omic o es (a ec i g Ca a i ae Scy mae i ae se a i ae Co y o i ae Ce am yci ae e i ae a Cu cu io i ae 7 e e e ces 5 a es i us a io s o e ese a i e s ecies o ami ies eco e a y ie 9 A e i ege a io o e Ma ukau coas a y ie Auck a ( y A E Es e I 1 Ma s a ae ia o og a s o e su ey a ea a use ege a io I a o omic i e (Co eo e a o y 111
C ECK IS O AMI IES
A e i ae 3 Cu cu io i ae 3 7 a ac i ae 31 57 Agy i ae 5 1 e mes i ae 3 53 ycoseci ae 3 55 A o ii ae 3 5 y isci ae os omi ae 33 A ici ae 3 5 E a e i ae 9 5 se a i ae 7
A i ae . 35 9 E o y i ae 31 5 i ii ae A c aeoc y ici ae 33 Euc emi ae 3 53 i o ac y i ae 9 5 e i ae 3 7 e o i ae 9 5 i i ae 3 5 e i ae 3 7 is e i ae 5 i i o i ae 33 3 u es i ae 9 5 y o i i ae 9 i o agi ae 31 5 y i ae 9 5 a gu ii ae 31 57 yso i ae 39 Ca a i ae 3 53 a i ii ae 3 Sa i gi ae 3 5 Ca a i ae 39 eio i ae 5 1 Sca i ii ae 5 3 Ce am yci ae 3 uca i ae 9 51 Sca a aei ae 9 51 Ce y i ae 3I 5 yci ae 3 53 Sc a ii ae 3 C ysome i ae 35 Me a yi ae 33 Scy mae i ae 5 Cii ae 33 Me y ae 3 55 S a y i i ae 5 3 C am i ae 9 5 Me o ysii ae 3 1 e e io i ae 34, C e i ae 3 55 Mo e i ae 33 3 ogossi i ae 3 5 Cocci e i ae 3 59 Myce o agi ae 3 I o e i ae 33 Co y ii ae 33 3 emo yc i ae 35 9 Co y o i ae 31 5 i i u i ae 31 55 C y o agi ae 31 57 3 ages ci e a e ose o w ic eac ami y Oe eme i ae 5 a ea s i e a u a Summa ies o S ecies Cucu i ae 31 5 e i ae 3 5 a a a i e A o a e is o S ecies I o uc o y Sec io s 7 INTRODUCTION I c easi g co ce is e esse y ci y we e s a o e a io a ace o e omes a ga e s w ic co e s e wo a ou e co i io o ei e i o me is e ig c i s a ee e essio s o the o si e o is e ec e i e ac a mo e a mo e a ge ci ies a e e Ma ukau a ou e a ea seeme o e a ac i e su ec o s u ies o e qua i y o ei i e s ace O o suc a su ey ecause o ea y accessi i i y o e s ecia i e es a e e g ee is a s — a ks ayi g au o e sui a i i y o e a ks a ga e s as ega s ie s ceme e ies a ga e s—sca e e a ou i a ma e ei e a i e y ig co e o a i e a s ecies a o s ee s a omes a so e a ai a i i y o some u a a A y u a e e o me is a mos i a ia y ece e e ages o is e o s ow a a g a ce a e ee e y a o a es uc io o e a a a ima i e o the au a o y ie is ic a a ie i o ma o ecog- si e e g ee a eas a e su seque y a sca e i o the ise a i a s e a i e us o o es w ic is e se y su u s y ome ow e s a ci ic a mi is a io s usi g ege a e a e ce a o s s e e agai s esicca io u ose u y se ec e a s e e is a y e e oom a e osu e; a e ga e s a a ks w ic usua y o a s o i e e o e o the si es Co seque y ea e e g ou e ose o wi ig a insolation. o y easi y is e se s ecies o wee s may gai a o ey a so emo s a e a mos i ige ous ee e s e- s o a eous y o a w i e e g ee a eas o ci ies a e cies a e o o e a ce o o y a e y ow o e a ce o e e o e ma -ma e a i icia a i imica o a y i g eco ogica c a ges a a e u a e o i e ou si e e o a e A ima s e ce i g e s a e as a u e o a a i e us w e eas a a ge o o io o e o eig o a e sc emes ey mus come i o ci ies i ci e - s ecies a e equa y a ome i o a i a s a y o o ei ow acco e e a es a yi g i sec s e Go wa ic agme a is ow ew ea a o ca i a e u a a eas wi e a i e ease u ig ess Ao ea oa as i e i e o a a au a e eme s om i sec s a o e sma i e e a es ge e a y i s ee s e massi e a cie co i e a occu ow e e e se i a mos im ossi e o a e se e wo ese e eme s a e a egacy om C e aceous I is ecise y the sma au a o i e e a es a e imes 7 o mo e mi io yea s ago i e e a o e s o a eous o a o wee s mosses a ic e s a a e i osau s w ic o y ew ea a i e i e a o y commo y c ose y esea c e s o mo i o ei e the ew ea a ca sa egua wi i i s u ique a imo y i e si y o io a o e eco ogy o i i i ua a a o s ecies o e sys ema ica y io ogica y a eco - a ima s ecies a icu a y o i us ia ci ies a a e ogica y a ua e i is o ecessa y a ey e as aci g o ems wi emissio s mi e ue s a a e co s icuous as o i s a ce a kau i ee o a aka e im ai i g e qua i y o ai a wa e s ( uck 1955 1959 Muc o the a i e o es au a is sma c y ic e usi e; S ee e 1973 awkswo 197 a is 197 a kie & a ma y s ecies we e a e o a s a i ew ea a E e 197 K aus i e & Sei e 19 e o e the a i a o e o y esia s a a e e Eu o- e s u y esc i e e e is a a e y e i s o i s ea s ey a e ow o e way o su e e i c io u ess ki u e ake i ew ea a I ea s wi a e a i e y e i ige ous ege a io w ic co ai s e s ecia we k ow g ou o i sec s the Co eo e a o ee es a i a s a a e so i a o ei su i a is ese e As is su ey seeks o e e mi e e s ecies i e si y i the is su ey as emo s a e e e e e a i e y sma ece y e e o e esi e ia a ea o y ie a su u us a c es a so a a e esca e es uc io may o M oski i Auck a Ci y a i e a u a ege- co ai au as o u sus ec e ic ess a i e si y
ACKNOWLEDGMENTS W A Su i ky ki y e mi e me ee om o access o a a o i C aw e o e muc ime a e o o a ocks a ima ya s a e Wai aki S eam us e imi a y ocessi g o a a om the e e si e co ec- My a ks o e e ey e e G ace a eggy e - io s om y ie A a Es e as ge e ous y o i e e i ia Ma i Ma ga e ocke a i A e e Wa ke a ou i e accou o e o a y o the y ie a ea (see o ocessi g e ma y sam es a s ecime s om a e i Ι am g a e u o o y a is a es e mo e y ie Ι g a e u y ack ow e ge e assis a ce wi o ei ai s aki g wo k i c ea i g mos o e habitus i e i ica io s o i e y ma y s ecia is s o a y E i us a io s; o Ma is essi e a K ys ei e o ig M o so S Ε ö y-Υου ga Εsρañο e a a io o e o og a s; a o e o C os y a a L.R. Gi og y Y Gomy G a s ea M C e e a u a o a ice o e co e a ese a- ammo C M o ayek A o oway K io o is e o a o maki g i a ea i y oug e ous o C o so J.F. aw e ce I ö Moo e medium o SI a o ec io s esk o u is i g A.F. ew o R.D. o e Se ma om so u i i a y my si ce e a ks o o o gwo I ow se C Wa G We e a W Wi me i ec o o SI a o ec io o e co i ui g o ee es; A a Es e o a e ogams a e s; essica ogis ica su o w ic as e a e me as a esea c ee e o mosses; a oa i g ey o u gi G ace Associa e o i g is s u y o com e io nf ld tl S rv S UC U E A ESE A IO O SU EY E O This report is essentially a statement of the background, Zealand record and its source, and present distribution, techniques, and findings of a rather simple but long-term both in New Zealand and worldwide. By means of super- biotic survey. It is not intended as a guide to identification, script key letters, attention is drawn to those species which although it will perhaps be found helpful in that regard. (a) are introduced, (b) are first reported for New Zealand Nor does it seek to draw any comprehensive ecological in this publication, (c) have Lynfield as their type locality, conclusions or speculations, even though there is a basis and (d) are involved in some taxonomic change. for this implicit in the findings. It is felt that the data of In the course of this study it has become apparent that themselves make a most eloquent statement regarding a number of taxonomic changes—new combinations, new species diversity in what might too easily be dismissed as synonymies, and new taxa — would be desirable. These `just a little bit of bush' in suburbia. have been grouped together under the section 'Taxo- The section `Collection Methods' (p. 9) outlines the nomic Notes' (p. 76), for the convenience of readers with techniques employed in obtaining beetles from the many a particular interest in such concerns. Only a very few new different kinds of habitat found in the survey area. It taxa are erected here; a substantial body of material necessarily mentions some of the floristic elements of remains to be formally described. Three pages of line certain habitats, but these are described in detail in the figures are given in support of the descriptions of new next section, `Habitat Types and their Beetle Faunas' (p. taxa. 11). Further information on the Lynfield flora may be The section devoted to References (p. 86) is by no found in Alan Esler's appendix to this report (p. 102), means comprehensive. It is confined to citations of first which includes a list of native plant species and a vegeta- records of foreign species in New Zealand, and to authors tion map. cited in the section `Taxonomic Notes'. Background In the two subsequent sections (pp. 20, 21), considera- information on established taxa may be sought in such tion is given to the influences which have shaped the standard works as Junk's Coleopterorum Catalogus. Lynfield area and its biota into their present-day form, and For each family in the Annotated List, a habitus draw- to some particular features and associations found there. ing is given near the family heading as an aid to visuali- Emphasis is given to the relative species diversity appar- sation. Note, though, that some of the illustrations used ent in the native element of the flora, the vertebrate fauna, are of species not recorded at Lynfield; they stand as and the Coleoptera. examples only. However, habitus drawings of many of the On p. 23 is a table of statistics which summarises and Lynfield species were already available at the time of contrasts quantitatively some of the biological features of writing. They are incorporated in this volume as a series the 982 beetle species found in the survey. This is fol- of plates (p. 90), in the hope that keen students will find lowed on p.24 by a comprehensive tabulation of species them useful in putting names to a number of common data, in systematic sequence of families. The legend at the beetles. Some duplication of figures between the plates foot of each left-hand page of this section indicates the and the Annotated List has been inevitable. meaning of the data codes used in the table, and it is A taxonomic index (to the Coleoptera only) has been possible to quickly gain an impression of the status of included with this report, for the convenience of readers. individual species and of larger taxonomic groupings for However, the alphabetical `Checklist of Families' (p. 6) a number of biological variables. and the alphabetical sequence of genera and species under The meat of the report is to be found in the section family-group headings provide easy alternate access. `Annotated List of Species' (p. 39), in which information The terms native, indigenous, and endemic are often is given regarding each of the beetle species recorded at used in the text, and may require clarification to prevent Lynfield during this survey. The same sequence of taxa is uncertainty or ambiguity. `Native' and `indigenous' are used as in the tabulation of species data. At the family- synonymous, and refer to species which may occur also group level this sequence, and the names used, represent outside the territory under consideration (in the present either the best current opinion available to me from context, New Zealand plus outlying islands) but are con- specialist collaborators, or — for the Curculionoidea in sidered to occur naturally in the area, i.e., their presence particular — my own view of the systematics of the group. is not the result of human traffic. `Endemic' is reserved for The genera and species of each family-group taxon are those species that occur solely in the territory under presented in alphabetical order. consideration. For all species, even those as yet undescribed, informa- The voucher specimens from the Lynfield beetle sur- tion is given on their apparent ecological affinities at vey are housed in the New Zealand Arthropod Collection Lynfield. For introduced species, this is supplemented (NZAC), DSIR Plant Protection, Mt Albert Research where possible by an indication of the earliest New Centre, 120 Mt Albert Rd, Auckland 3, New Zealand. I o uc o y Sec io s COLLECTION METHODS The principal collecting methods used to retrieve the into a holding bag, which is labelled with a numbered tag. beetle fauna for the Lynfield survey are outlined below. The particulars of the site are entered in a notebook: Beetles occur in practically every conceivable niche of relevant information on the vegetation that formed the both natural and modified environment, be they terrestrial canopy and the undergrowth, and the conditions on the or aquatic, endophytic or aerial, topical or interstitial. ground. Four main types of litter were sampled: (a) the Beetles may readily fly, or may be flightless and just loose or free litter of dropped twigs, leaves, flowers, fruits, crawl. Some are active in the daytime, others at night; a and seeds; (b) the litter and mould in hollows in the trunks few are almost indifferent to the time of day. The collect- and major limbs of living and dead trees; (c) the litter from ing of organisms of such immensely diverse lifestyles decaying wood, crumbled or chopped over and into a requires a whole series of techniques to be applied for sifter; and (d) the wrack by the sea-shore. The ground litter years through the four seasons, should a thorough survey in the area surveyed was as varied as the canopy and be attempted. topography were diverse, requiring large numbers of samples to ensure adequate coverage of its rich fauna. (1) Beating vegetation. This is done by holding a beat- Holes in trees were rather uncommon. A few were found ing sheet or tray under the foliage of trees, shrubs, lianes in Vitex lucens, Dysoxylum spectabile, and Sophora and higher ground plants while giving one or two sharp microphylla among the native plants, and only in Salix jarring blows with a stick. The sheet is then scanned, and fragilis among the foreign element. Scrapings of the either the specimens are picked up by hand, or with caudex of treeferns and necrotic areas of tree trunks aspirators or tweezers, or the entire beaten material is induced by Phellinus and other fungi were included as transferred to bags for processing through Tulgren extrac- litter samples. Litter was taken continually for seven tion funnels or for sorting by eye under shelter. The opera- years, amounting to 824 samples processed through tion may be repeated on the same plant species until Tulgren funnels, but only the first 104 were numbered and sufficient material has been secured to ascertain possible entered into the Systematicc Group litter book at Mt biological or ecological associations of individual beetle Albert Research Centre, Auckland. species with a particular plant species or niche. The mat- erial is then labelled and put aside before beating another (4) Sifting garden and farm litter. This litter may be plant species. Canopy beetles are more readily collected compost heaps of lawn clippings, plant prunings, vege- earlier in the morning and into the evening. The canopy tables, fruits and kitchen refuse, pure or combined, as well usually becomes more accessible for this operation at as hen-house straw and feed, chicken and sheep manure, bush margins, on regenerating vegetation around paths and stored foodstuffs. Bags of lawn clippings mixed with and clearings, and on the upper part of steep slopes, fermenting fruits and vegetables were frequently taken gullies, and cliffs. into the Wairaki Stream bush to test the adaptability of native species to this unusual litter, and to see whether the (2) Sweeping vegetation. The insects living or resting foreign beetles that were so common in the exposed on vegetation can be obtained by sweeping with a firm net. compost heaps in gardens would take to it even when This method is particularly practical for the low vegeta- placed in dense native bush. tion of meadows, roadsides, wastelands, and wetlands in the open and for grasses, sedges, and ferns in the bush. If (5) Floating from stream-bed deposits. Only the lower clean specimens are required from night sweeping, this 200 m or so of two streams have a bed, and only the must be done in the early hours of a breezy night, espe- Wairaki Stream has some gravel. The areas above and cially on the open vegetation of paddocks and roadsides, around the streams are urbanised, and storm-water run-off where slugs and snails commonly abound on wet and from properties and streets carries down litter, which is dewy nights. heaped up as flood debris in sedges and obstacles on the way to the sea. Stream-bed carabids and staphylinids are (3) Sifting natural litter. In this context litter is any easily disturbed by splashing the ground with the water organic matter breaking down on the ground under native nearby, or floated out by scooping gravel, sand, and silt vegetation or washed up onto the sea-shore. Included in i o a ucke ο wa e As oo e is ca o a so s this concept are twigs, branches, logs, fallen trees, and the o ee es om ou si e a i si e e us a ma y o lower part of the trunk of standing dead trees. Litter em a e a e e a gic e ma e ia is agge u si e samples are taken by scraping with a trowel ground debris i oo we o si e i yis o su seque ocessi g and the loose top layer of humus into a sifter made of tough oug u g e u e s material, which has a top handle on a firm ring, another handle below attached to a 1-cm-mesh sieve, and a col- (6) Washing and sifting soil. This method consists of lecting bag tied below this. The gathered litter is shaken washing soil through sieves of progressively finer mesh, about with a turning or tumbling action, and the finer and is extensively used to extract nematodes and other materials and insects fall through into the bag. The proce- small organisms from the soil. Although no species were dure is repeated on the same type of litter until 2 or 3 cubic extracted in this way that had not been obtained in deep decimetres have been sifted. The sample is then emptied pit-traps as well, the method was very handy to check
1 nf ld tl S v quick y o o e s ecies o o mo e s ecime s om ew (11) Malaise trapping. wo Ma aise a s we e se u si es a o co ec soi - we i g a ae a u ae a Wa e ay o e i e se us a e u e a o e mai ri u e ig esi e e s eam e a e o the i (7) Netting aquatic and splash zone species. e dytis- a ie a ea u e a canopy o Vitex lucens a yso y um cids a aqua ic hydrophilids a e easi y see ei e swim- spectabile. ey we e e e e o y om a ua y o mi g i c ea wa e s o w i s su aci g o ick u ai i Ma c 19 1 because o eque a serious i e e e ce mu ky wa e s e e is use ow -s eam om ou e s om asse s- y wo u e Ma aise tra s we e u u i ( is u e a e i e (was e a moss (s as e e Wairaki S tream us away om uma tra ic O e a a so o scoo u ea es a wigs om s i wa e s o e 3 m ig was ace i a sma c ea i g su ou ed e e e ma e ia is agge o su seque e ami a- y a asso me o a i e ees a s u s a a g ou io co e co sisti g i e mai o Schoenus tendo, wi a ai (8) Baiting with plant cuttings. Cu a c es o ees amou o Hakea sericea, Solanum mauritianum, Acacia s u s a lianes a e u e u a ie o o a c es verticiliata, a Ulex europaeus. e o e a was o ei ow ki i suc a way a a eas a o e ace o a c a e o i g o y a ick e o wa e a u e ouc es e a a owi g ig ess ee es o ge a ke y Pseudopanax Iessonii, Olearia furfuracea, o the cu ma e ia a a ea i g s ee ca e e Cyathodes fasciculata, Geniostoma u es e Hedycarya u e ea wi ou i a ce Eac u e is agge wi arborea, Coprosma o us a a Me icy us ami ο υs e ame o e a a e a e i was se u is was wi e o y owe i g s ecime i e e i e s u y a ea o e simu a eous y i e o es ed a eas a i ga e s o e a m Rhopalostylis sapida a ew me es ow - - 5 m away ac oss a a ock w e e e e same stream e o i g a s wi 75 e a o we e c ea e a s ecies was a ai a e i o e u es we e week y c ecke o ig y o as o g as ac i i y was o se e is may a e as e o a ew weeks o y as wi (12) Misting with repellents or insecticides. is Parsonsia heterophylla a Rubus cissoides; o mo s me o is use o is u i sec s i i g i woo issu es as o s ecies o Coprosma, Pittosporum, a Pseudo- stream- a k a c i - ace c acks a e e i y es a panax; o e e yea s as o So o a microphylla. c im e s o u ks a ogs o e ica a o e a gi g i a a coas a c i aces a o supralittoral a (9) Baiting for carrion beetles. is o es cooke i e i a ocks co e ed wi a gae o s u e wi a - a aw mea a sma ea mamma s a i s occa- ac es C o s ee s a e ai o the g ou be ow e a eas sio a y icked u om oa si es we e ace i si e o e su eye e aces a e s ay ed with a i e mis a i e ege a io and in a s ig y mo e e ose a ea i a Some i sec s eac i s a y a dro ; othe s ake se e a Acacia mearnsii g o e esi e a a c o a i e o es mi u es e o e ecomi g su icie y is u e o e (10) Pit-trapping. Shallow i -tra s a e use o g ou easi y o ice e i s ec ici e use is y e um- ase au a a ee i -tra s o hypogean (soil-inhabiting) wi a i knock-down u s o e sis e ce au as As s a ow i -tra s a e oo se ec i e a i e i- ( Rearing. Imma u e s ages may e co ec ed a cie i a us si ua io ey we e use a e a e y a ea e o secu e a u s ecime s o e usi e s ecies o e a mos so e y u e o e o ca io ai A o e e a e a ae a u ae wi a u s a o associa e a ee i -tra s we e se u a Wa tle ay a Wairaki ee e s ecies wi i s a icu a os o ic e o e - S eam a ke se ice o se e a yea s is y e o am e e a c -cu i g o g o s Gastrosarus i -tra is e a e y o i g a o e u o 50 cm ee i o nigricollis a Astetholida lucida we e commo u gi g e g ou wi a 1 cm soi auge A a co ai i g y e um e s o a c es o e g ou u ey we e e a o (75 is ace a e o om e ga s e wee a e y see a e e mo e a e y caug A oug e e a a e si es o e i a e sea e o wi k ea e u es o cu a c es (me o a o e we e c ecke c ay o a e a a e oi u e is owe e o e e a a eas o ce a o ig some ee e s ecies o caug ( e o me oce u e u e ou o e easie a c ea e w i e ovipositing we e ea e i um e s e o o e i is e ig y sea e o wi a a o ia e iece o oa o e e su ace au a om E e y o e o e i ee co ec i g me o s us ea e e i g a co e ed wi a a ge astic s ee o kee wi as yie e ee e s ecies o i e es A thoug a ai -wa e ou e si es o i - tra i g mus e oo - es s a ermi e co o ies we e o oug y sea c e o ee E a o was ou o e ac ica o o y ecause ee e gues s o e we e ou e u s o the a o eg igi e e a o a io a a y a y oss o concen- Mesoponera castanea, owe e a e qui e eque y i e Ijalion, u a so i o i g a e a ac i e o soi s ecies i y the i Carpelimus staphylinid, e ptiliids Noto- e tra s we e se ice o ce a mo i o ma wea e ptenidium crassum a N. oblongum, a e Austra ia a mo e o e i we s e s pselaphid immig a Gerallus punctipennis. I o uc o y Sec io s A I A Y ES A EI EE E AU AS (1 o es ca o y ea y a isease u s ea es banksii; a e ma s wee i Steriphus ascitus on Scirpus owe s a ui s o ees s u s a ia es oge e a Typha. e s ecies o e c ysome i ge us Aphilon wi e a asi ic a e i y ic o a a yi g a c es a e associa e wi i e wo s the wee i Bryocatus a wigs u i e c ow s o i e a ic sou ce o oo polytrychi wi a ious moss s ecies a e wee i s o ma y ee es a ee i ec y o a s o a ey Gromilus thoracicus, Agacalles integer, a Pogonorhinus u o o a asi ise ca o y- we i g a o o s e e is a opacus wi g ou e s e ig ess c e i Paupris eme ous amou o ac i i y goi g o ay a ig i aptera was ou o e i a ia y o i o esce ces o e ca o y es ecia y i e g owi g a owe i g Gahnia lacera, a e equa y ig ess c ysome i seaso Mos ca o y ee es a e ca a e o ig Mos Peniticus suffusus o ea es o e o es oo s ecies o i e a s e e e e u a ew — o e am e e Carex, Uncinia, a Gahnia. sca a aei s Stethaspis longicornis a e Costelytra (4) Live wood. Only si s ecies a acki g ea y ees s ecies — isi e o iage i e ea y e e i gs o y o s u s a su s u s we e ou a y ie wo a e e u o e g ou a e wa s O e 753 a i e ee e woo o e s wo a e a c cu e s o e is a i o e s ecies co ec e a y ie 371 ( 9 may e ou i a o e a ga make e woo o e s a a c cu e s a o e ca o y; o e 9 o eig s ecies ou o a e o g o s e o e s wee i s O e o e woo o e s occu a y ie 95 ( may i e i a o ee is Ochrocydus huttoni, e a a o w ic wo ks u a c ow s o wo e e a ea y a e o a ee e ow i e owe a o e u ks o Leptospermum au a o y ie cou e e ua y e o ai e y e scoparium a Kunzea ericoides, a ec i g mos se e e y ea i g me o a o e e o me e a ae kee e ga e ies c ea y is os- (2) Tree trunks. e u ks o ees a e o e i e wi i g o e ass oug i - o es e seco o e is e ia es Metrosideros perforata o M. fulgens o e Oemona hirta, ossi y e mos e sa i e o a ce am- e s Pyrrosia serpens a Phymatosorus s ecies o yci s o e wo as i is ca a e o a acki g com- co e e wi moss i e wo s ic e s a i my e s e e y ea y as we as ea ees s u s a e s o i e s e se wi Tmesipteris s ecies a Asplenium flac- co i e s ico s a mo oco s e a ae a so kee e cidum. Suc u ks a e eque e y a a some y u e s ee o ass y em yi g e s a i gs a saw- a e e co y o i o e ge us Holopsis, a y e us oug i - o es o e ou si e O se a io s e - ig ess a ici e c ysome i Trachytetra rugulosa. e a s wo y o me io a e a ae ou o i g a u ks o Leptospermum scoparium a ec e y soo y oi g we i i e a c es o ea y Chamaecyparis mou a e i a i e y e i i u i Soronia hystrix, a lawsoniana a Cupressus macrocarpa ees (Cu es- e cau e o ee e s y e c y o y c i e wee i saceae ; a ae i 3-yea -o a e ecaye cu a c es Agacalles formosus a e wo cosso i es o o e o Sophora microphylla ( a aceae ; a ae i ea wi - ge us Heteropsis. oug u ks ike ose o Pinus i g s ems o e liane Ripogonum scandens (Smi acaceae ; radiata, o i e i ea co i io s o e Aus a ia e i- a a ae i e g ee sou cu ms o e ussock se ge i e ca a i Philophloeus luculentus, e o e e e o e Ga ia se i ο ia (Cy e aceae (o e o em eye u o i s ime om ew ea a y e i i o i Allocinops brookesi). e a c cu - e s a e Gastrosarus nigricollis, w ic ca a ack a um e (3 Ground plants. e g ou o a is ma e u o mo oco a ico e s o - a o ea e s c u mosses o i e e ees u was o se e o y i Leptospermum mosses a i e wo s i a i e o es us acks scoparium; and Astetholida lucida, w ic a so a acks c ea i gs a we a s e a e ogamic e eme o ma y s ecies o ees u was u i g mos se e e y e g ou o a co sis s mai y o e g ass ge e a Tristania conferta and Eucalyptus s ecies e ga make Nyxetes bidens, Oplismenus, Microlaena, a Cortaderia, e se ge ge e a is a ae o w ic we e ea e om s ems Clematis paniculata Carex, Uncinia, Gahnia, Lepidosperma, Morelotia, o e i e ( a u cu aceae a Nestegis lanceolata Schoenus, Scirpus, a Cyperus, e u us o au o a c es o (O eaceae e i o e is Rhadinosomus acuminatus, w ic is os -s e- Typha orientalis, e i iaceous ge e a Astelia a Dian- ci ic o Haloragis erecta. e a a wo ks i s way ow ella, a e aga aceous ge us Phormium a s ecies Cordyline pumilio. Some ee es s e e i moss ma s i o e a y ig i ie s ems a i g s a e a mo e o e se c us e s o g asses se ges us es as e ias a ess e aceous u e a s a i e a ; o e s a e associa e wi i e a e a e (5) Sound wood. Sou woo is a o es y ea ases o a s o u e ea A ew a e ig y os - ees a s u s o a y i m woo a oes o s ow c ea s eci ic Amo g ose c ose y associa e wi s ecia sig s o ecay As u ga s o es a e a ways ese i a s may e me io e Adriopea pallidata o Gahnia; a u e ose o ascomyce es a es ecia y asi iomy- Psilocnaeia parvula, Novitas dispar, a Phloeophago- ce es s a e e o i g as soo as e ig co i io s a e soma pedatum on Phormium species; Psilocnaeia asteliae, gi e e w o e ocess o eg a a io o e woo is Microlamia pygmaea, a Microtribus huttoni o e g ea y acce e a e i i sec s a ec a k a woo i e same Phormium s ecies us Astelia banksii; e ea - ea y s ages e i sec s a o is ea y wo k a e a mos mi i g wee i Microcryptorhynchus s ecies 1 on Astelia e c usi e y Co eo e a i ci a y o g o s wee i s
1 nf ld tl S rv
and bark and ambrosia beetles. Bark- eetle adults work (7) Wood mould. The trunk and main limb cores of old their way into or through the bark to lay eggs. The larvae trees tend to ecome hollow in some species. Only a few feed in the cambium, engraving patterns that are charac- individuals of Vitex lucens, Dysoxylum spectabile, Soph- teristic for each species. These os -s eci ic beetles are ora microphylla,Metrosideros excelsa, and the adventive usually the first to arrive at a new site. Thus all nine Salix fragilis have provided this niche at y ie The oeo agous sco y i es occurring in the y ie area cen al part of the trunks that has broken down over the have a restricted host association, the ee Dendrotrupes years accumulates at the bottom of trunks or in limbs as species living on Pseudopanax (A a iaceae Chaetopte- humus or wood mould. This particular humus is mic oc i- lius mundulus on Pittosporum ( i os o aceae a species ma ica y different from that under the forest floor litter of Hypocryphalus on Sophora ( a aceae and another because it is 5 sheltered. To emphasise its distinctiveness species of the same genus on Parsonsia (A ocy aceae it is convenien y termed wood mould. Old hollow trees The three foreign bark-bee es are associated specifically are a important ecological element in any forest, as they with Ficus, Pinus, and Cupressus. Four species of weevil provide the unique conditions that some animal species tunnel into wood as adults. They lay their eggs in niches, equi e o survival. O e a dozen species of Scy mae i ae usually at -3-cm intervals around the galleries. The and se a i ae were found to be exclusively in wood larvae then mine away from the tunnel in a radiate fashion. mould or closely associate wi it, in particular the four Psepholax sulcatus and Strongylopterus hylobioides species o Neuraphoconnus, wo s ecies of Stenichnaphes, attack a variety of trees, but the two species of Mesoreda and the se a i species Eupines altula and E. punctata. are associated wi Pseudopanax species only. A further All the wood mould species are small and, with the sole group of beetles that tunnel into wood as adults are species exception of a still unnamed Neuraphoconnus species, that culture their own gardens of ambrosia fungi, on which fully winged. The rather large flightless a ici Co es adults and larvae graze. They are representatives of the o ima which is often seen on foliage at night, seems to sco y i e genera Amasa and Xyleborinus and the a y- prefer hollow trees for hide-outs in the daytime. o i e genus a y us Beetles that do not bore into wood but have y o agous larvae lay eggs in cracks, broken (8) Fungi. Mushrooms, oa s oo s e y and bracket or twigs, or lesions; or, in the case of weevils, prepare an o i- shelf fungi, gall-rusts, moulds, ambrosia fungi, and other osi io site by chewing a pit with the rostrum. The eu e omyce es are a rich source of energy which Co eo- perhaps more than any o er group of insects are families that contribute important woo - o i g species at e a y ie are Ce am yci ae Cu cu io i ae A i i ae able to exploit. The y ie area, and indeed New Zea- uca i ae E a e i ae e e io i ae and A o ii ae A land as a whole, has a large fauna of beetles associate number of weevils and longhorns of sound wood are host- with fungi, some species occu ing on a wide range of specific, as shown in the Annotated List of Sp ecies. fungal groups, others on specific fungi. Because of the usually sporadic occu ence and generally ephemeral ( Decayed wood. Once fungi, bacteria, and insects nature of fungi, myce o agous beetles are almost invari- have degraded wood to a degree that fruiting bodies of the ably full-winge and gregarious, frequently congregating fungi sprout from stems, stumps, and logs and the wood in the thousands at new sites. They tend to be small and crumbles easily, it has reache the stage of rotten or sparsely pubescent or glabrous. Most, for instance decayed wood. This wood is inhabited by all sorts of Sca i ii ae S a y i i ae eio i ae and a i ii ae insects, mites, and worms. Not all types of wood rot feed as adult and larva on the outside; some, for instance evenly. Some trees, like Sophora microphylla and Dyso- Cii ae and A i i ae live as adult and/or larva in the xylum spectabile, decay rapidly on the outside but leave a ui o ies The largest, most conspicuous fungus on tree hard core; others, like Knightia excelsa, Corynocarpus trunks, logs, arid stumps is the cosmopolitan and ubiquitous laevigarus, a Hedycarya arborea, may rot rather evenly Ganoderma applanatum, a polypore that has a ui o y or become pulpy if Ganoderma applanatum has been with a pure white underside. It is found on many live and through the wood while the trees were still alive. Beetles dead hardwoods and softwoods. In y ie it is particu- of decayed wood are almost invariably glabrous, flight- larly common on Knightia excelsa, killing trees prema- less, inconspicuous, and small, some actually exceed- turely, and Acacia mearnsii . Relatively young ui o ies ingly minute. There is a rich variety of fungus feeders and are attractive to many ec o y ic beetles, especially eio- predators. The families which prevail in decayed wood i ae and in particular the species Zearagytodes macu- are i ii ae S a y i i ae se a i ae Scy mae i ae Ι e the agile adults arid larvae of which often congregate Sca i ii ae is e i ae e i ae Co y ii ae and e e- in the hundreds on the bright white underside of the fungal io i ae Most species inhabiting decaye wood remain shelves. Older ui o ies are invaded by Cis zeelandicus unname in New Zealand except for the y ie i ii ae and various other cii s The trunk of a dying Melicytus and Scy mae i ae described from specimens obtaine ramiflorus lying waterlogged on seeping ground was during the beetle fauna survey. One of the smallest beetles found affected by Phellinus punctatus; the effused, esu- in the world, Microscydmus lynfieldi, which measures i a e ui o y of the fungus was cove ing part of the only 55- mm was discovered at y ie and trunk. This fungus was visited regularly over several named after this area of Auckland City. To this day it is years. Three beetle species — the sca i ii Scaphisoma known only from the Wai aki Stream Reserve a y ie species 1zealandicas a y i i sandtheSternotropa
I o uc o y Sec io s 13
Gyrophaena species 2 — were observed almost always families, particularly Staphylinidae, Pselaphidae, simultaneously through the seasons, but mostly in small Scydmaenidae, Leiodidae, Colydiidae, and Curculionidae. numbers. An extensive white-rot spot on a large, live Vitex Some species are so common and ubiquitous that no lucens tree apparently induced by a Phellinus species had matter how small an area they are invariably there. Thus unusually large numbers of three beetle species which it would be hard not to spot or collect in samples at least were found exclusively there or only exceptionally else- some specimens of the ptiliid Notoptenidium lawsoni, the where. The beetles were the rhysodid Kupeus arcuatus, an scydmaenid Euconnus calvus, the staphylinid Ocalea undescribed scydmaenid with small eyes of the genus fuscicornis, the pselaphid Eupines impar, the corylophid Microscydmus, and the large anobiid Hadrobregmus crow- Holopsis pallida, the lathridiidLithostygnus sinuosus, the soni, which was known before only from the Whangarei curculionid Dermothrius farinosus, and several others. If area. Mushrooms and toadstools that grow on the ground dropped flowers pile up under Alseuosmia macrophylla are highly attractive to all sorts of beetles, but especially or Sophora microphylla, or fruits gather under Corynocar- to the staphylinid Atheta atriceps and the mycetophagid pus laevigatus, the rather large aleocharine rove beetle Triphyllus punctulatus, two full-winged native species Tramiathaea cornigera, the male of which has on two found to be consistently present in the bush while always tergites a robust, prominent tubercle, may be present in absent only metres away in the open field. The gall-rust large numbers. The three species of the carabid genus Uromycladium notabile found on most Acacia species Ctenognathus are very active and common on the ground severely attacks A. mearnsii at Lynfield. The fungus is at night. host to a ac us u i ο mis igο ico awhile the galls are fresh, and Araecerus palmaris, alongside the voracious (10) Garden compost. Lawn clippings, prunings, vege- Australian pyralid moth Gauna aegusalis, in older galls. tables, and fruits gradually piled up in the open and in bins The powdery mildews, notably those on Cucurbitaceae, make up the organic matter for compost. This basic are intensively browsed by adults and larvae of Illeis compost material was taken into the native bush to test its galbula, an Australian ladybird vividly patterned in yel- suitability for the endemic fauna, and the ecological low and black, noticed in the country only since March versatility of the foreign beetles that are attracted to this 1985 but already very common in Auckland gardens. fermenting and decomposing vegetable matter in gar- Finally, there are a few beetles that culture their own dens. The biota in compost are extraordinarily rich and fungi, known as ambrosia. The fungus is carried by the varied, regardless of whether compost is in gardens, adult on the body in special cavities or mycetangia into the paddocks, or bush. The composition of the beetle fauna, tunnels the beetles bore. The ambrosia quickly thrives in owe e i e s ma ke y wi e oca io ο e com- the galleries, lining the walls with a fungal carpet that os o ewe a s ecies we e ou o e c ose y adults and larvae crop while moving to and fro. One associated with compost heaps in gardens and open fields. endemic platypodine and three foreign scoltyines have All without exception are foreign, and all but Thalycrodes this special life habit in the Lynfield area. Among these is australis are full-winged. These species comprise some Xyleborinus eucalypticus, whose presence in the country 35% of the foreign element registered for the Lynfield was discovered during the early stages of the Lynfield area. The family Staphylinidae has provided 33 species, beetle survey in March 1975. It had well established Nitidulidae 7, Ptiliidae 6, and Lathridiidae 5; the remain- populations in dead, sound Acacia decurrens, Corynocar- ing 29 species are distributed among 11 further families. pus laevigatus, and Me icy us ami ο us On an unusu- In a native bush situation, the same type of compost is ally warm and humid late winter day, 27 August I980, this capable of luring at least half the foreign species just species was observed swarming everywhere— in the bush, alluded to, in addition to many species from the endemic on beaches, open fields, and urban gardens, indeed over element. the entire Lynfield area and surrounding districts, getting into homes through every open door and window. (11) Open fields. The areas away from beaches and open stream beds not covered in forest or dense scrub are (9) Leaf litter. The leaves, flowers, and fruits shed and designated as open fields. Paddocks, urban parks, recrea- fallen from the treetops and undergrowth constitute the tional fields, and even densely planted gardens if not leaf litter. This organic matter may be sparsely scattered contiguous with native bush come under this heading. The all over or gathered together, in places forming thick beetles commonly found on such open grounds are asso- carpets, according to the topographic conditions at a ciated with grasses or herbs or are predators. Some are particular site. Apart from helping to retain soil moisture diurnal, others nocturnal in habit. Clay banks, slopes, and and to prevent raindrops from directly hitting the soil flats, especially if partly denuded, are often found to be surface, leaf litter is a food source for different organisms pitted with perfectly circular holes leading to burrows. as it gradually breaks down, and shelter for others, be they These are seen throughout the year, and are inhabited by scavengers or predators. Leaf litter beetles are usually larvae of the native tiger beetle Neocicindela tuberculata. drab, and are mostly active at night. Many are flightless, The extremely agile adults are easily recognised by their small, and because hardly noticed are still waiting to be distinctly patterned elytra, but are around only during the described and named. The list of species occurring in leaf summer months. A second species of the same genus, N. litter is appreciable, made up of representatives of many spilleri, although dull in appearance is commonly and 1 nf ld tl S rv easily seen on the ground even in the darkest interior of the dung areAphodiuspseudolividus, Cercyon haemorrhoid- native bush. As more specimens of this species than of Ν alis, C. atricapillus, Neohypnus andinus, several species turberculata were found in swimming pools -3 m o Philonthus, and all five species o Anotylus found in the from the native vegetation, it must be assumed that it lives area. The Australian dung beetle Onthophagus granula- also in gardens, although never actually sighted there at tus, known to be in the country at least since 1 7 was y ie Several scarab beetles are common in the gar- seen only occasionally at y ie All these species are dens and fields of y ie among them the native capable of flight, and all are adventive. The fowl manure Costelytra zealandica and Odontria xanthosticta, the holds a greater variety of beetles than the sheep dung, South African Heteronychus arator, the Australian mainly because of the mix with straw, fe ed, and damp Aphodius tasmaniae, and the South American Ataenius areas around the water trough. Thus Sitophilus oryzae, picinus, all but the last species doing considerable harm to Gnatocerus cornutus, Aiphitobius diaperinus, and pastures. It is possible to observe a lot of activity on the Ahasverus advena were there primarily because of the ground, in the daytime mainly because of the presence of chicken feed. Three species reported for y ie were five Australian species, the rove beetles Thyreocephalus obtained only in the hen-house—the rove beetles Omalium orthodoxus and Astenus guttula, the ladybird Diomus allardi and Crataraea suturalis, and the cryptic beetle notescens, and the ant beetles Anthicus hesperi and a. Cryptophaguspilosus; the first two are reported here for kreusleri, and at night principally because of the five the first time from New Zealand. Other species associated roaming ca a i s two Clivina and three Lecanomerus with the e - ouse were the siaphylinids Oxytelus scu us species; four of these are Australian, the other is allegedly andPhilonthussordidus, the hydrophilidDactylosternum e a ic Damper sites are favoured by the presumed abdominale, the is e i s Carcinopspumilio, Gnathoncus endemic s a y i i Carpelimus zealandicus, the rotundatus, and G. schmidti, and the ant beetle Anthicus European C corticinus, and the Australian ca a i Tachys jioralis. All the species associated with the domestic fowl australis, and a further foreign ca a i species of uncer- at y ie are fully winged and foreign. Trox scaber, a tain provenance, Paratachys crypticola. Other beetles beetle frequently present in hen-houses in New Zealand, feed on grasses and herbs. Among the weevils associated was strangely enough not found at y ie with varied herbs are the South American Graphognathus (13 C rr n Conditions in an urban environment nor- the leucoloma, Listroderes difficilis, and L. foveatus, mally do not allow carrion species to prosper. Bait for . sulcatus, European Otiorhynchus rugosostriatus and O luring this special fauna was therefore set in an isolated and the Australian Steriphus diversipes lineatus. Among small grove o Acacia mearnsii and adjacent native bush. the weevils associated with specific plants are the European Only six species out of the hundreds of endemic Coleo- and Sit- Gymnetron pascuorum on Plantago lanceolata piera obtained at y ie are attracted to carrion: the Rhin- ona discoideus on Medicago species, the Australian eio i s Mesocolon alacre, Paracatops lugubris, and the oncus australis on Rumex and Polygonum species, and Pseudonemadus lituratus, the agy i 'Necrophilus' South American Listronotus bonariensis on various grasses and Linogeraeus urbanus specifically on the trailing grass Paspalum distichum. To the list of pasture beetles may be added the Australian a i i Euciodes suturalis on r 1-7 ( pp t p grasses, and the presumably Australian c ysome i Ι dd th h p nd tt r d tr f Que cus Longitarsus fuliginosus, which is common in mea owy Euca y us ia a us Ace Acacia i us C y ome ia grasslands at y ie nd Cu essus nd th h n r p n n th M n lf r (1 n Sheep and free-range hens provided abun- Sh p p dd nd th W r Str b h t t t rn dant, steadily available dung at Lynfield. The sheep flock nd h t ll rtr t th l ft r h t ( ac yca us consisted of around adult animals and, in spring and ac y ioi es on th r ht r x t summer, their lambs. The sheep were grazing in approxi- nd n th M n lf r fl n d b r p mately 3 7 hectares of pasture subdivided into five fenced ( y a a n t v b h th tr -f rn p pr v l nt paddocks, each with a water-trough and some trees. One W t rn l p f th W ttl b h h n r n r - field had an Acacia mearnsii grove adjacent to native t n f nd n p n pl nt t n f d d nd bush, which was a highly favoured site for resting and for d n ttl (Acacia ecu e s A mea sii nd h lth sheltering from sun and stormy weather. The fowl manure p n ( i us i as e a ia a came from two or three dozen hens kept in a yard and hen- 5 W ttl b h t th l r r f th tr house not far from the paddocks on the same property. The ό r p r r (Vitexlucens)in th W ttl b h h tr n e - ouse was lined with a thick layer of straw, and had ff t d b h t -r t pr v d p rt l rl ttr t v t th besides perches, nesting boxes, and troughs of feed and rh d d Ku eus a cua us th n b d a o egmus water. Sheep manure is not greatly different from well c owso i nd r phth l p f th broken down compost made from lawn clippings. The d n d n Mic oscy mus faunas in lawn compost and sheep dung are, therefore, 7 r n h f t n h r l r p n ( y oc a us ic o- rather similar, but myce o agous beetles are absent or ma oi es cu nd p nd d b t f r b tl W r rarer in dung because it dries out faster. Common in sheep Str b h (
