© Zoological Institute, St.Petersburg, 2001
Ca_sebearers from Caucasus (Lepidoptera: Coleophoridae)
V.V. Anikin & V.I. Shchurov
Anikin, V.V. & Shchurov, V.1. 2001. ·casebearers from Caucasus (Lepidoptera: Coleo phoridae). Zoosystematica Rossica, 10(1): 171-179. A checklist of the Coledphoridae of Caucasus (North Caucasus of Russia, Georgia, Arme nia, Azerbaijan) is presented. 127 species of 37 genera are recorded. Among them, 43 spe cies are new to Caucasus, 22 to Azerbaijan,4 to Armenia, 4 to Georgia, and 11 to Russia. Females of Amselghia azishtella Anikin and Ecebalia monoceros (Falk) are described for the first time.
V. V. Anikin, Department of Biology, Saratov University, Astrakhanskaya ul. 83, Saratov 410026, Russia, e-mail: [email protected]
V.1. Shchurov, Krasnodar Forestry Board, ul. Krasnaya 22, Krasnodar 350640, Russia, e mail: [email protected]
The casebearer fauna of Caucasus is poorly The names of collectors, whose material is known. Fuchs (1903) described 4 species from _ kept in the Zoological Institute, St.Petersburg, Erevan, of which 3 remained nomina dubia and are abbreviated as follows: Re. - S.Ya. Reznik, were not included in our review: Coleophora Rj. - M.A. Rjabov, T. __: V.V. Tikhonov, Z. - angustiorella Fuchs, 1903, ·c. fuchsiella Oud A.K. Zagulajev. Extensive material from jeans, 1971 (metalicella Fuchs, 1903, nom. Kabardino-Balkaria was collected by A.A. praeocc.), and C. debilella Fuchs, 1903. Filip Bolov (!)is name is abbreviated as B.). The re jev (1926) recorded 19 species from the Stav maining material was collected by _Shchurov in ropol Terr.; they are included in our list,,except Krasnodar Terr., Adygeya,.Stavropol Terr. and• for "Coleophora punctulateJla Z.", which is, Kabardino-Balkaria using a 240-v quartz-lamp probably a misidentification; for 7 species, we in 1996-2000 and by Anikin in Krasnodar Terr. examined Filipjev's specimens and confirmed in 1999. The material collected by' Bolov, his identifications. A checklist of 30 species Anikin and Shchurov is now deposited in the fromCaucasus was published by Patzak ( 1977) Zoological Institute, St.Petersburg, except for and 20 species were recorded from North Cau some duplicate specimens collected by Shchu casus by Falkovitsh & Jalava (1997). Some ad rov and kept in his collection. ditfonal records or descriptions of new species In total, 127 species of 37 genera are. re were published . by Reznik (1977; 1989), corded. Among them, 43 species (marked Baldizzone .& Patzak (1991), Falkovitsh with *) are new to Caucasus. Four species are ( 1991 ), Baldizzone ( 1994), and Anikin ( I 998). new to Armenia: Orthographis brevipalpella, In the check-list, the information is given in Damophila trifolii, Nemesia chalcogramella, the followingsequence: species name (with the Klinzigedia wockeela. Twenty-two species are most often used or the most important syno new to Azerbajian: Casas zernyi, Frederickoe nyms), reference to the publications in which nigia jlavipennella; Haploptilia hemerobiella, this species was mentioned