Yorkshire Coleoptera Checklist 2020

Checklist of Yorkshire Coleoptera

Figures are based on records on the YNU database in February 2020

The following checklist and the Family order in which it is laid out are based on Checklist of of the British Isles, 3rd Edition (2018), edited by A.G. Duff, and published by Pemberley Books.

(To search for a species use Ctrl+F)

Species VC61 VC62 VC63 VC64 VC65 County

2 Gyrinidae Gyrinus aeratus Stephens, 1835 17 0 2 0 0 19 Gyrinus caspius Ménétries, 1832 12 11 2 0 0 25 Gyrinus distinctus Aubé, 1838 14 1 10 3 0 28 Gyrinus marinus Gyllenhal, 1808 95 13 55 20 2 185 Gyrinus minutus Fabricius, 1798 0 2 3 0 0 5 Gyrinus paykulli Ochs, 1927 9 4 14 6 0 33 Gyrinus substriatus Stephens, 1828 157 135 297 130 79 798 Gyrinus urinator Illiger, 1807 35 35 10 10 2 92 Orectochilus villosus (Müller, O.F., 1776) 12 55 6 99 56 228

3 Haliplidae Brychius elevatus (Panzer, 1793) 26 36 17 14 8 101 Peltodytes caesus (Duftschmid, 1805) 0 0 2 0 0 2 Haliplus ruficollis agg. 7 1 22 1 0 31 Haliplus apicalis Thomson, C.G., 1868 25 9 2 3 0 39 Haliplus flavicollis Sturm, 1834 60 28 74 45 33 240 Haliplus fluviatilis Aubé, 1836 67 27 137 24 11 266 Haliplus fulvus (Fabricius, 1801) 12 11 18 10 11 62 Haliplus heydeni Wehncke, 1875 52 16 86 46 2 202 Haliplus immaculatus Gerhardt, 1877 113 31 183 50 11 388 Haliplus laminatus (Schaller, 1783) 1 2 35 10 1 49 Haliplus lineatocollis (Marsham, 1802) 312 146 290 110 56 914 Haliplus lineolatus Mannerheim, 1844 54 18 59 32 18 181 Haliplus mucronatus Stephens, 1828 2 0 2 0 0 4 Haliplus obliquus (Fabricius, 1787) 55 35 41 46 33 210 Haliplus ruficollis (De Geer, 1774) 193 173 399 151 61 977 Haliplus sibiricus Motschulsky, 1860 389 193 359 145 91 1177 Haliplus variegatus Sturm, 1834 1 1 2 0 0 4 Haliplus confinis Stephens, 1828 50 36 83 37 18 224

4 Noteridae Noterus clavicornis (De Geer, 1774) 290 170 452 176 38 1126 Noterus crassicornis (Müller, O.F., 1776) 39 28 35 3 0 105

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Yorkshire Coleoptera Checklist 2020

5 Hygrobiidae hermanni (Fabricius, 1775) 50 27 88 32 14 211

6 Dytiscidae Boreonectes multilineatus (Falkenström, 1922) 0 0 0 8 12 20 Agabus affinis (Paykull, 1798) 26 80 79 35 22 242 Agabus arcticus (Paykull, 1798) 0 0 36 27 45 108 Agabus biguttatus (Olivier, 1795) 1 5 19 4 2 31 Agabus bipustulatus (Linnaeus, 1767) 667 687 1091 589 369 3403 Agabus congener (Thunberg, 1794) 9 21 25 48 63 166 Agabus conspersus (Marsham, 1802) 73 3 3 0 0 79 Agabus didymus (Olivier, 1795) 139 77 133 49 18 416 Agabus guttatus (Paykull, 1798) 15 136 208 126 100 585 Agabus labiatus (Brahm, 1790) 27 2 2 3 2 36 Agabus melanarius Aubé, 1837 2 89 10 1 1 103 Agabus nebulosus (Forster, 1771) 178 117 195 126 48 664 Agabus paludosus (Fabricius, 1801) 113 90 122 70 40 435 Agabus striolatus (Gyllenhal, 1808) 0 0 36 0 0 36 Agabus sturmii (Gyllenhal, 1808) 293 272 577 239 107 1488 Agabus uliginosus (Linnaeus, 1761) 88 54 43 48 4 237 Agabus undulatus (Schrank, 1776) 22 1 0 18 0 41 Agabus unguicularis (Thomson, C.G., 1867) 48 41 31 15 14 149 Ilybius aenescens Thomson, C.G., 1870 10 13 99 17 12 151 Ilybius ater (De Geer, 1774) 117 92 219 75 22 525 Ilybius chalconatus (Panzer, 1796) 47 79 44 61 10 241 Ilybius fenestratus (Fabricius, 1781) 36 21 49 8 1 115 Ilybius fuliginosus (Fabricius, 1792) 264 186 354 178 102 1084 Ilybius guttiger (Gyllenhal, 1808) 19 34 83 8 0 144 Ilybius montanus (Stephens, 1828) 97 144 140 40 62 483 Ilybius quadriguttatus (Lacordaire, 1835) 91 21 160 36 0 308 Ilybius subaeneus Erichson, 1837 9 4 24 6 0 43 Platambus maculatus (Linnaeus, 1758) 51 166 125 145 109 596 Colymbetes fuscus (Linnaeus, 1758) 335 173 388 145 39 1080 Rhantus bistriatus (Bergsträsser, 1778) 0 0 2 0 0 2 Rhantus exsoletus (Forster, 1771) 17 9 95 11 2 134 Rhantus frontalis (Marsham, 1802) 1 0 0 1 0 2 Rhantus grapii (Gyllenhal, 1808) 41 43 171 23 0 278 Rhantus suturalis (MacLeay, 1825) 126 57 152 62 7 404 Rhantus suturellus (Harris, T.W., 1828) 43 43 138 34 36 294 Liopterus haemorrhoidalis (Fabricius, 1787) 107 24 224 45 0 400 Acilius canaliculatus (Nicolai, 1822) 21 0 268 0 0 289 Acilius sulcatus (Linnaeus, 1758) 53 54 146 42 7 302 Dytiscus circumcinctus Ahrens, 1811 9 1 39 10 0 59 Dytiscus circumflexus Fabricius, 1801 36 16 53 11 1 117 Dytiscus dimidiatus Bergsträsser, 1778 1 1 3 11 0 16 Dytiscus marginalis Linnaeus, 1758 108 88 267 76 22 561

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Yorkshire Coleoptera Checklist 2020

Dytiscus semisulcatus Müller, O.F., 1776 11 32 30 7 9 89 Hydaticus seminiger (De Geer, 1774) 0 0 0 4 0 4 Hydaticus transversalis (Pontoppidan, 1763) 0 0 2 7 0 9 Bidessus unistriatus (Goeze, 1777) 0 0 1 0 0 1 Hydroglyphus geminus (Fabricius, 1792) 68 45 126 53 10 302 Deronectes latus (Stephens, 1829) 0 6 2 2 3 13 Graptodytes bilineatus (Sturm, 1835) 11 0 0 0 0 11 Graptodytes granularis (Linnaeus, 1767) 30 22 34 7 0 93 Graptodytes pictus (Fabricius, 1787) 195 42 289 51 0 577 Hydroporus angustatus Sturm, 1835 233 163 326 148 16 886 Hydroporus discretus Fairmaire & Brisout de Barneville, 1859 57 72 50 54 38 271 Hydroporus erythrocephalus (Linnaeus, 1758) 111 90 237 91 46 575 Hydroporus ferrugineus Stephens, 1829 0 14 11 11 5 41 Hydroporus glabriusculus Aubé, 1838 0 0 0 1 0 1 Hydroporus gyllenhalii Schiödte, 1841 157 361 492 308 257 1575 Hydroporus incognitus Sharp, 1869 142 211 376 163 69 961 Hydroporus longicornis Sharp, 1871 0 38 22 18 9 87 Hydroporus longulus Mulsant & Rey, 1861 5 56 21 20 14 116 Hydroporus marginatus (Duftschmid, 1805) 0 0 1 0 0 1 Hydroporus melanarius Sturm, 1835 31 102 147 115 92 487 Hydroporus memnonius Nicolai, 1822 161 244 179 135 82 801 Hydroporus morio Aubé, 1838 0 26 41 90 118 275 Hydroporus neglectus Schaum, 1845 65 66 136 25 2 294 Hydroporus nigrita (Fabricius, 1792) 45 150 107 119 129 550 Hydroporus obscurus Sturm, 1835 9 48 140 34 39 270 Hydroporus obsoletus Aubé, 1838 0 4 1 1 1 7 Hydroporus palustris (Linnaeus, 1761) 380 273 675 253 140 1721 Hydroporus planus (Fabricius, 1782) 584 402 506 285 79 1856 Hydroporus pubescens (Gyllenhal, 1808) 220 382 686 323 276 1887 Hydroporus rufifrons (Müller, O.F., 1776) 0 4 4 8 0 16 Hydroporus scalesianus Stephens, 1828 5 0 4 3 0 12 Hydroporus striola (Gyllenhal, 1826) 42 28 102 41 12 225 Hydroporus tessellatus (Drapiez, 1819) 161 94 137 59 3 454 Hydroporus tristis (Paykull, 1798) 81 175 303 193 168 920 Hydroporus umbrosus (Gyllenhal, 1808) 54 97 82 43 34 310 Nebrioporus depressus agg. 0 1 0 0 0 1 Nebrioporus assimilis (Paykull, 1798) 37 7 47 26 9 126 Nebrioporus depressus (Fabricius, 1775) 0 0 0 1 0 1 Nebrioporus elegans (Panzer, 1794) 145 157 160 126 111 699 Oreodytes davisii (Curtis, 1831) 0 11 18 31 46 106 Oreodytes sanmarkii (Sahlberg, C.R., 1826) 30 181 155 194 166 726 Oreodytes septentrionalis (Gyllenhal, 1826) 0 9 3 38 65 115 Porhydrus lineatus (Fabricius, 1775) 23 4 23 8 0 58 Scarodytes halensis (Fabricius, 1787) 10 0 14 5 0 29 Stictonectes lepidus (Olivier, 1795) 7 5 98 31 1 142

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Yorkshire Coleoptera Checklist 2020

Stictotarsus duodecimpustulatus (Fabricius, 1792) 36 42 50 40 26 194 Suphrodytes dorsalis (Fabricius, 1787) 2 0 1 3 0 6 Suphrodytes dorsalis (Fabricius, 1787) 84 24 36 47 7 198 Suphrodytes figuratus (Gyllenhal, 1826) 5 0 7 0 0 12 Hygrotus confluens (Fabricius, 1787) 52 27 45 21 19 164 Hygrotus decoratus (Gyllenhal, 1810) 25 40 34 12 1 112 Hygrotus impressopunctatus (Schaller, 1783) 279 130 367 149 25 950 Hygrotus inaequalis (Fabricius, 1777) 248 157 368 152 35 960 Hygrotus nigrolineatus (Steven, 1808) 11 0 14 4 8 37 Hygrotus novemlineatus (Stephens, 1829) 0 2 0 0 5 7 Hygrotus parallelogrammus (Ahrens, 1812) 12 0 0 0 0 12 Hygrotus quinquelineatus (Zetterstedt, 1828) 1 0 8 1 0 10 Hygrotus versicolor (Schaller, 1783) 80 5 99 28 10 222 Hyphydrus ovatus (Linnaeus, 1761) 167 60 370 79 18 694 Laccornis oblongus (Stephens, 1835) 12 3 0 9 0 24 Laccophilus hyalinus (De Geer, 1774) 95 21 81 39 6 242 Laccophilus minutus (Linnaeus, 1758) 215 129 345 169 58 916 Laccophilus poecilus Klug, 1834 0 0 6 0 0 6

7 Carabidae Cicindela campestris Linnaeus, 1758 16 94 269 19 19 417 crepitans (Linnaeus, 1758) 0 0 1 0 0 1 Calosoma inquisitor (Linnaeus, 1758) 0 0 1 0 2 3 Carabus arvensis Herbst, 1784 2 50 2 9 3 66 Carabus auratus Linnaeus, 1761 0 0 0 1 0 1 Carabus glabratus Paykull, 1790 0 14 1 0 3 18 Carabus granulatus Linnaeus, 1758 52 6 73 6 0 137 Carabus granulatus subsp. granulatus Linnaeus, 1758 0 0 1 0 0 1 Carabus monilis Fabricius, 1792 4 10 11 13 0 38 Carabus nemoralis Müller, O.F., 1764 16 58 53 38 6 171 Carabus nitens Linnaeus, 1758 2 86 22 9 26 145 Carabus problematicus Herbst, 1786 8 182 144 32 31 397 Carabus violaceus Linnaeus, 1758 10 96 82 34 17 239 Cychrus caraboides (Linnaeus, 1758) 7 36 134 43 14 234 Leistus ferrugineus (Linnaeus, 1758) 43 30 146 24 0 243 Leistus fulvibarbis Dejean, 1826 23 11 98 42 6 180 Leistus rufomarginatus (Duftschmid, 1812) 15 8 49 7 0 79 Leistus spinibarbis (Fabricius, 1775) 15 19 48 5 2 89 Leistus terminatus (Hellwig in Panzer, 1793) 28 122 131 54 8 343 Nebria brevicollis (Fabricius, 1792) 55 113 384 139 36 727 Nebria livida (Linnaeus, 1758) 12 6 0 0 0 18 Nebria rufescens Strøm, 1768 0 22 35 38 17 112 Nebria salina Fairmaire & Laboulbène, 1854 8 94 73 42 18 235 Notiophilus aesthuans Motschulsky, 1864 1 0 0 1 6 8 Notiophilus aquaticus (Linnaeus, 1758) 5 99 47 14 14 179 Notiophilus biguttatus (Fabricius, 1779) 123 105 456 124 41 849

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Yorkshire Coleoptera Checklist 2020

Notiophilus germinyi Fauvel, 1863 3 65 31 22 13 134 Notiophilus palustris (Duftschmid, 1812) 17 15 84 16 2 134 Notiophilus quadripunctatus Dejean, 1826 4 3 5 0 0 12 Notiophilus rufipes Curtis, 1829 2 2 14 3 0 21 Notiophilus substriatus Waterhouse, G.R., 1833 11 4 45 15 5 80 Blethisa multipunctata (Linnaeus, 1758) 12 0 6 6 0 24 Elaphrus cupreus Duftschmid, 1812 37 17 131 42 3 230 Elaphrus lapponicus Gyllenhal, 1810 0 1 0 1 0 2 Elaphrus riparius (Linnaeus, 1758) 66 16 64 39 11 196 Elaphrus uliginosus Fabricius, 1792 1 0 0 0 0 1 Loricera pilicornis (Fabricius, 1775) 97 87 286 102 30 602 Clivina collaris (Herbst, 1784) 7 7 6 19 2 41 Clivina fossor (Linnaeus, 1758) 68 41 103 59 14 285 Dyschirius aeneus (Dejean, 1825) 8 1 13 4 0 26 Dyschirius angustatus (Ahrens, 1830) 1 0 0 0 0 1 Dyschirius globosus (Herbst, 1784) 28 93 104 13 6 244 Dyschirius luedersi Wagner, 1915 18 3 30 2 0 53 Dyschirius nitidus (Dejean, 1825) 3 1 0 0 0 4 Dyschirius politus (Dejean, 1825) 8 1 7 0 1 17 Dyschirius salinus Schaum, 1843 9 0 1 0 0 10 Dyschirius thoracicus (Rossi, 1790) 13 1 0 0 0 14 Broscus cephalotes (Linnaeus, 1758) 17 6 11 0 0 34 Miscodera arctica (Paykull, 1798) 2 59 9 10 7 87 Aepus marinus (Strøm, 1783) 0 7 0 0 0 7 Aepus robinii (Laboulbène, 1849) 0 7 0 0 0 7 Trechus fulvus Dejean, 1831 0 1 0 0 0 1 Trechus obtusus Erichson, 1837 68 169 129 59 15 440 Trechus quadristriatus (Schrank, 1781) 88 74 194 37 14 407 Trechus rivularis (Gyllenhal, 1810) 0 0 0 6 0 6 Trechus rubens (Fabricius, 1792) 0 9 20 2 2 33 Trechus secalis (Paykull, 1790) 26 20 21 26 4 97 Trechus subnotatus Dejean, 1831 0 0 6 0 0 6 Blemus discus (Fabricius, 1792) 14 6 11 11 1 43 Trechoblemus micros (Herbst, 1784) 2 10 6 10 5 33 Tachys bistriatus (Duftschmid, 1812) 1 0 0 0 0 1 Tachys scutellaris Stephens, 1828 0 1 0 0 0 1 Elaphropus parvulus (Dejean, 1831) 0 0 27 0 0 27 Asaphidion flavipes agg. (Linnaeus, 1761) 1 0 7 1 0 9 Asaphidion curtum (Heyden, 1870) 14 6 40 14 0 74 Asaphidion flavipes (Linnaeus, 1761) 10 1 2 10 0 23 Asaphidion pallipes (Duftschmid, 1812) 0 2 0 0 0 2 Asaphidion stierlini (Heyden, 1880) 10 0 16 5 0 31 Ocys harpaloides (Audinet-Serville, 1821) 39 25 107 47 7 225 Ocys tachysoides (Antoine, 1933) 0 0 2 0 0 2 Ocys quinquestriatus (Gyllenhal, 1810) 1 1 11 8 0 21 Bracteon litorale (Olivier, 1790) 2 10 1 13 4 30

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Yorkshire Coleoptera Checklist 2020

Bembidion aeneum Germar, 1824 95 29 155 49 5 333 Bembidion articulatum (Panzer, 1795) 11 1 15 1 0 28 Bembidion assimile Gyllenhal, 1810 59 8 159 1 1 228 Bembidion atrocaeruleum (Stephens, 1828) 3 20 1 36 27 87 Bembidion biguttatum (Fabricius, 1779) 99 29 145 49 2 324 Bembidion bipunctatum (Linnaeus, 1761) 1 1 1 2 1 6 Bembidion bruxellense Wesmael, 1835 4 13 35 10 1 63 Bembidion clarkii (Dawson, 1849) 24 13 56 21 0 114 Bembidion cruciatum Dejean, 1831 14 8 3 0 2 27 Bembidion cruciatum subsp. bualei Jacquelin du Val, 1852 2 0 2 0 0 4 Bembidion decorum (Zenker in Panzer, 1800) 0 17 2 27 11 57 Bembidion deletum Audinet-Serville, 1821 15 9 74 22 4 124 Bembidion dentellum (Thunberg, 1787) 44 6 100 39 6 195 Bembidion doris (Panzer, 1796) 19 3 28 21 0 71 Bembidion femoratum Sturm, 1825 6 6 24 11 2 49 Bembidion fluviatile Dejean, 1831 0 0 0 3 0 3 Bembidion fumigatum (Duftschmid, 1812) 31 1 66 2 0 100 Bembidion geniculatum Heer, 1837/8 6 7 41 12 6 72 Bembidion gilvipes Sturm, 1825 50 22 26 17 0 115 Bembidion guttula (Fabricius, 1792) 87 54 294 82 12 529 Bembidion humerale Sturm, 1825 0 0 46 0 0 46 Bembidion illigeri Netolitzky, 1914 45 9 43 10 2 109 Bembidion iricolor Bedel, 1879 31 0 9 0 0 40 Bembidion lampros (Herbst, 1784) 88 72 342 94 18 614 Bembidion lunatum (Duftschmid, 1812) 13 3 13 6 0 35 Bembidion lunulatum (Geoffroy in Fourcroy, 1785) 34 4 114 25 0 177 Bembidion mannerheimii Sahlberg, C.R., 1827 9 67 30 19 5 130 Bembidion maritimum (Stephens, 1835) 7 2 4 0 0 13 Bembidion minimum (Fabricius, 1792) 62 0 4 1 0 67 Bembidion monticola Sturm, 1825 0 4 2 9 4 19 Bembidion nigricorne Gyllenhal, 1827 2 60 3 5 0 70 Bembidion normannum Dejean, 1831 6 0 0 0 0 6 Bembidion obliquum Sturm, 1825 10 7 92 16 5 130 Bembidion obtusum Audinet-Serville, 1821 61 25 103 32 1 222 Bembidion prasinum (Duftschmid, 1812) 0 3 2 15 10 30 Bembidion properans (Stephens, 1828) 24 30 81 16 2 153 Bembidion punctulatum Drapiez, 1821 0 15 3 26 10 54 Bembidion quadrimaculatum (Linnaeus, 1761) 47 16 85 35 5 188 Bembidion quadripustulatum Audinet-Serville, 1821 0 0 1 0 0 1 Bembidion saxatile Gyllenhal, 1827 2 10 3 1 1 17 Bembidion schuppelii Dejean, 1831 0 0 1 1 0 2 Bembidion stephensii Crotch, 1866 11 8 12 5 2 38 Bembidion stomoides Dejean, 1831 1 2 0 14 4 21 Bembidion testaceum (Duftschmid, 1812) 0 1 0 2 2 5 Bembidion tetracolum Say, 1825 47 42 146 75 33 343 Bembidion tibiale (Duftschmid, 1812) 0 24 14 36 19 93

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Yorkshire Coleoptera Checklist 2020

Bembidion varium (Olivier, 1795) 49 1 31 1 1 83 Pogonus chalceus (Marsham, 1802) 11 0 0 0 0 11 Pogonus littoralis (Duftschmid, 1812) 2 0 0 0 0 2 Patrobus assimilis Chaudoir, 1844 1 56 4 25 13 99 Patrobus atrorufus (Strøm, 1768) 12 14 42 50 11 129 Patrobus septentrionis Dejean, 1828 2 0 2 0 0 4 Stomis pumicatus (Panzer, 1795) 27 14 58 25 3 127 Poecilus cupreus (Linnaeus, 1758) 19 24 89 6 1 139 Poecilus lepidus (Leske, 1785) 3 22 0 0 0 25 Poecilus versicolor (Sturm, 1824) 56 42 122 10 1 231 Pterostichus nigrita agg. (Paykull, 1790) 17 16 80 34 11 158 Pterostichus adstrictus Eschscholtz, 1823 3 80 18 22 12 135 Pterostichus aethiops (Panzer, 1796) 0 3 1 9 4 17 Pterostichus anthracinus (Panzer, 1795) 15 0 9 0 0 24 Pterostichus cristatus (Dufour, 1820) 1 3 2 9 10 25 Pterostichus diligens (Sturm, 1824) 44 105 508 109 32 798 Pterostichus gracilis (Dejean, 1828) 7 0 18 6 0 31 Pterostichus longicollis (Duftschmid, 1812) 4 0 1 2 0 7 Pterostichus macer (Marsham, 1802) 13 10 7 5 1 36 Pterostichus madidus (Fabricius, 1775) 54 118 557 173 51 953 Pterostichus melanarius (Illiger, 1798) 82 24 167 86 15 374 Pterostichus minor (Gyllenhal, 1827) 38 29 201 33 2 303 Pterostichus niger (Schaller, 1783) 94 47 424 79 25 669 Pterostichus nigrita (Paykull, 1790) 48 76 595 68 5 792 Pterostichus oblongopunctatus (Fabricius, 1787) 16 25 52 4 0 97 Pterostichus quadrifoveolatus Letzner, 1852 8 2 56 11 0 77 Pterostichus rhaeticus Heer, 1837/8 8 18 91 26 30 173 Pterostichus strenuus (Panzer, 1796) 164 50 426 142 19 801 Pterostichus vernalis (Panzer, 1795) 33 10 73 27 0 143 Abax parallelepipedus (Piller & Mitterpacher, 1783) 25 25 157 76 13 296 Platyderus depressus (Audinet-Serville, 1821) 8 1 4 0 0 13 Synuchus vivalis (Illiger, 1798) 8 6 30 15 4 63 Calathus melanocephalus agg. (Linnaeus, 1758) 13 16 32 21 13 95 Calathus ambiguus (Paykull, 1790) 4 5 6 0 0 15 Calathus cinctus Motschulsky, 1850 1 2 1 0 0 4 Calathus erratus (Sahlberg, C.R., 1827) 4 82 8 2 0 96 Calathus fuscipes (Goeze, 1777) 26 24 89 24 13 176 Calathus melanocephalus (Linnaeus, 1758) 32 125 82 13 11 263 Calathus micropterus (Duftschmid, 1812) 0 55 6 17 17 95 Calathus mollis (Marsham, 1802) 26 12 1 0 0 39 Calathus rotundicollis Dejean, 1828 26 5 45 16 0 92 Sphodrus leucophthalmus (Linnaeus, 1758) 0 2 3 0 0 5 Laemostenus complanatus (Dejean, 1828) 1 1 1 1 0 4 Laemostenus terricola (Herbst, 1784) 8 9 10 3 1 31 Olisthopus rotundatus (Paykull, 1790) 4 128 59 14 5 210 Oxypselaphus obscurus (Herbst, 1784) 55 10 289 54 4 412

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Yorkshire Coleoptera Checklist 2020

Paranchus albipes (Fabricius, 1796) 37 46 214 92 33 422 Batenus livens (Gyllenhal, 1810) 7 0 19 0 0 26 Platynus assimilis (Paykull, 1790) 14 34 180 49 11 288 dorsalis (Pontoppidan, 1763) 57 19 105 31 1 213 Sericoda quadripunctata (De Geer, 1774) 0 1 0 0 0 1 emarginatum (Gyllenhal, 1827) 35 17 67 14 3 136 Agonum ericeti (Panzer, 1809) 0 10 49 8 2 69 Agonum fuliginosum (Panzer, 1809) 105 64 460 88 32 749 Agonum gracile Sturm, 1824 19 12 151 25 7 214 Agonum gracilipes (Duftschmid, 1812) 2 0 0 0 0 2 Agonum marginatum (Linnaeus, 1758) 41 8 108 18 11 186 Agonum micans Nicolai, 1822 15 3 27 11 1 57 Agonum moestum (Duftschmid, 1812) 1 0 4 2 0 7 Agonum muelleri (Herbst, 1784) 17 17 74 35 7 150 Agonum nigrum Dejean, 1828 0 0 1 0 0 1 Agonum piceum (Linnaeus, 1758) 13 5 29 5 0 52 Agonum sexpunctatum (Linnaeus, 1758) 0 0 3 0 0 3 Agonum thoreyi Dejean, 1828 47 8 142 26 4 227 Agonum versutum Sturm, 1824 10 0 2 0 0 12 Agonum viduum (Panzer, 1796) 6 0 11 7 0 24 Amara aenea (De Geer, 1774) 34 44 271 23 9 381 Amara anthobia Villa & Villa, 1833 0 0 2 0 0 2 Amara apricaria (Paykull, 1790) 13 10 52 9 9 93 Amara bifrons (Gyllenhal, 1810) 9 2 12 8 1 32 Amara communis (Panzer, 1797) 17 11 61 19 4 112 Amara consularis (Duftschmid, 1812) 4 2 2 2 0 10 Amara convexior Stephens, 1828 1 3 4 1 0 9 Amara curta Dejean, 1828 0 3 1 0 0 4 Amara equestris (Duftschmid, 1812) 0 1 0 0 0 1 Amara eurynota (Panzer, 1796) 7 6 26 3 1 43 Amara famelica Zimmermann, 1832 1 3 0 0 0 4 Amara familiaris (Duftschmid, 1812) 52 34 186 46 7 325 Amara fulva (Müller, O.F., 1776) 11 0 14 0 2 27 Amara fusca Dejean, 1828 0 0 1 0 0 1 Amara lucida (Duftschmid, 1812) 7 1 2 1 0 11 Amara lunicollis Schiødte, 1837 9 17 103 17 1 147 Amara ovata (Fabricius, 1792) 12 7 28 12 2 61 Amara plebeja (Gyllenhal, 1810) 48 47 210 57 6 368 Amara praetermissa (Sahlberg, C.R., 1827) 2 2 10 0 1 15 Amara similata (Gyllenhal, 1810) 29 12 102 24 7 174 Amara spreta Dejean, 1831 0 6 0 0 0 6 Amara tibialis (Paykull, 1798) 15 7 30 1 0 53 Curtonotus aulicus (Panzer, 1796) 40 23 105 34 5 207 Curtonotus convexiusculus (Marsham, 1802) 30 4 18 7 0 59 Harpalus affinis (Schrank, 1781) 37 17 95 22 8 179 Harpalus anxius (Duftschmid, 1812) 1 0 0 0 0 1

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Yorkshire Coleoptera Checklist 2020

Harpalus attenuatus Stephens, 1828 1 0 0 0 0 1 Harpalus calceatus (Duftschmid, 1812) 1 0 0 0 0 1 Harpalus froelichii Sturm, 1818 2 0 0 0 0 2 Harpalus latus (Linnaeus, 1758) 9 15 17 6 3 50 Harpalus neglectus Audinet-Serville, 1821 1 0 0 0 0 1 Harpalus rubripes (Duftschmid, 1812) 5 2 29 4 0 40 Harpalus rufipalpis Sturm, 1818 0 0 12 1 0 13 Harpalus rufipes (De Geer, 1774) 62 38 142 30 9 281 Harpalus servus (Duftschmid, 1812) 1 0 1 0 0 2 Harpalus smaragdinus (Duftschmid, 1812) 0 1 0 0 0 1 Harpalus tardus (Panzer, 1796) 32 1 10 1 0 44 Ophonus ardosiacus (Lutshnik, 1922) 5 1 0 0 0 6 Ophonus puncticeps Stephens, 1828 3 0 4 0 0 7 Ophonus puncticollis (Paykull, 1798) 2 0 2 0 0 4 Ophonus rufibarbis (Fabricius, 1792) 40 12 54 10 5 121 Ophonus rupicola (Sturm, 1818) 1 0 0 0 0 1 Ophonus schaubergerianus (Puel, 1937) 3 3 2 1 0 9 Anisodactylus binotatus (Fabricius, 1787) 4 3 95 4 0 106 Dicheirotrichus gustavii Crotch, 1871 16 0 1 0 0 17 Dicheirotrichus obsoletus (Dejean, 1829) 1 0 0 0 0 1 Trichocellus Ganglbauer, 1892 0 0 1 0 0 1 Trichocellus cognatus (Gyllenhal, 1827) 6 145 50 16 7 224 Trichocellus placidus (Gyllenhal, 1827) 48 25 193 67 4 337 Bradycellus caucasicus (Chaudoir, 1846) 2 17 19 10 7 55 Bradycellus distinctus (Dejean, 1829) 0 2 0 0 0 2 Bradycellus harpalinus (Audinet-Serville, 1821) 17 44 194 38 4 297 Bradycellus ruficollis (Stephens, 1828) 4 157 88 15 9 273 Bradycellus sharpi Joy, 1912 2 6 11 7 0 26 Bradycellus verbasci (Duftschmid, 1812) 18 6 73 12 5 114 Stenolophus mixtus (Herbst, 1784) 35 7 157 26 1 226 Acupalpus brunnipes (Sturm, 1825) 0 0 1 0 0 1 Acupalpus dubius Schilsky, 1888 28 24 292 44 1 389 Acupalpus exiguus Dejean, 1829 2 0 7 0 0 9 Acupalpus meridianus (Linnaeus, 1761) 5 0 13 2 0 20 Acupalpus parvulus (Sturm, 1825) 31 3 77 10 0 121 Anthracus consputus (Duftschmid, 1812) 9 1 11 4 0 25 Chlaenius nigricornis (Fabricius, 1787) 15 2 2 3 1 23 Chlaenius tristis (Schaller, 1783) 1 0 0 0 0 1 Chlaenius vestitus (Paykull, 1790) 19 1 5 0 1 26 Oodes helopioides (Fabricius, 1792) 1 1 0 1 0 3 Licinus depressus (Paykull, 1790) 9 1 1 0 0 11 Licinus punctatulus (Fabricius, 1792) 0 1 0 0 0 1 Badister bullatus (Schrank, 1798) 49 16 126 29 1 221 Badister sodalis (Duftschmid, 1812) 4 3 29 4 0 40 Badister unipustulatus Bonelli, 1813 4 0 15 0 0 19 Panagaeus bipustulatus (Fabricius, 1775) 13 0 0 0 0 13