(9
Ο (11
(1 (15 Intr d t r S t n prolongatus, the histerid Saprinus detritus, and the sta- that consists in the main of ferns, mosses, and liverworts. phylinid Stenomalium helmsi. The histerid was among the The two streams and the other trickles scattered about in very first New Zealand insects to be described, but Fabri- the small gullies are sheltered from direct sunlight by cius (1775) had listed it, in error, as an Australian species forest canopy. Clean, crumbly, and cracked banks devoid collected during Captain Cook's first voyage on the of vegetation have a few beetle species which are always `Endeavour'. All the common foreign carrion beetles al- there in numbers, some probably just hiding in the damp ready reported from New Zealand were present: Creo- cracks, others sheltering in the daytime. The two carrion philus oculatus, Philonthus politus, Necrobia ruficollis, beetles `Necrophilus' prolongatus and Colon hirtale and N. rufipes, Dermestes maculatus, and Omosita colon. the rove beetle Sepedophilus largulus, the latter usually (The large rove beetle Creophilus oculatus is an Austra- associated with the large shelf fungus Ganoderma appla- lian species, but was already in New Zealand at the time natum, occasionally come out of the crevices in startling of Captain Cook's visit in I769.) Five further species numbers when disturbed with repellent sprays. `N associated with carrion are reported here for the first time prolongatus was otherwise obtained only by using bait from New Zealand, two having been found first elsewhere appropriate for carrion beetles. Specimens of the in the country, the other three only at Lynfield. These are scarabaeids Saphobius edwardsi and Odontria xanthosticta Aleochara puberula, Gabronthus sulcifrons, Nitidula have occasionally emerged from the cracks too. O. carnaria, Omosita discoidea, and Litargus balteatus. A xanthosticta occurs the whole year round at Lynfield; it specimen of the European burying beetle Nicrophorus seems to like, particularly in the winter months, damp vespillo in the Brookes Collection (New Zealand Arthro- niches and coastal caves. Stream banks are frequented pod Collection, DSIR) was collected at St Heliers, Auck- also by the subglabrous Melanophthalma species 5, and land City, by A.T. Pycroft in 1928. As this beetle has not foremost by a ptilodactylid resembling closely turned up at Lynfield, and has not been collected in the 60 Byrrocryptus species but belonging to an undescribed years since it was picked up at St Heliers, the species is genus which is, in fact, widespread in New Zealand, and presumably not established in and around Auckland. has terrestrial instead of aquatic larvae. Common in the vegetation at the top of the banks and adjacent steep slopes ( 4 Stream banks. The lower part of the Wairaki Stream is the small cerambycid Spilotrogia elongata and the and the upper part of the Wattle Bay Stream have sand- tyrine pselaphid Zeatyrus lawsoni. Some of the rare stone banks and run on sandstone beds, but farther down species found at Lynfield were obtained only in this the Wattle Bay Stream has soil banks 0.5-1.5 m high as a vegetation, such as the ptiliid Actidium angulicolle, the result of carving through old landslides. Some sandstone staphylinids Agnosthaetus brouni, Hyperomma disper- banks, especially along the Wairaki Stream, are covered sum, and H. mandibulare, and the pselaphids Microtyrus in rather delicate vegetation, and the overhanging earth punctatus and Hamotulus mutandus. banks elsewhere have forest floor plants on top. The stream banks may be flanked by steep, sandy loam slopes, (15) Stream beds. Each of the only two permanent which are mostly seeping and covered in low vegetation streams has a bed approximately 300 m long by 1.5-4.0m wide covered by forest canopy. The flanking undergrowth is relatively sparse. The Wairaki Stream bed is sandstone with the odd small gravel patch. Sandstone bed surfaces are usually covered with a dense carpet of mosses made up r 8 - (opposite page) of Hypnodendron marginatum, Fissidens asplenioides, 8 Gr nd v r f nd r n j ( r d nt fl n n Thamnobryum pandum and Thuidium furfurosum at wet n r v f br ttl ll (S l x fr l W ttl areas and of Tridontium tasmanicum and Fissidens Wattle Bay sedge p (C r x S rp ph br n h rigidulus at the splash-zone or in the water. The Wattle f br h ttl (Alb z l ph nth t t p r ht Bay Stream bed is sandstone in its steeper upper course, 0 h W r Str nd t n b d xp d t n f t then gravel, and finally silt, since stormwater run-off from p l fl n d b r n t nd l hn a recent housing development above the northern heights M th f th W ttl tr t l t d th b nd f of Wattle Bay washed considerable amounts of soil into fl x ( h r t n x nd th M r p p r Alb z nd the bush. The stream ends up at a Carex swamp with S l x f rth r b Scirpus, Cyperus, Typha, Juncus, and naturalised Loni- 2 Small patch of halophilous plants, comprising Cotula cera japonica and a stand of Salix fragilis with a dense coronopifolia nt r p r d th S rp rn r l h n ground cover of Tradescantia fluminensis and some tr t nd S r rn n fl r W ttl Hedychium gardnerianum and Phormium tenax. As the W t d f W ttl h n t l b lt f l r Carex swamp prevented an easy outflow of flood waters, p h t tr (M tr d r x l the stream backed up above the swamp, depositing sand 4 A p t-tr p t t W ttl h d p p t v r d t and silt and raising the water-table at the lower 25 m or so pr v nt fl d n d r n h v r n of the bush stream by 60 cm. As a consequence several G rd n r f h p n p dd 20 fr th W r large trees of Vitex lucens, Dysoxylum spectabile, and Str b h h hl ttr t v t f r n b tl b t n t t Corynocarpus laevigatus and a number of Cyathea and n t v ρ n tree ferns became waterlogged and died within
1 nf ld tl S rv
3-5 yea s o e i g u e s eam e o i ec su ig esc i e i the sec io a o omic o es ( 7 wi e si e a o e s eam is a i y ei g i a e y e ame Myrtonymus zelandicus. It is e a e o a g ou Carex lessoniana a Cyperus eragrostis. e ee e o wee i s w ic a e associa e o ma y wi aqua ic a s ecies egu a y ou o e s eam e s a e a mos su aqua ic o se ai s e s a mo oco s u i ew e c usi e y ca a i s a s a y i i s e o me ami y ea a wi e es ia mosses O e suc moss wee i a i g e commo s ecies Notagonum lawsoni a as u e u i y ie a so a is ame Bryocatus Scopodes elaphroides a e ess commo Syllectus polytrychi i e a o omic sec io ume ous soi sam es anomalus a Pentagonica vittipennis. e s a y i i s ake om a ou oo e s o My aceae a o e ees o o e ee es a e e ese e y ee s ecies o Carpe- a o e the cou y i e as 15 yea s o ma e limus, s ecies 3 a s ecies o e u e e mi e Ge us is i u io o Myrtonymus zelandicus a e s ow o 1 a e Eu o ea Myllaena intermedia, e e e o e aces o o e e a e i s ecies e ge us Myrtonymus o e i s ime o ew ea a ; a ese i si o is o es ic e o ew ea a ; i a so occu s o e a ge amo gs mosses i e wo s a e s o a eas o sa - a eas o Quee s a ew Sou Wa es a Wes e s o e Mo e o ess co i e o s eam g a e is a u e Aus a ia As some o e Aus a ia s ecies a e c ose y Eu o ea o e ee e Aloconota sulcifrons, a es ic e e a e o M zelandicus, e ossi i i y a i oo is Aus - o wo mosses o e s as - o e Tridontium tasmanicum a ia ca o e u e ou and Fissidens rigidulus, is a e y agi e s ecies o e same (17 o e ee e ami y is e e e as Ge us s ecies 1 O eac sa e a e ock i e Wai ema a s ecime s o ma y o e a i e a o eig ee e s ecies o ma io o w ic e y ie a ea e o gs is sa - may e icke u o e s eam e s a icu a y s o y s o e a mu s o e Acco i g y the sea-s o e a o e a a e oo i gs e i the ig i e e e is i e sa u mi e wi a ia e amou s o s e a a i e ea i e a i - (1 Soi e soi s o y ie a e ia e oams om woo S e s o pipi (Paphies australis), ue musse s o g y wea e e sa s o e a si s o e o the owe (Mytilus edulis), aci ic oys e (Crassostrea gigas), a Mioce e Wai ema a o ma io ey a e im e ec y ock oys e (Saccostrea glomerata) ca accumu a e co - ai e o e ye ow- ow ea s o si oam e u e si e a y oca y was e u u i g the s o ms a i e - y ie soi s a e a ai amou o a ima i e i em; mi e y swee the Ma ukau a ou W e e sa s o e ea wo ms a wo ms ema o es Sym y a mea y- c i s a e o use y c acke a a ge ocks e o eak ugs cica a ym s s i g ai s a s a ee es Mos o the s o e is i e e wi ou e s e eac es om ee es i i g i o o i g i o soi o so i e a a s age Wa e ay o y ie Co e a e s e e e o a wa y o y e mai ee e g ou s wi soi -i a i i g a ae ig a ki g c i s a o seawa y e e si e mu - a s a y ie a e e ige ee e s ecies o Neocicindela, a o e a mos a ocke Ma ukau a ou wo s ecies e sca a aei s wi e e ce io o o e o wo a o ii es a e commo i e ay ime o e su ace o a e sa e e i o ac y i Ge us 1s ecies e1 e a e i s ecies o sma a ious y a e e eac ee e Phycosecis lim - Conoderus, the c ysome i s ecies o Peniticus a bata, a the ige ee e Neocicindela tuberculata. ese Eucolaspis, a e oa - ose wee i s ecies ee wo ee e s ecies we e o ice a icke u y e ee e s ecies occu i g i e y ie a ea a e w o y scie is s accom a yi g Ca ai Cook u i g is i s y ogea as a u a a a a a e e igme e a e - oyage o e E ea ou (17 9 a a e esc i e y ous a comρ e e y i O e is a s i u ame s ecies a icius (1775 17 1 u e ack amo g Pennisetum o e wi es ea s a y i i ge us Carpelimus, usua y clandestinum, Festuca arundinacea, Apium prosiratum, associa e wi am si y g ou Suc co i io s a e Haloragis erecta, Calystegia sepium, Atriplex prostrata, a e commo u e e e se ege a io o e a i e a a us a a is um ic is ec iοi es So c us as e us o y ie a a so a ou co o ies o u s o e uma ia mu a is a Veronica persica a e commo y a Mesoponera castanea. is a icu a a is k ow o ou e a k i g ee e Mimopeus elongatus, e a os se a i s ecies i some a eas o e cou y e ee e Anthicus s ecies 1wi a s iki g a e ye ow i Carpelimus s ecies a oug e a i e y commo e mi a a e a segme a e wee i s Cecyropa a easi y caug i ee i - a s was isco e e a discors, Mandalotus miricollis, a Mandalotus s ecies y ie a is k ow so e y om is oca i y e ce y i o ki g is e ee e Anommatus duodecimstri- atus, a o e i s ecies a as u e u a y ie A o e y ie isco e y is wo me io i g i is co - is Eu o ea i o igi I occu s s o a ica y i is u c ec io u i g a s u y o e e a ic a ea ic au as om a s o e g o e u ou e y ake o s i s wi soi o a u ia e osi s o e Waimea asi e so ( se e a o e a s o a as e eco om y ie is e i em i ii e ca a i s a e su e a ea seu osco io a a ioc o ius caeca us eie (197 we e o ai e om a i s a o y o e so a om ew ea a e i soi e o m o ca a i s we e co ec e a y ie u e ee e s ecies a so i s isco e e u i g e y ie same seu osco io s ecies as u e u wice i ee i - Su ey is a wee i a as si ce ee ou o e a a s om soi s a a e i e i commo wi ose o e wi es ea i coas a a eas a o is a s om Auck a Waimea ai s O e o e y ie s ecime s i c u e i e Ci y o wa s I has u e ou o e e sma es wee i o igi a esc i io as a a a y e o e s ecies came om a - us a k ow i e wo As i acke a ame i is cm- ee i a a se i e Wai aki S eam us
I o uc o y Sec io s 19
1. These endemic species are frequently mingled with the found to be present among them, and then only sporadi- naturalised weevils Listroderes foveatus, Listronotus cally so. Although the oxyteline rove beetles of wrack are bonariensis, and Steriphus diversipes lineatus. The wide- primarily land dwellers inhabiting cast-up algae, they are spread and, in open beaches, common beetles Thelyphassa occasionally found in numbers also in the intertidal zone. diaphana and Pericoptus truncatus associated with drift- By using the flotation collecting method a gravelly patch wood, and Cecyropa modesta and Lagrioida brouni asso- to the west of Wattle Bay was repeatedly checked during ciated with sand vegetation, are absent on the Manukau outgoing tides. The gravel was scooped into buckets of shores of Lynfield. The small driftwood weevil Macro- water and stirred. As soon as beetles that stayed in air rhyncolus littoralis was found only once. pockets under the gravel and boulders are disturbed, they come up to the surface, becoming very active. Of the three (18) Wrack and intertidal zone. Wrack is seaweed and wrack oxytelines listed above, only Teropalpus unicolor other marine algae cast ashore and accumulated against was found regularly in intertidal gravel, always in low the sandy beach at high tide level, or even further back in numbers not exceeding three per square metre. stormy weather. The intertidal zone extends over the shallow mudflats for the most part, and over a hard bottom (19) Bush streams. Only two streams have permanently of tilted sandstone and mudstone strata in the remaining running water, over a stretch of a few hundred metres. a eas The harder, less erodable sandstone layers stand out One, at the eastern end of the surveyed area, is 500 m long as ridges studded with oysters, mussels, and barnacles. and flows down the Wattle Bay valley. Its upper course is The splash zone is coated with algae, and here and there steep, with the water channelled over a sandstone bed. The are salt-tolerant plants such as Scirpus nodosus, Leptocar- watercourse suddenly flattens out, to continue partly on a pus similis, Apium prostratum, Lobelia anceps, Samolus gravelly bed but mostly on a muddy bed sheltered and repens, and two or three grass species. A small patch at shaded by a forest canopy of broadleaf trees and giant Wattle Bay has a community of halophilous plants con- tree-ferns until it opens up into a Carex lessoniana wet- sisting of Cotula coronopifolia, Scirpus cernuus, land. It then meanders through to a stand of the Palearctic Triglochin striatum, and Sarcocornia quinqueflora. Al- brittle willow (Salix fragilis) and then through Phormium though the mangrove Avicennia resinifera is common in tenax to the beach. The Wairaki Stream is situated at the parts of the Manukau Harbour, and fruits and seedlings western end of the surveyed area. Its various small tribu- often arrive in numbers with incoming tides on the Lynfield taries are piped, discharging into a 600-m-long stretch of shore, no plants become established beyond a few weeks. water that flows through native bush under a more open By far the main group of beetles represented in wrack is canopy than that at Wattle Bay. The upper 500 m of open the family Staphylinidae or rove beetles, with the oxytelines water flows on a sandstone bed with rills, slides, small Teropalpus coloratus, T. unicolor, and Blediotrogus gut- cascades, waterholes, and pools; the lower I00 m is tidal tiger, the omaliine Omaliomimus litoreus, and the and muddy. It is followed by a further I00 m stretch of staphylinines Cafius algophilus and C. quadriimpressus, piped waters discharging into Lynfield Cove. Apart from followed by the endemic feather-winged beetles Ptiliodes the two obvious streams, there are a few gullies and amplicollis and P. austerus, the European Ptenidium drainage lines with trickles of water at Wattle Bay, in the punctatum of the same family, and the European water Wairaki Stream bush, and elsewhere between Wattle Bay scavenger beetle Cercyon depressus, which was unknown and Lynfield Cove. No aquatic Dytiscidae or Hydrophili- from New Zealand until discovered during the Lynfield dae and no Elmidae are present in the bush waters of survey. The larvae of the Teropalpus species run to and fro Lynfield. Although the shady streams and trickles seem with the receding and advancing tidal ripples, behaving admirably suited to lowland hydraenid species, none was not unlike wrybills and plovers chasing outgoing waves found. This absence was somewhat unexpected, because and retreating from incoming surf on shores. Kelp dar- a rather common North Island species, Podaena latipalpis kling beetles of the genus Chaerodes, which seem to be Ordish (1984), occurs at Huia, only some 15 km SW of present on open shores at all times, are absent from the Lynfield on the same side of the Manukau. Elmidae and sheltered sands at Lynfield. A special effort was made to Hydraenidae require well oxygenated waters, which are secure certain intertidal beetle species at Lynfield, in case not available in the Lynfield area because the waters some were present in the particular conditions provided originate through seepage and percolation from headland by the estuarine waters of the Manukau Harbour. The hills, flowing too slowly and smoothly and for too short a trechine carabid Temnostega rangitotoensis, discovered distance to become sufficiently aerated. Two very simi a by A.E. Brookes in 1920 among plants of the eel-grass comb-waisted beetles (Ptilodactylidae) are present, both genus Zostera on Rangitoto Island in the Hauraki Gulf, of them chestnut-brown in colour. They are short-lived, and described by him a few years later (1932), was sought and stand out from other Lynfield beetles in having the without success. Several limnichid species of the genus hind margin of the prothorax distinctly crenulated, with Hyphalus occur from the Coromandel Peninsula to the far rounded off teeth. The smaller one, a species of Byrrocryp- north of Northland, apparently in association with com- tus, is 5.0-5.5 mm long and has a distinctly granulated munities of barnacles. The barnacle genera Chamaesipho pronotum and simple claws. It flies fr early February and Elminius are common at Lynfield, but only the inter- and has aquatic larvae. The other species belongs to a still tidal pseudoscorpion Maorichthonius mortenseni was undescribed genus; it is 5.5-6.5 mm long, has a relatively nf ld tl S rv smooth pronotum, and its claws have a more or less l t Apart from the bullrush ph r nt l the pond laminate swelling near the base. It flies from mid February itself contains El d n d n M r ph ll pr p n- to late March, and has terrestrial larvae in seeping soils. A Polygonum h dr p p r and Alt rn nth r ph l - further group of beetles associated with aquatic environ- x r d It supports a steady mixed population of one or ments are the marsh beetles (Helodidae). This family is two dozen mallard ducks (An pl t rh n h and grey represented by species at Lynfield. Adults visit flowers ducks (A. p r l ; these are sometimes joined by or rest on foliage, always quite close to water. The muscovy ducks (C r n h t The overflow from onisciform larvae of all the Lynfield species live in still the pond traverses the golf course into a channel to the waters among trapped plant debris. Waitemata Harbour, on the other side of the Auckland isthmus. Drinking troughs in nearby sheep paddocks and ( Ponds and troughs. As there is no pond anywhere domestic swimming pools proved to be attractive to some in the Lynfield study area, one nearby on the Maunga- aquatic beetles, too. The only two water beetles (Dytis- kiekie golf course was included in the survey. It is fed by cidae) collected at Lynfield were found in the pond and in a little stream that runs through a ph r nt l marsh, drinking troughs. The European species Rhantus t r l and is flanked on one side by C th d ll r C does occasionally get into chlorinated swimming pools, d Ιb t ρtο ρ r ορ r υ Μ Ι tυ r fΙοrυ where it was observed to stay for up t 3 or4 weeks if not tt p r n d M r n tr l C pr unduly disturbed. The only aquatic hydrophilid caught at robusta, G n t rupestre, and C th d f - Lynfield, En hr tr t is common in the pond.
RELATIVE BIOTIC DIVERSITY IN THE SURVEY AREA The Lynfield beetle survey has produced 753 endemic commutes annually between New Zealand and a number beetle species in the few years over which the systematic of Melanesian islands. These have been observed every collecting was undertaken. This number, although sub- year at Lynfield since the beetle survey started in 197 stantial and unexpectedly high, is by no means final. The paucity of native vertebrates in the Lynfield bush is Further species were still turning up at a rate of one or two compensated by a strong representation of introduced. per fortnight from litter and decayed wood samples during birds. The leaf litter is searched by blackbirds ( rd the months prior to ending the main field work. The r l thrushes ( ph l l and the occasional canopy of several tree species known to host special flock of Califomian quail ( ρhοrt x l f rn In the beetles — notably the crowns of r n p t f rr n canopy feed or perch greenfinches (Chl r hl r chaf- El rp d nt t Kn ht x l and t finches ( r n ll l b goldfinches (C rd l r- l n l t — could not be reached, and Malaise traps d l house sparrows ( r d t starlings could not be set long enough or high enough for rare and (St rn v l r mynas (A r d th r tr t and the elusive species to be eventually secured. It would be odd small flock of rosella parakeet ( l t r x reasonable to assume that the number of species thus far In the appendix to this report, Esler lists 1 species of obtained could be increased by up to 15% to a total of native phanerogams for Lynfield, which constitute roughly something like 5 endemic species. Beetle species gen- % of the or so gymnosperm and angiosperm erally amount to about one-quarter of the insect fauna on species of New Zealand and its outlying islands. My a site under normal conditions, which would mean that the estimate of the total native beetle fauna of New Zealand surveyed area at Lynfield ought to house approximately is around 1 to 1 5 species, so the Lynfield popu- 3 different indigenous insect species. lation would represent - 5% as for the plants. How- This area that holds literally thousands of invertebrate ever, the plant list includes a fair number of species that species has only nine native vertebrates, the copper skink occur naturally also outside New Zealand, whereas the (C l d n n and a bird population of seven resident figure for beetles excludes all the non-endemic species, and one migratory species. This latter comprises three or regardless of their mode of arrival. four pairs of the New Zealand pigeon ( ph n v - A number of Lynfield beetles were described from l nd two or three moreporks ( n x n v l n- New Zealand while the survey was in progress. Some of d a few tui ( r th d r n v l nd and these had their primary types drawn from specimens grey warblers (G r n t and moderate numbers collected at Lynfield. Thus, this small suburban area of of fantails ( h p d r f l n silvereyes ( t r p metropolitan Auckland is currently the type locality for l t r l and kingfishers ( l n n t Only in the species, and this number will undoubtedly increase once spring and summer months are there a few shining cuck- the 13 indigenous species as yet unnamed that occur at oos (Chr x l d a species that unfailingly Lynfield are described. i o uc o y Sec io s 2 OES E I IGE OUS AU A EASI Y A A O MO I IE E I O ME A CO I IO S? In order to decide whether the indigenous fauna is consti- were located in the Wairaki Stream bush and three in the tutionally able to overcome the challenge of environ- upper part of Wattle Bay valley. Now, 15 years later, the mental changes bought about by urban and rural develop- number of seedlings has increased manyfold in both ment, it is necessary first to have in mind the type of areas, owing presumably to the return of the native pigeon vegetation that was covering New Zealand in geological (Hemiphaga novaeseelandiae) to these sites. Pigeons like times, and then to consider apparent adaptations of the the small, brick-red fruit of nikau, digesting the pericarp native fauna to such changes. A few telling examples can and dropping the remainder with the seed. Most saplings be drawn from a simple survey such as that reported here, are sprouting under puriri (Vitex lucens), the tree these as follows. birds prefer to perch on. Only one palm plant has devel- oped a trunk as yet, and this has been flowering for nine (1) Geological background. It is generally believed or ten years. Part of the Lynfield area is of special interest that forests clothed the whole of New Zealand for more in that the missionary and botanist William Colenso than 100 million years before the Kaikoura Orogeny happened to visit the Wairaki Stream bush on 4 February gradually built up the present mountain chains, starting in 1842, on his way from Otahuhu to the Kaipara Harbour. the Pliocene (Fleming 1977). Mountain uplift interrupted A passage in his `Memoranda of an excursion' reads as the eastward-drifting cyclonic cycle of the weather, re- follows: "I ... discovered a shrub of a genus altogether ducing the rainfall on the leeward side of the ranges. Only unknown to me. This plant bears an oblong succulent a small part of the North Island, from Hawkes Bay to the crimson-coloured bacca [Latin for berry], containing Wairarapa, and nearly half the South Island, from Marlbor- several large angular and irregularly shaped seeds; its ough to Otago, were affected. The remainder of New growth is diffuse and slender with but few branches, and Zealand continued to be covered in forest except during its height is 5-9 feet. In habit alone it closely approaches the relatively short periods of the various Pleistocene to some species of the Coprosma genus. There were glaciations. As the northern half of the North Island was several of them here, on the immediate banks of a little hardly influenced by these events, forests persisted over rivulet [Wairaki Stream] which ran through this dell; I did all but the narrow littoral zones. Abundant, well distrib- not, however, observe it in any other locality" (Colenso uted rainfall and a lack of past climatic extremes have 1845). This particular plant species, afterwards recog- produced a rather uniform cover of evergreen forests with nised as Alseuosmia macrophylla, continues to thrive in very dense undergrowth, contrasting the conditions found the Wairaki Stream bush, where it is a rather showy and in the temperate forests of the Northern Hemisphere and important constituent of the undergrowth. the Magellanic forest of southern Chile. (3) Numbers of native species occurring in urban and (2) Historical background. As soon as the first Polyne- rural habitats. Out of the 753 indigenous beetle species sian people arrived, in the 11th century A or shortly collected at Lynfield, 10 or so are restricted to beach or before, the clearing of forests by fire began. At the time of wrack and 2 others to beach and open fields. The great arrival of the Europeans, at least half of the pristine forests majority of species are more at home in the native bush had disappeared. Burning off was done only partially by than anywhere else, with the possible exception of the the Maori, leaving a mosaic of native vegetation in place. branch-cutting longhorns Gastrosarus nigricollis and European settlers, however, tended to remove millable Astetholida lucida . Although 68 beetles, amounting to trees first and then clear everything else, not only from some 9% of the 744 non-littoral species, figure in the table arable flatlands but also from deep gullies and steep under the column `open fields of urban and rural habitats', hillsides. The native bush was frequently extirpated so the actual subsistence of some species of open environ- thoroughly that not a tree was left for miles on end in many ments was marginal or doubtful, and the endemic status of parts of the country. The Auckland isthmus has been others was uncertain. heavily populated by Maori and European alike since their respective arrival in New Zealand. It would be hard (4) The Pittosporum and Pseudopanax beetle fauna in to find a patch of indigenous forest that had not been gardens. Pseudopanax and Pittosporum species are heavily disturbed by milling, cutting, or burning in any known to be specific and highly attractive hosts to a whole span of 50 years or so. Useful trees such as matai (Prumno- series of beetle species that are agile on the wing. Both pitys spicata) and totara (Podocarpus totara) were almost plant genera are as common in gardens as in the bush, and certainly wiped out at Lynfield, and rimu (Dacrydium were repeatedly tested and checked at all times of the year cupressinum), another excellent timber species, is repre- for associated fauna. The test garden, on Tropicana Drive, sented by a single seedling at present. The absence of was only 225 m due north from the Wairaki Stream bush, mature nikau palm (Rhopalostylis sapida) in the Lynfield with 45 m of gardens and 180 m of paddocks bridging the area is rather puzzling. The palm is common elsewhere on gap. None of the six common foliage weevils of Pitto- the northern side of the Manukau Harbour. At the time the sporum and Pseudopanax were ever found on garden beetle survey commenced, in 1975, only six seedlings specimens, although some were occasionally seen on
nf ld tl S rv wi ow a es a c o es i es Cu a c es w ic (8) Ecological flexibility of indigenous and foreign o e e y a ac i e o se e wee i si o g o a species belonging to the same genus. e e a e 1 ou a k ee e s ecies i e us u i g e i s 1 ee e ge e a w ic a e a i e as we as adven ive weeks a e ei g se u a e ai e o yie a si gΙe s ecies a Lynfield, i o i g a oge e 55 s ecies A s ecime i e ga e a i e s ecies we e ou o y i the us ; a 33 o eig s ecies a e a ome i ga e s a ie s u 1 (5) The Coprosma beetle fauna in gardens. Coprosma o em occu qui e ea i y i a a i e us e i o me s ecies a e e e yw e e i e us a a e we e e- Muc e same si ua io is ou i a u o e s ecies o se e a so i ga e s is rubiaceous ge us as si os - Ptiliidae, a u o e s ecies o o -aqua ic Hydrophili- s eci ic ee e s ecies a a e commo i the Lynfield dae, a o -ca io Histeridae, a a entimine a us e o g o Hybolasius crista, e ea - ee e rhy irhinine Curculionidae — a i e s ecies occu so e y Pleuraltica cyanea, a ou s ecies o wee i s ee o i the us a o eig s ecies so e y o e omi a y em i e ge us ao e a a e o e ei g Tychanus i ga e s a ie s gibbus, e o y ig ess s ecies amo g e si o e Lynfield ge e a s a i g a i e a adven ive s ecies ee es we e ou o ga e Coprosma. wi eco ogica y co as i g e e e ces a e as o ows e es e am es a e Anabaxis, Sphaerothorax, Cono- (6) The Phormium and Cordyline beetle fauna in derus, a A isome is es gardens. A i g ou o a s wo y o me io com- CARABIDAE ises ee s ecies o Agavaceae w ic a e o u a wi Tachys — 1 a i e adventive ga e e s e ca age ee (Cordyline australis) a e Notagonum — 1 a i e adventive ( a so i us wo s ecies o ew ea a a Phormium tenax a Ρ STAPHYLINIDAE cookianum. e ca age ee su o s wo u y wi ge Lithocharis — a i e 1 adventive cossonine wee i s Eucossonus setiger a Tanysoma Atheta — a i e 9 adventive ( a so i us comatum. e a s ecies a e the u y wi ge cossonine Myllaena — 1 a i e 1 adven ive ( is a so i us Phloeophagosoma pedatum, e ig ess cossonines Oligota — 3 a i e 1 adven ive ( is a so i us Novitas dispar a Microtribus huttoni, a ee ig - PSELAPHIDAE ess lamiine o g o s Mic o amia ygmaea Psilocnaeia Anabaxis — 1 a i e 1 adventive parvula, and asteliae. O ce mo e o e o e i e CLAMBIDAE ee e s ecies w ic a e so commo u e a u a co i- Sphaerothorax — a i e 1 adventive io s as s ow u o ga e a s SCARABAEIDAE Ataenius — 1 a i e 1 adventive ( is a so i us (7 The pollen-feeding diurnal species of weevils and Saprosites — 1 a i e 1 adven ive ( is a so i us longhorns in gardens. e e a e e s ecies o eug om- ELATERIDAE me wee i s i e us w ic ee i isc imi a e y o e Conoderus — ve i adven3 a i e o e o a wi e a ge o owe i g a s A oug a A O AE e ce e ye s o e was ou o a i e o e o ic a s a o egmus — 1 a i e 1 adven ive i e ga e s O y ou ou o e i ee s ecies o NITIDULIDAE cerambycine o g o s occu i g a Lynfield a e e Epuraea — a i e adventive i e y ace e eyes c a ac e is ic o iu a a i s A CUCUJIDAE ou ca e see i ga e s Coptomma variegatum a C y amo a — 1 a i e 1 adventive (a so i us Gastrosarus nigricollis a e y Zorion mi u um a CORYLOPmDAE Navomorpha sulcata o o y a e eque y u a so i Anisomeristes — a i e 3 adven ive ( a so i us um e s o Ligustrum si e se Leptospermum scopar- ANTHICIDAE ium, a a ious wee s ecies o Rubus. Ant icus — 1 a i e adventive ( a so i us
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: b n (4) A omo a us i awa ae (Mac eay Ι AU W U BE C A omo a us a iega us Moo e Ι AU W CC U O GR eme i a asu a W i e Ε SS U CA i o oeus ucu e us (Newman) Ι AU W C U O CA : nt n n (2) e ago ica i i e is C au oi Ι AU W CC EU O CR Sco o es e a oi es (W i e Ε CC EU CR Y ISCI AE (2 y y us e ega s (Mo ou ie Ι AU W CC O CR a us su u a is (Mac eay Ι PA W CC OP CR I II AE ( 7 Ac o ic is i co s icua (Matthews) Ι AU(? W W W W W W W C O CR Ac o ic is mo a o i (A i e Ι PA SS O CR Ac o ic is su cog a a o so Ι NA C O CR Ac i ium a gu ico e o so E EU CR Ac i ium i ea e Ma ews Ε ' W SS E C Cissi ium o eo a um o so E CC U CR i e ium e a icum o so ' Ε C EU CR Kusc e i ium mao i o so E W EU CR e osa a g a is o so E W U CR ei osa a i e me ia o so E W SS U CR e osa a mi ima o so , Ε CC EU CR e a es i a ( ewma I A C O CR o o e i ium c assum o so E W C EU CR o o e i ium awso i (Matthews) Ε W C EU CR o o e i ium o o gum o so E W U CR O ige a o eo a a (Allibert) Ι PA C O CR e i ium ae iga um E ic so Ι PA W CC O CR e i ium u c a um (Cy e a Ι PA C O CR e i ium usi um (Cy e a Ι PA W CC O CR i io es am ico is o so E W ΕΕ C i io es aus e us o so E W CC ΕΕ C i e a acaciae o so E C EU CR i e a ca e i ( ou , Ε W EU CR i e ia co usa o so " E W SS U CR i e ia e ugi ea o so E . W C U ' CR
EGE r v n n E nd ; , Lynfield t p l l t ; I ntr d d f r n; AU A tr l n; A l r d ; Ν rth A r n; SA S th A r n; ΕΤ Eth p n; IM Indo-Malayan FR: f r t r p rt fr l nd W n ( nd t n W n d; fl htl ; W n d nd fl htl nd v d l Ab nd n SS n l p n; v r r r ( r 3 ; r r ( -1 ; CC n (11-5 ; C v r n ( r th n 5 Habitat: BU, b h; O p n; ΒΕ b h; G r nd; CA n p a u a Summa ies o S ecies a a 2
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SCA I II AE ( aeoce a scu e a is (Redtenbacher) Ε U C Ϊ ac y o us a us ( ou Ε C EU G Sca isoma ac uosum ou , Ε CC U C Sca isoma s e a e ou Ε EU C Sca isoma s ecies Ε EU G Sca isoma s ecies 2 Ε EU C Ge us s ecies Ε CC EU C
S A Y I I AE : r t n n (2 eso eus acu ice s (Ee aue Ε CC EU C Sii o eius i i us Β οu Ε C EU G : O l n (13) ou ie um i a um (Β οu Ε , EU CA Isc o e us ge a is ( ou Ε EU CA Isc o e us ec us (Β οu Ε C EU CA Mic osi a s Ε EU C Oma iomimus i o eus ( ou Ε ΕΕ C Oma ium a ia i ai mai e & isou Ι A SS O C a a oeos i a gay a e sis (Mac eay I AU C O C S e oma ium co ie i e aue Ε EU ΕΕ C S e oma ium e msi (Cameron) Ε EU ΕΕ C S e oma ium mo i i e um ( ou - Ε CC EU CA S e oma ium i o i ( ou Ε SS EU CA
26 nf ld tl S rv
x n (and no. of species) r v n n W n Ab nd n b t t S e oma ium s a i ( ou Ε C EU G S e oma ium su ci o a ( ou Ε CC EU G : Ox t l n (14) A o y us u ei e is (Mac eay Ι AU C O G A o y us com a a us (E ic so Ι PA W C O G A o y us semi u us ( au e Ι AU W C O ΕΕ C A o y us i so i (Cameron) Ι AU C EU O G A o y us s ecies Ι AU W C O G e io ogus gu ige S a Ε W SS ΒΕ G Ca e imus i i ea us (Stephens) Ι PA W O ΒΕ G Ca e imus co ici us (G a e o s Ι PA W C EU O ΕΕ C Ca e imus usi us (G a e o s Ι PA SS EU G Ca e imus ea a icus (S a Ε C EU O G Ca e imus s ecies Ε CC EU G O y e us scu us G a e o s Ι PA O G e o a us co o a us (S a Ε C ΒΕ G e o a us u ico o (S a Ε W CC ΒΕ G : O r n (1) a a o c us e i e is (Β οu Ε CC U C : El n r (1) eo eusis i gu a (Fauvel) . Ε EU CA : d r n (11) As e us gu u a au e Ι AU C O C y e omma is e sum ou Ε U G y e omma ma i u a e Β οu Ε EU G y ome o e i ico is ( o as o Ι PA CC O C i oc a is ig ice s K aa Ι PA CC O G i oc a is e a is (Β οu Ε EU G i oc a is i is (K aa Ι PA CC O G o a ium i a i um ( au e Ι AU O G Me o ee a icus e e ac e Ε EU C Scim a ium ae um ack u Ι AU W O G Su ius o i quus ( isou Ι PA O G : St ph l n n (19) Ca ius a go i us (Β οu Ε CC ΒΕ C Ca ius qua iim essus (W i e Ε C ΒΕ G C eo i us ocu a us ( a icius Ι AU CC O ΒΕ G Ga ius ig i u us (Gravenhorst) Ι PA W ΕΕ G Ga o us su ci o s (S a I A CC O G Gy o y us ac ico is (Mue e Ι PA O G e aci us usi us (Stephens) Ι PA O ΕΕ G eo y us a i us ( au e Ι SA CC O G eo a o i us ou i (Sharp) Ε EU CA o o i us socius ( au e Ι AU O G ac yco y us u i es ( ou Ε U G a aco y us a ecae ( ou Ε EU CA i o us o gico is S e e s Ι PA CC O G i o us a cus S a I A CC O G i o us o i us (Linnaeus) Ι PA W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W CC O C i o us y o e us K aa Ι SA CC O C i o us ec a gu us (Sharp) Ι PA C O G i o us so i us (G a e o s I A C O C y eoce a us o o o us (O i Ι AU CC O G : Ε th t n (2) Ag os ae us ou i e aue Ε EU C E a us ec e esi ei e Ι PA SS O C : h p r n (12) Se e o i us ace us (Β οu Ε EU C CA Se e o i us a ica i ius (Β οu Ε C EU CA Se e o i us au icomus (Β οu Ε C EU C CA Se e o i us aus e us (Β οu Ε C EU C Se e o i us e ico is ( ou Ε EU C Se e o i us ia i o a (Β οu Ε C U CA Se e o i us ae u us ( Β οu Ε U C Se e o i us a gu us ( ou Ε . CC EU C CA Se e o i us macu osus ( ou Ε U CA Se e o i us u i us (Broun) Ε EU C Se e o i us s ecies Ε EU C ac y o us i i u us ( a icius I A ‚ W CC O C : Al h r n (70) A eoc a a u e uia (Kiug Ι (a CC O C A oco o a su ci o s (Stephens) Ι PA C EU C A isc a a a is (Cravenhorst) Ι PA CC O C Amisc a eci ie s (S a Ι PA SS O C