from Caucasus pamophila mayrella, D. deauratella, · D. al and/or material examined, distribution outside cyonipennella; Symphypodaparthenica, Coleo Caucasus. phora albidella, C. bedella, Apista gallipen Countries and administrative subdivisions nella, Razowskia 'coronillae, Eupisia ornati are given in the list as follows: Ar, Armenia; pennella, E. lixella; Bourgogneja pennella, Az, Azerbaijan; Gg, Georgia;· RU, Russia Perygra alticolella, P. taeniipennella, Ecebalia (RuR, Rostov Prov.; RuKr, Krasnodar Terr.; therinella, E. versurella,. E. lassella, 1onescu RuA, Adygeya; RuKB, Kabardino-Balkaria; mia isomoera, Klinzigedia onopordiella,K. im R.uS, Stavropol Terr.; RuD, Daghestan). Lo plicitella. Four species are new to Georgia: calities are given in Fig. 1. Amselghia felixella, Damophila deauratella, 172 V. V.Anikin& V.I.Shchurov:ColeophoridaeofCaucasus•ZOOSYST. ROSSICA Vol. 10
48
I Russia II Georgia III Armenia IV Azerbaidzhan 0 150km TURKEY\
42
Fig. l. Map of Caucasus, localities: 1, Krasnodar, Lenin vii.; 2, Starokorsunskaya; 3, Seversk Distr., vie. Ubinskaya, Mt. Sober-Oachokh; 4, Mt. Papai; 5, Anapa, Mt. Lysaya; 6, Sukko; 7, Bolshoi Utrish; 8, Natukhaevskaya; 9, Gelendzhik Distr., Betta; 10, Sochi, Lazarevskoe Distr., Lygotkh; 11, Sochi; 12, vie. Krasnaya Polyana, Achishkho Range; 13, Ap- · sheronsk Distr., Chernigovskaya; 14, Eysk Distr., Shabelskoye; 15, Krnsnogar env., Enem; 16, Azish-tau Range; vie. Kamyshanova Polyana; 17, Lagonaki Plateau; 18, vie. Krasnaya Polyana, Aibga Range;.19, Tamansk Peninsula, Mt. Polyvadina, vie. Veselovka; 20, Tuapse Distr., Central Caucasus mt., Mt. Lysaya; 21,Novorossiisk Distr., Shirokaya Balka; 22, Kabardino-Balkar Nature Reserve, Bezengi River; 23, Sovetskoe; 24, Sukon gorge; 25, Prokhladninsk Distr., Novo-Poltavsk; 26, Proletarskoe; 27, Planovskoe; 28, Urvan River; 29, Zolsk Distr:;' Sarmakovo; 30, Nalchik; 31, vie. Makhaehkala, Mt. Talgi; 32; Derbent; 33, Akhtynsk Distr., Kurush; 34, Bala Hany; 35, Samursk Nature Reserve; 36, Batumi; 37, Abkhazia, Danisparetsli; 37, lien; 38, Megri; 40, Belab; 41, Zakatalsk Nature Reserve, Belokany; 42, Lenk- oran Distr., Alekseevka; 43, Astarinsk Distr., Alasha; 44, Lerik; 45, Gosmaljan; 46, Girkan Nature Reserve, Talysh Mts, Avrora; 47, Kubinsk Distr., Dustaur; 48, Kuba; 49, Chananab.
Coleophora albidella, Suireia alnifoliae. In the list, the modern nomenclature is used Among 50 species new to N Caucasus, 11 are (Anikiit & Falkovitsh, 1997; Falkovitsh, in new to the fauna of Russia: Casas zernyi, press). Haploptilia drymophila, H. kronee/la, Tuber culia albitarse/la, Amselghia rectilineella, Ar Check-list dania colutella, Multicoloria berlandella, M. polonicella, Klimeschja rudella, Ecebalia mo *Casas zernyi (Toll, 1944). Az: Belab, 24.V. 1957, 4 cl, 1 'i? (Z.). RuKr: Abrau Peninsula,. dry steppe, noceros, Casignetella trodlodytella. New re 27.V.2000, 2 'j?.