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Coleoptera: Tenebrionoidea: Salpingidae) from Baltic Amber (Tertiary, Eocene

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Coleoptera: Tenebrionoidea: Salpingidae) from Baltic Amber (Tertiary, Eocene Baltic J. Coleopterol. 13(2) 2013 ISSN 1407 - 8619 Description of a second species of the narrow-waisted bark bee- tles (Coleoptera: Tenebrionoidea: Salpingidae) from Baltic amber (Tertiary, Eocene) Vitaly I. Alekseev Alekseev V. I. 2013. Description of a second species of the narrow-waisted bark beetles (Coleoptera: Tenebrionoidea: Salpingidae) from Baltic amber (Tertiary, Eocene). Baltic J. Coleopterol., 13(2): 137 - 140. The second representative of the family Salpingidae Leach, 1815 and first extinct species of rostrate narrow-waisted bark beetles is described from Baltic amber. Salpingus henricusmontemini sp.nov. differs from other representatives of the genus by the length of rostrum, by the shape of pronotum, by the structure of antennae and punctuation of body surface. Key words: Coleoptera, Salpingidae, Salpingus henricusmontemini, new species, Baltic am- ber, Tertiary, Eocene. Vitaly I. Alekseev. Department of Zootechny, Kaliningrad State Technical University, Sovetsky av. 1. 236000 Kaliningrad, Russia. E-mail: [email protected] INTRODUCTION MATERIALS AND METODS The family narrow-waisted bark beetles The amber piece with the beetle inclusion was (Salpingidae Leach, 1815) includes about 300 ex- obtained from a commercial source in Kaliningrad tant species from 45 genera (Slipinski, Leschen, and registered in the collection of Christel and Lawrence, 2011). The family is poorly known Hans Werner Hoffeins in May 2013. Photos were taxonomically. Two extant genera belonging to taken with a Zeiss AxioCamICc 3 digital camera Salpinginae were reported from Baltic amber: mounted on a Zeiss Stemi 2000-stereomicroscope. Salpingus Illiger, 1802 and Lissodema Curtis, 1833 (Klebs, 1910). One extinct monotypical ge- nus is described from Baltic amber (Alekseev, SYSTEMATIC PART 2013) and one from lowermost Eocene French amber (Kirejtshuk et Nel, 2009). In the present Family Salpingidae Leach, 1815 paper, a new species of Salpingidae, from Eocene Subfamily Salpinginae Leach, 1815 Baltic amber, assigned to the recent genus Genus Salpingus Illiger, 1801 Salpingus Illiger, 1801, is described and illus- Salpingus henricusmontemini sp.nov. trated. (Figs. 1-4) 137 Anichtchenko A. Material examined: holotype Nr. 25-3 [CCHH], impressions on elytra. S. henricusmontemini sex unknown. The beetle inclusion is slightly sp.nov. can be reliably differentiated from the damaged: the small apical area of the head is black- other species of the genus Salpingus based on ened, because of thermal processing of the am- the following combination of characters: five- ber piece in an autoclave. The surface of left elytra segmented antennal club, deep and oval basal is not well enough visible. Segments 6-11 of the foveae of pronotum, short rostrum, confused right antenna are lost. The beetle inclusion is punctuation on elytra and dark monotonous color preserved in a polished piece of transparent am- of body, rostrum and appendages. ber with a yellowish shade. The amber piece is embedded in polyester resin (measurements of Description. the piece are 11.5 x 7.0 x 5.0 mm). In the studied Body length: ca. 3.7 mm (from apex of rostrum to piece syninclusions are absent, but the initial apex of elytra); body length/maximum body width amber nugget has included two female speci- 2.5. Middle-sized, elongated, flattened, equipped mens of Diptera (Dolichopodidae and with a short rostrum (Figs. 1-2), unicolorous: Chironomidae). The examined specimen is cur- dorsal surface, underside, head, head append- rently housed in the private collection of Christel ages and legs dark brown. and Hans Werner Hoffeins (Hamburg, Germany). The type will be deposited at the Senckenberg Head prognathous, finely and sparse punctured Deutsches Entomologisches Institut in (distance between punctures wider than diam- Müncheberg, Germany (SDEI) as part of the in- eter of puncture), weakly flattened dorsoventrally, stitute’s amber collection for permanent preser- without constricted neck, subparallel-sided pos- vation. terior to eyes, developed into a rostrum anteriorly. Rostrum 1.6 times as long as wide between eyes. Type strata. Baltic Amber. Eocene. The head with eyes as broad as long including rosrum, slightly narrower than pronotum. The Type locality: Baltic Sea coast, Yantarny settle- temples 1/3 of eye length, the distance between ment [formerly Palmnicken], the Kaliningrad re- eye and antennal insertion equal to one eye gion, Russia. length. Eyes slightly protuberant, without bris- tles, finely faceted, oval. Maxillary palps with Diagnosis. The new species may be referred to elongate (about 3 time longer than wide) and the Salpingus Ill. due to a combination of mor- spindle-shaped