LEGEND Provenance: E. endemic; nf ld type locality; I, introduced, foreign; AU, Australian; PA, l r t ; NA, North American; SA, South American; ΕΤ Ethiopian; IM Ind -Malayan FR: first report from New Zealand Wing (condition): WD, w ng d; FL, flightless; WD FL, winged and flightless individuals Abundance: SS, single specimen; , very rare (2 or 3); RR, rare (4-10); CC, common (i1-50); VC, very common (more than 50) Habitat: BU, bush; OP. open; ΒΕ beach; GR, ground; CA, canopy. b l r S r f Sp t 2
x n (a o o s ecies r v n n W n Ab nd n b t t
A y o us g aci is S a Ε W U G A y o us o osus ( ou E W U G A e a amicu a (S e e s Ι Ρa W CC O G A e a a ice s ( ou Ε W CC U G A e a co ia ia (K aa Ι Ρa W O G A e a co ie i Came o Ι AU W U O E G A e a o i u a au e I AU W CC U O G A e a ea a ica Came o Ι ΕΤ W C O E G A e a s ecies - Ι Ρa W CC U O G A e a s ecies 2 Ε W U G A e a s ecies I AU W ΟΡ G A e a s ecies 4 Ι AU W SS ΟΡ G A e a s ecies I AU W U G Aus oca ea ookesi (Came o Ι AU W CC O G o oma a u ca ica ( ou Ε W CC U G o oma a s ecies Ε W CC U G o oma a s ecies 2 Ε W SS U G C a a aea su u a is (Ma e eim I Ρa W O G ig ammus mi ico is ( au e Ε W U CA E ce a us a u us ( ou Ε W CC U G a ag ia co ci a (E ic so Ι Ρ W SS ΟΡ G Gy o o us u i e is ( ou Ε W CC E G Gy o ae a o igo i a Came o Ε W CC U G Gy o ae a u c a a ou Ε W U G Gy o ae a s ecies Ε W CC U G Gy o ae a s ecies 2 E W CC U G Gy o ae a s ecies Ε W CC U G Gy o ae a s ecies 4 Ε W CC U G Gy o ae a s ecies Ε W U G Gy o ae a s ecies 6 Ε W SS U G a o ec a a i es ( omso Ι Ρa W E G Isc og ossa i u e cu a a ( ou E W SS U G Isc og ossa ec i a a Came o Ε W U G My ae a i e me ia (E ic so I Ρa W CC U O G My ae a mag ico is (Came o Ε U G e emi o ia so i a (Ma s am Ι Ρ W CC ΟΡ G Oca ea uscico is ( ou Ε W C U G Oca ea socia is ( ou Ε W CC U CA O igo a u gico a Wi iams Έ W SS U CA O igo a a a K aa I Ρa W CC U O G O igo a se ige a Wi iams Ε W U CA O igo a s ecu ico is (Came o E W U G a a y o us s ecies Ε W U G o y o us s e a is ( ou Ε W C U G Si usa a a e a ( e aue E W U G Si usa u e ( ou Ε W U G Si usa s ecies Ε W SS U G Si usa s ecies 2 Ε W U G S e omas a e a a Came o Ε W SS U G S e o o a e sico o ( ou Ε W U G S e o o a ea a ica (Came o E W U G Sy us ae a ius ( ou Ε W U CA Sy us a esce s ( ou Ε W U CA Sy us g a i e ( ou Ε W CC U G Sy us s ecies Ε W U CA amia aea co ige a ( ou Ε W C U G Ge us s ecies Ι AU W CC O G Ge us s ecies 2 Ι AU W SS U G Ge us s ecies Ε W U G Ge us s ecies 4 Ε W U G Ge us s ecies Ι AU W CC ΟΡ G Ge us 2 s ecies Ε W U G Ge us s ecies Ε W U G Ge us 4 s ecies Ε W CC U G Ge us s ecies Ε W SS ΒΕ G Ge us 6 s ecies Ε W CC U G SE A I AE : r n n ( 4 E ei a a e igma ou Ε W U G E ei a a a icu a ( ou Ε W SS U G Sago a u ica a ou Ε W CC U G Sago a e ca a a ou Ε W SS U G Sago a ge a is ou Ε W U G Sago a ami a a ou Ε W C U G Sago a o gi e is ou Ε W CC U G Sago a ma o S a Ε W SS U G Sago a mo ico a ou Ε W U G Sago a o a i is ou Ε W C U G Sago a s ecies Ε W SS U G
n f ld tl S rv
x n (and no. of species) r v n n FR W n Ab nd n b t t
Sago a s ecies 2 Ε ΒΕ G Sago a s ecies Ε SS EU CA Sago a s ecies 4 Ε EU G : E pl t n (24 A o ec us c a ige (Β οu Ε SS EU CR A o ec us s ecies Ε SS EU G a o ec υs sρecies 2 Ε EU G E eusoma us ai oce a us ( ou Ε CC EU CR E eusoma us s ecies Ε SS EU G Eu ec o sis a iqua ( ou Ε EU CR Eu ec o sis o aca (S a Ε EU CR Eu ec o sis o ico is (Β οu Ε EU G Eu ec o sis u e ige a (Β οu Ε CC EU CR Eu ec o sis s ecies Ε C EU CR e o ec us s ecies Ε CC EU CR e o ec us s ecies 2 Ε EU CR Mic o y us i acu us ou Ε SS EU CR Mic o y us u c a us ou Ε SS EU G Mic o y us s ecies Ε SS EU CR a a ec us s ecies . Ε EU CR i ορsis s ecies Ι AU CC O CR ec omo us scu u a us (Β οu Ε CC EU CR ec omo us isu cico is( ou Ε EU CR W i ea s ecies Ε SS EU CR e a ius c e e o e sis( ou Ε CC EU CR e a ius o scu us (Β οu Ε EU CR e a ius sa age i (Β οu Ε CC EU CR CR Ge us s ecies . Ε CC EU : r h l t n ( A a a is e ec ica (Ki g Ι AU SS O CA A a a is o eo a a ( ou Ε EU CR Eu i es ai u a (Β οu Ε CC U CR Eu i es g a a a ( Β οu Ε CC EU CR CR Eu i es g a a (S a Ε . EU Eu i es im a (S a Ε C EU CR Eu i es u c a a ( ou Ε CC EU CR Eu i es s ecies Ε EU CR Eu i es s ecies 2 Ε EU CR Gas o o us a omi a is (Β οu Ε EU CR Gas o o us s a i (Β οu Ε CC EU CR yso ya is i a a (S a Ε SS EU CR S a es scu u a :a E ou Ε EU CR : r n ( Ge ai us u c i e is Sc au uss I AU CC EU CR amo u us mu a us (S a Ε EU CR ea y us awso i S a Ε CC EU CR : l ph n (4 W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W W se a us ci imus ( ou Ε U CR se a us e ica us ou Ε CC EU CR se a us uicis Β οu Ε SS EU CR se a us au e S a Ε CC EU CR HYDROPHILIDAE ( 2 A o o us ai u us ( ou Ε C EU CR Ce cyo a aiis ( ayku Ι PA CC O ΒΕ C OPCe cyo a ica i ius (Ma s am Ι PA C CR Ce cyo e essus S e e s Ι PA CC ΒΕ C OPCe cyo aemo oi a is ( a icius Ι PA C GR Cy oma iawso a S a Ε CC EU CR OP ac y os e um a omi a e ( a icius Ι SA C CR OP ac y os e um ma gi a e (S a Ι AU SS CR OPE oc us i us (Β οu Ι AU CR EUE y us a ico is ( ou Ε CC O CR y os yg us o a is ( ou Ε CC EU CR ygmo us mo es us Wh t Ε CC EU CR
HISTERIDAE (8 OPAc i us ig ico is ( o ma Ι PA CR OPCa ci o s umi io (E ic so Ι PA C CR .G a o cus o u a us (Kuge a Ι PA CC O CR OPG a o cus sc mi i ei e Ι PA CC . CR a e ie us a oga us ( ou Ε SS U CR a e ie us u us ( Β οu Ε C EU CR OP a ysoma akewe i Ma seu Ι AU C CR
EGE r v n n E, nd ; *, nf ld t p l l t ; I ntr d d f r n; AU A tr l n; A l r t ; Ν rth A r n; SA S th A r n; ΕΤ Eth p n; IM, Ind -M l n FR: f r t r p rt &om l nd W n ( nd t n : WD, n d; fl htl ; WD n d nd fl htl nd v d l Ab nd n SS n l p n; VR, v r r r (2 r 3); r r (4-10); CC n (11-SΟ ; C v r n ( r th n 50) bIt t: BU, b h; O p n; ΒΕ b h; GR, r nd; CA n p ,
b l r S r f Sp t 2
x n (a o ο s ecies r v n n W n Ab nd n b t t
Sa i us e i us ( a icius E W CC U O ΒΕ G C AM I AE ( C am us omes icus ou I AU W CC U O G S ae o o a kusc e i E ö y-You ga Ε W U G S ae o o a su usus ( ou Ε W U G S ae o o a ie e sis ( ack u I AU W O G E O I AE ( Cy a us u c a us S a Ε W U CA Cy o ge a is S a Ε W SS U CA Cy o g a ige S a Ε W U CA Cy o u o i S a Ε W CC U CA Cy o osci a s S a Ε W U CA Cy o a ice s S a E W C U CA Cy o emo us ou Ε W CC U CA Cy o su usus S a Ε W U CA Cy o i i i e is ou Ε W CC U CA Cy o waika oe sis ou Ε W U CA Cy o ea a icus S a Ε W U CA Cy o s ecies Ε W CC U CA Cy o s ecies 2 Ε W U CA Cy o s ecies Ε W U CA Cy o s ecies 4 Ε W CC U CA Cy o s ecies E W CC U CA Cy o s ecies 6 Ε W U CA Cy o s ecies Ε W SS U CA Cy o s ecies 8 Ε W SS U CA Cy o ius i i us S a E W SS U CA Ge us s ecies Ε W CC U CA Ge us 2 s ecies Ε W SS U CA UCA I AE ( Ce a og a us i o a us ( a y Ε W C U CA Ce a og a us a ia us (Wes woo E W C U CA
SCA A AEI AE : Aph d n ( A o ius g a a ius ( i aeus I Ρa W ΟΡ G A o ius seu o i i us ( a asa Ι SA W CC ΟΡ G A o ius asma iae o e Ι AU W C O G A ae ius ou i (S a Ε W U G A ae ius ici us a o Ι SA W CC U O G Sa osi es commu is ( ou Ε W CC U G Sa osi es me a ( ack u Ι AU W CC U O G : S r b n ( O o agus g a u a us o ema I AU W ΟΡ G Sa o ius e wa si S a Ε U G Sa o ius squamu osus ou Ε C U G
: M l l nth n (6 Cos e y a mac o u ea Gi e Ε W ΟΡ G Cos e y a ea a ica (W i e Ε W CC U O G O o ia a os ic a W i e Ε W CC U O G y o o a ae a ( a icius Ε W CC U CA Se icos i us cos e a ( ou Ε W U CA S e as is o gico is (A ow Ε W CC U G CA
: n t n ( e e o yc us a a o ( a icius Ι ΕΤ W CC ΟΡ G Y I AE (1 Sy o us o u us ( ou Ε U G Sy o us se a ius ( ou E CC U G U ESΤI AE (1 Mao a ia e emi a (W i e E W U CA I O AC Y I AE ( y oc y us s ecies Ε W U G Ge us s ecies Ε W CC U G E A E I AE ( Ag o us mo es us S a Ε W SS U CA Ag y us a ia i is (Ca è e I AU W C O G CA Am i a ys awso i S a Ε C U G CA Co o e us e su (S a Ι AU W C O G CA Co o e us ma i imus ( ou Ε W CC U G CA Co o e us os icus (Esc sc o I SA W CC ΟΡ G Co o e us su ma mo a us (Mac eay I AU W O CA C e ice a a i o a (Ca è e Ε W SS U CA C e ice a s a gu a a (W i e Ε W U G CA omemus e ega s S a Ε W U CA omemus a i es S a Ε W U CA
3 nf ld tl S rv
x n (a o o s ecies r v n n W n Ab nd n b t t omemus uscico is ou E W U CA omemus a e is iga us (W i e E W SS U CA omemus simi is S a E W C U CA omemus su usus S a Ε W SS U CA Me a a ci c ige (W i e Ε W CC U CA Me a a a o ima s (W i e Ε W U CA Oc os e us ea a icus (W i e Ε W CC U CA a s oeus gu a us S a E W U CA a i us i osus (S a Ε W SS U CA o amus e a us ou Ε W SS U CA o amus wake ie i S a Ε W SS U G EUC EMI AE (2) omaeo us ige us (W i e Ε W SS U CA a e a is a s S a Ε W U CA YCI AE ( 1 o os oma u i e e ( a icius I AU W CC O CA CA A I AE ( 3 Asi is ug a ei Wi me Ε W U CA Asi is i i e e ( ou Ε W U CA Ma o es umi us ( e isso I Ρa W SS ΒΕ G E MES I AE ( A e oce us aus a is ( o e I AU W ΟΡ CA A e us e asci ( i aeus I Ρa W ΟΡ CA e mes es aemo oi a is Kues e I Ρa W ΟΡ G e mes es macu a us eGee I Ρa W CC O G ic e o es u ga a ( ou E W CC U CA ogo e ma g a u a um ou E W SS U CA ogo e ma maes um ou Ε W U CA ogo e ma sig a um S a Ε W CC U O CA ΝΟΒΙΙ Ε ( A o ium u c a um ( eGee Ι Ρa W C ΟΡ CA o ca oma i osa (W i e Ε W U CA a o egmus aus a ie sis ic I AU W CC U O CA a o egmus c owso i Es año Ε W U CA o co ius wa i Es año E? W U O CA Me emus g isei i us ( ou Ε W U CA Mic os e omo us o o gus ( ou Ε W U CA e oce a se icea ( ou Ε W U CA I I AE (1 i us s eciosus ou E W U G CA OGOSSI I AE (2 e i o e y ou i ( ascoe Ε W CC U G CA e i o e y ig os a sa (W i e E W SS U CA E I AE ( Aus a io es es i us ( ou Ε W C U CA e o e um s ecies Ε CC U G e o e um s ecies 2 Ε U G e o e um s ecies Ε U G e o ium s ecies Ε W U G e o ium s ecies 2 Ε SS U G Ge us s ecies Ε U G Ge us 2 s ecies E W C U CA C E I AE (5) ec o ia u ico is ( a icius I Ρa W C O G ec o ia u i es ( eGee I Ρa W CC O G a mius o gi es S a Ε W SS U CA au is a e a S a Ε U CA yma o aea o i oi es ascoe E W CC U CA YCOSECI AE (1 ycosecis im a a ( a icius Ε CC ΒΕ G ME Y I AE (7) asy es a ice s ou Ε W SS U CA asy es a i o s ( ou Ε W C U CA asy es s ewa i ou Ε W U CA asy es su cya eus ou E W SS U CA asy es s ecies Ε W U G
LEGEND Provenance: E nd ; *, nf ld t p l l t ; I, introduced, f r n; AU A tr l n; Ρ l r t ; Ν rth A r n; SA S th A r n; ΕΤ Eth p n; IΜ Ind -M l n f r t report fr Zealand Wing (condition): W winged; fl htl ; W winged nd fl htl individuals Abundance: SS n l p n; v r r r ( r 3 ; RR, r r ( -10); CC n (11-5 ; VC, v r n ( r th n 5 Habitat: BU, b h; O p n; ΒΕ b ach; GR, r nd; CA n p a u a Summa ies o S ecies a a 31
Taxon (and no. of species) r v n n FR W n Ab nd n b t t
a y es ig esce s ou Ε W C U CA a y es semi i u us ou Ε W W W W U CA I I U I AE (15) Ca o i us a i so i o so Ι AU W O G Ca o i us imi ia us ( a icius Ι A W O G Ca o i us ga e i o so Ι AU C O C Ca o i us emi e us ( i aeus Ι A W C O G Ca o i us ma gi e us Mo sc u sky Ι A W C O C E i aea a a c ica (W i e Ε W U C E u aea im e ia is ( ei e Ι AU W C O G E u aea scu e a is ( ou Ι AU W C O C E u aea sig a a ou Ε W U O ome u aea amoe a ( ou Ε W U CA i i u a ca a ia (Schaller) Ι A W C O G Omosi a co o ( i aeus Ι A W O C Omosi a iscoi ea ( a icius Ι A W O C a i i ia as e e ia ou Ε CC U CA So o ia ys i S a Ε CC U CA a yc o es aus a is ack u Ι AU SS O C I O AGI AE (4 Mo o oma o gico is Cy e a Ι A CC O C Mo o oma ici es e s Ι A W W CC O C Mo o oma s i ico is au é Ι A W C O C Mo o oma es acea Mo sc u sky Ι A SS O C CUCU I AE ( 3 A as e us a e a (Wa Ι A W CC O C C y amo a e ico is (W i e Ε W CC U CA C y amo a es a i si (Gué i Ι AU C U O CA A AC I AE (1) a ac us u i ο mis igo ico a om so Ι AU C O CA C Y O AGI AE (13 A oma ia ewisi ei e Ι A W C O C C y o agus i osus Gy e a Ι A W O C E is emus g o u us ( ayku Ι A W C U O C Mic am i a iscoi ea ( ou Ε W U CA Mic am i a e msi ei e Ε W SS U C CA Mic am i a i sig is ei e Ε W C U O CA Mic am i a u i a ( ou Ε W C U CA Mic am i a si a a ( ou Ε W U CA Mic am i a umi a ( ou Ε W U CA Mic am i a s ecies Ε W CC U C Mic am i a s ecies 2 Ε SS U CA Sa ius u ice s ( ou Ε CC U C Ge us s ecies Ε C U O CA A GU II AE (4) Ca a oc y us macu osus ( ou Ε W U CA a a i s o i us (S a Ε W C U CA o e us e essus (S a Ε W O CA o e us i e s (S a Ε U G E O Y I AE ( C y o ac e ou i ( ascoe Ε C U C C y o ac e sy e ica S a Ε CC U C CA CE Y I AE (2 A omma us uo ecims ia us (Mueller) Ι A U C y o ac e u i es ( ei e Ε C U G CO Y O I AE ( A isome is es a ica is ( ea Ι AU W O C A isome is es a e Ma ews Ε W C U C CA A isome is es s a i Ma ews Ε W C U C A isome is es o acicus (E ic so Ι AU W C U O C CA A isome is es s ecies Ι AU W C U O C CA Co y o us s ecies Ε W U G o o sis a i a ou Ε C U G o o sis o u a a Β οu Ε CC U C o o sis s ecies Ε CC U G o o sis s ecies 2 Ε C U G CA o o sis s ecies Ε C U C o o sis s ecies 4 Ε C EU C CA O o e us aequa is S a Ι A W O C O o e us a oma ius ( ee I A W CC U O C O o e us s ecies Ι A W SS EU C Saci a o o ga ou Ε W C EU CA Sacium u c e um ( ea Ι AU CC O CA
2 nf ld tl S rv
x n (a o o s ecies r v n n W n Ab nd n b t t Ge us s ecies E W U CA Ge us s ecies 2 Ε W U CA Ge us 2 s ecies Ε CC U G
COCCI E I AE ( 27) A a ia i u c a a ( i aeus Ι Ρa W CC O CA A o e us a i i us ( ou Ε CC ΒU G Cocci e a u ecim u c a a i aeus I Ρa W CC O CA Coe o o a i aequa is ( a icius I AU W O CA C y o aemus mo ou ie i Mu sa I AU W ΟΡ CA iomus o esce s ( ack u I AU W O G iomus s ecies Ι AU W O CA a mus c a y eus ( ois u a I AU W C U O CA a mo ia co o mis ( ois u a Ι AU W CC O CA I eis ga u a (Mu sa Ι AU W CC O CA Mi us ygmaeus ack u Ι AU W CC O CA y o ius acce us ( ou Ε W CC U CA y o ius co so s ( ou Ε W CC U CA y o ius agus ( ou Ι AU W CC O CA y o ius o es ie i (Mu sa Ι AU W CC O CA y o ius mi u u us ( ou Ε C U G CA y o ius a us ( ou Ε C U G y o ius su usus ( ou E CC U G y o ius is is ( ou Ε W U CA y o ius e a is (E ic so Ι AU W SS O CA y o ius s ecies Ε CC U G y o ius s ecies 2 Ε U G y o ius s ecies Ε U G Scym us oewi (Mu sa Ι Νa W CC O CA S e o us i i us Ka u Ε W O CA S e o us g iseus C a eau Ε W SS ΟΡ CA S e o us is io C a eau I AU W ΟΡ CA A I II AE (3 ) A i ius i ascia us ( ei e Ι AU W CC U O G A i ius cos a us (E ic so I AU W C U O G A i ius o i e (Wes woo Ι AU W C U O G CA ica a i us is ( ei e E W CC U CA ica a a u i e is ( ou Ε W CC U G ica a e ico a ( ou Ε W CC U CA ica a a iega a ( ou Ε W C U CA Co ica ia e o ga a (Gy e a I Ρa W U G CA Co ica ia e es a is ( i aeus I Ρa W SS O G Co i ica a i a is ( ou I AU W C U O G Co i ica a me i ia a o so Ι AU W CC O G ie e e a i um (Au é I Ρa W SS U CA E icmus i o ea us ( ou Ε W CC U O G E icmus ca ice s ( ou Ε W CC U G E icmus o i us ( ou Ε W CC U CA E icmus o ea us e o Ε W CC U CA E icmus s a i e o E W CC U O G i os yg us mi o ou Ε U G i os yg us si uosus ( e o Ε C U G i os yg us s ecies Ε U G Me a o a ma a ac is ( ou E CC U G Me a o a ma i e sico is e o Ε W U CA Me a o a ma u e ( ou Ε W C U CA Me a o a ma u i u a ( ou Ε W CC U CA Me a o a ma a sa is ( ou E W CC U CA Me a o a ma e a ica e o Ε W C U CA Me a o a ma s ecies Ε W U CA Me a o a ma s ecies 2 Ε W C U CA Me a o a ma s ecies Ε W CC U CA Me a o a ma s ecies 4 Ι AU W O G Me a o a ma s ecies Ε W CC U G CA Ge us s ecies Ι AU W CC U G ME O YSII AE ( 2) o o a amecus e uis ei e Ε W C U G CA o o a amecus s ecies Ε W U G
MYCE O AGI AE ( 8) i a gus a ea us eCo e Ι Νa W ΟΡ G i a gus es i us (S a Ι AU W C U O G CA i y us a s e sus ( ou Ε W CC U CA
LEGEND Provenance: E, endemic; *, Lynfield type locality; I, introduced, foreign; AU, Australian; PA, Palearctic; NA, North American; SA, South American; ET, Ethiopian; IM, Indo-Malayan FR: first report from New Zealand Wing (condition): WD, winged; FL, flightless; WD FL, winged and flightless individuals Abundance: SS, single specimen; VR, very rare (2 or 3); RR, rare (4-10); CC, common (11-50); VC, very common (more than 50) Habitat: BU, bush; OP, open; BE, beach; GR, ground; CA, canopy. b l r S r f Sp t
x n (and no. of species) r v n n FR W n Ab nd n b t t i y us u igi osus (Β οu Ε W CC EU CA i y us is i ei us ( ou Ε W EU O CA i y us u c u a us ( ou Ε W C EU C i y us se a us ( ou Ε W CC EU G CA y aea s e co ea ( i aeus I A CC O G A C EOC Y ICI AE (1) A c eoc y icus ο a i Κas a Ι SA W C O C CII AE ( EU Cis as e imus Β οu Ε W CA Cis assimi is ou Ε W EU CA Cis oe ge i ( ei e Ε C EU C CA Cis i us is Β οu Ε EU C Cis i eico is ou Ε W CC EU CA Cis mi u us ay o Ε C EU C CA Cis u u us Β οu Ε W CC U C CA Cis ee a icus ei e Ε C EU C CA Cis s ecies Ε CC EU C Cis s ecies 2 Ε W EU G Cis s ecies Ε W CC U CA Cis s ecies 4 Ε W EU CA Cis s ecies Ε W SS EU G Έ ea ο sρecies Ε W EU O CA O o ius i u i es ack u Ι AU W EU CA Sco y ocis s ecies Ε W SS EU C y og a us u o i ( ou Ε CC U CA ME A YI AE (8) A o e us o a us ( ou Ε W CC EU CA C e o ec o ascia um ( e e ac e Ε EU O CA y o ia ca i a Β οu Ε C EU C CA y o ia s ecies Ε W CC EU CA y e oc a is s ecies Ε CC EU C Meco c esia s ec a i is ou Ε EU CA eo c esia i e ge s Β οu Ε CC EU C Ge us s ecies Ε CC U C MO E I AE (6) W EU CA Mo e a a a c ica i e Ε W SS Mo e a e ac a ascoe Ε W CC EU CA Mo ei a ucu a ( ou Ε W CC EU CA Mo e is e a s ecies Ι AU W EU CA S e omo e a ia eg ec a (Β οu Ε W C EU CA eamo e a mo ac a Β οu Ε EU CA I I O I AE (1) W A oci o s ookesi ou Ε SS . EU CA CO Y II AE (24) EU O CA i oma i su a is W i e Ε W C i oma a a S a Ε W EU CA i oma ugosa S a Ε W VC EU O CA i oma ici a S a Ε CC EU CA Co e us u ius S a Ε CC EU C Co e us simi is S a Ε EU O C E a sus akewe i ascoe Ε CC EU C E is a us i a is ou Ε EU C E is a us awso i S a Ε C EU C G e e e a se a a Β οu Ε EU C I is g aci is S a Ε W EU CA o ou us c assu us ou Ε W CC EU C CA is o e us as e (S a Ε W CC EU CA is o e us sca e ( a icius Ε CC EU O CA o a ius eco us ou Ε CC EU C yc ome us e essiuscu us (W i e Ε W C EU O C yc ome us a i a s S a Ε EU C yc ome us mi o S a Ε W CC U CA yc ome us sim e ou Ε C U C ecy us u e cu a us (Β οu Ε . U C i o ium a iquum S a Ε W CC U CA Sy ca us ys i S a Ε CC U C Sy ca us o a us S a Ε CC EU CR a iomimus i e a us Wo as o Ε C EU CA OS OMI AE (1) yoco a owi i ascoe Ε W CC EU C O E I AE (1) ou i y a squamige ( ou Ε U C
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x n (a o o s ecies Ρr v n n W n Ab nd n b t t
E E IO I AE ( 1 A i o ius ia e i us ( a e I ΙΜ W ΟΡ G Ama osoma simu a s e e ac e Ε U G Ama ygmus is is se su ack u I AU W CC U O CA A o a u i es a es Ε W CC U O G A c aeog e es cos i e is ou Ε U G A ys o a ugice s a es Ε CC U CA E a us o acus ou Ε SS U G G a oce us co u us ( a icius Ι ΙΜ W CC ΟΡ G o e us c assico is ou E W C U CA o e us a u us ou Ε W U G CA o e us iscus S a E W C U CA Me imus a esi S a Ε CC U G Me imus caecus S a Ε C U G Me imus c assus S a Ε C U G Mimo eus e o ga us ( me E CC U E G e o eis auck a icus ( ou Ε U G a yc i us me a icus W i e E W SS U CA a yc i us so o ae ou Ε W U CA i o ium a a e ( a icius Ι ΙΜ W O G U oma e e io oi es (W i e E W C U O G y oc us i ia is ou Ε W CC U CA OE EME I AE ( a iso a us mac eayi (C am io Ι AU W SS ΒΕ G a iso a us ig o o a us ( o ema I AU W ΒΕ G SA I GI AE ( Sa i gus a gus ico is ou Ε W C U O CA Sa i gus i u a us ascoe E W C U CA Sa i gus i us ou E W C U CA Sa i gus e u c a us ou Ε W C U CA Sa i gus quisqui ius ou E W CC U O CA Sa i gus e uc us ( ai Ε W U CA Sa i gus swa ei ( ai Ε W U CA Sa i gus s ecies Ε W CC U CA A ICI AE (1 A icus o a is ( i aeus I Ρa W CC O G A icus g a e Ki g Ι AU W ΟΡ G CA A icus es e i Ki g Ι AU W CC U O G A icus k eus e i Ki g Ι AU W CC U O G A icus e uci i es ou Ε W C U CA A icus s ecies Ε C U O E G Co es u a a ( ou Ε SS U G Co es c is i ( ou Ε CC U G Co es o sa is ou Ε CC U CA Co es o ima ou Ε CC U CA Mac a ia e i is ascoe Ε W CC U CA ic a a ca s ecies Ε W U CA A E I AE (5 y o i us i i us ( ou Ε W SS U CA y o i us s ecies Ε W U CA y o i us s ecies 2 Ε W SS U CA y o i us s ecies Ε W U CA y o i us s ecies 4 Ε W SS U CA SC A II AE (3 o o e us ige us ( ou Ε W CC U CA o o e us usi a us ( ou Ε W U CA o o e us s ecies Ε W CC U CA CE AM YCI AE : r n n ( io o us e icu a is W i e Ε W C U O CA : A n ( A o a us is is ( a icius Ι Ρa W C ΟΡ CA : C r b n ( Am eo o us is is ( a icius Ε W CC U O CA BUAs e o ea au e a es Ε W C O CA As e o i a uci a ou Ε W C ΟΡ CA e e ium sig i e um ( ewma I AU W C ΟΡ CA Ca i io is scu e a is ( a icius I AU W SS ΟΡ CA BUCo omma a iega um ( a icius Ε OPW CC CA BUE u i a su i ea a W i e Ε OPW CC CA BUGas osa us ig ico is a es Ε W ΟΡ CA
LEGEND Provenance: E e emic; y ie y e ocaii y; I i o uce o eig ; AU Aus a ia ; Ρa a ea c ic; Νa o Ame ica ; SA Sou Ame ica ; ΕΤ E io ia ; ΙΜ I o-Ma aya FR: i s e o om ew ea a Wing (condition): WD, wi ge ; ig ess; W wi ge a ig ess i i i ua s Abundance: SS si g e s ecime ; e y a e ( o 3 ; a e ( -1 ; CC commo (11-5 ; C e y commo (mo e a 5 Habitat: BU, us ; OP, o e ; E eac ; G g ou ; CA ca o y b l r S r f Sp t
x n (and no. of species) r v n n W n b nd n b t t a omo a su ca a ( a icius Ε W C U O CA Oemo a i a ( a icius Ε W C U O CA Oc ocy us u o i ascoe Ε W CC EU CA o aca a semi u c a a ( a icius Ι AU W C O CA S e o o es a i us ascoe Ε W C EU O CA essa omma u a um ewma Ι AU W CC O CA o um mu um ou Ε W U CA u o es u c i e is ascoe Ε W CC EU O CA o io mi u um ( a icius Ε W C EU CA : n (37) A io ea a i a a Β οu Ε CC U CA y o asius c is a ( a icius Ε W C U CA y o asius mo es us ou Ε W U CA y o asius e a o a es Ε W C U CA y o asius omissus (Β οu Ε W U CA y o asius sim e a es Ε W C EU CA y o asius s ic icus ( a es Ε W CC EU CA y o asius ege s ou Ε W CC EU O CA y o asius i i esce s a es Ε W C EU O CA Mic o amia ygmaea a es Ε CC EU CA o u osoma a gus um ( ou Ε CC EU G o yaca ia a i es (W i e Ε W EU CA si oc aeia as eiiae Kusc e E C EU CA si oc aeia ou i a es Ε W C EU CA si oc aeia i ea is a es Ε W CC U CA si oc aeia a a (Bates) Ε C EU O CA si oc aeia a u a (W i e Ε - CC U CA i osoma co e um ( ou Ε CC U C i osoma i oi es ( a es Ε C U C Soma i ia a a c ica (White) Ε CC EU O G CA S i o ogia e o ga a ( ou Ε CC EU G CA S i o ogia agi is (Bates) Ε W CC EU CA S i o ogia macu a a a es Ε W CC U CA S i o ogia ic u a (Bates) Ε SS EU CA S e e i sis imacu a a (White) Ε W CC EU CA S e e i sis g aci is (White) Ε W EU CA S e e i sis a i e is a es Ε W CC EU CA e e osoma a icoma ( ou Ε CC EU G e e osoma e e icum ( ou Ε CC EU C e e osoma e es e ( ou Ε EU C e o ea ci i es W i e Ε W CC EU CA y o o es cos i e is (Breuning) Ε EU CA y o o es g iseus (Fabricius) Ε C EU O G CA y o o es ume a us a es Ε W CC EU CA y o o es ae us W i e Ε C EU O CA y o o es u us a es Ε CC EU CA y o o es ugico is a es Ε CC U CA C YSOME I AE : Cr pt ph l n (2 A omus ou i S a Ε W CC U CA Oc oso sis su ascia a (Saunders) Ι AU W CC O CA : Ε οlρ n (6) Euco as is u ea ( a icius Ε W EU O CA Euco as is co o a a ou Ε W C EU O CA Euco as is ucu a ( ou Ε W C EU O CA Euco a sis a i i e is (W i e Ε W EU O CA Euco as is ic ico is ou Ε W C EU O E CA e i icus su usus S a Ε C EU C : Chr l n (6) A i o e igma S a Ε CC EU G A i o mi u um ou Ε C EU C A i o mo s osum ou Ε CC EU G A i o s ecies Ε SS EU C y goi es s ecies Ι AU W CC O CA a o sis c a y is S å Ι AU W RR O CA : G l r n (2) A o ia u c ico is (S a Ε W EU CA A o ia u ga is (Β οu Ε W CC EU CA : Alt n (4) A ema a a o a (Sharp) Ε W C U CA o gi a sus u igi osus ( ou Ι AU W C O C eu a ica cya ea ( ou Ε W C EU CA ac y e a ugu osa ( ou Ε C EU CA EMO YC I AE ( i o y c us u u us ( ou Ε W C U CA A I I AE : Anthr b n ( 0 Cace a us u o i (Sharp) Ε W C U O CA asya i us u u eus ( ou Ε CC EU CA
6 nf ld tl S rv
x n (a o o s ecies r v n n W n Ab nd n b t t E a is s i ico is S a Ε W U CA Eucio es su u a is ascoe Ι AU W CC O G Ga yus a us (S a Ε W CC U O CA awso ia a ia i is S a Ε W CC U CA o us u is (S a Ε W C U CA yma us e ae a (S a Ε W C U O CA eos o ius u a us (S a Ε W C U O CA S a ius ou i (Sharp) Ε W C U O CA : Ch r n ( A aece us a ma is ( ascoe Ι AU W CC O CA ys oc y us ig us ( ou Ε CC U G ys oc y us i a us (S a Ε C U O G ys oc y us ugosus (S a Ε C U G CA Mic a i us a omus (S a Ε C U G o oc o agus c assus (S a Ε W CC U CA o oc o agus o acicus ( ou Ε CC U G
E I AE : A l d r n ( A a ius wo as o i (S a Ε W CC U CA E I AE : Ap n n (2 A io u icis ( o s e I Ρa W C O CA eocy a me osi e os ( ou Ε W U CA CU CU IO I AE : r h r n : Ent n ( 6 Asy o yc us ce i us ( o ema Ι SA ΟΡ G ac yo us s ecies Ε U G CA Ca o es i o is (W i e Ε U CA Cecy o a isco s ou Ε ΒΕ G G a og a us euco oma ( o ema Ι SA O G I e imus com essus ( ou E CC U ΒΕ CA y e o a es as e ou Ε CC U G Ma a o us i i us ( ascoe Ε C U G Ma a o us mi ico is ( ou Ε CC O ΒΕ G Ma a o us s ecies Ε ΒΕ G O io y c us ugosos ia us (Goe e I Ρa • ΟΡ G O io y c us su ca us ( a icius I Ρa O G ae oc a is co u e us ( ou Ε C U G ae oc a is se i e ( ou Ε CC U G yc i us ca osus o ema I E CC O G Si o a iscoi eus Gy e a Ι Ρa W O G : r h r n : At rp n ( a i osomus acumi a us ( a icius Ε W CC U O G : r h r n : G n pt r n ( Go i e us scu e a us Gy e a I AU W SS O CA : r h r n : b thr r n (4 Geoc us i aequa is ( ou Ε CC U G Geoc us o i us (Broun) Ε C U G Geoc us simi is ou Ε ΒΕ G Geoc us s ecies Ε SS U G : r h r n : h t rh n n ( G omi us se osus ( ou Ε CC U G G omi us o acicus ( ou Ε CC U G is o e es i ici is Ge mai Ι SA W SS ΟΡ G is o e es o ea us ( ea Ι SA W SS ΒΕ G is o o us o a ie sis (Kusc e I SA W C O ΒΕ G S e i us asci us ( ascoe Ε ΟΡ G S e i us i e si es i ea us ( ascoe I AU W SS O G : C r l n n : Er rh n n (2 yoca us o y ic i Kusc e Ε CC U G My o ymus e a icus Kusc e E C U O G : C r l n n : E n n ( Go o o e us s i ico is ou Ε W SS U CA o oc eme ooke i W i e Ε W U CA o oc eme squamosa ou Ε W U CA y e es i e s ( a icius Ε W U CA ac o a emissa ascoe Ε W C U CA ac o a a ia i is ascoe Ε W C U CA o a ome us ucosus ( ascoe Ε W SS U CA Sco o e us aequus ou Ε W CC U CA Sco o e us e ici a us W i e Ε W CC U CA S e a o y c us c assus ou Ε W CC U CA S e a o y c us cu i es W i e Ε W CC U CA
LEGEND Provenance: E nd ; *, nf ld t p l l t ; I ntr d d f r n; AU A tr l n; Ρ l r t ; Ν rth A r n; SA S th A r n; ΕΤ Eth p n; ΙΜ Ind -M l n f r t report fr l nd W n (condition): W winged; fl htl ; W winged nd fl htl individuals Ab nd n SS n l p n; v r r r ( r 3 ; r r ( -10); CC n (11-5 ; VC, v r n ( r th n 5 Habitat: U b h; O open; E beach; GR, r nd; CA canopy. b l S r f Sp t
x n (and no. of species) r v n n W n Ab nd n b t t S e a o y c us awso i S a Ε W CC U CA ysius ico is ( a icius Ε W CC U CA : C r l n n : C r l n n ( A euma ascia um ( ou Ε W CC U CA A euma u i es ascoe Ε W CC U CA A euma ii ica e (Broun) Ε W C U CA Gym e o ascuo um (Gy e a I A W CC U O ΒΕ G eomyc a u i a ou Ε W U CA e is o eiis c ucige (Broun) Ε W U CA e is o eus u us ( ou Ε W U CA e is o eus usco o a us ( ou Ε W SS U CA e is o eus eucocomus ( ou Ε W U CA e is o eus mao i us ( ou Ε W U CA e is o eus se macuia us ( ou Ε W CC U CA e is o eus su us ( ou Ε W SS U CA e is o eus i i i e is ( ou Ε W U CA ao e a u o i ou Ε W U CA aoie a i usca ou Ε W CC U CA aoie a squamosa ou Ε W U CA S o eus a osig a us ack u I AU W C O CA : C r l n n : C t rh n h n ( i o cus aus a is Oke I AU W CC O CR : C r l n n : r d n ( i oge acus u a us ( o ema Ι SA W O GR : C r l n n : M d l n n ( eo aemosaccus a i us ( ascoe I AU W CC O CA : C r l n r : Cr pt rh n h n (62 Aca es ce us ou Ε CC U G Aca es ou ci gu a us Ε CC U C CA Aca es ou co ico is Ε SS U G Aca es ou o sa is Ε CC U G Aca es ascoe ys icu us Ε C U CA Aca es ou a i os is Ε U G Aca es Β οu sci us Ε U G Aca es e ico a ou Ε CC U C Aca es s ecies Ε SS U C Agaca es o mosus ou Ε C U CA Agaca es i ege (Broun) Ε U G Agaca es s ecies Ε SS U G A a a cis a ios e us (Broun) Ε U G A ia a cis i cu us ( ou Ε U C A aca es o i us ( ou Ε C U CA A aca es i i us ( ou Ε CC U CA A acai es s ecies Ε U C A acai es s ecies 2 Ε U CA C y eo us ookesi ( ou - Ε SS U CA Ciy eoius ac ymosus ( ou Ε CC U CA C y eo us ascoei ( ou Ε CC U C CA C y eo us sig a us ( ou Ε CC U C CA C isius ascicu a us ( ou Ε CC U C C isius a iega us ou Ε CC U CA e mo ius a i osus ( ou Ε C U C e mo ius u ico is ( ou Ε CC U C e mo ius sa gui eus ( ou Ε U C i ymus e o eus ( ascoe Ε C U CA i ymus i u us (Pascoe) Ε CC U CA i ymus s ecies Ε C U O CA Eu y i us squamige W i e Ε W U CA Ge aca es e a is ( ou Ε CC U C I ece ia u i a ou Ε U CA Mecis os y us ouei aco ai e Ε W C U CA Meso e a su ci o s ou Ε W CC U CA Meso e a s ecies Ε W CC U CA Me aca es as e sus ou Ε C U C Me acai es s ecies Ε U C Mic oc y o y c us k o ei (Kirsch) Ε C U O CR CA Mic oc y o y c us e usi us ( ascoe Ε C U O CA Mic oc y o y c us a e ( ou Ε C U CA Mic oc y o y c us s ecies Ε CC U C Mic oc y o y c us s ecies 2 Ε CC U CA o aca es e icu us ( ou Ε C U CA o aca es s ecies Ε CC U CA o aca ies s ecies 2 Ε CC U CA o aca es s ecies Ε U CA Omoeaca ies c isioi es ( ou Ε C U CA Omoeacaiies o a e us ( ou Ε CC U CA a oma ia se ige a ou Ε CC EU G a oma ia es i a ou Ε EU C se o a su ca us W i e Ε W C EU CA
8 nf ld tl S rv
x n (and o o s ecies r v n n W n Ab nd n b t t Sce o o ic us a u us ou Ε C U G Sce o o ic us i a is ou Ε U G Sce o o ic us i ei o a ( ou Ε C U G S o gy o e us y o ioi es (W i e Ε C U CA Sym e ius e i us ou Ε CC U CA Sym e ius es u o ascoe Ε CC U CA Sym e ius e a us ascoe Ε CC U CA yc a us e uga us ascoe Ε U G yc a us gi us ascoe E - CC U CA yc a us e ucosus ascoe Ε SS U CA : C r l n n : M l t n ( A eco aga a ia r n Ε W U CA E ym eus s a i ascoe Ε CC U G I us s ecies Ε U G ae a e us is i us ascoe Ε C U G o i a sim e ( ou Ε C U G y i us as u us ascoe Ε C U G y i us mo icus ou Ε CC U G ogo o i us o acus ( ou Ε W CC U G Sosge es o gico is Β ου Ε U G : h n h ph r n (2 yo o us s ecies Ι AU W U ΟΡ G Si o i us o y ae ( i aeus Ι ΙΜ W C ΟΡ G : C n n ( 4 Agas eg us ae eo iceus ( ou Ε W CC U CA Agas eg us simu a s (S a Ε W CC U CA A ao us ugosus ( ou Ε C U G Cam osca us a iuscu us ( ou Ε W U CA Ei a us o a us ou Ε W CC U G Ei a us a u us ascoe Ε W C U CA Eucosso us se ige (S a Ε W CC U CA Euo yum u um ( ou Ε W C U O ΒΕ CA E omesi es o imus ou Ε U G e e o sis a i os is Ma s a Ε W SS U CA e e o sis awso i Wo as o Ε W C U CA e e o sis s ecies Ε W CC U CA Mac o y co us i o a is ( ou I AU? SS ΒΕ G Mac oscy a us a ico is (S a Ε W C U CA Mac oscy a us emo us (S a Ε W CC U CA Mic o i us ou i (Wo as o Ε U CA Mic o i us u o i Wo as o Ε C U G Mic o i us s ecies Ε U G o i as is a ou Ε CC U ΒΕ G ac yo s u ius (Wo as o Ε W C U CA e a um ea a icum Wo as o Ε W CC U CA oeo agosoma co i um Wo as o E W CC U G oeo agosoma i u um Wo as o Ε W C U O CA oeo agosoma e a um Wo as o Ε W CC ΒΕ G Se ico ogus su ae esce s Wo as o Ε W C U O CA S e o u is e i is (S a E W U CA S e o u is wo as o ia us (S a Ε W SS U CA a ysoma coma um ( ou E W U CA o os oma a ica e ( ou Ε W CC U CA ou a o gi os is (Wo as o Ε W CC U CA ou a s a ia a (Wo as o Ε W CC U CA ou o sis e i os is (S a Ε W CC U CA ou o sis s ecies Ε W SS U CA U as co i os is Ma s a Ε W U CA : S l t n ( 2 Amasa u ca a (E ic so Ι AU W O ΒΕ CA C ae o e ius mu u us ( ou Ε W C U CA C y a us wa e i (Eic o Ι AU W O CA e o u es cos ice s ou Ε W C U CA e o u es es i us ou Ε W CC U CA e o u es s ecies Ε W CC U CA y u gus ig i e a ( a icius Ι Ρa W O CA y oc y a us s ecies Ε W C U CA y oc y a us s ecies 2 Ε W C U O CA oeosi us cu essi o ki s Ι Νa W CC O CA y e o i us euca y icus (Sc e I AU W C U O ΒΕ CA y e o i us sa ese i ( a e u g I Ρa W O CA : Ρl t p d n ( a y us a ica is W i e Ε W CC U CA
EGE r v n n E nd ; nf ld t p l l t ; I ntr d d f r n; AU A tr l n; A l r t ; A rth A r n; SA S th A r n; ΕΤ Eth p n; IM Ind -M 1 γ n f r t r p rt fr l nd W n ( nd t n W n d; fl htl ; W n d nd fl htl nd v d l Ab nd n SS n l p n; v r r r ( r 3 ; r r ( -1 ; CC n (11-5 ; C v r n ( r th n 5 b t t U b h; O p n; E b h; G r nd; CA n p
Ann t t d t f Sp 39 ANNOTATED LIST OF SPECIES
RHYSODIDAE REMARKS: In tr b d rd d fr AK S h r ld b tl Clivina vagans Putzeys, 1866 a FIRST REPORTED I N.Z.: T. Broun, 1880, nd r th misi e - i ie name C. rugithorax Putzeys. EARLIEST N.Z. RECORD: Auckland, AK, n th 1870s, T. Broun. REMARKS: In gardens, paddocks, h n rd nd tr beds. Widespread in the North Ι ; l n nd MC
CARABIDAE: b d n 'ground beetles' (assisted by B. P. M r 16 Rhyzodiastes Paratachys crypticola ( r tt n 1960) a Rhysodidae proprius FIRST REPORTED IN N.Z.: E r tt n 1960. EARLIEST N.Z. RECORD: Okauia, WO—BP, b 1924, t l ht A.E. r Kaveinga orbitosa (Broun, 1880) REMARKS: An ntr d d p f n rt n r n b t REMARKS: U ll f nd n d d d b t th n l pr b bl Palearctic. r nt n AK WO CL, G WI, nf ld p n xtr t d fr l l pl nd WN. 13-23 d p Tachys antarcticus t 1874 Kupeus arcuatus (Chevrolat, 1873) REMARKS: ln ft d d d d REMARKS: r f nd n l n d d l f Kn htia nd th r tr th r nd r n p n h t r t d b Tachys australis S h 1863 a Phellinus f n (Hymenochaetaceae) n l r n r t I S REPORTED IN N.Z.: T. Broun, 1893b, nd r th n r f l v Vitex tr Bembidium tersatum Broun. EARLIEST N.Z. RECORD: Mokohinau I., ND, b f r 1893, S nd r CARABIDAE REMARKS: C n n d p r nd rd d fr r nd b tl AK WI nd WN. Tachys captus l b rn 1888 b Ε IES Ν Ζ RECORD: Hikurangi, ND, 6 A 1927, Ε r- b rn REMARKS: In h p p dd pr nt d n NZAC b p n fr nd AK
CARABIDAE: Psydrinae r nd b tl ( t d b Β M r Mecyclothorax rotundicollis (W i e 1 a 17 Ctenognathus FIRST REPORTED IN N.Z.: A Wh t 1846. Carabidae novaezelandiae EARLIEST N.Z. RECORD: f I l nd n th rl 1840s, S n l r CARABIDAE: Cicindelinae REMARKS: In rd n nd p dd r t d ew ea- a bl n f M puncripennis (Macleay) b M r (I984 t r b tl nd pers. comm.), b t i s occu e ce in p n f ld makes Neocicindela spilleri n Nidek, 1965 i n nl l nd d t f r nd t t n th ntr REMARKS: rt l rl n in th b h nd in h d b h tr ; nl n ll n th p n in rd n nd CARABIDAE: Pterostichinae p dd r nd b tl Neocicindela tuberculata (Fabricius, 1775) Fig. 117 Aulacopodus calathoides (Broun, 1886) REMARKS: In p n tr b d nd b h tr ; n in REMARKS: Und r l n th d t l r f p dd rd n r d t l l p Aulacopodus sharpianus (Broun, 1893) CARABIDAE: Scaritinae REMARKS: In tr b d in b h r nd b tl Laemostenus complanatus (Dejean, 1828) ( t d b Β M r FIRST REPORTED IN N.Z.: W tt n 19 EARLIEST N.Z. RECORD: Tairua, CL, rl 1870s, T. Broun. Clivina basalis Chaudoir, 1843 b REMARKS: More nl f nd n r h nd r h p I S REPORTED IN N.Z.: P.J. C r n nd C.F. t h r f d r rd n l pp n 1980. Rhytisternus r EARLIEST N.Z. RECORD: Ohia, ND, 4 Feb 1917, Τ Chaudoir, 1865 a Broun. FIRST REPORTED IN N.Z.: T. Broun, 1880, nd r th nam Holcaspis (Rhytisternus) rugifrons r n 1880. K t b l ft r p n EARLIEST N.Z. RECORD: A l nd AK b f r 1880, Βr n a ntr d d p b f r t r p rt f th p n l nd REMARKS: In C p n b h tr fr tr b n nd b d nf ld th t p l l t d t x n h n pp 6-8 l tt r hr K n Is, ND, AK, CL, WO, TK, nd WN.
y ie ee e Su ey
CARABIDAE: Agoninae REMARKS: Commo i ga e e use a com os I AC g ou ee es e ese e y s ecime s om AK WO W a (assis e y Β Moo e Demetrida nasuta White, 1846 Ctenognathus bidens (C au oi 1 7 REMARKS: ea e om eyci e ia a ksii REMARKS: Commo a ig o e us oo Philophloeus luculentus ( ewma 1 a b Ctenognathus cardiophorus (C au oi 1 7 EARLIEST N.Z. RECORD: Katikati, BP, 8 Oct 193I, a.E. REMARKS: S eam e s a a ks a us oo Brookes. Ctenognathus novaezelandiae ai mai e 1 3 d ig 17 11 REMARKS: on plants, particularly in trunks with rough or REMARKS: O coas a s o es a c i s a icu a y i ege- loose bark. Represented in NZAC y s ecime s om AK a io o e ma i ime o e C a K Notagonum lawsoni ( a es 1 7 REMARKS: Commo i s eam e ege a io and Carex CARABIDAE: Pentagonicinae swamps. g ou ee es Notagonum submetallicum (W i e 1 a Pentagonica vittipennis C au oi 1 77 FIRST REPORTED I a W i e 1 u e e ame FIRST REPORTED IN N.Z.: T. Β οu 1 u e e ame Colpodes su me a icus W i e Wake ie ia i a a Broun. EARLIEST RECORD: " ew ea a " i e ea y 1 s EARLIEST N.Z. RECORD: o I (" i e e oca i ies" i REMARKS: On e g ou a o ege a io e 1 7 s T. ou REMARKS: In s eam e s o Ga ia se i o ia a i Ma - CARABIDAE: Harpalinae aise a e ese e i AC y s ecime s om g ou ee es AK, CL, BP, SD, MB, NN, WD, MC, and FD. (assis e y Β Moo e Scopodes elaphroides (W i e 1 Haplaner species 1 a b REMARKS: In s eam e s EARLIEST N.Z. RECORD: Mt Te A ua a a a a 1 m ua- i e a ge HB, e 197 G W amsay Y ISCI AE REMARKS: O e i s eam e o e i Ma aise a I AC wa e ee es a sο om HB, WI, a W A u esc i e species from Aus a ia ( Moo e e s comm Hypharpax australis ( e ea 1 9 a FIRST REPORTED I e e ac e 1 REMARKS: e ese e i AC y s ecime s om AK CL, WO, BP, GB, HB, RI, WN, SD, NN, MB, MC, CO, and Chatham Is. Lecanomerus sharpi (Csiki 193 REMARKS: Commo i Ga ia se i o ia c us e s o coas a s o es; occasio a y i ga e s a a ocks Lecanomerus vestigialis (E ic so 1 a FIRST REPORTED I Ν Ζ T. ou 1 u e e ame Fig. 1 Liodessus eca ome us s e o us Broun. y isci ae plicatus EARLIEST N.Z. RECoRD: owick AK i e ea y 1 s T. Broun. Hyphydrus elegans (Mo ou ie 1 a REMARKS: I w ack e i eac es a i e ise um u FIRST REPORTED I Ν Ζ ou 1 u e e ame i a ock e ese e i AC y s ecime s om y o o us i i ico is ou AK C G W a EARLIEST N.Z. RECoRD: Ma s e oi i e 1 7 s Mu o Lecanomerus atriceps (Mac eay 1 71 REMARKS: In wa e oug s a o s o I. FIRST REPORTED IN N.Z.: C i g im 19 3 EARLIEST N.Z. RECORD: Swa so AK A 191 a E Rhantus suturalis (Mac eay I 5 a ookes FIRST REPORTED IN S a 1 u e e ame R. REMARKS: Widespread in ND, AK, and WN. pulverosus (S e e s EARLIEST N.Z RECORD: " ew ea a " i e 1 7 s Syllectus anomalus ( a es 1 7 REMARKS: In o s a wa e oug s REMARKS: In s eam e CARABIDAE: Lebiinae PTILIIDAE g ou ee es ea e -wi ge ee es (assis e y Β Moore) (assis e y C Johnson) Anomotarus illawarrae (Mac eay 1 73 a Acrotrichis inconspicua (Matthews, 1 7 FIRST REPORTED IN N.Z.: Sco I9 3 FIRST REPORTED IN N.Z.: a Ma ews 1 7 EARLIEST N.Z. RECORD: Swa so AK I ec 1955, J.C. EARLIEST N.Z. RECORD: Auck a AK i e ea y 1 7 s Watt. awso REMARKS: In Gahnia se i o ia and coas a ege a io A REMARKS: I ga e e use Aus a ia s ecies so a ou o y i AK a WN. Acrotrichis montandoni (A i e 1 Anomotarus variegatus Moore, 19 7 FIRST REPORTED IN N.Z.: C. Johnson, 1982. FIRST REPORTED IN J.C. Wa 19 3 EARLIEST N.Z. RECORD: y ie AK 1 May 197 G EARLIEST N.Z. RECORD: Auck a AK Ma 19 I Kusc e S i e REMARKS: In mi e com os Ann t t d t f Sp 41
EARLIEST RECORD: y ie AK 3 Oc 197 C Kusc e REMARKS: In ga e s a a ocks; commo i com os ea s Ptenidium punctatum (Gy e a I 7 a FIRST REPORTED IN C o so I9 EARLIEST RECORD: y ie AK 3 u 197 C Kusc e REMARKS: a o i ous; ou i seas o e w ack a occa- sio a y i ga e com os Ptenidium pusillum (Gy e a 1 a ig 19 o o e i ium FIRST REPORTED IN N.Z.: C o so I9 i ii ae a wso i EARLIEST RECORD: y ie AK 3 u 197 C Kusc e Acrotrichis subcognata o so 1975 REMARKS: In ga e com os Ι SΤ E O E I C 1975a o so C EARLIEST RECORD: Auck a Is 19 A 19 7 So - Ptiliodes amplicollis o so 19 e se REMARKS: In w ack o seas o e REMARKS: I ga e com os Ptiliodes austerus o so I9 C Actidium angulicolle o so I9 C REMARKS: I w ack o seas o e REMARKS: I moss ma s f y o e o a cua um a Ptinella acaciae o so 19 Ec i o ium is i um o s ee s o es y s eams K ow REMARKS: In ecaye Acacia mea sii a K ig ia e ce sa
o y om y ie i es e y Ga o e ma a a a um ( o y o aceae « Actidium lineare Ma ews 1 7 Ptinella cavelli ( ou 1 93 REMARKS: O e seas o e e i e s as - o e REMARKS: O Co y i e a ksii Cissidium foveolatum o so I9 c Ptinelki confusa o so 19 C REMARKS: I i e a ecaye woo REMARKS: In ecaye woo K ow om y ie o y Dipentium zelandicum o so 19 Ptinellaferruginea o so 19 c REMARKS: I υmi ιe ogs a soi REMARKS: I ecaye woo Kuschelidium maori o so I9 Ptinella taylorae o so 1977 REMARKS: O e s ecime ou i Ga ia se i o ia e o e REMARKS: In ecaye woo A ew ea a s ecies a u- e ac e om a moss ma y ie is e o y ace e a ise i E g a a I e a ge us as we as e s ecies is k ow om C Ptinella watti o so I9 : Nellosana grandis o so I9 REMARKS: I ecaye woo REMARKS: I ea i e K ow om y ie o y Nellosana intermedia o so 19 AGYRTIDAE REMARKS: I ecaye woo K ow om y ie o y u yi g ee es Nellosana n C o so I9 (assis e y A ew o REMARKS: I ecaye woo a o a c o K ig ia e - ce sa wi ame es ci a a i us ( o y o aceae
Nephanes t t n ( ewma 1 3 FIRST REPORTED IN C o so I9 EARLIEST RECORD: y ie AK Ma 1975 G Kusc e REMARKS: I ga e com os Notoptenidium crassum o so 19 REMARKS: Nearly a co ec e i associa io wi Meso- o e a a u s K ow om y ie o y Notoptenidium lawsoni (Ma ews 1 73 Fig. 19 I19 ig REMARKS: I ea i e a ecaye woo 'Necrophilus' Agy i ae o o ga us Notoptenidium oblongum o so I9 REMARKS: wo o ai e i ee i - a s se o y ogea ec o i us o o ga us (S a 1 I Fig. 1 au a a o e i Meso o e a cas a ea a u s K ow om y ie o y REMARKS: O ca io A u s i e away i ee c acks a o ows o s ee s a y i si es a s eam a ks i us Oligella foveolata (A i e 1 FIRST REPORTED IN C o so 19 EARLIEST RECORD: y ie AK 15 Se 197 G LEIODIDAE Kusc e ca io ee es REMARKS: In ga e com os (assis e y A ew o Ptenidium laevigatum E ic so 1 5 FIRST REPORTED IN C o so I9 LEIODIDAE: Leiodinae Colenisia p
K t b l ft r p n REMARKS: In us - oo i e . a i o uce s ecies b i s e o o is s ecies i ew ea a Isocolon hilare 1 93 C Β οu y ie is e y e oca i y d a o omic c a ge; see 7 - 1 REMARKS: In amo gs g ou a s a us - oo i e 42 y ie ee e Su ey
EIO I AE : EIO I AE (cont.) Pseudonemadus lituratus (Broun, 1880) EMA KS In carrion, but much less frequent than the other two cholevine species.
