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SYSTEMATICS of the MEGADIVERSE SUPERFAMILY GELECHIOIDEA (INSECTA: LEPIDOPTEA) DISSERTATION Presented in Partial Fulfillment of T
SYSTEMATICS OF THE MEGADIVERSE SUPERFAMILY GELECHIOIDEA (INSECTA: LEPIDOPTEA) DISSERTATION Presented in Partial Fulfillment of the Requirements for The Degree of Doctor of Philosophy in the Graduate School of The Ohio State University By Sibyl Rae Bucheli, M.S. ***** The Ohio State University 2005 Dissertation Committee: Approved by Dr. John W. Wenzel, Advisor Dr. Daniel Herms Dr. Hans Klompen _________________________________ Dr. Steven C. Passoa Advisor Graduate Program in Entomology ABSTRACT The phylogenetics, systematics, taxonomy, and biology of Gelechioidea (Insecta: Lepidoptera) are investigated. This superfamily is probably the second largest in all of Lepidoptera, and it remains one of the least well known. Taxonomy of Gelechioidea has been unstable historically, and definitions vary at the family and subfamily levels. In Chapters Two and Three, I review the taxonomy of Gelechioidea and characters that have been important, with attention to what characters or terms were used by different authors. I revise the coding of characters that are already in the literature, and provide new data as well. Chapter Four provides the first phylogenetic analysis of Gelechioidea to include molecular data. I combine novel DNA sequence data from Cytochrome oxidase I and II with morphological matrices for exemplar species. The results challenge current concepts of Gelechioidea, suggesting that traditional morphological characters that have united taxa may not be homologous structures and are in need of further investigation. Resolution of this problem will require more detailed analysis and more thorough characterization of certain lineages. To begin this task, I conduct in Chapter Five an in- depth study of morphological evolution, host-plant selection, and geographical distribution of a medium-sized genus Depressaria Haworth (Depressariinae), larvae of ii which generally feed on plants in the families Asteraceae and Apiaceae. -
Maternal Care in Acanthosomatinae (Insecta: Heteroptera: Acanthosomatidae)̶Correlated Evolution with Title Morphological Change
Maternal care in Acanthosomatinae (Insecta: Heteroptera: Acanthosomatidae)̶correlated evolution with Title morphological change Author(s) Tsai, Jing-Fu; Kudo, Shin-ichi; Yoshizawa, Kazunori BMC Evolutionary Biology, 15, 258 Citation https://doi.org/10.1186/s12862-015-0537-4 Issue Date 2015-11-19 Doc URL http://hdl.handle.net/2115/63251 Rights(URL) http://creativecommons.org/licenses/by/4.0 Type article File Information 10.1186_s12862-015-0537-4.pdf Instructions for use Hokkaido University Collection of Scholarly and Academic Papers : HUSCAP Tsai et al. BMC Evolutionary Biology (2015) 15:258 DOI 10.1186/s12862-015-0537-4 RESEARCH ARTICLE Open Access Maternal care in Acanthosomatinae (Insecta: Heteroptera: Acanthosomatidae)— correlated evolution with morphological change Jing-Fu Tsai1,3*, Shin-ichi Kudo2 and Kazunori Yoshizawa1 Abstract Background: Maternal care (egg-nymph guarding behavior) has been recorded in some genera of Acanthosomatidae. However, the origin of the maternal care in the family has remained unclear due to the lack of phylogenetic hypotheses. Another reproductive mode is found in non-caring species whose females smear their eggs before leaving them. They possess pairs of complex organs on the abdominal venter called Pendergrast’s organ (PO) and spread the secretion of this organ onto each egg with their hind legs, which is supposed to provide a protective function against enemies. Some authors claim that the absence of PO may be associated with the presence of maternal care. No study, however, has tested this hypothesis of a correlated evolution between the two traits. Results: We reconstructed the molecular phylogeny of the subfamily Acanthosomatinae using five genetic markers sequenced from 44 species and one subspecies with and without maternal care. -