9

Yorkshire Coleoptera Checklist 2020

Panagaeus cruxmajor (Linnaeus, 1758) 1 0 7 0 0 8 Perigona nigriceps (Dejean, 1831) 1 0 0 2 0 3 Lebia chlorocephala (Hoffmann, J., 1803) 3 7 2 0 0 12 Lebia cyanocephala (Linnaeus, 1758) 0 0 0 1 0 1 Demetrias atricapillus (Linnaeus, 1758) 95 22 161 25 1 304 Demetrias monostigma Samouelle, 1819 8 1 0 0 0 9 Cymindis vaporariorum (Linnaeus, 1758) 0 11 3 0 0 14 Paradromius linearis (Olivier, 1795) 136 43 287 53 7 526 Paradromius longiceps (Dejean, 1826) 8 0 8 0 0 16 Dromius agilis (Fabricius, 1787) 2 4 14 11 0 31 Dromius angustus Brullé, 1834 1 7 2 2 0 12 Dromius meridionalis Dejean, 1825 13 5 8 8 0 34 Dromius quadrimaculatus (Linnaeus, 1758) 10 21 56 26 1 114 Calodromius spilotus (Illiger, 1798) 18 13 35 11 1 78 Philorhizus melanocephalus (Dejean, 1825) 78 13 153 28 0 272 Philorhizus notatus (Stephens, 1827) 12 4 0 1 0 17 Philorhizus quadrisignatus (Dejean, 1825) 0 0 1 0 0 1 Philorhizus sigma (Rossi, 1790) 0 1 7 8 0 16 Microlestes maurus (Sturm, 1827) 19 0 3 0 0 22 Lionychus quadrillum (Duftschmid, 1812) 0 1 0 0 0 1 Syntomus foveatus (Geoffroy in Fourcroy, 1785) 24 8 20 1 0 53 Syntomus obscuroguttatus (Duftschmid, 1812) 0 0 2 0 0 2 Syntomus truncatellus (Linnaeus, 1761) 8 1 15 1 0 25 Odacantha melanura (Linnaeus, 1767) 0 0 1 0 0 1

12 Hydrophilidae aequalis Thomson, C.G., 1868 222 271 333 221 107 1154 Helophorus alternans Gené, 1836 0 0 1 1 0 2 Helophorus arvernicus Mulsant, 1846 7 27 10 15 29 88 Helophorus brevipalpis Bedel, 1881 349 365 598 331 178 1821 Helophorus dorsalis (Marsham, 1802) 11 25 2 3 2 43 Helophorus flavipes Fabricius, 1792 26 312 262 224 200 1024 Helophorus fulgidicollis Motschulsky, 1860 40 2 0 0 0 42 Helophorus grandis Illiger, 1798 362 306 336 270 100 1374 Helophorus granularis (Linnaeus, 1761) 6 4 5 8 3 26 Helophorus griseus Herbst, 1793 175 73 93 50 23 414 Helophorus longitarsis Wollaston, 1864 2 2 1 0 0 5 Helophorus minutus Fabricius, 1775 486 254 337 201 51 1329 Helophorus nanus Sturm, 1836 48 13 21 8 3 93 Helophorus nubilus Fabricius, 1777 5 3 8 9 0 25 Helophorus obscurus Mulsant, 1844 260 201 184 150 51 846 Helophorus porculus Bedel, 1881 1 4 0 2 0 7 Helophorus rufipes (Bosc d'Antic, 1791) 1 0 0 0 0 1 Helophorus strigifrons Thomson, C.G., 1868 19 11 33 25 0 88 Helophorus tuberculatus Gyllenhal, 1808 2 10 1 3 0 16 Georissus crenulatus (Rossi, 1794) 18 8 0 1 0 27

10

Yorkshire Coleoptera Checklist 2020

Hydrochus angustatus Germar, 1824 1 0 0 1 0 2 brevis (Herbst, 1793) 8 0 2 1 0 11 Hydrochus crenatus (Fabricius, 1792) 0 0 1 0 0 1 Hydrochus elongatus (Schaller, 1783) 3 1 55 3 0 62 emarginatus (Schaller, 1783) 0 1 0 1 0 2 Anacaena limbata agg. (Fabricius, 1792) 9 1 12 3 1 26 Anacaena bipustulata (Marsham, 1802) 45 4 18 5 1 73 Anacaena globulus (Paykull, 1798) 384 627 1064 555 397 3027 Anacaena limbata (Fabricius, 1792) 506 191 606 233 35 1571 Anacaena lutescens (Stephens, 1829) 188 183 632 170 47 1220 Paracymus scutellaris (Rosenhauer, 1856) 1 3 0 0 0 4 Berosus affinis Brullé, 1835 3 1 4 1 0 9 Berosus fulvus Kuwert, 1888 0 1 0 0 0 1 Berosus luridus (Linnaeus, 1761) 3 0 9 0 0 12 Berosus signaticollis (Charpentier, 1825) 5 1 21 2 0 29 Chaetarthria seminulum (Herbst, 1797) 28 25 24 12 1 90 Chaetarthria seminulum (Herbst, 1797) 0 0 3 0 0 3 Chaetarthria simillima Vorst & Cuppen, 2003 3 0 1 0 0 4 Cymbiodyta marginellus (Fabricius, 1792) 188 55 148 84 2 477 Enochrus affinis (Thunberg, 1794) 66 66 215 13 0 360 Enochrus bicolor (Fabricius, 1792) 34 1 14 0 0 49 Enochrus coarctatus (Gredler, 1863) 32 61 106 30 8 237 Enochrus fuscipennis (Thomson, C.G., 1884) 2 6 1 5 2 16 Enochrus halophilus (Bedel, 1878) 30 0 1 1 0 32 Enochrus melanocephalus (Olivier, 1792) 17 9 30 11 3 70 Enochrus ochropterus (Marsham, 1802) 30 34 7 16 6 93 Enochrus quadripunctatus (Herbst, 1797) 3 2 0 6 0 11 Enochrus testaceus (Fabricius, 1801) 131 56 183 47 1 418 Helochares lividus (Forster, 1771) 118 25 157 42 3 345 Helochares obscurus (Müller, O.F., 1776) 3 1 7 0 0 11 Helochares punctatus Sharp, 1869 66 95 246 28 0 435 Hydrobius fuscipes (Linnaeus, 1758) 664 527 880 419 207 2697 Hydrochara caraboides (Linnaeus, 1758) 0 0 2 2 0 4 Hydrophilus piceus (Linnaeus, 1758) 0 0 3 0 0 3 Laccobius atrocephalus Reitter, 1872 2 0 0 0 0 2 Laccobius atratus Rottenberg, 1874 1 30 0 13 8 52 Laccobius bipunctatus (Fabricius, 1775) 305 212 406 179 86 1188 Laccobius colon (Stephens, 1829) 90 37 105 35 6 273 Laccobius minutus (Linnaeus, 1758) 92 25 96 40 25 278 Laccobius sinuatus Motschulsky, 1849 61 24 74 15 4 178 Laccobius striatulus (Fabricius, 1801) 25 39 13 51 32 160 Laccobius ytenensis Sharp, 1910 0 0 1 0 0 1 Coelostoma orbiculare (Fabricius, 1775) 51 58 59 38 26 232 Cercyon analis (Paykull, 1798) 29 5 76 9 0 119 Cercyon bifenestratus Küster, 1851 0 0 2 0 0 2 Cercyon convexiusculus Stephens, 1829 171 54 197 71 16 509

11

Yorkshire Coleoptera Checklist 2020

Cercyon depressus Stephens, 1829 4 0 1 0 0 5 Cercyon granarius Erichson, 1837 2 1 0 0 0 3 Cercyon haemorrhoidalis (Fabricius, 1775) 37 15 106 22 15 195 Cercyon impressus (Sturm, 1807) 21 14 116 26 13 190 Cercyon laminatus Sharp, 1873 0 0 1 0 0 1 Cercyon lateralis (Marsham, 1802) 33 27 113 22 15 210 Cercyon littoralis (Gyllenhal, 1808) 18 7 0 0 0 25 Cercyon marinus Thomson, C.G., 1853 48 7 46 8 4 113 Cercyon melanocephalus (Linnaeus, 1758) 37 19 188 38 15 297 Cercyon nigriceps (Marsham, 1802) 3 4 4 2 0 13 Cercyon obsoletus (Gyllenhal, 1808) 4 3 17 2 0 26 Cercyon pygmaeus (Illiger, 1801) 17 8 38 8 3 74 Cercyon quisquilius (Linnaeus, 1761) 7 5 26 3 12 53 Cercyon sternalis Sharp, 1918 0 0 0 1 0 1 Cercyon terminatus (Marsham, 1802) 11 3 11 2 1 28 Cercyon tristis (Illiger, 1801) 97 40 73 56 8 274 Cercyon unipunctatus (Linnaeus, 1758) 17 11 33 7 7 75 Cercyon ustulatus (Preyssler, 1790) 44 19 74 22 7 166 Megasternum concinnum (Marsham, 1802) 111 38 321 95 40 605 Cryptopleurum crenatum (Kugelann, 1794) 0 0 0 3 0 3 Cryptopleurum minutum (Fabricius, 1775) 35 14 131 18 8 206 Cryptopleurum subtile Sharp, 1884 3 1 1 1 0 6 Sphaeridium bipustulatum Fabricius, 1781 8 5 20 10 1 44 Sphaeridium lunatum Fabricius, 1792 21 15 41 5 2 84 Sphaeridium marginatum Fabricius, 1787 3 0 4 0 0 7 Sphaeridium scarabaeoides (Linnaeus, 1758) 12 6 50 10 7 85 Enochrus quadripunctatus agg. (Herbst, 1797) 2 0 3 1 0 6

13 Sphaeritidae Sphaerites glabratus (Fabricius, 1792) 0 4 0 1 0 5

14 Histeridae Abraeus granulum Erichson, 1839 0 0 2 0 0 2 Abraeus perpusillus (Marsham, 1802) 3 9 8 1 1 22 Plegaderus vulneratus (Panzer, 1797) 0 0 3 1 0 4 Acritus nigricornis (Hoffmann, J., 1803) 4 3 7 1 0 15 Aeletes atomarius (Aubé, 1842) 0 1 1 0 0 2 Saprinus aeneus (Fabricius, 1775) 5 2 5 0 0 12 Saprinus planiusculus Motschulsky, 1849 1 0 0 0 0 1 Saprinus semistriatus (Scriba, 1790) 10 9 19 5 1 44 Saprinus virescens (Paykull, 1798) 1 0 0 2 0 3 Hypocaccus dimidiatus (Illiger, 1807) 2 2 0 0 0 4 Hypocaccus rugiceps (Duftschmid, 1805) 2 0 0 0 0 2 Hypocaccus rugifrons (Paykull, 1798) 0 1 1 0 0 2 Gnathoncus buyssoni Auzat, 1917 0 0 1 0 0 1 Gnathoncus nannetensis (Marseul, 1862) 3 5 2 2 0 12

12

Yorkshire Coleoptera Checklist 2020

Gnathoncus rotundatus (Kugelann, 1792) 1 0 0 2 0 3 Myrmetes paykulli Kanaar, 1979 0 5 0 0 0 5 Dendrophilus punctatus (Herbst, 1792) 0 1 0 0 0 1 Dendrophilus pygmaeus (Linnaeus, 1758) 0 0 2 0 0 2 Kissister minimus (Aubé, 1850) 0 0 1 0 0 1 Carcinops pumilio (Erichson, 1834) 9 2 16 4 1 32 Paromalus flavicornis (Herbst, 1792) 0 4 11 3 0 18 Paromalus parallelepipedus (Herbst, 1792) 0 0 2 0 0 2 Onthophilus punctatus (Müller, O.F., 1776) 0 0 1 0 0 1 Onthophilus striatus (Forster, 1771) 18 11 75 11 1 116 Margarinotus brunneus (Fabricius, 1775) 14 6 27 9 1 57 Margarinotus marginatus (Erichson, 1834) 4 2 1 0 0 7 Margarinotus merdarius (Hoffmann, J., 1803) 3 0 17 1 0 21 Margarinotus neglectus (Germar, 1813) 0 1 0 1 0 2 Margarinotus purpurascens (Herbst, 1792) 8 1 4 1 0 14 Margarinotus striola (Sahlberg, C.R., 1819) 2 9 18 11 5 45 Margarinotus ventralis (Marseul, 1854) 9 6 28 7 2 52 Hister unicolor Linnaeus, 1758 8 5 9 0 2 24 Atholus bimaculatus (Linnaeus, 1758) 3 1 9 2 0 15 Atholus duodecimstriatus (Schrank, 1781) 4 5 19 4 1 33

15 Hydraena britteni Joy, 1907 27 27 29 28 9 120 Hydraena flavipes Sturm, 1836 0 2 2 1 0 5 Hydraena gracilis Germar, 1824 16 171 156 192 166 701 Hydraena nigrita Germar, 1824 5 47 27 24 18 121 Hydraena palustris Erichson, 1837 11 1 3 2 1 18 Hydraena pulchella Germar, 1824 0 2 2 2 0 6 Hydraena pygmaea Waterhouse, G.R., 1833 0 4 2 4 1 11 Hydraena riparia Kugelann, 1794 146 104 116 81 8 455 Hydraena rufipes Curtis, 1830 0 5 0 15 16 36 Hydraena testacea Curtis, 1830 28 10 53 6 2 99 Limnebius aluta Bedel, 1881 1 2 1 3 0 7 Limnebius nitidus (Marsham, 1802) 22 5 51 13 0 91 Limnebius papposus Mulsant, 1844 1 0 8 1 0 10 Limnebius truncatellus (Thunberg, 1794) 20 183 132 94 139 568 Enicocerus exsculptus (Germar, 1824) 3 6 2 13 18 42 Ochthebius auriculatus Rey, 1885 15 0 0 0 0 15 Ochthebius bicolon Germar, 1824 22 14 19 15 3 73 Ochthebius dilatatus Stephens, 1829 116 18 35 8 0 177 Ochthebius marinus (Paykull, 1798) 47 2 0 0 0 49 Ochthebius minimus (Fabricius, 1792) 407 150 302 181 43 1083 Ochthebius nanus Stephens, 1829 0 0 1 0 0 1 Ochthebius viridis Peyron, 1858 2 0 0 0 0 2

16 Ptiliidae

13

Yorkshire Coleoptera Checklist 2020

Nossidium pilosellum (Marsham, 1802) 2 0 0 0 0 2 Ptenidium formicetorum Kraatz, 1851 0 0 2 0 0 2 Ptenidium fuscicorne Erichson, 1845 18 7 31 4 1 61 Ptenidium gressneri Erichson, 1845 0 0 0 1 0 1 Ptenidium intermedium Wankowicz, 1869 6 1 9 5 1 22 Ptenidium laevigatum Erichson, 1845 8 8 23 9 1 49 Ptenidium nitidum (Heer, 1841) 22 24 132 26 15 219 Ptenidium punctatum (Gyllenhal, 1827) 4 5 7 0 2 18 Ptenidium pusillum (Gyllenhal, 1808) 30 5 113 24 8 180 Ptenidium turgidum Thomson, C.G., 1855 0 2 0 1 0 3 Euryptilium gillmeisteri Flach, 1889 0 0 0 1 0 1 Euryptilium saxonicum (Gillmeister, 1845) 0 0 2 0 0 2 Ptiliola kunzei (Heer, 1841) 1 3 11 3 0 18 Ptiliolum fuscum (Erichson, 1845) 5 15 21 9 4 54 Ptiliolum marginatum (Aubé, 1850) 0 0 1 0 0 1 Ptiliolum sahlbergi (Flach, 1888) 0 0 0 1 0 1 Ptiliolum schwarzi (Flach, 1887) 0 0 3 0 0 3 Ptiliolum spencei (Allibert, 1844) 0 4 18 4 3 29 Actidium aterrimum (Motschulsky, 1845) 0 0 0 1 0 1 Oligella foveolata (Allibert, 1844) 1 0 10 2 0 13 Oligella intermedia Besuchet, 1971 0 0 1 0 0 1 Ptilium horioni Rosskothen, 1934 0 0 4 0 0 4 Ptilium myrmecophilum (Allibert, 1844) 0 5 5 0 0 10 Millidium minutissimum (Ljungh, 1804) 0 1 3 0 1 5 Ptinella aptera (Guérin-Méneville, 1839) 0 1 2 0 0 3 Ptinella britannica Matthews, A., 1858 0 1 0 1 0 2 Ptinella cavelli (Broun, 1893) 2 4 10 3 0 19 Ptinella denticollis (Fairmaire, 1858) 0 2 3 2 0 7 Ptinella errabunda Johnson, 1975 0 2 15 3 1 21 Pteryx suturalis (Heer, 1841) 0 3 9 7 2 21 Nephanes titan (Newman, 1834) 12 6 28 7 7 60 Smicrus filicornis (Fairmaire & Laboulbène, 1855) 0 1 0 0 0 1 Baeocrara variolosa (Mulsant & Rey, 1861) 0 0 0 1 0 1 Acrotrichis arnoldi Rosskothen, 1935 1 0 0 0 0 1 Acrotrichis atomaria (De Geer, 1774) 13 14 65 21 4 117 Acrotrichis cephalotes (Allibert, 1844) 0 0 1 0 0 1 Acrotrichis cognata (Matthews, A., 1877) 25 15 126 31 18 215 Acrotrichis danica Sundt, 1958 7 3 32 16 3 61 Acrotrichis dispar (Matthews, A., 1865) 0 0 2 0 0 2 Acrotrichis fascicularis (Herbst, 1793) 43 27 172 50 13 305 Acrotrichis grandicollis (Mannerheim, 1844) 22 20 85 18 6 151 Acrotrichis henrici (Matthews, A., 1872) 12 0 33 4 0 49 Acrotrichis insularis (Mäklin, 1852) 6 8 19 4 0 37 Acrotrichis intermedia (Gillmeister, 1845) 9 13 99 28 2 151 Acrotrichis josephi (Matthews, A., 1872) 3 0 0 0 0 3 Acrotrichis lucidula Rosskothen, 1935 0 1 4 0 0 5

14

Yorkshire Coleoptera Checklist 2020

Acrotrichis montandonii (Allibert, 1844) 6 12 27 8 0 53 Acrotrichis norvegica Strand, A., 1941 0 0 2 2 0 4 Acrotrichis parva Rosskothen, 1935 1 0 2 1 0 4 Acrotrichis pumila (Erichson, 1845) 1 0 1 0 0 2 Acrotrichis rosskotheni Sundt, 1971 0 0 17 6 0 23 Acrotrichis rugulosa Rosskothen, 1935 1 2 2 1 0 6 Acrotrichis sanctaehelenae Johnson, 1972 0 1 1 0 0 2 Acrotrichis sericans (Heer, 1841) 12 6 46 11 3 78 Acrotrichis silvatica Rosskothen, 1935 1 2 8 0 0 11 Acrotrichis sitkaensis (Motschulsky, 1845) 27 4 80 26 4 141 Acrotrichis strandi Sundt, 1958 5 2 17 6 1 31 Acrotrichis thoracica (Waltl, 1838) 2 7 15 9 3 36

17 Leiodidae Sogda suturalis (Zetterstedt, 1828) 0 4 0 0 0 4 Hydnobius latifrons (Curtis, 1840) 0 0 2 0 0 2 Hydnobius punctatus (Sturm, 1807) 0 3 4 0 0 7 Hydnobius spinipes (Gyllenhal, 1813) 0 0 3 0 0 3 Triarthron maerkelii Märkel, 1840 0 2 0 0 0 2 Leiodes badia (Sturm, 1807) 0 2 4 0 0 6 Leiodes calcarata (Erichson, 1845) 3 8 44 7 3 65 Leiodes ciliaris (Schmidt, W.L.E., 1841) 0 1 0 0 0 1 Leiodes cinnamomea (Panzer, 1793) 1 2 2 0 0 5 Leiodes ferruginea (Fabricius, 1787) 4 3 2 1 1 11 Leiodes furva (Erichson, 1845) 1 0 0 0 0 1 Leiodes gallica (Reitter, 1884) 0 1 0 0 0 1 Leiodes gyllenhalii Stephens, 1829 0 2 0 0 0 2 Leiodes litura Stephens, 1832 1 4 2 1 0 8 Leiodes lucens (Fairmaire, 1855) 0 0 6 0 0 6 Leiodes lunicollis (Rye, 1872) 1 1 0 0 0 2 Leiodes nigrita (Schmidt, W.L.E., 1841) 0 1 0 0 0 1 Leiodes obesa (Schmidt, W.L.E., 1841) 1 0 12 5 0 18 Leiodes picea (Panzer, 1797) 0 1 0 0 0 1 Leiodes rufipennis (Paykull, 1798) 0 0 4 0 1 5 Leiodes rugosa Stephens, 1829 0 2 0 1 0 3 Leiodes strigipennis Daffner, 1983 0 0 11 2 0 13 Leiodes triepkii (Schmidt, W.L.E., 1841) 0 0 1 0 0 1 Liocyrtusa minuta (Ahrens, 1812) 1 1 0 1 0 3 Liocyrtusa vittata (Curtis, 1840) 2 1 2 1 2 8 Agaricophagus cephalotes Schmidt, W.L.E., 1841 0 0 1 0 0 1 Colenis immunda (Sturm, 1807) 1 2 5 4 0 12 Anisotoma glabra (Fabricius, 1792) 0 0 1 0 0 1 Anisotoma humeralis (Fabricius, 1792) 5 9 80 10 5 109 Anisotoma orbicularis (Herbst, 1792) 1 5 15 1 0 22 Amphicyllis globus (Fabricius, 1792) 0 3 12 1 0 16 Agathidium arcticum Thomson, C.G., 1862 0 0 1 0 0 1

15

Yorkshire Coleoptera Checklist 2020

Agathidium atrum (Paykull, 1798) 1 4 28 3 0 36 Agathidium convexum Sharp, 1866 2 0 2 1 0 5 Agathidium laevigatum Erichson, 1845 19 4 28 5 1 57 Agathidium marginatum Sturm, 1807 3 1 0 0 0 4 Agathidium nigrinum Sturm, 1807 2 3 21 4 0 30 Agathidium nigripenne (Fabricius, 1792) 2 7 34 14 3 60 Agathidium pisanum Brisout de Barneville, 1872 0 0 0 1 0 1 Agathidium rotundatum (Gyllenhal, 1827) 0 5 5 0 0 10 Agathidium seminulum (Linnaeus, 1758) 0 2 5 5 1 13 Agathidium varians Beck, 1817 10 6 35 5 1 57 Colon appendiculatum (Sahlberg, C.R., 1822) 0 1 0 0 0 1 Colon brunneum (Latreille, 1807) 1 2 3 3 0 9 Colon dentipes (Sahlberg, C.R., 1822) 0 1 0 1 0 2 Colon latum Kraatz, 1850 0 2 1 0 0 3 Colon serripes (Sahlberg, C.R., 1822) 0 2 2 1 0 5 Nemadus colonoides (Kraatz, 1851) 1 1 0 1 0 3 Nargus anisotomoides (Spence, 1813) 1 4 16 0 0 21 Nargus velox (Spence, 1813) 24 6 116 20 7 173 Nargus wilkinii (Spence, 1813) 3 11 67 5 0 86 Choleva agilis (Illiger, 1798) 2 3 17 1 0 23 Choleva angustata (Fabricius, 1781) 11 2 15 3 0 31 Choleva cisteloides (Frölich, 1799) 0 0 1 0 0 1 Choleva fagniezi Jeannel, 1922 0 0 16 5 0 21 Choleva glauca Britten, 1918 1 1 4 5 1 12 Choleva jeanneli Britten, 1922 7 3 10 7 0 27 Choleva oblonga Latreille, 1807 1 1 6 5 0 13 Choleva spadicea (Sturm, 1839) 0 3 7 0 0 10 Sciodrepoides fumatus (Spence, 1813) 3 8 21 11 2 45 Sciodrepoides watsoni (Spence, 1813) 19 19 101 22 3 164 Catops chrysomeloides (Panzer, 1798) 9 4 13 8 0 34 Catops coracinus Kellner, 1846 7 5 29 6 2 49 Catops fuliginosus Erichson, 1837 16 9 77 17 0 119 Catops fuscus (Panzer, 1794) 3 1 3 1 0 8 Catops grandicollis Erichson, 1837 11 5 35 15 6 72 Catops kirbii (Spence, 1813) 7 7 23 7 2 46 Catops longulus Kellner, 1846 0 5 9 3 1 18 Catops morio (Fabricius, 1787) 22 8 50 18 4 102 Catops nigricans (Spence, 1813) 18 4 48 10 2 82 Catops nigrita Erichson, 1837 0 6 58 4 2 70 Catops tristis (Panzer, 1793) 13 14 109 20 6 162 Catopidius depressus (Murray, 1856) 1 0 1 0 0 2 Ptomaphagus medius Rey, 1889 3 3 13 3 1 23 Ptomaphagus subvillosus (Goeze, 1777) 15 24 135 43 4 221 Ptomaphagus varicornis (Rosenhauer, 1847) 1 0 0 0 0 1 Leptinus testaceus Müller, P.W.J., 1817 1 5 8 3 0 17

16

Yorkshire Coleoptera Checklist 2020

18 Silphidae Necrodes littoralis (Linnaeus, 1758) 6 6 20 21 66 119 Thanatophilus dispar (Herbst, 1793) 0 1 0 2 0 3 Thanatophilus rugosus (Linnaeus, 1758) 27 8 62 27 10 134 Thanatophilus sinuatus (Fabricius, 1775) 9 5 28 6 3 51 Oiceoptoma thoracicum (Linnaeus, 1758) 10 38 48 11 0 107 Aclypea opaca (Linnaeus, 1758) 5 7 7 6 0 25 Aclypea undata (Müller, O.F., 1776) 0 1 0 0 0 1 Dendroxena quadrimaculata (Scopoli, 1772) 3 2 21 2 0 28 Silpha atrata Linnaeus, 1758 16 36 111 21 8 192 Silpha laevigata Fabricius, 1775 0 2 3 0 0 5 Silpha obscura Linnaeus, 1758 2 0 1 0 0 3 Silpha tristis Illiger, 1798 12 3 103 7 1 126 Silpha tyrolensis Laicharting, 1781 0 3 5 2 3 13 Nicrophorus humator (Gleditsch, 1767) 35 5 113 73 337 563 Nicrophorus interruptus Stephens, 1830 2 1 4 0 0 7 Nicrophorus investigator Zetterstedt, 1824 25 8 40 70 382 525 Nicrophorus vespillo (Linnaeus, 1758) 17 10 60 15 4 106 Nicrophorus vespilloides Herbst, 1783 12 25 201 17 8 263 Nicrophorus vestigator Herschel, 1807 0 4 0 0 0 4

19 Staphylinidae Pella limbata (Paykull, 1789) 1 0 2 0 0 3 Acidota crenata (Fabricius, 1792) 3 12 23 15 2 55 Acidota cruentata Mannerheim, 1830 0 4 10 7 0 21 Anthobium atrocephalum (Gyllenhal, 1827) 15 19 183 22 0 239 Anthobium unicolor (Marsham, 1802) 37 30 218 57 7 349 Anthophagus alpinus (Paykull, 1790) 0 0 0 4 0 4 Anthophagus caraboides (Linnaeus, 1758) 0 34 4 10 5 53 Deliphrum tectum (Paykull, 1789) 2 3 17 4 2 28 Eucnecosum brachypterum (Gravenhorst, 1802) 0 2 6 16 8 32 Geodromicus nigrita (Müller, P.W.J., 1821) 0 3 4 7 4 18 Lesteva hanseni Lohse, 1953 0 0 0 3 0 3 Lesteva longoelytrata (Goeze, 1777) 62 54 200 63 32 411 Lesteva monticola Kiesenwetter, 1847 0 4 2 13 13 32 Lesteva pubescens Mannerheim, 1830 1 13 12 5 2 33 Lesteva punctata Erichson, 1839 3 9 10 6 3 31 Lesteva sicula Erichson, 1840 4 4 63 7 1 79 Lesteva sicula subsp. heeri Fauvel, 1871 53 45 228 64 17 407 Olophrum consimile (Gyllenhal, 1810) 0 2 0 0 0 2 Olophrum fuscum (Gravenhorst, 1806) 2 3 22 12 7 46 Olophrum piceum (Gyllenhal, 1810) 23 44 206 52 20 345 Philorinum sordidum (Stephens, 1834) 15 15 30 6 5 71 Phyllodrepoidea crenata Ganglbauer, 1895 0 9 12 3 0 24 Eusphalerum luteum (Marsham, 1802) 1 9 19 6 1 36 Eusphalerum minutum (Fabricius, 1792) 8 13 5 39 7 72