SCYDMAENIDAE `stone beetles' (assisted by H. Franz)
Fig. 21 Colon Leiodidae hirtale Zeadolopus maoricus Daffner, I985 EMA KS In bush litter. Zeadolopus spinipes Broun, 1903 EMA KS In bush litter. Fig. 22 Adrastia Zeadolopus validipes Daffner, 1985 Scydmaenidae clarkei EMA KS In bush litter. Genus 1 species 1 Euconnus calvus (Broun, 1880) d EMA KS In leaf litter. EMA KS From grass mats of Oplismenus and Microlaena, d and especially from an agaric sample taken in the bush. Euconnus setosus (Sharp, 1874) Genus 1 species 2 REMARKS: At base of C r x lambertiana, C. dissita, and Uncinia banksii on bush floor, and in Ptychomnion acicu- EMA KS In decayed Knightia log. lare moss mats. LEIODIDAE: Camiarinae Euconnus species 1 Agyrtodes disparatus Szymczakowski, 1966 EMA KS In a hollow Sophora microphylla. EMA KS On Auricularia polytricha. Occasionally aiso on Magellanoconnus galerus (Broun, 1885) d Phellinus f n nd n r b h flοοr l tt r EMA KS From sifted wood by Mesoponera castanea a runs under a large Metrosideros excelsa tree Agyrtodes hunuensis (Broun, 1893) d EMA KS In litter and amongst Gahnia plants. Few under Maorinus angulatus (Broun, 1893) logs and in the cracks of stream banks. EMA KS In Gahnia lacera litter on coastal cliffs and at bases of large, old Metrosideros excelsa trees. Agyrtodes nebulosus (Broun, 1880) d EMA KS On Ganoderma applanatum and in decayed Microscydmus lynfieldi Franz, 1977 wood. EMA KS Obtained from decayed wood of various trees. Lynfield is also the type locality for the synonym M. Baeosilpha rufescens Broun, 1895 omahutensis Franz, 1977. The holotypes were found to- EMA KS In decayed Melicytus ramiflorus wood. gether in fairly old, decaying M l t r flοr still pro- Camiarites convexus (Sharp, 1876) tected by its bark. EMA KS In bush litter. Microscydmus species 1 Camiarus thoracicus Sharp, 1876 EMA KS In decayed wood. A series was found in spongy EMA KS Amongst ground plants and in litter. white rot caused by a Phellinus fungus (Hymenochaeta- Inocatops concinnus (Broun, 1880) ceae) at a large, necrotic area on a live Vitex lucens. EMA KS In b h flοοr l tt r nd Ptychomnion moss mats. Microscydmus species 2 Zearagytodes maculifer (Broun, 1880) EMA KS In Ptychomnion aciculare moss mat in amongst EMA KS Confined to Ganoderma applanatum in Lynfield Gahnia setifolia under Leptospermum scoparium canopy. bush. Adults and larvae in large numbers in the white Microscydmus species 3 underside of this bracket fungus at all times. EMA KS One in decayed Acacia mearnsii in native bush, Genus 1 species 1 and one in hollow Sophora microphylla. EMA KS In mixed bush litter, under rotten Pinus radiata Neuraphoconnus kuscheli Franz, 1977 washed into the reserve by flooding, and in decayed EMA KS Extracted from leaf mould in hollow of an old, Knightia logs. diseased Knightia excelsa. Known from Lynfield only. LEIODIDAE: Colinae Neuraphoconnus species 1 EMA KS In decayed wood and hollow logs. Colon hirtale (Broun, 1880) Fig 21, 121 EMA KS Occasionally in bush litter; common in cracked, Neuraphoconnus species 2 undermined stream banks; numerous in Malaise traps. EMA KS In decayed wood and mould in hollow logs. Neuraphoconnus species 3 LEIODIDAE: Cholevinae EMA KS In decayed wood and Salix fragilis mould. Mesocolon alacre (Broun, 1880) Sciacharis fulva Broun, 1893 d EMA KS In carrion, and easily attracted with bait. EMA KS In leaf litter. Paracatops lugubris (Sharp, 1882) Sciacharis lanosa (Broun, 1886) d EMA KS In carrion, and readily baited. EMA KS At the base of Astelia banksii on coastal cliffs. Ann t t d t f Sp 43
Sciacharis puncticollis (Broun, 1880) d Scaphisoma p EMA KS In d d d nd n th l f ld f tr EMA KS On Ama i a musca ia nd r i us a ia a n h ll b h n ll Ca e a U ci ia pl nt n us oo Sciacharis sannia ( r nz 1977) a i Maiaise tr p EMA KS In d d d p ll n l r t p r t n 1 p 1 b r d n l EMA KS In d d d Sciacharis taranakii r nz I9 I d EMA KS In l f l tt r t b f l r i e uce s tr STAPHYLINIDAE Stenwhnaphes newtoni r nz 1985 r v b tl EMA KS On fr Sa i agi is d ld n Carex P. esso ia a p n th r in M l tr p Kn n l ( t d b Μ nd fr Tawhiti Rahi ( r Kn ht I p r HB, nd Pelorus r d MB (th t p l l t Stenichnaphes urbanus r nz I980 C d EMA KS r ntl f nd t b f As e ia a ksii nd Phormium tenax; n ll in Phellinus h t r t n l nd in h ll tr n f l v Co y oca us ae iga us E l ht in M l tr p G n nd p d r b d fr Lynfield p n nl rd d fr th Moko- h nau I t Pelorus r d MB. Stenichnaphes p 1 EMA KS C ht n M l tr p 24 o o o c us Staphylinidae Stenkhnus kuschelianus r nz 1977C ferrugineus EMA KS A n t Ga ia ace a pl nt (holotype); - nd p n f nd n d h ll l v So o a mic o- STAPHYLINIDAE: Proteininae phylla. Kn n nl fr Lynfield. Nesoneus acuticeps rr h r 1939 Stenichnus p 1 EMA KS In d p b tl frass nd r b r f Me icy us EMA KS On f nd n d h ll Me osi e os n th r n ami o us nd Co y i e a ksii a o s a i g d d d e yca ya a o ea t nf t d y G oeo o us a usi us f n (C rt . Silphotelus nitidus Broun, 1895 SCAPHIDIIDAE EMA KS In f rl dr l f l tt r f A seuosmia ac o y a ff b tl a ac yg o is e a a b t p rt l rl a u a n ( t d b I l bl b h fl r A r l n l d n Ama i a musca ia Occa- n ll n ic o oma essu a um f n nd p - ll n th p l p r d Irpex e is
STAPHYLINIDAE: Omaliinae . Broiiniellum hilarum (Broun, 191 EMA KS On pl nt nd p ll n d - ld t br n h f Sc e e a igi a a h n n tr Ischnoderus genalis (Broun, 1880) EMA KS ea e si g y fr e yca ya Me icy us a i e ht n M l tr p 23 Βaeοce a Ischnoderus tectus (Broun, 1 Scaphidiidae scu e a is EMA KS On ays - o cu a c es o Schefflera digirara a i os o um c ass οΙiυm n d d o a os y is sap-
`Baeocera' scutellaris (Redtenbacher, 1867) Fig. 23,122 ida p l l v ; n ll nd r b r ; r d l ht n EMA KS In d d Corynocarpus laevigcitus,Hedycarya M l tr p arborea, nd Melicope ternata. Microsilpha sp. Brachynopus kitus (Broun, 1881) EMA KS In l f l tt r f i e uce s th r d b t n r t EMA KS C n n b h fl r l tt r n ll l in b ttr d d d Omaliomimus litoreus ( ou 1 Scaphisoma actuosum rrn 1881 EMA KS In r n t r s o es EMA KS b In d d d Omalium allardi ai mai e & isoui 1 59 a Scaphisoma sternale (Broun, 1914) EA IES ECO y ie AK 5 O t 1974, G. EMA KS In l f l tt r f t l s o es Kuschel, s aw sam e 7 / EMA KS In p ltr tr f hen-house. Scaphisoma p 1 b EMA KS On usco o ia yo i a (Hymenochaetaceae) Paraphloeostiba gayndahensis (Mac eay 1871) thr h t th r EA IES ECO A l nd C t AK 26 M r 1944, n d n r l n D. Spiller. K t b l ft r p n a EMA KS On f r nt n fr t in fr h l n l pp n n ntr d d p b f r t r p rt f th p n l nd p t b n n U omyc a ium ϋ a iΙe ll f Acacia C nf ld th t p l l t d t x n h n pp 6-8 mearnsii, nd n ld r br t f M nfl r n
44 y ie ee e Su ey
S A Y I I AE OMA II AE (co REMARKS: In si o s a y s eam e Stenomalium cottieri e aue 1939 Carpelimus zealandicus (S a 19 REMARKS: REMARKS: O e om moss a i e wo s a o e a s eam I ga e s a ocks a c icke ya s u a - a k o e a ack o seas o e icu a y commo i si o s eam e s a see ages o coas a mu s o e c i s Es a is e i i is Isies Stenomalium helmsi (Came o 19 5 Carpelimus species 1 ig 1 3 REMARKS: O ea is a eac a s ee ca case a us ma gi REMARKS: e igme e a e ye ow; ou i see i g si y oam i e o cm o soi some imes a g ou e ei i Stenomalium moniliferum ( ou 1 93 u e a ick ma o yc om io acicu a e moss o O is- REMARKS: I a gi g ea o s o Cya ea me u a is me us im eci us g ass ea i y o ai e i ee i - a s a so i wigs o Sc e e a igi a a e o y i s a y i i o ai e a y ie Stenomalium philpotti ( ou 1 9 Oxytelus sculptus G a e o s 1 a REMARKS: O e o ai e om si e ea e o s o FIRST REPORTED IN N.Z.: T. ou 1 u e e ame Cya ea me u a is a gi g agai s a ee- e u k Oma ium u um Broun. Stenomalium spadix ( ou 1 EARLIEST N.Z. RECORD: Parua, ND, in the 1870s, T. Broun. REMARKS: I o es oo i e a icu a y i ea e -u REMARKS: I com os i s a ou y s aw es i e Teropalpus coloratus (Sharp, 1900) Stenomalium sulcithorax ( ou 1 REMARKS: O seas o e A u s a a ae mee i comi g REMARKS: I us oo i e es ecia y i e ic e wi i es o o ow ou goi g i es o a ou 1 m om e es am o c icke o es ea e -u w ack mo e so a ig a i e ay ime Νο i e i a a u s o a ae o se e STAPHYLINIDAE: Oxytelinae Teropalpus unicolor (S a 19 Anotylus brunneipennis (Mac eay 1 73 a REMARKS: a o i ous A eas e a u s o is s ecies ca FIRST REPORTED IN Μ Came o 195 u e e ame e u y i e i a o wo o mo e ou s i coa se y g a e y O y e us c ookesi Came o a eas m o mo e om e ig i e w ack Es a is e i EARLIEST N.Z. RECORD: M oski AK Ma 19 A E e U i e Ki g om ookes REMARKS: I e me i g ui s ga e e use c i i gs a STAPHYLINIDAE: Osoriinae u i gs a u g Paratorchus brevipennis ( ou 1 93 Anotylus complanatus (E ic so 1 39 REMARKS: I i e u a icu a y i e ea mou o FIRST REPORTED IN N.Z.: A au e 19 o ow ees e g Sophora mic o y a EARLIEST N.Z. RECORD: Otago, 1 77 W u o REMARKS: In fermenting fruit, lawn clippings, garden com- STAPHYLINIDAE: Eleusininae post, and sheep and chicken manure. Common i ci ies a Zeoleusis virgula ( au e 1 9 a m a oug ou e cou y REMARKS: I i ous issues o oke a c es o Me icy us A o y us semirufus ( au e 1 77 a ami o us a Co y oca us ae iga us a ea s ems o FIRST REPORTED I N.Z.: ou 1 u e e ame Co y i e a ksii a eyci e ia aue ia a Oma ium i ia e ou STAPHYLINIDAE: Paederinae EARLIEST N.Z. RECORD: ai ua C i e 1 7 s ou b REMARKS: I e me i g ui s ga e com os u g a Astenus guttula au e 1 77 a yis o ca cases EA IES ECO : ee y s K o 19 9 E S Gou ay Anotylus vinsoni (Cameron, 1936) a REMARKS: E cee i g y commo i ga e s a as u es FIRST REPORTED IN N.Z.: P.M. ammo 197 EARLIEST N.Z. RECORD: Auckland, AK, in the 1860s, Hyperomma dispersum ou 1 93 Lawson. REMARKS: In us oo i e REMARKS: In garden refuse, chicken yard, and sheep pad- Hyperomma mandibulare ou 1 93 dock; quite common in native vegetation too, especially in REMARKS: In amo gs moss a e s οf am s o es y shady stream beds. sma s eams Anotylus species 1 a b Hypomedon debilicornis (Wo as o I 57 b EARLIEST N.Z. RECORD: a me s o o WI—W 5 u e EARLIEST N.Z. RECORD: Auck a AK u 19 9 i ack- 193I W Co ie i g s aw M eace REMARKS: I e me i g ui s aw c i i gs ga e com- REMARKS: I aw a mi e ga e com os s ee a - os ea a u g ock a e - ouse Blediotrogus guttiger S a 19 Lithocharis nigriceps K aa I 59 a b REMARKS: a o i ous; i w ack EARLIEST N.Z. ECO : a me s o o WI-W 7 a - Carpelimus bilineatus (S e e s 1 3 b ua y 1975 M Esso EARLIEST N.Z. RECORD: Mt Te Aroha, BP, undated, Broun REMARKS: In lawn clippings, compost, and sheep paddock. Coll. (NZAC). Lithocharis ventralis ( ou 1 REMARKS: I compost bins and at back of beach. REMARKS: O us oo a s eam e Carpelimus corticinus (G a e o s 1 b Lithocharis vilis (K aa I 59 b EARLIEST N.Z. RECORD: Takaka Hill, NN, 762 m, 7 Oct 1964, EARLIEST N.Z. RECORD: y ie o ica a i e AK G. Kuschel. Ma 1975 sam e 75/1 G Kusc e REMARKS: In chicken yard, sheep paddock, and back of sea- REMARKS: I ga e com os a s ee a ock shore, but especially in bare silt of shady stream beds. Lobrathium bipartitum ( au e 1 7 b Carpelimus pusillus (Gravenhorst, 1802) a b EARLIEST N.Z. ECO : Auck a AK u 19 1 Μ EARLIEST N.Z. RECORD: o ica a i e y ie AK 1 May. Se 197 G Kusc e REMARKS: In ga e a a ock
Ann t t d t f Sp 5
Medon zeelandicus Redtenbacher, 1867 Fig. 124 Philonthus longicornis St ph n 1832 a REMARKS: In l tt r n b h fl r I S REPORTED IN N.Z.: A Fauvel, 1900. b EARLIEST RECORD: A l nd AK n th 1860s, aw- Scimbalium laetum l b rn 1888 EARLIEST RECORD: Waiheke I., AK, 18 A 1941, C. so Ch b rl n REMARKS: On e me i g ege a es a i a ock b REMARKS: In l n l pp n nd h p p dd Philonthus parcus Sh rp 1874 EARLIEST RECORD: Utiku, RI, 27 l 1I917, J. o Sunius propinquus (Brisout, 1867) a REMARKS: In l n p t p ltr tr nd ld ca cases FIRST REPORTED I N.Z.: Ν M rt n 1983. EARLIEST RECORD: Methven, MC 1911, T. ll Philonthus politus (Linnaeus, 1758) REMARKS: In h p p dd nd r Acacia mea sii FIRST REPORTED I N.Z.: C.A.A. Fauvel, 1900, ae eus auct. STAPHYLINIDAE: Staphylininae EARLIEST RECORD: Auckland, AK, in the 1860s, Cafius algophilus (Broun, 1894) n REMARKS: In r nd n nd nd r l REMARKS: On rr n b Cafius quadriimpressus (Wh t 1846) Philonthus pyropterus Kraatz, 1859 REMARKS: In r nd n nd nd r l EARLIEST RECORD: Lynfield, AK, 5 n 1975, C. Kuschel. Creophilus oculatus (Fabricius, 1775) REMARKS: In l n l ppin rd n p t a s ee FIRST REPORTED IN N.Z.: J.C. Fabricius, 1775. a ock EARLIEST RECORD: " v Zelandia", 17691770, J. b n & D. Solander. Philonthus rectangulus (S a 1 7 REMARKS: On rr n in p dd nd t h r EARLIEST RECORD: Upp r Mou e e Oc 197 M Esso Gabrius nigritulus (Gravenhorst, 1 REMARKS: In com os i s a i s ee a ocks es e- REPORTED I N.Z.: T. Broun, 1880, nd r th n I S cia y a s e e i g a eas u e ees Staphylinus o ico is Broun. EARLIEST RECORD: Whangarei Harbour, ND, in the Philonthus sordidus (Gravenhorst, 1802) 1 7 Broun. FIRST REPORTED IN N.Z.: T. Broun, 1880, u e the ame im essi o s Broun. REMARKS: On h r . EARLIEST RECORD: r n th 1870s, T. ou Gabronthus sulcifrons (Sh rp 1889) b REMARKS: In l n nd x d com os a ocks a e - EARLIEST RECORD: r p n r v Lynfield, AK, 1 h n 1975, C. Kuschel. REMARKS: In h p p dd a icu a y h r h p h l- Thyreocephalus orthodoxus (Olliff, 1 7 a t r nd r an Acacia mea sii g o e I S REPORTED Came o & C u c e 19 EARLIEST RECORD: a a oe oe AK A 1955 C Gyrohypnus fracticornis (M ll r 1776) Wa FIRST REPORTED C.M. W f ld 1874, a o- REMARKS: In ga e s a a ocks occasio a y i us i us u c u a us Paykull. 1860s, EARLIEST RECORD: A l nd AK n th STAPHYLINIDAE: Euaesthetinae n REMARKS: In h p p dd nd r Acacia mea sii Agnosthaetus brouni Bernhauer, 1939 1833) b REMARKS: On tr b n b ll t rf l nd in Leptacinus pusillus (St ph n r nd-pl nt l tt r EARLIEST RECORD: r p n r v Lynfield, AK 2 b M r 1975, pl 75/16, G. Kuschel. Edaphus beczedesi Reitter, 1914 REMARKS: In lawn l pp n p t bin h p a ock EARLIEST RECORD: Lynfield, AK I8 n 19 1 Ma aise nd r n h r a C Kusc e b ΕΜ ΚS In ll p n s ace in e us 1 m om (Fauvel, 1866) Neohypnus andinus ma gi close to gardens. EARLIEST RECORD: Kaitaia, A 1973, J. Gou es REMARKS: I aw a ga e com os c icke coo a STAPHYLINIDAE: Tachyporinae s ee a ock Sepedophilus acerbus ( ou 1 Neoxantholinus brouni (S a 1 7 REMARKS: Associa e wi Ga o e ma a a a um REMARKS: In i us a ia a t p a o ow Sa i agi is Sepedophilus a'ricapillus ( ou 1 Notolinus socius (Fauvel, 1877) REMARKS: On t nd n t p th Ga o e ma a a- FIRST REPORTED IN N.Z.: T. Broun, 1880 u e e ame a um l n r tt n Co y i e a ksii a o i us a aci us Broun, 1880. EARLIEST E O Whangarei rb r n th Sepedophilus auricomus (Broun, 1 1870s, T. Broun. REMARKS: On s a i g s um s i es e y Ga o e ma a - REMARKS: In x d rd n p t nd h n rd a a um ame es ci a a i a I e e is Au icu a ia o y ic a nd e i us spp.; l on Amanita musca ia Pachycorynusfulvipes (Broun, 1880) REMARKS: One fr d ld in o ow o o K ig ia Sepedophilus austerus (Broun, 1 . e ce sa n th r fr p rtl b r d t p f e yca ya REMARKS: In i e ; a so o a i o e ogs a o ea Sepedophilus brevicornis (Broun, 1 93 Paracorynus arecae (Broun, 1880) REMARKS: In ecaye woo REMARKS: In d d br n h p rt l rl th tt d b Sepedophilus flavithorax ( ou 1 Fig. 1 5 l t p a ica is REMARKS: On a s es ecia y a ksia nd eyci e ia
nd n fr hl br n h n n br n h . K t b l ft r p n Sepedophilus laetulus (Broun, I91 a ntr d d p b f r t r p rt f th p n l nd REMARKS: In litter of r nd pl nt e.g., Ca e U ci ia C nf ld th t p l l t d t x n h n pp 6-8 Sc oe us . 46 y ie ee e Su ey
S A Y I I AE : AC Y O I AE (C nt Atheta politula Fauvel, 1878 FIRST REPORTED IN N.Z.: M. Bernhauer, 1943, as A. splen- Sepedophilus largulus (Broun, 1880) didicollis Bernhauer. REMARKS: On Ganoderma applanatum and in hollow trees; REMARKS: In paddock under Acacia mearnsii trees and in also in cracks of stream banks. stream bed in bush. Sepedophilus maculosus (Broun, 1880) Atheta zealandica Cameron, 1945 a REMARKS: In litter and decayed wood, and on Auricularia FIRST REPORTED IN M Β rnh r 19 3 A pseudo- polytricha. coriaria Bernhauer. Sepedophilus nubilus (Broun, 1880) EARLIEST N.Z. RECORD: Methuen, MC, Dec 1911, T. Hall. REMARKS: In decayed wood. REMARKS: In fermenting fruits, lawn clippings, garden com- Sepedophilus species 1 post, henhouse straw, sheep paddock, a ea is o sea- REMARKS: In a buried tree-fern log. s o e; rarely on Gloeoporus adustus on ea Hedycarya. b Tachyporus nitidulus (Fabricius, 1781) a Atheta species 1 a FIRST REPORTED IN N.Z.: N.A. Martin, 1983. EARLIEST N.Z. ECO Tropicana Drive, Lynfield, AK, I EARLIEST N.Z. RECORD: Owairaka, AK, 14 Oct 1941, D. Jan 1975, G. Kuschel. Spiller. REMARKS: In compost heaps and bins, c icke ya a REMARKS: In garden and paddocks, especially in mixed sheep paddock; also on beach. compost and prunings. Atheta species 2 REMARKS: Under Ptychomnion aciculare moss and at base of STAPHYLINIDAE: Aleocharinae small Carex in bush. Aleochara puberula (Klug, 1833) a b Atheta species 3 b EARLIEST N.Z. RECORD: Haumoana, HB, 26 Jan 1959 Β Μ EARLIEST N.Z. RECORD: Lynfield, AK, 12 Sep 1974, sample May. 74/50, G. Kuschel. REMARKS: On dry carcases. REMARKS: In garden compost. b Aloconota sulcifrons (Stephens, 1832) Atheta species 4 a FIRST REPORTED IN N.Z.: A. Fauvel, 1900, as Atheta pavens EARLIEST N.Z. ECO Tropicana Drive, Lynfield, AK, 2 Erichson. Feb 1975, G. Kuschel. EARLIEST N.Z. RECORD: Auckland, AK, in the 1860s, REMARKS: In paddock under Acacia mearnsii trees. Lawson. Atheta species 5 a b REMARKS: In gravel of stream bed. EARLIEST N.Z. ECO Lynfield, AK, 11 Dec 1976, in Amischa analis (Gravenhorst, 1802) a stream gravel, G. Kuschel. FIRST REPORTED IN N.Z.: A. Fauvel, 1900. REMARKS: In stream bank and gravel. EARLIEST N.Z. RECORD: Riccarton, MC, in the 1870s, C.M. Austrocalea brookesi (Cameron, 1950) a Wakefield. FIRST REPORTED I Μ C r n 195 REMARKS: In compost heaps. EARLIEST N.Z. ECO Avondale, AK, 16 May 1939, ex Amischa decipiens (Sharp, 1869) b h lf f n K Η rrο EARLIEST N.Z. ECO Otaki, WN, 25 Mar 1959, A.C. EMA KS In ga e in compost bins, and in paddock and Eyles. chicken ya REMARKS: In paddock under Acacia mearnsii tree. Botromana vulcanica (Broun, I894) Aphytopus gracilis Sharp, 1886 EMA KS In leaf litter. REMARKS: In litter. Botromana species 1 Aphytopus porosus (Broun, 1912) REMARKS: Οn t nd n d d Hedycarya arborea i es e y REMARKS: In litter at base of bush Carex species and on Gloeoporus adustus; easily baited with garden refuse and Hymenochaete fungus. vegetables placed on bush floor. Atheta amicula (Stephens, 1832) a Botromana species 2 FIRST REPORTED IN N.Z.: A. Fauvel, 1900. EMA KS On rr οn EARLIEST N.Z. RECORD: Auckland, AK, in the 1860s, Crataraea suturalis (Mannerheim, 1830) a b Lawson. EARLIEST N.Z. ECO Timaru, SC, 5 Dec 1941, D. S i e REMARKS: In lawn clippings and garden compost, and in EMA KS In henhouse straw. sheep paddock. Digrammus miricollis ( au e 19 Atheta atriceps (Broun, 1880) EMA KS On tt p r t nυ fοl a i ous a ea o REMARKS: In bush floor litter, especially when enriched with broken branch. garden compost; common on Amanita muscaria in bush Encephalus latulus growing under a Pinus radiata tree. ( ou 1 9 EMA KS In decayed wood and bush floor litter. Atheta coriaria (Kraatz, 1858) Falagria concinna (Erichson, 1 b FIRST REPORTED IN N.Z.: A. Fauvel, 1900. EARLIEST N.Z. RECORD: Auckland City, AK, 6 Jul 1949, in EARLIEST N.Z. RECORD: Auckland, AK, in the 1860s, Law- son. packing straw, ?N.R. Peace. REMARKS: A single specimen on ground in garden. REMARKS: In compost bins, chicken yard, and inflorescence of an ornamental Musa species. Gyronotus rufipennis (Broun, 1880) EMA KS Halophilous; on seashore. Atheta cottieri Cameron, 1945 FIRST REPORTED IN Μ C r n 19 5 Gyrophaena oligotina Came o 19 5 EARLIEST N.Z. RECORD: Palmerston North, WI—WN, Feb EMA KS In decayed wood. 1930, W. Cottier. Gyrophaena punctata Βrο n 1 REMARKS: In garden compost, sheep paddock, and stream EMA KS In stumps of Sophora microphylla a Melicope bed, and on beach. ternata, and in epiphytes o Salix fragilis. Ann t t d t f Sp 47
Gyrophaena p 1 . Silusa paralkk ( rnh r 1943) REMARKS: In d d d REMARKS: In l f l tt r Gyrophaena p Silusa puber (Broun, 1880) REMARKS: On d d d th Fuscoporia dryophila nd REMARKS: In l f l tt r nd l nd r Dacrycarpus ac y- Phellinus pp ioi es
Gyrophaena p 3 Silusa p 1 . REMARKS: In d d d nd n Ca e nd Uncinia l tt r REMARKS: In l f l tt r n b h fl r Silusa p 2 Gyrophaena p REMARKS: In d ld n d h ll Dysoxylum s ec a- REMARKS: In d d Melicytus ramiflorus d nd n i e Alseuosmia, Brachyglottis, nd Myrsine l tt r Stenomastax dentata C r n 1945 Gyrophaena p 5 REMARKS: In d d d REMARKS: In d d d Sternotropa versicolor (Broun, 1880) Gyrophaena p 6 REMARKS: r G oeo o us a us us on Hedycarya arborea REMARKS: On r tt n l th Ganoderma applanatum. nd fr Tricholoma pessundatum. Halibrecta flavipes ( h n 1861) a Sternotropa zealandica (C r n 1947) EARLIEST RECORD: r p n r v Lynfield, AK 27 REMARKS: O usco o ia yo i a nd e i us sp. 1976, n r C. Kuschel. Sytus aerarius (Broun, 1880) REMARKS: ln r n h r REMARKS: On yso y u s ec a i e a Me icy us ami- Ischnoglossa bituberculata (Broun, 1894) ο us s ems REMARKS: In r tt n Pinus radiata t p Sytus flavescens (Broun, 1880) Ischnoglossa pectinata C r n 1945 REMARKS: On r tt d Acacia ecu e s . REMARKS: In d d d . Sytus granifer (Broun, 1894) Myllaena intermedia (Erichson, 1837) a REMARKS: On Hedycarya s em i es e y G oeo o us EARLIEST RECORD: S xt n Molesworth, 1067 m, adastus, n Knightia th ame es ci a a i a a n MB, 17 A 1966, pl 66/249, n n t b n d n eyci e ia a so n r v l f s eam b d A.K. W l r REMARKS: In tr b d n v r d p l tt r f t r h n- Sytus p n l ; r d l bt n d n M l tr p REMARKS: On Ganoderma applanatum nd t i es e MyIlaena magnicollis (C r n 1946) y Irpex e is . REMARKS: In b h fl r l tt r Tramiathaea cornigera (Broun, 1 REMARKS: Nehemitropia sordida (Marsham, 1802) In l f i e ; es ecia y commo w e i e FIRST REPORTED IN A Fauvel, 1900. mi e with i e ui s o Co y oca us ae iga us r EARLIEST RECORD: " l nd" b f r 1900. baited with garden refuse and ege a es REMARKS: In h p p dd p ll nd r Acacia G n p a . mearnsii tr EARLIEST ECO r p n r v Lynfield, AK 14 Ocalea fuscicornis (Broun, 1880) M r 1975, pl 75/ C Kusc e
REMARKS: In l f l tt r . REMARKS: In l n l pp n th f r nt n fr t Ocalea socialis (Broun, 1880) G n p 2 b REMARKS: In l r n p r r l n r l t r nd EARLIEST ECO Lynfield, AK 21 I980, G. Oligota fungicola W ll 1976 Kuschel. . REMARKS: t n fr v t t n REMARKS: tt d n M l tr p
. Oligota parva (Kraatz, 1858) G n p 3 FIRST REPORTED IN S A W ll 1976. EMA KS In h d s eam b d . EARLIEST RECORD: l n NN, 2 M 1966, M.J. G n 1 p 4 Esson. . REMARKS: In stream e nd n p ph t r n n Sa i REMARKS: In rd n p t r d h p p dd ; l n agi is b h fl r l tt r u e A seuosmia a ac yg o is nd G n p 5 a in nd l v r rt b v tr b n nd r d n n p REMARKS: In rd n p t nd n c icke ya Oligota setigera (W ll 1976) G n 2 p REMARKS: t n n l fr v t t n REMARKS: In t pl h z n f h d b h stream Oligota speculicollis (C r n 1945) G n 3 p REMARKS: One n d d d nd n th r n l tt r fr REMARKS: Mos o aine om si e ecaye woo o ac- t l l p yca us ac y ioi es wo associa e wi ame es ci - Paraphytopus p 1 a a i a n K ig ia e ce sa . REMARKS: On s eam b d nd n d p l tt r G n 4 p Polylob sternalis (Broun, 1 EMA KS In ecaye woo a ase o sma Carex i us REMARKS: On I e e is Ga o e ma a a a um a a i ga e ea i e Hygrophoraceae. G n 5 p 1 REMARKS: b h K t b l ft r p n a G n p i o uce s ecies b i s e o o is s ecies i ew ea a REMARKS: C I ecaye So o a Me icy us yso y um nd y ie e y e oca i y d a o omic c a ge; see 7 - 1 Cyathea; l n Hymenachaete f n nf ld tl S rv
PSELAPHIDAE El t species . `d rf b tl REMARKS: Und r Gahnia setifolia. Euplectopsis nt ( r n 1 93 REMARKS: In d ld f h ll Sophora microphylla nd Metrosideros excelsa tr Euplectopsis opaca (Sh rp 1 7 EMA KS In l f l tt r Euplectopsis ovicollis ( r n 1 EMA KS In tr b d nd n Carex lessoniana p Euplectopsis tuberigera ( r n 1 EMA KS In d d d nd n l nd r r tt n l Euplectopsis p EMA KS In n d p d n d 5 Sagola l ph d laminata Leptoplectus species REMARKS: In d d d f Dacrycarpus dacrydioides PSELAPHIDAE: Faroninae nd t b f ld Cupressus macrocarpa t p v r d b Exeirarthra enigma Βr n 1 93 Pennisetum clandestimum r n r b h r n REMARKS: In d d Knightia excelsa, nd n d ld Leptoplectus species l t d n h ll f l v Dysoxylum spectabile. EMA KS In d d Vitex. Exeirarthra planicula ( r n 19 1 Microtyrus iracundus ( r n 1 93 REMARKS: In l r nd Hedycarya arborea stump. EMA KS In l f l tt r Sagola duplicata r n 1 Microtyrus punctatus Βr n 1 93 REMARKS: In l r nd Sophora microphylla t p nd REMARKS: A n nd l v r rt n t t p l p nd r Gahnia lacera. b v tr b n Sagola excavata Βrο n 1 M r t r species REMARKS: In d d d EMA KS Und r Gahnia setifolia. Sagola genalis r n 1 1 r pl t species EMA KS Und r t l Gahnia lacera nd t b f Carex EMA KS In d ld f h ll Sophora microphylla flagellifera. tr b Sagola laminata Broun, 1 93 5 1 Philiopsis species 1 a REMARKS: In l f l tt r; r n n t d f th EARLIEST N.Z. RECORD: r p n r v nf ld AK 12 n r Gahnia, Carex, Morelotia, nd Uncinia. S p 197 n x d p t G K h l Sagola longipennis Βrο n 1911 REMARKS: In rd n p t nd p l d- p pr n n n EMA KS In d d d n h ll tr nd n t l h n rd nd nd r Pinus radiata n b h l tt r Plectomorphus sculpturatus (Broun, 1880) Sagola major Sh rp 1 7 REMARKS: In d d d f n rl ll tr p EMA KS Und r Phormium tenax n r th f W ttl Β Plectomorphus trisulcicollis ( r n 1 stream. REMARKS: In Dendroligotrichum nd Schoenus, Uncinia, and Carex Sagola monticola Broun, 1912 d REMARKS: In d d d Whitea species 1 . Sagola notabilis r n 1 REMARKS: In d d d EMA KS In d d d Zelandius clevedonensis (Βr n 1 93 Sagola species REMARKS: In l tt r nd n d ld f h ll tr Dysoxylum spectabile. REMARKS: In d d wood. Sagola species 2 Zelandius obscurus ( r n 1 93 REMARKS: In d ld f h ll Dysoxylum tr REMARKS: On b h nd n n t l t l v t t n Gahnia lacera l tt r nd tr b d Sagola species 3 Zelandius sandageri ( r n 1 93 REMARKS: t n fr d d tr -f rn fr nd EMA KS In d ld f l r h ll Metrosideros Sagola species 4 excelsa, nd n Gahnia setifolia l tt r REMARKS: Und r Gahnia setifolia. Genus 1 species 1 EMA KS In l tt r f t r h nn l nd n nd r PSELAPHIDAE: Euplectinae Alseuosmia and Brachyglottis. Alloplectus claviger ( r n 1 93 EMA KS In l f l tt r PSELAPHIDAE: Brachyglutinae Alloplectus species 1 Anabaxis electrica (K n 1 3 d EMA KS r l tt r pl f ll Carex lambertiana, I S REPORTED IN r n 19 1 nd r th n n C. dissita, nd Uncinia banksii nd r d n n p . A minor Broun. Alloplectus species 2 EA IES RECORD: r C n th 1 7 r n EMA KS A n t nd l h n n tr tr n nd n REMARKS: In a p dd l tt r l t d nd r G hn t f l ndMetrosideros Anabaxis foveolata (Broun, 1880) d excelsa. REMARKS: In Oplismenus imbecillus and Microlaena ave- El t ll ph l (Broun, 1893) nacea t Dendroligotrichum dendroides l f REMARKS: In leaf litter. l tt r nd n Gahnia setifolia. Ann t t d t f Sp 49
Eupines altula (Broun, 1880) REMARKS: M t n d ld f l r h ll Vitex lucens tr ; n h ll Salix fragilis b W ttl tr Eupines glabrata (Broun, 1886) REMARKS: In l f l tt r nd r tt n d; p ll n n l tt r f t l Astelia banksii nd Ga ia lacera. Eupines r t (Sh rp 1874) REMARKS: In tr b d Eupines impar (Sh rp ?1874) REMARKS: Ev r h r n th r nd n l f nd br n h l tt r nd n d nd r r t Eupines punctata (Broun, 1886) 26 Adolopus Hydrophilidae REMARKS: In h ll i e uce s tr nd in d d p r sp. 1 Macropiper excelsum. t nd n t of b Cercyon analis (Paykull, 1798) Eupines species 1 EARLIEST RECORD: Ardagh, Methven, MC 14 b REMARKS: In lt d b C r x lessoniana p 1912, ll Eupines species REMARKS: In l n l pp n rd n p t p ltr tr REMARKS: In t l Gahnia lacera l tt r h n rd nd h p p dd Gastrobothrus abdominalis (Broun, 1880) b Cercyon atricapillus (Marsham, 1802) REMARKS: In (Carex, l f l tt r n t ll r nd pl nt EARLIEST RECORD: Mangere, AK Apr 1931 Κ Ρ U ci ia O isme us nd r Gahnia se i ο ia nd n b K ig ia e ce sa d d REMARKS: In fr h h p d n Gastrobothrus sharpi (Broun, 1880) Cercyon depressus St ph n 1829 b REMARKS: On th r nd n l tt r r nd d ; EARLIEST RECORD: r p n r v Lynfield, 27 l in h ll tr nd d d d 1976, G. Kuschel. Physobryaxis inflata (Sh rp 1874) REMARKS: Ιn r nd nd r d d n l n beach. REMARKS: r r tt n K ig ia l Cercyon haemorrhoidalis (Fabricius, 1775) Startes sculpturata Broun, 1886 FIRST REPORTED IN N.Z.: C.M. W f ld 1874, nd r th REMARKS: In d ld f h ll Me osi e os e ce sa n n C a i es (F.). EARLIEST RECORD: Greymouth, PSELAPHIDAE: Tyrinae A l nd nd n th b 1870s, l Gerallus punctipennis Schaufuss, 1879 REMARKS: In rd n p t n th p n nd n bins, s ee EARLIEST RECORD: r p n r v Lyrifield, AK 2 u g i a ock a us i e e ic e wi ui s a A 1975, n l nd r ac yca us ac y ioi es G. ga e e use Kuschel. Cyloma lawsona REMARKS: In fr bl l ft n usi g Meso o e a cas a ea S a 1 7 r n b t n t t d d r tl th th nt l n REMARKS: In leaflitter, p ll f e ic e with i e Cory- oca us fr t r b t d th eac es i ea e -u us Hamotulus mutandus (Sh rp 1874) oo i e ; a so i us a swam se ges REMARKS: In tr l tt r nd n tr b n Dactylosternum bd n l (Fabricius, 1792) Zeatyrus lawsoni Sh rp 1881 FIRST REPORTED Ν N.Z.: D.H. dd 1961. REMARKS: In d nd r ; l in d d EARLIEST RECORD: Owairaka, AK, S p 1940, nd r d nd h ll tr . n Spiller. PSELAPHIDAE: Pselaphinae REMARKS: In rd n com os ou y s aw i e - ouse a c icke ya a o a ima ca cases Pselaphus citimus (Broun, 1893) REMARKS: In nd nd r r tt n l Dactylosternum marginale (S a 1 7 a FIRST REPORTED Ν N.Z.: D. Sh rp 1 7 Pselaphus delicatus Broun, 1886 EARLIEST RECORD: A l nd AK n th ea y 1870s, REMARKS: In Sc oe us e o d nd ec um ca e se awso . f rn l tt r ,. REMARKS: In p t b n Pselaphus dulcis Broun, 1881 Enochrus tritus (Broun, 1880) REMARKS: At th b f l r i e uce s tr n 127 FIRST REPORTED ΙΝ Ν r n 1 Pselaphus p p r Sh rp 1874 EARLIEST RECORD: " n d ff r nt oca i ies" [ e wee REMARKS: In t f o y ic a e us mage a icus Whangarei ND a ai ua C ] n th 1870s, T. Broun. nd yc om io acicu a e n ll l n l tt r REMARKS: In a p nd Kn n l fr th K r d I S r t n nd h t HYDROPHILIDAE Exydrus flavicornis (Broun, 1880) t r v n r b tl REMARKS: In l f nd t l tt r fr bl l nd c acks n tr b n Adolapus altulus (Broun, 1880) Hydrostygnus frontalis REMARKS: In l f l tt r ll r nd pl nt fr bl l ( ou 1880) nd d ld th r d n h ll tr REMARKS: In decayed wood, particularly i i es e y Irpex brevis f n ( l p r K t b l ft r p n Rygmodus modestus a Wh t 1 ntr d d p b f r t r p rt f th p n l nd REMARKS: C On fl r n Cya o es ascicu a a b t t nf ld th t p l l t d t x n h n pp 6-8 ht n M l tr p 5 y ie ee e Su ey
HISTERIDAE i ee es (assis e y Y Gomy nd G a s ea
28 Clambus Cl b d domesticus
ig 7 Tomogenius Sphaerothorax suffusus ( ou 1 is e i ae latipes EMA KS I i e a Acritus nigricornis ( o ma 1 3 a Sphaerothorax tierensis ( ack u 19 I S E O E I EA IES ECO o ica a i e y ie AK 19 S E ö y-You ga 199 u 197 G Kusc e EA IES ECO a a gi M Se 19 9 i e sam- i os o um Me icy us EMA KS I com os i e 9/1 9 ake i a sc u a e oo o a ig coas a c i G Kusc e Carcinops pumilio (E ic so 1 3 EMA KS I ea o Euca y us a c es a cu am oo i I S E O E I Wa ke 19 u e e sy o- s ee a ock e ese e i AC y s ecime s om ym C qua uo ecims ia us (S e e s a i a i io o AK a M EA IES ECO y e o MC o 19 1 Wa ke EMA KS I aw c i i gs ga e com os ou y s aw HELODIDAE i e - ouse a c icke ya a o ca case i s ee ma s ee es a ock (assis e y y o m Gnathoncus rotundatus (Kuge a 179 a I S E O E I Wa ke 19 as G. nanneten- sis (Ma seu EA IES ECO akaia MC Ma 19 Wa ke EMA KS I e - ouse s aw; a so i ie ca case Gnathoncus schmidti ei e 1 9 a EA IES ECO Kumeu AK 31 u 1975 i ow ma u e C Wa EMA KS I e - ouse a c icke ya s aw a ma u e Parepierus abrogatus ( ou 1 EMA KS U e a iece o woo Parepierus purus ( ou 1 ig 9 Veronatus e o i ae tricostellus EMA KS I us oo i e i ecaye woo mou i si e o ow ees o u ks o e yca ya a o ea i es e y G oeo o us a us us a o u ks o My si e aus a is Cyphanus punctatus S a 1 7 wi Ga o e ma a a a um EMA KS O ai e y ea i g Platysoma bakewelli Ma seu 1 a Cyphon genalis S a 1 7 I S E O E I S a 1 7 u e e sy o ym EMA KS O a s ea es wa e cog a um S a Cyphon graniger S a 1 7 EA IES ECO Auck a AK i e 1 7 s EMA KS O a s y s eams awso Cyphon huttoni S a 1 7 EMA KS I ga e com os EMA KS O a s ea es wa e Saprinus detritus ( a icius 1775 Cyphon oscillans S a 1 7 EMA KS O ea a ima s i c u i g is o eac es EMA KS O a s y s eams Cyphon parviceps S a 1 7 CLAMBIDAE EMA KS O a s ea s eams; a icu a y a u a o c am ee es o a os y is sa i a owe s (assis e y S E ö y-You ga Cyphon remotus ou 1 EMA KS O a s ea s eams Clambus domesticus Broun, 1 a Fig. 28, 128 Cyphon suffusus S a 1 7 I S E O E I ou 1 EMA KS O a s ea es wa e EA IES ECO ai ua C i e ea y 1 s "o - e wi ows o my ouse" ou Cyphon viridipennis ou 1 EMA KS I aw c i i gs ga e u i gs a com os ; EMA KS O a s ea s eams occasio a y i a i e us i e Cyphon waikatoensis ou 1 EMA KS O ege a io y s eam Sphaerothorax kuscheli Endröd -Y n EMA KS O o ee s um s a i oose moss ma s o Cyphon zealandicus S a 1 7 o y ic aceae EMA KS O ege a io y es wa e Ann t t d t f Sp 51
Cyphon p 1 REMARKS: On pl nt n r fresh water. Cyphon p 2 REMARKS: On pl nt n r tr Cyphon p 3 REMARKS: On pl nt n r fr h t r Cyphon p 4 REMARKS: On pl nt n r tr Cyphon p 5 REMARKS: On pl nt n r tr
Cyphon p 6 31 Costelytra REMARKS: On pl nt b fr h t r Scarabaeidae zealandica Cyphon p 7 REMARKS: In fr h h p d n REMARKS: On plants near streams. Aphodius tasmaniae p 1847 Fig. I30 Cyphon p 8 Ι SΤ REPORTED IN N.Z.: W. Cottier, 1956, nd r th n - REMARKS: On pl nt b fr h t r n A howitti Hope. Cyprobius nitidus Sh rp 1878 EARLIEST RECORD: Mt Gr MC 9 S p 1916, S. REMARKS: On v t t n n r tr nd G n 1 p 1 REMARKS: In rd n p r and paddocks. REMARKS: On plants near tr Ataenius brouni (Sharp, 1876) G n 2 p 1 REMARKS: In tr b d REMARKS: S p n t n ht Ataenius picinus r ld 1861 FIRST REPORTED IN A Μ Richards, 1959, under the LUCANIDAE synonym Saprosites rugosus Richards. EARLIEST RECORD: Auckland, 4 1939, on 'stag beetles' AK M decaying banana squash, D. Spiller. (assisted by Β A. Holloway) REMARKS: In dung, garden compost, and fermenting fruit . Saprosites communis (Broun, 1880) REMARKS: In r tt n d f M l t r flοrυ - x l p t b l t x l n n r d t n th b h Dacrycarpus dacrydioides, Hedycarya arborea, and Pseudopanax arboreus.
