ROOSTING HABITS of the IRISH GOAL-TIT, with SOME OBSERVATIONS on OTHER HABITS (Plate 43)
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(326) ROOSTING HABITS OF THE IRISH GOAL-TIT, WITH SOME OBSERVATIONS ON OTHER HABITS BY ROBERT F. RUTTLEDGE. (Plate 43). THE observation in The Handbook, Vol. i, p, 256, stating that data on normal roosting habits of the Coal-Tit were scanty, led me to pay particular attention to this subject. My observations are confined to the Irish Coal-Tit (Parus ater hibemicus). The results are those of over three years' study of a limited number of birds in Co. Ma}'o. The locality where observations were made is fairly exposed, but with considerable shelter in the form of gardens, outhouses, trees and scrubby woods. There are a number of difficulties in the way of detailed study of this species, chief of which is the extreme difficulty of seeing rings on marked birds in thick foliage and on such a restless bird. In summer, in particular, it is very hard to trace birds to their roosts, with the result that there is a paucity of observations at this season. When working alone, or with only one companion, it is extremely difficult to follow up the rapid movements of the birds as they travel through thickets or in high tree-tops prior to going to rcost. Observations have, therefore, at times been rather disjointed, especially when a bird changes its roost, thus necessitating renewed search. Although primarily the work was in connexion with roosting- habits, other habits have claimed attention and some notes on them are given. I am much indebted to John Barlce for the photographs of two typical roost-sites. BEHAVIOUR PRIOR TO ROOSTING. This varies to some extent. Coal-Tits are inclined to become silent about half-an-hour before roosting-time, whereas Great Tits and Blue Tits are inclined to call and scold up to roosting-time. Coal-Tits often go up into the tops of trees or high bushes and there sit silently, perhaps preening. They then suddenly fly off or travel rapidly through the tree-lops or lower bushes for an appreciable distance to arrive at the vicinity of the roost. - Often birds arrive and hang about in the vicinity of the roost either to preen or sit quietly for 7-10 minutes, even 20 minutes or exceptionally 25 minutes, before entering the roost. Very rarely have I seen a bird come from a distance and go straight into the roost. Such instances seem to occur only when a bird is very much behind the normal time of going to roost (e.g. a bird which arrived from a distance, calling and seeming agitated, 33 minutes after sunset, went straight into its roost.). Coal-Tits are normally very suspicious prior to entering their roost. There is individual variation, but on the whole their anxiety is more pronounced than at their VOL. xxxix] ROOSTING HABITS OF IRISH COAL-TIT. 327 nesting-site. Most birds, before entering finally, will either sit just outside the roost for a few moments, or will enter the roost, then re-appear just outside, perhaps several times. If conscious of being watched they delay entering and may even depart to an alternative roost. Such enforced delays must be avoided when timing birds into their roosts. Timings have been taken from the birds' final entry into the roost. Once in the roost individuals vary in their alertness, even in similar forms of roost. Some are very wakeful 20 minutes or even 30 minutes after entry; others are not easily disturbed even a short time after entry. At night they are not very responsive to noise or even to light, but they awake immediately and leave the roost if the tree-trunk or branch, etc., is lightly tapped or jerked. (cf. Dunsheath and Doncaster [1]), ROOSTS. The following have come under my notice. Holes in rotting tree-trunks—usually of ash-trees—-(illustration No. 1). Frequent. These holes are frequently enlarged from Tree-Creepers' scrapings as Messrs. Rankin [2] have pointed out, but are also made by the tits themselves. I have found them to be very uniform in shape, as the following dimensions of three typical ones show. Defth; entrance to back. Diameter at entrance Vertical Horizontal 3 ins. 2 ins. if ins. 3 ins. 2I ins. " 2| ins. 2 ins. 2 ins. 2 ins. The difference in depth of the last one may be due to the roost not having been fully excavated at the time of measurement. Ivy-covered hawthorn trees {Illustration No. 2). Frequent. Ivy on gable of house. Three. Ivy-covered fork of sapling. Three. Ivy-covered fork of tree. Two. Ivy-covered stump. One. Between thick ivy stems around a branch. One. Short ivy around a sapling. One. Garden arch covered with ivy and creepers. One. Crevice between masonry and corrugated-iron roof. Two. Haystack. One. Hole c. 3 ins. deep. Hole in hollow end of snapped-off branch. One. Hole excavated by bird in dry mat of dead ivy on a gable. One. Depth 2| ins. Entrance 21 ins. x z\ ins. On ledge in split of rotting stump. One. Ledge 3 ins. front to rear, 2 ins. broad. 