17

Yorkshire Coleoptera Checklist 2020

Eusphalerum primulae (Stephens, 1834) 3 12 17 4 0 36 Eusphalerum sorbi (Gyllenhal, 1810) 4 21 13 17 4 59 Eusphalerum torquatum (Marsham, 1802) 3 4 16 12 8 43 Acrolocha minuta (Olivier, 1795) 0 1 2 0 0 3 Acrolocha sulcula (Stephens, 1834) 7 8 13 4 2 34 Acrulia inflata (Gyllenhal, 1813) 3 6 47 8 0 64 Dropephylla devillei Bernhauer, 1902 0 6 10 5 0 21 Dropephylla gracilicornis (Fairmaire & Laboulbène, 1856) 0 1 0 0 0 1 Dropephylla ioptera (Stephens, 1832) 7 4 10 7 2 30 Dropephylla vilis (Erichson, 1840) 3 11 51 9 0 74 Hapalaraea pygmaea (Paykull, 1800) 0 2 7 3 0 12 Micralymma marinum (Strøm, 1783) 4 9 1 0 0 14 Omalium allardi Fairmaire & Brisout de Barneville, 1859 0 3 6 4 0 13 Omalium caesum Gravenhorst, 1806 41 12 103 18 2 176 Omalium excavatum Stephens, 1834 32 8 41 15 2 98 Omalium exiguum Gyllenhal, 1810 0 1 2 0 0 3 Omalium italicum Bernhauer, 1902 8 2 38 4 0 52 Omalium laeviusculum Gyllenhal, 1827 7 6 0 0 0 13 Omalium laticolle Kraatz, 1858 0 0 2 4 0 6 Omalium oxyacanthae Gravenhorst, 1806 3 5 9 8 2 27 Omalium riparium Thomson, C.G., 1857 11 3 0 0 0 14 Omalium rivulare (Paykull, 1789) 43 24 225 36 7 335 Omalium rugatum Mulsant & Rey, 1880 0 2 9 3 0 14 Omalium rugulipenne Rye, 1864 0 2 0 0 0 2 Omalium septentrionis Thomson, C.G., 1856 0 6 8 3 0 17 Phloeonomus punctipennis Thomson, C.G., 1867 3 6 26 14 3 52 Phloeonomus pusillus (Gravenhorst, 1806) 3 5 19 2 1 30 Phloeostiba plana (Paykull, 1792) 0 2 1 0 0 3 Phyllodrepa floralis (Paykull, 1789) 14 6 17 5 0 42 Phyllodrepa puberula Bernhauer, 1903 0 1 1 0 0 2 Phyllodrepa salicis (Gyllenhal, 1810) 0 1 0 0 0 1 Xylodromus concinnus (Marsham, 1802) 14 17 17 8 2 58 Xylodromus depressus (Gravenhorst, 1802) 5 3 8 6 1 23 Xylostiba monilicornis (Gyllenhal, 1810) 0 0 6 1 0 7 Coryphium angusticolle Stephens, 1834 3 3 30 6 0 42 Eudectus whitei Sharp, 1871 0 0 0 3 0 3 Megarthrus bellevoyei (Saulcy, 1862) 5 5 21 0 0 31 Megarthrus denticollis (Beck, 1817) 11 12 100 23 6 152 Megarthrus depressus (Paykull, 1789) 20 19 155 39 3 236 Megarthrus prosseni Schatzmayr, 1904 2 0 14 0 0 16 Metopsia clypeata (Müller, P.W.J., 1821) 27 15 74 32 1 149 Proteinus atomarius Erichson, 1840 0 1 9 2 0 12 Proteinus brachypterus (Fabricius, 1792) 14 26 100 23 3 166 Proteinus crenulatus Pandellé, 1867 0 3 13 2 0 18 Proteinus laevigatus Hochhuth, 1871 8 1 37 11 1 58 Proteinus ovalis Stephens, 1834 42 15 217 37 3 314

18

Yorkshire Coleoptera Checklist 2020

Arrhenopeplus tesserula Curtis, 1828 0 0 2 1 1 4 Micropeplus fulvus Erichson, 1840 15 6 53 19 0 93 Micropeplus porcatus (Paykull, 1789) 4 10 1 4 0 19 Micropeplus staphylinoides (Marsham, 1802) 19 9 38 9 1 76 Trichonyx sulcicollis (Reichenbach, 1816) 0 1 1 0 0 2 Trimium brevicorne (Reichenbach, 1816) 0 2 1 1 0 4 Bibloporus bicolor (Denny, 1825) 0 2 7 0 0 9 Bibloporus minutus Raffray, 1914 0 3 1 1 0 5 Bibloplectus ambiguus (Reichenbach, 1816) 1 1 1 7 0 10 Bibloplectus pusillus (Denny, 1825) 0 0 0 1 0 1 Bibloplectus tenebrosus (Reitter, 1880) 0 0 0 3 0 3 Euplectus bescidicus Reitter, 1881 0 1 0 0 0 1 Euplectus karstenii (Reichenbach, 1816) 0 0 3 3 0 6 Euplectus nanus (Reichenbach, 1816) 1 1 3 1 1 7 Euplectus piceus Motschulsky, 1835 0 2 3 2 0 7 Euplectus sanguineus Denny, 1825 8 5 23 5 2 43 Euplectus signatus (Reichenbach, 1816) 0 2 12 0 0 14 Plectophloeus nitidus (Fairmaire, 1857) 0 0 2 0 0 2 Batrisodes venustus (Reichenbach, 1816) 1 1 0 1 0 3 Bryaxis bulbifer (Reichenbach, 1816) 13 9 26 37 2 87 Bryaxis curtisii (Leach, 1817) 0 5 7 7 0 19 Bryaxis puncticollis (Denny, 1825) 5 14 24 19 1 63 Bythinus burrellii Denny, 1825 2 11 8 16 3 40 Bythinus macropalpus Aubé, 1833 3 2 10 4 0 19 Tychus niger (Paykull, 1800) 16 20 23 13 4 76 Brachygluta fossulata (Reichenbach, 1816) 10 12 32 22 0 76 Brachygluta helferi (Schmidt-Göbel, 1836) 21 0 0 0 0 21 Fagniezia impressa (Panzer, 1803) 0 2 2 3 0 7 Reichenbachia juncorum (Leach, 1817) 5 4 3 5 0 17 Rybaxis longicornis (Leach, 1817) 16 0 81 8 1 106 Pselaphaulax dresdensis (Herbst, 1792) 2 0 1 2 0 5 Pselaphus heisei Herbst, 1792 6 10 11 15 3 45 Claviger testaceus Preyssler, 1790 0 2 0 0 0 2 Phloeocharis subtilissima Mannerheim, 1830 4 9 34 9 2 58 Cilea silphoides (Linnaeus, 1767) 9 9 18 6 0 42 Lamprinodes saginatus (Gravenhorst, 1806) 8 3 6 0 0 17 Sepedophilus constans (Fowler, 1888) 0 1 1 0 0 2 Sepedophilus immaculatus (Stephens, 1832) 16 2 19 1 0 38 Sepedophilus littoreus (Linnaeus, 1758) 4 6 24 2 0 36 Sepedophilus lusitanicus Hammond, 1973 0 0 1 0 0 1 Sepedophilus marshami (Stephens, 1832) 83 26 271 27 6 413 Sepedophilus nigripennis (Stephens, 1832) 98 26 266 36 7 433 Sepedophilus pedicularius (Gravenhorst, 1802) 1 0 11 11 0 23 Sepedophilus testaceus (Fabricius, 1792) 4 0 0 2 0 6 Tachinus corticinus Gravenhorst, 1802 12 11 25 39 4 91 Tachinus elongatus Gyllenhal, 1810 0 16 10 7 4 37

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Yorkshire Coleoptera Checklist 2020

Tachinus humeralis Gravenhorst, 1802 10 11 54 10 3 88 Tachinus laticollis Gravenhorst, 1802 20 20 110 14 4 168 Tachinus lignorum (Linnaeus, 1758) 0 1 3 3 0 7 Tachinus marginellus (Fabricius, 1781) 39 40 278 62 17 436 Tachinus pallipes (Gravenhorst, 1806) 5 12 60 9 2 88 Tachinus proximus Kraatz, 1855 2 5 32 7 1 47 Tachinus rufipennis Gyllenhal, 1810 0 1 0 1 1 3 Tachinus rufipes (Linnaeus, 1758) 117 76 630 132 17 972 Tachinus subterraneus (Linnaeus, 1758) 11 4 36 8 1 60 Tachyporus Gravenhorst, 1802 0 0 1 0 0 1 Tachyporus chrysomelinus agg. 5 13 21 8 9 56 Tachyporus atriceps Stephens, 1832 11 11 20 10 0 52 Tachyporus chrysomelinus (Linnaeus, 1758) 30 8 332 44 5 419 Tachyporus dispar (Paykull, 1789) 83 81 411 90 16 681 Tachyporus formosus Matthews, A.H., 1838 0 1 1 1 1 4 Tachyporus hypnorum (Fabricius, 1775) 188 109 790 113 19 1219 Tachyporus nitidulus (Fabricius, 1781) 64 47 307 45 7 470 Tachyporus obtusus (Linnaeus, 1767) 122 73 541 83 24 843 Tachyporus pallidus Sharp, 1871 47 24 139 29 2 241 Tachyporus pusillus Gravenhorst, 1806 8 4 34 10 3 59 Tachyporus quadriscopulatus Pandellé, 1869 0 0 0 1 0 1 Tachyporus solutus Erichson, 1839 27 8 268 3 1 307 Tachyporus tersus Erichson, 1839 0 10 8 7 1 26 Tachyporus transversalis Gravenhorst, 1806 8 6 76 4 0 94 Bolitobius castaneus (Stephens, 1832) 8 6 28 7 2 51 Bolitobius cingulatus (Mannerheim, 1830) 5 8 50 6 3 72 Ischnosoma splendidum (Gravenhorst, 1806) 22 16 61 14 3 116 Lordithon exoletus (Erichson, 1839) 2 5 11 1 0 19 Lordithon lunulatus (Linnaeus, 1761) 4 12 37 10 6 69 Lordithon thoracicus (Fabricius, 1777) 5 14 70 19 2 110 Lordithon trinotatus (Erichson, 1839) 9 17 83 11 1 121 Mycetoporus angularis Mulsant & Rey, 1853 0 1 1 0 0 2 Mycetoporus baudueri Mulsant & Rey, 1875 0 1 0 0 0 1 Mycetoporus bimaculatus Lacordaire, 1835 0 0 0 1 0 1 Mycetoporus clavicornis (Stephens, 1832) 3 14 33 8 2 60 Mycetoporus despectus Strand, A., 1969 0 0 6 1 2 9 Mycetoporus lepidus (Gravenhorst, 1806) 9 28 56 26 3 122 Mycetoporus longulus Mannerheim, 1830 4 8 10 10 2 34 Mycetoporus nigricollis Stephens, 1835 3 1 7 2 1 14 Mycetoporus piceolus Rey, 1883 4 1 0 0 0 5 Mycetoporus punctus (Gravenhorst, 1806) 2 4 6 0 0 12 Mycetoporus rufescens (Stephens, 1832) 2 12 21 8 2 45 Parabolitobius inclinans (Gravenhorst, 1806) 0 9 7 4 0 20 Trichophya pilicornis (Gyllenhal, 1810) 0 3 4 0 0 7 Habrocerus capillaricornis (Gravenhorst, 1806) 35 15 131 44 8 233 Gymnusa brevicollis (Paykull, 1800) 1 8 27 3 3 42

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Yorkshire Coleoptera Checklist 2020

Gymnusa variegata Kiesenwetter, 1845 1 4 11 0 2 18 Deinopsis erosa (Stephens, 1832) 4 0 16 3 0 23 Myllaena brevicornis (Matthews, A.H., 1838) 3 12 30 13 10 68 Myllaena dubia (Gravenhorst, 1806) 10 5 47 5 3 70 Myllaena elongata (Matthews, A.H., 1838) 1 3 3 0 0 7 Myllaena gracilis (Matthews, A.H., 1838) 3 0 4 2 0 9 Myllaena infuscata Kraatz, 1853 11 4 18 12 1 46 Myllaena intermedia Erichson, 1837 4 0 18 1 3 26 Myllaena kraatzi Sharp, 1871 1 8 10 0 1 20 Myllaena minuta (Gravenhorst, 1806) 4 1 15 12 3 35 Amarochara umbrosa (Erichson, 1837) 0 2 1 0 0 3 Calodera aethiops (Gravenhorst, 1802) 19 3 16 6 2 46 Calodera nigrita Mannerheim, 1830 2 2 4 1 1 10 Calodera protensa Mannerheim, 1830 0 0 0 2 0 2 Calodera riparia Erichson, 1837 2 3 12 3 0 20 Chilomorpha longitarsis (Thomson, C.G., 1867) 0 13 1 0 0 14 Cousya nitidiventris (Fagel, 1958) 0 0 1 0 0 1 Crataraea suturalis (Mannerheim, 1830) 1 1 5 4 1 12 Haploglossa gentilis (Märkel, 1844) 1 1 0 0 0 2 Haploglossa nidicola (Fairmaire, 1852) 2 1 2 2 1 8 Haploglossa villosula (Stephens, 1832) 0 11 1 3 1 16 Ilyobates bennetti Donisthorpe, 1914 4 0 4 3 0 11 Ilyobates nigricollis (Paykull, 1800) 0 0 3 1 0 4 Ilyobates propinquus (Aubé, 1850) 2 0 5 0 0 7 Ischnoglossa prolixa (Gravenhorst, 1802) 0 0 10 2 0 12 Ischnoglossa turcica Wunderle, 1992 0 0 1 0 0 1 Mniusa incrassata (Mulsant & Rey, 1852) 1 8 21 9 5 44 Ocalea badia Erichson, 1837 0 3 1 1 0 5 Ocalea latipennis Sharp, 1870 0 2 2 1 0 5 Ocalea picata (Stephens, 1832) 3 23 53 19 8 106 Ocalea rivularis Miller, L., 1851 0 1 0 0 0 1 Ocyusa maura (Erichson, 1837) 11 2 80 12 1 106 Ocyusa picina (Aubé, 1850) 1 1 5 3 0 10 Oxypoda acuminata (Stephens, 1832) 5 7 25 4 3 44 Oxypoda alternans (Gravenhorst, 1802) 8 23 106 20 3 160 Oxypoda annularis Mannerheim, 1830 2 6 22 5 0 35 Oxypoda brachyptera (Stephens, 1832) 9 0 3 1 0 13 Oxypoda brevicornis (Stephens, 1832) 28 21 83 12 4 148 Oxypoda carbonaria (Heer, 1841) 2 5 18 2 0 27 Oxypoda elongatula Aubé, 1850 26 19 270 46 7 368 Oxypoda exoleta Erichson, 1839 6 1 4 0 0 11 Oxypoda flavicornis Kraatz, 1856 3 0 10 1 0 14 Oxypoda formiceticola Märkel, 1841 0 2 14 0 0 16 Oxypoda haemorrhoa (Mannerheim, 1830) 28 17 157 18 9 229 Oxypoda induta Mulsant & Rey, 1861 1 2 2 0 0 5 Oxypoda islandica Kraatz, 1857 0 0 0 3 0 3

21

Yorkshire Coleoptera Checklist 2020

Oxypoda lentula Erichson, 1837 15 1 7 0 0 23 Oxypoda longipes Mulsant & Rey, 1861 2 0 1 0 0 3 Oxypoda nigricornis Motschulsky, 1860 0 0 5 1 0 6 Oxypoda opaca (Gravenhorst, 1802) 19 11 71 18 1 120 Oxypoda procerula Mannerheim, 1830 1 3 26 6 3 39 Oxypoda soror Thomson, C.G., 1855 0 0 1 0 0 1 Oxypoda spectabilis Märkel, 1844 1 7 2 2 0 12 Oxypoda vittata Märkel, 1842 0 0 1 0 0 1 Tetralaucopora longitarsis (Erichson, 1837) 14 12 37 6 5 74 Tetralaucopora rubicunda (Erichson, 1837) 0 2 0 0 1 3 Phloeopora corticalis (Gravenhorst, 1802) 1 2 8 1 1 13 Phloeopora testacea (Mannerheim, 1830) 3 5 29 5 0 42 Thiasophila angulata (Erichson, 1837) 0 4 11 0 0 15 Dinarda maerkeli Kiesenwetter, 1843 1 7 2 0 0 10 Meotica exilis (Knoch in Gravenhorst, 1806) 3 0 1 2 1 7 Meotica exillima Sharp, 1915 2 1 3 1 0 7 Meotica filiformis (Motschulsky, 1860) 1 2 9 2 0 14 Dacrila fallax (Kraatz, 1856) 5 0 5 0 0 10 Dasygnypeta velata (Erichson, 1837) 2 0 0 0 0 2 Gnypeta caerulea (Sahlberg, C.R., 1831) 0 0 0 5 5 10 Gnypeta carbonaria (Mannerheim, 1830) 8 6 22 4 3 43 Gnypeta ripicola (Kiesenwetter, 1844) 0 1 3 0 0 4 Gnypeta rubrior Tottenham, 1939 3 2 14 0 0 19 Ischnopoda leucopus (Marsham, 1802) 1 9 5 2 0 17 Ischnopoda umbratica (Erichson, 1837) 0 2 1 0 0 3 Tachyusa coarctata (Erichson, 1837) 0 1 1 0 0 2 Tachyusa constricta (Erichson, 1837) 0 6 1 6 0 13 Thinonoma atra (Gravenhorst, 1806) 18 4 41 8 1 72 Acrotona aterrima (Gravenhorst, 1802) 17 13 136 17 8 191 Acrotona benicki (Allen, 1940) 0 1 0 0 0 1 Acrotona exigua (Erichson, 1837) 0 0 1 0 0 1 Acrotona muscorum (Brisout de Barneville, 1860) 7 7 36 21 6 77 Acrotona obfuscata (Gravenhorst, 1802) 0 0 3 0 0 3 Acrotona parvula (Mannerheim, 1830) 13 5 44 14 9 85 Acrotona pygmaea (Gravenhorst, 1802) 1 1 26 4 0 32 Adota maritima (Mannerheim, 1843) 1 0 0 0 0 1 Alaobia gagatina (Baudi, 1848) 3 11 10 3 1 28 Alaobia pallidicornis (Thomson, C.G., 1856) 2 2 19 11 1 35 Alaobia sodalis (Erichson, 1837) 1 12 17 7 2 39 Alaobia taxiceroides Munster, 1932 0 1 5 0 0 6 Alaobia trinotata (Kraatz, 1856) 30 19 125 23 5 202 Alianta incana (Erichson, 1837) 6 1 42 9 2 60 Aloconota cambrica (Wollaston, 1855) 0 3 0 1 1 5 Aloconota coulsoni (Last, 1952) 0 0 1 0 0 1 Aloconota currax (Kraatz, 1856) 2 9 4 0 0 15 Aloconota gregaria (Erichson, 1839) 58 33 140 37 7 275

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Yorkshire Coleoptera Checklist 2020

Aloconota insecta (Thomson, C.G., 1856) 3 2 13 3 2 23 Aloconota languida (Erichson, 1837) 4 0 1 2 0 7 Aloconota longicollis (Mulsant & Rey, 1852) 2 0 0 0 0 2 Aloconota sulcifrons (Stephens, 1832) 4 2 28 1 1 36 Amidobia talpa (Heer, 1841) 1 11 13 0 0 25 Amischa analis (Gravenhorst, 1802) 52 49 277 49 16 443 Amischa bifoveolata (Mannerheim, 1830) 14 9 21 6 2 52 Amischa decipiens (Sharp, 1869) 10 7 41 7 4 69 Amischa forcipata Mulsant & Rey, 1873 5 0 4 1 0 10 Amischa nigrofusca (Stephens, 1829) 1 1 12 3 2 19 Anopleta corvina (Thomson, C.G., 1856) 0 4 8 0 2 14 Anopleta puberula (Sharp, 1869) 0 0 1 0 0 1 Atheta aeneicollis (Sharp, 1869) 17 27 68 22 2 136 Atheta aquatica (Thomson, C.G., 1852) 1 8 37 5 1 52 Atheta aquatilis (Thomson, C.G., 1867) 1 3 34 1 2 41 Atheta brunneipennis (Thomson, C.G., 1852) 1 4 23 5 1 34 Atheta castanoptera (Mannerheim, 1830) 2 25 132 24 4 187 Atheta graminicola (Gravenhorst, 1806) 41 29 238 57 17 382 Atheta hypnorum (Kiesenwetter, 1850) 1 14 18 5 0 38 Atheta incognita (Sharp, 1869) 0 10 9 0 0 19 Atheta triangulum (Kraatz, 1856) 20 4 24 12 2 62 Atheta xanthopus (Thomson, C.G., 1856) 7 7 45 5 3 67 Atheta basicornis (Mulsant & Rey, 1852) 0 0 1 3 0 4 Atheta britanniae (Bernhauer & Scheerpeltz, 1926) 3 10 27 6 4 50 Atheta crassicornis (Fabricius, 1792) 24 55 302 66 15 462 Atheta diversa (Sharp, 1869) 0 0 2 1 0 3 Atheta divisa (Märkel, 1844) 1 0 2 4 0 7 Atheta euryptera (Stephens, 1832) 3 3 3 2 0 11 Atheta fungicola (Thomson, C.G., 1852) 0 5 7 1 0 13 Atheta harwoodi (Williams, 1930) 10 5 60 12 4 91 Atheta intermedia (Thomson, C.G., 1852) 1 1 3 3 0 8 Atheta liturata (Stephens, 1832) 1 7 9 11 4 32 Atheta nidicola (Johansen, 1914) 0 1 2 2 0 5 Atheta nigritula (Gravenhorst, 1802) 1 0 2 0 0 3 Atheta oblita (Erichson, 1839) 1 2 4 0 0 7 Atheta paracrassicornis Brundin, 1954 0 0 0 1 0 1 Atheta pilicornis (Thomson, C.G., 1852) 0 1 0 1 0 2 Atheta ravilla (Erichson, 1839) 4 7 42 9 3 65 Atheta strandiella (Brundin, 1954) 1 0 0 0 0 1 Atheta vaga (Heer, 1839) 6 12 46 13 1 78 Badura macrocera (Thomson, C.G., 1856) 3 3 16 6 2 30 Badura puncticollis (Benick, 1938) 0 0 1 0 0 1 Bessobia excellens (Kraatz, 1856) 1 7 3 8 0 19 Bessobia fungivora (Thomson, C.G., 1867) 3 7 4 6 0 20 Bessobia monticola (Thomson, C.G., 1852) 1 3 4 3 0 11 Bessobia occulta (Erichson, 1837) 0 3 3 0 1 7

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Yorkshire Coleoptera Checklist 2020

Boreophilia eremita (Rye, 1866) 4 10 18 5 6 43 Brundinia marina (Mulsant & Rey, 1853) 19 0 0 0 0 19 Brundinia meridionalis (Mulsant & Rey, 1853) 4 0 0 0 0 4 Cadaverota cadaverina (Brisout de Barneville, 1860) 0 9 16 1 0 26 Callicerus obscurus Gravenhorst, 1802 4 3 9 2 0 18 Callicerus rigidicornis (Erichson, 1839) 1 2 16 2 0 21 Chaetida longicornis (Gravenhorst, 1802) 21 22 147 19 11 220 Coprothassa melanaria (Mannerheim, 1830) 2 2 0 0 0 4 Dadobia immersa (Erichson, 1837) 1 1 4 6 0 12 Dalotia coriaria (Kraatz, 1856) 8 2 44 13 0 67 Datomicra canescens (Sharp, 1869) 0 0 5 0 0 5 Datomicra celata (Erichson, 1837) 24 16 227 26 12 305 Datomicra dadopora (Thomson, C.G., 1867) 4 5 49 12 4 74 Datomicra nigra (Kraatz, 1856) 35 12 151 35 11 244 Datomicra sordidula (Erichson, 1837) 11 8 51 6 3 79 Datomicra zosterae (Thomson, C.G., 1856) 2 1 0 2 0 5 Dilacra luteipes (Erichson, 1837) 5 1 12 4 0 22 Dilacra vilis (Erichson, 1837) 2 0 16 6 0 24 Dimetrota aeneipennis (Thomson, C.G., 1856) 0 0 4 0 0 4 Dimetrota atramentaria (Gyllenhal, 1810) 25 14 138 22 16 215 Dimetrota cauta (Erichson, 1837) 6 0 2 1 0 9 Dimetrota cinnamoptera (Thomson, C.G., 1856) 0 17 32 4 6 59 Dimetrota ischnocera (Thomson, C.G., 1870) 2 2 32 7 1 44 Dimetrota laevana (Mulsant & Rey, 1852) 0 5 10 1 2 18 Dimetrota marcida (Erichson, 1837) 0 3 61 2 1 67 Dimetrota nigripes (Thomson, C.G., 1856) 8 4 48 5 7 72 Dimetrota setigera (Sharp, 1869) 4 3 34 3 1 45 Dinaraea aequata (Erichson, 1837) 17 18 93 23 1 152 Dinaraea angustula (Gyllenhal, 1810) 22 2 55 3 1 83 Dinaraea linearis (Gravenhorst, 1802) 0 1 3 1 0 5 Dochmonota clancula (Erichson, 1837) 0 0 19 6 0 25 Enalodroma hepatica (Erichson, 1839) 1 2 8 9 4 24 Geostiba circellaris (Gravenhorst, 1806) 36 21 310 55 13 435 Halobrecta algae (Hardy, 1851) 3 0 0 0 0 3 Halobrecta flavipes Thomson, C.G., 1861 4 3 0 0 0 7 Hydrosmecta delicatula (Sharp, 1869) 0 1 0 0 0 1 Hydrosmecta eximia (Sharp, 1869) 0 2 0 1 0 3 Hydrosmecta fragilis (Kraatz, 1854) 0 2 0 0 0 2 Hydrosmecta subtilissima (Kraatz, 1854) 0 0 0 1 0 1 Liogluta alpestris (Heer, 1839) 2 34 16 2 3 57 Liogluta granigera (Kiesenwetter, 1850) 1 2 18 2 2 25 Liogluta longiuscula (Gravenhorst, 1802) 6 9 19 2 1 37 Liogluta microptera Thomson, C.G., 1867 1 6 74 13 3 97 Liogluta pagana (Erichson, 1839) 1 3 0 2 0 6 Lyprocorrhe anceps (Erichson, 1837) 0 6 5 1 0 12 Microdota Mulsant & Rey, 1873 0 0 1 0 0 1

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Yorkshire Coleoptera Checklist 2020

Microdota amicula (Stephens, 1832) 22 25 107 29 11 194 Microdota atricolor (Sharp, 1869) 4 4 14 3 1 26 Microdota benickiella (Brundin, 1948) 0 0 2 2 0 4 Microdota boreella (Brundin, 1948) 0 0 2 1 0 3 Microdota excelsa (Bernhauer, 1911) 0 0 3 0 0 3 Microdota glabricula (Thomson, C.G., 1867) 0 0 1 0 1 2 Microdota indubia (Sharp, 1869) 5 8 38 15 2 68 Microdota liliputana (Brisout de Barneville, 1860) 0 0 2 0 0 2 Microdota minuscula (Brisout de Barneville, 1860) 0 0 1 0 0 1 Microdota spatuloides (Benick, 1939) 0 1 0 0 0 1 Microdota subtilis (Scriba, 1866) 0 0 2 3 0 5 Mocyta amplicollis (Mulsant & Rey, 1873) 2 2 16 2 2 24 Mocyta clientula (Erichson, 1839) 2 5 16 4 0 27 Mocyta fungi (Gravenhorst, 1806) 155 84 964 178 47 1428 Mocyta orbata (Erichson, 1837) 5 1 34 8 4 52 Mocyta orphana (Erichson, 1837) 1 0 4 0 0 5 Mycetota laticollis (Stephens, 1832) 39 15 241 44 11 350 Nehemitropia lividipennis (Mannerheim, 1830) 16 13 58 14 1 102 Neohilara subterranea (Mulsant & Rey, 1853) 0 0 2 0 0 2 Notothecta confusa (Märkel, 1844) 0 0 1 0 0 1 Notothecta flavipes (Gravenhorst, 1806) 0 6 14 0 0 20 Oreostiba tibialis (Heer, 1839) 0 0 0 8 4 12 Ousipalia caesula (Erichson, 1839) 0 1 0 0 0 1 Pachnida nigella (Erichson, 1837) 24 2 88 8 1 123 Pachyatheta mortuorum (Thomson, C.G., 1867) 0 0 0 1 0 1 Parameotica difficilis (Brisout de Barneville, 1860) 17 1 1 0 1 20 Philhygra arctica (Thomson, C.G., 1856) 0 8 2 8 5 23 Philhygra britteni Joy, 1913 6 2 9 0 0 17 Philhygra debilis (Erichson, 1837) 7 0 6 0 1 14 Philhygra deformis (Kraatz, 1856) 0 1 0 0 1 2 Philhygra elongatula (Gravenhorst, 1802) 17 23 56 29 14 139 Philhygra fallaciosa (Sharp, 1869) 1 2 3 0 0 6 Philhygra gyllenhalii (Thomson, C.G., 1856) 5 2 18 3 0 28 Philhygra hygrobia (Thomson, C.G., 1856) 7 1 18 3 0 29 Philhygra hygrotopora (Kraatz, 1856) 2 6 14 3 3 28 Philhygra luridipennis (Mannerheim, 1830) 2 3 7 3 0 15 Philhygra malleus (Joy, 1913) 6 5 31 6 2 50 Philhygra melanocera (Thomson, C.G., 1856) 6 5 36 5 0 52 Philhygra obtusangula (Joy, 1913) 1 0 15 2 2 20 Philhygra palustris (Kiesenwetter, 1844) 6 5 7 1 0 19 Philhygra terminalis (Gravenhorst, 1806) 4 0 0 0 0 4 Philhygra volans (Scriba, 1859) 3 1 13 3 1 21 Plataraea brunnea (Fabricius, 1798) 13 6 34 2 1 56 Pycnota paradoxa (Mulsant & Rey, 1861) 3 0 0 1 0 4 Rhagocneme subsinuata (Erichson, 1839) 1 4 15 2 0 22 Schistoglossa curtipennis (Sharp, 1869) 0 0 7 2 0 9

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Yorkshire Coleoptera Checklist 2020

Schistoglossa gemina (Erichson, 1837) 3 0 3 2 0 8 Thinobaena vestita (Gravenhorst, 1806) 9 6 0 0 0 15 Trichiusa immigrata Lohse, 1984 10 0 47 0 1 58 Thamiaraea cinnamomea (Gravenhorst, 1802) 0 0 1 0 0 1 Thamiaraea hospita (Märkel, 1844) 0 1 0 0 0 1 Aleochara bilineata Gyllenhal, 1810 2 4 6 0 0 12 Aleochara binotata Kraatz, 1858 1 0 0 0 0 1 Aleochara bipustulata (Linnaeus, 1761) 29 10 54 10 1 104 Aleochara brevipennis Gravenhorst, 1806 3 0 1 0 1 5 Aleochara cuniculorum Kraatz, 1858 3 0 3 2 1 9 Aleochara curtula (Goeze, 1777) 12 15 61 18 12 118 Aleochara discipennis Mulsant & Rey, 1853 0 0 2 0 0 2 Aleochara fumata Gravenhorst, 1802 0 0 3 0 0 3 Aleochara funebris Wollaston, 1864 1 2 20 0 2 25 Aleochara grisea Kraatz, 1856 9 4 0 0 0 13 Aleochara inconspicua Aubé, 1850 0 0 1 0 0 1 Aleochara intricata Mannerheim, 1830 2 6 12 2 0 22 Aleochara kamila Likovský, 1984 2 1 2 3 0 8 Aleochara lanuginosa Gravenhorst, 1802 15 24 115 23 17 194 Aleochara lygaea Kraatz, 1862 0 2 0 0 0 2 Aleochara moerens Gyllenhal, 1827 0 0 4 0 0 4 Aleochara obscurella Gravenhorst, 1806 13 11 0 0 0 24 Aleochara punctatella Motschulsky, 1858 3 4 0 0 0 7 Aleochara ruficornis Gravenhorst, 1802 6 8 18 3 1 36 Aleochara spadicea (Erichson, 1837) 6 4 2 6 0 18 Aleochara sparsa Heer, 1839 6 8 5 6 2 27 Aleochara tristis Gravenhorst, 1806 0 0 1 0 0 1 Aleochara verna Say, 1836 5 3 5 3 1 17 Aleochara villosa Mannerheim, 1830 0 2 0 0 0 2 Tinotus morion (Gravenhorst, 1802) 8 11 28 10 2 59 Drusilla canaliculata (Fabricius, 1787) 38 37 294 18 8 395 Lomechusa emarginata (Paykull, 1789) 0 0 6 0 0 6 Lomechusa paradoxa Gravenhorst, 1806 0 1 0 0 0 1 Cordalia obscura (Gravenhorst, 1802) 0 3 2 1 0 6 Falagria caesa Erichson, 1837 0 5 17 1 0 23 Falagria sulcatula (Gravenhorst, 1806) 0 0 4 0 0 4 Falagrioma thoracica (Stephens, 1832) 0 1 2 0 0 3 Myrmecocephalus concinnus (Erichson, 1839) 0 0 1 0 0 1 Myrmecopora brevipes Butler, 1909 0 0 1 0 0 1 Myrmecopora sulcata (Kiesenwetter, 1850) 4 2 0 0 0 6 Myrmecopora uvida (Erichson, 1840) 0 1 0 0 0 1 Autalia impressa (Olivier, 1795) 4 15 122 16 3 160 Autalia longicornis Scheerpeltz, 1947 4 2 31 3 3 43 Autalia puncticollis Sharp, 1864 0 0 0 3 1 4 Autalia rivularis (Gravenhorst, 1802) 19 16 153 25 9 222 Agaricochara latissima (Stephens, 1832) 0 6 52 7 2 67