last palpomere (Fig.4). Antenna phological characters, such as: well-developed 11-segmented (Fig.2), reaching humeri, with short (longer than wide) rostrum, which is not bordered and dense hairs on antennomeres. Antennal club laterally between eyes and antennal insertions; loose, 5-segmented: antennomeres 7-10 dilated, pronotum narrowed to base with basal impres- obconical; antennomere 11 ovate. The basal sions and without lateral denticulation; anten- antennomeres (1-6) thin, oblong, subequal in size. nae with loose club; elytra punctate-striate, shin- ing and naked . Salpingus henricusmontemini Pronotum irregularly and comparatively dense sp.nov. is easily distinguishable from the other punctured, obviously narrowed to base, elon- extinct amber species (Eopeplus stetzenkoi gate (proportion “pronotal length/maximum Kirejtshuk et Nel, 2009 and Protolissodema pronotal width” approximately 1.2). The punc- ulrikae Alekseev, 2013) by the presence of a ros- tuation of pronotum medially fine and sparse (like trum. The new species is similar to recent S. punctuation on the forehead), coarse and deeper ruficollis (Linnaeus, 1761) and S. caucasicus at margins (the distance between punctures less Reitter, 1905 in the presence of pronotal impres- than diameter of each puncture). Side margin of sions and by the form and comparatively fine pronotum bordered anteriorly, laterally in first punctuation of the elytra, to S. planirostris third and also finely bordered posteriorly. The (Fabricius, 1787) by the short rostrum and basal basal impressions of pronotum in form of two 138 New species of Platyderus Stephens, 1827 (Coleoptera, Carabidae) from North Spain elytral length/maximum width across the elytra - 1.5; the proportion elytral length/pronotal length - 3.0. Elytra with well-developed humeri, distinctly wider than pronotum, with confused rows of regular punctures (reaching from base to apex), in basal third punctuation is more confused. Scutellar striole present. The interspaces are ap- proximately 2-3 of each puncture diameter, the interstices are approximately 3-4 of the puncture diameter. The punctures are round, deep, 2-3 times larger than eye facet diameter. Basal trans- Figure 1. Salpingus henricusmontemini sp.nov. versal impression on elytra present in basal third, Habitus: dorso-lateral view, left side. shallow, but well visible. Epipleuron present, wid- est at base, reaching the second ventrite. Metathoracic wings fully developed. Legs (Fig.3) with tarsal formula 5-5-4. Tarsi and tibiae slender, visibly pubescent. Claws well-de- veloped, long (approximately 1/3 of the ultimate tarsomere length), simple. All tarsomeres not bilobed. Last metatarsomere equal in length to first metatarsomere and approximately as long as 2 and 3 metatarsomeres combined. Tibiae subequal in length, non-widened distally. Femora Figure 2. Salpingus henricusmontemini sp.nov. comparatively wide, rounded. Pro- and Habitus: dorso-lateral view, right side. mesocoxae cylindrical, rounded, metacoxae trans- verse. The procoxal separation is not visible, meso- and metacoxae distinctly separated. Abdomen with five ventrites. The first ventrite 1.3 times longer than each of the subequal in Figure 3. Salpingus henricusmontemini sp.nov. Habitus: ventro-lateral view. distinct, deep, rounded and closed slightly trans- verse foveae. Scutellum rounded, almost trian- gular. Base of prothorax narrower than the com- bined elytral bases. Prosternum and mesoster- num irregularly punctured like pronotum. Elytra (Fig.2) shining, elongate, covering abdo- men; maximum width of elytra (1/2 of its length) – Figure 4. Salpingus henricusmontemini sp.nov. 1.5 mm; length of elytra – 2.3 mm. The proportion Habitus: frontal view 139 Anichtchenko A. length 2-4 ventrites; the last ventrite is the short- Kirejtshuk A.G., Nel A. 2009. New genera and est. All ventrites distinctly punctured. Ventrites species of Cucujiformia (Coleoptera, 2-4 with isolated long erect hairs in median part. Polyphaga) from lowermost Eocene French Last ventrite and last visible tergite with narrow amber. Denisia 26: 103-118. combs of hairs along distal margins. Klebs R., 1910. Über Bernsteineinschlüsse im Derivatio nominis. The epithet of this new spe- allgemeinen und die Coleopteren meiner cies is devoted to Henricus Montemin (Prussian: Bernsteinsammlung. Schriften der Erkus Mants, Herkus Monte; Lithuanian: Herkus Physikalisch-ökonomischen Gesellschaft Mantas; German: Heinrich Monte), who was the zu Königsberg i. Pr, 51: 217-242. most famous leader of the Great Prussian Upris- ing (1260-1274) against the Teutonic order. Slipinski S.A., Leschen R. A. B., Lawrence J. F., 2011. Order Coleoptera Linnaeus, 1758. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An DISCUSSION outline of higher-level classification and survey of taxonomic richness. Zootaxa, The genus Salpingus Ill. includes more as
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