- S Europe,N Africa,Syria, Iran. cords from provinces of Russia outside Cau Sclcriductia ochripennella (Zeller, 1849). RuKr: casus are marked with"!". Seversk · Distr., Ubinskaya, Mt. Papai, · 700 m, ZOOSYST. ROSSICA Vol. 10• V.V.,4.nikin & V.1.Shchurov:Coleophoridaeof Caucasus 173 8.VI.1996, 1 d'; Gelendzhik Distr., Betta; 13.IX.1996, 1 0. chamaedriella (Bruand, 1852). RuKB: Teberda ; Seversk Distr., Ubinskaya, Mt. Papai laearctic, introduced in N America and New Zealand. 800 m, 8.Vl.1997, l er. RuKB (Falkovitsh & Jalava, D. dcauratclla (Lienig & Zeller, 1846). Az: Nakhi 1997): Teberda (Patzak, 1977). RuS (Filipjev, 1926): � chevan Prov., Chananab, 28.IV.1955, Id'(Z.); Gel-Gel, Russia (Saratov, Ulyanovsk,"!Kaluga, and !Ore! Prov., 1700 m, 19.VI.1964, 2 d' (Z.). Gg: Abkhazia, Danis Ural, Altai), Europe, !tan, Mongolia. paretsli, 26.Vl.1969, 1 d' (Z.). RuKr: Anapa, Mt. *C. albidella (Denis & Schiffermliller, 1775). Az: Lysaya, steppe, 9.Vl.1998, I <;>; Sochi, Aibga Range, Zakatalsk Nature Reserve, forest zone, 12.Vl.1962, I o' 1850 m, alpine meadows, 10.VII.1999, 1 rl. RuKB: (Z.). Gg: Adzharia, Kintrishsk Nature Reserve, Hiknara, Upper Balkar, 21.Vl.1987, 1 d' (Z.); Kabardino-Balkar 7.Vll.1974, I er(Z.). RuKr: Seversk Distr., Ubinskaya, Nature Reserve, 3.V.1997, Id'(B.); Kenzhe, 6.Vl.1997, Mt. Papai, oak-beech forest, 700 m, 11: V.1996 and 3 d', l 'i' (B.). RuS (Filipjev, 1926): Kislovodsk (Falk 8.Vl.1997; 2 er;Sochi, Lygotkh, oak forest, 450 m, ·1 r;>. ovitsh & Jalava, 1997). - Russia (Astrakhan, Vol RuA: Sober-Oachokh Mt., oak forest, 450 •. m, gograd, Saratov, Ulyanovsk, Samara, Kaluga, Tula, 8.Vl.1998, 1 er. - Russia (Saratov Prov., Ural, Altai). Smolensk, and Novosibirsk Prov., Altai, Ural). Transpa Transpalaearctic. laearctic, introduced in N America. C. kuehnella (Goezi:;, 1783) (= pal/iatella Zincken, . *D. alcyonipennella (Kollar, 1832) (= cuprariella 1813; = pallipennella Treitschke, 1833). RuKB: Te Zeller, 1847). Az: 20 km E ofLerik, 4-5.VIl.1964, 8 d' berda (Patzak, 1977). -,, Russia (Saratov, Samara, and (Z.); Lenkoran Distr., Alekseevka, 29.Vl.1962, 1 d' !Kaluga Prov., Ural); Europe, Asia Minor. (Z.); Kuba, garden, 21.VII.1962, 1 er(Z.); Talysh Mts, C. ibipeunella Zeller, 1849 (= nemon,m Heinemanh, Avrora, 12.VIl.1964, 2 er (Z.). RuKr: Adler, 1854). RuA: Sober-Oachokh Mt., oak forest, 450 m, 28.Vl.1978, 1 d', 2 'i' (Re.); Seversk Distr., Ubinskaya, 8. VI.I 998, I d'. - Russia (Saratov, Ulyanovsk, and Mt. Papai, 800 m, 18-19.Vl.1997, 1 <;>. RuKB: Upper Samara Prov., Ural); Europe, N Africa,Lebanon. Balkar, l.VIII.1989, 1 er(Z.); Kabardino-BalkarNature *C. betulella Heinemann & Wocke, 1877. RuKr: Reserve, 2200 m, 23.Vl.1990, 2 d' (Z.); Planovskoe, Seversk Distr., Ubinskaya, Mt. Papai, 800 m, forest� 8.VI. and 9.VII.1997, 3 d' (B.). RuS (Filipjev, 1926). steppe, 18-19.VI.1998, I <;>; Mt. Markotkh, 460 m, RuD: Kurush, 2700 m, 5.VIl.1990, 10 d' (Z.). - Russia 28.Vl.2000, 2 er. - Russia (Samara and !Kaluga Prov.); (Astrakhan, Volgograd, Saratov, Ulyanovsk, Samara, Europe. !Kaluga, Novosibirsk, and Irkutsk Prov., Altai, Ural, *C. zelleriella Heinemann, 1854 (= pannonice/la Yakutia). Transpalaearctic, introduced in N America, Gozmany, 1956; = p/atyphyllae .Oku, 1965). RuKr: Australia, and New Zealand. Seversk Distr., Ubinskaya,'"Upper reaches of Papai D. frischella (Linnaeus, 1758) (= auronitella Toll, River, forest-steppe, 700 m, 18-19.Vl.1998, 3 d', I.