Adult Postabdomen, Immature Stages and Biology of Euryommatus Mariae Roger, 1856 (Coleoptera: Curculionidae: Conoderinae), a Legendary Weevil in Europe
insects Article Adult Postabdomen, Immature Stages and Biology of Euryommatus mariae Roger, 1856 (Coleoptera: Curculionidae: Conoderinae), a Legendary Weevil in Europe Rafał Gosik 1,*, Marek Wanat 2 and Marek Bidas 3 1 Department of Zoology and Nature Protection, Institute of Biological Sciences, Maria Curie–Skłodowska University, Akademicka 19, 20-033 Lublin, Poland 2 Museum of Natural History, University of Wrocław, Sienkiewicza 21, 50-335 Wrocław, Poland; [email protected] 3 ul. Prosta 290 D/2, 25-385 Kielce, Poland; [email protected] * Correspondence: [email protected] Simple Summary: Euryommatus mariae is a legendary weevil species in Europe, first described in the 19th century and not collected through the 20th century. Though rediscovered in the 21st century at few localities in Poland, Austria, and Germany, it remains one of the rarest of European weevils, and its biology is unknown. We present the first descriptions of the larva and pupa of E. mariae, and confirm its saproxylic lifestyle. The differences and similarities between immatures of E. mariae and the genera Coryssomerus, Cylindrocopturus and Eulechriopus are discussed, and a list of larval characters common to all Conoderitae is given. The characters of adult postabdomen are described and illustrated for the first time for diagnostic purposes. Our study confirmed the unusual structure of the male endophallus, equipped with an extremely long ejaculatory duct enclosed in a peculiar fibrous conduit, not seen in other weevils. We hypothesize that the extraordinarily long Citation: Gosik, R.; Wanat, M.; Bidas, and spiral spermathecal duct is the female’s evolutionary response to the male’s extremely long M. -
Hymenoptera: Eulophidae) 321-356 ©Entomofauna Ansfelden/Austria; Download Unter
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Entomofauna Jahr/Year: 2007 Band/Volume: 0028 Autor(en)/Author(s): Yefremova Zoya A., Ebrahimi Ebrahim, Yegorenkova Ekaterina Artikel/Article: The Subfamilies Eulophinae, Entedoninae and Tetrastichinae in Iran, with description of new species (Hymenoptera: Eulophidae) 321-356 ©Entomofauna Ansfelden/Austria; download unter www.biologiezentrum.at Entomofauna ZEITSCHRIFT FÜR ENTOMOLOGIE Band 28, Heft 25: 321-356 ISSN 0250-4413 Ansfelden, 30. November 2007 The Subfamilies Eulophinae, Entedoninae and Tetrastichinae in Iran, with description of new species (Hymenoptera: Eulophidae) Zoya YEFREMOVA, Ebrahim EBRAHIMI & Ekaterina YEGORENKOVA Abstract This paper reflects the current degree of research of Eulophidae and their hosts in Iran. A list of the species from Iran belonging to the subfamilies Eulophinae, Entedoninae and Tetrastichinae is presented. In the present work 47 species from 22 genera are recorded from Iran. Two species (Cirrospilus scapus sp. nov. and Aprostocetus persicus sp. nov.) are described as new. A list of 45 host-parasitoid associations in Iran and keys to Iranian species of three genera (Cirrospilus, Diglyphus and Aprostocetus) are included. Zusammenfassung Dieser Artikel zeigt den derzeitigen Untersuchungsstand an eulophiden Wespen und ihrer Wirte im Iran. Eine Liste der für den Iran festgestellten Arten der Unterfamilien Eu- lophinae, Entedoninae und Tetrastichinae wird präsentiert. Mit vorliegender Arbeit werden 47 Arten in 22 Gattungen aus dem Iran nachgewiesen. Zwei neue Arten (Cirrospilus sca- pus sp. nov. und Aprostocetus persicus sp. nov.) werden beschrieben. Eine Liste von 45 Wirts- und Parasitoid-Beziehungen im Iran und ein Schlüssel für 3 Gattungen (Cirro- spilus, Diglyphus und Aprostocetus) sind in der Arbeit enthalten. -