26

Yorkshire Coleoptera Checklist 2020

Encephalus complicans Stephens, 1832 6 3 18 9 2 38 Gyrophaena Mannerheim, 1830 0 0 1 0 0 1 Gyrophaena affinis Mannerheim, 1830 7 5 20 3 0 35 Gyrophaena bihamata Thomson, C.G., 1867 2 0 9 1 0 12 Gyrophaena fasciata (Marsham, 1802) 1 6 23 10 0 40 Gyrophaena gentilis Erichson, 1839 6 16 38 13 2 75 Gyrophaena joyi Wendeler, 1924 1 0 0 2 0 3 Gyrophaena joyioides Wüsthoff, 1937 0 0 7 5 0 12 Gyrophaena manca Erichson, 1839 3 8 33 14 1 59 Gyrophaena minima Erichson, 1837 1 6 6 5 1 19 Gyrophaena munsteri Strand, A., 1935 0 1 0 1 1 3 Gyrophaena nana (Paykull, 1800) 2 0 1 1 1 5 Gyrophaena poweri Crotch, 1867 1 0 0 0 0 1 Gyrophaena strictula Erichson, 1839 0 1 4 0 0 5 Gyrophaena williamsi Strand, A., 1935 0 0 1 1 0 2 Anomognathus cuspidatus (Erichson, 1839) 0 5 7 0 1 13 Homalota plana (Gyllenhal, 1810) 4 8 16 7 1 36 Bolitochara Mannerheim, 1830 0 1 0 0 0 1 Bolitochara bella Märkel, 1844 3 0 1 1 0 5 Bolitochara lucida (Gravenhorst, 1802) 0 7 14 7 0 28 Bolitochara mulsanti Sharp, 1875 0 3 6 0 1 10 Bolitochara obliqua Erichson, 1837 9 22 129 18 4 182 Bolitochara pulchra (Gravenhorst, 1806) 0 1 1 0 0 2 Leptusa fumida (Erichson, 1839) 10 31 180 33 5 259 Leptusa norvegica Strand, A., 1941 0 3 2 0 0 5 Leptusa pulchella (Mannerheim, 1830) 1 8 14 3 2 28 Leptusa ruficollis (Erichson, 1839) 1 12 35 6 1 55 Heterota plumbea (Waterhouse, G.R., 1858) 1 0 0 0 0 1 Thecturota marchii (Dodero, 1922) 0 0 1 0 0 1 Placusa depressa Mäklin, 1845 0 1 3 0 0 4 Placusa tachyporoides (Waltl, 1838) 0 0 0 1 0 1 Arena tabida (Kiesenwetter, 1850) 2 0 0 0 0 2 Phytosus balticus Kraatz, 1859 2 4 0 0 0 6 Phytosus spinifer Curtis, 1838 1 2 0 0 0 3 Diglotta mersa (Haliday, 1837) 3 1 0 0 0 4 Hygronoma dimidiata (Gravenhorst, 1806) 27 7 85 16 5 140 Cypha discoidea (Erichson, 1839) 4 0 2 0 1 7 Cypha hanseni (Palm, 1949) 0 1 0 1 0 2 Cypha laeviuscula (Mannerheim, 1830) 8 7 15 7 3 40 Cypha longicornis (Paykull, 1800) 33 27 99 17 2 178 Cypha pulicaria (Erichson, 1839) 5 3 4 1 0 13 Cypha punctum (Motschulsky, 1857) 1 2 0 0 0 3 Cypha seminulum (Erichson, 1839) 1 0 0 0 0 1 Holobus apicatus (Erichson, 1837) 3 8 7 3 2 23 Oligota inflata (Mannerheim, 1830) 4 13 19 3 0 39 Oligota parva Kraatz, 1862 12 1 25 0 0 38

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Yorkshire Coleoptera Checklist 2020

Oligota picipes (Stephens, 1832) 4 2 4 0 0 10 Oligota pumilio Kiesenwetter, 1858 0 1 1 0 0 2 Oligota punctulata Heer, 1839 7 1 54 13 3 78 Oligota pusillima (Gravenhorst, 1806) 6 2 8 3 0 19 Scaphidium quadrimaculatum Olivier, 1790 1 19 41 3 1 65 Scaphisoma agaricinum (Linnaeus, 1758) 3 14 53 14 2 86 Scaphisoma boleti (Panzer, 1793) 8 10 15 6 0 39 Siagonium quadricorne Kirby, W., 1815 4 6 29 6 1 46 Deleaster dichrous (Gravenhorst, 1802) 2 6 4 5 5 22 Syntomium aeneum (Müller, P.W.J., 1821) 6 23 65 21 1 116 Coprophilus striatulus (Fabricius, 1792) 5 6 9 3 0 23 Aploderus caelatus (Gravenhorst, 1802) 1 4 5 1 1 12 Bledius annae Sharp, 1911 2 3 4 12 1 22 Bledius bicornis (Germar, 1822) 0 1 0 0 0 1 Bledius defensus Fauvel, 1872 0 2 0 10 2 14 Bledius dissimilis Erichson, 1840 10 0 0 0 0 10 Bledius erraticus Erichson, 1839 0 2 0 0 2 4 Bledius femoralis (Gyllenhal, 1827) 0 3 0 0 0 3 Bledius fergussoni Joy, 1912 1 3 0 0 0 4 Bledius fuscipes Rye, 1865 0 0 0 1 0 1 Bledius gallicus (Gravenhorst, 1806) 5 6 6 1 1 19 Bledius limicola Tottenham, 1940 10 0 8 0 0 18 Bledius longulus Erichson, 1839 2 3 0 2 2 9 Bledius occidentalis Bondroit, 1907 1 0 1 0 0 2 Bledius opacus (Block, 1799) 9 4 4 0 1 18 Bledius pallipes (Gravenhorst, 1806) 0 0 0 1 0 1 Bledius praetermissus Williams, 1929 6 0 1 0 0 7 Bledius spectabilis Kraatz, 1857 0 0 1 0 0 1 Bledius subterraneus Erichson, 1839 0 13 3 8 3 27 Bledius terebrans (Schiødte, 1866) 0 1 0 0 0 1 Bledius tricornis (Herbst, 1784) 3 1 0 0 0 4 Carpelimus bilineatus Stephens, 1834 27 13 38 17 2 97 Carpelimus erichsoni (Sharp, 1871) 4 0 2 1 0 7 Carpelimus corticinus (Gravenhorst, 1806) 32 8 108 24 13 185 Carpelimus elongatulus (Erichson, 1839) 24 10 76 31 4 145 Carpelimus foveolatus (Sahlberg, C.R., 1832) 11 0 0 0 0 11 Carpelimus fuliginosus (Gravenhorst, 1802) 1 0 4 0 0 5 Carpelimus gracilis (Mannerheim, 1830) 1 0 3 0 1 5 Carpelimus halophilus (Kiesenwetter, 1844) 2 0 0 1 0 3 Carpelimus impressus (Lacordaire, 1835) 1 1 4 1 0 7 Carpelimus lindrothi Palm, 1942 0 0 1 0 0 1 Carpelimus manchuricus subsp. subtilicornis (Roubal, 1946) 2 0 1 8 1 12 Carpelimus obesus (Kiesenwetter, 1844) 1 0 0 0 0 1 Carpelimus pusillus (Gravenhorst, 1802) 8 8 18 3 1 38 Carpelimus rivularis (Motschulsky, 1860) 34 8 73 19 9 143 Carpelimus zealandicus (Sharp, 1900) 6 2 12 3 0 23

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Yorkshire Coleoptera Checklist 2020

Ochthephilus andalusiacus (Fagel, 1957) 0 1 0 0 1 2 Ochthephilus angustior (Bernhauer, 1943) 0 0 0 1 0 1 Ochthephilus aureus (Fauvel, 1871) 0 3 2 2 1 8 Ochthephilus omalinus (Erichson, 1840) 0 14 1 8 3 26 Thinobius bicolor Joy, 1911 0 2 0 1 0 3 Thinodromus arcuatus (Stephens, 1834) 1 9 1 5 3 19 Anotylus complanatus (Erichson, 1839) 14 20 44 8 4 90 Anotylus fairmairei (Pandellé, 1867) 0 1 3 0 0 4 Anotylus insecatus (Gravenhorst, 1806) 0 1 1 0 0 2 Anotylus inustus (Gravenhorst, 1806) 9 11 62 2 1 85 Anotylus maritimus Thomson, C.G., 1861 3 2 0 0 0 5 Anotylus mutator (Lohse, 1963) 3 5 24 5 0 37 Anotylus nitidulus (Gravenhorst, 1802) 1 8 8 4 1 22 Anotylus rugosus (Fabricius, 1775) 142 61 482 117 33 835 Anotylus saulcyi (Pandellé, 1867) 0 1 0 0 0 1 Anotylus sculpturatus (Gravenhorst, 1806) 51 39 259 79 17 445 Anotylus tetracarinatus (Block, 1799) 40 57 241 61 32 431 Oxytelus fulvipes Erichson, 1839 0 0 6 7 0 13 Oxytelus laqueatus (Marsham, 1802) 19 25 139 42 14 239 Oxytelus migrator Fauvel, 1904 0 0 2 0 0 2 Oxytelus piceus (Linnaeus, 1767) 0 1 1 0 0 2 Oxytelus sculptus Gravenhorst, 1806 6 4 25 3 2 40 Platystethus arenarius (Fourcroy, 1785) 28 29 126 17 13 213 Platystethus cornutus (Gravenhorst, 1802) 35 12 56 15 4 122 Platystethus degener Mulsant & Rey, 1878 1 0 1 0 0 2 Platystethus nitens (Sahlberg, C.R., 1832) 15 2 11 1 0 29 Platystethus nodifrons (Mannerheim, 1830) 28 3 5 2 4 42 Oxyporus rufus (Linnaeus, 1758) 13 6 21 5 0 45 Stenus aceris Stephens, 1833 2 5 11 1 2 21 Stenus argus Gravenhorst, 1806 10 2 0 6 0 18 Stenus asphaltinus Erichson, 1840 0 0 1 0 0 1 Stenus ater Mannerheim, 1830 3 0 4 2 0 9 Stenus atratulus Erichson, 1839 4 5 10 5 0 24 Stenus bifoveolatus Gyllenhal, 1827 16 23 148 39 5 231 Stenus biguttatus (Linnaeus, 1758) 14 11 14 9 6 54 Stenus bimaculatus Gyllenhal, 1810 60 25 127 62 9 283 Stenus binotatus Ljungh, 1804 4 10 34 3 2 53 Stenus boops Ljungh, 1810 53 13 132 33 8 239 Stenus brevipennis Thomson, C.G., 1851 1 9 8 10 12 40 Stenus brunnipes Stephens, 1833 82 27 246 76 21 452 Stenus canaliculatus Gyllenhal, 1827 23 2 40 15 4 84 Stenus carbonarius Gyllenhal, 1827 24 2 23 8 0 57 Stenus cicindeloides (Schaller, 1783) 36 15 100 32 0 183 Stenus clavicornis (Scopoli, 1763) 76 45 309 60 10 500 Stenus comma LeConte, 1863 6 4 9 5 0 24 Stenus crassus Stephens, 1833 4 1 16 5 0 26

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Yorkshire Coleoptera Checklist 2020

Stenus europaeus Puthz, 1966 23 3 12 9 1 48 Stenus flavipes Stephens, 1833 18 30 42 45 6 141 Stenus formicetorum Mannerheim, 1843 1 2 3 0 0 6 Stenus fulvicornis Stephens, 1833 37 19 155 41 6 258 Stenus fuscicornis Erichson, 1840 2 2 19 2 1 26 Stenus fuscipes Gravenhorst, 1802 3 0 2 1 0 6 Stenus geniculatus Gravenhorst, 1806 0 0 1 0 0 1 Stenus guttula Müller, P.W.J., 1821 15 17 10 10 8 60 Stenus guynemeri Jacquelin du Val, 1850 0 5 19 7 10 41 Stenus impressus Germar, 1824 53 72 405 99 28 657 Stenus incrassatus Erichson, 1839 1 0 3 0 0 4 Stenus juno (Paykull, 1789) 116 38 336 93 13 596 Stenus kiesenwetteri Rosenhauer, 1856 0 0 1 0 0 1 Stenus latifrons Erichson, 1839 18 14 60 10 1 103 Stenus lustrator Erichson, 1839 3 2 10 0 0 15 Stenus melanarius Stephens, 1833 4 0 18 4 0 26 Stenus melanopus (Marsham, 1802) 9 0 3 1 0 13 Stenus morio Gravenhorst, 1806 1 1 0 0 0 2 Stenus nanus Stephens, 1833 8 13 63 12 4 100 Stenus nigritulus Gyllenhal, 1827 9 3 7 1 1 21 Stenus nitens Stephens, 1833 3 0 35 2 0 40 Stenus nitidiusculus Stephens, 1833 17 22 131 61 16 247 Stenus niveus Fauvel, 1865 2 0 1 1 0 4 Stenus ochropus Kiesenwetter, 1858 0 0 1 1 0 2 Stenus oscillator Rye, 1870 0 0 8 0 0 8 Stenus ossium Stephens, 1833 17 7 28 6 0 58 Stenus pallipes Gravenhorst, 1802 9 1 15 2 0 27 Stenus pallitarsis Stephens, 1833 47 8 57 20 5 137 Stenus palustris Erichson, 1839 11 4 8 5 0 28 Stenus picipennis Erichson, 1840 8 5 43 8 2 66 Stenus picipes Stephens, 1833 20 13 69 32 7 141 Stenus providus Erichson, 1839 27 28 116 41 9 221 Stenus pubescens Stephens, 1833 11 4 48 12 3 78 Stenus pusillus Stephens, 1833 21 12 88 27 6 154 Stenus similis (Herbst, 1784) 68 67 265 53 7 460 Stenus solutus Erichson, 1840 9 2 38 4 0 53 Stenus subaeneus Erichson, 1840 6 3 12 1 0 22 Stenus tarsalis Ljungh, 1810 27 6 10 6 1 50 Stenus umbratilis Casey, 1884 0 0 0 9 2 11 Dianous coerulescens (Gyllenhal, 1810) 1 18 23 18 19 79 Euaesthetus bipunctatus (Ljungh, 1804) 2 1 11 15 2 31 Euaesthetus laeviusculus Mannerheim, 1844 3 4 27 4 1 39 Euaesthetus ruficapillus Lacordaire, 1835 20 4 47 7 3 81 Pseudopsis sulcata Newman, 1834 1 3 4 3 0 11 Achenium depressum (Gravenhorst, 1802) 1 0 0 0 0 1 Achenium humile (Nicolai, 1822) 1 0 0 0 0 1

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Yorkshire Coleoptera Checklist 2020

Astenus lyonessius (Joy, 1908) 18 1 7 4 0 30 Astenus procerus (Gravenhorst, 1806) 0 0 1 0 0 1 Astenus pulchellus (Heer, 1839) 7 1 13 2 0 23 Lathrobium Gravenhorst, 1802 0 0 2 0 0 2 Lathrobium brunnipes (Fabricius, 1792) 59 40 389 113 32 633 Lathrobium elongatum (Linnaeus, 1767) 9 4 21 10 4 48 Lathrobium fovulum Stephens, 1833 1 1 7 6 0 15 Lathrobium fulvipenne (Gravenhorst, 1806) 39 23 98 26 6 192 Lathrobium geminum Kraatz, 1857 16 9 72 10 5 112 Lathrobium impressum Heer, 1841 7 0 19 14 0 40 Lathrobium longulum Gravenhorst, 1802 9 2 13 1 1 26 Lathrobium pallidipenne Hochhuth, 1851 1 4 2 1 0 8 Lathrobium quadratum (Paykull, 1789) 17 3 26 5 0 51 Lathrobium rufipenne Gyllenhal, 1813 0 0 2 0 0 2 Lathrobium terminatum Gravenhorst, 1802 7 10 109 8 4 138 Lathrobium zetterstedti Rye, 1872 0 1 2 2 3 8 Lithocharis nigriceps Kraatz, 1859 18 3 46 9 1 77 Lithocharis ochracea (Gravenhorst, 1802) 12 2 24 6 2 46 Lobrathium multipunctum (Gravenhorst, 1802) 1 6 5 1 1 14 Medon apicalis (Kraatz, 1857) 1 0 0 0 0 1 Medon brunneus (Erichson, 1839) 0 1 3 0 0 4 Medon castaneus (Gravenhorst, 1802) 0 0 1 0 0 1 Medon fusculus (Mannerheim, 1830) 0 4 0 0 0 4 Medon pocofer (Peyron, 1857) 0 1 0 0 0 1 Ochthephilum fracticorne (Paykull, 1800) 5 9 19 6 1 40 Paederus fuscipes Curtis, 1826 0 0 1 0 0 1 Paederus littoralis Gravenhorst, 1802 1 3 1 1 0 6 Paederus riparius (Linnaeus, 1758) 4 13 4 1 0 22 Pseudomedon obscurellus (Erichson, 1840) 0 0 2 1 0 3 Pseudomedon obsoletus (Nordmann, 1837) 0 0 7 0 0 7 Rugilus erichsonii (Fauvel, 1867) 15 9 48 19 0 91 Rugilus orbiculatus (Paykull, 1789) 48 19 186 33 2 288 Rugilus rufipes Germar, 1836 61 12 121 33 4 231 Rugilus subtilis (Erichson, 1840) 0 1 0 0 0 1 Scopaeus gracilis (Sperk, 1835) 0 0 2 0 0 2 Scopaeus sulcicollis (Stephens, 1833) 0 0 2 0 0 2 Sunius bicolor (Olivier, 1795) 4 0 1 0 0 5 Sunius melanocephalus (Fabricius, 1792) 3 1 0 1 0 5 Sunius propinquus (Brisout de Barneville, 1867) 21 3 27 0 0 51 Bisnius cephalotes (Gravenhorst, 1802) 12 8 58 15 2 95 Bisnius fimetarius (Gravenhorst, 1802) 43 25 319 41 11 439 Bisnius nigriventris (Thomson, C.G., 1867) 4 1 11 3 2 21 Bisnius parcus (Sharp, 1874) 5 0 14 2 0 21 Bisnius puella (Nordmann, 1837) 2 4 35 11 2 54 Bisnius sordidus (Gravenhorst, 1802) 25 5 73 11 0 114 Bisnius subuliformis (Gravenhorst, 1802) 1 4 3 2 0 10

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Yorkshire Coleoptera Checklist 2020

Cafius fucicola Curtis, 1830 1 1 0 0 0 2 Cafius xantholoma (Gravenhorst, 1806) 15 10 0 0 0 25 Erichsonius cinerascens (Gravenhorst, 1802) 2 5 13 0 0 20 Gabrius appendiculatus Sharp, 1910 1 4 11 5 1 22 Gabrius astutoides (Strand, A., 1946) 1 0 0 0 0 1 Gabrius bishopi Sharp, 1910 13 0 4 5 1 23 Gabrius breviventer (Sperk, 1835) 32 24 141 39 3 239 Gabrius nigritulus (Gravenhorst, 1802) 6 7 15 4 0 32 Gabrius osseticus (Kolenati, 1846) 0 1 0 1 0 2 Gabrius piliger Mulsant & Rey, 1876 5 0 44 5 2 56 Gabrius splendidulus (Gravenhorst, 1802) 6 16 88 18 6 134 Gabrius trossulus (Nordmann, 1837) 17 6 33 28 1 85 Gabrius velox Sharp, 1910 0 1 0 0 0 1 Gabronthus thermarum (Aubé, 1850) 1 0 0 0 0 1 Neobisnius lathrobioides (Baudi, 1848) 0 0 1 0 0 1 Neobisnius procerulus (Gravenhorst, 1806) 1 0 1 0 0 2 Neobisnius prolixus (Erichson, 1840) 0 0 1 0 0 1 Neobisnius villosulus (Stephens, 1833) 0 1 1 2 0 4 Philonthus addendus Sharp, 1867 2 5 9 2 2 20 Philonthus albipes (Gravenhorst, 1802) 10 3 28 3 1 45 Philonthus atratus (Gravenhorst, 1802) 4 4 6 0 0 14 Philonthus carbonarius (Gravenhorst, 1802) 45 63 225 47 7 387 Philonthus cognatus Stephens, 1832 64 47 206 63 10 390 Philonthus concinnus (Gravenhorst, 1802) 13 1 25 5 0 44 Philonthus corruscus (Gravenhorst, 1802) 0 0 0 1 0 1 Philonthus corvinus Erichson, 1839 3 2 0 0 0 5 Philonthus cruentatus (Gmelin in Linnaeus, 1790) 4 2 8 2 0 16 Philonthus debilis (Gravenhorst, 1802) 4 2 39 3 0 48 Philonthus decorus (Gravenhorst, 1802) 11 29 210 48 16 314 Philonthus discoideus (Gravenhorst, 1802) 4 5 10 2 0 21 Philonthus ebeninus (Gravenhorst, 1802) 0 4 4 2 0 10 Philonthus fumarius (Gravenhorst, 1806) 0 1 0 6 0 7 Philonthus intermedius (Lacordaire, 1835) 13 11 48 5 2 79 Philonthus jurgans Tottenham, 1937 2 2 7 2 0 13 Philonthus laminatus (Creutzer, 1799) 22 25 119 32 10 208 Philonthus longicornis Stephens, 1832 11 5 9 5 1 31 Philonthus mannerheimi Fauvel, 1869 0 2 0 0 0 2 Philonthus marginatus (Müller, O.F., 1764) 17 18 84 43 8 170 Philonthus micans (Gravenhorst, 1802) 22 17 7 5 2 53 Philonthus micantoides Benick & Lohse, 1956 13 3 28 11 0 55 Philonthus nigrita (Gravenhorst, 1806) 3 12 9 6 1 31 Philonthus parvicornis (Gravenhorst, 1802) 0 0 2 2 0 4 Philonthus politus (Linnaeus, 1758) 15 6 49 9 3 82 Philonthus quisquiliarius (Gyllenhal, 1810) 27 10 54 18 6 115 Philonthus rectangulus Sharp, 1874 4 4 24 8 2 42 Philonthus rotundicollis (Ménétries, 1832) 2 11 26 4 0 43

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Yorkshire Coleoptera Checklist 2020

Philonthus rubripennis Stephens, 1832 0 20 0 2 0 22 Philonthus rufipes (Stephens, 1832) 0 0 2 0 0 2 Philonthus sanguinolentus (Gravenhorst, 1802) 10 14 33 2 0 59 Philonthus spinipes Sharp, 1874 1 0 6 0 0 7 Philonthus splendens (Fabricius, 1792) 15 11 41 14 11 92 Philonthus succicola Thomson, C.G., 1860 17 10 117 24 10 178 Philonthus tenuicornis Mulsant & Rey, 1853 18 8 91 13 2 132 Philonthus umbratilis (Gravenhorst, 1802) 10 3 34 2 0 49 Philonthus varians (Paykull, 1789) 56 36 205 58 19 374 Philonthus ventralis (Gravenhorst, 1802) 1 3 8 3 0 15 Remus sericeus Holme, 1837 0 1 0 0 0 1 Creophilus maxillosus (Linnaeus, 1758) 14 8 30 4 7 63 Dinothenarus pubescens (De Geer, 1774) 3 1 5 3 0 12 Ocypus aeneocephalus (De Geer, 1774) 7 16 48 45 11 127 Ocypus brunnipes (Fabricius, 1781) 30 19 93 16 4 162 Ocypus fortunatarum (Wollaston, 1871) 0 1 2 0 0 3 Ocypus fuscatus (Gravenhorst, 1802) 2 2 8 1 0 13 Ocypus nitens (Schrank, 1781) 1 1 5 4 0 11 Ocypus olens (Müller, O.F., 1764) 41 14 160 33 1 249 Ocypus ophthalmicus (Scopoli, 1763) 0 0 17 0 0 17 Ontholestes murinus (Linnaeus, 1758) 6 0 17 1 0 24 Ontholestes tessellatus (Geoffroy, 1785) 2 2 21 7 1 33 Platydracus fulvipes (Scopoli, 1763) 1 0 1 2 0 4 Platydracus latebricola (Gravenhorst, 1806) 0 3 0 0 0 3 Platydracus stercorarius (Olivier, 1795) 11 8 33 5 5 62 Staphylinus caesareus Cederhjelm, 1798 0 0 1 0 0 1 Staphylinus dimidiaticornis Gemminger, 1851 0 1 0 0 1 2 Staphylinus erythropterus Linnaeus, 1758 3 14 0 1 3 21 Tasgius ater (Gravenhorst, 1802) 27 4 36 5 0 72 Tasgius globulifer (Geoffroy, 1785) 10 7 38 6 3 64 Tasgius melanarius (Heer, 1839) 10 7 63 22 3 105 Tasgius morsitans (Rossi, 1790) 10 4 28 1 2 45 Tasgius pedator (Gravenhorst, 1802) 0 1 0 1 0 2 Tasgius winkleri Bernhauer, 1906 4 4 4 0 0 12 Euryporus picipes (Paykull, 1800) 0 0 0 1 1 2 Heterothops binotatus (Gravenhorst, 1802) 5 2 0 2 0 9 Heterothops dissimilis (Gravenhorst, 1802) 6 3 12 1 0 22 Heterothops minutus Wollaston, 1860 4 0 26 6 0 36 Heterothops niger Kraatz, 1868 3 1 2 2 0 8 Heterothops praevius Erichson, 1839 1 1 4 2 0 8 Quedius auricomus Kiesenwetter, 1850 0 6 6 6 5 23 Quedius boopoides Munster, 1923 4 0 7 2 3 16 Quedius boops (Gravenhorst, 1802) 15 20 53 20 10 118 Quedius brevicornis (Thomson, C.G., 1860) 4 0 0 1 0 5 Quedius brevis Erichson, 1840 0 4 3 0 0 7 Quedius cinctus (Paykull, 1790) 10 3 28 13 0 54

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Yorkshire Coleoptera Checklist 2020

Quedius cruentus (Olivier, 1795) 11 12 72 19 3 117 Quedius curtipennis Bernhauer, 1908 24 32 230 78 16 380 Quedius fulgidus (Fabricius, 1792) 1 2 2 1 0 6 Quedius fuliginosus (Gravenhorst, 1802) 15 11 83 32 7 148 Quedius fulvicollis (Stephens, 1833) 0 5 6 7 1 19 Quedius fumatus (Stephens, 1833) 3 12 51 14 2 82 Quedius humeralis Stephens, 1832 7 0 4 1 0 12 Quedius invreae Gridelli, 1924 0 2 9 2 0 13 Quedius lateralis (Gravenhorst, 1802) 1 6 33 8 1 49 Quedius levicollis (Brullé, 1832) 12 6 56 14 11 99 Quedius longicornis Kraatz, 1857 3 3 3 1 0 10 Quedius maurorufus (Gravenhorst, 1806) 36 12 172 49 8 277 Quedius maurus (Sahlberg, C.R., 1830) 3 10 18 3 0 34 Quedius mesomelinus (Marsham, 1802) 17 12 123 15 4 171 Quedius molochinus (Gravenhorst, 1806) 17 27 126 41 14 225 Quedius nemoralis Baudi, 1848 5 0 24 1 0 30 Quedius nigriceps Kraatz, 1857 1 14 41 4 1 61 Quedius nigrocaeruleus Fauvel, 1874 3 0 0 2 0 5 Quedius nitipennis (Stephens, 1833) 8 8 14 21 7 58 Quedius persimilis Mulsant & Rey, 1876 3 0 32 5 2 42 Quedius picipes (Mannerheim, 1830) 8 11 25 9 1 54 Quedius plagiatus Mannerheim, 1843 0 5 9 11 1 26 Quedius puncticollis (Thomson, C.G., 1867) 1 4 0 7 1 13 Quedius riparius Kellner, 1843 1 1 0 0 0 2 Quedius schatzmayri Gridelli, 1922 5 6 21 7 0 39 Quedius scintillans (Gravenhorst, 1806) 11 2 71 23 3 110 Quedius scitus (Gravenhorst, 1806) 0 1 0 1 0 2 Quedius semiaeneus (Stephens, 1833) 10 5 7 2 1 25 Quedius semiobscurus (Marsham, 1802) 19 9 31 10 0 69 Quedius simplicifrons Fairmaire, 1861 16 0 0 0 0 16 Quedius truncicola Fairmaire & Laboulbène, 1856 0 2 4 1 0 7 Quedius umbrinus Erichson, 1839 5 13 12 19 10 59 Quedius xanthopus Erichson, 1839 0 12 5 1 1 19 Atrecus affinis (Paykull, 1789) 5 24 99 21 6 155 Othius angustus Stephens, 1833 6 5 37 11 6 65 Othius laeviusculus Stephens, 1833 14 3 20 4 3 44 Othius punctulatus (Goeze, 1777) 20 22 159 51 10 262 Othius subuliformis Stephens, 1833 18 34 120 21 12 205 Gauropterus fulgidus (Fabricius, 1787) 2 1 3 3 0 9 Gyrohypnus angustatus Stephens, 1833 20 13 44 19 1 97 Gyrohypnus atratus (Heer, 1839) 0 3 0 0 0 3 Gyrohypnus fracticornis (Müller, O.F., 1776) 26 12 123 21 4 186 Gyrohypnus punctulatus (Paykull, 1789) 1 1 49 8 1 60 Leptacinus batychrus (Gyllenhal, 1827) 0 4 15 0 0 19 Leptacinus formicetorum Märkel, 1841 0 5 8 0 0 13 Leptacinus intermedius Donisthorpe, 1936 1 0 5 2 0 8