<;>; 1962). Ar: Erevan (Patzak, 1977). Gg: Tbilisi (Patzak, Anapa, Mt. Lysaya, oak forest,7. and 28.VII.1999, 3 cl; 1977). RuKr: Mzymta River, 1750 m, 30.VII.2000, 2 Mt. Aibga, 950 m, 26.Vll.2000, 2 er.-· Russia (Saratov d'. RuKB (Falkovitsh & Jalava, 1997): Teberda Prov., Far East); Europe, Iran, Japan. (Patzak, 1977); Kabardino-Balkar Nature Reserve, 2200 C. bedella Falkovitsh, 1976. Az: Kuba, 23.Vll.1962, m, 22.Vl.1990, 2 d' (Z.). RuS (Filipjev, 1926). -Russia I er(Z.). - Iran. (Saratov and Ulyanovsk Prov., Altai). W Palaearctic. *C. currucipennclla Zeller, 1839 (= nemorum He D. trifolii Curtis, 1832 (= melilotel!a Scott, 1861 ). inemann, 1854; = cristinae Capu�e, 1971). RuKr: Ar: Erevan, the city zoo, IO.Vl.1960, I 'i' (Z.); 10 km Seversk Distr., Ubinskaya, Mt. Papai, forest, 700 m, NE Megri, Kaladash Gorge, l 5.VIl.1960, I 'i' (Z.). 18.Vl.1998, I <;>; Anapa, Mt. Lysaya, oak forest, RuKr: Krasnodar, Lenin, l.Vl.1998, l er; Anapa, Mt. 28.VII.1999, I er;Mt. Aibga, 950 m, 26.VII.2000, I o'. Lysaya, steppe, 9.VI.1998, I <;>. RuKB: Proletarskoe, - Russia (Ulyanovsk, Samara, !Moscow, !Belgorod, steppe, 16.VI.1998, I 'i' (B.). RuS: Kislovodsk (Falk !Kaluga, and !Novosibirsl (Z.); Lerik, stamm, 1839). RuKB: Proletarskoe, steppe, l 7.Vl.1998, l r;> (B.). (Filipjev, 1926; examined). - Russia 2.VIl.1964, 1 er(Z.). RuKB: Kabardino-Balkar Nature RuS Reserve, 8.VI.1980, I 'i' (Z.). -Russia (Saratov, Sam (Volgograd, Saratov, and !Novosibirsk Prov.); C and S ara, !Tula, and !Novosibirsk Prov.); Europe, Asia Mi Europe, Asia Minor. nor, Mongolia. ,- (Meyrick, 1891) (= Symphypoda parthenica Mctapista fcrruginea (Baldizzone, 1994). Ar cygnipennel!a Toll, 1956; = candidel/a Toll, 1959; = (Baldizzone, 1994). -Asia Minor, Iran. Toll, I 959). Erevan (Patzak, 1977). transcaspica Ar: Razowskia coronillae (Zeller, 1849). Az: Lerik, Az: Lerik, 5-6.Vl.1962, I d' (Z.). -Russia (Astrakhan, 2.VIl.1964, I d' (Z.). RuKr: Anapa, Mt. Lysaya, Volgograd, and Saratov Prov., Ural). S Palaearctic from 22.Vl.1999 and 15.VIl.2000, '850 m, 4 o'; Mt. Mark Algeria and Greece to Mongolia. otkh, 28.VI.2000, 4 d', I <;>. �uA: Sober-Oachokh Mt., *Ocdicaula serinipennella (Christoph, 1872). steppe, 8.VII.1998, I er; Azish-tau Range, 1400 ni, RuKB: Proletarskoe, steppe, 17.Vl.1998, I 'i' (B.). - 15.VII.1998, I d'. RuKB: Teberda (Patzak, 1977). - Russia (Astrakhan, Volgograd, Saratov, and Ulyanovsk Russia (Saratov, Samara, and Ulyanovsk Prov.). W Pa Prov., Ural). Transpalaearctic (southern part). laearctic. ZOOSYST. ROSSICA Vol. 10• V.V.Anikin & V.l.Shchurov:Co/eophoridaeof Caucasus 175 Stabilaria univittclla (Staudinger, 1880). Ar Eupista ornatipennella (Hubner, 1796). Az: Belab, (Baldizzone, 1994): � Crimea, Asia Minor, Turkmenis- 24.V.1957, 1 'i' (Z.); Gosmaljan, 29.Vl-1.VII.1964, 5 d', tan, Iran. . · · 4 'i' (Z.). Ru KB: Teberda (Patzak, 