The Curculionoidea of the Maltese Islands (Central Mediterranean) (Coleoptera)
BULLETIN OF THE ENTOMOLOGICAL SOCIETY OF MALTA (2010) Vol. 3 : 55-143 The Curculionoidea of the Maltese Islands (Central Mediterranean) (Coleoptera) David MIFSUD1 & Enzo COLONNELLI2 ABSTRACT. The Curculionoidea of the families Anthribidae, Rhynchitidae, Apionidae, Nanophyidae, Brachyceridae, Curculionidae, Erirhinidae, Raymondionymidae, Dryophthoridae and Scolytidae from the Maltese islands are reviewed. A total of 182 species are included, of which the following 51 species represent new records for this archipelago: Araecerus fasciculatus and Noxius curtirostris in Anthribidae; Protapion interjectum and Taeniapion rufulum in Apionidae; Corimalia centromaculata and C. tamarisci in Nanophyidae; Amaurorhinus bewickianus, A. sp. nr. paganettii, Brachypera fallax, B. lunata, B. zoilus, Ceutorhynchus leprieuri, Charagmus gressorius, Coniatus tamarisci, Coniocleonus pseudobliquus, Conorhynchus brevirostris, Cosmobaris alboseriata, C. scolopacea, Derelomus chamaeropis, Echinodera sp. nr. variegata, Hypera sp. nr. tenuirostris, Hypurus bertrandi, Larinus scolymi, Leptolepurus meridionalis, Limobius mixtus, Lixus brevirostris, L. punctiventris, L. vilis, Naupactus cervinus, Otiorhynchus armatus, O. liguricus, Rhamphus oxyacanthae, Rhinusa antirrhini, R. herbarum, R. moroderi, Sharpia rubida, Sibinia femoralis, Smicronyx albosquamosus, S. brevicornis, S. rufipennis, Stenocarus ruficornis, Styphloderes exsculptus, Trichosirocalus centrimacula, Tychius argentatus, T. bicolor, T. pauperculus and T. pusillus in Curculionidae; Sitophilus zeamais and -
Insecta, Diptera): Proposed Conservation of Usage by Designation of Musca Plebeja Linnaeus, 1758 As the Type Species of Thereva
198 Bulletin of Zoological Nomenclature 60(3) September 2003 Case 3251 Thereva Latreille, 1797 and Phasia Latreille, 1804 (Insecta, Diptera): proposed conservation of usage by designation of Musca plebeja Linnaeus, 1758 as the type species of Thereva Kevin C. Holston Department of Entomology, University of Illinois, Urbana, IL 61801, U.S.A. (e-mail: [email protected]) Michael E. Irwin Department of Natural Resources and Environmental Sciences, University of Illinois, Urbana, IL 61801, U.S.A. (e-mail: [email protected]) F. Christian Thompson Systematic Entomology Laboratory, PSI, BARC, ARS, USDA, do Smithsonian Institution, Washington, D.C. 20560, U.S.A. (e-mail: [email protected]) Abstract. The purpose of this application, in relation to Article 67.2 of the Code, is to conserve the usage of the name Thereva Latreille, 1797 for a cosmopolitan genus of stiletto flies (family THEREVIDAE) that currently includes 201 species, and also the usage of Phasia Latreille, 1804 and PHASIINAE for a group of tachinid flies (family TACHINIDAE), some of which are economically important as parasites of plant bugs (Heteroptera). It is proposed that Musca plebeja Linnaeus, 1758 should be designated as the type species of the therevid genus Thereva. M. plebeja was not one of the nominal species that were first associated with the name Thereva by Fabricius (1798). Fabricius used Thereva for a group of tachinid flies that are now referred to by the name Phasia Latreille, 1804. Keywords. Nomenclature; taxonomy; Diptera; THEREVIDAE; TACHINIDAE; Thereva; Phasia; Thereva plebeja; stiletto flies; tachinids. 1. Latreille (1797, p. 168) described a genus of stiletto fries (now in the family THEREVIDAE) and named it Thereva. -