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Yorkshire Coleoptera Checklist 2020

Leptacinus pusillus (Stephens, 1833) 13 7 83 12 1 116 Megalinus glabratus (Gravenhorst, 1802) 40 13 36 10 0 99 Nudobius lentus (Gravenhorst, 1806) 2 10 12 8 1 33 Phacophallus parumpunctatus (Gyllenhal, 1827) 6 2 11 1 0 20 Xantholinus elegans (Olivier, 1795) 12 2 20 1 1 36 Xantholinus gallicus Coiffait, 1956 0 3 3 0 1 7 Xantholinus laevigatus Jacobsen, 1849 0 0 1 3 0 4 Xantholinus linearis (Olivier, 1795) 102 59 390 65 20 636 Xantholinus longiventris Heer, 1839 31 27 115 28 0 201 Xantholinus tricolor (Fabricius, 1787) 1 0 0 1 0 2 Eutheia schaumii Kiesenwetter, 1858 0 2 1 0 0 3 Eutheia scydmaenoides Stephens, 1830 0 2 3 1 1 7 Cephennium gallicum Ganglbauer, 1899 2 0 5 0 0 7 Euconnus denticornis (Müller, P.W.J. & Kunze, 1822) 0 2 0 0 0 2 Euconnus duboisi Méquignon, 1929 2 0 1 0 0 3 Euconnus fimetarius (Chaudoir, 1845) 0 1 2 0 0 3 Euconnus hirticollis (Illiger, 1798) 1 2 1 8 0 12 Microscydmus nanus (Schaum, 1844) 0 1 0 1 0 2 Neuraphes angulatus (Müller, P.W.J. & Kunze, 1822) 0 3 1 0 0 4 Neuraphes elongatulus (Müller, P.W.J. & Kunze, 1822) 0 10 34 7 1 52 Neuraphes plicicollis Reitter, 1879 0 0 0 3 0 3 Neuraphes talparum Lokay, 1921 0 4 0 1 0 5 Scydmoraphes helvolus (Schaum, 1844) 4 0 8 1 0 13 Scydmoraphes sparshalli (Denny, 1825) 0 2 0 1 0 3 Stenichnus bicolor (Denny, 1825) 0 2 9 3 0 14 Stenichnus collaris (Müller, P.W.J. & Kunze, 1822) 10 24 54 19 3 110 Stenichnus scutellaris (Müller, P.W.J. & Kunze, 1822) 8 4 11 1 0 24 Scydmaenus tarsatus Müller, P.W.J. & Kunze, 1822 21 10 127 19 2 179

20 Geotrupidae Odonteus armiger (Scopoli, 1772) 0 0 1 0 0 1 Typhaeus typhoeus (Linnaeus, 1758) 14 12 5 1 0 32 Anoplotrupes stercorosus (Scriba, 1791) 0 6 6 0 3 15 Geotrupes mutator (Marsham, 1802) 2 0 0 0 0 2 Geotrupes spiniger (Marsham, 1802) 13 1 13 3 9 39 Geotrupes stercorarius (Linnaeus, 1758) 8 13 46 7 12 86 Trypocopris vernalis (Linnaeus, 1758) 1 5 0 0 1 7

21 Trogidae Trox sabulosus (Linnaeus, 1758) 0 0 0 1 0 1 Trox scaber (Linnaeus, 1767) 0 1 11 2 0 14

22 Lucanidae Sinodendron cylindricum (Linnaeus, 1758) 9 30 128 29 5 201 Lucanus cervus (Linnaeus, 1758) 1 2 3 3 0 9 Dorcus parallelipipedus (Linnaeus, 1758) 31 7 47 11 2 98

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Yorkshire Coleoptera Checklist 2020

23 Scarabaeidae Onthophagus fracticornis (Preyssler, 1790) 0 0 1 0 0 1 Onthophagus joannae Goljan, 1953 0 0 0 1 0 1 Onthophagus medius (Kugelann, 1792) 0 0 1 0 0 1 Onthophagus nuchicornis (Linnaeus, 1758) 0 1 0 0 0 1 Onthophagus similis (Scriba, 1790) 0 0 4 0 0 4 Aegialia arenaria (Fabricius, 1787) 15 8 0 0 0 23 Aegialia insularis Pittino, 2006 1 6 1 4 4 16 Euheptaulacus villosus (Gyllenhal in Schönherr, 1806) 0 1 1 2 0 4 Heptaulacus testudinarius (Fabricius, 1775) 0 2 0 2 0 4 Oxyomus sylvestris (Scopoli, 1763) 0 2 4 0 0 6 Aphodius (Eupleurus) Mulsant, 1842 0 0 1 0 0 1 Aphodius fimetarius/pedellus agg. 7 17 41 6 2 73 Aphodius arenarius (Olivier, 1789) 1 0 4 2 1 8 Aphodius ater (De Geer, 1774) 21 22 74 28 11 156 Aphodius borealis Gyllenhal, 1827 0 5 5 2 2 14 Aphodius coenosus (Panzer, 1798) 0 0 1 0 0 1 Aphodius conspurcatus (Linnaeus, 1758) 0 3 5 6 0 14 Aphodius constans Duftschmid, 1805 0 5 3 6 3 17 Aphodius contaminatus (Herbst, 1783) 13 12 48 11 4 88 Aphodius depressus (Kugelann, 1792) 14 11 37 16 4 82 Aphodius distinctus (Müller, O.F., 1776) 5 4 8 1 0 18 Aphodius erraticus (Linnaeus, 1758) 3 3 0 1 0 7 Aphodius fasciatus (Olivier, 1789) 0 8 3 1 3 15 Aphodius fimetarius (Linnaeus, 1758) 2 0 4 0 3 9 Aphodius foetens (Fabricius, 1787) 5 6 12 5 0 28 Aphodius foetidus (Herbst, 1783) 5 3 0 0 1 9 Aphodius fossor (Linnaeus, 1758) 22 21 48 9 9 109 Aphodius granarius (Linnaeus, 1767) 9 6 11 2 1 29 Aphodius haemorrhoidalis (Linnaeus, 1758) 6 5 11 2 1 25 Aphodius ictericus (Laicharting, 1781) 2 0 0 1 0 3 Aphodius lapponum Gyllenhal, 1808 0 6 23 7 3 39 Aphodius luridus (Fabricius, 1775) 4 5 14 2 0 25 Aphodius merdarius (Fabricius, 1775) 3 4 2 1 0 10 Aphodius nemoralis Erichson, 1848 0 0 1 0 0 1 Aphodius obliteratus Panzer, 1823 1 2 2 7 0 12 Aphodius paykulli Bedel, 1907 1 0 7 0 0 8 Aphodius pedellus (De Geer, 1774) 0 0 1 2 0 3 Aphodius plagiatus (Linnaeus, 1767) 4 0 0 0 0 4 Aphodius porcus (Fabricius, 1792) 0 1 0 1 0 2 Aphodius prodromus (Brahm, 1790) 20 21 72 19 5 137 Aphodius pusillus (Herbst, 1789) 2 2 7 4 2 17 Aphodius rufipes (Linnaeus, 1758) 337 22 117 92 588 1156 Aphodius rufus (Moll, 1782) 18 6 27 4 23 78 Aphodius sordidus (Fabricius, 1775) 0 1 4 0 0 5

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Yorkshire Coleoptera Checklist 2020

Aphodius sphacelatus (Panzer, 1798) 9 19 84 12 7 131 Aphodius sticticus (Panzer, 1798) 1 0 3 3 1 8 Aphodius subterraneus (Linnaeus, 1758) 1 1 3 2 0 7 Aphodius zenkeri Germar, 1813 1 0 5 0 0 6 Psammodius asper (Fabricius, 1775) 0 1 0 0 0 1 Melolontha hippocastani Fabricius, 1801 0 0 1 0 0 1 Melolontha melolontha (Linnaeus, 1758) 6 15 44 56 251 372 Polyphylla fullo (Linnaeus, 1758) 0 0 1 0 0 1 Amphimallon solstitiale (Linnaeus, 1758) 3 4 1 0 0 8 Serica brunnea (Linnaeus, 1758) 13 13 53 36 106 221 Hoplia philanthus (Füessly, 1775) 1 8 20 12 4 45 Anomala dubia (Scopoli, 1763) 19 4 1 0 0 24 Phyllopertha horticola (Linnaeus, 1758) 17 4 26 1 1 49 Gnorimus nobilis (Linnaeus, 1758) 0 0 1 0 0 1 Gnorimus variabilis (Linnaeus, 1758) 0 0 1 0 0 1 Oxythyrea funesta (Poda, 1761) 0 0 2 0 0 2 Cetonia aurata (Linnaeus, 1758) 1 3 1 0 0 5 Protaetia metallica (Herbst, 1782) 0 1 0 0 0 1

25 Calyptomerus dubius (Marsham, 1802) 15 3 22 3 0 43 Clambus armadillo (De Geer, 1774) 8 8 5 4 2 27 Clambus evae Endrödy-Younga, 1960 0 1 0 0 1 2 Clambus gibbulus (LeConte, 1850) 0 0 3 0 0 3 Clambus nigrellus Reitter, 1914 0 0 0 4 1 5 Clambus nigriclavis Stephens, 1832 0 0 0 4 0 4 Clambus pubescens Redtenbacher, 1849 13 4 18 2 1 38 Clambus punctulum (Beck, 1817) 3 3 16 1 0 23 Clambus simsoni Blackburn, 1902 0 0 4 0 0 4

26 Elodes elongata Tournier, 1868 2 0 2 2 1 7 Elodes minuta (Linnaeus, 1767) 4 12 12 12 3 43 Elodes pseudominuta Klausnitzer, 1971 1 14 32 1 3 51 Elodes tricuspis Nyholm, 1985 0 0 0 1 0 1 Odeles marginata (Fabricius, 1798) 2 21 43 7 12 85 Microcara testacea (Linnaeus, 1767) 44 10 106 32 2 194 Cyphon coarctatus Paykull, 1799 68 64 237 81 13 463 Cyphon hilaris Nyholm, 1944 6 12 78 7 2 105 Cyphon laevipennis Tournier, 1868 60 7 113 11 2 193 Cyphon ochraceus Stephens, 1830 22 14 45 12 1 94 Cyphon padi (Linnaeus, 1758) 25 23 146 38 6 238 Cyphon palustris Thomson, C.G., 1855 4 6 24 8 4 46 Cyphon pubescens (Fabricius, 1792) 1 3 3 1 0 8 Cyphon punctipennis Sharp, 1873 1 6 0 8 16 31 Cyphon variabilis (Thunberg, 1787) 32 21 175 36 12 276

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Yorkshire Coleoptera Checklist 2020

Prionocyphon serricornis (Müller, P.W.J., 1821) 0 10 10 2 0 22 Hydrocyphon deflexicollis (Müller, P.W.J., 1821) 1 5 0 1 6 13 Scirtes hemisphaericus (Linnaeus, 1758) 20 13 31 18 6 88

27 Dascillidae Dascillus cervinus (Linnaeus, 1758) 5 12 13 15 6 51

28 Buprestidae Agrilus angustulus (Illiger, 1803) 3 2 1 1 0 7 Agrilus laticornis (Illiger, 1803) 2 7 8 2 0 19 Agrilus viridis (Linnaeus, 1758) 0 2 0 0 0 2 Aphanisticus pusillus (Olivier, 1790) 0 1 0 0 0 1 Trachys minuta (Linnaeus, 1758) 2 0 0 0 0 2 Trachys troglodytes Gyllenhal in Schönherr, 1817 1 1 0 0 0 2

29 Byrrhidae Simplocaria maculosa Erichson, 1847 1 0 0 0 0 1 Simplocaria semistriata (Fabricius, 1794) 11 12 46 14 2 85 Morychus aeneus (Fabricius, 1775) 1 0 0 0 2 3 Cytilus sericeus (Forster, 1771) 3 16 73 16 7 115 Byrrhus arietinus Steffahny, 1842 0 1 6 1 0 8 Byrrhus fasciatus (Forster, 1770) 2 7 18 0 3 30 Byrrhus pilula (Linnaeus, 1758) 8 17 98 27 8 158 Byrrhus pustulatus (Forster, 1770) 3 5 18 1 1 28 Porcinolus murinus (Fabricius, 1794) 1 1 1 0 0 3 Curimopsis maritima (Marsham, 1802) 2 0 0 0 0 2 Curimopsis nigrita (Palm, 1934) 0 0 35 0 0 35 Curimopsis setigera (Illiger, 1798) 1 0 0 0 0 1

30 Elmidae Elmis aenea (Müller, P.W.J., 1806) 121 326 272 377 344 1440 Esolus parallelepipedus (Müller, P.W.J., 1806) 13 98 40 213 187 551 Limnius volckmari (Panzer, 1793) 48 240 157 320 262 1027 Normandia nitens (Müller, P.W.J., 1817) 0 1 0 0 0 1 Oulimnius major (Rey, 1889) 1 0 0 0 0 1 Oulimnius troglodytes (Gyllenhal, 1827) 0 0 0 9 1 10 Oulimnius tuberculatus (Müller, P.W.J., 1806) 15 131 60 193 171 570 Riolus cupreus (Müller, P.W.J., 1806) 0 4 1 11 2 18 Riolus subviolaceus (Müller, P.W.J., 1817) 12 24 9 29 20 94

31 Dryopidae Pomatinus substriatus (Müller, P.W.J., 1806) 1 4 1 0 0 6 Dryops auriculatus (Fourcroy, 1785) 13 2 2 1 2 20 Dryops ernesti des Gozis, 1886 8 25 21 38 11 103 Dryops griseus (Erichson, 1847) 4 0 0 1 0 5 Dryops luridus (Erichson, 1847) 44 17 60 13 11 145

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Yorkshire Coleoptera Checklist 2020

Dryops nitidulus (Heer, 1841) 0 0 1 5 4 10 Dryops similaris Bollow, 1936 4 0 0 0 0 4

32 Limnichidae Limnichus pygmaeus (Sturm, 1807) 0 1 9 0 0 10

33 Heteroceridae Augyles maritimus (Guérin-Méneville, 1844) 7 0 4 1 0 12 Heterocerus fenestratus (Thunberg, 1784) 24 3 7 7 4 45 Heterocerus flexuosus Stephens, 1828 4 1 5 1 0 11 Heterocerus fusculus Kiesenwetter, 1843 0 0 0 1 0 1 Heterocerus marginatus (Fabricius, 1787) 12 9 8 8 0 37

34 Psephenidae Eubria palustris Germar, 1818 1 15 4 10 1 31

36 Eucnemidae Melasis buprestoides (Linnaeus, 1761) 0 2 37 13 0 52 Epiphanis cornutus Eschscholtz, 1829 0 1 0 0 0 1 Microrhagus pygmaeus (Fabricius, 1792) 0 4 0 0 0 4 Eucnemis capucina Ahrens, 1812 0 0 2 0 0 2

37 Throscidae Trixagus carinifrons (de Bonvouloir, 1859) 1 4 19 1 0 25 Trixagus dermestoides (Linnaeus, 1767) 4 19 88 9 3 123 Trixagus obtusus (Curtis, 1827) 0 0 4 0 2 6

38 Elateridae Agrypnus murinus (Linnaeus, 1758) 24 4 4 1 0 33 Hypnoidus riparius (Fabricius, 1792) 18 28 104 35 26 211 Actenicerus sjaelandicus (Müller, O.F., 1764) 1 7 1 8 0 17 Anostirus castaneus (Linnaeus, 1758) 0 0 0 6 0 6 Ctenicera cuprea (Fabricius, 1775) 1 25 120 27 15 188 Ctenicera pectinicornis (Linnaeus, 1758) 5 10 15 7 7 44 Aplotarsus incanus (Gyllenhal, 1827) 30 57 105 26 14 232 Paraphotistus impressus (Fabricius, 1792) 0 2 0 2 0 4 Paraphotistus nigricornis (Panzer, 1799) 25 2 2 1 0 30 Prosternon tessellatum (Linnaeus, 1758) 9 6 36 1 0 52 Selatosomus aeneus (Linnaeus, 1758) 4 2 20 7 1 34 Cidnopus aeruginosus (Olivier, 1790) 12 4 6 3 2 27 Kibunea minuta (Linnaeus, 1758) 29 18 110 7 2 166 Denticollis linearis (Linnaeus, 1758) 10 34 158 25 15 242 Athous bicolor (Goeze, 1777) 13 4 41 4 0 62 Athous campyloides Newman, 1833 0 0 7 2 0 9 Athous haemorrhoidalis (Fabricius, 1801) 94 92 727 77 25 1015 Athous subfuscus (Müller, O.F., 1764) 0 0 2 1 0 3

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Yorkshire Coleoptera Checklist 2020

Athous vittatus (Fabricius, 1792) 4 13 42 8 5 72 Stenagostus rhombeus (Olivier, 1790) 0 0 3 1 0 4 Hemicrepidius hirtus (Herbst, 1784) 29 38 87 22 1 177 Adrastus pallens (Fabricius, 1792) 28 28 106 22 11 195 Agriotes acuminatus (Stephens, 1830) 31 52 202 32 8 325 Agriotes lineatus (Linnaeus, 1767) 31 14 17 5 0 67 Agriotes obscurus (Linnaeus, 1758) 69 56 244 42 17 428 Agriotes pallidulus (Illiger, 1807) 39 103 643 87 35 907 Agriotes sordidus (Illiger, 1807) 0 0 1 0 0 1 Agriotes sputator (Linnaeus, 1758) 35 15 40 8 4 102 Dalopius marginatus (Linnaeus, 1758) 16 99 281 56 15 467 Ampedus balteatus (Linnaeus, 1758) 10 18 63 4 2 97 Ampedus nigerrimus (Lacordaire, 1835) 0 0 2 0 0 2 Ampedus nigrinus (Herbst, 1784) 0 1 2 1 0 4 Ampedus pomorum (Herbst, 1784) 0 4 2 0 0 6 Ischnodes sanguinicollis (Panzer, 1793) 0 1 0 0 0 1 Elater ferrugineus Linnaeus, 1758 1 0 0 0 0 1 Sericus brunneus (Linnaeus, 1758) 2 4 14 5 1 26 Melanotus Eschscholtz, 1829 0 2 5 0 0 7 Melanotus castanipes (Paykull, 1800) 1 9 9 0 0 19 Melanotus villosus (Geoffroy in Fourcroy, 1785) 0 0 2 1 0 3 Fleutiauxellus maritimus (Curtis, 1840) 1 1 0 3 6 11 Oedostethus quadripustulatus (Fabricius, 1792) 13 4 10 1 2 30 Zorochros minimus (Lacordaire, 1835) 0 6 13 20 27 66

40 Lycidae Pyropterus nigroruber (De Geer, 1774) 3 7 32 2 0 44 Platycis minutus (Fabricius, 1787) 0 20 8 6 1 35

41 Lampyridae Lampyris noctiluca (Linnaeus, 1758) 7 14 100 27 6 154

42 Cantharidae Podabrus alpinus (Paykull, 1798) 2 16 36 13 8 75 Ancistronycha abdominalis (Fabricius, 1798) 2 14 19 3 0 38 Cantharis cryptica Ashe, 1947 54 48 396 57 42 597 Cantharis decipiens Baudi, 1871 24 37 244 29 10 344 Cantharis figurata Mannerheim, 1843 4 0 8 6 0 18 Cantharis fusca Linnaeus, 1758 2 0 6 3 1 12 Cantharis lateralis Linnaeus, 1758 48 19 103 12 1 183 Cantharis livida Linnaeus, 1758 33 17 152 14 5 221 Cantharis nigra (De Geer, 1774) 77 41 307 44 7 476 Cantharis nigricans (Müller, O.F., 1776) 52 54 380 46 25 557 Cantharis pallida Goeze, 1777 7 9 37 9 1 63 Cantharis paludosa Fallén, 1807 1 16 25 15 6 63 Cantharis pellucida Fabricius, 1792 15 39 199 18 16 287

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Yorkshire Coleoptera Checklist 2020

Cantharis rufa Linnaeus, 1758 74 28 166 20 4 292 Cantharis rustica Fallén, 1807 36 21 107 16 8 188 Cantharis thoracica (Olivier, 1790) 25 6 34 10 3 78 Rhagonycha elongata (Fallén, 1807) 0 0 2 0 0 2 Rhagonycha fulva (Scopoli, 1763) 286 134 1205 111 80 1816 Rhagonycha lignosa (Müller, O.F., 1764) 11 39 177 18 9 254 Rhagonycha limbata Thomson, C.G., 1864 50 39 316 46 11 462 Rhagonycha lutea (Müller, O.F., 1764) 2 6 0 1 0 9 Rhagonycha testacea (Linnaeus, 1758) 5 32 78 29 12 156 Rhagonycha translucida (Krynicki, 1832) 0 8 4 1 0 13 Crudosilis ruficollis (Fabricius, 1775) 4 0 4 0 0 8 Malthinus flaveolus (Herbst, 1786) 13 27 114 15 2 171 Malthinus frontalis (Marsham, 1802) 2 6 12 8 1 29 Malthinus seriepunctatus Kiesenwetter, 1852 5 6 14 1 0 26 Malthodes crassicornis (Mäklin, 1846) 0 7 0 1 0 8 Malthodes dispar (Germar, 1824) 10 5 6 15 4 40 Malthodes fibulatus Kiesenwetter, 1852 0 4 0 3 0 7 Malthodes flavoguttatus Kiesenwetter, 1852 0 8 5 6 0 19 Malthodes fuscus (Waltl, 1838) 2 9 15 7 0 33 Malthodes guttifer Kiesenwetter, 1852 0 5 5 3 1 14 Malthodes marginatus (Latreille, 1806) 7 27 60 19 11 124 Malthodes maurus (Laporte, 1840) 0 4 1 2 1 8 Malthodes minimus (Linnaeus, 1758) 33 26 54 10 4 127 Malthodes mysticus Kiesenwetter, 1852 0 39 27 13 5 84 Malthodes pumilus (Brébisson, 1835) 7 9 31 7 1 55

43 Laricobius erichsonii Rosenhauer, 1846 0 2 0 0 0 2

44 Dermestidae Dermestes ater De Geer, 1774 0 0 1 0 0 1 Dermestes frischii Kugelann, 1792 0 2 1 0 0 3 Dermestes haemorrhoidalis Küster, 1852 0 0 2 0 0 2 Dermestes lardarius Linnaeus, 1758 4 7 22 2 2 37 Dermestes maculatus De Geer, 1774 3 3 7 3 0 16 Dermestes murinus Linnaeus, 1758 7 2 5 1 0 15 Dermestes peruvianus Laporte, 1840 0 0 4 0 0 4 Dermestes undulatus Brahm, 1790 2 1 0 0 0 3 Attagenus pellio (Linnaeus, 1758) 10 6 35 3 2 56 Attagenus unicolor (Brahm, 1791) 0 0 0 1 0 1 Trogoderma granarium Everts, 1898 0 1 0 0 0 1 Trogoderma inclusum LeConte, 1854 0 0 1 0 0 1 Reesa vespulae (Milliron, 1939) 2 0 0 0 0 2 Megatoma undata (Linnaeus, 1758) 7 7 6 4 0 24 Ctesias serra (Fabricius, 1792) 0 13 6 4 0 23 Anthrenus fuscus Olivier, 1789 8 10 53 6 0 77

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Yorkshire Coleoptera Checklist 2020

Anthrenus museorum (Linnaeus, 1761) 1 3 6 0 0 10 Anthrenus verbasci (Linnaeus, 1767) 7 22 69 5 0 103 Trinodes hirtus (Fabricius, 1781) 0 0 2 0 0 2

45 Bostrichidae Bostrichus capucinus (Linnaeus, 1758) 1 0 2 1 0 4 Rhyzopertha dominica (Fabricius, 1792) 2 2 2 0 0 6 Lyctus brunneus (Stephens, 1830) 0 3 6 0 0 9 Lyctus linearis (Goeze, 1777) 0 3 1 0 0 4 Lyctus planicollis LeConte, 1858 0 0 1 0 0 1

46 Ptinidae Tipnus unicolor (Piller & Mitterpacher, 1783) 4 3 7 2 0 16 Hedobia imperialis (Linnaeus, 1767) 8 10 21 8 0 47 Gibbium aequinoctiale Boieldieu, 1854 0 0 6 0 0 6 Gibbium psylloides (de Czenpinski, 1778) 1 1 7 0 0 9 Stethomezium squamosum Hinton, 1943 0 0 1 0 0 1 Mezium affine Boieldieu, 1856 0 1 1 1 0 3 Trigonogenius globulus Solier, 1849 1 2 1 1 0 5 Niptus hololeucus (Faldermann, 1835) 4 8 15 5 0 32 Pseudeurostus hilleri (Reitter, 1877) 0 0 1 0 0 1 Ptinus clavipes Panzer, 1792 0 0 1 0 0 1 Ptinus fur (Linnaeus, 1758) 12 8 15 1 0 36 Ptinus lichenum Marsham, 1802 0 0 2 0 0 2 Ptinus palliatus Perris, 1847 0 0 2 0 0 2 Ptinus pusillus Sturm, 1837 0 0 1 0 0 1 Ptinus sexpunctatus Panzer, 1792 0 3 6 1 0 10 Ptinus subpilosus Sturm, 1837 0 4 2 0 0 6 Ptinus tectus Boieldieu, 1856 5 16 20 3 0 44 Grynobius planus (Fabricius, 1787) 4 19 10 7 1 41 Dryophilus pusillus (Gyllenhal, 1808) 5 22 18 7 7 59 Ochina ptinoides (Marsham, 1802) 0 2 5 5 0 12 Xestobium rufovillosum (De Geer, 1774) 6 11 9 6 1 33 Ernobius mollis (Linnaeus, 1758) 5 8 13 4 2 32 Ernobius nigrinus (Sturm, 1837) 1 2 0 0 0 3 Stegobium paniceum (Linnaeus, 1758) 2 8 32 7 0 49 Gastrallus immarginatus (Müller, P.W.J., 1821) 0 0 2 0 0 2 Anobium fulvicorne (Sturm, 1837) 9 9 36 9 0 63 Anobium inexspectatum Lohse, 1954 1 0 4 1 1 7 Anobium punctatum (De Geer, 1774) 22 15 45 15 3 100 Hadrobregmus denticollis (Creutzer in Panzer, 1796) 0 0 2 0 0 2 Ptilinus pectinicornis (Linnaeus, 1758) 19 42 103 22 3 189 Lasioderma serricorne (Fabricius, 1792) 0 0 2 0 0 2 Dorcatoma chrysomelina (Sturm, 1837) 1 5 5 9 0 20 Dorcatoma dresdensis Herbst, 1792 4 3 2 0 0 9 Dorcatoma flavicornis (Fabricius, 1792) 2 6 5 5 0 18

42

Yorkshire Coleoptera Checklist 2020

Dorcatoma substriata Hummel, 1829 0 4 1 0 0 5 Caenocara bovistae (Hoffmann, J., 1803) 0 0 2 0 0 2 Anitys rubens (Hoffmann, J., 1803) 0 1 3 3 0 7

47 Hylecoetus dermestoides (Linnaeus, 1761) 8 19 38 12 2 79

48 Phloiophilidae Phloiophilus edwardsii Stephens, 1830 1 5 0 1 0 7

49 Trogossitidae Thymalus limbatus (Fabricius, 1787) 0 5 4 6 0 15 Lophocateres pusillus (Klug, 1832) 0 0 1 0 0 1 Tenebroides mauritanicus (Linnaeus, 1758) 1 2 4 0 0 7 Nemozoma elongatum (Linnaeus, 1761) 1 0 1 0 0 2

51 Cleridae Tillus elongatus (Linnaeus, 1758) 0 3 0 0 0 3 Opilo mollis (Linnaeus, 1758) 0 1 0 0 0 1 Thanasimus formicarius (Linnaeus, 1758) 10 9 11 4 0 34 Tarsostenus univittatus (Rossi, 1792) 0 0 1 0 0 1 Korynetes caeruleus (De Geer, 1775) 2 1 2 1 0 6 Necrobia ruficollis (Fabricius, 1775) 3 3 4 2 0 12 Necrobia rufipes (De Geer, 1775) 8 6 11 1 0 26 Necrobia violacea (Linnaeus, 1758) 15 4 17 5 2 43

52 Melyridae Aplocnemus impressus (Marsham, 1802) 0 1 1 0 0 2 Aplocnemus nigricornis (Fabricius, 1792) 0 6 0 1 0 7 Dasytes aeratus Stephens, 1830 6 21 12 11 3 53 Dasytes niger (Linnaeus, 1761) 0 0 2 0 0 2 Dasytes plumbeus (Müller, O.F., 1776) 0 2 7 0 0 9 Dasytes virens (Marsham, 1802) 0 0 2 0 0 2 Psilothrix viridicoeruleus (Geoffroy in Fourcroy, 1785) 0 0 1 0 0 1 Dolichosoma lineare (Rossi, 1794) 5 0 0 0 0 5 Ebaeus pedicularius (Linnaeus, 1758) 0 0 1 0 0 1 Cordylepherus viridis (Fabricius, 1787) 1 0 1 3 0 5 Malachius bipustulatus (Linnaeus, 1758) 62 44 299 43 10 458 Anthocomus fasciatus (Linnaeus, 1758) 0 1 0 0 0 1 Anthocomus rufus (Herbst, 1784) 12 0 93 1 0 106

53 Byturidae Byturus ochraceus (Scriba, 1790) 10 9 56 9 2 86 Byturus tomentosus (De Geer, 1774) 38 70 368 33 13 522

54 Sphindidae

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Yorkshire Coleoptera Checklist 2020

Sphindus dubius (Gyllenhal, 1808) 0 7 1 0 0 8 Aspidiphorus orbiculatus (Gyllenhal, 1808) 2 12 19 2 0 35

55 Biphyllidae Biphyllus lunatus (Fabricius, 1787) 14 8 26 8 1 57 Diplocoelus fagi (Chevrolat in Guérin-Méneville, 1837) 0 0 0 1 0 1

56 Erotylidae Dacne bipustulata (Thunberg, 1781) 11 27 50 24 2 114 Dacne rufifrons (Fabricius, 1775) 4 11 9 8 2 34 Triplax aenea (Schaller, 1783) 2 13 80 18 7 120 Triplax russica (Linnaeus, 1758) 2 2 21 0 0 25 Triplax scutellaris Charpentier, 1825 0 2 0 0 0 2 Tritoma bipustulata Fabricius, 1775 0 2 4 0 0 6