1977). RuS (Filipjev, Multicoloria partitella (Zeller, 1849). Az: Lenkoran 1926). - Russia (Astrakhan, Volgograd, Saratov, Sam (Reznik, 1989). -. Russia '(Volgograd, Saratov Distr., ara, Ulyanovsk, !Tula, and !Omsk Prov., Ur11l, Altai); Altai); Europe, Asia Minor. Middle and South Europe, Asia Minor, Middle Asia *M. conspicuella (Zeller, 1849). RuKr: Anapa, (Kopetdagh). Sukko, 14.VI.1997, larvae on Gailium, ex.I. 9.V.1998, 1 E. lixella (Zeller, 1849). Ar (Baldizzone, 1994): lien, d', I <;>. - Russia (Astrakhan, Volgograd, Saratov, and 22.V.1957, 2 'i' (Z.). Az: Kubinsk Distr., Dustaur, 1400 !Kaluga Prov., Ural, Altai). Transpalaearctic. m, 1 d', l <;> (Z.). RuA: Azisli-tau Range, 1400 m, M. vicinella (Zeller, 1849). Ar (Baldizzone, 1994): mixed forest, 16.VII.1998, l d'. RuKB (Falkovitsh & Erevan (Patzak, 1977). RuR (Reznik, 1977). - Russia Jalava, 1997): Nalchik, 3.VII. 1989, 1 d' (Z.); (Volgograd, Saratov, and Ulyanovsk Prov.). W Pa Kabardino-Balkar Nature Reserve, Bezengi River, 1700 laearctic (southernhalf). m, 15-16.VII.1989, 8 d', 2 'i' (Z.); same locality, M. berlandella (Toll, 1956). Az: Lenkoran (Reznik, 17.VIl.1989, 8 d', 1 'i' (B.). RuS: Kislovodsk, Essentuki 1989). RuKr: Black Sea coast, Gelendzhik Distr., (Falkovitsh & Jalava, 1997). RuD: Dagestan, Akhtynsk Betta, 12.V.1996, on Astragalus asper Jacq., 2 d'; same Distr., Kurush, 2600 m, I d' (Z.). - Russia (Saratov, locality, 14.IV.1998, ex.I. 29.V.1998, I <;>.- N Africa, S Samara, Ulyanovsk, and !Tula Prov.); Europe, Asia Mi Europe, Asia Minor, Middle Asia. nor. *M. polonicella (Zeller, 1865). RuKr: Anapa, Mt. E. caucasica (Stainton, 1867). RuKr: Gelendzhik Lysaya, 6.V.1997, on Onobrychis, ex I., 1.Vl.1997, 2 d', I Distr., Betta, 22.VI.1996, 2 <;>; Anapa, Mt. Lysaya, <;>;Anapa, Sukko, 24.V.1997, 1 <;>.-Poland, Germany. 22.Vl.1999, I c;>; Mt. Markotkh, 460 m, 26.VI.2000, 2 M. geghardella (Baldizzone, 1994). Ar (Baldizzone, d', 3 'i'. RuS (Filipjev, 1926; examined). GG: Akhalt 1994). sikhe (Stainton, 1867). M. symphistropha Reznik, I 976. Az: Lenkoran *Klimeschja rudella (Toll, 1944). RuKB: (Reznik, 1989). -Turkey. Kabardino-Balkar Nature Reserve, Bezengi River, M. dubiella (Baker, 1888) (= reisseri Rebe.l, 1926). 22.Vl.1990, I d' (Z.). -Europe. Az: Lenkoran, (Reznik, 1989). - S Europe, N Africa, *Bourgogneja pennella (Denis & Schiffermiiller, Asia Minor. 1775) (= onosmella Brahm, 1791). Az: Gosmaljan, I 'i', M. guttella Reznik, 1971. Az: Lenkoran (Reznik, ex.I. on Artemisia 10.VI.1988 (Re.). RuD: Akhtynsk 1989). -Middle Asia. Distr., Kurush, 2500 m, 4.Vl.1994, 1 'i' (Z.). - Russia M.�fuscociliella (Zeller, 1849) (= medicaginis Her (Samara and Ulyanovsk Prov., Ural). W Palaearctic rich-Schilffer, 1861). RuKr: Krasnodar, Anapa, (middle and southern parts). 12.Vl.1997, I d'. RuD (Reznik, 1977): Khodzhal *Perygra murinipennella (Duponchel, 1844) (= Makhi, 26.VI.1926, 1 d' (Rj.). - Russia (Astrakhan, otidipenne/la Hubner, 1817). RuKB: Kabardino-Balkar Volgograd, and Ulyanovsk Prov., Altai). W Palaearctic Nature Reserve, 28.V.1988, 1 'i' (Z.); same locality, (southern part fromItaly to Altai). 