Saprosites mendax (Blackburn, 1892) a FIRST REPORTED IN N.Z.: A Μ Richards, 1959, under the synonym S punctatus Richards. EARLIEST RECORD: Owairaka, AK, 7 May 1943, D. McKenzie. REMARKS: In garden compost nd prunings; common in 30 Ceratogna us rotten wood of Pseudopanax, Hedycarya, Melicytus, and Lucanidae parrianus Dysoxylum, often together with the endemic S communis. Ceratognathus irroratus ( rr 1845) SCARABAEIDAE: Scarabaeinae . REMARKS: Ad lt n l nd tr n l rv n d n d h d lt r l tr pp d n M l n tt n 'dung beetles' . Ceratognathus parrianus (W t d 1863) Fig. 30, 129 Onthophagus granulatus h n 1858 a FIRST REPORTED IN N.Z.: C.M. Wakefield, 1873. REMARKS: Ad lt n tr n nd br n h l rv n nd r t d d d Ad lt l M l tr pp d EARLIEST RECORD: l n rl 1872, R.W. Fereday. REMARKS: I sheep dung. Saphobius edwardsi Sharp, 1873 SCARABAEIDAE REMARKS: In stream bed and bank. `scarab beetles' Saphobius squamulosus Broun, 1886 SCARABAEIDAE: Aphodiinae REMARKS: In leaf litter, wood mould of hollow trees and 'lesser dung beetles' tr n nd r d aying logs, and in old, dry sheep carcases. Aphodius granarius ( nn 1767) SCARABAEIDAE: Melolonthinae FIRST REPORTED IN C.M. W f ld 1874. 'chafers' EARLIEST RECORD: C nt rb r 1872, C.M. W f ld REMARKS: In h p p dd (assist d by J. C. W tt Aphodius pseudolividus (Balthasar, 1941) a Costelytra mac obrunnea Given, 1952 FIRST REPORTED I N.Z.: J.G. r n 1967. REMARKS: In gardens, and Malaise trapp d n native bush. EARLIEST RECORD: Karekare, AK b 1916, A Ε Costetytra zealandica (White, 1846) ' Fig. 31 r REMARKS: More common in paddocks and gardens than in
native bush. . . K t b l ft r p n Odontria xanthosticta White, 1846 a ntr d d p b f r t r p rt f th p n l nd REMARKS: C Adults year-round n bush, paddocks, and garden; nf ld th t p l l t d t x n h n pp 6-8 occasionally hiding in numbers in coastal caves. 2 y ie ee e Su ey
SCARABAEIDAE ME O O I AE (co
Pyronota laeta (Fabricus, 1775) REMARKS: Particularly common on Leptospermum scopar- ium; active in the daytime. Sericospilus costella (Broun, 1880) REMARKS: One beaten, two in Malaise trap. Stethaspis longicornis (Arrow, 1924) REMARKS: Flying in the early evening.
SCARABAEIDAE: Dynastinae `horned scarabs' Fig. 34 y oc y us Ptilodactylidae u qu a i Heteronychus arator (Fabricius, 1775) Fig. 131 FIRST REPORTED IN N.Z.: G.H. Cunningham, 1940, under the Genus 1 species 1 name Pentodon australe. REMARKS: Underneath overhanging stream banks and EARLIEST N.Z. RECORD: Waiheke I., AK, Mar 1937, G. amongst fern and moss on a steep, seeping slope. It flies Chamberlain (fide D. Spiller and E.G. Turbott, 1944). between 15 Feb and 25 Mar. Larvae terrestrial, in soil of REMARKS: In garden and paddocks. seeping banks covered with ferns, moss, and liverworts.
BYRRHIDAE ELATERIDAE `moss beetles' `click beetles' ( t d b C Μ v n nd C W tt
Fig. 32 Liochoria Byrrhidae huttoni Fig. 35 o amus Synorthus rotundus (Broun, 1881) Elateridae wake ie i REMARKS: On old logs. Synorthus setarius (Broun, 1880) Aglophus modestus Sharp, 1877 REMARKS: On fairly damp ground and stream beds with moss REMARKS: Beating vegetation. and liverworts. Agrypnus variabilis (Candèze, 1857) a I S E O E I Sh rp 1 77 BUPRESTIDAE EARLIEST N.Z. RECORD: Auckland, AK, in the early 1870s. REMARKS: Associated in the main with Acacia and Eucalyp- `jewel beetles' tus species. Amphiplatys lawsoni Sharp, 1877 REMARKS: Common in bush litter and on low-growing plants at night. Conoderus exsul (Sharp, 1877) FIRST REPORTED IN N.Z.: D. Sharp, 1877. EARLIEST N.Z. RECORD: Wellington, WN, Feb 1875, Wake- field. REMARKS: In paddocks and gardens, sometimes coming to light in large numbers. Conoderus maritimus (Broun, 1893)
Fig. 33 Nascioides REMARKS: Readily Malaise trapped. . Buprestidae e ysi Conoderus posticus (Eschscholtz, 1822) a EARLIEST N.Z RECORD: "Bayleys Coast" [Baylys Beach], Maoraxia eremita (White, 1846) r v ll v 19 Β G v n REMARKS: Attracted to cut branches of Myrsine australis and REMARKS: In paddocks and gardens, leaf litter and prunings, successfully reared from these. and under bark. Conoderus submarmoratus (Macleay, 1872) a b PTILODACTYLIDAE EARLIEST N.Z. RECORD: Grey Lynn, AK, Feb 1958, R.W. `comb-waist beetles' Taylor. Byrrocryptus species 1 REMARKS: In gardens. REMARKS: Malaise trapped at stream; caught only between 11 Ctenicera antipoda (Candèze, 1863) Jan and 3 Feb. Larvae aquatic. REMARKS: In Malaise trap. Ann t t d t f Sρ 53
Ctenicera strangulata (Wh t 1 EMA KS t n ff v t t n M l tr pp d nd t n n d p l d- ff p t-tr p Lomemus elegans Sh rp 1 77 EMA KS On fr ft d d n d n b t n ff v t t n Lomemus flavipes Sh rp 1 77 EMA KS On v t t n Lomemus fuscicornis r n 1 93 EMA KS On b t n v t t n th r M l tr pp d Lomemus lateristrigatus (Wh t 1 Fig. 37 o os oma EMA KS r d fr Knightia excelsa d Lycidae u i e e Lomemus similis Sh rp 1 77 EA IES ECO Drury, Pokeno, and Clevedon, AK, EMA KS S b b t n v t t n t n M l tr p n th 1 T. r n Lomemus suffusus Sh rp 1 77 EMA KS S n fl n n th d t th frequency. EMA KS In M l tr p Metablax cinctiger (Wh t 1 CANTHARIDAE EMA KS Off v t t n nd n M l tr p ` ld r beetles' Metablax approximans (Wh t 1 EMA KS C ht n fl ht rl n th rn n nd n ( t d b W W tt r M l tr p Ochosternus zealandicus (Wh t 1 EMA KS Ad lt n th n p l rv n d n d Panspoeus guttatus Sh rp 1 77 EMA KS S p n t n ht C n n th A l nd r AK nd n ll t bl h d n En l nd (f r t d - v r d 19 l 19 1 W nd r Gr t r r h r Parinus villosus (Sh rp 1 77 EMA KS t n ff v t t n Thoramus perblandus r n 1 EMA KS t n ff v t t n Fig. 38 Asi is Thoramus wakefieldi Sh rp 1 77 35 13 Cantharidae fulvithorax EMA KS r d fr i us a ia a b rd b nd n d n n t v b h Asilis dugdalei W tt r 1979 EMA KS On o e ia o u ea and in Malaise trap. EUCNEMIDAE Asi is piliventer ( ou 1881) `f l l b tl EMA KS On b t n ff v t t n n v t n igus um si e se fl r n swam t b f W ttl Malthodes pumilus ( r b n 1 35 I S E O E I W W tt r 1979 as Ma o es p EA IES ECO rt n h Chr t h r h MC
1 19 1 E S G rl EMA KS Sweeping grass nd d t b f W ttl b h Kn n nl fr f l n l nd (AK HB, MC, and CO
DERMESTIDAE `h d beetles' Fig. 36 eoc a is Eucnemidae sim e
Dromaeolus nigellus (Wh t 1 EMA KS Und r b r f i us a ia a l l n n th b h Talerax distans Sh rp 1 77 EMA KS In d d d
LYCIDAE `net-winged b etles'
Porrostoma rufipenne (Fabricius, 1801) Fig. 37, I33 FIRST REPORTED I Ν r n 1 93b nd r th n - Fig. 39 ogo e ma n Metriorrhynchus erraticus Broun. Dermestidae maes um
b K t b l ft r p n Anthrenocerus australis ( o e 1843) a i o uce p b f r t e o o is sρecies i ew ea a EARLIEST N.Z. ECO Dunedin, DN, 22 Feb 1942, D. c nf ld th t p l l t a o omic c a ge; see 7 - 1 Spiller. 4 nf ld tl S rv
E MES I AE (co Methemus griseipilus (Broun, 1881) .. REMARKS: On akea se icea and Acacia mea sii n n t v ΕMA KS Ιn rd n n Pseudopanax arboreus fl r b h Anthrenus verbasci (Linnaeus, 1767) Microsternomorphus oblongus (Broun, 1880) FIRST REPORTED IN N.Z.: F.W. tt n 1904, as A. museorum REMARKS: E ht p n r r d fr Acacia ecu e s (n Linnaeus, 1761). Xenocera sericea Broun, 1880 EARLIEST RECORD: Chr t h r h n th 1880s, F.W. REMARKS: On As e ia a ksii tt n REMARKS: O n ll n fl r f Zantedeschia, Dau- cus, Foeniculum, Achillea. PTINIDAE Dermestes haemorrhoidalis Kuester, 1852 b p d r b tl EARLIEST RECORD: Eltham, TK, b 1932, fr t r d lv t h REMARKS: Ind r Dermestes maculatus DeGeer, 1774 a FIRST REPORTED IN N.Z.: F.W. tt n 1904. EARLIEST RECORD: Chr t h r h MC n th 1880s. REMARKS: On ld r rr n Τr h l d v l t (Broun, 1880) REMARKS: On t t o Parsonsia nd Carmichaelia; n r r d fr d d culms f Gahnia lacera. Trogoderma granulatum Broun, 1881 REMARKS: In M l tr p 41 i us Ptinidae Trogoderma maestum Broun, 1880 Fig. 39, I34 s eciosus REMARKS: On Ligustrum sinense fl r nd n l Ptinus speciosus Broun, 1880 Fig. 41, 135 Trogoderma signatum Sh rp 1877 REMARKS: Und r Co a e ia u a a n y o s o REMARKS: On fl r f Pseudopanax arboreus a Lepto- Cya ea me u a is nd n l f l tt r o Alseuosmia macro- spermum scoparium; n l nd t p phylla.
TROGOSSITIDAE - ANOBIIDAE - ` cadelle ee es b r r b tl ( t d b Ε ρ ñ l
42 p d pt r x Trogossitidae n r p r 40 Hadrobregmus Anobiidae magnus p d pt r x br n ( 1876) EMA KS In d d d n td r pl d nd n Anobium punctatum (DeGeer, 1774) h ll Salix fragilis. FIRST REPORTED IN N.Z.: T. Broun, 1880, nd r th synonym p d pt r x n r p r 1846) A amplicolle Broun. (Wh t Fig. 42, 136 REMARKS: On t Phyllocladus trichomanoides br n h EARLIEST RECORD: Tairua, CL, n th 1870s, T. Broun. REMARKS: In th llb rd f ld h d nd h ; n Cu - essus macrocarpa r d C. o u osa n p r PELTIDAE Dorcatoma pilosa (Wh t 1846) s ie ee es REMARKS: r pped t n ht Hadrobregmus australiensis 1901 a Australiodes vestitus (Broun, 1881) FIRST REPORTED IN N.Z.: F. Español, 1976. REMARKS: On v t t n; ttr t d b ece y cu a c es EARLIEST RECORD: Titirangi, AK 14 v 1937, A J. h n n tr Hipwell. Rentonellum p 1 REMARKS: Ad lt nd l rv n d f Cu essus o u osa REMARKS: In decayed wood of ac yca us ac y ioi es Acacia mea sii i us a ia a Me osi e os e ce sa and Me icy us am Ιο υs nd e yca ya a o ea Cya o es asicu a a l n td r plywood. ntοn ΙΙυ p 2 Hadrobregmus crowsoni Español, 1976 REMARKS: In i e om g ou co e e wi O isme us REMARKS: In n r t r f l r i e uce s ee im eci us a Mic o ae a a e acea g asses Holcobius watti Español, I982 Rentonellum p 3 REMARKS: t n ff Pinus a ia a and Co a e ia u a a REMARKS: In r tt n d Ann t t d t f Sp
PHYCOSECIDAE `beach beetles'
Fig. 43 Rentonidium Peltidae costiventre 5 Phycosecis Rentonium species 1 h d limbata EMA KS Under Gahnia lacera, in decayed wood. Rentonium species 2 Phycosecis limbata (Fabricius, 1781) 45, 137 EMA KS On beach sand. EMA KS In stream bed moss. Genus 1 species 1 EMA KS In leaf litter from under Phyllocladus trichoman- MELYRIDAE oides and Olearia furfuracea, and in rotten wood. `flower beetles' Genus 2 species 1 EMA KS On bark of thick branches of trees, p ll
Dysoxylum spectabile, Vitex lucens, and Schefflera digi- tata, often congregating.
CLERIDAE `checkered beetles'
Fig. 46 Dasytes Melyridae subcyaneus
Dasytes laticeps Broun, 1 . EMA KS On v t t n Dasytes planifrons (Broun, 1 3 EMA KS On v t t n; v t r to fl r nd ttr t d n n b r t Cyathodes fasiculata fl r h n Arthracanthus Βrο n 1 3 pr p d b S h rd
Fig. 44 Phymatophaea 1 5 . Cleridae violacea Dasytes stewarti Broun, 1881 EMA KS Two taken in Malaise trap. Necrobia ruficollis (Fabricius, 1775) a Ι SΤ E O E IN N.Z.: F.W. Hutton, 1904. Dasytes subcyaneus Broun, 1880 Fig. 46, 138 EA IES ECO Christchurch, MC, in the 1880s, EMA KS On vegetation. F.W. Hutton. Dasytes species 1 EMA KS O carrion. EMA KS On Carex virgata in swamp at back of Wattle B Necrobia rufipes (DeGeer, 1775) a nd r n p f Salix fragilis. I S E O E I N.Z.: J.J. Walker, 19 Hatyles nigrescens Broun, 1883
EA IES ECO Auckland, AK, 1875, T. r n EMA KS On v t t n . EMA KS On rr n Halyles semidilutus Broun, 1 3 Parmius longipes Sharp, 1877 EMA KS On Hoheria populnea.
EMA KS On a plant. . Paupris aptera Sharp, 1877 NITIDULIDAE EMA KS On Gahnia lacera at night; one reared from culms `sap beetles' of this sedge species. (assisted by R. M. Dobson, L. R. G ll l Phymatophaea opiloides Pascoe, 1876 R. D. Pope, and R. T. Thompson) EMA KS Scattered on vegetation. Carpophilus davidsoni Dobson, 1952 a EA IES ECO Avondale, Auckland, AK, 14 b 1941, D. Spiller. EMA KS On flowers of Ligustrum sinense and rotten wood. Carpophilus dimidiatus (Fabricius, 179 b K t b l ft r p n EA IES ECO Greymouth, WD, in th 1 7 R. a ntr d d p b f r t r p rt f th p n l nd Helms. nf ld th t p l l t d t x n h n pp 6-8 EMA KS On almonds. 6 nf ld tl S rv
b I I U I AE (cont.) Thalycrodes australis Blackburn, 1891 a EARLIEST N.Z. RECORD: Lynfield, AK, 17 Jun 1978, G. Kuschel. REMARKS: On fermenting fruits in garden.
RHIZOPHAGIDAE 'root beetles'
Fig. 47 Platipidia Nitidulidae asperella Carpophilus gaveni Dobson, 1964 a b EARLIEST RECORD: Whangarei, ND, 12 Apr 1929, Ε Fairbum.
REMARKS: On fermenting fruits and vegetables. Fig. 48 M n t Rhizophagidae Carpophilus hemipterus (Linnaeus, 1758) spinicollis FIRST REPORTED IN N.Z.: F.W. Hutton, 1904. Monotoma longicollis Gyllenhal, 1827 EARLIEST RECORD: Christchurch, MC, in the 1880s, FIRST REPORTED IN G. Kuschel, 1979. F.W. Hutton. EARLIEST RECORD: Mt Roskill, AK, 10 Jan 19 Ε REMARKS: On fermenting fruits and vegetables. Brookes. b Carpophilus marginellus Motschulski, 1858 REMARKS: In lawn and mixed compost, arid n sheep shelters EARLIEST RECORD: Nelson, NN, 4 May 1935 Ε S under an Acacia mearnsii grove. Gourlay. M n t p p Herbst, I793 REMARKS: On fermenting fr it and vegetables. FIRST REPORTED IN N.Z.: F.W. Hutton, 1904. Epuraea antarctica (White, 1846) EARLIEST RECORD: Chr t h r h MC, in the 1880s, REMARKS: On old sheep carrion at bush margin. F.W. Hutton. REMARKS: In garden compost and litter sheep paddock, Epuraea imperialis (Reitter, 1877) a b n particularly n heaps of drying Pennisetum clandestinum EARLIEST RECORD: Whangarei, ND, 12 Apr 1929, Ε M. spinicollis. Fairburn. grass, but considerably less frequent than
REMARKS: In garden compost and fermenting fruits and veg- Monotoma spinicollis aubé, 1837 a Fig. 48, 140 etables. Common from ND to NN; also on the Kermadec Is. FIRST REPORTED a F.W. Hutton, 1904. EARLIEST RECORD: Christchurch, MC, in the 1880s, Epuraea scutellaris (Broun, 1880)a F.W. Hutton. FIRST REPORTED IN T. Broun, 1880. REMARKS: Very common in lawn clippings, garden litter, EARLIEST RECORD: Heads, ND, the Whangarei n 1870s, and compost, in paddocks where sheep shelter, and in T. Broun. chicken straw. REMARKS: In fermenting fruits and vegetables, garden litter, lawn clippings, paddocks, and parks. Monotoma testacea Motschulsky, 1845 FIRST REPORTED IN N.Z.: F.W Hutton, 1904, under the Epuraea signata Broun, 1880 synonym A. ιιb υ drfον οΙ t (Watson for) Waterhouse. REMARKS: On old logs. EARLIEST RECORD: Christchurch, MC, in the 1880s, Homepuraea amoena (Broun, 1880) F.W. Hutton. REMARKS: On sap oozing from Vitex lucens trunks. REMARKS: In hen-house straw. b t d l carnaria (Schaller, 1783) EARLIEST RECORD: Takapuna, AK, 22 Mar 1941, K. CI CI I AE Harrow. `fΙ t b tl
REMARKS: On dry carrion. Omosita colon (Linnaeus, 1758) FIRST REPORTED IN N.Z.: J.J. Walker, 1904. EARLIEST RECORD: Christchurch, MC, in the 1880s, F.W. Hutton. REMARKS: On dry sheep carrion. Omosita discoidea (Fabricius, 1775) a b EARLIEST RECORD: Tropicana Drive, Lyrifield, AK, 14 Mar 1975, sample 75/20 (week-old lawn clippings with fermenting peaches), C. Kuschel. REMARKS: In mixed compost and dry sheep carrion.
l t p d asperella Broun, 1893 Fig. 47, 139 Fig. 49 Platisus Cucujidae sp. REMARKS: On trunks and thicker branches of trees, particu- larly at areas of oozing sap. Ahasverus advena (Waltl, 1834) a b Soronia hystrix Sharp, 1876 EARLIEST RECORD: Matamata, WO, Mar 19 Ε REMARKS: On sooty mould of Leptospermum scoparium. Brookes. Ann t t d t f Sp 57
REMARKS: In garden compost and henhouse straw. Cryptophagus pilosus Gyllenhal, I827 FIRST REPORTED IN N.Z.: E.S. GOurlay, 1954. Cryptamorpha brevicornis (White, 1846) EARLIEST N.Z. RECORD: Morrinsville, WO, I0 Dec 194I, D. REMARKS: On Rhopalostylis sapida, Cordyline australis, and Spiller. G hn t fοl REMARKS: In hen-house straw. Cryptamorpha desjardinsi (G ér n 1844) a Ephistemus globulus (Paykull, 1786) a I S E O E I ΝΙ Βrο n 1 nd r th n C. FIRST REPORTED IN N.Z.: N.A. Martin, I983. suturalis White. EARLIEST N.Z. RECoRD: Okauia, WO BP, Mar 1924, A.E. Whangarei ND, Auckland AK, and EARLIEST N.Z. RECORD: Brookes. Tairua CL, in the 1870s, T. Broun. REMARKS: In garden and paddocks, lawn clippings, compost Found on all sorts of vegetation, but seems to REMARKS: heaps, and leaf litter of gardens and bush. prefer monocots, especially Gahnia, Phormium, Frey- cinetia, Cordyline, Cortaderia, Musa, and Pennisetum. Micrambina discoidea (Broun, 1893) REMARKS: On freshly cut branches of i os o um e ui- folium.
PHALACRIDAE — Micrambina helmsi Reitter, 1880 `bald beetles' REMARKS: On Phellinus fungus (assisted by R. T. Thompson) Micrambina insignis Reitter, 1880 REMARKS: Common on vegetation, especially on freshly cut branches of Sophora, Dysoxylum, e yca ya Me ico e and Acacia on Gahnia se i ο ia in hollow i e trees, and on Auricularia polytricha and Ama i a musca ia found on the latter fungus up to 40 m outside the native bush. Micrambina rutila (Broun, 1880) REMARKS: As common as M i sig is Reitter; attracted to cut branches of Rubus cissoi es Sc e e a igi a a a so sia Euca y us Cu essus Mac o i e and agu a ia on fallen o a os y is sa i a leaves; and occasionally on plants in gardens, especially a e esc ia ae io ica Fig. 50 a ac us Phalacridae u i o mis igo ico a Micrambina silvana (Broun, 1880) REMARKS: Some beaten from tree ferns; most obtained in h l r n fοr fr r l Thompson, 1980 a Malaise traps. Fig. 50, 14I Micrambina tumida (Broun, 1893) FIRST REPORTED IN N.Z.: R.T. Thompson and J.E. Marshall, REMARKS: One from beaten Cya ea me u a is fronds, the 1980. others in Malaise trap. Tropicana Drive, Lynfield, AK, 14 EARLIEST N.Z. RECORD: Micrambina species 1 Sep 1974, G. Kuschel. REMARKS: Associated with the sedge Gahnia setifolia. REMARKS: Extremely common in Acacia galls caused by the rust fungus Uromycladium. Micrambina species 2 REMARKS: On a ee fern. Salltius ruficeps (Broun, 1880) CRYPTOPHAGIDAE REMARKS: Associated with ground plants, especially P o - `cryptic beetles' mium cookia um a Ga ia se i οΙia by seaside cliffs. (assisted by C. Johnson) Genus 1 species 1 REMARKS: On monocots and dicots, attracted particularly to days-old cut branches hung in ees; common also on U o- myc a ium o a i e gall, alongside Phalacrus u i o mis igo ico a Thompson (Phalacridae).
LANGURIIDAE `slender beetles'
Fig. 51 C y o agus Cryptophagidae acutangulus
Atomaria lewisi Reitter, 1877 FIRST REPORTED IN N.Z.: C. Johnson, 1971. EARLIEST N.Z. RECoRD: Whangarei, ND, I6 Mar 1927, A.E. Brookes. REMARKS: In gardens and paddocks; particularly abundant in heaps of prunings, lawn clippings, and garden compost. 2 a a i s n r d prolixus K t b l ft r p n a i o uce s ecies b i s e o o is sρecies i ew ea a Cathartocryptus maculosus (Broun, 1881) C y ie is e y e oca i y d a o omic c a ge; see 7 - 1 REMARKS: On Dysoxylum spectabile and on dead branches. 8 Lynfield tl S rv
A GU II AE (co Hapalips prolixus (S a 1 7 Fig. 5 1 REMARKS: O Rhopalostylis sapida, Cordyline banksii, Cyathea meduliaris, Cyathea dealbata, a Dickronia squarrosa. Loberiis depressus (S a 1 7 ΕΜ ΚS I a ga e -3 m om e us ma gi Loberus nitens (S a 1 7 REMARKS: O g ou a ase o coas a c i a s (Phor- mium, Gahnia, g asses
ig 55 Anisomeristes EROTYLIDAE C r l ph d sharpi a some ee es REMARKS: On n Cortaderia jubata, a o e amo gs Carex lessoniana i e Wa e ay swam Occu s i AK S nd b t e y r r Anisomeristes ater M tth 1 REMARKS: I ea i e o Cortaderia jubata, Gahnia setifolia, a Freycinetia baueriana, a a icu a y o ea e a si e ea o s o e ee- e s Cya ea dealbata a C. medullaris. Anisomeristes sharpi Ma ews 1 Fig. 55 I REMARKS: I ea i e o a u e Ga ia ace a G. ig 53 Thallis set ο ia U ci ia a k sii U. u ci a a Ca e am e ia a C. issi a and o mium cookia um mo e commo i E o y i ae o i ta coas a c i ege a io Cryptodacne brouni ( ascoe 1 7 Anisomeristes thoracicus (E ic so 1 a b d REMARKS: I mois ecaye woo EARLIEST RECORD: E som Auck a Ci y AK Oct Cryptodacne synthetica S a 1 7 191I ou ( M REMARKS: I ecaye woo a i woo mou o a ge REMARKS: e y a u a i ga e a ock a us o ow Vitex ees i e com os ea s a mou y a eas o a s Occu s i e o th I South I C atham Is a Ke ma ec Is CERYLIDAE Anisomeristes s ecies 1 FIRST REPORTED IN Ν Ma in 19 3 ki g is e ee es EARLIEST RECORD: Owai aka AK May 1939 o ce e y i g ass ouse Whiri. REMARKS: Usua y a o gsi e A. o acicus a us as a υ a Occurs i e o I Sou I a C a am Is Coryl.ophus p 1 REMARKS: I ecaye woo a a m a ee- e i e Ηο ρ ρ 1Ι d Broun, 1 3 REMARKS: I ea i e a o Co a e ia u a a Ga ia set ι ο ia As e ia a ksii a o a os y is sa i a Holopsis rotundata ou 1 93 REMARKS: Metrosideros e ce sa ig 5 I i e u e a ge ee i A omma us e us with Ga ia ace a c us e s Cerydidae uo ecims ia s ΗοΙϋρsis p 1 REMARKS: yc om io acicu a e Ορ is Anommatus duodecimstriatus (Mue e 1 1 a ig 5 1 3 I moss ma s menus im eci us g ass ma s a moss a i e wo s o FIRST REPORTED G Kusc e 1979 . EARLIEST RECORD: o ica a i e y ie AK 5 am s o es ea s eams A 1975 G Kusc e oki sis p REMARKS: A e igme e i y ogea immig a οm REMARKS: O ogs a u ks usua y o e g ow wi Eu o e o ai e i ee i - a s sea e o a e o Me osi e os e o a a Hypodacne rubripes ( ei e 1 Ηο ρ p 3 REMARKS: I ic mois umus u e ogs a oa s i REMARKS: In l f l tt r t th ase o g ou a s (Ca e o ow ees a i ecaye woo U ci ia As e ia Ga ia o mium a o mou y u ks a ogs Holopsis p CORYLOPHIDAE REMARKS: I ea i e u e s u s a ees a e ases o oo e ee es g ou a s a o ogs a u ks d b Anisomeristes apicalis ( ea 1 95 a Orthoperus aequalis S a 1 5 a FIRST REPORTED n Ν Ζ Ν Ma i 19 3 EARLIEST RECORD: o ica a i e y ie AK EARLIEST RECORD: D'Urville Ι S I A 19 I Ε S u 197 G Kusc e Gou ay REMARKS: Ιιι com os ea s i ga e a a ock Ann t t d t f Sp
a b Orthoperus atomarius ( r 1 1 REMARKS: Sp r d ll n rd n pl nt pr nt d n EARLIEST RECORD: r p n r v nf ld AK 9 AC fr AK nd C (Gr t rr r I nl n 197 n Tricholoma pesundatum f n G K h l Diomus notescens ( l b rn 1 9 a REMARKS: E ll n n th b h rd n nd p d- FIRST REPORTED IN N.Z.: E S G rl 19 d n f r nt n pl nt ; l n d d d EARLIEST N.Z. RECORD: l r t n rth WI—W 9 Apr Orthoperus species 1 a b 1933 n ppl tr W C tt r EARLIEST N.Z. RECORD: nf ld AK A 1977 G REMARKS: On d l n r nt n AK WI K h l nd BR. EMA KS On p n xtr t d fr ft d l tt r nd Diomus p 1 a b d d d pl EARLIEST N.Z. RECoRD: r p n r v nf ld AK 1 Sacina oblonga r n 1 93 S p 197 G K h l REMARKS: On f l ; n r t n n l r n b r n r - REMARKS: On Leptospermum scoparium in paddock along- ntl t br n h p nd d fr th n p ; l ο n side bush margin. ld tr n nd t p r bl n l r nd z Halmus chalybeus ( d v l 1 35 a Sacium pulchellum ( 1 95 a b FIRST REPORTED IN N.Z.: W W r tt 1902. EARLIEST N.Z. REcoRD: t r n AK 1915 A E EARLIEST N.Z. RECORD: I p rt d n 1 99 from New South r Wales, Australia. REMARKS: One on Phormium cookianum n rd n r n- REMARKS: h t l-bl l d b rd is visible v r h r n d r n tr n f b rnt Pinus radiata tr W d pr d n n t v nd introduced plants, n native bush as well as in AK nd n th K r d I gardens and parks. Represented in NZAC from ND, AK, CL, BP, TO, and NN. G n 1 p 1 REMARKS: On tr n f d d Dysoxylum spectabile tr Harmonia conformis (Boisduval, 1835) a G n 1 p FIRST REPORTED IN N.Z.: Kirk, 1896. EARLIEST N.Z. RECORD: I p rt d in 1896. REMARKS: On tr tr n REMARKS: On Eucalyptus nd Albizia, in ND and AK G n p 1 Itleis galbula (Mulsant, I850) a REMARKS: In tr b d Gahnia l tt r nd d ld f Dysoxylum l FIRST REPORTED IN N.Z.: r I9 5 EARLIEST N.Z. RECORD: Pt Chevalier, Auckland City, AK, 15 Mar 1985, on Cucurbita pepo, B.A. Philip. COCCINELLIDAE REMARKS: On powdery mildews, especially those of Cucur- `ladybirds' bitaceae, in gardens. . a b (assisted b Κ t n nd p `Midus' pygmaeus Blackburn, 1892 EARLIEST N.Z. RECORD: r p n Drive, Lynfield, AK, I3 Oct 1974, on Acacia mearnsii, G. Kuschel. REMARKS: Some on Acacia mearnsii, most on So o a mi- crophylla in garden. Rhyzobius acceptus (Broun, 1880) REMARKS: On e yca ya a o ea and Schefflera digitata, but most Malaise trapped. Rhyzobius consors (Broun, 1880) REMARKS: Some beaten off plants, others trapped. Rhyzobius fagus ( r n 1880) a FIRST REPORTED IN r n 1880. Fig. 56 Coccinella EARLIEST N.Z. RECORD: r C ιn th 1 7 r n Coccinellidae leonina REMARKS: On Acacia mearnsii.
Adalia bipunctata (Linnaeus, 1758) a Rhyzobius forestieri (Mulsant, 1853) a FIRST REPORTED IN N.Z.: W. Cottier, 1956. FIRST REPORTED IN N.Z.: Sharp, 1889, under the synonym EARLIEST N.Z. RECORD: Christchurch, MC, 30 Mar 1936, E. Scymnus ci cu a is Sharp, 1889. Fairburn. EARLIEST N.Z. RECoRD: Picton, SD, 1884, R. Helms. REMARKS: In gardens and parks on plants, especially willows REMARKS: Incorrectly referred to under the name R. ventralis (Salix pp (Erichson) in the New Zealand literature. Adoxellus flavihirtus (Broun, 1880) Rhyzobius minutulus (Broun, 1880) REMARKS: On Astelia banksii nd Phormium cookianum f REMARKS: On bush floor sedges, cut branches, and old trunks t l cliffs. of Sophora microphylla, Nestegis lanceolata, Hedycarya arborea, Me icy us ami o us arid Me osi e os e o - Coccinella undecimpunctata nn 175 a a FIRST REPORTED IN N.Z.: W. Colenso, 1888, under the syno- nym C. novaezealandiae C l n 1 Rhyzobius rarus (Broun, 1880) EARLIEST N.Z. RECoRD: I p rt d n 1 7 REMARKS: Largely associated with Gahnia and Ca e spe- REMARKS: Sp r d ll n rd n plants, preying on aphids. cies growing on th b h fl r and coastal cliffs. Coelophora inaequalis (Fabricius, 1775) a b Rhyzobius suffusus (Broun, 1880) EARLIEST N.Z. RECORD: C rn ll AK 5 n 19 Β M REMARKS: On bush floor sedges (Ga ia Mo e o ia Ca e May. (U ci ia and on the trunk of a dying So o a mic o y a Rhyzobius tristis (Broun, 1880) K t b l ft r p n REMARKS: On bush canopy. a introduced species b first report of this species in New Zealand Rhyzobius ventralis (Erichson, 1843) a Lynfield is the type locality d taxonomic change; see pp. 76-81 FIRST REPORTED IN N.Z.: W Kirk, 1900.
60 y ie ee e Su ey
COCCI E AE (coni.) Bicava illustris ( ei e 1 REMARKS: O usco o ia yo i a a Ga o e ma a a- EARLIEST ι ο Auck a 1 9 T. ou a um a so i o o ow Sa i agi is REMARKS: O E l pt e ese e i AC om AK ica a a u i e is ( ou 191 WO BP, HB, TK, NN, MB, a MC u a e u commo REMARKS: O ecayi g u ks a s um s i woo mou o eys o E iococcus co iaceus a so o o ow Sa i agi is a a ase o o mium cooki- Rhyzobius species 1 a um and As e ia a ksiio coas a c i s REMARKS: O As e ia a ksii o mium cookia um a Bicava terricola ( ou 1 93 Gahnia lacera o coas a c i s REMARKS: On dead trunks with Au icu a ia o y ic a u - y ο iυs ρ gus a u e cu a c es o i os o um c ass οΙiυm REMARKS: O coas a c i mo oco s some o Ga ia se i o ia Acacia mea sii a So o a Rhyzobius species 3 mic o y a REMARKS: A eig s ecime s ou oge e o a Μe icορe Bicava variegata ( r n 1880) e a a s um REMARKS: ou o mos a s o e co g ega i g i um- b Ga o e ma a a a um I e e is G oeo o us Scymnus kewi (Mu sa 1 5 e s o EARLIEST ECO Auckland, 2I Feb 1941, swept om a us us a e i us u gi ca o s C S W ei Corticaria elongata (Gy e a 1 7 a REMARKS: O Aucu a Euca y us a o mium i ga - FIRST REPORTED IN M e o 1 e a i a ea o Euca y us a am usa u i gs i EARLIEST RECORD: Auck a AK i e 1 s a ock awso Stethorus bifidus Kapur, 1948 REMARKS: I us oo ea i e a o Ga o e ma a a- REMARKS: On Phormium a A ge ica i ga e e e - a um acke u gus om a My si e aus a is ee emic s a us o e s ecies mus e ques io e Corticariafenestralis ( nn 175 Stethorus griseus C a eau 1979 FIRST REPORTED IN ou 191 u e e sy o ym REMARKS: I a ga e C. o gu a r n 1910. EARLIEST RECORD: Wai ake e a ge AK 1 75 T. Stethorus histrio Ch z 1974 ou FIRST REPORTED I W i e ea 19 7 REMARKS: In garden. EARLIEST ECO We i g o W i e 19 s Ε Co ye Cortinicara hirtalis ( ou 1 REMARKS: On garden plants. FIRST REPORTED Π Ν Ζ ou 1 EARLIEST RECORD: Auck a AK i e 197 s T. ou LATHRIDIIDAE REMARKS: In ga e com os o g ass a wee s o ga e s a a ocks i s a i g es ma e ia s a occasio a y i mi ew ee es e us o Co a e ia u a a (assis e y C o so Cortinicara meridiana o so 1975 FIRST REPORTED IN C o so 1975 EARLIEST RECORD: M Moe au C 195 G e e - g as REMARKS: O o mium g asses a ee s i ga e s a a ocks b Dienerellafilum (au é I 5 EARLIEST RECORD: ami o WO 7 Aug 197 i
s o es G ey . REMARKS: i a o ow So o a mic o y a ee i e us E icmus ονeaΙυs ( ou 1 Fig. 57 Enicmus REMARKS: I us ea i e ; a so i s a y co e o ga e thr d d caviceps u e a o a ge a sia ρaρy e a Enicmus caviceps ( ou 1 93 Fig. 57 1 5 Ar d b f tυ ( tt r 1877) REMARKS: In ea i e es ecia y a ou Ca e U ci ia FIRST REPORTED IN T. 1914, r n under the synonym a Sc oe us se ges Lathridius dualis r n 1914. EARLIEST ECO M thv n MC, 1912, T. Hall. Enicmusjloridus ( r n 1 REMARKS: In lawn clippings and garden compost; occasio - REMARKS: Some i moss a i e wo s o ee u ks o a y on plants (Phormium, Cortaderia, Euca y us ogs; mos Ma aise a e
Enicmusfoveatus e o 1 . . Aridius costatus (Er h n 1842) REMARKS: O Ku ea e icoi es a e os e mum sco a - I S REPORTED IN T. r n 1880, under the synonym am i e uce s Lathridius costulatus r n 1 ou a so i um e s i a o o ow ee EARLIEST RECORD: r CL, in e 1 7 s T. r n REMARKS: In lawn and mixed compost, ou y s aw cu Enicmus sharpi e o 1 branches, and pr n n in gardens, a ocks a us REMARKS: Mos ou i ga e oge e wi E. i o ea us i e ea i e o a a sia ρaρy e a g owi g i s a e Aridius nodifer (Westwood, 1839) I S REPORTED IN A. White, 1 6, under the synonym Lithostygnus minor ou 1 93 Lathridus antipodus White, 1 6. REMARKS: In ecaye woo es ecia y of Me osi e os e - EARLIEST RECORD: North I., in the 1840s, S Parry. ce sa
REMARKS: In lawn c i i gs garden litter, arid compost, es- LiIhostygnus sinuosus ( e o 1 . pecially in heaps of pr n n REMARKS: I ea i e o us oo Ann t t d t f Sp 1
Lithostygnus p 1 REMARKS: In d d d p rt l rl f t l l ff Melanophthalma alacris ( r n 1 REMARKS: At b f Carex, Uncinia, nd t l l ff t - Melanophthalma diversicollis l n 1 REMARKS: On f ll n fr nd f Rhopalostylis sapida nd n Carmichaelia aligera. Melanophthalma puber ( r n 1 REMARKS: On v r pl nt p ll Parsonsia hetero- phylla and Pseudopanax arboreus. Melanophthalma pudibunda ( r n 1 5 Holoparamecus REMARKS: On Melicytus ramiflorus tr n nd t br n h M r ph d tenuis tr n nd n h ll Vitex lucens and Salix fragilis. Holoparamecus p 1 Melanophthalma tarsalis ( r n 1 REMARKS: At b f r nd pl nt (As e ia Ca e REMARKS: On t br n h f Beilschmiedia nd Acacia n U ci ia nd n tr b d l tt r nd d d d Rhopalostylis sapida nd Cyathea dealbata fr nd ; r th r n n Cortaderia jubata n d n t v b h Melanophthalma zetandica l n 1 MYCETOPHAGIDAE REMARKS: S n l n n pl nt b b t n ; n n b r n t `f n b tl br n h f Schefflera, seu o a a e yca ya Me o- si e os Me icy us u us Ca mic ae ia Acacia ac y- g o is i os o um Mac o i e nd G oeo o us a us us f n f e yca ya a o ea Melanophthalma p 1 , REMARKS: In p ph t n Sa i agi is t W ttl p Sp n r M zelandica l n Melanophthalma p REMARKS: M b f nd n So o a Me osi e os Sa i a i e b t n b v r n n n t h Cortaderia jubata and Freycinetia baueriana. Melanophthalma p 3 59 Triphyllus REMARKS: On v r pl nt p ll n Cya ea ea - M t ph d hispidellus a a t ei sc mie ia awa nd Freycinetia baueriana. b Sp n r M. puber ( r n Litargus balteatus C nt I 5 EARLIEST N.Z. RECORD: r p n r v nf ld AK 3 Melanophthalma p b M 1975 n h p r G K h l EARLIEST N.Z. RECORD: r p n r v nf ld AK 3 REMARKS: In r pr n n n ld h p r nd n O t 197 G K h l t rl n (Sturnus vulgaris) n t ll n p dd REMARKS: In x d rd n com os nd M l tr p t b b h r n t n t v f r t v r ll f r t nt nn l Litargus vestitus (Sh rp 1 79 l b nt r b bl n A tr l n r nt EARLIEST N.Z. RECORD: nr A l nd AK 1 l 195 r tt n d h d f Cy a a sco ymus A M Melanophthalma p 5 REMARKS: In pr n n l n l pp n nd p t n REMARKS: On e i us f n nd n f n d ld Sa i r nd; n v r h r n tr nd hr b agi is r th r n n v rt l r v rh n n tr Triphyllus adspersus b n nd r d n n p f Vitex lucens. ( r n 1 REMARKS: On ac yg o is e a a o ea ia u u acea
G n 1 p 1 a nd Ga ia n t l l ff EARLIEST N.Z. RECORD: r p n r v nf ld AK 1 l 197 G K h l Triphyllus fuliginosus ( r n 1 REMARKS: On o u gus-i es e s ems o Me icy us ami- REMARKS: S n ld l ; t n Ga o e ma a a- o us Me ico e e a a a Co y oca us ae iga us a um r n n ld l f Acacia mea sii n a i e b h A tr n b tl th h n t f r t b r l n Triphyllus hispidellus ( ou 1 ig 59 1 7 th l tr An r nt fr A tr l h r t r REMARKS: On e yca ya a o ea Me ico e e a a a fr Q n l nd t t r e os e mum sco a ium n th b h; n p n ou o e os e mum sco a ium i rd n MEROPHYSIIDAE Triphyllus punctulatus ( r n 1 ` t d ld b tl REMARKS: e y n n t d t l n th b h o y; n t f nd n f n r n n th p n (p dd rd n Holoparamecus tenuis tt r 1 79 5 1 Triphyllus serratus ( r n 1 REMARKS: On tr n l nd t p v r d th fr t n REMARKS: Knightia e ce sa yso y um s ec- b d f l r r f n ; n d p r tt d d nd d On tr n f ld n d h ll tr nd l ; nd n h nd r a i e i e uce s So o a mic o y a a Me ico e Ama i a muscaria Pinus d d l e a a occasio a y o nd r a ia a tr g owi g i e us K t b l ft r p n Typhaea stercorea ( i aeus 175 a a ntr d d p b f r t r p rt f th p n l nd FIRST REPORTED IN r n 1 nd r th n Í nf ld th t p l l t d t x n h n pp 6-8 curvipes r n nd i a r n 62 Lynfield tl S rv
ΜΥCE ΟΡΗAGΙ AΕ ( nt Cis zeelandkus tt r 1880 Fig. 61, 148 REMARKS: In Ganoderma applanatum r n n Myrsine EARLIEST N.Z. RECORD: A l nd AK nd Wh n r australis and Acacia mearnsii. d n th 1870s, T. r n Cis p 1 REMARKS: In p t h p nd b n p t r nd h REMARKS: In Phyllocladus trichomanoides nd Dacrycar- pus dacrydioides l tt r Cis p 2 ARCHEOCRYPTICIDAE REMARKS: ln r tt n d n nt f n b tl Ιι Cis p 3 REMARKS: In Gloeoporus adustus n Hedycarya arborea stem. Cis p REMARKS: t n Cis p 5 REMARKS: In fl d d br Έnn rthrοn p 1 REMARKS: On r tt n Dacrycarpus dacrydioides, Melicytus ramiflorus, nd Cupressus macrocarpa n rd n n r b h 60 Archeocrypticus b Ar h r pt d topali Orophius dilutipes l b rn 1891 EARLIEST N.Z. REcoRD: K r h p r AK 30 A
Archeocrypticus topali K z b 1964 60 1977, n r tt n d G. K h l I S REPORTED IN N.Z.: Z. K z b 1981. EMA KS On Trametes cinnabarina r n n Knightia EARLIEST N.Z. RECORD: 3 Ν f p 16 Apr excelsa. 1942, nd r Eucalyptus. Scolytocis p 1 REMARKS: In rd n p r nd p dd EMA KS On d n Knightia excelsa. Xylographus fultoni ( r n 1886) REMARKS: On d d pr t d Metrosideros excelsa tr CIIDAE h lf f n b tl ( t d b J. F. r n MELANDRYIDAE `l p n b tl
61 Cis C d zeelandicus 62 Hylobia M l ndr d nubeculosa Cis asperrimus r n 1880 REMARKS: On Oleariafurfuracea nd v r d d d Allopterus ornatus (Broun, 1880) Cis assimilis r n 1880 REMARKS: a v r l h rd-t - t h d rn l b tl r REMARKS: On Muehienbeckia australis nd Hedycarya ft n n th n ht; tl M l tr pp d S arborea. f nd n Ligustrum sinense fl r th r b t d th t Cis boettgeri ( tt r 1880) br n h f Pseudopanax arboreus. REMARKS: On t f Hedycarya, Melicope, Melicytus, Ctenopkctronfasciatum ( dt nb h r 1867) and S ph r r d n Ganoderma applanatum nd Gloeo- An EMA KS: l h rd-t - t b tl ft n n n r porus adustus. pr n n n p dd On r r d fr Ulex europaeus Cis illustris Βrο n 1880 dj nt t n t v b h REMARKS: On n b r d d n fr l tt r pl Hylobia calida r n 1880 Cis lineicollis r n 1880 REMARKS: On tr n f d d r d n tr (S ph r EMA KS On t br n h f Parsonsia heterophylla nd Knightia, Vitex, Acacia); l n l f l tt r nd n l Pinus radiata n th b h Hylobw p 1 Cis minutus f rd 1931 REMARKS: On t br n h f Pittosporum tenuifolium. REMARKS: In d d d f t tr nd n Ganoderma Lyperocharis p 1 applanatum. REMARKS: S tt r d n v r l f l tt r Cis rufulus Βrο n 1880 Mecorchesia spectabilis r n I914 EMA KS ln l f l tt r nd n olearia furfuracea, Acacia REMARKS: On r r d fr Dracophyllum sinclairei, - mearnsii, nd Astelia bankrii. nd n f nd n n Astelia banksii r tt Ann t t d t f Sp 3
Neorchesia divergens Β οu 191 COLYDIIDAE EMA KS I ea i e ; mos om i e ga e e u e a oug mou ee es amo gs Gahnia setifolia a G. lacera. (assis e y o e Ge us 1 s ecies 1 EMA KS Co sis e y ou a ase o us oo se ges (Uncinia, Carex, Schoenus).