76 ©Kreis Nürnberger Entomologen; Download Unter
ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Galathea, Berichte des Kreises Nürnberger Entomologen e.V. Jahr/Year: 1997 Band/Volume: 13 Autor(en)/Author(s): Dunk Klaus von der Artikel/Article: Ecological studies on Pipunculidae (Diptera) 61-76 ©Kreis Nürnberger Entomologen; download unter www.biologiezentrum.at galathea 13/2 Berichte des Kreises Nürnberger Entomologen1997 • S. 61 -76 Ecological studies on Pipunculidae (Diptera) K laus von der D unk Zusammenfassung: Es wird über Freilandbeobachtungen an Augenfliegen berich tet. Räumlich begrenzte Vorkommen erwiesen sich als erstaunlich artenreich. Sie werden im einzelnen vorgestellt, sowie eine bemerkenswerte Begleitfauna genannt. Betrachtungen von Verhaltensweisen runden das Bild ab, zeigen aber gleichzeitig die Notwendigkeit für weitere Studien. Abstract: Studies on Pipunculid flies in their natural environment are presented. Certain places are described, which proved to be astonishingly rieh in species. Some remarkable associating insect species are listed. As far as investigated comments on the behaviour of the adult flies are added. Key words: Diptera, Pipunculidae, behaviour, ecology Introduction Pipunculid flies are rather small mostly black insects, developing as parasitoids inside leafhoppers, with the ability of hovering (relationship to Syrphidae) and with enormous compound eyes, useful for males in search for females, and for females in search for a potential victim, a cicad larva. Most specimen of Pipunculidae studied so far were collected by Malaise traps. This material allows to describe the existing species, to secure their systematical stand, and to mark their distribution. Many questions in this chapter are still open. On the other hand the development as parasitoids in leafhoppers show fascinating aspects of adaptations to this life and even has an ecological/economical content regarding pest control. -
Zum Sommeraspekt Der Zikadenfauna Des Kyffhäusergebirges (Hemiptera: Auchenorrhyncha)
Beiträge zur Zikadenkunde 9: 15-22 (2007) 15 A food plant study of the Auchenorrhyncha of the Częstochowa upland, southern Poland (Insecta, Hemiptera) Dariusz Świerczewski1 Kurzfassung: Auf der Hochebene von Tschenstochau in Süd-Polen wurde 2004 bis 2006 die Zikadenfauna von 31 Kräuter- und Grasarten untersucht. Das Material umfasst insgesamt 1970 adulte Individuen aus 44 Arten. Für 21 Pflanzenarten werden Zikadengilden beschrieben und diskutiert. Abstract: Auchenorrhyncha were sampled on 31 species of forbs and grasses in the years 2004-2006 in the area of Częstochowa Upland in southern Poland. Altogether, the material includes 1970 adult individuals belonging to 44 species. Auchenorrhyncha guilds for 21 plants are described and discussed. Key words: Hemiptera, Auchenorrhyncha guilds, food plants, Poland 1. Introduction The knowledge of Auchenorrhyncha food plants in Poland, unlike those published recently from Germany (Nickel 2003) or Britain (e.g. Waloff & Solomon 1973; Cook 1996), are far from complete. Smreczyński (1954) and Nast (1976) provided records mainly for arboricolous taxa, i.e. Macropsinae, Idiocerinae, Jassinae and Typhlocybinae, but there is still no information on the food preferences of the bulk of the leafhopper fauna. The aim of this work was to determine Auchenorrhyncha guilds associated with forbs and grasses, which are notable constituents of grasslands on limestone and sandy substrates of the Częstochowa upland in southern Poland. 2. Study area, methods and material A survey was carried out over a 3-year period (2004-2006) near the villages of Olsztyn and Mstów, UTM CB 72, CB 73 (Fig. 1). This region is characterized by Upper Jurassic limestone formations dissected by valleys filled with Pleistocene sands. -