2200 m, 22.Vl.1990, I d' (Z.). - Russia (Far East, Al M. ditella (Zeller, 1849) (= anatolica Toll, 1952). Ar tai); Europe. (Baldizzone, 1994). RuKB: Teberda (Patzak, 1977); *P. alticolella (Zeller, I 849) (= caespititie/la auct.). Karagach, 27.V.1998, I 'i' (B.). -' Russia (Astrakhan, Az: Kuba, 16.Vll.1962, 1 d' (Z.). RuKr: Krasnodar, Volgograd, Saratov, Samara, and Ulyanovsk Prov·., Lenin, 30.VIl.1996, I c;,. RuA: Azish-tau Range, 1400 Ural); Europe, N Africa, Middle Asia, Iran, Mongolia. m, meadow, 5.VIl.1996, I c;>. RuKB (Falkovitsh & M. cracella (Vallot, 1835). RuR (Reznik, 1977). - Jalava, 1997). - Russia (Saratov, Samara, Ulyanovsk, Russia (Volgograd and Saratov Prov.); S Europe. and !Kaluga Prov., Altai, Chukotka). Holarctic. *Suireia milvipennis (Zeller, 1839). RuKr: Seversk *P. caespititiella (Zeller, 1839). RuKr: Seversk Distr., Ubinskaya, Mt. Papai, 18.VI.1998, I <;>. RuA: Distr., Ubinskaya, Mt. Papai, 800 m, steppe, 8.VI.1996, Sober-Oachokh Mt., oak forest, 450 m, 26.IV.1998, 3 d'. RuA: Azish-tau Range, 1400 m, alpine meadows, ex.I. on Carpinus betulus, 27.V.1998; I d'; same local 13.Vl.1999, I d' .. � Russia (Samara and Ulyanovsk ity, 8.Vl.1998, 1 d'. - Russia (Leningrad, Ulyanovsk, Prov.); Europe. !Kaluga, and Irkutsk Prov.). Transpalaearctic. P. glaucicolella (Wood, 1892). Gg: Tbilisi (Patzak, *S. badiipennella (Duponchel, 1843). RuKB: Nal 1977). RuKr: Krasnodar, Lenin, 23.V and 1.Vl.1996, 2 chik, Nalchik River, 22.Vl.1986, I 'i' (Z.); Nalchik, bo d'; Seversk Distr., Ubinskaya, Mt Papai, open wood tanical garden, 1.VIl.1986, 1 d' (Z.). - Russia (Samara land of juniper, 1 l.Vl.1996, I d'; same locality, step Prov., Ural); Europe, Iran. pized meadow, I l.VII.1996, I d'. RuA: Azish-tau °' *S. alnifoliae (Barasch, 1934). Gg: Batumi, Dzho Range, 1400 m, meadow, 6.VII.1996, I d', 1 c;>; same Dzho, 9.VI.1974, 1 'i' (Z.). - Russia (!Kaluga Prov.); locality, 26.VII.1997, 2 d'. RuD: Samursk Nature Re Europe, Canada. serve, 17.V.1992, I 'i' (Z.). - Russia (Volgograd, Sara " S. limosipennella (Duponchel, 1842). Ar (Baldiz tov, Samara, Ulyanovsk, !Kaluga, !Smolensk, and No- zone, 1994): Erevan (Patzak, 1977). RuS (Filipjev, vosibirsk Prov.). Holarctic. 1926). � Russia (SamaraProv.); Europe, Middle Asia. P. taeniipennella (Herrich-Schliffer, 1855). Ar: Ere S. trigeminella (Fuchs, 1881). Ar (Baldizzone, van (Patzak, 1977). Az: Gosmaljan, old garden, 1994). -Europe, Turkey. 1.VII.1964, I d' (Z.). RuKB:'Teherda (Patzak, 1977). - * Argyractinia ochrea (Haworth, 1828). RuKr: Russia (Samara and Ulyanovsk Prov:). W Palaearctic. Krasnodar, Lenin, 6.VIII.1996, 1. <;>. - Russia (lower *Nemesia chalcogrammella (Zeller, 1839). Ar: Volga); Europe, Asia Minor: Araks River, Nyuvadi, 1.IX.1932, I 'i' (Rj.).- Europe. 