57 Monotomidae Rhizophagus bipustulatus (Fabricius, 1792) 10 18 79 25 2 134 Rhizophagus cribratus Gyllenhal, 1827 0 0 5 4 0 9 Rhizophagus depressus (Fabricius, 1792) 1 3 11 3 0 18 Rhizophagus dispar (Paykull, 1800) 12 34 199 35 2 282 Rhizophagus fenestralis (Linnaeus, 1758) 0 0 3 0 0 3 Rhizophagus ferrugineus (Paykull, 1800) 2 7 5 2 0 16 Rhizophagus nitidulus (Fabricius, 1798) 0 3 28 6 0 37 Rhizophagus oblongicollis Blatch & Horner, 1892 0 1 0 0 0 1 Rhizophagus parallelocollis Gyllenhal, 1827 1 4 5 0 0 10 Rhizophagus perforatus Erichson, 1845 3 4 32 8 0 47 Rhizophagus picipes (Olivier, 1790) 1 0 8 3 0 12 Cyanostolus aeneus (Richter, 1820) 0 1 0 0 0 1 Monotoma angusticollis (Gyllenhal, 1827) 0 3 0 0 0 3 Monotoma bicolor Villa & Villa, 1835 5 2 32 6 1 46 Monotoma brevicollis Aubé, 1838 0 0 1 0 0 1 Monotoma conicicollis Guérin-Méneville, 1837 0 4 7 0 0 11 Monotoma longicollis (Gyllenhal, 1827) 8 7 13 4 0 32 Monotoma picipes Herbst, 1793 13 8 37 9 2 69 Monotoma quadrifoveolata Aubé, 1838 0 1 0 0 0 1 Monotoma spinicollis Aubé, 1838 3 0 4 3 0 10 Monotoma testacea Motschulsky, 1845 0 0 6 2 0 8

58 Cryptophagidae Cryptophagus acutangulus Gyllenhal, 1827 7 5 18 3 0 33 Cryptophagus badius Sturm, 1845 0 1 0 4 0 5 Cryptophagus cellaris (Scopoli, 1763) 1 4 6 1 0 12 Cryptophagus corticinus Thomson, C.G., 1863 0 0 1 0 0 1 Cryptophagus dentatus (Herbst, 1793) 10 10 64 12 4 100 Cryptophagus denticulatus Heer, 1841 4 4 22 9 3 42 Cryptophagus distinguendus Sturm, 1845 9 1 19 7 1 37

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Yorkshire Coleoptera Checklist 2020

Cryptophagus immixtus Rey, 1889 1 3 2 2 0 8 Cryptophagus labilis Erichson, 1846 0 0 1 0 0 1 Cryptophagus lapponicus Gyllenhal, 1827 0 1 0 0 0 1 Cryptophagus laticollis Lucas, 1846 7 6 21 2 4 40 Cryptophagus lycoperdi (Scopoli, 1763) 5 1 33 5 0 44 Cryptophagus obsoletus Reitter, 1879 1 0 0 0 0 1 Cryptophagus pallidus Sturm, 1845 4 4 4 1 0 13 Cryptophagus pilosus Gyllenhal, 1827 11 4 56 7 0 78 Cryptophagus populi Paykull, 1800 0 1 4 1 0 6 Cryptophagus pubescens Sturm, 1845 3 6 13 4 2 28 Cryptophagus ruficornis Stephens, 1830 5 0 13 1 0 19 Cryptophagus saginatus Sturm, 1845 2 1 4 0 0 7 Cryptophagus scanicus (Linnaeus, 1758) 3 16 24 8 2 53 Cryptophagus schmidtii Leunis in Sturm, 1845 0 2 4 0 0 6 Cryptophagus scutellatus Newman, 1834 5 4 22 3 0 34 Cryptophagus setulosus Sturm, 1845 8 15 19 5 4 51 Cryptophagus subdepressus Gyllenhal, 1827 0 1 0 0 0 1 Telmatophilus caricis (Olivier, 1790) 6 7 52 7 0 72 Telmatophilus schoenherrii (Gyllenhal, 1808) 0 0 3 0 0 3 Telmatophilus typhae (Fallén, 1802) 23 6 96 12 3 140 Paramecosoma melanocephalum (Herbst, 1793) 0 6 9 11 2 28 Henoticus serratus (Gyllenhal, 1808) 2 7 14 3 0 26 Micrambe pilosula (Erichson, 1846) 7 0 1 0 0 8 Micrambe ulicis (Stephens, 1830) 51 22 106 5 16 200 Micrambe abietis (Paykull, 1798) 1 0 0 0 0 1 Antherophagus pallens (Linnaeus, 1758) 4 8 42 4 1 59 Antherophagus similis Curtis, 1835 3 6 36 8 1 54 Caenoscelis ferruginea (Sahlberg, C.R., 1820) 0 4 4 0 0 8 Caenoscelis sibirica Reitter, 1889 0 0 1 0 0 1 Caenoscelis subdeplanata Brisout de Barneville, 1882 2 0 19 1 0 22 Atomaria apicalis Erichson, 1846 11 6 31 4 0 52 Atomaria atra (Herbst, 1793) 0 1 0 0 0 1 Atomaria atricapilla Stephens, 1830 29 21 138 16 6 210 Atomaria barani Brisout de Barneville, 1863 1 0 1 2 0 4 Atomaria basalis Erichson, 1846 29 1 6 1 0 37 Atomaria clavigera Ganglbauer, 1899 1 0 0 0 1 2 Atomaria diluta Erichson, 1846 0 0 2 5 0 7 Atomaria fimetarii (Fabricius, 1792) 2 1 6 2 0 11 Atomaria fuscata (Schönherr, 1808) 70 33 166 36 6 311 Atomaria fuscipes (Gyllenhal, 1808) 4 5 2 2 1 14 Atomaria hislopi Wollaston, 1857 0 1 0 0 0 1 Atomaria impressa Erichson, 1846 0 1 0 1 1 3 Atomaria lewisi Reitter, 1877 77 31 304 46 10 468 Atomaria linearis Stephens, 1830 23 18 59 13 7 120 Atomaria lohsei Johnson & Strand, A., 1968 0 1 2 0 0 3 Atomaria mesomela (Herbst, 1792) 18 2 39 11 3 73

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Yorkshire Coleoptera Checklist 2020

Atomaria morio Kolenati, 1846 2 0 0 0 0 2 Atomaria munda Erichson, 1846 1 1 8 3 0 13 Atomaria nigripennis (Kugelann, 1794) 0 0 2 0 0 2 Atomaria nigrirostris Stephens, 1830 9 7 64 16 3 99 Atomaria nigriventris Stephens, 1830 0 2 3 1 0 6 Atomaria nitidula (Marsham, 1802) 86 33 339 39 13 510 Atomaria peltata Kraatz, 1853 0 1 0 0 0 1 Atomaria pulchra Erichson, 1846 0 5 5 1 0 11 Atomaria puncticollis Thomson, C.G., 1868 1 0 0 0 0 1 Atomaria punctithorax Reitter, 1888 2 0 1 1 0 4 Atomaria pusilla (Paykull, 1798) 3 7 7 4 2 23 Atomaria rhenonum Kraatz, 1853 12 0 0 0 0 12 Atomaria rubella Heer, 1841 22 7 55 11 1 96 Atomaria scutellaris Motschulsky, 1849 0 0 3 0 0 3 Atomaria strandi Johnson, 1967 0 1 4 0 0 5 Atomaria testacea Stephens, 1830 91 23 226 39 19 398 Atomaria umbrina (Gyllenhal, 1827) 0 1 0 1 0 2 Ootypus globosus (Waltl, 1838) 5 5 9 4 0 23 Ephistemus globulus (Paykull, 1798) 46 13 155 20 7 241

59 Silvanidae Uleiota planatus (Linnaeus, 1761) 6 0 0 0 0 6 Cryptamorpha desjardinsii (Guérin-Méneville, 1844) 0 0 1 0 0 1 Psammoecus bipunctatus (Fabricius, 1792) 5 0 14 0 0 19 Nausibius clavicornis (Kugelann, 1794) 0 0 1 0 0 1 Ahasverus advena (Waltl, 1834) 8 2 19 3 0 32 Oryzaephilus mercator (Fauvel, 1889) 1 0 1 0 0 2 Oryzaephilus surinamensis (Linnaeus, 1758) 2 0 4 0 0 6 Silvanus bidentatus (Fabricius, 1792) 0 0 3 0 0 3 Silvanus unidentatus (Olivier, 1790) 0 2 2 0 0 4

60 Cucujidae Pediacus depressus (Herbst, 1797) 0 0 1 0 0 1 Pediacus dermestoides (Fabricius, 1792) 0 12 5 2 1 20

61 Phalacridae Phalacrus caricis Sturm, 1807 6 4 10 1 0 21 Phalacrus corruscus (Panzer, 1796) 1 0 4 0 0 5 Phalacrus fimetarius (Fabricius, 1775) 27 2 19 3 0 51 Phalacrus substriatus Gyllenhal, 1813 0 1 2 0 0 3 Olibrus aeneus (Fabricius, 1792) 44 7 107 10 1 169 Olibrus affinis (Sturm, 1807) 3 0 1 0 0 4 Olibrus corticalis (Panzer, 1796) 9 0 7 0 0 16 Olibrus liquidus Erichson, 1845 4 0 10 1 0 15 Olibrus pygmaeus (Sturm, 1807) 1 0 0 0 0 1 Stilbus oblongus (Erichson, 1845) 6 0 14 1 0 21

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Yorkshire Coleoptera Checklist 2020

Stilbus testaceus (Panzer, 1796) 126 46 196 28 9 405

62 Laemophloeidae Cryptolestes ferrugineus (Stephens, 1831) 3 5 18 3 0 29 Cryptolestes pusillus (Schönherr, 1817) 0 0 2 0 0 2

63 Kateretidae Kateretes pedicularius (Linnaeus, 1758) 1 3 6 7 0 17 Kateretes pusillus (Thunberg, 1794) 18 12 2 6 2 40 Kateretes rufilabris (Latreille, 1807) 28 14 39 15 7 103 Brachypterus glaber (Stephens, 1832) 73 36 329 38 13 489 Brachypterus urticae (Fabricius, 1792) 79 45 398 50 21 593 Brachypterolus linariae (Stephens, 1830) 0 0 5 0 0 5 Brachypterolus pulicarius (Linnaeus, 1758) 4 0 30 2 1 37 Brachypterolus vestitus (Kiesenwetter, 1850) 0 0 2 0 0 2

64 Nitidulidae Carpophilus dimidiatus (Fabricius, 1792) 1 0 3 0 0 4 Carpophilus hemipterus (Linnaeus, 1758) 2 1 4 2 0 9 Carpophilus ligneus Murray, 1864 0 1 4 1 0 6 Carpophilus marginellus Motschulsky, 1858 8 0 30 1 1 40 Carpophilus mutilatus Erichson, 1843 0 0 2 0 0 2 Carpophilus obsoletus Erichson, 1843 0 0 2 0 0 2 Carpophilus sexpustulatus (Fabricius, 1792) 1 2 16 5 0 24 Epuraea aestiva (Linnaeus, 1758) 42 58 190 32 13 335 Epuraea angustula Sturm, 1844 0 2 3 1 0 6 Epuraea biguttata (Thunberg, 1784) 15 33 115 19 7 189 Epuraea fuscicollis (Stephens, 1835) 0 1 0 1 0 2 Epuraea guttata (Olivier, 1811) 0 0 3 0 0 3 Epuraea limbata (Fabricius, 1787) 2 1 0 0 0 3 Epuraea longula Erichson, 1845 0 1 0 1 0 2 Epuraea marseuli Reitter, 1872 3 12 13 3 0 31 Epuraea melanocephala (Marsham, 1802) 19 22 159 18 7 225 Epuraea melina Erichson, 1843 4 7 12 1 0 24 Epuraea pallescens (Stephens, 1835) 9 5 19 2 0 35 Epuraea rufomarginata (Stephens, 1830) 0 3 11 0 0 14 Epuraea silacea (Herbst, 1783) 3 13 13 8 1 38 Epuraea terminalis Mannerheim, 1843 0 2 0 0 0 2 Epuraea thoracica Tournier, 1872 0 3 0 1 0 4 Epuraea variegata (Herbst, 1793) 0 1 0 0 0 1 Pria dulcamarae (Scopoli, 1763) 10 1 25 6 0 42 Meligethes erythropus (Marsham, 1802) 0 0 1 0 0 1 Meligethes aeneus (Fabricius, 1775) 245 169 1330 135 57 1936 Meligethes atramentarius Förster, 1849 0 1 0 1 0 2 Meligethes atratus (Olivier, 1790) 7 15 78 13 1 114 Meligethes bidens Brisout de Barneville, 1863 1 0 0 0 0 1

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Yorkshire Coleoptera Checklist 2020

Meligethes brevis Sturm, 1845 6 4 0 1 0 11 Meligethes brunnicornis Sturm, 1845 3 7 10 2 0 22 Meligethes carinulatus Förster, 1849 23 11 53 2 2 91 Meligethes difficilis (Heer, 1841) 15 3 33 1 0 52 Meligethes flavimanus Stephens, 1830 5 10 9 4 2 30 Meligethes fulvipes Brisout de Barneville, 1863 5 3 1 1 0 10 Meligethes gagathinus Erichson, 1845 0 1 0 0 0 1 Meligethes incanus Sturm, 1845 0 0 1 0 0 1 Meligethes kunzei Erichson, 1845 0 0 0 1 0 1 Meligethes lugubris Sturm, 1845 2 0 1 0 0 3 Meligethes morosus Erichson, 1845 11 4 11 1 0 27 Meligethes nigrescens Stephens, 1830 47 16 92 13 2 170 Meligethes obscurus Erichson, 1845 3 7 9 4 0 23 Meligethes ochropus Sturm, 1845 0 0 2 0 0 2 Meligethes ovatus Sturm, 1845 5 2 7 0 0 14 Meligethes pedicularius (Gyllenhal, 1808) 7 2 17 2 2 30 Meligethes planiusculus (Heer, 1841) 0 2 6 1 0 9 Meligethes rotundicollis Brisout de Barneville, 1863 0 0 1 0 0 1 Meligethes ruficornis (Marsham, 1802) 1 0 0 0 0 1 Meligethes solidus (Kugelann, 1794) 0 0 1 3 0 4 Meligethes subrugosus (Gyllenhal, 1808) 7 0 0 0 0 7 Meligethes viridescens (Fabricius, 1787) 4 15 47 6 2 74 Nitidula bipunctata (Linnaeus, 1758) 4 5 10 5 2 26 Nitidula flavomaculata Rossi, 1790 0 1 0 0 0 1 Nitidula rufipes (Linnaeus, 1767) 1 0 6 3 0 10 Omosita colon (Linnaeus, 1758) 13 1 15 6 1 36 Omosita depressa (Linnaeus, 1758) 0 3 12 3 0 18 Omosita discoidea (Fabricius, 1775) 11 9 36 13 1 70 Soronia grisea (Linnaeus, 1758) 7 8 27 9 1 52 Soronia punctatissima (Illiger, 1794) 0 4 7 6 0 17 Cychramus luteus (Fabricius, 1787) 1 11 13 2 0 27 Pocadius adustus Reitter, 1888 1 4 8 0 0 13 Pocadius ferrugineus (Fabricius, 1775) 3 3 16 2 1 25 Thalycra fervida (Olivier, 1790) 0 0 1 2 0 3 Cryptarcha strigata (Fabricius, 1787) 0 0 3 2 0 5 Cryptarcha undata (Olivier, 1790) 0 0 2 3 0 5 Glischrochilus hortensis (Geoffroy in Fourcroy, 1785) 43 30 132 43 3 251 Glischrochilus quadriguttatus (Fabricius, 1777) 1 4 6 3 0 14 Glischrochilus quadripunctatus (Linnaeus, 1758) 4 8 9 3 1 25 Pityophagus ferrugineus (Linnaeus, 1761) 0 2 1 0 0 3

65 Bothrideridae Teredus cylindricus (Olivier, 1790) 0 0 3 0 0 3 Anommatus duodecimstriatus (Müller, P.W.J., 1821) 4 1 5 0 0 10

66 Cerylonidae

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Yorkshire Coleoptera Checklist 2020

Cerylon fagi Brisout de Barneville, 1867 1 0 0 1 0 2 Cerylon ferrugineum Stephens, 1830 1 20 85 21 4 131 Cerylon histeroides (Fabricius, 1792) 4 4 33 15 4 60

67 Sphaerosoma pilosum (Panzer, 1793) 0 0 1 0 0 1

68 Endomychidae Endomychus coccineus (Linnaeus, 1758) 4 13 31 11 1 60 Symbiotes latus Redtenbacher, 1849 0 0 1 0 0 1 Mycetaea subterranea (Fabricius, 1801) 28 5 43 3 1 80

69 Coccinellidae Coccidula rufa (Herbst, 1783) 133 57 426 63 17 696 Coccidula scutellata (Herbst, 1783) 12 2 44 2 0 60 Rhyzobius litura (Fabricius, 1787) 92 56 300 35 15 498 Rhyzobius lophanthae (Blaisdell, 1892) 2 0 8 0 0 10 Clitostethus arcuatus (Rossi, 1794) 0 0 4 0 0 4 Stethorus punctillum (Weise, 1891) 1 1 13 0 0 15 Nephus redtenbacheri (Mulsant, 1846) 15 10 26 7 1 59 Scymnus auritus Thunberg, 1795 8 4 19 4 0 35 Scymnus femoralis (Gyllenhal, 1827) 3 2 4 0 0 9 Scymnus frontalis (Fabricius, 1787) 2 1 9 0 0 12 Scymnus haemorrhoidalis Herbst, 1797 3 4 18 2 0 27 Scymnus interruptus (Goeze, 1777) 0 0 1 0 0 1 Scymnus limbatus Stephens, 1832 2 0 5 1 0 8 Scymnus nigrinus Kugelann, 1794 1 2 1 2 0 6 Scymnus schmidti Fürsch, 1958 3 0 1 0 0 4 Scymnus suturalis Thunberg, 1795 13 20 24 4 2 63 Cryptolaemus montrouzieri Mulsant, 1853 1 0 0 0 0 1 Hyperaspis pseudopustulata Mulsant, 1853 2 4 6 0 0 12 Chilocorus bipustulatus (Linnaeus, 1758) 5 23 38 1 0 67 Chilocorus renipustulatus (Scriba, 1791) 31 32 106 24 3 196 Exochomus nigromaculatus (Goeze, 1777) 0 0 2 0 0 2 Exochomus quadripustulatus (Linnaeus, 1758) 24 17 188 17 2 248 Anisosticta novemdecimpunctata (Linnaeus, 1758) 61 10 319 43 7 440 Tytthaspis sedecimpunctata (Linnaeus, 1761) 17 1 67 3 0 88 Myzia oblongoguttata (Linnaeus, 1758) 4 15 12 8 2 41 Myrrha octodecimguttata (Linnaeus, 1758) 13 13 19 10 0 55 Propylea quattuordecimpunctata (Linnaeus, 1758) 174 128 1384 108 40 1834 Calvia quattuordecimguttata (Linnaeus, 1758) 42 67 371 71 37 588 Halyzia sedecimguttata (Linnaeus, 1758) 15 15 106 26 28 190 Psyllobora vigintiduopunctata (Linnaeus, 1758) 92 34 509 41 8 684 Anatis ocellata (Linnaeus, 1758) 7 28 74 14 5 128 Aphidecta obliterata (Linnaeus, 1758) 11 41 52 16 16 136 Hippodamia tredecimpunctata (Linnaeus, 1758) 1 2 3 0 0 6

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Yorkshire Coleoptera Checklist 2020

Hippodamia variegata (Goeze, 1777) 31 7 142 14 5 199 Coccinella hieroglyphica Linnaeus, 1758 6 20 67 4 2 99 Coccinella magnifica Redtenbacher, 1843 0 0 4 0 0 4 Coccinella quinquepunctata Linnaeus, 1758 0 2 0 0 0 2 Coccinella septempunctata Linnaeus, 1758 349 150 1965 128 71 2663 Coccinella undecimpunctata Linnaeus, 1758 53 10 161 8 9 241 Adalia bipunctata (Linnaeus, 1758) 88 58 886 71 29 1132 Adalia decempunctata (Linnaeus, 1758) 58 60 433 55 24 630 Harmonia axyridis (Pallas, 1773) 85 19 511 41 56 712 Harmonia quadripunctata (Pontoppidan, 1763) 4 1 39 2 0 46 Subcoccinella vigintiquattuorpunctata (Linnaeus, 1758) 58 10 38 1 0 107

70 Corylophidae Sericoderus brevicornis Matthews, A., 1890 0 0 1 0 0 1 Sericoderus lateralis (Gyllenhal, 1827) 11 1 31 2 2 47 Corylophus cassidoides (Marsham, 1802) 4 0 3 0 0 7 Corylophus sublaevipennis Jacquelin du Val, 1859 6 2 0 0 0 8 Orthoperus aequalis Sharp, 1885 0 0 1 0 0 1 Orthoperus atomus (Gyllenhal, 1808) 0 0 3 1 0 4 Orthoperus brunnipes (Gyllenhal, 1808) 9 2 0 0 0 11 Orthoperus corticalis (Redtenbacher, 1849) 1 1 4 0 0 6 Orthoperus nigrescens Stephens, 1829 1 0 2 0 0 3

71 Latridiidae Lithostygnus serripennis Broun, 1914 14 0 14 1 0 29 Stephostethus angusticollis (Gyllenhal, 1827) 1 0 2 0 0 3 Stephostethus lardarius (De Geer, 1775) 76 40 194 41 5 356 Cartodere bifasciata (Reitter, 1877) 133 48 496 52 10 739 Cartodere constricta (Gyllenhal, 1827) 0 0 3 0 0 3 Cartodere nodifer (Westwood, 1839) 84 48 399 49 10 590 Cartodere norvegica (Strand, 1940) 0 0 1 0 0 1 Latridius assimilis Mannerheim, 1844 3 2 17 3 1 26 Latridius porcatus Herbst, 1793 17 1 10 2 1 31 Latridius consimilis Mannerheim, 1844 1 0 1 3 1 6 Latridius minutus (Linnaeus, 1767) 1 4 19 1 1 26 Thes bergrothi (Reitter, 1880) 0 1 2 2 0 5 Enicmus brevicornis (Mannerheim, 1844) 3 0 4 0 1 8 Enicmus fungicola Thomson, C.G., 1868 1 0 13 2 0 16 Enicmus histrio Joy & Tomlin, 1910 9 8 13 5 2 37 Enicmus rugosus (Herbst, 1793) 0 4 2 1 0 7 Enicmus testaceus (Stephens, 1830) 2 1 9 2 0 14 Enicmus transversus (Olivier, 1790) 79 26 219 38 13 375 Dienerella elongata (Curtis, 1830) 0 1 6 1 0 8 Dienerella filiformis (Gyllenhal, 1827) 0 1 2 0 0 3 Dienerella filum (Aubé, 1850) 1 0 3 0 0 4 Dienerella ruficollis (Marsham, 1802) 8 1 6 1 0 16

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Yorkshire Coleoptera Checklist 2020

Adistemia watsoni (Wollaston, 1871) 0 13 0 0 0 13 Corticaria crenulata (Gyllenhal, 1827) 19 3 11 0 0 33 Corticaria dubia Dajoz, 1970 0 0 1 0 0 1 Corticaria elongata (Gyllenhal, 1827) 12 10 35 5 0 62 Corticaria fagi Wollaston, 1854 0 0 1 0 0 1 Corticaria ferruginea Marsham, 1802 1 1 3 0 0 5 Corticaria fulva (Comolli, 1837) 1 1 5 1 0 8 Corticaria impressa (Olivier, 1790) 85 11 149 15 0 260 Corticaria inconspicua Wollaston, 1860 0 0 2 0 0 2 Corticaria punctulata Marsham, 1802 13 2 34 3 0 52 Corticaria rubripes Mannerheim, 1844 0 1 2 0 0 3 Corticaria serrata (Paykull, 1798) 1 1 0 0 0 2 Corticaria umbilicata (Beck, 1817) 1 6 1 1 0 9 Corticarina minuta (Fabricius, 1792) 91 66 304 62 17 540 Corticarina similata (Gyllenhal, 1827) 0 4 1 1 0 6 Cortinicara gibbosa (Herbst, 1793) 122 81 436 41 8 688 Melanophthalma transversalis (Gyllenhal, 1827) 5 0 0 0 0 5

72 Mycetophagidae Pseudotriphyllus suturalis (Fabricius, 1801) 12 23 23 14 4 76 Triphyllus bicolor (Fabricius, 1777) 0 13 3 1 0 17 Litargus balteatus LeConte, 1856 1 0 0 0 0 1 Litargus connexus (Geoffroy in Fourcroy, 1785) 8 6 31 17 1 63 Mycetophagus atomarius (Fabricius, 1787) 3 3 24 11 4 45 Mycetophagus multipunctatus Fabricius, 1792 6 0 15 4 0 25 Mycetophagus piceus (Fabricius, 1777) 4 12 18 6 1 41 Mycetophagus populi Fabricius, 1798 0 0 0 2 0 2 Mycetophagus quadriguttatus Müller, P.W.J., 1821 0 0 1 0 0 1 Mycetophagus quadripustulatus (Linnaeus, 1761) 8 22 79 24 7 140 Typhaea stercorea (Linnaeus, 1758) 17 11 51 11 2 92

73 Ciidae Cis bidentatus (Olivier, 1790) 6 11 16 19 3 55 Cis bilamellatus Wood, 1884 23 16 92 30 0 161 Cis boleti (Scopoli, 1763) 15 22 173 39 9 258 Cis fagi Waltl, 1839 2 1 3 1 0 7 Cis festivus (Panzer, 1793) 0 5 7 3 1 16 Cis hispidus (Paykull, 1798) 3 0 1 0 0 4 Cis lineatocribratus Mellié, 1849 0 0 2 0 0 2 Cis micans (Fabricius, 1792) 0 0 2 0 0 2 Cis nitidus (Fabricius, 1792) 6 13 30 7 0 56 Cis pygmaeus (Marsham, 1802) 0 1 2 1 0 4 Cis vestitus (Mellié, 1849) 0 12 3 3 1 19 Cis villosulus (Marsham, 1802) 1 0 4 1 0 6 Ennearthron cornutum (Gyllenhal, 1827) 2 1 7 1 0 11 Orthocis alni (Gyllenhal, 1813) 1 7 10 3 1 22

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Yorkshire Coleoptera Checklist 2020

Sulcacis affinis (Gyllenhal, 1827) 0 4 4 0 0 8 Octotemnus glabriculus (Gyllenhal, 1827) 7 11 116 30 5 169

74 Tetratomidae Hallomenus binotatus (Quensel, 1790) 0 3 18 11 2 34 Tetratoma ancora Fabricius, 1790 0 2 0 2 0 4 Tetratoma desmarestii Latreille, 1807 1 4 3 0 0 8 Tetratoma fungorum Fabricius, 1790 10 14 74 16 4 118

75 Melandryidae Orchesia micans (Panzer, 1793) 4 7 15 6 0 32 Orchesia minor Walker, 1836 0 10 7 0 3 20 Orchesia undulata Kraatz, 1853 3 19 56 7 4 89 Anisoxya fuscula (Illiger, 1798) 0 3 2 0 0 5 Abdera flexuosa (Paykull, 1799) 0 9 1 1 1 12 Abdera quadrifasciata (Curtis, 1829) 0 2 0 0 0 2 Phloiotrya vaudoueri Mulsant, 1856 1 0 6 0 0 7 Hypulus quercinus (Quensel, 1790) 0 4 4 0 0 8 Melandrya caraboides (Linnaeus, 1761) 1 11 14 5 1 32 Conopalpus testaceus (Olivier, 1790) 0 3 7 1 0 11 Osphya bipunctata (Fabricius, 1775) 0 1 0 0 0 1

76 Mordellidae Mordellistena neuwaldeggiana (Panzer, 1796) 0 1 4 1 0 6 Mordellistena pumila (Gyllenhal, 1810) 2 0 28 1 0 31 Mordellistena variegata (Fabricius, 1798) 2 0 1 0 0 3 Mordellochroa abdominalis (Fabricius, 1775) 0 5 4 0 0 9

77 Ripiphoridae Metoecus paradoxus (Linnaeus, 1761) 5 14 13 7 2 41

78 Zopheridae Orthocerus clavicornis (Linnaeus, 1758) 0 1 0 0 0 1 Synchita humeralis (Fabricius, 1792) 1 0 9 1 0 11 Synchita separanda (Reitter, 1882) 0 0 2 0 0 2 Bitoma crenata (Fabricius, 1775) 6 5 34 7 0 52 Colydium elongatum (Fabricius, 1787) 0 0 2 0 0 2

79 Tenebrionidae Lagria hirta (Linnaeus, 1758) 77 41 216 46 98 478 Eledona agricola (Herbst, 1783) 5 13 19 3 0 40 Tenebrio molitor Linnaeus, 1758 9 10 18 4 1 42 Tenebrio obscurus Fabricius, 1792 0 1 0 0 0 1 Alphitobius diaperinus (Panzer, 1796) 1 1 7 1 1 11 Alphitobius laevigatus (Fabricius, 1781) 1 1 2 0 0 4 Tribolium castaneum (Herbst, 1797) 3 5 11 4 1 24

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Yorkshire Coleoptera Checklist 2020

Tribolium confusum Jacquelin du Val, 1863 0 1 4 0 0 5 Palorus ratzeburgii (Wissmann, 1848) 1 1 0 1 0 3 Phylan gibbus (Fabricius, 1775) 7 1 0 0 0 8 Melanimon tibialis (Fabricius, 1781) 0 1 6 1 0 8 Nalassus laevioctostriatus (Goeze, 1777) 0 3 10 0 0 13 Xanthomus pallidus (Curtis, 1830) 4 0 0 0 0 4 Blaps mortisaga (Linnaeus, 1758) 0 1 1 0 0 2 Blaps mucronata Latreille, 1804 3 11 7 1 1 23 Prionychus melanarius (Germar, 1813) 0 0 2 0 0 2 Gonodera luperus (Herbst, 1783) 0 0 2 2 0 4 Pseudocistela ceramboides (Linnaeus, 1758) 0 1 0 0 0 1 Isomira murina (Linnaeus, 1758) 10 11 17 4 2 44 Mycetochara humeralis (Fabricius, 1787) 0 0 2 0 0 2 Crypticus quisquilius (Linnaeus, 1760) 18 0 0 0 0 18 Phaleria cadaverina (Fabricius, 1792) 9 0 0 0 0 9 Myrmechixenus subterraneus Chevrolat, 1835 0 0 1 0 0 1 Myrmechixenus vaporariorum Guérin-Méneville, 1843 0 0 1 1 0 2 Corticeus bicolor (Olivier, 1790) 0 5 13 1 0 19 Corticeus fraxini (Kugelann, 1794) 0 0 2 0 0 2 Corticeus linearis (Fabricius, 1790) 1 0 0 0 0 1 Corticeus unicolor Piller & Mitterpacher, 1783 6 0 21 1 1 29 Scaphidema metallicum (Fabricius, 1792) 26 11 32 3 0 72 Alphitophagus bifasciatus (Say, 1824) 3 1 5 0 0 9 Gnatocerus cornutus (Fabricius, 1798) 2 4 3 0 0 9 Diaperis boleti (Linnaeus, 1758) 0 1 0 0 0 1