MORDELLIDAE i ai ee es
ig 5 Pristoderus Co y ii ae antarcticus Bitoma insularis W i e 1 EMA KS O So o a microphylla, Corynocarpus laeviga- tus, a Acacia mearnsii; eque y o e acke u gus Ganoderma applanatum. Bitoma nana S a 1 7 Mordella ig 3 EMA KS Some ea e o Leptospermum sco a ium a Mo e i ae antarctica Pseudopanax arboreus, mos o ai e i Ma aise a Mordella antarctica W i e 1 ig 3 I 9 Bitoma rugosa S a 1 7 EMA KS O owe i g e os e mum sco a ium EMA KS O a c es a u ks o Sophora mic o y a i os o um c assi ο ium Sc e e a igi a a yso y um Mordella detracta ascoe 1 7 s ec a i e Acacia mea sii a i us radiata; qui e com- EMA KS O e os e mum sco a ium owe s mo i ga e s a a ks a so Mordella jucunda ( ou 1 Bitoma vicina S a 1 7 EMA KS ea e om ea s em o Olearia furfuracea a EMA KS O cu a c es o Pseudopanax arboreus a Ga ia ace a ea s a ks o Pittosporum tenuifolium. Mordellistena s ecies 1 a b Coxelus dubius S a 1 7 EA IES ECO y ie AK a 19 I caug i EMA KS O ecayi g ogs (K ig ia Acacia yso- Ma aise a G Kusc e y um o Fuscoporia dryophila u gus EMA KS wo s ecime s Ma aise a e i us si e c ose o a o c a Coxelus similis S a 1 7 EMA KS I us a ga e ea i e Stenomordellaria neglecta ( ou 1 EMA KS O owe i g e os e mum sco a ium a Enarsus bakewelli ascoe 1 Schefflera digitata; o e e ac e om U e eu o aeus EMA KS I ea i e a o o i g ogs o e i g a i e us Epistranus hirtalis ou 1 93 Zeamordella monacha ou 1 EMA KS I g ou swa s a s eam- e i e EMA KS ea e om Ulex europaeus a us ma gi Epistranus lawsoni S a 1 7 EMA KS I ecaye woo o a ious ki s; es ecia y a u a i s a i g s ems o e yca ya a o ea i es e RHIPIPHORIDAE y o y o aceous u gi a e e ee es Glenentela serrata ou 1 93 EMA KS I Vitex lucens ea i e Ithris gracilis S a 1 7 EMA KS O So o a mic o y a Notoulus crassulus ou 191 EMA KS I g ou i e a ea ees (So o a K ig ia Pristoderus asper (S a 1 7 EMA KS O a ious ees a s u s i us ( e os e - mum sco a ium So o a mic o y a Acacia mea sii U e eu o aeus Euca y us s ig Rhipistena i i o i ae lugubris Pristoderus scaber ( a icius 1775 EMA KS O yi g o ea Corynocarpus laevigatus, Soph- Allocinops brookesi Β οu 19 1 ora microphylla, a Me ico e ternata. Aa Acacia EMA KS u a mearnsii was e ac e om Protarphius decorus ou 191 og mi e y e ce am yci s Co omma a iega um a EMA KS I si e g ou i e i c u i g ecaye woo Ois a ae emo aemona a e a o i a o A oci o s O Pycnomerus depressiusculus (W i e 1 EMA KS I ecayi g woo o Dacrycarpus dacrydioides, K t b l ft r p n Cupressus a mac oca a Acacia ecu e s Me icy us i o uce s ecies b i s e o o is sρecies i ew ea a ami o us a e yca ya a o ea a so i o i g Co y- y ie is e y e oca i y a o omic c a ge; see 7 - 1 i e a ksii 64 Lynfield Beetle Survey
CO Y II AE (cont.) TENEBRIONIDAE
d r l n b tl Pycnomerus l t t n Sh rp 1886 ( t d b J. C. W tt EMA KS bl nd b tl th p t f r r d nt r - t d In d d M l t ramiflorus wood nd n tr - b d l tt r Pycnomerus n r Sh rp 1876 EMA KS In th d n d f h ll S l x fr l b C r x p Pycnomerus simplex r n 1880 ΕΜA ΚS In l f l tt r nd d d d f Dacrycarpus dacrydioides, Melicytus r fl r Hedycarya arborea, nd A mearnsii; l n d n n p n t r n n th r nd n n t v b h Recyntus tuberculatus ( r n 1880) ιt κ In d n d; l n t th lrp x br v 68 Mimopeus nd n Gahnia l tt r n br n d elongatus Rhizonium antiquum 1876 Fig. 150 Sh rp Alph t b d p r n us ( nz r 1797 REMARKS: In d n d f Hedycarya arborea and I S REPORTED IN G d n 1934. S ph r microphylla; l n d d fr nd f Cyathea EARLIEST RECORD: A l nd AK 19 O t 1909, T. medullaris. r n Syncalus hystrix Sh rp 1876 REMARKS: In h n rd REMARKS: In l f l tt r Amarosoma simulans dt nb h r 1867 Syncalus optatus Sh rp 1876 REMARKS: Und r t l Gahnia l r REMARKS: In l f l tt r Amarygmus tristis n l b rn 1893 Tarphiomimus nd nl t W ll t n 1873 Ι SΤ REPORTED IN M M 1963, A. tristis EMA KS On br n h nd tr n f n d t nd n - br t p EA IES RECORD: St M r d A l nd AK 20 v 1956, S A PROSTOMIDAE REMARKS: On ld d pl nt (Solanum mauritianum, Ευ Ι ρt Virgilia capensis), n r d l b tl Acacia mearnsii, pp h l r d r b r n th d t ; r th r n n rd n Aphtora rufipes t 1872 REMARKS: In d d d f v r tr ; l n Cupres- sus macrocarpa n p r Archaeoglenes costipennis r n 1893 REMARKS: In d d d Artystona rugiceps t 1874 REMARKS: On nd n ld tr p ll Myrsine australis nd Metrosideros excelsa, f d n n l h n t n ht Edalus opacus r n 1880 Dryocora 66 REMARKS: In b h fl r l tt r r t d howitti Gnatocerus corn t ( br 1798) a FIRST REPORTED IN tt n 1904. Dryocora howitti 1868 66, 151 W EARLIEST RECORD: Chr t h r h n th 1860s, W - REMARKS: In d d d f ld REMARKS: In h n-h nd h n rd - ZOPHERIDAE Lorelus crassicornis r n 1880 f l d r l n b tl REMARKS: On Cyathea medullaris and C. dealbata. Lorelus latulus Βrο n 1910 REMARKS: On Astelia banksii nd Cordyline australis. r l pr Sh rp 1876 REMARKS: On Cordyline australis, Macropiper excelsum, Rubus cissoides, Hedycarya arborea, S h ffl r digitata, nd Pseudopanax arboreus. Menimus batesi Sh rp 1876 REMARKS: In d n d f Vitex tucens, C r n rp l v t Melicyt r fl r nd S l x fr l h l r t nd l t n f th thr Menimus p bt n d n th rv 67 Syrphetodes ph r d marginatus Menimus caecus Sh rp 1876 REMARKS: In r tt n d f Hedycarya arborea, Vitex lu- r n ph l x r (Βrο n 1880) cen , Melicytus r fl r Acacia decurrens, and a. REMARKS: In d d l p r n d n l mearnsii; l n d ld f h ll S ph r micro- A o a e is o S ecies 5
phylla. A small, blind, depigmented species with eyes SALPINGIDAE reduced to a few rudimentary ommatidia. a k mou ee es Menimus crassus S a 1 7 REMARKS: I ecayi g woo o a ious ico ees u a - icu a y commo i e yca ya a o ea A sma ig- me e s ecies wi eyes a ia e i si e a i g 5-15 o ma omma i ia Mimopeus elongatus ( me 1 ig 15 REMARKS: I g ou ege a io a ack o eac es a o coas a s o es a c i s `Pheloneis' aucklandicus ( ou 1 REMARKS: I amo gs o mium cookia um Ga ia se i ο- ia a us oo U ci ia a Ca e s ecies Fig. 70 Sa i gus Salpingidae i u a us `Tanychilus' metallicus White, 1846 REMARKS: Caught in Malaise trap. Salpingus angusticollis ou 1 `Tanychilus' sophorae Broun, 1880 EMA KS Ο Acacia mea sii yso y um s ec a i e e yca ya a o ea Me ico e e a a Euca y us REMARKS: O K ig ia e ce sa Acacia mea sii a Sc e - s e a igi a a A i ia o a a acacia e ici a a i os o um c assi - - o ium Co y oca us ae iga us seu o a a a o eus Tribolium castaneum ( e s 1797 i os o um e ui o ium a i us a ia a FIRST REPORTED I C a icius 1775 u e e Salpingus bilunatus ascoe 1 7 ig 7 153 se io u o icia y su esse ame a a e ( a icius EMA KS Ο Acacia mea sii Euca y us s u us cis- EARLIEST N.Z. RECORD: " o a e a ia" o oa s i soi es yso y um s ec a i e ac yca us ac y ioi es 1769-70, J. Banks. Me icy us ami o us seu o a a a o eus Co a e ia REMARKS: I oo s i is oo a a mo s u a a a So a um mau i ia um . Uloma tenebrionoides (W i e 1 Salpingus hirtus ou 1 REMARKS: I ecaye woo o mos ees a s u s EMA KS Ο Me icy us ami o us Me ico e e a a i c u i g i us a Cu essus s ecies e yca ya a o ea Co y oca us ae iga us yso y um Xylochus tibialis ou 1 s ec a i e i e uce s a So a um mau i ia um REMARKS: O es y cu u us cissoi es Me osi e os Salpingus perpunctatus ou 1 e o a a a Euca y us s a us ma gi ; a so o Co o- EMA KS O Ca e esso ia a yso y um s ec a i e s e mum as a um e i y ic o Me osi e os e ce sa Co os e mum as a um e yca ya a o ea seu o- a a a o eus U ci ia a Ca e s ecies o e us oo Co a e ia u a a u us cissoi es i os o um OEDEMERIDAE e ui o ium Sc e e a igi a a Co y oca us ae iga us a ee es So o a mic o y a a So a um mau i ia um Salpingus quisquilius ou 1 EMA KS Ο So o a mic o y a i ga e a Acacia mea sii i a ock y us ma gi Salpingus reductus ( ai 19 5 EMA KS In Ga ia i e a Ma aise a Salpingus swailei ( ai 19 5 REMARKS: Ο Acacia mea sii a yso y um s ec a i e and in I e e is o e yca ya a o ea s em Salpingus species 1 EMA KS Ο a Me icy us ami o us s um Fig. 69 e y assa Oedemeridae i ea a AΝ ICI AE Parisopalpus macleayi (C am io 1 95 a a ee es FIRST REPORTED IN N.Z.: T. Broun, 1917, under the synonym Sessinia ookesi Βrο n (assis e y G We e EARLIEST N.Z. RECORD: Wai u eac ea W a ga ei Anthicus floralis (Linnaeus, 175 a 10 Jan 1914, A.E. Brookes. FIRST REPORTED IN N.Z.: ou 1 93 under the REMARKS: A ig sy o ym A a a ou Parisopalpus nigronotatus (Boheman, 1858) a EARLIEST N.Z. RECORD: owick AK e o e 1 93 FIRST REPORTED IN N.Z.: T. Broun, 1914, under the synonym Broun. Sessi ia s ic icus Βrο n REMARKS: In poultry straw of e - ouse i ga e com os EARLIEST N.Z. RECORD: Motueka, NN, in the 1900s. and i ea s o u i gs i a ock REMARKS: At light. Anthicus glaber King, 1869 a b EARLIEST N.Z. RECORD: o ica a i e y ie AK 5 K t b l ft r p n Jan 1975, on i us i a ock G Kusc e a ntr d d p b f r t e o f th ρ n l nd REMARKS: Ina a ock o i us a amo gs Co a e ia C nf ld th t p l l t d t x n h n see pp 6-8 a e ise um 66 LynfieΙ ee e Su ey
A ICI AE (co
ig 7 y o i us i idus ig 71 A icus A e i ae A ici ae es e i
A icus es e i Ki g 1 9 b ig 71 I5 SCRAPTIIDAE EA IES ECO Owai aka AK I May 19 I so ea i g ee es S i e EMA KS I a ock ea e -u coa se u i gs a some imes i s eam e i e us A icus k eus e i Ki g 1 9 a b EA IES ECO o ica a i e y ie AK 17 Aug 197 u e o mium cookia um i ga e G Kusc e EMA KS Commo i aw s; occasio a y i s eam e s i e us A icus e uci i es ou 1 EMA KS Mo e o ess co i e o mo oco s suc as eyci e ia aue ia a Ga ia se i o ia G lacera, a ig 73 o o e us Carex a U ci ia s ecies o e us oo ; a icu a y Sc a ii ae usi a us a u a o e Sou Ame ica Co a e ia u a a g ow- i g i e us Nothotelus nigellus ( ou 1 A icus s ecies 1 Fig. 155 REMARKS: On owe i g Mue e eckia aus a is a o EMA KS a icu a y commo o e g ou u e Ga ia ege a io i ge e a O e ea e om O ea ia u u acea se i o ia a Acacia mea sii a a ack o eac es woo Co es u a a ( ou 19 3 Nothotelus usitatus ( ou 1 Fig. 73 1 EMA KS I ea i e om sma us oo U ci ia a EMA KS O e o eyci e ia aue ia a a o e o mi e Carex s ecies ege a io Co es c is i ( ou 1 Fig. 15 o o e us s ecies 1 EMA KS U e o mium e a As e ia a ksii a REMARKS: O mi e ege a io ; mos i Ma aise a Ga ia se i οΙia a ew i ee i - a s Co es o sa is ou 1 93 EMA KS O As e ia a ksii a o ege a io a ig Co es o ima ou 1 93 EMA KS I o ow Me osi e os e ce sa a o a s a ig CERAMBYCIDAE o g om ee es Mac a ia e i is ascoe 1 77 Fig. 157 EMA KS O ege a io ea s eams ic a a ca s ecies Fig. 15 EMA KS O e o e os e mum sco a ium a us ma - gi ; o e o Sa i agi is a mou o Wa e ay s eam
ADERIDAE u e ee es
y o i us i i us ( ou 1 93 ig 7 159 EMA KS O mi e ege a io ig 7 1 y o i us s ecies io o us Ce am yci ae e icu a is EMA KS Ma aise a y o i us s ecies CE AM YCI AE io i ae EMA KS Ma aise a Prionoplus reticularis W i e 1 3 Fig. 7 y o i us s ecies 3 EMA KS a ae a u a i es y ea a e y e- EMA KS O e i Ma aise a o e o mi e ege a io caye woo o i us a ia a a i as e u y g ow y o i us s ecies 4 a a ou y C Wa i a o og o Acacia mea sii i EMA KS ea e om mi e ege a io Wa e ay ese e Ann t t d t f Sp 67
CERAMBYCIDAE: Aseminae tis, Casuarina, Chamaecyparis, Citrus, Coriaria, Cu es- Fraxinus, akea e yca ya Arhopalus tristis ( a icius 17 7 a sus, Freycinetia, Gahnia, Juglans, Ligustrum, Macropiper, M l FIRST REPORTED I Mi iga 197 Hoheria, Idesia, Melicytus, Muehlenbeckia, Olearia, Parsonsia, Pitto- EARLIEST N.Z. RECORD: O ua ay AK a 19 3 sporum, r n Punica, Pyrus, Quercus, Ripogonum, REMARKS: I ea Pinus radiata; a icu a y commo i a a c o ees c a e y i e Salix, Schefflera, Solanum, S ph r Tabebuia, Ulex, Ulmus, a Virgilia. CERAMBYCIDAE: Cerambycinae Phoracantha semipunctata ( a icius 1775 Ambeodontus tristis ( a icius 1775 I S REPORTED IN C Μ Wake ie 1 7 u e e REMARKS: I Chamaecyparis lawsoniana, Cupressus ame P. recurva ewma macrocarpa, C. torulosa, a Dacrycarpus dacrydioides. EARLIEST RECORD: Mi e o MC e 1 73 u y is ew ea a ce am yci is es a is e i sou e REMARKS: a ae co ica su co ica a y o agous; C i e (co ec e i ue o Varas y a eac e i a sc oo a u a in ea u ks a a c es o Eucalyptus s ui i g i e 195 a si ce ou a so i a i ia 1 Stenopotes pallidus ascoe 1 75 km o e o REMARKS: ea e a ea e om ac yca us ac y ioi- Astetholea pauper a es 1 7 es Me asequoia g y os o oi es y oc a us ic o- REMARKS: A ac e i um e s o es y cu Rubus ma oi es i us a ia a a Prumnopitys ferruginea. cissoides ( a i e a R. u icosus (adventive). e su - Tessaromma undatum ewma 1 co ica a ae a e simi a o ose o Zorion. Ι SΤ E O E I Wa ke 19 Astetholida lucida ou 1 EARLIEST RECORD: Auck a AK May 19 REMARKS: Se e a ea e om a ae O e a u caug in W l r ig in the ay ime u o e o ai e y ea i g o i REMARKS: O Euca y us s ecies Ma aise a a a cu s i e a c es -3 mm ick o Votum mundum ou 1 amage o se e in a i e us a s a oug ooke REMARKS: wo caug i Ma aise a o os s Acme a smi ii e u a e u a Ca is emo Xuthodes punctipennis ascoe 1 75 ci i us C s eciosus Came ia a o ica Euca y us s REMARKS: Mos y Ma aise a e os s e u a s a - e ma ii a icι οΙiυs Me osi - (o a ki s i c u i g i us e ce sa Me icy us ami o us a o u us ig a e os ke a ece sis a a us ace i o ia u us e sica si ium ca eya um y us commu is Sa i ca ea S Zorion minutum ( a icius 1775 o uosa i ouc i a g a u osa a Tristania co e a REMARKS: I a ge um e s o owe i g igus um si e se in the us a Wa e ay A ema e o i osi i g i cu Bethelium signiferum ( ewma 1 a c es o Euca y us s a us ma gi Mos s ecime s FIRST REPORTED IN A W i e 1 u e e ame a e o e io e variant ('batesi' Ca i ium i e sico e W i e EARLIEST RECORD: o I i e ea y 1 s CERAMBYCIDAE: amii ae REMARKS: O ea Acacia ecu e s A mea sii A. poda- Adriopea pallidata lyriifolia, A. verticillata, and Albizia lophantha. ou 191 REMARKS: O se ges o e ge e a Ca e Ga ia a i Callidiopis scutellaris ( a icius 1 1 Sci us Some a ae i cu ms o Ga ia ace a ea e FIRST REPORTED I o ag 1957 oug o a u s EARLIEST RECORD: Waka a a Wai awa ec Hybolasius crista ( a icius 1775) 1935 Ga e REMARKS: O Co osma uci a C mac oca a a C o - REMARKS: O Euca y us s u e oose a k us a Coptomma variegatum ( a icius 1775 Hybolasius modestus ou 1 REMARKS: O a in ea Acacia ecu e s A mea sii REMARKS: ai e wi cu a c es o i os o um Ie ui- A i ia o a a a So o a microphylla. o ium Eburida sublineata W i e 1 d Hybolasius pedator a es 1 7 REMARKS: A ac e o cu Me osi e os e ce sa a Euca- REMARKS: Co i e o seu o a a a icu a y cοmmο y us s Some a ae om Euca y us ea e oug o a u s o ea a o eus Gastrosarus nigricollis a es 1 7 y o asius omissus ( ou 1 REMARKS: Iso a e s ecime ea e o a ious ege a io REMARKS: O e Ma aise a e Se e a a ae see i i ge e os e mum sco a ium a c es a o e o Hybolasius simplex a es 1 7 ese ea e oug o a u REMARKS: In i os o um c assi o ium a e υ ο iυm ai e i a ge um e s wi cu a c es o e os s i e Navomorpha sulcata ( a icius 1775 us u o i ga e s a oug a u s ou a imes o REMARKS: A u s i um e s o owe i g a s a icu- wi ows a ouse wa s a y Mue e eckia aus a is os s M aus a is O ea ia u u acea a Phyllocladus trichomanoides. Hybolasius sticticus ( a es 1 7 REMARKS: I i os o um e u ο ium ea i y a ac e y Ochrocydus huttoni ascoe 1 7 cu a c es o e os REMARKS: a ae se e e y a ack Leptospermum sco a ium u ks a o a esse eg ee ose o Kunzea ericoides. Hybolasius vegetus ou 1 1 REMARKS: O i osi i g o cu a c es o My si e aus a is Oemona hirta ( a icius 1775 Fig. I I i os o um e ui o ium a i us radiata. REMARKS: th e mos e sa i e o e a i e o g o s os ge e a Acacia, Acer, Albizia, Azara, Betula, Brachyglot- Hybolasius viridescens a es 1 7 REMARKS: Easi y ai e wi cu a c es os s yso - y um s ec a i e e yca ya a o ea i iscus si e sis K t b l ft r p n o e ia o u ea a so sia e e o y a seu o a a a i o uce s ecies b i s e o o is s ecies i ew ea a a o eus So a um mau i ia um U e eu o aeus a C d y ie is e y e oca i y a o omic c a ge; see 7 - 1 i e uce s y ie ee e Su ey
CE AM YCI AE AMII AE (cont.) Tenebrosoma tenebricum (Β οu 1 93 d EMA KS I i e a amo gs o mium cookia um Co - Microlamia pygmaea a es 1 7 a e ia u a a Ga ia se i o ia a us oo U ci ia EMA KS O o mium e a cookia um a As e ia a Ca e s ecies; a ae ike y o e i mo oco a ico a ksii A a e y k ow om mo oco s o y a s a a ce i ecay Nodulosoma angustum ( ou 1 d Tenebrosoma terrestre ( ou 1 EMA KS Usua y o ai e om g ou i e a ae EMA KS In g ou i e ; o e ea e om a a om a ea e oug om ea a so sia e e o y a a Me icy us ami ο us s um o e s um o e os e mum sco a ium Tetrorea cilipes W i e 1 Polyacanthia flavipes (W i e 1 EMA KS I seu o a a a o eum esso ii a i o- EMA KS ou i Ma aise a ; o e eac ea e o cu a - s o um e ui o ium Sc e e a igi a a c es o Mue e eckia aus a is a Xylotoles costipennis ( eu i g 19 d Fig. 90-93 d si oc aeia as e iae Kusc e 19 7 Fig. 85-89 EMA KS O a i a ious mo oco a ico a s EMA KS ou o As e ia a ksii Co os e mum as a- os s Ca e esso ia a Co a e ia u a a C s e e s u Co y i e aus a is o mium cookia um a a Ga ia se i o ia e a Xylotoles griseus ( a icius 1775 Psilocnaeia brouni a es 1 7 d EMA KS We a us e o us a ga e s; a ae i EMA KS O seu o a a a o eus a esso ii woo y a e aceous a s os s Aeo ium s A au ia si oc aeia i ea is a es 1 7 d o o um Aucu a a o ica ego ia s C ysa emum EMA KS A ac e i a ge um e s o cu a s o s ems o s Co y oca us ae iga us Euca y us s a sia a- e i e a so sia e e o y a o ica e yca ya a o ea agu a ia a e so ii Ma- c o i e e ce sum Me y a si c ai ii a so sia e e o- si oc aeia a a ( a es 1 7 d y a i os o um c assi o ium e ui o ium seu- EMA KS On a a ie y o os s e a ae i woo o i y o a a a o eus u us cissoi es Sa i ca ea S issues; a so i e y y ui ( agu a ia os s A au ia agi is Sc e e a igi a a Se ecio c ue us (ci e a ia o o um Co y i e aus a is Euca y us s o e ia So a um mau i ia um a i e uce s o u ea agu a ia a e so ii Me icy us ami o us a so sia e e o y a i os o um c assi o ium a Xylotoles humeratus a es 1 7 seu o a a esso ii EMA KS O a i ece y cu a so sia e e o y a Psilocnaeia parvula (W i e 1 d y o o es laetus W i e 1 EMA KS O o mium e a a cookia um REMARKS: Usually o u ks a icke a c es o ea ees a s u s os s Acacia mea sii Cu essus ma- i osoma co e um ( ou 1 93 d c oca a yso y um s ec a i e Euca y us s EMA KS Mos s ecime s o ai e y si i g i e u e e yca ya a o ea o e ia o u ea a so sia e e o- o mium cookia um Ga ia se i o ia G ace a a y a y oc a us ic oma oi es i us a ia a i o- Co a e ia u a a y coas a s o es a c i s s o um c assi o ium e ui o ium u us cissoi es Ptinosoma ptinoides ( a es 1 7 d u icosus i e uce s a Wei ma ia si ico a EMA KS Commo i i e o mo oco s (Co a e ia Xylotoles nudus a es 1 7 Ga ia Ca e U ci ia a ae i cu ms a o i e s EMA KS Mos o Co a e ia u a a i e us Somatidia antarctica (W i e 1 Xylotoles rugicollis a es 1 7 ig 9 -9 EMA KS I i e a o u ks; a ae i ea woo os s EMA KS O a i a ious mo oco a ico a s ac yca us ac y ioi es Euca y us e ma ii e y- os s yso y um s ec a i e i os o um e ui o ium ca ya a o ea e os e mum sco a ium a so sia seu o a a a o eus Sci us u ia i is a So o a e e o y a i os o um e ui o ium u us cissoi es mic o y a Spilotrogia elongata ( ou 1 3 d EMA KS O e ea e o eyci e ia aue ia a se e a o ai e y ae oso s o -s ayi g o s eam a ks C YSOME I AE Spilotrogia fragilis ( a es 1 7 d ea ee es EMA KS ea e om ege a io a e e Spilotrogia maculata a es 1 7 d EMA KS O cu u us cissoi es a Mue e eckia aus- a is u mos ea e o e e Spilotrogia pictula ( a es 1 7 d EMA KS O e ea e Stenellipsis bimaculata (W i e 1 d EMA KS O e ea e om a so sia e e o y a o e caug o ouse wa a ew swe om ege a io mos Ma aise a e `Stenellipsis gracilis' (W i e 1 d ig 75 Longitarsus EMA KS A ew ea e om ege a io C ysome i ae jacobaeae `Stenellipsis latipennis' a es 1 7 d EMA KS Some o cu a c es o Co osma o us a a CHRYSOMELIDAE: Cryptocephalinae e os e mum sco a ium mos Ma aise a e Arnomus brouni S a 1 7 Tenebrosoma albicoma ( ou 1 93 d EMA KS Some ea e o owe i g e os e mum sco - EMA KS A u s i us oo i e ; a ae ea e oug a ium mos Ma aise a e o a u om ecaye co es o i us i as e o us oo Ochrosopsis subfasciata (Sau e s 1 5 b a om o e ase o e os e mum sco a ium EA IES ECO y ie AK e 19 a wi - Ann t t d t f Sp 69
ow C A Kusc e CHRYSOMELIDAE: Alticinae REMARKS: I um e s o Eucalyptus nicholsii. Alema paradoxa (S a 1 7 REMARKS: Ma aise a e i a ge um e s; os a u - CHRYSOMELIDAE: Eumolpinae ce ai (assis e y C aw C Longitarsus fuliginosus ( ou 1 Ευ οl ρ br nn ( a icius 179 Fig. I Ι SΤ REPORTED IN ou 1 REMARKS: Ma aise a e e sy o ymy u is e y EARLIEST RECORD: ai ua C i e 1 7 s ou S aw (1957 is es ig o e as e au o i a ecia e REMARKS: In p t r a aw s; os u k ow e is i c i e e ces i e ma e ge i a ia Pleuraltica cyanea ( ou 1 Eucolaspis colorata Β οu 1 93 REMARKS: Commo i e summe o Co osma s ecies REMARKS: O e i um e s o owe i g e os e mum a icu a y C. o us a sco a ium oge e wi E. ucu a ( ou r h t tr rυ υl ( ou 1 Eucolaspis jucunda ( ou 1 REMARKS: a e commo o ee u ks wi e s e i- REMARKS: Es ecia y commo i e summe o owe i g y es o c im i g Me osi e os e o a a e os e mum sco a ium Eucolaspis pallidipennis (Wh t 1 REMARKS: In Ma aise a s NEMONYCHIDAE Eucolaspis picticornis Β οu 1 93 i e ca ki wee i s REMARKS: Commo a ig o u e s o ey ege a io i e us ; some imes i um e s o Co osma a Halor- agis; a so o Ca ys egia se ium o sa y eac es Peniticus suffusus S a 1 7 REMARKS: ea i y o ai e y si i g i e i us oo associa io s o Ga ia Ca e a U ci ia s ecies; c im - i g o o u e g ow a ig
CHRYSOMELIDAE: Chrysomelinae (assis e y Se ma Aphilon n S a 1 7 7 i o y c us REMARKS: O moss a i e wo s emo yc i ae u u us Aphilon minutum ou 1 Rhinorhynchus rufulus ( ou 1 Fig. 7 1 3 REMARKS: O moss a i e wo s REMARKS: On y oc a us ic oma oi es um o i ys Aphilon monstrosum ou 1 e ugi ea a es ecia y ac yca us ac y ioi es REMARKS: O moss a i e wo s is a e wo ece - i g s ecies a e e y s i y u e o c - ig a e ce easi y s o e a ig ANTHRIBIDAE u gus wee i s Aphilon s ecies 1 REMARKS: On moss (assis e y Β A o oway Pyrgoides p 1 b EARLIEST RECORD: o ica a i e y ie AK 11 ec 197 o e g ou u e an Acacia mea sii g o e REMARKS: a e co mmo a imes o Acacia mea sii
Paropsis charybdis S å 1 FIRST REPORTED IN G M omso 19 as a o sis sp. EARLIEST RECORD: Coo e s K o y e o MC o 191 ummo REMARKS: A Eucalyptus ea ee e w ic is a e s a se y r pr nt d a y ie Gy a c aeus Anthr b d o a us CHRYSOMELIDAE: Galerucinae ANTHRIBIDAE: Anthribinae Adoxia puncticollis (S a 1 REMARKS: E cee i g y co mmo i So o a mic o y a i Cacephatus huttoni (S a 1 7 Fig. I s i g ime REMARKS: O ea a s os s Co osma s Co y o- ca us laevigatus, Eucalyptus s e yca ya a o ea Adoxia vulgaris ( ou 1 Mac o i e e ce sum Me icy us ami o υs a so sia REMARKS: On ac yg o is e a a a O ea ia rani, u e e o y a seu o a a a o eus i ogo um scan- some imes so commo as o e ou es i g o o e dens, nd u us cissoi es pl nt Dasyanthribus purpureus ( ou 1 Fig. 1 5 REMARKS: ea e om eyci e ia aue ia a Sa i K t b l ft r p n agi is a Vitex uce s a b i o uce s ecies i s e o o is s ecies i ew ea a Etnalis spinicollis S a 1 73 Fig. I C y ie is e y e oca i y d a o iomic c a ge; see 7 - 1 REMARKS: O o e ia o u ea 0 Lynfield Beetle Survey
A I I AE : A I I AE (cont.) Notochoragus thoracicus (Broun, 1893) EMA KS In bush litter. Euciodes suturalis 1 a Fig. 167 FIRST REPORTED IN A C b r 1959 as E i is sp. EARLIEST N.Z. RECORD: Hastings, v 19 1 G BELIDAE M r Aglycyderinae REMARKS: In p dd ; l rv n ld l f r Garyus altus (Sh rp 1 7 1 REMARKS: t n ff d d br n h f Acacia mearnsii, A. verticillata, Corynocarpus laevigatus, Cupressus macro- carpa, nd Hedycarya arborea. Lawsonia variabilis Sh rp 1 73 1 9 REMARKS: On Parsonsia heterophylla nd Mue e eckia com e a Lophus rudis (Sh rp 1 7 17 REMARKS: On tr n nd th r br n h f d d tr nd n l r n b r f r v r l nth n d d yso y um s ec a i e 1 r ft r t d th thr h t r- 7 A a ius l n l d wo as o i Phymatus hetaera (Sh rp 1 7 REMARKS: On d d tr n nd br n h f Corynocarpus Aralius wollastoni (Sh rp 1 7 d 7 175 laevigatus, Hedycarya arborea, Parsonsia heteropylla, REMARKS: S b rt l n d n r d d br n h f seu o- Pseudopanax arboreus, and Sophora microphylla. a a a o eus a esso ii -5 th h ft n Pleosporius bullatus (Sh rp 1 7 171 t d t t nt f d n (193 th t th p - REMARKS: On d d br n h f v r tr t t d th th tr f rn Cya ea ea a a n rr t ei sc mie ia awa Co y oca us ae iga us Euca y us e ma ii a so sia e e o y a i os o um e ui o- nd ium U e eu o aeus So a um mau i ia um BRENTIDAE Sharpius brouni (Sh rp 1 7 Apioninae REMARKS: d l b t n ff v r d d br n h t Acacia mea sii A e ici a a A i ia o a a Co a- e ia u a a Co y oca us ae iga us yso y um s ec a- i e Euca y us pp Fraxinus excelsa, Gahnia lacera, Hedycarya arborea, Melicytus ramiflorus, Muehlenbeckia complexa, Nestegis lanceolata, O ea ia u u acea i us a ia a Populus nigra, Pseudopanax arboreus, Schefflera digitata, Sophora microphylla, nd U e eu o aeus ANTHRIBIDAE: Choraginae Araecerus palmaris ( 1 a Fig. I72 FIRST REPORTED IN N.Z.: T.W. K r 1 95 nd r th n 79 Apion Doticus pestilens Oll ff r nt d u icis EA IES ECO W ll n t n W 1 9 n hr v-
ll d ppl W K r Apion ulicis ( r t r 177I a 79 I7 REMARKS: nd n Acacia mea sii A e ici a a nd FIRST REPORTED IN N.Z.: D. M ll r 1931
Lagunaria patersonii. rv b rv d n U omyc a ium EARLIEST N.Z. RECORD: b r t d n l n 5 b 1931 o a i e ll f Acacia p nd n v r ld dr fr t and in Al x ndr 7 b 1931 agu a ia p l f REMARKS: I p rt d f r th ntr l f U e europaeus.