Insects of Larose Forest (Excluding Lepidoptera and Odonates)
Insects of Larose Forest (Excluding Lepidoptera and Odonates) • Non-native species indicated by an asterisk* • Species in red are new for the region EPHEMEROPTERA Mayflies Baetidae Small Minnow Mayflies Baetidae sp. Small minnow mayfly Caenidae Small Squaregills Caenidae sp. Small squaregill Ephemerellidae Spiny Crawlers Ephemerellidae sp. Spiny crawler Heptageniiidae Flatheaded Mayflies Heptageniidae sp. Flatheaded mayfly Leptophlebiidae Pronggills Leptophlebiidae sp. Pronggill PLECOPTERA Stoneflies Perlodidae Perlodid Stoneflies Perlodid sp. Perlodid stonefly ORTHOPTERA Grasshoppers, Crickets and Katydids Gryllidae Crickets Gryllus pennsylvanicus Field cricket Oecanthus sp. Tree cricket Tettigoniidae Katydids Amblycorypha oblongifolia Angular-winged katydid Conocephalus nigropleurum Black-sided meadow katydid Microcentrum sp. Leaf katydid Scudderia sp. Bush katydid HEMIPTERA True Bugs Acanthosomatidae Parent Bugs Elasmostethus cruciatus Red-crossed stink bug Elasmucha lateralis Parent bug Alydidae Broad-headed Bugs Alydus sp. Broad-headed bug Protenor sp. Broad-headed bug Aphididae Aphids Aphis nerii Oleander aphid* Paraprociphilus tesselatus Woolly alder aphid Cicadidae Cicadas Tibicen sp. Cicada Cicadellidae Leafhoppers Cicadellidae sp. Leafhopper Coelidia olitoria Leafhopper Cuernia striata Leahopper Draeculacephala zeae Leafhopper Graphocephala coccinea Leafhopper Idiodonus kelmcottii Leafhopper Neokolla hieroglyphica Leafhopper 1 Penthimia americana Leafhopper Tylozygus bifidus Leafhopper Cercopidae Spittlebugs Aphrophora cribrata -
ZGRUPOWANIA PIEWIKÓW (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) WYBRANYCH ZBIOROWISK ROŚLINNYCH BABIOGÓRSKIEGO PARKU NARODOWEGO Monografia
ZGRUPOWANIA PIEWIKÓW (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) WYBRANYCH ZBIOROWISK ROŚLINNYCH BABIOGÓRSKIEGO PARKU NARODOWEGO Monografia LEAFHOPPER COMMUNITIES (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) SELECTED PLANT COMMUNITIES OF THE BABIA GÓRA NATIONAL PARK The Monograph ROCZNIK MUZEUM GÓRNOŚLĄSKIEGO W BYTOMIU PRZYRODA NR 21 SEBASTIAN PILARCZYK, MARCIN WALCZAK, JOANNA TRELA, JACEK GORCZYCA ZGRUPOWANIA PIEWIKÓW (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) WYBRANYCH ZBIOROWISK ROŚLINNYCH BABIOGÓRSKIEGO PARKU NARODOWEGO Monografia Bytom 2014 ANNALS OF THE UPPER SILESIAN MUSEUM IN BYTOM NATURAL HISTORY NO. 21 SEBASTIAN PILARCZYK, MARCIN WALCZAK, JOANNA TRELA, JACEK GORCZYCA LEAFHOPPER COMMUNITIES (HEMIPTERA: FULGOROMORPHA ET CICADOMORPHA) SELECTED PLANT COMMUNITIES OF THE BABIA GÓRA NATIONAL PARK The Monograph Bytom 2014 Published by the Upper Silesian Museum in Bytom Upper Silesian Museum in Bytom Plac Jana III Sobieskiego 2 41–902 Bytom, Poland tel./fax +48 32 281 34 01 Editorial Board of Natural History Series: Jacek Betleja, Piotr Cempulik, Roland Dobosz (Head Editor), Katarzyna Kobiela (Layout), Adam Larysz (Layout), Jacek Szwedo, Dagmara Żyła (Layout) International Advisory Board: Levente Ábrahám (Somogy County Museum, Kaposvar, Hungary) Horst Aspöck (University of Vienna, Austria) Dariusz Iwan (Museum and Institute of Zoology PAS, Warszawa, Poland) John Oswald (Texas A&M University, USA) Alexi Popov (National Museum of Natural History, Sofia, Bulgaria) Ryszard Szadziewski (University of Gdańsk, Gdynia, Poland) Marek Wanat (Museum -
Les Tachinidae De La Manche