80 Oedemeridae Nacerdes melanura (Linnaeus, 1758) 19 4 8 6 0 37 Ischnomera caerulea (Linnaeus, 1758) 0 4 0 1 0 5 Ischnomera cinerascens (Pandellé, 1867) 0 3 0 0 0 3 Ischnomera cyanea (Fabricius, 1792) 0 9 0 3 0 12 Ischnomera sanguinicollis (Fabricius, 1787) 0 6 0 0 2 8 Oedemera lurida (Marsham, 1802) 57 11 329 14 10 421 Oedemera nobilis (Scopoli, 1763) 1 4 47 1 0 53 Oedemera virescens (Linnaeus, 1767) 0 38 0 0 0 38

81 Meloidae Meloe proscarabaeus Linnaeus, 1758 1 8 5 5 2 21 Meloe violaceus Marsham, 1802 2 6 32 7 3 50

84 Pyrochroidae Pyrochroa coccinea (Linnaeus, 1761) 1 14 15 0 0 30 Pyrochroa serraticornis (Scopoli, 1763) 50 35 75 31 18 209

85 Salpingidae

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Yorkshire Coleoptera Checklist 2020

Aglenus brunneus (Gyllenhal, 1813) 0 0 1 0 0 1 Lissodema cursor (Gyllenhal, 1813) 0 0 0 2 0 2 Lissodema denticolle (Gyllenhal, 1813) 0 1 0 0 0 1 Rabocerus foveolatus (Ljungh, 1824) 0 1 2 0 0 3 Rabocerus gabrieli Gerhardt, 1901 2 4 9 6 0 21 Sphaeriestes castaneus (Panzer, 1796) 1 8 6 0 1 16 Sphaeriestes reyi (Abeille de Perrin, 1874) 4 4 3 1 1 13 Vincenzellus ruficollis (Panzer, 1794) 9 8 41 11 1 70 Salpingus planirostris (Fabricius, 1787) 23 56 111 22 6 218 Salpingus ruficollis (Linnaeus, 1761) 2 18 20 9 3 52

86 Anthicidae Cordicollis instabilis (Schmidt, 1842) 0 1 0 0 0 1 Stricticollis tobias (Marseul, 1879) 1 0 6 0 0 7 Notoxus monoceros (Linnaeus, 1760) 41 4 0 3 1 49 Anthicus antherinus (Linnaeus, 1760) 2 0 3 0 0 5 Cyclodinus constrictus (Curtis, 1838) 2 1 0 0 0 3 Omonadus floralis (Linnaeus, 1758) 50 16 67 19 3 155 Omonadus formicarius (Goeze, 1777) 15 9 37 6 5 72

87 Aderidae Aderus populneus (Creutzer in Panzer, 1796) 2 0 9 0 0 11 Euglenes oculatus (Paykull, 1798) 2 14 3 6 0 25 Vanonus brevicornis (Perris, 1869) 0 0 2 0 0 2

88 Scraptiidae Anaspis costai Emery, 1876 2 3 11 0 0 16 Anaspis fasciata (Forster, 1771) 18 29 87 22 5 161 Anaspis frontalis (Linnaeus, 1758) 13 56 171 27 10 277 Anaspis garneysi Fowler, 1889 3 8 5 5 0 21 Anaspis lurida Stephens, 1832 0 1 2 3 1 7 Anaspis maculata (Geoffroy in Fourcroy, 1785) 116 98 490 71 24 799 Anaspis pulicaria Costa, A., 1854 0 4 0 2 0 6 Anaspis regimbarti Schilsky, 1895 33 25 131 15 4 208 Anaspis rufilabris (Gyllenhal, 1827) 13 49 145 16 9 232 Anaspis thoracica (Linnaeus, 1758) 1 2 9 1 0 13

89 Cerambycidae Monochamus sartor (Fabricius, 1787) 1 0 4 0 0 5 Monochamus sutor (Linnaeus, 1758) 1 0 6 3 0 10 Rhagium bifasciatum Fabricius, 1775 2 34 67 12 8 123 Rhagium inquisitor (Linnaeus, 1758) 2 4 2 0 0 8 Rhagium mordax (De Geer, 1775) 3 30 41 20 5 99 Stenocorus meridianus (Linnaeus, 1758) 6 15 22 15 3 61 Grammoptera abdominalis (Stephens, 1831) 1 0 4 1 0 6 Grammoptera ruficornis (Fabricius, 1781) 59 57 364 58 14 552

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Yorkshire Coleoptera Checklist 2020

Leptura quadrifasciata Linnaeus, 1758 15 25 57 5 0 102 Anastrangalia sanguinolenta (Linnaeus, 1761) 0 0 1 0 0 1 Stictoleptura rubra (Linnaeus, 1758) 1 0 3 0 0 4 Stictoleptura scutellata (Fabricius, 1781) 0 0 3 0 0 3 Anoplodera sexguttata (Fabricius, 1775) 0 9 0 0 0 9 Pachytodes cerambyciformis (Schrank, 1781) 0 0 5 0 0 5 Alosterna tabacicolor (De Geer, 1775) 0 31 7 2 0 40 Pseudovadonia livida (Fabricius, 1777) 0 0 6 0 0 6 Rutpela maculata (Poda, 1761) 34 28 170 18 3 253 Stenurella melanura (Linnaeus, 1758) 0 6 2 0 0 8 Asemum striatum (Linnaeus, 1758) 3 3 2 2 0 10 Tetropium gabrieli Weise, 1905 1 4 4 2 0 11 Arhopalus ferus (Mulsant, 1839) 1 1 0 0 0 2 Arhopalus rusticus (Linnaeus, 1758) 2 0 7 0 2 11 Cerambyx cerdo Linnaeus, 1758 1 0 0 0 0 1 Gracilia minuta (Fabricius, 1781) 2 0 4 2 0 8 Nathrius brevipennis (Mulsant, 1839) 0 0 3 1 0 4 Molorchus minor (Linnaeus, 1758) 2 1 0 3 0 6 Glaphyra umbellatarum (von Schreber, 1759) 0 0 0 2 0 2 Aromia moschata (Linnaeus, 1758) 2 2 2 4 0 10 Hylotrupes bajulus (Linnaeus, 1758) 0 0 2 0 0 2 Callidium violaceum (Linnaeus, 1758) 1 0 1 0 0 2 Pyrrhidium sanguineum (Linnaeus, 1758) 1 0 0 1 0 2 Phymatodes testaceus (Linnaeus, 1758) 2 10 1 3 0 16 Poecilium alni (Linnaeus, 1767) 0 0 2 0 0 2 Clytus arietis (Linnaeus, 1758) 11 37 128 18 1 195 Plagionotus arcuatus (Linnaeus, 1758) 0 0 1 1 0 2 Anaglyptus mysticus (Linnaeus, 1758) 6 2 14 4 0 26 Mesosa nebulosa (Fabricius, 1781) 0 0 3 0 0 3 Agapanthia villosoviridescens (De Geer, 1775) 33 2 103 8 4 150 Pogonocherus fasciculatus (De Geer, 1775) 0 1 0 1 0 2 Pogonocherus hispidulus (Piller & Mitterpacher, 1783) 5 18 4 4 3 34 Pogonocherus hispidus (Linnaeus, 1758) 2 10 2 4 2 20 Acanthocinus aedilis (Linnaeus, 1758) 2 1 6 1 0 10 Leiopus nebulosus (Linnaeus, 1758) 0 1 0 0 0 1 Leiopus nebulosus (Linnaeus, 1758) 6 23 42 22 2 95 Saperda carcharias (Linnaeus, 1758) 0 3 7 0 0 10 Saperda populnea (Linnaeus, 1758) 5 6 23 11 0 45 Saperda scalaris (Linnaeus, 1758) 0 9 5 0 0 14 Stenostola dubia (Laicharting, 1784) 0 7 8 11 5 31 Phytoecia cylindrica (Linnaeus, 1758) 5 0 0 1 0 6 Tetrops praeustus (Linnaeus, 1758) 6 7 34 6 0 53

92 Chrysomelidae Zeugophora flavicollis (Marsham, 1802) 0 0 1 0 0 1 Zeugophora subspinosa (Fabricius, 1781) 4 3 21 1 0 29

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Yorkshire Coleoptera Checklist 2020

Orsodacne cerasi (Linnaeus, 1758) 0 34 7 9 1 51 Bruchidius cisti (Fabricius, 1775) 6 11 10 3 0 30 Bruchidius varius (Olivier, 1795) 4 1 8 0 0 13 Bruchidius villosus (Fabricius, 1792) 4 2 79 9 0 94 Bruchus loti Paykull, 1800 43 27 162 15 0 247 Bruchus pisorum (Linnaeus, 1758) 2 1 2 0 0 5 Bruchus rufimanus Boheman, 1833 20 10 57 1 0 88 Bruchus rufipes Herbst, 1783 0 0 4 0 0 4 Callosobruchus chinensis (Linnaeus, 1758) 0 1 1 0 0 2 Macroplea appendiculata (Panzer, 1794) 4 1 0 9 0 14 Macroplea mutica (Fabricius, 1792) 4 0 4 0 0 8 Donacia aquatica (Linnaeus, 1758) 0 1 1 1 0 3 Donacia bicolora Zschach, 1788 0 2 0 0 0 2 Donacia cinerea Herbst, 1784 7 5 6 1 0 19 Donacia clavipes Fabricius, 1792 16 2 5 3 0 26 Donacia crassipes Fabricius, 1775 0 0 2 0 0 2 Donacia dentata Hoppe, 1795 1 0 0 0 0 1 Donacia impressa Paykull, 1799 7 1 1 0 0 9 Donacia marginata Hoppe, 1795 1 1 1 5 0 8 Donacia semicuprea Panzer, 1796 15 2 28 3 0 48 Donacia simplex Fabricius, 1775 20 17 53 21 12 123 Donacia sparganii Ahrens, 1810 7 0 0 2 0 9 Donacia thalassina Germar, 1811 4 9 4 0 0 17 Donacia versicolorea (Brahm, 1791) 15 9 25 14 9 72 Donacia vulgaris Zschach, 1788 16 18 38 11 0 83 Plateumaris bracata (Scopoli, 1772) 4 0 0 0 0 4 Plateumaris rustica (Kunze, 1818) 0 3 10 3 1 17 Plateumaris discolor (Panzer, 1795) 1 28 29 27 9 94 Plateumaris sericea (Linnaeus, 1758) 21 18 24 9 1 73 Lema cyanella (Linnaeus, 1758) 3 6 5 3 0 17 Oulema melanopus 6 0 11 3 3 23 Oulema obscura (Stephens, 1831) 14 37 19 20 2 92 Oulema rufocyanea (Suffrian, 1847) 51 14 79 16 6 166 Crioceris asparagi (Linnaeus, 1758) 1 2 37 0 0 40 Lilioceris lilii (Scopoli, 1763) 10 2 39 6 2 59 Labidostomis tridentata (Linnaeus, 1758) 0 1 0 0 0 1 Clytra quadripunctata (Linnaeus, 1758) 0 13 17 0 0 30 Cryptocephalus aureolus Suffrian, 1847 4 11 0 0 0 15 Cryptocephalus biguttatus (Scopoli, 1763) 0 2 0 0 0 2 Cryptocephalus bipunctatus (Linnaeus, 1758) 0 1 0 1 0 2 Cryptocephalus fulvus (Goeze, 1777) 2 2 27 2 0 33 Cryptocephalus hypochaeridis (Linnaeus, 1758) 1 2 0 0 0 3 Cryptocephalus labiatus (Linnaeus, 1761) 4 15 93 12 0 124 Cryptocephalus moraei (Linnaeus, 1758) 0 1 5 0 0 6 Cryptocephalus parvulus Müller, O.F., 1776 0 0 14 1 0 15 Cryptocephalus punctiger Paykull, 1799 0 0 2 0 0 2

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Yorkshire Coleoptera Checklist 2020

Cryptocephalus pusillus Fabricius, 1777 18 14 56 8 1 97 Oomorphus concolor (Sturm, 1807) 0 1 0 1 0 2 Timarcha goettingensis (Linnaeus, 1758) 4 1 40 8 1 54 Timarcha tenebricosa (Fabricius, 1775) 0 0 8 3 223 234 Leptinotarsa decemlineata (Say, 1824) 0 0 1 0 0 1 Chrysolina banksii (Fabricius, 1775) 1 0 2 0 0 3 Chrysolina sturmi (Westhoff, 1882) 0 1 4 2 0 7 Chrysolina americana (Linnaeus, 1758) 3 5 8 0 0 16 Chrysolina brunsvicensis (Gravenhorst, 1807) 0 8 0 3 2 13 Chrysolina fastuosa (Scopoli, 1763) 0 3 33 0 1 37 Chrysolina graminis (Linnaeus, 1758) 9 18 2 12 0 41 Chrysolina herbacea (Duftschmid, 1825) 0 0 1 1 0 2 Chrysolina hyperici (Forster, 1771) 4 8 36 2 1 51 Chrysolina marginata (Linnaeus, 1758) 1 1 5 0 2 9 Chrysolina oricalcia (Müller, O.F., 1776) 4 1 16 7 5 33 Chrysolina polita (Linnaeus, 1758) 41 33 111 33 6 224 Chrysolina sanguinolenta (Linnaeus, 1758) 1 0 7 0 0 8 Chrysolina staphylaea (Linnaeus, 1758) 14 17 39 12 6 88 Chrysolina varians (Schaller, 1783) 2 13 3 5 3 26 Gastrophysa polygoni (Linnaeus, 1758) 26 12 53 31 3 125 Gastrophysa viridula (De Geer, 1775) 45 26 202 42 9 324 Phaedon armoraciae (Linnaeus, 1758) 30 29 63 18 3 143 Phaedon cochleariae (Fabricius, 1792) 40 13 52 24 9 138 Phaedon concinnus Stephens, 1831 10 0 0 0 0 10 Phaedon tumidulus (Germar, 1824) 68 36 195 26 4 329 Hydrothassa glabra (Herbst, 1783) 3 6 4 9 2 24 Hydrothassa hannoveriana (Fabricius, 1775) 0 0 0 10 2 12 Hydrothassa marginella (Linnaeus, 1758) 20 43 49 33 5 150 Prasocuris junci (Brahm, 1790) 14 12 8 7 1 42 Prasocuris phellandrii (Linnaeus, 1758) 6 2 2 3 1 14 Plagiodera versicolora (Laicharting, 1781) 2 3 7 0 0 12 Chrysomela aenea Linnaeus, 1758 1 43 11 9 7 71 Chrysomela populi Linnaeus, 1758 10 13 9 1 0 33 Gonioctena decemnotata (Marsham, 1802) 0 0 2 0 0 2 Gonioctena olivacea (Forster, 1771) 0 4 8 0 1 13 Gonioctena pallida (Linnaeus, 1758) 0 34 3 9 5 51 Gonioctena viminalis (Linnaeus, 1758) 0 3 0 0 0 3 Phratora laticollis Suffrian, 1851 13 6 14 3 0 36 Phratora vitellinae (Linnaeus, 1758) 15 6 39 28 14 102 Phratora vulgatissima (Linnaeus, 1758) 53 14 144 33 18 262 Galerucella calmariensis (Linnaeus, 1767) 14 3 37 7 0 61 Galerucella lineola (Fabricius, 1781) 10 12 100 18 8 148 Galerucella nymphaeae (Linnaeus, 1758) 10 7 69 10 0 96 Galerucella pusilla (Duftschmid, 1825) 2 0 11 8 0 21 Galerucella sagittariae (Gyllenhal, 1813) 41 7 31 12 0 91 Galerucella tenella (Linnaeus, 1761) 15 21 21 33 2 92

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Yorkshire Coleoptera Checklist 2020

Pyrrhalta viburni (Paykull, 1799) 14 10 45 5 0 74 Xanthogaleruca luteola (Müller, O.F., 1766) 0 0 1 0 0 1 Galeruca tanaceti (Linnaeus, 1758) 8 3 17 3 0 31 Lochmaea caprea (Linnaeus, 1758) 8 14 132 14 3 171 Lochmaea crataegi (Forster, 1771) 9 21 91 14 6 141 Lochmaea suturalis (Thomson, C.G., 1866) 11 34 251 9 8 313 Phyllobrotica quadrimaculata (Linnaeus, 1758) 3 7 15 15 0 40 Luperus flavipes (Linnaeus, 1767) 1 6 19 0 0 26 Luperus longicornis (Fabricius, 1781) 9 25 27 16 3 80 Calomicrus circumfusus (Marsham, 1802) 0 1 1 0 0 2 Agelastica alni (Linnaeus, 1758) 12 2 189 3 0 206 Sermylassa halensis (Linnaeus, 1767) 29 14 45 2 2 92 Phyllotreta atra (Fabricius, 1775) 3 9 11 0 0 23 Phyllotreta consobrina (Curtis, 1837) 0 2 0 0 0 2 Phyllotreta cruciferae (Goeze, 1777) 0 3 0 0 0 3 Phyllotreta diademata Foudras, 1860 0 0 6 0 0 6 Phyllotreta exclamationis (Thunberg, 1784) 9 3 10 5 1 28 Phyllotreta flexuosa (Illiger, 1794) 2 3 2 10 2 19 Phyllotreta nemorum (Linnaeus, 1758) 6 4 18 14 7 49 Phyllotreta nigripes (Fabricius, 1775) 11 8 72 6 0 97 Phyllotreta nodicornis (Marsham, 1802) 5 1 24 1 1 32 Phyllotreta ochripes (Curtis, 1837) 8 3 5 0 0 16 Phyllotreta punctulata (Marsham, 1802) 1 1 1 0 0 3 Phyllotreta tetrastigma (Comolli, 1837) 13 7 28 15 1 64 Phyllotreta undulata Kutschera, 1860 36 47 116 29 5 233 Phyllotreta vittula (Redtenbacher, 1849) 1 3 9 3 0 16 Aphthona ?atratula Allard, 1859 1 1 9 1 0 12 Aphthona atrocaerulea (Stephens, 1829) 5 3 14 1 0 23 Aphthona euphorbiae (Schrank, 1781) 88 52 402 26 6 574 Aphthona herbigrada (Curtis, 1837) 3 7 0 5 1 16 Aphthona lutescens (Gyllenhal, 1808) 1 0 3 0 0 4 Aphthona melancholica Weise, 1888 1 1 9 2 0 13 Aphthona nigriceps (Redtenbacher, 1842) 2 2 2 11 2 19 Aphthona nonstriata (Goeze, 1777) 15 12 56 37 1 121 Longitarsus aeneicollis (Faldermann, 1837) 2 2 3 1 0 8 Longitarsus anchusae (Paykull, 1799) 0 2 4 0 0 6 Longitarsus atricillus (Linnaeus, 1761) 3 5 5 0 1 14 Longitarsus brunneus (Duftschmid, 1825) 0 2 2 3 0 7 Longitarsus dorsalis (Fabricius, 1781) 3 1 4 0 0 8 Longitarsus exoletus (Linnaeus, 1758) 3 2 15 1 0 21 Longitarsus flavicornis (Stephens, 1831) 0 0 2 0 0 2 Longitarsus ganglbaueri Heikertinger, 1912 2 3 2 0 2 9 Longitarsus gracilis Kutschera, 1864 23 9 14 5 2 53 Longitarsus holsaticus (Linnaeus, 1758) 0 1 0 0 0 1 Longitarsus jacobaeae (Waterhouse, G.R., 1858) 30 8 70 13 6 127 Longitarsus kutscherae (Rye, 1872) 5 2 16 0 0 23

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Yorkshire Coleoptera Checklist 2020

Longitarsus luridus (Scopoli, 1763) 73 40 297 52 9 471 Longitarsus melanocephalus (De Geer, 1775) 12 5 52 10 1 80 Longitarsus membranaceus (Foudras, 1860) 2 5 3 0 4 14 Longitarsus nasturtii (Fabricius, 1792) 0 1 0 1 0 2 Longitarsus nigerrimus (Gyllenhal, 1827) 0 1 0 0 0 1 Longitarsus nigrofasciatus (Goeze, 1777) 1 0 0 0 0 1 Longitarsus obliteratus (Rosenhauer, 1847) 2 0 1 0 0 3 Longitarsus ochroleucus (Marsham, 1802) 3 2 7 2 0 14 Longitarsus parvulus (Paykull, 1799) 24 29 83 6 0 142 Longitarsus pellucidus (Foudras, 1860) 2 1 7 0 0 10 Longitarsus pratensis (Panzer, 1794) 22 8 28 5 0 63 Longitarsus reichei (Allard, 1860) 3 0 12 0 0 15 Longitarsus rubiginosus (Foudras, 1860) 13 0 21 2 0 36 Longitarsus succineus (Foudras, 1860) 20 8 49 6 3 86 Longitarsus suturellus (Duftschmid, 1825) 37 19 131 25 2 214 Longitarsus tabidus (Fabricius, 1775) 0 1 1 0 0 2 Altica carinthiaca Weise, 1888 0 0 1 0 0 1 Altica brevicollis Foudras, 1860 0 1 1 0 0 2 Altica ericeti (Allard, 1859) 1 12 26 3 3 45 Altica helianthemi (Allard, 1859) 3 5 9 4 0 21 Altica lythri Aubé, 1843 71 52 363 38 23 547 Altica oleracea (Linnaeus, 1758) 14 10 55 5 0 84 Altica palustris Weise, 1888 23 13 96 16 2 150 Hermaeophaga mercurialis (Fabricius, 1792) 0 0 1 0 0 1 Batophila aerata (Marsham, 1802) 0 0 0 1 0 1 Batophila rubi (Paykull, 1799) 8 24 50 11 3 96 Lythraria salicariae (Paykull, 1800) 0 1 2 0 0 3 Ochrosis ventralis (Illiger, 1807) 0 0 1 0 0 1 Neocrepidodera ferruginea (Scopoli, 1763) 75 35 271 49 12 442 Neocrepidodera impressa (Fabricius, 1801) 1 0 0 0 0 1 Neocrepidodera transversa (Marsham, 1802) 29 25 149 23 7 233 Derocrepis rufipes (Linnaeus, 1758) 9 26 23 16 6 80 Hippuriphila modeeri (Linnaeus, 1761) 14 13 19 6 5 57 Crepidodera aurata (Marsham, 1802) 21 4 25 4 0 54 Crepidodera aurea (Fourcroy, 1785) 0 2 16 2 2 22 Crepidodera fulvicornis (Fabricius, 1792) 108 46 450 61 25 690 Crepidodera plutus (Latreille, 1804) 5 0 16 1 0 22 Epitrix atropae Foudras, 1860 2 0 1 0 0 3 Epitrix pubescens (Koch, J.D.W., 1803) 4 0 27 3 0 34 Podagrica fuscicornis (Linnaeus, 1767) 0 3 0 0 0 3 Mantura matthewsii (Curtis, 1833) 5 7 0 10 0 22 Mantura obtusata (Gyllenhal, 1813) 0 4 1 7 0 12 Mantura rustica (Linnaeus, 1767) 6 5 15 8 0 34 Chaetocnema conducta (Motschulsky, 1838) 0 2 0 0 0 2 Chaetocnema arida Foudras, 1860 1 2 0 0 0 3 Chaetocnema concinna (Marsham, 1802) 68 40 246 46 10 410

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Yorkshire Coleoptera Checklist 2020

Chaetocnema confusa (Boheman, 1851) 4 0 1 0 0 5 Chaetocnema hortensis (Fourcroy, 1785) 17 15 57 21 8 118 Chaetocnema picipes Stephens, 1831 1 0 1 0 0 2 Chaetocnema sahlbergii (Gyllenhal, 1827) 0 1 0 0 0 1 Chaetocnema subcoerulea (Kutschera, 1864) 0 1 0 0 0 1 Sphaeroderma rubidum (Graëlls, 1858) 20 13 35 2 1 71 Sphaeroderma testaceum (Fabricius, 1775) 35 15 116 20 6 192 Apteropeda globosa (Illiger, 1794) 0 3 1 2 0 6 Apteropeda orbiculata (Marsham, 1802) 2 8 7 1 1 19 Mniophila muscorum (Koch, J.D.W., 1803) 0 4 0 3 1 8 Psylliodes affinis (Paykull, 1799) 39 5 137 25 1 207 Psylliodes attenuata (Koch, J.D.W., 1803) 0 0 4 1 0 5 Psylliodes chalcomera (Illiger, 1807) 0 0 1 0 0 1 Psylliodes chrysocephala (Linnaeus, 1758) 41 23 223 9 8 304 Psylliodes cuprea (Koch, J.D.W., 1803) 3 10 21 2 2 38 Psylliodes dulcamarae (Koch, J.D.W., 1803) 1 0 6 0 0 7 Psylliodes hyoscyami (Linnaeus, 1758) 0 1 0 0 0 1 Psylliodes laticollis Kutschera, 1864 11 5 16 0 0 32 Psylliodes marcida (Illiger, 1807) 9 4 0 0 0 13 Psylliodes napi (Fabricius, 1792) 10 12 29 14 6 71 Psylliodes picina (Marsham, 1802) 17 7 33 7 0 64 Cassida flaveola Thunberg, 1794 5 12 31 14 1 63 Cassida hemisphaerica Herbst, 1799 1 1 0 0 0 2 Cassida nobilis Linnaeus, 1758 0 1 3 0 0 4 Cassida prasina Illiger, 1798 0 0 1 0 0 1 Cassida rubiginosa Müller, O.F., 1776 62 32 223 38 16 371 Cassida vibex Linnaeus, 1767 1 0 0 1 0 2 Cassida viridis Linnaeus, 1758 5 4 18 3 0 30

93 Nemonychidae Cimberis attelaboides (Fabricius, 1787) 0 10 0 1 0 11

94 Anthribidae Anthribus fasciatus (Forster, 1771) 0 0 4 0 0 4 Anthribus nebulosus (Forster, 1771) 6 14 10 11 3 44 Platyrhinus resinosus (Scopoli, 1763) 10 1 36 3 0 50 Platystomos albinus (Linnaeus, 1758) 0 0 1 1 0 2 Choragus sheppardi Kirby, W., 1819 7 2 1 1 0 11 Araecerus fasciculatus (De Geer, 1775) 0 0 0 1 0 1 Bruchela rufipes (Olivier, 1790) 3 0 22 1 0 26

95 Rhynchitidae Caenorhinus mannerheimii (Hummel, 1823) 9 22 11 9 0 51 Involvulus cupreus (Linnaeus, 1758) 0 30 1 0 0 31 Lasiorhynchites cavifrons (Gyllenhal, 1833) 0 2 6 4 0 12 Lasiorhynchites olivaceus (Gyllenhal, 1833) 0 1 0 0 0 1

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Yorkshire Coleoptera Checklist 2020

Neocoenorrhinus minutus (Herbst, 1797) 0 12 4 0 0 16 Neocoenorrhinus aequatus (Linnaeus, 1767) 14 6 52 1 0 73 Neocoenorrhinus germanicus (Herbst, 1797) 16 8 23 7 0 54 Neocoenorrhinus interpunctatus (Stephens, 1831) 0 0 3 0 0 3 Neocoenorrhinus pauxillus (Germar, 1824) 0 0 0 1 0 1 Temnocerus coeruleus (Fabricius, 1798) 2 1 8 1 0 12 Temnocerus longiceps (Thomson, C.G., 1888) 4 4 19 9 0 36 Temnocerus nanus (Paykull, 1792) 18 12 73 15 0 118 Byctiscus betulae (Linnaeus, 1758) 0 2 0 0 0 2 Deporaus betulae (Linnaeus, 1758) 19 30 78 18 1 146

96 Attelabidae Attelabus nitens (Scopoli, 1763) 13 19 61 12 0 105 Apoderus coryli (Linnaeus, 1758) 0 3 1 1 0 5

97 Apionidae Apion cruentatum Walton, J., 1844 8 5 14 5 1 33 Apion frumentarium (Linnaeus, 1758) 61 16 113 17 7 214 Apion haematodes Kirby, W., 1808 17 7 42 6 3 75 Apion rubens Stephens, 1839 3 4 2 2 8 19 Apion rubiginosum Grill, 1893 1 0 3 0 0 4 Perapion affine (Kirby, 1808) 2 3 4 1 0 10 Perapion curtirostre (Germar, 1817) 45 24 136 26 7 238 Perapion hydrolapathi (Marsham, 1802) 39 16 48 11 0 114 Perapion marchicum (Herbst, 1797) 14 6 34 3 0 57 Perapion violaceum (Kirby, 1808) 41 27 175 31 19 293 Aspidapion aeneum (Fabricius, 1775) 9 6 28 0 1 44 Aspidapion radiolus (Marsham, 1802) 18 9 36 2 2 67 Aspidapion soror (Rey, 1895) 0 0 0 1 0 1 Ceratapion carduorum (Kirby, W., 1808) 9 7 25 6 0 47 Ceratapion gibbirostre (Gyllenhal, 1813) 58 4 78 15 6 161 Ceratapion onopordi (Kirby, W., 1808) 26 4 23 5 4 62 Diplapion confluens (Kirby, W., 1808) 11 0 2 0 0 13 Diplapion stolidum (Germar, 1817) 1 0 1 0 0 2 Omphalapion hookerorum (Kirby, W., 1808) 16 3 20 3 1 43 Omphalapion laevigatum (Paykull, 1792) 0 0 0 1 0 1 Exapion fuscirostre (Fabricius, 1775) 1 0 18 0 0 19 Exapion ulicis (Forster, 1771) 78 18 98 9 10 213 Kalcapion pallipes (Kirby, W., 1808) 7 25 10 11 3 56 Melanapion minimum (Herbst, 1797) 0 4 2 0 0 6 Squamapion atomarium (Kirby, W., 1808) 5 2 0 0 0 7 Squamapion vicinum (Kirby, W., 1808) 0 0 1 1 0 2 Malvapion malvae (Fabricius, 1775) 24 0 57 2 1 84 Pseudapion rufirostre (Fabricius, 1775) 5 1 13 2 1 22 Rhopalapion longirostre (Olivier, 1807) 0 0 0 1 0 1 Cyanapion afer (Gyllenhal, 1833) 6 2 8 4 1 21

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Yorkshire Coleoptera Checklist 2020