Dysnocryptus dignus ( r n 1 Neocyba metrosideros ( r n 1 177 REMARKS: On nd nd r b h flοοr n t f th n r REMARKS: On Me osi e os e ce sa Ca e Co a e ia Ga ia nd U ci ia Dysnocryptus inflatus (Sh rp 1 7 REMARKS: In l tt r f n t nd d t nd n l nd r- CURCULIONIDAE r th f Ca e Ga ia e i os e ma Sc oe us nd 'weevils' Uncinia; r r d fr l f Ga ia ace a CURCULIONIDAE: r h r n Ent n Dysnocryptus rugosus (Sh rp 1 7 REMARKS: App r ntl r tr t d t n t rr n n `br d-n d v l b h flοοr Ca e a U ci ia p Co a e ia u a a Asynonychus cervinus ( h n 1 ad Fig. 178 Cy e us us u a us eyci e ia aue ia a Ga ia ace a FIRST REPORTED IN N.Z.: Ε S G rl 19 nd r th n - nd Phormium tenax. n A go ma i Cr t h Micranthribus atomus (Sh rp 1 7 I73 EARLIEST N.Z. RECORD: l n 1937 E S G rl REMARKS: C n n l tt r nd r th n t Astelia REMARKS: rth n n t n l nd O n ll banksii, Carex pp Cortaderia jubata, C. splendens, n n rd n pl nt r r l n n th r nd r p ll Cyperus ustulatus, Gahnia lacera, G se i o ia o mium Brachyolus p 1 cookia um nd e a REMARKS: In r p n r f th b h th Sc oe us o oc o agus crassus (Sh rp 1 7 17 e o d n nt n th r nd n a e os e mum sco a - REMARKS: On d d d f v r tr ium r v Ann t t d t f Sp 71
Sitona discoideus Gyllenhal, 1 3 1 Ι SΤ E O E I M E n 1975 S humeralis. EARLIEST RECORD: A t t p r l rv S p nd d lt O t 197 M.J. Esson nd M May ( AC REMARKS: In rd n ( n h n n l th l n b 1977 B.A. ll nd p n p t r A t d th Medicago p
CURCULIONIDAE: Brachycerinae: Aterpini Rhadinosomus acuminatus (Fabricius, 1775 REMARKS: A t d th Haloragis e ec a th a ae Arecophaga mi i g i e s ems Curculionidae varia Catoptes binodis (Wh t 1 CURCULIONIDAE: Brachycerinae: Gonipterini REMARKS: r nd th r n f l f dicots. Gonipterus scutellatus G ll nh l 1 33 Fig. I 5 Cecyropa discors Broun, 1 1 FIRST REPORTED IN Broun, 1 93 Gonipterus sp. REMARKS: In r t v t t n f nd b h EARLIEST RECORD: W ll n t n WN, 1 9 W.M. Mas-
Graphognathus leucoloma (Boheman, 1 179 kell. I S REPORTED IN R.A. rr n 1955 REMARKS: A t d th Eucalyptus s ecies EARLIEST RECORD: t n Η v 19 REMARKS: rth n n t n l nd Sp r d ll n CURCULIONIDAE: Brachycerinae: b thr r n dicots, rd n nd p dd d n n v r b t th Geochus inaequalis ( ou 1 • pr f r n f r Fabaceae, p ll Medicago Trifo- nd REMARKS: In us oo i e es ecia y Ga ia lacera e - hum p is om u e Me osi e os e ce sa ees u a so u e G. Irenimus compressus (Broun, 1 se i οΙia REMARKS: On v r nd r t r dicot pl nt Geochus politus (Broun, 1 1 Lyperobates asper r n 1 93 REMARKS: C n n t b h fl r l tt r o c y ogams REMARKS: S tt r d n th b h b t r n n ico s a mo oco s l ht r d nd r Phyllocladus trichomanoides and Geochus simiIis . Olearia furfuracea. ou 1 93 REMARKS: In l tt r nd r t b h Mandalotus irritus ( 1 77 Geochus REMARKS: Ιn b h fl r l tt r p 1 REMARKS: A si g e s ecime om a i e sam e (7 / 7 Mandalotus miricollis (Broun, 1917 Fig. 1 ake u e ac yca us ac y ioi es REMARKS: Sa y b h nd coas a ege a io ; occasio a y i ga e s i c us e s o Phormium e a Co si e e y Kusc e (197 as a ike y a e i e s ecies om Aus a- CI CU IO I AE Brachycerinae: Rhytirhinini ia u is ow seems im o a e Gromilus setosus (Β οu 1 93 Mandalotus p 1 REMARKS: I i e mos y om Ga ia a Ιec υm o REMARKS: In ma i ime ege a io a Wa e ay ee i g o se e a e e ou o ege a io a ig Otiorhynchus rugosostriatus (Goe e 1777 a Fig. I I Gromilus thoracicus (Broun, 1 93 FIRST REPORTED IN N.Z.: E.S. Gourlay, 1955 REMARKS: A t d th f rn p ll th s ecies o EARLIEST RECORD: p r M 19 l Blechnum, Polystichum, and Pteris; d lt n t h n th (NZAC). fr nd l rv n th l REMARKS: In rd n nd p t r rth n n t Listroderes difficilis G r ln 1 95 a Fig. 1 Otiorhynchus sulcatus (Fabricius, 1775 a Fig. I FIRST REPORTED IN N.Z.: Muggeridge, 1933 u e e FIRST REPORTED N.Z.: I I K n l 1 9 ame Listoderes cos i os is EARLIEST RECORD: Mt Ed n A l nd C t AK 1 EARLIEST RECORD: We i g o W ec 19 a ack- Broun (BMNH). i g oma oes ( AC REMARKS: I ga e s a as u es a e oge e ic REMARKS: O y a e oge e ic o u a io s i ew ea- Paelocharis corpulentus ( ou 1 a A u s a a ae ee i g o a wi e a ge o e - REMARKS: In d p r r f th b h p ll b tr aceous ico s i a ocks a ga e s nd p n l p Listroderes foveatus ( ea 19 a Paelocharis setifer (Broun, 1 93 d FIRST REPORTED IN N.Z.: G Kuschel, 197 REMARKS: In b h fl r l tt r EARLIEST RECORD: ook a s ew ymou K Phlyctinus callosus Boheman, 1 3 a Fig. 1 3 ec 195 ( AC FIRST REPORTED IN N.Z.: Broun, 1 93 nd r th n - REMARKS: A n t r t v t t n t W ttl All n Rhyncogonus germanus r jn l nd p p l t n r p rth n n t EARLIEST RECORD: Mt E nt n r Str tf rd K b - n n t b d pr d n th rth I. f r 1 93 A Ur h rt (BMNH). REMARKS: Sp r d n r nd nd l pl nt n rd n nd Listronotus bonariensis (Kusc e 1955 a Fig. 1 7 p dd FIRST REPORTED IN G A K M r h ll 1937 as y e - o es g iseus EARLIEST N.Z. RECORD: e so 9 A 19 7 E S Gou - K t b l ft r p n a ay ntr d d p b f r t r p rt f th p n l nd c REMARKS: In as u es a rb n aw s ee i g as a u s a nf ld th t p l l t t x n h n pp 6-8 l rv n v r r p ll Lolium p
2 y ie ee eSu ey
CU CU IO I AE : Y I I I I ( nt CU CU IΟΝI AE C r l n n C r l n n Aneuma fasciatum ( ou 1 d Steriphus ascitus ( ascoe 1 7 d REMARKS: O i os o um e ui o ium e a ae mi i g REMARKS: I Carex lessoniana swam wi i e s e se e ea es Typha orientalis a Scirpus fluviatilis; a u s ou mo e commo y o Scirpus. e a ae ee o oo s a a e Aneuma fulvipes ascoe 1 7 a a e o a aqua ic su s a e ( M May 197 REMARKS: Associa e wi Pittosporum tenuifolium, e a a i owe u s Steriphus diversipes lineatus ( ascoe 1 73 d ig 1 Aneuma rubricale FIRST REPORTED IN A C a k 193 as Desiantha ( ou 1 maculata. REMARKS: Associa e wi Pittosporum tenuifolium, e EARLIEST N.Z. RECORD: e so o 191 ou Co a a i o e owe s i ookes Co ( AC Gymnetron pascuorum (Gy e a 1 13 a Fig. 191 REMARKS: S ee a ock ee s o a ious e aceous di- FIRST REPORTED IN N.Z.: G Kusc e 197 co s i as u es a ga e s EARLIEST N.Z. RECORD: Auck a Ci y AK u 1971 u o ( AC REMARKS: Associa e wi Plantago a ceo a a CURCULIONIDAE: Curculioninae: Erirhinini commo i was e a s a ocks ga e s a ks a oa si es Bryocatus polytrichi Kusc e d Fig. 97-105, 189 REMARKS: Associa e wi mosses (Polytrichadelphus, Neomycta rubida Broun, 1 Campylopus, Fissidens) o us acks us ma gi s a REMARKS: Associa e wi Metrosideros excelsa. s eam a ks Peristoreus cruciger ( ou 1 1 d REMARKS: es egis Myrtonymus zelandicus Kusc e d Fig. 106-116 Associa e wi lanceolata. REMARKS: In soi om g ou su ace o a ou cm ee Peristoreus fulvus ( ou 1 e e i g o mois u e e e s a ou oo e s o my a- REMARKS: Associa e wi Muehlenbeckia australis. ceous ees suc as Metrosideros excelsa, M. perforata, Peristoreus fusconotatus ( ou 1 d Leptospermum scoparium, Kunzea e icoi es a Lopho- REMARKS: Associa e wi O ea ia u u acea a a i e myrtus bullata. ea s Peristoreus leucocomus ( ou 19 1 CURCULIONIDAE: Curculioninae: Eugnomini REMARKS: Associa e wi O ea ia u u acea a a ee - Gonoropterus spinicollis ou 19 9 d i g o ac e es i owe ea s REMARKS: Associa e with Nestegis s ecies e a ae i e Peristoreus maorinus ( ou 1913 ui REMARKS: Associa e wi aco y um si c ai ii o Hoplocneme hookeri W i e 1 coas a c i s; a a ee i g o owe s REMARKS: o e ee e s as a u s o a ious owe i g Peristoreus sexmaculatus ( ou 1 1 a s; a ae i ea a c es o Coprosma. REMARKS: Associa e wi Ca mic ae ia a ige a a a e- Hoplocneme squamosa ou 1 e o i g i owe u s REMARKS: A ew a u s Ma aise a e Peristoreus sudus ( ou 1 1 Nyxetes bidens ( a icius 1775 REMARKS: Associa e wi Ca mic ae ia a ige a a a i REMARKS: o e ee e as a a u o a ious a s; ga - o s i uci g a a i some ia es a ees Peristoreus viridipennis ( ou 1 Pactola demissa ascoe 1 7 REMARKS: Associa e wi Mue e eckia aus a is a M REMARKS: a ge um e s a ac e y cu a c es o Par- com e es sonsia heterophylla. Praolepra fultoni ou 1 Pactola variabilis ascoe 1 7 REMARKS: O Co osma s ecies REMARKS: Associa e wi Schefflera digitata a Pseudo- Praolepra infusca ou 1 panax s ecies; a ae su co ica i yi g a c es REMARKS: O Co osma s ecies Rhopalomerus fucosus ( ascoe 1 77 Praolepra squamosa ou 1 ig 19 REMARKS: A u ee i g o o e a ae i ecayi g woo REMARKS: O Co osma s ecies Scolopterus aequus ou 1 d Storeus albosignatus ack u 1 9 Fig. 193 REMARKS: o e ee e o a mos e e y owe i g a ; FIRST REPORTED I G Kusc e 197 a a i ecayi g woo a e o s EARLIEST N.Z. RECORD: i oki W a ga ei 1 Oc Scolopterus penicillatus W i e 1 d 1935 E ai u ( AC REMARKS: o e ee e o a so s o mo oco s a ico s; REMARKS: Associa e wi Acacia e ici a a a es e- a a i ecayi g woo cia y eque o A mea sii Stephanorhynchus crassus ou 1 d Fig. 190 REMARKS: Associa e wi i os o um a a i e ui CURCULIONIDAE: Curculioninae: Ceutorhynchini Stephanorhynchus curvipes W i e 1 Rhinoncus australis Oke 1931 a d Fig. 194 REMARKS: A u s ee i g o o e o ma y ico s a FIRST REPORTED IN N.Z.: G Kusc e 197 as uc oi es mo oco s; a a i ea a k o seu o a a s ecies (i e o Stephanorhynchus lawsoni S a 1 7 EARLIEST N.Z. RECORD: I uma ao Ma ge e AK 9 Ma 19 M May ( AC REMARKS: A u s ee i g o o e o a wi e a ge of a gio- REMARKS: I gardens a a ocks co g ega i g a imes a s e ms; a a su co ica i ea a c es o Coprosma. ases o Phormium tenax. a a i uci g ga s i s em o Tysius bicornis ( a icius 17 1 Polygonum persicaria i ew ea a ( M May I9 7 ; REMARKS: o e ee e o ma y owe i g a s; es e- a u s a a e y associa e wi ume i Aus a ia (E C cia y commo o es egis lanceolata. immme ma e s comm Ann t t d t f Sp 73
CURCULIONIDAE: Curculioninae: Baridini Andracalles horridus ( ou 1 I Fig. 19
Linogeraeus urbanus ( o ema 1 59 a ig 195 REMARKS: On a ious ees a s u s os s ei - FIRST REPORTED I Ν Ζ B.M. May 19 sc mie ia awa Ca mic ae ia a ige a Co y oca us EARLIEST N.Z. RECORD: y ie Auck a Ci y AK e ae iga us yso y um s ec a i e o e ia o u ea 1975 A o oway ( AC Ku ea e icoi es e os e mum sco a ium Mue e - REMARKS: Associa e wi Paspalum distichum (sy pas- eckia aus a is es egis a ceo a a i os o um e ui- palodes); a a mi i g i e s o o s A u s a ea i sec- o ium a So o a mic o y a o a o a ua y; e y ac i e i u u i g wa me ou s Andracalles vividus (Broun, 1 o ay ime is ise ua s ecies a i e i ew ea a REMARKS: O Me osi e os e ce sa and i e uce s ia Aus a ia Andracalles p 1 REMARKS: In us oo i e CURCULIONIDAE: Curculioninae: Magdalinini Andracalles p d Neolaemosaccus narinus ( ascoe 1 7 a Fig. 19 REMARKS: O Me osi e os e ce sa FIRST REPORTED IN N.Z.: R.H. M ll n 197 Clypeolus brookesi ( ou 19 3 d EARLIEST N.Z. RECORD: Auck a C t AK 7 Oc 1939 REMARKS: A si g e s ecime ea e om a ea a c o S i e ( AC Co osma macrocarpa. REMARKS: Associa e wi Acacia mearnsii and Albizia Clypeolus lachrymosus d lophantha; a a n d d br n h ( ou 1 1 REMARKS: Mos s ecime s associa e wi So a um mau i- ia um i us a Wa e ay CURCULIONIDAE: Curculioninae: Cryptorhynchini Clypeolus pascoei ( ou 1 ΆcaΙΙes certus ou 1 REMARKS: In us oo i e es ecia y u e Ga ia se ge REMARKS: In mo oco a ico i e o us oo es e- a Co a e ia g ass Some ea e om a ae o ey- cia y om a Gahnia lacera / G. se i o ia associa io u e ci e ia aue ia a and O ea ia u u acea . a ge Metrosideros excelsa ees Some ea e om e y Clypeolus signatus ecaye Pinus pinaster n a u ie i us oo ( ou 1 REMARKS: Usua y o ai e om i e o i e se ges (Ca e Ga ia U ci ia Sc oe us e i os e ma some ea e A ΙΙ n l t ( ou 1 om we ecaye cu ms o Ga ia ace a REMARKS: Some i a so sia e e o y a mos om i e Crisius fasciculatus o se ges es ecia y coas a Gahnia lacera a Blechnum ( ou 1 93 `capense'. REMARKS: O eyci e ia aue ia a ecaye woo a amo gs us oo se ges (Ca e Ga ia U ci ia A ΙΙ n ll ou 1913 Crisius variegatus u REMARKS: om a i e sam e o 1 REMARKS: O So o a mic o y a and a so sia e e o- A ΙΙ d r l ou 1 1 y a a ae i i e a s ea i y a acki g e e ose REMARKS: I i e u e A seuosmia mac o y a a oo s o a u oo e So o a mic o y a ee amo g us oo Ca e a Uncinia. Dermothrius farinosus (Broun, 1 9 d A ΙΙ h tr l ascoe 1 7 REMARKS: I ea i e REMARKS: A ac e i um e s o cu u us cissoi es a i Dermothrius ruficollis d Parsonsia heterophylla; a so o a e i e u us u i- ( ou 1 9 Fig. 199 cosus a us ma gi s REMARKS: om ac yg o is e a a A seuosmia mac o- y a My si e aus a is and ec um ca e se i e A ΙΙ l t r tr ou 1 3 Dermothrius sanguineus (Broun, 1 1 REMARKS: Amo gs As e ia banksii c us e s REMARKS: U e ec um ca e se e ea c ay a k A ΙΙ t ou 1 Didymus erroneus ( ascoe REMARKS: om ecaye Dacrycarpus dacrydioides woo 1 7 REMARKS: O a a ie y o woo y a s os s a Sa i agi is woo mou a Wa e ay Carex swam Acacia mea sii Eucalyptus s i us o e ia o u ea e - A ΙΙ t rr l ou 1 3 os e mum sco a ium es egis a ceo a a O ea ia u - REMARKS: I i e a woo mou ake om a o u acea a so sia e e ορ yΙΙa i os o um e ui o - o ow Me osi e os e ce sa u k hum, seu o a a a o eus a u us cissoi es A ΙΙ p 1 Didymus intutus ( ascoe 1 7 Fig. REMARKS: ou a ase o Blechnum 'capense'. REMARKS: O O ea ia u u acea some ea e oug Agacalles formosus ou 1 Fig. 197 om a ae i ea woo o is s u REMARKS: ea i y o ai e y sc a i g a c o i g ee Didymus s ecies 1 e u ks (Cyathea and Dicksonia). REMARKS: d Commo i ea u ks a a c es o ees a Agacalles integer ( ou 1 93 s u s os s Acacia mea sii A. e ici a a A i ia REMARKS: O i omes o Blechnum 'capense'. o a a Co y oca us ae iga us K ig ia e ce sa Agacalles p 1 Me osi e os e ce sa Mue e eckia aus a is i ο- REMARKS: om a sam e o si e ecaye woo sρο um e ui o ium o u us ig a So o a mic o y a Allanalcis allostethus ( ou 1 93 and Wei ma ia si ico a REMARKS: I si e i e uce s ea i e a ecaye woo Euthyrhinus squamiger W i e 1 Fig. 1 Allanalcis incultus ( ou 1 93 REMARKS: Only e emai s o s ecime s a e y e REMARKS: In ea i e o Vitex lucens. acke u gus Ga o e ma a a a um i a i g e u k a ick a c es o K ig ia e ce sa May e us a ew ea a a ia o e commo a wi es ea Ε me i- K t b l ft r p n a u us (Fabricius) om Aus a ia a e SW aci ic a i o uce s ecies b i s e o o is s ecies i ew ea a c Getacalles ventralis (Broun, 1 5 Fig. y ie is e y e oca i y d a o omic c a ge; see 7 - 1 REMARKS: Mos y i coas a mo oco i e 4 nf ld tl Su ey
CU CU lO I AE C Y O Y C I I (co Omoeacalles ovatellus (Broun, 1881) Fig. 206 EMA KS On trees and shrubs. Hosts: Corynocarpus laevi- Indecentia nubila Broun, 1880 Fig. 203 gatus, Dysoxylum spectabile, Hedycarya arborea, Vitex EMA KS On Schefflera digitata. lucens, and Cordyline australis. Mecistostylus douei rd r 1866 Fig. 204 Paromalia setigera Broun, 1880 Fig. 207 EMA KS Restricted to Pseudopanax species; particularly EMA KS In bush and coastal cliff litter; also in a a ge abundant in P. arboreus. hollow Metrosideros excelsa. . Mesoreda sulcifrons Broun, 1909 Paromalia vestita Broun, 1880 EMA KS On Pseudopanax lessonii. EMA KS M tl among bush floor sedges (Gahnia, Carex, Mesoreda species 1 Uncinia). EMA KS On Pseudopanax arboreus. Psepholax sulcatus White, 1846 Fig. 208 Metacalles aspersus Broun, 1893 Fig. 205 EMA KS Attacking various dead and dying trees. Hosts: EMA KS In mixed bush floor litter. Albizia lophantha, Coprosma robusta, Cordyline australis, Corynocarpus laevigatus, Elaeocarpus dentatus, Melicope Metacalles species 1 ternata, Melicytus ramiflorus, Myrsine australis, a Pitto- EMA KS In Gahnia setifolia litter. sporum tenuifolium. . Microcryptorhynchus kronei (Kirsch, 1877) Scelodolichus altulus ou 1 EMA KS In bush floor litter and on vegetation, principally EMA KS In leaf litter, especially of Gahnia se i o ia on dead trunks, branches, and twigs. Hosts: Carmichaelia aligera, Coprosma macrocarpa, C. robusta, Eucalyptus Scelodolichus hilaris Broun, 1893 sp., Leptospermum scoparium, Pinus radiata in paddock, EMA KS In b h flοοr l tt r and Solanum mauritianum. Scelodolichus lineithorax (Broun, 1881) Microcryptorhynchus perpusillus (Pascoe, 1877) d EMA KS A t d th b h flοοr d (Carex, EMA KS On dead trunks, branches, and twigs of a variety Gahnia, Lepidosperma, Schoenus, Uncinia), and of trees and shrubs, especially on the more exposed ones at Freycinetia baueriana. the bush margin, but readily extending to adjacent garden Strongylopterus hylobioides (White, 1846) Fig. 209 and orchard plants, and getting on to monocots too. Hosts: EMA KS Common o trees of the coas a e a c i s Acacia decurrens, A. verticillata, Astelia banksii, Citrus es ecia y o Metrosideros e ce sa Me icy us ami o us spp., Collospermum hastatum on Metrosideros excelsa, and Olearia furfuracea. Cortaderia splendens, Cupressus torulosa, Eucalyptus spp., and Phormium cookianum. Sympedius lepidus Βrο n 1 5 EMA KS On a ious ege a io ; some a ac e y cu Pit- Microcryptorhynchus vafer (Broun, 1881) tosporum e ui o ium EMA KS On dead branches of trees and shrubs. os s Beilschmiedia tawa, Dysoxylum spectabile, Knightia ex- Sympedius testudo ascoe 1 7 ig I and Sophora micro- l pt p r p r M tr d r x l Μ EMA KS On cut es egis a ceo a a perforata, Nestegis lanceolata, Olearia furfuracea, Pinus phylla. radiata at bush margin, and Sophora microphylla. Sympedius vexatus Pascoe, 1876 Easily baited with cut branches of Parsonsia Microcryptorhynchus species 1 EMA KS heterophylla and Rubus cissoi es EMA KS On Astelia banksii and adjacent Phormium cookianum. A leaf miner on Astelia (Β M M 19 7 Tychanus ferrugatus ascoe 1 7 Microcryptorhynchus species 2 EMA KS In bush floor i e EMA KS On various dicots and monocots. Hosts: Carex Tychanus gibbus ascoe 1 7 ig I1 lessoniana, Dysoxylum spectabile, Melicytus ramiflorus, EMA KS On i os o um e ui o ium Parsonsia heterophylla, Salix fragilis, and Sophora micro- Tychanus verrucosus ascoe 1 7 phylla. EMA KS Common to the sou o Auck a u e eme y Notacalles leviculus (Broun, 1881) a e a y ie a e ace e e y Sym e ius e a us EMA KS On old trunks of Elaeocarpus dentatus, Hoheria ascoe o e os a ( a i e Rubus s ecies populnea, Pseudopanax arboreus, and So o a micro- phylla; also cut branches of Parsonsia heterophylla. CURCULIONIDAE: Curculioninae: Molytini Notacalles species 1 Arecophaga varia Broun, 1880 ig 1 EMA KS On Carex lessoniana, Sophora microphylla, Co- EMA KS ee s ecime s Malaise trapped. Usually com- rynocarpus laevigatus, and Salix fragilis by the Carex mo w e e Rhopalostylis sa i a occu s u a e Wai aki swamp in Wattle Bay. S eam a Wa e ay e os a m as o y ece y Notacalles species 2 established from u es is e se y e a i e igeo EMA KS On tree ferns and Hoheria populnea. Hemiphaga o aesee a iae Notacalles species 3 Erymneus sharpi ascoe 1 77 EMA KS On Carmichaelia aligera and Olearia furfur- EMA KS In forest oo i e acea. Idus species 1 Omoeacalles crisioides (Broun, 1880) EMA KS A small, blind, bush-floor litter s ecies EMA KS On branches and trunks of most trees and shrubs. Paedaretus hispidus ascoe 1 7 ig 13 os s Acacia mearnsii in bush, Co y oca us laevigatus, EMA KS In decaying wood of conifers, monocots, and di- Cupressus torulosa at bush margin, Dysoxylum spectabile, cots; extracted or ea e om Acacia eccu e s a Eucalyptus sp. in bush, Freycinetia baueriana, Hedycarya mearnsii, Brachyglottis repanda, Cordyline aus a is arborea, Macropiper excelsum, Melicope ternata, Meli- e yca ya a o ea Me osi e os e ce sa O ea ia u - t r flοr Ol r f rf r r n h t r - u acea i us a ia a oa s o us oo seu o a a ph ll d p n x rb r S h ffl r d t t esso ii Sa i agi is in Carex swamp, and So o a mic- Solanum mauritianum in bush, and Sophora microphylla. o y a Ann t t d t f Sp 75
Phronira pl x (Broun, 1881) Fig. 214 Heteropsis kiwsoni W ll t n 1873 REMARKS: In l tt r nd d n d REMARKS: On caudex f d d Cya ea r d n tr Phrynixus astutus 1876 f rn REMARKS: In b h fl r l tt r t r p p 1 Phrynixus modicus r n 1880 REMARKS: On caudex f d d C th nd Dicksonia tr REMARKS: M tl fr ft d d n d f seu o- f rn
a a a o eus nd ac yca us ac y ioi es n r r d Macrorhyncolus littoralis (Broun, 1880) 2I7 fr th l tt r FIRST REPORTED IN N.Z.: T. Broun, 1880, n nd n Pogonorhinus opacus (Broun, 1880) p REMARKS: Nearly ll ht n M l tr p t r pp r EARLIEST RECORD: Tairua, CL, n th l t 1870s, T. W ttl tr nd r d n n p f i e uce s nd Broun (BMNH). yso y um s ec a i e th t l x r nt th f rn A - REMARKS: In dr ft d t d th f rn Macroscytalus parvicornis (Sh rp 1878) d Sosge es longicollis r r 1914 EMA KS t n fr tr f rn nd n t REMARKS: tt r fr b f b h fl r U ci ia Ca e nd M r t l r t (Sh rp 1878) Fig. 1 Sc oe us p A bl nd r nd-l tt r p REMARKS: t n fr v r tr ; r r d fr Co osma mac oca a CURCULIONIDAE: Rhynchophorinae Microtribus brouni (W ll t n 1874) Dryophthorus b p 1 Fig. 215 REMARKS: On th p ph t Co os e mum as a um EARLIEST RECORD: Lynfield, AK 22 b 1975, G. Kuschel. Mic o i us u o i W ll t n 1873 REMARKS: In p rtl d d i us a ia a b rd d p t d REMARKS: On As e ia a ksii Co os e mum as a um a b v tr b n b fl d t r n b h; nd n Ca e gemi a a es ecia y commo o o mium cooki- Cu essus mac oca a t p v r r n b e ise um a um a e a c a es i um l n d a e ce Microtribus p 1 REMARKS: Sitophilus oryzae ( r I7 3 Fig. 216 th n h b n o e ega i g is wee i FIRST REPORTED iN N.Z.: J.C. Fabricius, 1781, nd r th Novitas dispar Broun, 1 93 n Curculio i u e cu a us Fabricius. REMARKS: Associa e wi o mium e a a cooki- EARLiEST N.Z. RECORD: C ll t d d r n C pt n C a um v b t n O t 1769 nd b 1777. Pachyops dubius (W ll t n 1873) REMARKS: In h n f d REMARKS: In dr d d d f v r tr nd l n ; p ll n n a so sia e e o y a CURCULIONIDAE: Cossoninae Pentarthrum zealandicum W ll t n 1 73 Agastegnus aeneopiceus (Broun, 1880) d REMARKS: In Cο y οca ρυs ae iga us a i us e ce sa REMARKS: r d fr d d Co osma mac oca a ea us ma gi Mac o i e e ce sum and Me icy us Agastegnus d simulans (Sh rp 1878) ami o us . REMARKS: On Co y oca us ae iga us a so sia e e ο oeo agosoma co i um Wo as o 1 73 y a nd Sc e e a igi a a REMARKS: In d d yso y um s ec a i e Ga ia ace a Allaorus rugosus (Broun, 1893) Mac o i e e ce sum Me icy us ami Ιο us Sa i agi is REMARKS: A ase o U ci ia nd Ca e s ecies i us a and i e uce s i mi e i e o i e yso y um Co osma My si e Phloeophagosoma dilutum W ll t n 1874 e yca ya Cya ea nd ickso ia REMARKS: Acacia ecu e s A i ia o a a d In d d Camptoscapus planiusculus (Broun, 1880) ei sc mie ia awa Co osma mac oca a Co y i e REMARKS: t n n l n d ff r nt pl nt a ksii Cu essus o u osa ac yca us ac y ioi es Eiratus ornatus r n 1886 Mac o i e e ce sum Me icy us ami o us i us a i- REMARKS: In d d Co y oca us ae iga us Ku ea en- a a seu o a a a o eus So o a mic o y a nd coi es nd i us a ia a i e uce s Ei a us parvulus 1876 oeo agosoma e a um Wo as o 1 7 REMARKS: A t d th seu o a a a o eus nd REMARKS: Associa e wi o mium e a esso ii a a su co ica i ea a c es Sericotrogus su ae esce s Wo as o 1 73 Eucosso us se ige (S a 1 7 REMARKS: U iqui ous o e s co i e s mo oco s a i- REMARKS: A t d th Co y i e aus a is nd C co s; eco e a so om Ke ma ec o o k a o a ksii owe Is a s Euo yum u um (Broun, 1880) Stenotrupis debilis (Sh rp 1 7 REMARKS: In d d Mac o i e e ce sum Me osi e os e - REMARKS: On ea ea es o o a os y is sa i a ce sa i us a ia a and Sa i agi is In d p fl r nd Stenotrupis wollastonianus (S a 1 7 th rb rd n En l nd w e e es a is e si ce 193 REMARKS: On d d l v f o a os y is sa i a a r p rt d b n 1948. Tanysoma comatum (Broun, 1886) d Exomesites optimus r n 1886 REMARKS: A t d th Co y i e aus a is REMARKS: In tr b d and t ase o us oo Ca e spp. Heteropsis latirostris Ma s a 19 d o os oma a ica e (Broun, 1880) REMARKS: Acacia ecu e s REMARKS: r pl f si e woo In ea i us a Cu essus o u osa a us ma gi K t b l ft r p n ou a o gi os is (Wo as o 1 73 d a i o uce s ecies b i s e o o is s ecies i ew ea a C REMARKS In ea o s o Cyai ea me u a is a C ea - y ie is e y e oca i y d a o omic c a ge; see 7 - 1 a a 6 nf ld tl S rv
Toura sharpiana (W ll t n 1 7 d Dendrotrupes p 1 REMARKS: In d d fr nd f Cyathea medullaris nd C REMARKS: A t d th r ntl d d nd d n Pseudo- dealbata. panax arboreus; ll n br n h l th n 15 th Touropsis brevirostris (Sh rp 1 7 h thr Dendrotrupes p ft n x t n th EMA KS In d d br n hl t f Pittosporum tenuifolium. br n h f tr D. costiceps n th th r p rt D. p 1 n D. vestitus Touropsis p 1 th th n br n h nd n th r n b t n REMARKS: t n x d v t t n Hylurgus ligniperda ( br 179 a Fig. 221 FIRST REPORTED I n 1977 Unas conirostris M r h ll 1953 EARLIEST N.Z. RECORD: Wh tf rd AK 197 ( REMARKS: In Phellinus-i uce n r t r n tr n f REMARKS: In Pinus radiata t p l r Vitex lucens tr Hypocryphalus p 1 CU CU IO I AE S l t n EMA KS A t d th Parsonsia heteropylla. Hypocryphalus Amasa truncata (Er h n 1 a Fig. 219 p Sophora microphylla. FIRST REPORTED IN M ll n 19 9 EMA KS A t d th EARLIEST N.Z. RECORD: C nt rb r 193 Phloeosinus cupressi p n 19 3 a Fig. 222 REMARKS: Ad lt xtr t d fr Kunzea ericoides and FIRST REPORTED IN nd 1953 Metrosideros excelsa. EA IES N.Z. RECORD: A l nd AK 19 3 ( Chaetoptelius mundulus ( r n 1 1 REMARKS: On Cupressus torulosa. EMA KS A t d th d n r r ntl d d Pitto- Xyleborinus eucalypticus (S h dl 193 a b Fig. 223 sporum crassifolium and P. tenuifolium. EARLIEST N.Z. RECORD: nf ld AK 3 M r 1975 G K h l (NZAC). Cryphalus wapleri (E hh ff 1 71 a Fig. 220 REMARKS: Acacia decurrens, Cory- FIRST REPORTED IN M ll n 197 Ad lt nd l rv n nocarpus laevigatus, and Melicytus ramiflorus. EA IES N.Z. RECORD: K r r 3 S p 19 Or h In tr (NZAC). Xyleborinus saxeseni ( tz b r 1 37 a REMARKS: A t d th Ficus p b t th nf ld I S REPORTED IN nd 19 p n r ll t d n n M l tr p th th r EARLIEST N.Z. RECORD: M r n n 19 3 dr n d n p l ( Dendrotrupes costiceps r n 1 1 REMARKS: tt d n M l tr p REMARKS: A t d th d n nd fr hl d d Pseudo- panax arboreus, n th r br n h th n D. vestitus. CU CU IO I AE l t p d n Dendrotrupes vestitus r n 1 1 d Platypus apicalis Wh t 1 EMA KS A t d th d n nd fr hl d d Pseudo- REMARKS: In Corynocarpus laevigatus nd Dysoxylum panax arboreus and P. lessonii. spectabile t rl d n t r d tr l v l
TAXONOMIC OTES CARABIDAE Ctenognathus novaezelandiae Fairmaire, 1843 = r - Microscydmus lynfieldi Franz, 1977 = M. h t n h l v ll Motschulsky, 1864 n.syn. Franz, 1977 n.syn. M t h l t p p n x n d h p r p- Sciacharis fulva Broun, 1893c = S r th (Franz, r nt d b t n n th nf ld b h nd n n n 1975) n.syn., S p d fr l (Franz, 1975) n.syn. th ll t n b th n r n f C. novaezelandiae. Sciacharis lanosa (Broun,1886) = S nr (Broun, 1893b) SCYDMAENIDAE n.syn., S n l n (Franz, 1975) n.syn., S p d l n (Franz, 1975) n.syn. Adrastia Broun, 1881, reinstated genus =Austroscydmaenus (Franz, Franz, 1975 n.syn., r Lhoste, 1938 n.syn. Sciacharis puncticollis (Broun, 1880) = S n l 1977) n.syn. Adrastia clarkei (Franz, 1975) (S d n n.comb., Sciacharis taranakii Franz, 1980 = S p Franz, 1980 n.emend. (named after C.E. Clarke, but spelled l r n.syn. Euconnus calvus (Broun, 1880) = E l t l ( r n 1 93b n n E pl n p (Broun, 1893b) n.syn. Stenichnaphes urbanus Franz, 1980 = S p l r n Franz, 1985 n.syn. Euconnus setosus (Sharp, 1874) = E r t (Broun, 1893b) n.syn, E t p n Franz, 1975 n.syn. PSELAPHIDAE Magellanoconnus Franz, 1967 new status (from subgenus Anabaxis electrica (King, 1863) = A. n r Broun, 1921 of E nn Thomson) n.syn. Magellanoconnus galerus (Broun, 1885) n.comb. (E n- Anabaxis foveolata (Broun, 1880) n.comb. (E pl t a. n pl t d (Broun, 1 93b n n
Maorinus Franz, 1980 new status (from subgenus of CORYLOPHIDAE E nn Thomson) An r t p l (Lea, 1895) n.comb. (S r d r Maorinus angulatus (Broun, 1893b) n.comb. (E nn = An r t th r (Erichson, 1842) n.comb. (Seri- E nn tr p n Franz, 1975 n.syn. d r x n t
CERAMBYCIDAE 5- 3 l n th p d t l r tt h nt r h rt n n rth rn p p l t n l n r n th th rn nd Ch th I l nd Eburida White, 1 r n t t d (n t syn. with d n- p p l t n tha Newman, 1 EMA E r nd spermatheca in 9; p l t n b l modestus Broun, 1 = Η n l Broun, h b r t nd d r ll p l n n p nt t n 19 3 n.syn., d t r r n 19 n.syn., thora- cicus Broun, 1 93b n.syn. LENGTH 3 -5 3 ; dth 1 -1 5
b l vegetus Broun, 1 1 = br v ll Broun, MATERIAL EXAMINED. l nd ( 79 p n in ll 1 3 n.syn., fasciatus Broun, 1 1 n.syn., n r AK Lynfield, Mt Roskill, A l nd C t (5 G Kuschel); Broun, 1 93b n.syn., r l p Broun, 19 3 n.syn., Η Gr n A l nd C t (7 G Kuschel); Huia (1 B.M. r f n Broun, 1893b n.syn., n t f t Breun- M ; Waitakere n (13 R.A. rr n B.A. ll G ing, 19 n.syn. Kuschel, Spiller); Matuku rv Bethells (3 C Cr WO r t (1 J.C. W tt ; Okauia, Matamata (1 Ε b l viridescens Bates, 1 7 = l t ll Broun, r ; M h n (1 G W C M r I l nd (1 19 3 n.syn. J.C. W tt l t r (1 J.I. n nd nd Zondag). G Te Ar r (1 Μ ; Waikaremoana (5 Nodulosoma Breuning, 195 n t t (fr b n of G Kuschel). Wh t n h Tangoio (1 J.C. W tt ; be Ο Somatidia Thomson) = Laevisoma Breuning, 195 n.syn., W p n (1 C C r Oh n (1 r- r WI W n n (1 S Dugdale); (3 C C b r ; Sp n Βr n n 195 n n xt n(1 WN: Ot h(1 R.M. ll ; W n (1 ; d d l p r t d fr S t d b th b n f p r n ll ο ( Sp ll r ;I l nd (5 E S G rl M St ph n th t b nd b th nt r rl n n- r n t pronotum. I l nd (1 E.S. Gourlay, G Kuschel, McBurney, G W ; Urv ll I l nd (1 1 E Fairburn, E.S. Gourlay, G W (Broun, n.comb. d l n t 1 (Somatidia) ; Ch t d l l nd(1 E.S. Gourlay); r n h ( G = Somatidia ο t f r S discoidea1 93bBroun,n.syn., Kuschel, McBurney, M r G W S W ld ; Sh p Broun, 1913 n.syn., S n d l r Broun, 1913 n.syn., S C v (5I G K h l McBurney). l n (5 E.S. Gour- b t r n r n 19 9 n.syn. lay). n (1 E.S. Gourlay). W O r t ( G Kuschel). DN: n d n (7 CHATHAM IS A S W t n Psilocnaeia Bates, 1 7 r n t t d (fr n n with (3 E.S. Gourlay, G Kuschel, G W A.K. W l r ; St n ll p Bates, 1 7 p p b monotypy Ρ (5 G Kuschel); tt I - r t ( G Kuschel), Gl r linearis t (3 G Kuschel); S th E t I l nd (1 J.I. n nd h n h r th St n ll p loop-like r b t ll TYPE DATA. ΗοΙοt p Ι 3 × 1 15 W ttl Lynfield, b l sclerite nt n d n th n t b l r t n f th nt rn l A l nd C t AK 1 M r h 1979 n Astelia banksii, G nd p l t n h b r t th nd f th b r b t Kuschel (NZAC); paratypes in th n f d ff r fr t n h v n th l b rr t d b br d n t l nd d r th n th d t r f ommatidia t th n rr t p nt a l n t d prothorax, nd l tr d r ll fl tl t r lin v OS A S b f nd n p o Astelia nd Collosper- nd h l d th p r S p h r tr n f rr d t l n mum (Liliaceae), Phormium a Cordyline (Agavaceae). r pr v l n l t l l l b f fl htl n REMARKS. Psilocnaeia asteliae h rd t d t n h fr Ρ aegrota ( t n xt rn l h r t r l n h r t r t th t n a egrota (Bates) n.comb. ( l t l Fig. 1- b f th nt rn l f th l nd th f t r f th t n Psilocnaeia t l n.sp. Fig. 85-89 h b r n th b r f th f l p r t th t p r d l h t p nd parvula (Wh t h r th Int nt r dd h br n nt nn nd l p l r b n h t p rv h r fr th hr K n t St rt d r t l d n l v n t f nt nt xp d r h I l nd aegrota n rthl nd ( n l v f th r Kn ht elytra, th bd d ft n l nt d r h d t n th p l n nd Ch n nd Mokoh nau l nd n rth rn A l nd pronotum, p b n nl l htl d n r t rd d f (Mt A l nd r d C r nd l asteliae t b b nt th r f r n l n r p tt d alutaceous, v r f n l fr th C r nd l th th x pt n f M r I l nd h r nd h ll l p n t t l br r n rl b h nd l r p rt ll thr p r f nd d -b - d h p d t l r p - f pp r l b E br d n t n f n l r f r th r nt d tt h nt p nt f th b l r t v r n l n th l ommatidia. Prothorax l htl l n r th n br d aluta- h rt n 5 f r th p p l t n fr th n rth rn h lf f th ceous, v r f n l nd h ll l punctate; p l r d r ll rth I l nd l n r f r th p p l t n fr th th rn h lf n pl t b l r f n pl t l t ; subapical nd f th rth I l nd S th I l nd nd th Ch th I l nd bb l n tr t n th 1 setiferous p t n r p l Elytra d t t ddl v r n l nd t rd b l n l l n brouni Bates (from subsequent placement in l htl t p r n t rd b nd p x p br dl r nd d Stenellipsis). h r l r n rl ff d; 1 rr f p r xt nd n - b nd ddl d b h nd h r l r th p r l n bullata (Bates, 1876) n.comb. (St n ll p = W n tr n l r d d b t n -th rd elytral l n th l n r St n ll p v r d p r n n 1940 n.syn. br n nd f r n t xt nd n b nd v ntr t
MALE. Aedeagus h t v r bl n d r l v n Psilocnaeia linearis Bates, 1874 = Sl n ll p p r ll l 5 r l ttl l nd r r l sclerite n 7 th l p r n n 19 n.syn. • 8 nf ld tl S rv
Psilocnaeia n n ( a es com (Xylotoles) n r n nt t r S. cuneata Sh rp S. gracilis (Wh t S. latipennis t nd S. sculpturata ( r n h Psilocnaeia parvula (W i e 1 com (Xylotoles) = ll S. longula r n n nd th C l d n n p Xylotoles apicicauda eu i g 19 3 sy n d b r n n t Stenellipsis, r l ft n rt d n th n f r th t b n Ptinosoma eu i g 195 ew s a us ( om su ge us o Somatidia omso = Odontosoma eu i g 195 sy 'Stenellipsis' gracilis (W i e 1 = S. mediofasciata S p r t d fr ndb SomatidiaTenebrosoma th b n f eu i g 19 sy p r n th f r t b nd from Nodulosoma y th pr n f 1 r d t n t p r n th h nd t b Tenebrosoma eu i g 195 ew s a us ( om su ge us o Somatidia omso Ptinosoma convexum ( ou 1 93 n.comb. (Soma i ia = Soma i ia a u a Broun, 1 93 n.syn., S e u a a Broun, Ch r t r d b h v n f r t b th p r n n-f l t 19 1 n.syn. l tr nd r n l tr th t nd l t th b l r n f th l tr Ptinosoma ptinoides (Bates, 1 7 n.comb. (Soma i ia = (Broun, S u ico is191 Broun,n.syn.,S se ico o a Tenebrosoma albicoma 1 93 n.comb. (Soma- 19 9 n.syn. i ia Tenebrosoma tenebricum (Broun, 1 93 n.comb. (Soma- Spilotrogia a es 1 7 ei s a e ( om sy o ymy wi i ia Stenellipsis a es 1 7 = Eurychaena a es 1 7 ( om sy o ymy wi Stenellipsis), Trichocnaeia eu i g 1959 Tenebrosoma terrestre (Broun, 1 n.comb. (Soma i ia sy y e s ecies Spilotrogia maculata a es = Soma i ia usca a Broun, 1 93 n.syn. Spilotrogia d ff r fr Stenellipsis nd Psilocnaeia b h v n l n fl ll f r b l l r t r nf r d th n n-d fl t- Xylotoles costipennis ( eu i g 19 com (Stenellip- n b r nd 1 r d n l r t n th nt rn l b sis) ig 9 -93 l n p l t n h b r t th nd f th b r nd b E ESC I IΟΝ S l r t nd nf d th X. rugicollis h v n th p r th l d t nt r n th b r t th bl nd nd t ( 9 -9 b t d ff r n n h v n ll hr p t th n b l n th f ll n p (l t n r p t n n th fr n n t th nd n th r p t n th n n p r nth n t th l r -l b ; th fr n n t b d nd r th hr p t; th l tr ll th l ht n tr t n Spilotrogia elongata (Broun, 1883) n.comb. (Co es e a = b h nd th b th r p nt d nd xt nd d t p nd th y o o es ygmaeus Broun, 19 3 n.syn. nt r tr 3 5 7 nd 9 l br nd r d t l b t d t n t t ; th pp r p rt f th p t rn th f nt hr - Spitotrogia fragilis (Bates, 1 7 n.comb. (S e e i sis p t t t; nd th h nd t b l d n l p b nt
Spilotrogia hilarula Broun ( ic oc aeia MALE. n th nt p r r l l b b r n l n tr n t t p x nd n th nt r nd r d f l b Spilotrogia maculata a es 1 7 = Stenellipsis maculipen- A d d r ll nd l t r ll n 9 nd 91 b n - nis eu i g 19 l rl xp nd d n h d t p x th 3 h rp r n n nd r d
Spilotrogia pictula (Bates) n.comb. ( y o o es FEMALE. Sp r th n 9 b t v r bl ; n b d f ll tr t d p r th 75 × nd v r l n l ndr - Spilotrogia pulchella (Bates, 1 7 n.comb. ( y o o es = l l nd 11 × y o o es ascia us Sharp, 1 n.syn. Χ ge ma us Sharp, 1 n.syn., Χ g aci is Broun, I91 n.syn., Χ e simi is LENGTH 10.0-14.0 dth - Breuning, 19 n.syn. M t r l x n d rt -n n x pl fr AK C nd Stenellipsis a es 1 7 y e s ecies S. bimaculata (Wh t OS A S In d d p rt f n t nd d t ; th r r d h n d ff r fr Xylotoles n nd Spilotrogia r fr ld l of Gahnia (C p r nd d d br n h t b th l f fl ll n th nt rn l nd th of Nothofagus menziesii ( pr n f p l t n h b r t th nd f th b r nd fr Psilocnaeia t b h v n th r h ll l REMARKS. Xylotoles costipennis and X. rugicollis r n t nl r n t th l b nn t d b t l t r f p tr v r t f th r r n b t r ft n n th pl nt t d t th n rr t p nt th pr th r x tr n v r nd t th t p rt l rl n th adventive Cortaderia jubata th l tr d r ll nv x n th l t r l v b nd th b l r n n th r p n p rt f th f r t nd th n t v C. th rd th t p r h h r t r j t l t d r nd r th n splendens. A th rd p f th n X. nudus t n b r p f th l nd f n h v f fr nt p n n f th t p n th Cortaderia simi a t th t f S. bimaculata b t h v n t d f l p-l r X. rugicollis is n v r ll r 7 5-1 5 ; t b l l r t l n f n t b xt nd d b nd th n p rt f r pr nt d fr AK C nd WO th nt rn l r nf r d b p r f v r f n fl x bl p r ll l b r d fl t d b f r th nd t f r n rr -l p x l r Xylotoles laetus W i e 1 = Somatidia (Laevisoma) t th t f Xylotoles ( 1 h p n rn d th t metallica eu i g 19 sy x n t
BELIDAE MATERIAL EXAMINED. ew Zealand. AK y ie M oski 1 ma es 33 ema es co ec e e wee 1 u 197 a 7 Ma Aralius new name for Plalycephala Moiurouzier, 1 1 (not 1979 e ac e om moss o us acks us ma gi s a gender masculine aΙΙé 1 ; s eam a ks G Kusc e ; i a 1 ema e om moss o s eam e ame is e i e om A a iaceae e os - a ami y o aiik G Kusc e C o gio a e A e me 1 ema e 11 e ew Ca e o ia a ew ea a s ecies o 197 e i e sam e 7 /19 Me o
Aralius olivieri (Mo I ou ie 1 1 cοm (Platycephala) TYPE DATA. o o y e ma e 1 35 × 719 y ie AK mm Ma 1977 i moss om s eam a k G Kusc e ( AC ; Aralius wollastoni (Sharp, 1 7 n.comb. (Platycephala) = a a y es i AC Aglycyderes badius Broun, 1 n.syn., Platycephala jean- neli Paulian, 19 n.syn. ECO OGICA NOTE. o e ou i e oose co o ies o wo mosses o e ami y o y ic aceae oiy ic a e us mage - a icus and ogo a um su u a um g owi g si e- y-si e o a e a e c ay a ks a o si o us ack si es mo e o ess CURCULIONIDAE e ose o ig Agacalles integer (Broun, 1 93c n.comb. (Acalles) REMARKS. A o u ose s ecies o e ge e a a ea a ce o Β Agastegnus Broun, 1 3 = Glyphorhamphus Broun, 1 o ico is ou a ugosus ou om Mi Ca e u y Sou Is a o y ic i is c ose y e a e o o ico is n.syn., Selocomis Broun, 19 9 n.syn. om w ic i i e s y a a ica y oa y ou e ae eagus Agastegnus aeneopiceus (Broun) n.comb. (Selocomis) e sus a e i g a a e o a oi a s o e a e- ike sc e i e a a ai o sma squa e sc e i es a a a e a u y a owe
Agastegnus simulans (Sharp, Ι 7 =a biimpressus Broun, i e a sac 19 9 n.syn., A gratus (Broun, 1 n.syn., A. longipes (Broun, C. conicollis (Broun, 1 n.syn., A nitidirostris Broun, 1 93 n.syn., A Camptoscapus planiuscutus 1 = ruficollis Broun, 1 3 n.syn. (Β οu 1 1 n.syn., C. sanguineus Broun, 1 93c n.syn. Clypeolus brookesi (Broun) n.comb. (Acalles) Aneuma fasciatum (Broun) n.comb. (Peristoreus)
Clypeolus lachrymosus (Broun) n.comb. (Tychanus Asynonychus cervinus (Boheman) As A A a e i s (19 g ou s o ei s a i g A go i a i Dendrotrupes Broun C o c 1 7 o a e oge e ic o u a io s a e a e u co - e e a e ee a e commo s ecies o is ge us i ew ea- i ci g e s a ce is o a o e a a ee associa e wi seu o a a s ecies (A a i- aceae ey o e occu simu a eous y o e same a Bryocatus polytrichi n.sp. Fig. 97 - 1 5 s ecime u i i e e a eas o e ee Β ou as ame wo I egume o o y os um a e ae a i a si a k ow o s ecies vesti us and cos ice s e use o w i e i e i i- co ae emo a i iae a a ica i o e y a e is ow ca io a e s o o e s ecime o y a e o e s o e se ies es i u e co sis i g o a e se coa i g o agg u i a e g eyis go us e s ecies um e O igi a s ecime s o s ecies sca es u sca es ee o mi e o mesos emum a a o esc i e i 1 3 o e o e y ou a e s ecies um e o co ae; i o os um u o a e a i se io s e a su ace o g ee a e s ose o a e yea s o w i e a e s I u y 197 I os um a ea a e ae a si a mos o as e i e sca e- e ami e e e o u es s ecime s i e a ious ou ee o a s e se y im esse Eyes o o u i g os- co ec io s k ow o e is O y ou o e ma y s ecime s um cu e su cy i ica wi ou a e a g oo e o o a see a g ee s ecies um e a e s a us qua i ie as wi oa ocu a o es u e e wi a ge wi e y se a a e sy y es ma es eac ea i g e um e 131 o vestitus, o u es o mi e o isc a sma e su co iguous o u es a ema es e um e 1315 o cos ice s e e was o y c ose o ase ese o u es a o e s ig e e a io s o e o e a -w i e i e i ica io a e e s ecies e s ecime wi o e o mo e a ge s o g y cu e squami o m se ae wi e i e i ica io a e was se ec e as ec o y e a us Scu e um isi e u c i o m E y a s ia e is i c y o u ose e wo ames e e o wo is i c s ecies co a y o s a e- o u e e i e s iae i iae o e icu a e wi ou emuc o me s y Sc e (19 a Woo (19 Sc e a i g a sa segme 3 is i c y wi e a segme sy o ymise cos ice s u e νes i us Woo e o e way ou es i us u e cos ice s MALE. e i e 5 a S e i ei 99;9as ig s e i e i ig 1 1 1 Ae eagusas oa y i ou e 1 ig a a e ; Dendrotrupes costiceps Broun reinstated (from synonomy i e a sac a u y co s ic e us ce a a om a sc e i e with vestitus Broun) co sis i g o a ow y co ec e a es ese si ua e a ou ec o y e ema e × 95 mm o 1315 o g ee a e e e wi i u ca io o a o emes; o o e a ma u e i e - ai ua wi i e i ica io a e ( M a a ec o y e e- ose a o i e a sac ma e 1 × 5 o e a a as a o e u o i e i ica io a e FEMALE. e i e 5 a s ig y im esse e gi e as i ig (ΒΜΝΗ (= es i us ou i 1 u as ma gi a e icu a io qui e a ia e S emi e ea i y se a a e om es i us Β ou y a i g e w i is ig 1 5 Ge i a ia as i ig ; s e ma eca wi ou a o g i ge sca es s ea a so o e si es o e me as emum a s a k a uc i se io ; uc i s o i se e e a y a o e mos o e e i es a y a i g e es i u e o e a ica i o u sa e y a ec i i y o a a u e I a acks mai y e u k a LENGTH 1 3 -I 5 mm wi 7- 9 mm ick a c es o e os a
80 nf ld tl S rv
Dendrotrupes vestitus Broun constriction at ou e ase s a se y u c a e; se ae i ec e ec o y e ma e × 5 mm o 131 o g ee a e ai ua owa s me ia i e ose ea a e o ge Scu e um is i c wi i e i ica io a e ( M Η a a ec o y e ma e × E y a o o g-e i ic oi y co ca e y ou e a ase; a e a 95 mm o e a a as a o e u o i e i ica io a e ( M Η ma gi s cu e u e a y em aci g u e si e wi ou a (= cos ice s i e a ge; e y a i use y s ia e- u c a e o o sum ess so owa si es u s ia 9 qui e we e i e ; se ae o o sum e w i is - i ge sca es a e mo e o ess es ic e o e i ec e s aig ackwa s ea su u e s ig y ou wa s u - eu i es a e es i u e o e ec i i y is is i c y u e e o e si es i wi gs e i e y a se os e um o ou wa s a eas o i e s iae a 3 is s ecies goes mai y ema gi a e o ca a icu a e; a ica ma gi esce i g ea y o e me ium-si e a c es o e os a e ica y om o o um Mes as e um mo e a e y o g o ge a a mi -co a iame e ; me e is e um is i c w e o Gonoropterus Broun, 1904 = Pseudancistropterus i e u e e y a ma gi e i es 1 a use wi o 193 n.syn. wi ou a a y isce i e su u e; e i e 3 com e e y ee; Gonoropterus spinicollis Broun, 1904 = Pseudancistro- e i es a 5 use o a ia e eg ees e su u e a imes pterus horni 193 n.syn. is i c a imes o so esce a wi a ow a ge o oo o a omi a ca i y o co ae co iguous; mi e co ae a - Heteropsis latirostris Marshall, 1926 = Pentarthrum planum ow y se a a e oc a e s wi a o ge se a emo a e u - Marshall, 1937 n.syn. cu a e c a a e u a me o su ca e u e ea i iae wi s u s o a a e y so wi ou muc o o u cus; o i iae Macroscytalus Broun, 1 I = Baeorhopalus Broun, 1883 wi ou a o sa com u wi a e a com ; mi e i iae n.syn., Rhinanisus Broun, 1883 n.syn. wi a o g o sa com o a ica a o mo e; i i iae Macroscytalus parvicornis (Sharp, 1878) n.comb. (Rhinan- wi ou a o sa com u wi a asce i g i e o o sa com a si s ou ; first 3 segme s su equa i e g a wi isus) = Rhinanisus contiguus (Broun, 1 n.syn., R. wi i ose so es; c aw segme o us o ec i g we eyo fulvicornis (Broun, 1 n.syn., R. subconvexus Broun, segme 3 wi s o g o g i e gi g c aws i gu wi a 19 9 n.syn., R. sulcirostris Broun, 19 9 n.syn., R. suturalis ec a i g Broun, 19 9 n.syn. MALE. e gi e 7 wi ou ui ose a c es (wi g- o i g s uc- Macroscytalus remotus (Sharp, 1878) = M. badius (Broun, u e e gi e we e ose eyo e gi e 7 isi e u e - 1 n.syn., M. crenatus Broun, 1909 n.syn., M. depressus ea eyo as e i e S e i e a y sc e o ise is a Broun, 1883 n.syn., M. frontalis Broun, 1 93 n.syn., Μ u i i e wi a is i c s o g y sc e o ise a o eme S e - laticollis Broun, 1881 n.syn., M. russulus Broun, 1 1 i e 9 s ig y asymme ic wi qui e s o g y i e gi g a us n.syn. wi ou sc e o ise o es egme a ge ea y as o g as ae eagus wi a s e c e e o ga e i g a wi oi y oa Microcryptorhynchus perpusillus (Pascoe) n.comb. (Mio- se i e ous a ame es Ae eagus wi se a a e e o a ec- calles) um; a o emes o us o ge a ae eaga o y wi ou e we sc e o ise a ms o e use o u e a g e o e o e Microtribus W ll t n 1 73 = Mesoxenophasis W ll - o e o ec um I e a sac o g wi a a ge asa sc e i e t n 1 7 n.syn. a y o u i g a ase o sac wi a u u a o age i o m Microtribus brouni (W ll t n n b (Mesoxenophasis) e e sio ; e acu a o y uc i se i g a ase o asa sc e i e; wi 3 ma o a i io a sc e i es 1 me ia a a ica ea Myrtonymus n.gen. (Curculioninae: Erirhinini) os ium a occasio a y wi o e sma e ai y igme e e i a io o ame om G eek my os = my e a o yma = sc e i es as we ; wa s o sac wi i e y s i u ose es i u e ame o esig a io ; ge e mascu i e FEMALE. e gi e e i e a a ica ma gi S e i e wi e esse e igme e i wi i e s o e ec o su e ec s o g y i e gi g a ms; a o eme usua y i ec e s ig y o se so y se ae ea a os um o ec o y s ig y i e e me ia i e o e ig emis e i es weak y igme e wi ow wa om a is o o y co e gi g i o sa iew i a sma e o ga e a ica s y us; u sa a ge; s e ma eca a ge s aig i e om ase o ea o i se io oi o a e ae a u a e oug ou wi a e y sma g a c ose o uc wi ou a y co s ic io e wee ea a os um ea smoo i se io ; s e ma eca uc o igme e a ia e i wi g a ous wi ou a is i c o a im essio Eyes a se a e g i se i g o soa ica y o u sa os um ick as o g as o s o e a o o a e y s ig y TYPE SPECIES: My o ymus e a icus s cu e s o g y co e gi g i o sa iew o a ica i a i e wi e a a ica o io co e gi g i a e a iew o a e ; se ae ΙSΤ I U IOΝ l nd nd A tr l i ec e ackwa s o os os um o wa s o e os um ΗOS A U k ow o larvae; adults found consistently i Sc o es s o ea y o ei o m i ec e o ique y ow wa s; soi a ou oo e s o My aceae i ew ea a a Aus a ia seco a y sc o es o si es o os um a se A e ae s e e ; sca e is i c y i a e a a ica i i a e a iew e e i g REMARKS. My o ymus differs from the hypogean Erirhinini of o us eyo ase o os um; u ic e 7-segme e 1s seg- Eu o e a o Ame ica i e su i e aymo io ymi a i me icke a co si e a y o ge a segme s -7 a i g e os um s e c e a mos s aig o wa e ea mo i i o m e y s ig y wi e i g owa s c u ; c u wi e a os um iewe o sa y oi y a e i g owa s e i se - e o ga e-o a 1s segme a ou as o g as emai e wi a io oi s o e a e ae a e scu e um am y e ose asa a a me ia w i o se ae e as some imes s ig y ese ea u es a e iag os ic o Myrtonymina new subtribe, co s ic e o o a is i c y o ge a wi e wi ou ocu a w ic is wi es ea a a a e y we e ese e i Aus a- o es wi o wi ou a a ica co s ic io wi a u a ia a oug o s ecies a e ye esc i e
x n t 8
Myrtonymus zelandicus n.sp. Fig. 1 -11 o e Wai ema a o ma io w ic co sis s o i e e e i e a e ye owis o e is ow ; a emiae a egs usua y a sa s o e si s o e a mu s o e I as su seque y ee s a e a e ; a eima c u a e a u ic e os um wi i e es a is e a ese mi u e i wee i s a e co sis e y e o ga e a y co ue s a se u c a o o a 1 1-1 31× ese a ou e i e oo s o a a i e s ecies o My aceae o ge a wi e usua y 1 1-1 5× sο i ema e a 1 -1 31× om w ic sam es a e ee ake e was i g-sie i g i ma e a is o isc a ia y u a ways s a se y u c a e ec ique was a ie o ea y a soi sam es e a a wi some coa se u c a ea me ia i e E y a a is ac oss emai s u k ow i e oo s we e sc ee e o a ae o ei o sum wi a ia e i e u c a io i e u o o so esce wo ki gs wi ou success A u s we e ke o suc oo s u s iae e a su ace e y i e y u c a e occasio a y wi o ac ua ee i g was o se e iso a e a ge u c a o me as e um a e i es; a ome Hustache, 1937 Saccolaemus Kuschel, o ma e s i y oug ou a is u o im esse a ase a Neokiemosaccus = a e egs mo e a e y c a a e ( ig 1 7-1 9 ; i i iae wi 197 n.syn. o g se ae o i e o o sa com Neolaemosaccus narinus (Pascoe) n.comb. (Saccolaemus) MALE. S e i e wi sc e o ise a a a o eme as i ig 11 S e i e 9 as i ig 113 egme a ae eagus as i ig O moeacalles crisioides (Broun, 1 n.comb. (Acalles) 11 ; e o oa y ou e a a e ; asa sc e i e o us wi ai c igi ow cu i g u u a e e sio ; me ia sc e i e is Omoeacalles ovatellus (Β ou 1 1 n.comb. (Acalles) a e- ike wis e ; a ica sc e i es is i c Paelocharis setifer (Broun) n.comb. (Notiopatae) FEMALE. S e i e as i ig 115 Ge i a ia as i ig 11 ; s e - ma eca uc ess a mm o g e a e a u sa e Peristoreus cruciger ( ou 1 I = Ρ anchoralis(Broun, 1 1 n.syn., gracilirostris ( ou 1 1 n.syn. LENGTH 7 - mm wi 3- 7 mm
MATERIAL EXAMINED. ew ea a M Came ea Peristoreus fusconotatus (Broun, 1 = asperirostris Ηoυ ο a ma es 1 ema e Oc 19 e soi a ou oo e s (Broun, 1 1 n.syn., varius (Broun, 1 1 n.syn. o Me osi e os e ce sa C u c e ; Wai a gi agme s o y o 19 1 e soi o Μ e ce sa G Kusc e ; aw i i Peristoreus maorinus (Broun, 1913 =Ρ consonus (Broun, a i oo K ig s Is ma y agme s 1 ec 19 e soi o 1913 n.syn. M e ce sa G Kusc e ; u ukaka ma y agme s 3 o Oke reinstated (not synonymous with 19 e soi o M e ce sa G Kusc e AK Kawau I ag- Rhinoncus australis me s o y e 19 e soi o M e ce sa G Kusc e ; bruchoides (Herbst)) Waiwe a agme s o y 7 o 19 e soi o M e ce sa G Scolopterus aequus Broun, 1 = S aeneorufus Broun, Kusc e ; uia 1115197 1I9 ema e u a ma e a 1 1 n.syn., submetallicus Colenso, 1 n.syn. i yc om io acicu a e moss-ma u e e os e mum sco- S a ium M May; y ie M oski Auck a Ci y om Scolopterus penicillatus White, 1 = S.pectoralis Broun, 39 ma es 3 ema es Aug Wai aki S eam o Wa e ay 1 n.syn. 1975 o 7 May 19 e soi a ou oo e s o Me osi e os e ce sa M e o a a e os e mum sco a ium Ku ea e - Stephanorhynchus crassus Broun, 1 = S.fatuus Broun, coi es a o omy us u a a G Kusc e C e Ma ae oa 1 1 n.syn. i e a ie I 1719 ma eA e soi ( i sam e o y /59 o M e ce sa C u a Steriphus Erichson, 1 = Desiantha Pascoe, 1 7 n.syn. TYPE DATA. o o y e ma e 77 × 5 y ie mm fide E.C. Zimmerman (pers. comm.) Augus 1975 G Kusc e ( AC ; a a y es i AC Steriphus ascitus (Pascoe) n.comb. (Desiantha) OS Ρ AΝ S A u s associa e wi Me osi e os e os e - mum Ku ea a o omy us s ecies a My aceae Steriphus diversipes (Pascoe) n.comb. (Desiantha)
REMARKS. M e a icus is e sma es wee i so a k ow i Tanysoma Broun, 19 9 = Eutassa Broun, 19 9 n.syn. e wo I s wi e is i u io occυ e ce i some a e emo e aces a some geog a ica a ia io i e s a e o Tanysoma comatum ( ou n.comb. (Eutassa) e asa sc e i e sugges a is s ecies cou e e emic o ew ea a o a eas a i as ee es a is e o some Toura Broun, 19 9 = Merisma Broun, 19 9 n.syn., Protog- co si e a e ime ike e o e i e i i i e wee i s onum Broun, 19 9 n.syn. My o ymus e a icus a u s a e o e ou mai y i e o 1 cm o soi u i g e we e seaso a a g ea e e s Toura longirostris (Wollaston, 1 73 = fulva Broun, ow o 3 a cm u i g a o o ge y e io e i s 1 93 n.syn. s ecime s we e o ai e i y ie om e se ma s o e moss yc om io acicu a e associa e wi e os e mum Toura sharpiana (Wollaston, 1 73 n.comb. (Merisma) = sco a ium o a i e ec y ai e o e ye ow- ow Merisma aurantiaca (Broun, 1 93 n.syn., Stenotoura ea o si oam e u e o me om s o g y wea e e ocks prolixa Broun, 19 9 n.syn.