Les Tachinidae de la Manche : début de l’enquête et première liste (Diptera Brachycera) La famille de mouches que nous allons aborder dans cet article offre le paradoxe d’être à la fois l’une des plus riches en espèces et l’une des plus mal connues des entomologistes. Discrètes, souvent de petite taille et peu colorées, elles passent facilement inaperçues. Totalement ignorées du profane, elles n’ont reçu aucun nom vernaculaire et c’est à peine si les guides de vulgarisation représentent quelques espèces, 6 par exemple dans CHINERY (1986) contre 22 syrphes, famille d’ampleur comparable, soit moins de 1% de la faune européenne ! Un guide spécialisé dans les diptères, J. & H. HAUPT (1998), dont l’éditeur suisse annonce sans vergogne en couverture « l’identification des espèces européennes », illustre 13 espèces, soit 1,5% de la faune d’Europe. Il est certain que l’ampleur de la tâche a pu rebuter les diptéristes de notre pays, d’autant plus qu’aujourd’hui encore aucun ouvrage complet de détermination n’est actuellement disponible en langue française. Et pourtant ces mouches ne manquent pas d’attrait ! De nombreuses espèces sont d’une taille respectable, sous la loupe beaucoup même offrent un aspect séduisant par leurs couleurs veloutées, leurs reflets pruineux, leur système organisé de poils et de soies, toutes surtout ont une biologie complexe, captivante et encore mal étudiée. Pour tout dire, cette enquête limitée à la Manche (voir L’Argiope No 67) s’annonce passionnante car la page est totalement vierge et le champ de recherche immense. Bien sûr, les obstacles sont tels que l’inventaire devra être abordé avec prudence. -
Duffield Scene May 2#1C0C79.Qxd 16/12/2015 12:38 Page 1
Duffield Scene January 2016.qxp_Duffield Scene may 2#1C0C79.qxd 16/12/2015 12:38 Page 1 DUFFIELD SCENE JANUARY 2016 Edition 279 Published Monthly IMPROVEMENTS TO THE VILLAGE ON THE WAY... AND SOON Look out for a number of website has been updated with by improvements to the village in details of local facilities to hire, Jackie Clements 2016 and beyond – that’s the “what’s on” in the village and message from Duffield Parish links to Facebook and Twitter Council and local volunteers. accounts to keep local people In the six months since the up-to-date; Duffield Action Plan was • Improvements to facilities for published, progress has been elderly people, including made behind the scenes, with streamlining communications working groups being formed and outreach work. and meetings taking place to Town Street; • A plan for new recreational The Duffield Action Plan was discuss future improvements to • Sites being sought out to build facilities for young people will published in June 2015 after the public toilets, parking, a new public toilet; be made, with the involvement close consultation with local communication and local • A car parking review and of pupils from Ecclesbourne people. It sets out the priorities recreational facilities, to name a traffic report are to be School; for services and amenities in the few of the Plan’s priorities. undertaken; • Duffield Community village for the next five years. Current and future • Litter bins have been Association to re-publish the The Plan can be viewed on the developments include: purchased for Gray Rec and Village Guide and map; Parish Council website: • Cycle stands to be erected on Hazelwood Road; • Duffield Parish Council www.duffieldparishcouncil.gov.uk Duffield Scene January 2016.qxp_Duffield Scene may 2#1C0C79.qxd 17/12/2015 10:17 Page 2 DUFFIELD All information, letters, SCENE news, articles, diary Established events, photographs and August 1992 suggestions should be marked “Duffield Scene” and sent to: Duffield News, Town Street, Duffield.