328 BRITISH BIRDS. [VOL. xxxix. Branches of Cupressus macrocarpa. One. Old Wren's nest. One. Thick hedgerow. One. Reference to the use of holes in Sequoias has been made by Rankin [2] and in cedars and old nests of birds of the British race, by Dunsheath and Doncaster [1]. Height of roosts from ground.—The average height of twenty roosts was 9 feet 10 inches. Maximum, 21 feet in one case, in ivy on the gable of a building. Minimum, between two and three feet in two cases, in ivy-covered forks of holly bushes. In addition there were several at c. 15 feet in ivy-covered trees in which it was not possible to obtain completely accurate measure ment. In one instance a marked bird was found to roost in an excavated hole 8 feet from the ground in May and June. In the following November it was roosting at c. 15 feet in an ivy-covered hawthorn tree c, 300 yards away. Aspect.—It seems immaterial what aspect is used. Prevailing winds (S.W. here) and weather conditions do not seem to have any marked effect. I have note of the following aspects. N. 2. S. 2. E. 3. W. 3. N.W. 2. S.W. 2. S.E. 1. TIME OF GOING TO ROOST. From 13 roosts at which I have made observations, the following four are given in detail. They were used fairly consistently (roosts are very frequently changed or deserted, often only temporarily), and they bring out one or two points of interest. As in no case was a marked bird constantly observed there is no proof that the same bird would return after a temporary desertion of a roost, though there was in most cases evidence pointing 'to the probability. Marking a bird at the roost always entails desertion with consequent breaking Of any sequence of observations, and with the added difficulty of finding the new roost. All times given are in relation to local sunset. ROOST I. Date Time Weather Conditions 10-12-42 2 rains, before Fair. 9-4-43 3 mins. before Fair. 13-4-43 5 mins. before Very fine; mild. 22-4-43 16 mins. before Very fine; warm. 23-4-43 14 mins. before Very fine; mild. 19 mins. before 25-4-43 Full gale; rain. This roost was not seen to be in use again until 1945. Impossible to say if the same bird returned, but the roost was in exactly the same place. The difference of time in relation to sunset is, however, strange, though on the September dates sunset is approximately half an hour earlier than on the April dates given. VOL. xxxix] ROOSTING HABITS OF IRISH COAL -TIT. 329 Date Time Weather Conditions 14-9-45 10 mins. after Bright, warm, occasional showers. 15-9-45 Not in use Gloomy, drizzle, strong wind, S.E. 18-9-45 19 mins. after Calm, warm, bright. 20-9-45 16 mins. after Cool, strong wind, S. 10-10-45 33 mins. after Bright, warm, calm. This is an exception ally late record. 12-10-45 Not in use. ROOST II. Date Time Weather Conditions 4-^-43 5 mins. before Fair. 5-1-43 5 mins. before Bright, Fine. 6-1-43 2 mins. before Dull, foggy. 12-1-43 Not in use Dull, foggy. 14-1-43 In roost before sunset. Thick fog. 26-1-43 5 mins. after Bright. 3I-I-43 Already in roost Fine, bright. Great Tits and Blue Tits were 19 mins. before still abroad 30 mins. later. sunset. 2-2-43 Roost finally vacated. The same bird used this roost throughout. Note that on the evenings oi January 26th and 31st, there was considerable variation in time of roosting under similar weather conditions. ROOST III. Date Time Weather Conditions 10-10-45 23 mins. after Calm, bright, cool. 11-10-45 24 mins. after Calm, bright, cool. 12-10-45 23 mins. after Calm, overcast, warm, I3-IO-45 20 mins. after Overcast, rather cool, light breeze, S.E. A bird roosting 4 paces distant entered its roost 12 mins. after sunset on 13th. 16-10-45 Not in use Overcast, cool, light breeze, S.E. 19-10-45 19 mins. after Very gloomy, wind moderate, S.E. 24-10-45 Not in use Gale W. High wind and rain on succeeding days. Roost very sheltered. From. October 28th, weather was fine, calm. 4-i i-45 30J mins.