176 V.V.Anikin & V.!Shchurov:Coleophoridae ofCaucasus•ZOOSYST. ROSSICA Vol. 10 Ecebalia therinella (Tengstriim, 1848). Az: *E. lassella (Staudinger, 1859) (= telavivella Toll, Zakatalsk Nature Reserve, forest zone, 6.VIII.1962, I 1942). Az: Lenkoran Distr., Alekseevka, 29.VI.1962, I d', I 'i? (Z.). RuKB (Falkovitsh & Jalava, 1997): Nal ,'i? (Z.); same locality, 11.V.1964, I cl' (Z.). - Europe, N chik, botanical garden, 26.VI.1989, I 'i? (Z.); Africa, Asia Minor, Canary Is. Kabardino-Balkar Nature Reserve, 1350 m, 11- E. salinella (Stainton, · 1858). Ar (Baldizzone, 30.VII.1989, 7 'i? (Z.); vie. Elbrus, 26.Vl.1996, 1 'i? (B.). 1994). - Europe. RuS: Essentuki (Falkovitsh & Jalava, 1997). - Russia *E. adspersella (Benander, 1939). RuKr: (Astrakhan, Volgograd, Saratov, Samara, !Kaluga, Gelendzhik Distr., Betta, 13 and 18.IX.1996,.2 <;?. - !Ore!, and Irkutsk Prov., Altai, Yakutia). Transpa Russia (Volgograq, Saratov, Samara, and Ulyanovsk laearctic. Prov.); Europe. *E. monoceros (Falkovitsh, 1975). RuKB: vie. Bu E. tamesis (Waters, 1929}. RuKB: Teberda (Pafzak, lungu, 8.VIII.1998, 1 d', I '?(B.) (femaleswere hitherto 1977). - Russia (Far East); Europe. unknown). - Mongolia. E. virgaureae (Stainton, 1857). RuKr: Sochi, forest E. vestianella (Linnaeus, 1758) (= laripennella Zet alpine zone of Aibga Range, 1780 mm, 6.VIII.1998, I' terstedt, 1839). Ar (Baldizzone, 1994). RuKB (Falk '?. RuKB: Teberda (Patzak, 1977). RuD: Dagestan, Ku ovitsh & Jalava, 1997): Teberda (Patzak, 1977); rush, 2500 m, 5.VIl.1990, I· cl' (Z.). - Russia (Vol Kabardino-Balkar Nature Reserve, 1700 m, gograd, Saratov, Ulyanovsk, and Irkutsk Distr., Altai); 15.VII.1989, 3 cl' (Z.); same reserve, Bezengi River, Europe, Kazakhstan. ' 22.VII.1997, 4 cl', 3 'i? (B.); same locality, 16.VII.1998, E. sp. (pr. persana Baldizzone, 1990). RuA: Azish I cl',2 'i? (B.); Sarmakovo, 17.VIIl.1989, I cl', l '? (B.). tau Range, alpine meadows, 1400 m, 13.VI.1999, I cl'. RuS (Filipjev, 1926). - Russia (Astrakhan, Volgograd, Casignctella absinthii (Heinemann & Wocke, · Saratov, Samara, Ulyanovsk, !Kaluga, !Tula, Ural, No 1877). RuKB (Falkovitsh & Jalava, 1997). - Russia vosibirsk, and Irkutsk Prov., Altai, Buryatia, Yakutia). (Saratov, Samara Distr.); Europe. Transpalaearctic. *C. troglodytella (Duponchel, 1843). RuKr: Sochi, E. squamosella (Stainton, 1856) (= podolensis Toll, Achishkho Range, 1800 m, 28, VI.I 998, I d'.- Europe. 1938; = sabulicola Benander, 1939). RuKr: Anapa, *C. argentula (Stephens, 1834) (= cothurnella Du Natukhaevskaya, 14.IX.1997, 1 d'. RuKB (Falkovitsh ponchel, 1843). RuKr: Sochi, Achishkho Range, 1800 & Jalava, 1997): Kabardino-Balkar Nature Reserve, m, 28.Vl.1998, I cl'. - Russia (Astrakhan, Volgograd, 1700 m, 15.VIl.1989, 1 cl', I '? (Z.); same locality, Saratov, Samara, Ulyanovsk, and Novosibirsk Prov., 17.Vll.1998, 1 'i? (B.). - Russia (Altai, Irkutsk Prov., Ural, Bashkiria, Altai). W Palaearctic. Chukotka).Transpalaearctic. *C. artemisiella (Scott, 1861). RuKB: Kabardino0 *E. gaviaepennella (Toll, 1952). Ar (Baldizzone, Balkar Nature Reserve, IO.Vl.1980, 1 d' (Z.). - Russia 1994). RuKB: Sukon gorge, 6.VIII.1997, I cl' (B.); Bu (Astrakhan, Volgograd, Saratov, Samara, and Ulya lungu, 8.VIII.1998, 2 cl' (B.). - Russia (Astrakhan, Vol novsk Prov., Ural, Bashkiria). Transpalaearctic. gograd, Saratov, Samara, Ulyanovsk, !Kaluga, and No *C. granulatella (Zeller, 1849)'(= artemisiae Muh vosibirsk Prov., Altai, Buryatia). W Palaearctic (south lig, 1864). RuKr: Gclendzhik Distr., Betta, 13.IX.. ern part), Mongolia. 