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(1 1 11 (1 ig. 0 l t l cos i e is ( eu i g uia 9 91 ae eagus o sa a a e a ; 9 s e ma eca wi uc ; 93 age i o m asa sc e i e ig. 8 84 l n r t ( a es oo K ig s Is ig. 4 6 l t l r ll a es y ie 9 95 1 ae eagus o sa a a e a ; 3 asa sc e i e o sa ; ae eagus o sa a a e a ; 9 s e ma eca wi uc s e ma eca us u sa Sca e a s = mm A e ia io s Ic a e a ca i a; e e o ; e ec um ig. 8 8 l n t l s y ie 5 - ae eagus o sa a a e a ; 7 asa sc e i e o sa a ig. 0 r t p l tr h s y ie 97 a - a e a ; 9 s e ma eca us u sa e a; 9 o e eg; 99 1 s e i es a 9 ma e e a ; 1 1 Sca e a s = mm A e ia io s c u sa co u a i ; c 1 ae eagus o sa a a e a ; 1 3 ema e ge i a ia ma i g c am e ; o w i e o aque a ea o ma i g c am e ; a e a ; 1 s e i e ema e e a ;1 5 e gi e e icu- o commo o i uc ; e e o ; igme e i g; s a io ema e o sa s e ma eca; s s e ma eca uc ; s g s e ma eca Sca e a s = 1 mm A e ia io s a a o eme; a a o- g a ; agi a ema i ge; sc a e- ike sc e i e
84 y ie ee e Su ey
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06- 6 M rt n z l nd s y ie 1 11 s e i e ema e; 115 11 ge i a ia ma e a ema e a i us; 1 7 ig o e eg; 1 e mi e eg; 1 9 e i a e a eg; 11 a ome ma e s owi g e gi e a e ; 111 e a ig 1 11 111 o same sca e 1 7-1 9 o same sca e su ace ema e; 11 s e i e ma e; 113 s e i e 9 ma e; 11 -11 o same sca e f r n 85 E E E CES ai 1977 y u gus ig i e a ( a icius (Co eo e a 1886 Manual of the New Zealand Co eo e a Parts Sco y i ae Forest and timber insects in N.Z. 18. III & IV. Wellington, Colonial Museum and Geologi- cal Survey Department. i- ii 7 5-973 Baker, 19 5 New ladybird. y ie Plant Protection Centre circular, entomology, March 1985: 1-3 1893a Remarks on the Ca a i ae of New Zealand. Tra sactions and proceedings of the N.Z. Bates, W 1 7 On the o gico Co eo e a of New I stitute 25: 19 -19 Zealand. Annals a magazine of natural history (4) 14: 1 - 11 -131 1893b Manual of the New Zealand Co eo e a Parts - II Wellington, Institute. - ii 1876 New species of o gico Co eo e a from New Zealand. Entomologists' monthly magazine 13: 975-15 5 -5 1893c Descriptions of new Co eo e a from New Zealand. Annals magazine of natural history (6)12: Beier, Μ 197 The seu osco io s of New Zealand, Norfolk, and Lord Howe. .Z. journal of zoology 3: 1 1-195 -3 37 -39 199- 1903 Descriptions of new genera and species of New Zealand Co eo e a Annals a magazine of Belon, M 1 Revision des a i ii ae de Nouvelle- natural history (7) 11: 5 - 5 - 1 ; 7 (12): éIa e Revue e omo ogie Caen, 3 - 9- e aue Μ 19 3 Wei e e eue S a y i i e aus Neu- 1904 Descriptions of new genera and species of See a (New Zealand). o ia zoologica et hydrobi- New Zealand Co eo e a Annals a magazi e of o ogica 12: 17 -1 natural history (7) 14: 1-59 1 5-1 7 eu i g S 19 Novae species Ce am yci a um iii- 1909a Descriptions of new genera and species of o ia zoologica et y o io ogica 10: 37- 5 New Zealand Co eo e a Annals a magazi e of 115- 1 7- 37 natural history (8)4: 51-71 13 -1 1 75- 91 1943 Novae species Ce am yci a um ii o ia 1909b Revision of the New Zealand Cosso- zoologica et y o io ogica 12: 1 - i ae Tra sactio s of the N.Z. i stitute 41: 151- 15 1950 Revision des a me i i o gicο ia 1: 1910 Desc iptions of new genera and species of 9-159 Co eo e a Bulletin of the N.Z. i stitute 1: 1-7 1959 Nouveaux genres de amii ae (Co eo e a 1913 Desc iptions of new genera and species of Ce am yci ae Bulletin e a a es de la Sοcié é οy- Tra sactio s a proceedings of the N.Z. ale Ε omοΙogie de Belgique 95: 7 - Co eo e a i stitute 45: 97-1 3 19 Diagnoses ρ éΙimi ai es de nouveaux 1914 Desc iptions of new genera and species of amii ae du Μυséυm National is oi e a u e e de Bulletin of the N.Z. Institute 1 (3): Paris (Co eo e a Ce am yci ae A a es de a Soci- Co eo e a 1 - 9- 9 é éE omo ogiquede18: France (n.s.) 1917 Desc iptions of new genera and species of Britton, E 19 New Zealand cave Ca a i ae (Co eo- Bulletin of the N.Z. institute 1 (5): e a Annals a magazi e of natural history (13)3: Co eo e a 3 7- 7 1-1 Desc iptions of new genera and species of Brookes, A Ε 193 A new genus and six new species of 19 1 Co eo em u e i o e i s i u e 1 (6): 75-59 Co eo e a Tra sactio s a proceedings ο e Ν Ζ i stitute 63: 5-33 1923 Desc iptions of new genera and species of Bulletin of the N.Z. institute 1 ou Τ 1 Manual of the New Zealand Co eo e a Co eo e a ( Wellington, Colonial Museum and Geological Survey 7-7 Department. i + 51 Brown, G 19 7 Notes and records of New Zealand N.Z. entomologist 3: -5 1881 Manual of the New Zealand Co eo e a Part Sca a aei ae ΙI We i g o Co o ia Museum a Geo ogica Buck, F.D. 1955 A provisional list of the Co eo e a of Survey Department. i- iii 53-7 Epping Forest. The e omo ogis s mo y magazi e 1883 Revision of the New Zealand Cosso i ae 91: 17 -19 wi descriptions of new species. N.Z. journal of sci- 1959 Notes on the Co eo e a of Monks Wood, e ce 1: 7- 99 Hums. Entomologists' gazette 10: 3-93 Abstract of paper on New Zealand 1 5 Scy - Cameron, Μ 19 5 Some observations on the S a y i i ae .Z. journal of scie ce 3 -3 7 mae i ae of the ou Collection of Co eo e a in the British Museum, with descriptions of new genera and O E I c u es o y ci a io s o i s eco s o o eig s ecies i ew Zealand, and authors cited i the o uc io a a o- species. Annals a magazi e of natural history (11) omic Notes. 12: 15 -1
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Came o M 195 ew s ecies o S a y i - a icius C 1775 Sys ema e omo ogiae e s u gi i ae Entomologists' monthly magazine 86: 3- i siae [ ] + 3 Came o a C u c e 19 i a a i g a 17 1 S ecies i sec o um 1 am u gi e Ki o ii soi sam i g o Sca a aei ae (Co eo e a a os- iii + 55 si e e a o s i some o a a Auck a as- ai mai e 1 3 esc i io e ois ou e es es èces u es 113-117 i Τ K C os y a o i ge e Cο éορ è es e Océa ie Annales de la Société (e s ocee i gs o e Aus a asia Co e e ce Entomologique de France (2) 1: 11-15. o G ass a I e e a e Eco ogy au e A 19 Su es o y e ie s e ou e e e a e C a k A 193 I sec s i es i g Pinus radiata i ew Revue d' entomologie 19: 1 1-1 9 ea a .Z. journal of science and technology 13: 35- 3 emi g C A 1977 e is o y o i e i ew ea a o es s N.Z. journal of forestry 9- Co e so W 1 5 Memo a a o a e cu sio ma e i e o e Is a o ew ea a i e summe o a kie G W ; E e E 197 Eco ogy o i sec s i u a 1 1- ; i e e as a co i u io owa s e asce - e i o me s Annual review of entomology 23: ai i g o e a u a o uc io s o e ew ea a 3 7-3 7 g ou ; wi a icu a e e e ce o ei o a y Tas- a 19 7 u Ke is e Scy mae i e au a o manian journal of natural science 2: 1 - 3 1-3 a ei ame ika Biologie de l'Amerique Australe 3: 1 O some ew a u esc i e s ecies o 11-7 ew ea a i sec s o e o e s O o e a a 1975 Revision der Scydmaeniden von Aus a ie Co eo e a Transactions and proceedings of the N.Z. eusee a u e e ac a e I se Denkschrif- Institute 14: 77- ten, Österreichische Akademie der Wissenschaften, 1 A esc i io o a ew s ecies o Coccinella Mathematisch-Naturwissenschaftliche Klasse 118: ou i ew ea a Transactions and proceed- 1-31 ings of the N.Z. Institute 20: - 1977 eue Scy mae i e (Co eo e a aus Co ie W 195 I sec es s 9- 1 i A ki so eusee a Samoa e o ga-I se u Cook- e a a o ec io i ew ea a We i g o I se Koleopterologische Rundschau 53: 15- 5 Go e me i e 19 Wei e e ei ag u Ke is e Scy - Cum e A 1959 e i sec com e o sow as u es i mae i e au a eusee a s Sitzungsberichte der e o Is a I e Co eo e a as e ea e y Österreichischen Akademie der Wissenschaften, Mathe- summe swee -sam i g ou a o agricultural matisch-Naturwissenschaftliche Klasse, Abt. 1, 189: research 7 3-77 9-313 Cu i g am G 19 a iseases i isio Owai- 1985 4. ei ag u Ke is e Scy mae - aka Auck a DSIR Annual Report 14: 17- i e au a eusee a s Sitzungsberichte der Öster- reichischen Akademie der Wissenschaften, Mathema- tisch-Naturwissenschaftliche Klasse, Abt. 1, 194: a is K 197 U a isa io a e i e si y o i - 197- sec s 1 -13 i A Mou & Wa o (e s ogga W W 19 Aus a ia a y i ee es The agri- i e si y o i sec au as Sym osia o e oya cultural gazette of New South Wales 13: 95-911 E omo ogica Socie y o o o 9 o o ack- we Scie i ic u ica io s Gou ay E S 19 u e s ose wee i Asynonychus (Pantomerus) godmani C o c i ew ea a N.Z. E ö y-You ga S 199 C am i ae o ew ea a journal of science and technology a - 5 (Co eo e a Euci e oi ea N.Z. journal of zoology 17: 119-13 1954 Records of introduced Coleoptera and notes o e 1953-5 co ec i g seaso .Z. entomologist E ic so W 1 ei ag u au a o a ieme s a 1 (4): -1 mi eso e e ücksic au ie geog a isc e e - ei u g e I sec e Archiv für Naturgeschichte 8: 1955 Notes on exotic and New Zealand insects. N.Z. 3- 7 entomologist 1 (5): 3-5 Es año 197 Cο i uciό a co ocimie o e os 1964 Notes on New Zealand insects and records of Anobium y ge e os a i es e Aus a ia y ue a Zel- i o uce s ecies N.Z. entomologist 3 (3): 5-51 anda. Misce á ea οο όgica 3 1 1-1 Esso M 1975 o es o e io ogy a is i u io o ammo M 197 A e iew o e ge us Anotylus ee ece y isco e e e o ic wee i es s i awkes C G omso (Co eo e a S a y i i ae Bulletin ay - 1 i ocee i gs o e of the British Museum (Natural History), entomology Wee a es Co o Co e e ce 33: 139-1 7
f r n 8
a iso R.A. 1955 o es a eco s o some es s o 1979 e ge e a Mo o oma e s (Rhizo- eco omic im o a ce Ν Ζ e οmο οgis (5): 1 -13 phagidae) a A omma us Wesmae (Cerylidae) i Hawksworth, (ed.) 197 e c a gi g o a a au a ew ea a (Coleoptera). N e omo ogis 7(1): o i ai Sys ema ics Associa io s ecia o ume o - o o Aca emic ess 1 u so G.V. 19 3 A i e o ew ea a ee es a e i A A 19 eνisiό eI gé e ο A sy ο yc υs C o c a sac io s a ocee i gs o e I s i u e 54: (Coleoptera: Curculionidae). e is a e a Asocia- 353-399 ciό e Cie cias Naturales e i o a 17: 1 1-17 193 ew ea a ee es a ei a ae We - ea A Μ 1 95 esc i io s o ew s ecies o Aus a ia i g o e guso & Os o e 3 Coleoptera. ocee i gs o e i ea Socie y o Hustache, A 1937 Mag a i ae (Co eo e a Curculionidae) ew Sou Wa es (2) 10: -319 e Amé iqυe Μé i io aΙe A a s a maga i e o Ma s a G A K 19 Some ew Curculionidae om ew a u a is o y (10) 19: 19 - ea a (Co A a s a maga i e o a u a is- u o W 19 I e au ae o ae Zealandiae. o - o y (9) 18: 1-1 o Dulau viii + 37 1937 ew Curculionidae (Co om ew ea a a sac io s a ocee i gs o e oya o so C 1971 A oma ii ae (Co C y o agi ae om Socie y o 67: 3I -3 e o e a s o e I ia su co i e wi es- Ma i Ν A 19 3 Misce a eous o se a io s o a as u e c i io s o se e ew s ecies E omo ogis s mo y au a a a o a e s ecies is SI E omo ogy maga i e 106: - 3 i isio e o o 3 9 1975a A o o a o e su a a ic is a s o Ma ews A 1 7 o es o Aus a asia a o ew ea a Coleoptera: Ptiliidae. N ou a o Ame ica ic o e ygia Cis u a e omo ogica 1: oo ogy 9-1 95- 99 1975 Co i ica a a ew ge us o Co ica ii ae May B.M. 19 3 ew e omo ogica eco s N.Z. e o- (Coleoptera: Lathridiidae). E omo ogica sca i a- mo ogis 3 ( -53 ica 6: 3- 5 19 e imma u e s ages o i oge aeus u a us 19 A i o uc io o e Ptiliidae (Coleoptera) (Co eo e a Curculionidae: Baridinae) a i s io - o ew ea a N ou a o oo ogy 9: 333-37 ogy i Auck a ew ea a e omo ogis 8: 9-35 Kaszab, Ζ 19 1 ie Ga u ge u A e e i us Mi e 1931 o ious wee s co o esea c ρ 19- A c eoc y ici i (Coleoptera: Tenebrionidae). o ia i e o e a me o Scie i ic a I us ia e omo ogica u ga ica 42 (34) 1: 95-115 esea c Ki g 1 3 O e Pselaphidae o Aus a ia a sac- Mi iga 19 9 I sec amage o euca y s Ρ i io s o e E omo ogica Socie y o ew Sou Wa es o es esea c I s i u e e o 19 1: 37-5 197 O e seas woo - a a k- o i g i sec s i e - Ki gs ey I 1 9 O e occu e ce o e ack i e ce e a ew ea a o s N o es Se ice (O io y c us su ca us i e so a sac- wee i ec ica a e o 57 79 io s a ocee i gs ο e Ν Ζ I s i u e 33 -3 Montrouzier, Χ 1 1 Essai su a au e e omo ogique e Ki k T.W. 1 95 i isio o io ogy a omo ogy a Νoυνe e-CaΙ o ie ( a a e e es î es es i s 1 1-1 i e o e a me o Ag icu u e ew A i u e c CoΙéo è es A a es e a Socié é ea a E omo ogique e a ce (4)1: 5-3 1 9 i isio o io ogy a omo ogy Motschulsky, 1 E umé a iο es ou e es es èces 11 -177 i e o e a me o Ag icu u e ew e Co éο è es a ρο és e ses oyages u e i e ea a a Sοcié é es a u a is es e Moscou 37 97-355 19 i isio o io ogy a omo ogy Muggeridge, 1933 Some ece y eco e e o ic i sec 7 -3 i e o e a me o Ag icu u e ew es s ou a o ag icu u e 47: 1- ea a K aus i e ; Sei e (e s 19 OkoΙogie u a e Sys eme Wisse sc a ic e ei sc i e Ka -Ma - Ordish, G 19 Hydraenidae (Insecta: Coleoptera). U i e si ä ei ig Mathematisch-Naturwissen- au a o ew ea a 6. 5 schaftliche Reihe 35(6): 591- Kuschel, G e o eig Cυιcu iο oi ea es a is e i ew ascoe F.P. 1 7 esc i io s o some ge e a a s ecies ea a (Insecta: Coleoptera). Ν Ζ οu a o scie ce ο Aus a ia Cu cu iο i ae a sac io s ο e Ε ο- 15: 73- 9 mο οgica Socie y o o o 1870: 1 1- 1
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Paulian, R. 1944 Les Aglyceridae, une famille relicte Australian beetle now established in New Zealand. (Col.). v r nç d nt l 1 1I3-119. j rn l f z l 7: 407-416. Pilgrim, R.L.C. 1963 An d t l b n t t br a cara- Thomson, G.M. 1922 Naturalisation of animals and plants bid beetle new to New Zealand, and a review of the in New Zealand. Cambridge University Press. x + 607 exotic carabid fauna. f n t 5: 837-847. P. Todd, D.H. 1961 First record of the occurrence of t - Redtenbacher, L. 1867 Reise der österreichischen Fregatte l t rn bd n l (F.) (Coleoptera: Hydrophil- Novara urn die Erde, Zoologie 2 (Coleopteren). Wien. idae) in New Zealand. nt l t ( 25-27. iv + 249 p. Richards, A.M. 1959 Revision of the genus S pr t Voss, E. 1936 Über die Tribus Scolopterini sowie einige Redtenbacher, 1858 (Coleoptera: Aphodiinae) in New verwandte Gattungen und Gruppen von Neu-Seeland. Zealand. r n t n f th l S t f 87: Arb t n üb r rph l h nd t x n h 35-50. Ent l rl n- hl 3 602, 110-12I.
Schedl, K.E. 1962 Scolytidae und Platypodidae aus dem Wakefield, C.M. 1873 Remarks on the Coleoptera of Can- australisch-polynesischen Raum. Ent l h terbury, New Zealand. r n t n nd pr d n Arb t n d M G r 13 72-78. f th n t t t 5 294-304. Scott, L. 1983 Riverhead Biological Reserve. h W t 1874 Note on the occurrence f r t l rd r- 26-36. and h r nth r rv in Canterbury, New Sharp, D. 1874 Descriptions of new genera and species of Zealand. r n t n nd pr d n f th Pselaphidae and Scydmaenidae from Australia and n t t t 153-154. New Zealand. r n t n οf th Ent l l Walker, J.J. 1904 Antipodean field notes. II. A year's insect S t f nd n 1 7 438-518. hunting in New Zealand. h nt l t nthl 1876a On the Colydiidae of New Zealand. Ann l z n 24-28, 68-77, 115-126, 149-154. nd z n f n t r l h t r ( 1 17-29. Welch, R.C. 1973 Coleoptera. Pp. 212-233 in R.C. Steele 1876b Descriptions of some new genera and spe- & R.C. Welch (eds), Monks Wood - a nature reserve cies of New Zealand Coleoptera. h nt l t record. Huntingdon, U.K., The Nature Conservancy. nthl z n 13 20-28, 70-77, 97-102. White, A. 1846 Insects of New Zealand. Pp. 1-24 in J. 1877 On the Elateridae of New Zealand. Ann l Richardson and J.E. Gray (eds), The zoology of the nd z n f n t r l h t r ( 19 396-413, voyage of H.M.S. Erebus and Terror, under the com- 469-487. mand of Captain Sir James Clark Ross, during the 1878 Descriptions of eight new species, and a new years 1839 to 1843, Part XI. London, Longman, genus of Cossonides from New Zealand. r n t n Brown, Green, & Longmans. f th Ent l l S t f nd n 1 7 9-14. Williams, S.A. 1976 The genus Ol t (Coleoptera: Sta- 1882 On aquatic carnivorous Coleoptera or phylinidae) in New Zealand. j rn l f z l Dytiscidae. S nt f tr n t n f th l b- 3 247-255.
l n S t ( 179-1003. Wittmer, W. 1979 Zur Kenntnis der Cantharidae (Col.) 1884 Revision of the Hydrophilidae of New Zealand. Neuseelands. Ent l l n 275-325. r n t n f th Ent l l S t f nd n Wollaston, T.V. 1873 On the genera of the Cossonidae. 1 465-480. r n t n f th Ent l l S t f nd n 1886 On New Zealand Coleoptera, with descriptions 1 73 427-657. of new genera and species. S nt f tr n t n f 1874 Diagnostic characters of undescribed Cos- th l bl n S t ( 3 351-454. sonidae, no. 1. C t l nt l 1 I99-208. 1889 Two new species of S n n t l f 1 Wood, S.L. 1966 New synonymy in the Platypodidae and 364-365. Scolytidae (Coleoptera). Gr t n n t r l t Spiller, D. and E.G. Turbott 1944 The occurrence of some 17-33. Australian insects and a spider in New Zealand. - rd f th A l nd In t t t nd M 3 79-83. Zondag, R. 1953 Forest insect survey. r t In t S r- v n l tt r 3: 1-21. Thompson, R.T. and J.E. Marshall 1980 A taxonomic study 1957 Forest insect survey and life history. r t of h l r n fοr (Coleoptera: Phalacridae), an In t S rv n l tt r 1-32. a es 9
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222 oeosi us cu essi 22 y e o i us euca y icus 224 a y us a ica is App nd x 101 App nd x EGE A IO O E MANUKAU COAS A LYNFIELD, AUCKLAND a Ε Es e Native vegetation Olearia furfuracea, and Cyathodes fasciculata. There is a The vegetation of the coast between Wairaki Stream and dense ground cover of rank S h n t nd and tussocks Wattle Bay is predominantly indigenous, some quite of G hn ιfοΙ Gumland vegetation grew with Aca- modified, and some in a more or less natural state. The cia rn nearby on the Sylvania block, until recently Auckland isthmus in 1 was mainly scrub-covered it was cleared for housing. At the upper edge of Subritzky's with scattered remnants of forest in gullies. Most of the bush by the Wairaki Stream the presence of M l t usable land was thereafter cleared for farming. In many r flοrυ suggests a more fertile soil, but p d p r places along the steep Manukau coast the farmed land laterale, S h n t nd and other sedges indicate its reverted to a wild state with a predominance of native gumland origin. Near Strathnaver St up till the late 197 plants. In parts the present vegetation dates from the aban- there remained some poor pasture with Ul x r p donment of farming, in others from the last fire, distur- and G hn t f l Scrub οn t d b v th ll bance from the laying of the sewer line, or slumping of the grew d rr kumeraho, P. ph l f l and M r - land surface. l t affinis,alleth typical gumland species. Much of The underlying sandstone with bands of siltstone is gumland has now been taken over for housing because of fairly uniform in composition, but topographic differ- the easy topography. ences promoted vegetation of several kinds. This pattern has persisted through a long period of depletive Maori Gully forest zone occupation and the post-1 changes. Many centuries Ribbons of gully forest along streams have some large ago kauri (Agathis austraiis) grew on the easy topography t x l n mid x l p t b l trees which are of the ridges and left behind in the soil lumps of resin relics of a former vegetation. The epiphytic asteliad Co - which have given rise to the name 'gumland' for pt - lospermum h t t is mostly confined to this older p r scrub which replaced the kauri. The valley sides vegetation. In this moist atmosphere are many ferns and favoured broadleaf forest of t x l n and x l the vines p n nd n and r n t b r - p t b l The soil here is more fertile because gravita- n This is the prime habitat of the tall tree fern C th tional movement and mixing allows the subsoil to yield d ll r and the palm h p l t l p d This kind more of its nutrients. Examples of forest of this kind of vegetation almost certainly covered the hillsides. Its remain in deep gullies where it was reached less fre- canopy would have contained also Beilschmiedia tawa, B. quently by fires, was burnt less intensively, and recovered t r r and C r n rp l v t and above them more quickly. After the fires, shrubs and small trees scattered emergent r d pr n r n - established. On the exposed slopes above the shore, forest p t f rr n and M tr d r r b t The tall of a special kind developed, with M tr d r x l as Dacrycarpus dacrydioides grew along the streams mainly, its dominant tree. In the splash zone on the shore are and is still represented there. pockets of salt-resistant herbs. At the mouths of the larger A single burn would have killed many large trees and streams erosion debris accumulated behind sandy beaches promoted a phase of tree ferns, shrubs, and small trees. A to form small wet patches which developed into open new succession began from bare ground after repeated swamps and semi-swamp forest. burns, and from pasture. pt p r p r may have been the first woody pioneer. Little of this can now pt p r zone be seen because of its replacement by M r n tr l The nature of the original kauri forest can only be assumed d r rb r a C th d lb t a in the from the structure and composition of other kauri forest in more fertile places by M l t r fΙοr and C th the region, but some typical gumland persisted into the d ll r On some of the ridges are groves of S ph r 197 It derived much of its character from the very r ph ll mid here and there isolated Metrosideros ex- impoverished soil and the frequent burning which pro- l The gully forest contains many minor elements moted pt p r p r as its dominant plant. such as Dodonaea v tt p r tenuifolium, After a burn gumland has some colourful shrubs, and is Schefflera igitata, Alectryon x l El rp d n- rich in sedges, ferns, and orchids until sombre pt p r- t t and K nz r d The role of K nz is not arid the rush-like sedge S h n t nd take over a clear. Most likely it established in minor amounts at the decade or so later. When Colenso passed through here in same time as Leptospermum and, being longer-lived, has 1 1 he saw "open barren heaths" and "dreary sterile persisted. There is also the possibility that it replaced wilds". There is some advanced gumland vegetation in pt p r locally, or was more general and was Manukau Domain. In the canopy with pt p r removed for firewood. lt is less tolerant of poor soils a and the naturalised r are M r n tr l pt p r and was not prolific on the gumlands.
102 nf ld tl S rv
Coastal forest zone triche muelleri, Juncus spp., Scirpus chlorostachyus, S. Pohutukawa forest develops directly by establishing on reticularis), and where water seeps down sandstone slopes open ground, or less often after a series of vegetational (Machaerina sinclairii, Blechnum sp. `black spot', Cot- stages which stabilise a slope, at least temporarily. The ula membranacea). succession that began on slips with Hebe stricta, Coriaria arborea, Astelia banksii, Entelea arborescens, Cordyline Exotic vegetation banksii, and Brachyglottis repanda now has fewer of Plantations. Some of the plant communities are man- these species and more alien plants. If further slipping made or man-induced. Most conspicuous are plantations does not occur these are joined by Pseudopanax lessonii, of Pinus and Acacia on the Sylvania block, established Pittosporum crassifolium, Metrosideros excelsa, Sophora perhaps 40 or 50 years ago. The vegetation at that time was microphylla, and others, sometimes Corynocarpus laevi- probably low scrub. Acacia mearnsii reached maturity, gatus. The lower layer usually contains Macropiper ex- and many of the large trees collapsed. Seedlings have celsum, Coprosma macrocarpa, Polystichum richardii, established to a limited extent. Native scrub persists Gahnia lacera, and Carex, flagellifera. beneath this open canopy. In the gaps Leptospermum The seaward fringe provides a habitat for other species. scoparium, Hakea sericea, Cyathodes fasciculata, and Near the splash zone Poa anceps and Scirpus nodosus Ulex europaeus are prominent. Elsewhere there are con- have a foothold, and in the wind funnels at the mouths of spicuous amounts of Cyathea dealbata, Geniostoma some minor streams there is a local abundance of Phormium rupestre, Gahnia setifolia, and Schoenus tendo. tenax, Coprosma robusta, and Cortaderia splendens, These communities were reduced in size when much of usually growing with Pseudopanax lessonii. the easier topography was taken over for housing in the A stand of Metrosideros excelsa near Wattle Bay is the mid and late 1980s. On the steeper slopes vigorous gully most natural piece of forest vegetation along this coast, forest prevented any re-establishment of Acacia mearnsii and is the finest example of Auckland's coastal forest. and of A. decurrens except in some disturbed places. The Pinus radiata plantation on the spur in this region Splash zone is quite open beneath and, typical of pine plantations, has Small portions of the splash zone are stable enough to an understorey of native plants — mostly of species with support higher plants. They are species commonly seen in bird-borne seeds, particularly Melicytus ramiflorus and salt marshes (Sarcocornia quinqueflora, Stipa stipoides, Myrsine australis. Selliera radicans, Samolus repens) or coastal sand and estuary margins (Leptocarpus similis, Scirpus nodosus, Grassy areas. Little pasture remains, and the grassy Apium prostratum, Triglochin striatum, Lilaeopsis areas are mainly lawns and playing fields. These consis- novaezelandiae). Lobelia anceps, the most versatile of ted of a wide range of species until the invasion of Axono- these splash zone plants, grows in many diverse habitats. pus affinis in the 1970s. The overwhelming advance of Missing from this part of the coast are Senecio lautus Pennisetum clandestinum is further reducing the variety. and Coprosma repens, which require a more stable sub- Domestic gardens. Absence of severe drought and of strate. very low winter temperatures make the climate favour- able for plants from many parts of the world. Although Wetlands temperate plants predominate, Northern Hemisphere Where accumulating beach sand restricted the stream broadleaved trees (Quercus, Ulmus, Fagus, Fraxinus) outfall at Wattle Bay there would have been a wetland of and conifers (Pinus, Cupressus, Cedrus) are less favoured Dacrycarpus dacrydioides, Cordyline australis, Phor- than in cooler parts of New Zealand. More prominent are mium tenax, Scirpus fluviatilis, Typha orientalis, and the Australian genera (Eucalyptus, Acacia, Acmena, Syzygium, native form of Calystegia sepium. The Dacrycarpus has Tristanea) and New Zealand trees (Agathis, Dacrydium, gone, and in its place grows Salix fragilis. Silt carried by Podocarpus, Sophora, Metrosideros), and a few from the stream from disturbed land at the head of the catch- warmer climates (Jacaranda, Melia). ment, and bulldozing, altered the flat in the late 1970s, Among smaller plants are cultivars of native plants reducing the relic native community but promoting Carex (Hebe, Pittosporum, Coprosma, Phormium) and many lessoniana and the exotic Lonicera japonica. Festuca species from Australia (Callistemon, Grevillea, Banksia, arundinacea and Pennisetum clandestinum grow in abun- Telopea) and from southern Africa (Protea, Leucosper- dance on the stable beach sand, and Atriplex prostrata mum). Some of the most popular garden subjects are where it is disturbed. A similar but smaller delta may have citrus, Nerium oleander, and Hibiscus rosa-sinensis. Fruit existed at the mouth of Wairaki Stream. trees, herbaceous perennials, bedding plants, and vege- Other wetlands are very small. A depression contains tables are predominantly of Northern Hemisphere origin. ph r nt l h tt n t r t fοl C l - t p and Isachne globosa A few individuals of Escapes from cultivation. Introduced plants which have wetland species grow on open stream margins (Carex escaped from cultivation are a conspicuous part of the geminata, Cyperus ustulatus, Polygonum decipiens, Auckland landscape. However, the Manukau coast is not Scirpus fluviatilis, S. medianus), on wet tracks (Calli- as seriously infested as some other districts, because of the
App nd x 0 competition offered by the native vegetation and the low d l t t ( r t f S rtz G incidence of dumping of garden refuse. fl b ll t b ll G There were two waves of invasion. The first was in the Η r r Βr G farming period when h lt r pl nt escaped. Among them r h n l n t A C nn G v n r G r Alb z l ph nth A rn r Η l fοΙ n r d t and Ul x r p At the YCO O IACEAE same time birds carried the seeds of S l n r- p d rn L. d t r d n rt r t n and b fr t In the second wave in the last few decades were Asparagus p r d A n- MA A IACEAE d n d h rdn r n tr l d and M r tt l n E S th G L. n n with bird-borne seeds, and Ar j r f r ΟSΜUΝ ACΕAΕ x tt t and C rt d r pp part cularly C pt pt r h n ph l d C h (A r l G j b t with seeds carried by wind. Τr d nt flυ n- O Y O IACEAE n almost certainly arr ved as garden refuse. h t r d v r f l (W lld h S r G nd n ( r t f h S r G rr rp n ( r t f Ch n G C SI O ACEAE h n t v fl r pt r l n t A n rd G The principal habitats are: C t l f r tz n ; G gully l n l t A n rd G forest zone; pt p r zone; S splash zone; W t nn n (Spr n l rnh G wetland; O other — track marg ns, etc. E I ACEAE
E I O ES Ad nt nn n h G A f lν l G AS E IACEAE ll r t nd f l ( r t f Hook. G A pl n b lb f r r t f G t r l nt A h G A fl d r t f G tr l r G A h r n C I G A I pr ph ll C r G SC I AEACEAE A bl n f l C l G d rt l t A h G A p l d n r t f G E Y E I ACEAE
EC ACEAE n t pt r p nn r ( r t f ltt G l hn h b r nd l G f l f rrn (A. C nn Ett n h G GYM OS E MS br n ( M tt G O OCA ACEAE Β n ( r Ett n h I r rp d r d d (A h b nf G p (Gr n f r G r d pn n Lambert G p (bl p t f r G I W h ll l d tr h n d n G d d r p tr l rr G r n p t f rr n ( n b nf G ll rr G CYA EAC ΕAΕ MO OCO Y E O S C th d lb t ( r t f S rtz I G AGA ACEAE C d ll r ( r t f S rtz G C rd l n tr l ( r t f End G W E S AE IACEAE C b n f G C t pt r n ( h nb S th G C p l f G I nd l n r S rtz h r n l C G b r A h (K hn G t n x t G r t C W t r d l nt ( r t f C n I CY E ACEAE
ICKSO IACEAE h tt n (K r l W n rr ( r t f S rtz G t r t f l ( r ll W C r x d t tt in f G YO E I ACEAE C fl ll f n C l C G tr p l b ll (A. C nn nd l G C n t S h hr W h p d (Swartz) nd l G C l b rt n tt n f G r r nd l C l n n St d W p p nt n l r (C l nd l G C hr (Ch bl n G v l t n (A h nd l G C l ndr tt n f G l t h r h rd ( S th C C p n r tr C l C hr d nt f r ( r t f Ch n G C v r t tt in f W G EIC E IACEAE C p r t l t A h W St h r nn n h ( rd x Ch n El h r t r W YME O Y ACEAE G hn l n St d C G n ph ll d ( r t f S rtz G G p fl r K r G
04 nf ld tl S v
G t f l (A h f I G ICO Y E O S
G x nth rp ( f f G A SEUOSMIACEAE p d p r tr l (A h f Al r ph ll A C nn G l t r l r M h r n n l r ( f K W A IACEAE Ap pr tr t b ll x nt S M r l t ff n ( r n l C nt ll n fl r (C l nnf S h n br v f l r dr t l h t r t f Ο S h l n t S h lt W l p n v -z l nd (G nd A W ll S S t nd ( f f S rp rn hl S A OCY ACEAE S hl r t h v n W r n h t r ph ll A C nn G S fl v t l ( rr Gr W A A IACEAE S n nd t ( r Spr n W d p n x rb r (M rr h l p n G S d n C W Ρ r f l S l x A C nn G S n d ttb S Ρ l n ( C C K h C S r t l r (C l Ed r W Ρ l n x r f l G Un n b n tt n f G S h ffl r d t t t G r t G
U n n t ( nn f K G AS E ACEAE I I ACEAE r h l tt r p nd t G r t C G b rt x d ( r t f Spr n G C t l r n p f l S U CACEAE C br n l d W n tr l f Ο Gn ph l l t lb Ο pl n f l r W G ph r W lld Ο pr t rp r W Ol r f rf r ( h f O r n (A C nn r G U CAG Ι ACEAE S n l r t f x r t Ο r l h n tr t z t v S S h p d l A h Ο EM ACEAE n nοr W CA I IC ACEAE C ll tr h ll r S nd W I IACEAE Arthr p d rr t ( r t f r C CAM A U ACEAE W hl nb r r l ( r t f S hr d 0 A t l b n A C nn C A l ndr A C nn G C E O O IACEAE C ll p r h t t (C l S tt b G S r rn n fl r (Un -St rnb A S tt S n lI n r C l G CO O U ACEAE O C I ACEAE C l t p ( r W t r t l l b l ( t h M r G C t r r ( r t f r x f G Ρ b n A C nn n G CO IA IACEAE A MAE: A ECACEAE C r r rb r nd C h p l t l p d W ndl t r d G CO Y OCA ACEAE A A ACEAE C r n rp l v t t G r t G r n t b r n Endl CU O IACEAE p b n (C nn St n G W n nn lv l S l x A C nn G
OACEAE OSE ACEAE C rt d r pl nd n C nn r C r r p lt t h nb x b ll rd r K nth S p r l t ( l n h n C nn dr t nth E AEOCA ACEAE h l hn r n t f C El rp d nt t ( t G r t hl G I hn l b ( h nb K ntz W E AC I ACEAE hn r t f lf r ( r t f r n S C th d f l t ( r t f All n G M r l n v n ( l f G C j n p r n ( t G r t r M t p d ( b r 0 r ph ll n l r Ch Opl n b ll G n p r t f C ESCA O IACEAE t d p r b nn l r ( t v C nn r & Ed r C rp d t rr t t G r t G St p t p d ( f ld S A ACEAE ES IO ACEAE C r h l l r S p n G pt rp l Ed r S S ph r r ph ll A t G C
SMI ACACEAE GES E IACEAE p n nd n t G r t G h bd th n l ndr A C nn G
Y ACEAE GOO E IACEAE ph r nt l C r l W S ll r r d n C v S App nd x 105
A O AGACEAE O YGO ACEAE Go oca us mic a us h nb I Mue ie eckia aus a is ( r t f M n G G mo a us ( f Or h Κ M com e a (A C nn M n G a o agis e ec a ( n x M rr E hl Ο o ygo um eci ie s r W
AU ACEAE IMU ACEAE ei sc mie ia awa (A C nn nth Samo us e e s ( t G r t r S x f x K r G O EACEAE O E I AC ΕAΕ K ig ia e ce sa r G o e ia a ce s nn f S O
OG A IAC ΕAΕ A U CU ACEAE Ge ios oma u es e r t t r t f C ema is a icu a a G l G v r igus um (C nn C nn G C a u cu us r fl x G rn - n G MA ACEAE AM ACEAE o e ia o u ea A C nn G oma e is kume a o A C nn agia us i a ica us t G r t S (Waika i y ici o ia dd x l n ME IACEAE OSACEAE yso y um s ec a i e ( r t f f G u us cissoi es A C nn G MO IMIACEAE U IACEAE e yca ya a o ea t G r t G Co osma aus a is (A h b n n G MO ACEAE C uci a t G r t G a a o is mic o y a ( l C n G C mac oca a Ch C MYO O ACEAE C am oi es A C nn G Myo o um ae um r t f C C o us a l G Κ Ο
M Y SI ACEAE U ACEAE My si e aus a is (A h All n G I Me ico e sim e A C nn G MY ACEAE Μ e a a t G r t G Ku ea e icoi es (A h h p n G SA I ACEAE e os e mum sco a ium t G r t G A ec yo e ce sus G rtn G o omy us u a a (S l x A C nn rr t G o o aea iscosa acqui G Me osi e os i usa ( r t f S th G M e ce sa S l x G rtn C SA O ACEAE M e ce sa o us a G a c o e a cos a a (Endl rr x C M u ge s S l x G rtn G SC O U A IACEAE . Μ e o a a ( t G r t A h G e e mac oca a ( hI C n t All n C O EACEAE s ic a ( nth M r C es egis a ceo a a ( f hn n G SO A ACEAE ASSI O ACEAE So a um ame ica um M ll r assi o a e a a n t 0 x C G I IACEAE I E AC ΕAΕ E e ea a o esce s r C Mac o i e e ce sum ( r t f M G C
e e omia u i ea a A h C VERBENACEAE . K r I OS O ACEAE t x l n G i os o um c assi o ium n t S l x A C nn C VIOLACEAE
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