Cyanapion gyllenhalii (Kirby, W., 1808) 1 2 2 1 0 6 Cyanapion spencii (Kirby, W., 1808) 1 4 13 11 2 31 Eutrichapion ervi (Kirby, W., 1808) 54 31 132 21 5 243 Eutrichapion punctigerum (Paykull, 1792) 1 7 4 2 2 16 Eutrichapion viciae (Paykull, 1800) 19 11 85 16 5 136 Eutrichapion vorax (Herbst, 1797) 18 5 26 5 0 54 Hemitrichapion reflexum (Gyllenhal, 1833) 0 1 1 0 0 2 Holotrichapion aethiops (Herbst, 1797) 8 3 6 3 2 22 Holotrichapion ononis (Kirby, 1808) 25 8 2 8 2 45 Holotrichapion pisi (Fabricius, 1801) 14 8 17 6 4 49 Oxystoma cerdo (Gerstaecker, 1854) 21 8 61 4 1 95 Oxystoma craccae (Linnaeus, 1767) 2 0 6 2 0 10 Oxystoma pomonae (Fabricius, 1798) 1 0 1 0 0 2 Oxystoma subulatum (Kirby, W., 1808) 38 39 96 19 2 194 Pirapion immune (Kirby, W., 1808) 2 3 10 0 0 15 Catapion curtisii (Stephens, 1831) 1 0 0 0 0 1 Catapion pubescens (Kirby, W., 1811) 2 1 8 4 1 16 Catapion seniculus (Kirby, W., 1808) 25 7 7 7 4 50 Ischnopterapion loti (Kirby, 1808) 58 47 165 46 8 324 Ischnopterapion modestum (Germar, 1817) 1 0 2 1 0 4 Ischnopterapion virens (Herbst, 1797) 41 15 31 13 3 103 Protopirapion atratulum (Germar, 1817) 2 7 9 0 3 21 Stenopterapion meliloti (Kirby, 1808) 11 3 11 3 0 28 Stenopterapion scutellare (Kirby, 1811) 0 1 5 1 0 7 Stenopterapion tenue (Kirby, 1808) 15 2 14 5 0 36 Synapion ebeninum (Kirby, W., 1808) 0 0 0 1 0 1 Betulapion simile (Kirby, W., 1811) 42 5 90 13 0 150 Protapion apricans (Herbst, 1797) 67 44 219 41 21 392 Protapion assimile (Kirby, W., 1808) 27 36 80 11 1 155 Protapion difforme (Germar, 1818) 0 0 3 0 0 3 Protapion dissimile (Germar, 1817) 0 0 3 0 0 3 Protapion filirostre (Kirby, W., 1808) 0 0 4 0 0 4 Protapion fulvipes (Geoffroy in Fourcroy, 1785) 101 72 242 46 9 470 Protapion nigritarse (Kirby, W., 1808) 27 15 92 18 7 159 Protapion ononidis (Gyllenhal, 1827) 11 2 8 5 0 26 Protapion schoenherri (Boheman, 1839) 0 1 0 1 0 2 Protapion trifolii (Linnaeus, 1768) 19 5 42 4 2 72 Protapion varipes (Germar, 1817) 0 0 1 0 0 1 Pseudoprotapion astragali (Paykull, 1800) 0 0 0 1 0 1

98 Nanophyidae Nanophyes marmoratus (Goeze, 1777) 14 4 47 10 0 75

99 Dryophthoridae Dryophthorus corticalis (Paykull, 1792) 0 0 3 0 0 3 Sitophilus granarius (Linnaeus, 1758) 3 4 7 1 0 15

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Yorkshire Coleoptera Checklist 2020

Sitophilus oryzae (Linnaeus, 1763) 5 6 4 2 0 17 Sitophilus zeamais Motschulsky, 1855 0 0 2 0 0 2

100 Erirhinidae Grypus equiseti (Fabricius, 1775) 23 37 41 23 5 129 Notaris acridulus (Linnaeus, 1758) 55 15 120 48 5 243 Notaris aethiops (Fabricius, 1792) 0 1 0 1 0 2 Notaris scirpi (Fabricius, 1792) 6 0 45 10 0 61 Thryogenes festucae (Herbst, 1795) 8 0 0 0 0 8 Thryogenes nereis (Paykull, 1800) 0 3 0 1 0 4 Thryogenes scirrhosus (Gyllenhal, 1836) 3 0 1 0 0 4 Tournotaris bimaculatus (Fabricius, 1787) 34 3 16 1 0 54 Stenopelmus rufinasus Gyllenhal, 1836 5 0 7 1 0 13 Tanysphyrus lemnae (Paykull, 1792) 22 14 34 22 0 92

102 Curculionidae Andrion regensteinense (Herbst, 1797) 25 12 124 8 3 172 Charagmus griseus (Fabricius, 1775) 4 11 6 0 0 21 Coelositona cambricus (Stephens, 1831) 1 2 9 9 1 22 Coelositona puberulus (Reitter, 1903) 0 0 1 0 0 1 Taphrorychus bicolor (Herbst, 1794) 0 0 0 2 0 2 Archarius pyrrhoceras (Marsham, 1802) 28 27 115 16 3 189 Archarius salicivorus (Paykull, 1792) 32 16 117 27 11 203 Curculio villosus (Fabricius, 1781) 2 2 19 2 0 25 Curculio betulae (Stephens, 1831) 5 1 34 1 1 42 Curculio glandium Marsham, 1802 18 3 58 6 4 89 Curculio nucum Linnaeus, 1758 0 0 9 1 0 10 Curculio rubidus (Gyllenhal, 1836) 2 2 11 1 0 16 Curculio venosus (Gravenhorst, 1807) 4 1 42 4 0 51 Anthonomus bituberculatus Thomson, C.G., 1868 6 3 13 4 3 29 Anthonomus brunnipennis (Curtis, 1840) 1 1 0 0 0 2 Anthonomus conspersus Desbrochers, 1868 0 7 1 0 0 8 Anthonomus pedicularius (Linnaeus, 1758) 18 21 69 13 2 123 Anthonomus piri Kollar, 1837 0 1 1 0 0 2 Anthonomus pomorum (Linnaeus, 1758) 5 6 5 2 0 18 Anthonomus rectirostris (Linnaeus, 1758) 0 4 0 0 0 4 Anthonomus rubi (Herbst, 1795) 36 17 78 11 4 146 Anthonomus ulmi (De Geer, 1775) 0 3 4 1 4 12 Cionus alauda (Herbst, 1784) 13 11 33 8 1 66 Cionus hortulanus (Fourcroy, 1785) 0 0 2 0 0 2 Cionus scrophulariae (Linnaeus, 1758) 39 21 133 35 13 241 Cleopus pulchellus (Herbst, 1795) 1 7 8 6 0 22 Ellescus bipunctatus (Linnaeus, 1758) 0 3 0 0 0 3 Dorytomus dejeani Faust, 1882 7 3 37 4 0 51 Dorytomus filirostris (Gyllenhal, 1836) 7 0 5 3 0 15 Dorytomus hirtipennis Bedel, 1884 3 0 1 0 0 4

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Yorkshire Coleoptera Checklist 2020

Dorytomus longimanus (Forster, 1771) 7 7 19 4 0 37 Dorytomus melanophthalmus (Paykull, 1792) 18 9 19 8 0 54 Dorytomus rufatus Bedel, 1888 12 12 39 6 0 69 Dorytomus salicinus (Gyllenhal, 1827) 1 0 4 10 0 15 Dorytomus salicis Walton, J., 1851 6 1 4 0 0 11 Dorytomus taeniatus (Fabricius, 1781) 27 20 101 24 11 183 Dorytomus tortrix (Linnaeus, 1761) 7 3 29 3 1 43 Dorytomus tremulae (Fabricius, 1787) 1 0 12 1 0 14 Cleopomiarus graminis (Gyllenhal, 1813) 8 0 8 2 0 18 Cleopomiarus plantarum (Germar, 1824) 0 0 1 0 0 1 Gymnetron beccabungae (Linnaeus, 1761) 0 0 1 0 0 1 Gymnetron melanarium (Germar, 1821) 3 0 0 0 0 3 Gymnetron rostellum (Herbst, 1795) 0 0 1 0 0 1 Gymnetron veronicae (Germar, 1821) 4 2 0 1 0 7 Gymnetron villosulum Gyllenhal, 1838 24 0 4 2 1 31 Mecinus circulatus (Marsham, 1802) 0 0 1 0 0 1 Mecinus labilis (Herbst, 1795) 1 7 2 1 0 11 Mecinus pascuorum (Gyllenhal, 1813) 51 35 108 16 2 212 Mecinus pyraster (Herbst, 1795) 38 20 67 12 2 139 Miarus campanulae (Linnaeus, 1767) 2 8 9 0 0 19 Rhinusa antirrhini (Paykull, 1800) 6 0 43 7 0 56 Rhinusa linariae (Panzer, 1795) 0 3 2 0 0 5 Isochnus sequensi (Stierlin, 1894) 12 1 3 2 0 18 Isochnus foliorum (Müller, O.F., 1764) 1 2 4 4 1 12 Orchestes alni (Linnaeus, 1758) 8 0 3 3 0 14 Orchestes fagi (Linnaeus, 1758) 29 47 128 56 12 272 Orchestes pilosus (Fabricius, 1781) 1 8 4 0 0 13 Orchestes quercus (Linnaeus, 1758) 34 29 45 16 0 124 Orchestes rusci (Herbst, 1795) 20 20 77 16 4 137 Orchestes signifer (Creutzer, 1799) 13 7 11 4 0 35 Orchestes testaceus (Müller, O.F., 1776) 0 0 1 0 0 1 Pseudorchestes pratensis (Germar, 1821) 0 0 2 1 0 3 Rhamphus oxyacanthae (Marsham, 1802) 16 9 49 5 2 81 Rhamphus pulicarius (Herbst, 1795) 13 18 66 4 2 103 Tachyerges salicis (Linnaeus, 1758) 11 15 19 8 1 54 Tachyerges stigma (Germar, 1821) 0 1 2 1 0 4 Smicronyx reichi (Gyllenhal, 1836) 0 0 0 1 0 1 Orthochaetes setiger (Beck, 1817) 4 7 5 3 0 19 Sibinia primita (Herbst, 1795) 0 0 3 0 0 3 Tychius junceus (Reich, 1797) 0 0 1 0 0 1 Tychius lineatulus Stephens, 1831 2 1 0 0 0 3 Tychius meliloti Stephens, 1831 8 2 21 1 0 32 Tychius picirostris (Fabricius, 1787) 53 34 252 35 15 389 Tychius schneideri (Herbst, 1795) 1 1 0 0 0 2 Bagous alismatis (Marsham, 1802) 3 1 17 2 0 23 Bagous limosus (Gyllenhal, 1827) 1 3 6 1 0 11

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Yorkshire Coleoptera Checklist 2020

Bagous lutulosus (Gyllenhal, 1827) 1 0 0 0 0 1 Bagous nodulosus Gyllenhal, 1836 0 0 0 2 0 2 Bagous petro (Herbst, 1795) 0 0 0 3 0 3 Bagous subcarinatus Gyllenhal, 1836 0 1 0 0 0 1 Bagous tempestivus (Herbst, 1795) 3 2 2 1 0 8 Aulacobaris lepidii (Germar, 1824) 0 3 0 1 0 4 Aulacobaris picicornis (Marsham, 1802) 0 0 2 0 0 2 Melanobaris laticollis (Marsham, 1802) 0 1 0 0 0 1 Limnobaris dolorosa (Goeze, 1777) 17 18 30 11 0 76 Limnobaris t-album (Linnaeus, 1758) 0 6 2 5 1 14 Amalorrhynchus melanarius (Stephens, 1831) 2 0 9 0 0 11 Calosirus terminatus (Herbst, 1795) 1 0 0 0 0 1 Ceutorhynchus assimilis 134 66 184 46 9 439 Ceutorhynchus assimilis (Paykull, 1792) 3 8 9 1 1 22 Ceutorhynchus alliariae Brisout de Barneville, H., 1860 1 5 5 4 2 17 Ceutorhynchus atomus Boheman, 1845 1 0 0 1 0 2 Ceutorhynchus chalybaeus Germar, 1824 0 0 2 1 0 3 Ceutorhynchus cochleariae (Gyllenhal, 1813) 3 2 2 2 0 9 Ceutorhynchus constrictus (Marsham, 1802) 1 4 4 3 0 12 Ceutorhynchus contractus (Marsham, 1802) 26 19 23 17 3 88 Ceutorhynchus contractus var. pallipes Crotch, 1865 0 0 1 0 0 1 Ceutorhynchus erysimi (Fabricius, 1787) 37 34 44 30 5 150 Ceutorhynchus hirtulus Germar, 1824 0 1 0 1 0 2 Ceutorhynchus obstrictus (Marsham, 1802) 40 14 41 11 4 110 Ceutorhynchus pallidactylus (Marsham, 1802) 109 30 179 39 22 379 Ceutorhynchus parvulus Brisout de Barneville, Ch., 1869 0 0 1 0 0 1 Ceutorhynchus pectoralis Weise, 1895 0 0 0 1 0 1 Ceutorhynchus picitarsis Gyllenhal, 1837 9 2 14 0 0 25 Ceutorhynchus pumilio (Gyllenhal, 1827) 1 0 0 0 0 1 Ceutorhynchus pyrrhorhynchus (Marsham, 1802) 8 1 11 2 0 22 Ceutorhynchus resedae (Marsham, 1802) 2 0 3 0 0 5 Ceutorhynchus sulcicollis (Paykull, 1800) 1 0 0 1 0 2 Ceutorhynchus turbatus Schultze, 1903 3 0 0 0 0 3 Ceutorhynchus typhae (Herbst, 1795) 86 61 77 37 13 274 Coeliodes rana (Fabricius, 1787) 11 19 24 8 1 63 Coeliodes ruber (Marsham, 1802) 0 0 1 0 0 1 Coeliodes transversealbofasciatus (Goeze, 1777) 1 1 0 0 0 2 Coeliodinus rubicundus (Herbst, 1795) 10 18 36 10 1 75 Datonychus angulosus (Boheman, 1845) 2 0 1 0 0 3 Datonychus melanostictus (Marsham, 1802) 0 0 5 4 0 9 Drupenatus nasturtii (Germar, 1824) 9 0 12 0 0 21 Glocianus punctiger (Sahlberg, C.R., 1835) 8 0 3 1 0 12 Glocianus distinctus (Brisout de Barneville, Ch., 1870) 1 0 3 1 0 5 Hadroplontus litura (Fabricius, 1775) 27 12 59 16 6 120 Hadroplontus trimaculatus (Fabricius, 1775) 1 4 0 0 0 5 Micrelus ericae (Gyllenhal, 1813) 5 21 76 6 6 114

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Yorkshire Coleoptera Checklist 2020

Microplontus melanostigma (Herbst, 1795) 35 3 12 7 0 57 Microplontus campestris (Gyllenhal, 1837) 1 0 13 3 0 17 Mogulones asperifoliarum (Gyllenhal, 1813) 16 6 13 5 2 42 Nedyus quadrimaculatus (Linnaeus, 1758) 132 76 326 133 46 713 Parethelcus pollinarius (Forster, 1771) 69 23 145 30 3 270 Poophagus sisymbrii (Fabricius, 1777) 28 9 29 24 1 91 Sirocalodes depressicollis (Gyllenhal, 1813) 2 1 0 3 0 6 Sirocalodes mixtus (Mulsant & Rey, 1858) 5 0 2 0 0 7 Sirocalodes quercicola (Paykull, 1792) 0 0 2 0 0 2 Stenocarus ruficornis (Stephens, 1831) 3 7 0 4 0 14 Thamiocolus viduatus (Gyllenhal, 1813) 5 0 0 1 0 6 Trichosirocalus dawsoni (Brisout de Barneville, Ch., 1869) 1 1 0 0 0 2 Trichosirocalus horridus (Panzer, 1801) 0 2 1 0 0 3 Trichosirocalus thalhammeri (Schultze, 1906) 1 0 0 0 0 1 Trichosirocalus troglodytes (Fabricius, 1787) 96 57 161 28 1 343 Zacladus exiguus (Olivier, 1807) 0 0 0 1 0 1 Zacladus geranii (Paykull, 1800) 21 12 37 34 14 118 Amalus scortillum (Herbst, 1795) 8 1 8 6 0 23 Eubrychius velutus (Beck, 1817) 0 1 8 2 0 11 Neophytobius muricatus (Brisout de Barneville, Ch., 1867) 0 1 0 0 0 1 Neophytobius quadrinodosus (Gyllenhal, 1813) 1 2 0 0 0 3 Pelenomus canaliculatus (Fåhraeus, 1843) 2 2 3 2 1 10 Pelenomus comari (Herbst, 1795) 4 3 2 13 0 22 Pelenomus quadricorniger (Colonnelli, 1986) 0 0 2 0 0 2 Pelenomus quadrituberculatus (Fabricius, 1787) 17 11 17 12 0 57 Phytobius leucogaster (Marsham, 1802) 10 5 13 0 3 31 Rhinoncus bruchoides (Herbst, 1784) 3 2 3 0 0 8 Rhinoncus castor (Fabricius, 1792) 19 8 37 9 2 75 Rhinoncus inconspectus (Herbst, 1795) 27 1 18 4 0 50 Rhinoncus pericarpius (Linnaeus, 1758) 66 43 141 40 13 303 Rhinoncus perpendicularis (Reich, 1797) 34 9 52 14 5 114 Rhopalomesites tardyi (Curtis, 1825) 0 9 5 1 0 15 Pselactus spadix (Herbst, 1795) 17 0 0 0 0 17 Pseudophloeophagus aeneopiceus (Boheman, 1845) 0 1 0 0 0 1 Euophryum confine (Broun, 1881) 31 21 105 13 1 171 Euophryum rufum (Broun, 1880) 0 0 5 2 0 7 Pentarthrum huttoni Wollaston, 1854 3 0 20 2 0 25 Phloeophagus lignarius (Marsham, 1802) 3 0 9 1 0 13 Rhyncolus ater (Linnaeus, 1758) 0 2 3 0 0 5 Cryptorhynchus lapathi (Linnaeus, 1758) 3 3 25 3 2 36 Acalles misellus Boheman, 1844 8 3 2 0 0 13 Acalles ptinoides (Marsham, 1802) 1 7 20 2 5 35 Kyklioacalles roboris (Curtis, 1835) 0 13 14 6 1 34 Gronops lunatus (Fabricius, 1775) 2 0 5 0 0 7 Graptus triguttatus (Fabricius, 1775) 10 8 2 10 3 33 Neliocarus faber (Herbst, 1784) 2 3 3 1 0 9

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Yorkshire Coleoptera Checklist 2020

Neliocarus nebulosus (Stephens, 1831) 4 21 13 5 0 43 Neliocarus sus (Stephens, 1831) 18 30 20 3 3 74 Strophosoma capitatum (De Geer, 1775) 1 3 7 4 0 15 Strophosoma melanogrammum (Forster, 1771) 61 54 184 22 10 331 Attactagenus plumbeus (Marsham, 1802) 11 8 8 3 1 31 Philopedon plagiatum (Schaller, 1783) 40 20 5 0 0 65 Barynotus moerens (Fabricius, 1792) 4 17 47 28 5 101 Barynotus obscurus (Fabricius, 1775) 19 13 40 14 2 88 Barynotus squamosus Germar, 1824 0 5 14 15 5 39 Omiamima mollina (Boheman, 1834) 11 5 19 2 0 37 Otiorhynchus atroapterus (De Geer, 1775) 0 15 1 0 0 16 Otiorhynchus clavipes (Bonsdorff, 1785) 0 1 0 0 0 1 Otiorhynchus desertus Rosenhauer, 1847 0 0 3 0 1 4 Otiorhynchus ligneus (Olivier, 1807) 1 5 2 3 0 11 Otiorhynchus ligustici (Linnaeus, 1758) 0 0 1 0 0 1 Otiorhynchus nodosus (Müller, O. F. 1764) 0 6 3 9 2 20 Otiorhynchus ovatus (Linnaeus, 1758) 48 22 25 13 1 109 Otiorhynchus porcatus (Herbst, 1795) 8 7 2 5 2 24 Otiorhynchus raucus (Fabricius, 1777) 1 1 1 2 0 5 Otiorhynchus rugifrons (Gyllenhal, 1813) 1 5 5 6 3 20 Otiorhynchus rugosostriatus (Goeze, 1777) 6 4 12 1 0 23 Otiorhynchus singularis (Linnaeus, 1767) 47 78 259 46 22 452 Otiorhynchus sulcatus (Fabricius, 1775) 30 22 95 13 7 167 Caenopsis fissirostris (Walton, J., 1847) 1 0 0 1 0 2 Phyllobius argentatus (Linnaeus, 1758) 29 62 243 35 22 391 Phyllobius glaucus (Scopoli, 1763) 13 57 126 30 16 242 Phyllobius maculicornis Germar, 1824 22 16 86 16 2 142 Phyllobius oblongus (Linnaeus, 1758) 12 28 172 38 19 269 Phyllobius pomaceus Gyllenhal, 1834 70 44 158 71 28 371 Phyllobius pyri (Linnaeus, 1758) 104 79 354 69 28 634 Phyllobius roboretanus Gredler, 1882 98 65 262 62 28 515 Phyllobius vespertinus (Fabricius, 1792) 2 2 1 0 0 5 Phyllobius virideaeris (Laicharting, 1781) 30 23 68 15 4 140 Phyllobius viridicollis (Fabricius, 1801) 16 13 4 42 8 83 Liophloeus tessulatus (Müller, O.F., 1776) 36 17 47 9 1 110 Pachyrhinus lethierryi (Desbrochers, 1875) 0 1 5 0 0 6 Polydrusus cervinus (Linnaeus, 1758) 39 35 120 15 3 212 Polydrusus confluens Stephens, 1831 1 3 1 0 0 5 Polydrusus flavipes (De Geer, 1775) 2 0 2 0 0 4 Polydrusus formosus (Mayer, 1779) 8 0 10 0 0 18 Polydrusus mollis (Strøm, 1768) 0 33 9 7 4 53 Polydrusus pilosus Gredler, 1866 1 11 5 2 2 21 Polydrusus pterygomalis Boheman, 1840 10 53 21 23 9 116 Polydrusus pulchellus Stephens, 1831 3 0 0 0 0 3 Polydrusus tereticollis (De Geer, 1775) 0 52 69 5 1 127 Barypeithes araneiformis (Schrank, 1781) 19 25 51 21 9 125

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Yorkshire Coleoptera Checklist 2020

Barypeithes pellucidus (Boheman, 1834) 47 68 357 56 15 543 Barypeithes pyrenaeus Seidlitz, 1868 0 0 2 0 0 2 Barypeithes sulcifrons (Boheman, 1843) 6 5 8 0 0 19 Brachysomus echinatus (Bonsdorff, 1785) 13 12 15 12 1 53 Sciaphilus asperatus (Bonsdorff, 1785) 15 22 58 32 3 130 Sitona ambiguus Gyllenhal, 1834 0 3 1 1 0 5 Sitona cylindricollis (Fåhraeus, 1840) 10 3 12 0 1 26 Sitona hispidulus (Fabricius, 1777) 40 34 85 24 7 190 Sitona humeralis Stephens, 1831 17 6 22 9 1 55 Sitona lepidus Gyllenhal, 1834 46 48 144 44 15 297 Sitona lineatus (Linnaeus, 1758) 301 134 646 123 33 1237 Sitona lineellus (Bonsdorff, 1785) 6 14 4 2 0 26 Sitona macularius (Marsham, 1802) 2 2 1 0 0 5 Sitona ononidis Sharp, 1866 0 1 3 0 0 4 Sitona puncticollis Stephens, 1831 10 10 11 3 0 34 Sitona striatellus Gyllenhal, 1834 11 4 14 6 1 36 Sitona sulcifrons (Thunberg, 1798) 19 15 31 10 3 78 Sitona suturalis Stephens, 1831 31 40 78 29 5 183 Sitona waterhousei Walton, J., 1846 0 1 1 0 0 2 Tanymecus palliatus (Fabricius, 1787) 7 5 1 1 0 14 Trachyphloeus scabricul (Linnaeus, 1771) 2 0 1 0 0 3 Trachyphloeus alternans Gyllenhal, 1834 0 2 0 0 0 2 Trachyphloeus angustisetulus Hansen, 1915 0 2 2 0 0 4 Trachyphloeus bifoveolatus (Beck, 1817) 1 3 4 0 0 8 Cathormiocerus aristatus (Gyllenhal, 1827) 1 1 4 0 0 6 Cathormiocerus spinosus (Goeze, 1777) 2 2 1 0 0 5 Cathormiocerus myrmecophilus (Seidlitz, 1868) 1 0 0 0 0 1 Tropiphorus elevatus (Herbst, 1795) 0 1 1 0 0 2 Tropiphorus obtusus (Bonsdorff, 1785) 4 2 1 4 0 11 Tropiphorus terricola (Newman, 1838) 9 10 14 7 5 45 Hypera arator (Linnaeus, 1758) 10 2 6 3 0 21 Hypera dauci (Olivier, 1807) 3 1 0 0 0 4 Hypera diversipunctata (Schrank, 1798) 4 0 2 3 0 9 Hypera fuscocinerea (Marsham, 1802) 1 2 0 0 0 3 Hypera meles (Fabricius, 1792) 4 1 1 2 0 8 Hypera nigrirostris (Fabricius, 1775) 35 19 64 19 5 142 Hypera ononidis Chevrolat, 1863 0 0 1 0 0 1 Hypera plantaginis (De Geer, 1775) 8 14 22 17 0 61 Hypera pollux (Fabricius, 1801) 14 2 4 3 0 23 Hypera postica (Gyllenhal, 1813) 22 14 24 8 1 69 Hypera rumicis (Linnaeus, 1758) 9 4 14 4 1 32 Hypera suspiciosa (Herbst, 1795) 2 4 5 3 3 17 Hypera venusta (Fabricius, 1781) 2 2 5 0 0 9 Hypera zoilus (Scopoli, 1763) 9 10 11 17 1 48 Limobius borealis (Paykull, 1792) 4 1 0 9 2 16 Larinus planus (Fabricius, 1792) 0 0 1 0 0 1

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Yorkshire Coleoptera Checklist 2020

Lixus angustatus (Fabricius, 1775) 0 1 0 0 0 1 Lixus paraplecticus (Linnaeus, 1758) 3 0 1 0 0 4 Cleonis pigra (Scopoli, 1763) 22 2 0 0 0 24 Rhinocyllus conicus (Frölich, 1792) 2 0 17 0 1 20 Magdalis armigera (Fourcroy, 1785) 3 11 17 9 2 42 Magdalis barbicornis (Latreille, 1804) 2 0 3 0 0 5 Magdalis carbonaria (Linnaeus, 1758) 4 4 9 5 1 23 Magdalis cerasi (Linnaeus, 1758) 4 3 20 9 0 36 Magdalis duplicata Germar, 1818 0 0 0 1 0 1 Magdalis phlegmatica (Herbst, 1797) 1 0 0 3 0 4 Magdalis ruficornis (Linnaeus, 1758) 6 5 8 7 1 27 Liparus coronatus (Goeze, 1777) 0 1 0 0 0 1 Leiosoma deflexum (Panzer, 1795) 21 40 167 44 16 288 Leiosoma oblongulum Boheman, 1842 0 0 2 2 0 4 Anoplus plantaris (Naezen, 1794) 7 17 34 5 0 63 Anoplus roboris Suffrian, 1840 0 3 2 2 0 7 Hylobius abietis (Linnaeus, 1758) 16 10 13 6 1 46 Pissodes castaneus (De Geer, 1775) 8 2 1 1 0 12 Pissodes pini (Linnaeus, 1758) 10 8 11 4 0 33 Trachodes hispidus (Linnaeus, 1758) 0 0 2 0 0 2 Orobitis cyaneus (Linnaeus, 1758) 2 8 2 10 0 22 Scolytus intricatus (Ratzeburg, 1837) 5 16 19 14 0 54 Scolytus laevis Chapuis, 1873 0 1 0 0 0 1 Scolytus mali (Bechstein & Scharfenberg, 1805) 1 3 5 5 0 14 Scolytus multistriatus (Marsham, 1802) 1 0 13 0 1 15 Scolytus ratzeburgi Janson, E.W., 1856 0 2 2 0 0 4 Scolytus rugulosus (Müller, P.W.J., 1818) 2 0 4 3 0 9 Scolytus scolytus (Fabricius, 1775) 3 11 32 7 1 54 Pityophthorus lichtensteinii (Ratzeburg, 1837) 1 0 3 0 0 4 Pityophthorus pubescens (Marsham, 1802) 18 1 0 0 0 19 Cryphalus asperatus (Gyllenhal, 1813) 7 5 0 3 0 15 Ernoporicus caucasicus (Lindemann, 1876) 0 3 0 2 0 5 Ernoporicus fagi (Fabricius, 1798) 0 0 0 3 0 3 Ernoporus tiliae (Panzer, 1793) 0 0 1 2 0 3 Dryocoetes alni (Georg, 1856) 0 0 5 6 2 13 Dryocoetes autographus (Ratzeburg, 1837) 3 14 1 5 1 24 Dryocoetes villosus (Fabricius, 1792) 4 14 29 11 0 58 Ips acuminatus (Gyllenhal, 1827) 1 1 0 0 0 2 Ips cembrae (Heer, 1836) 0 3 0 0 0 3 Ips sexdentatus (Boerner, 1766) 0 0 1 0 0 1 Ips typographus (Linnaeus, 1758) 2 2 0 0 0 4 Orthotomicus laricis (Fabricius, 1792) 2 1 0 1 0 4 Pityogenes bidentatus (Herbst, 1784) 5 15 10 4 1 35 Pityogenes chalcographus (Linnaeus, 1760) 1 4 1 0 0 6 Pityogenes quadridens (Hartig, 1834) 0 0 3 0 0 3 Xyleborinus saxesenii (Ratzeburg, 1837) 8 4 6 2 0 20

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Yorkshire Coleoptera Checklist 2020

Xyleborus dispar (Fabricius, 1792) 0 1 8 1 0 10 Xyleborus dryographus (Ratzeburg, 1837) 0 0 2 0 0 2 Trypodendron domesticum (Linnaeus, 1758) 4 13 28 16 4 65 Trypodendron lineatum (Olivier, 1795) 0 1 4 3 0 8 Trypodendron signatum (Fabricius, 1792) 0 3 9 0 0 12 Hylesinus crenatus (Fabricius, 1787) 3 6 11 5 3 28 Hylesinus toranio (Danthoine in Bernard, 1788) 5 3 5 5 0 18 Hylesinus varius (Fabricius, 1775) 11 11 29 7 5 63 Hylastinus obscurus (Marsham, 1802) 2 7 10 0 1 20 Kissophagus hederae (Schmitt, 1843) 0 3 1 0 0 4 Pteleobius vittatus (Fabricius, 1792) 1 0 4 1 0 6 Hylastes ater (Paykull, 1800) 7 6 8 2 3 26 Hylastes brunneus Erichson, 1836 2 4 1 2 1 10 Hylastes cunicularius Erichson, 1836 0 2 0 1 0 3 Hylastes opacus Erichson, 1836 6 2 4 0 0 12 Hylurgops palliatus (Gyllenhal, 1813) 10 19 12 4 0 45 Phloeotribus rhododactylus (Marsham, 1802) 8 5 7 0 0 20 Polygraphus poligraphus (Linnaeus, 1758) 0 1 0 0 0 1 Tomicus piniperda (Linnaeus, 1758) 7 52 11 10 1 81 Xylechinus pilosus (Ratzeburg, 1837) 0 1 0 0 0 1 Platypus cylindrus (Fabricius, 1792) 1 1 3 0 0 5

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