1996, I d'. RuKB: Nalchik, botanical garden, LVII. 1986, I d' (Z.). - Russia (Saratov, Samara, and !Kaluga E. eichleri (Patzak, 1977). Ar: Erevan (Patzak, 1977). - Russia (Volgograd Prov.), Middle Asia. Prov., !Bashkiria, W Siberia); Europe, Mongolia. (Zetterstedt, 1839) (= C. inulae (Heinemann & Wocke, 1877). RuS E. sternipennella jlavagine/la (Filipjev, 1926). - Europe. Liening & Zeller, 1846). RuKr: Krasnodar, Lenin, 20.VII.1996, 1 <;?; same locality, 31.VIII.1996, I cl', l <;?; C. inulae transcaucasica (Patzak, 1977). Ar: Gami (Patzak, 1977). same locality, 21.Xl.1996, ex.I., 7.X.1997, I'?. RuKB: Teberda (Patzak, 1977); Kabardino-Balkar Nature Re C. peribcnanderi (Toll, 1943) (= benanderi Toll, serve, Bezengi River, 19.VII.1997, 3 cl', I '? (B.); same 1942). RuKr: Krasnodar, Lenin, I 1.VU998, 3 9. locality, l 7.VII.1998, I cl' (B.). - Russia (Astrakhan, RuKB: Novo-Poltavsk, 7.V.1998, 2 'i? (B.). RuS: Es Volgograd, Saratov, Samara, Ulyanovsk, !Tula, and sentuki (Falkovitsh·-& Jalava, 1997). - Russia (Astra !Kaluga Prov., Ural, Altai). W Palaearctic. khan, Saratov, !Kaluga, and !Tu.la Prov.); Europe. *E. versurella (Zeller, I 849) (= agricole/la Fuchs, C. tamara (Baldizzone, 1994)'.' "Caucasus mer." 1886; = amaranthivora Oku, 1965). Az: Zakatalsk Na (Baldizzone, 1994). ture Reserve, forest zone, 10.VI.1962, I cl' (Z.); Lenk C. gardcsanella (Toll, 1953) (= machinella Bradley, oran Distr., Alekseevka, 29.Vl.1962, I cl' (Z.). RuKr: 1971). Ar (Baldizzone, 1994). - Russia (Saratov Prov.); Adler, 29.VI.1972, 3 '? (Re.); Krasnodar, Lenin, Europe. 11.Vl.1999, I <;?. RuKB: Nalchik, botanical garden, ·C. nutantella (Muhlig & Frey, 1857). Ar (Baldiz 23.Vl.1989, 1 '? (Z.); Sovetskoe, 19.VI.1997, Id' (B.); zone, 1994). - Russia (Altai,. Novosibirsk Prov.); Kabardino-Balkar Nature Reserve, Bezengi River, Europe, Asia Minor. 5:VI.1998, 2 cl', 5 'i? (B.); Proletarskoe, 16.Vl.1998, I cl' C. graminicolella (Heinemann, 1876). Gg: Tbilisi • (B.). RuS (Filipjev, 1926). The species is very common (Patzak, 1977). RuD: vie. Makhachkala, Mt. Talgi, in the region in anthropogenic landscapes. - Russia (As 13.V.1991, I 9 (Z.). - Russia (UlyanovskProv.); Europe. trakhan, Volgograd, Saratov, Samara, Ulyanovsk, C. dianthi (Herrich-Schaffer, 1855). Ar (Baldizzone, !Kaluga, !Orel, and Novosibirsk Prov., Ural, Altai). Ho 1994): lien, 22.V.1957, I cl' (Z.).-- Russia (Volgograd; larctic; re·corded also fromArgentina. Saratov, and Samara Prov.). W Palaearctic. *E. pseudolinosyris Kasy, 1979. RuKr: Anapa, Mt. *C. pseudociconiella (Toll, I 952). RuKr: Gelen Lysaya, vie. Su-Psekh, 14.IX.1999, 4 cl', I <;?. - Russia dzhik Distr., Betta, 22.VII.1997, I cl'. - Russia (Vol (Volgograd, Saratov, Samara, and Ulyanovsk Prov., Al- gograd, Saratov, and !Novosibirsk Prov.); S Europe, tai, Baikal); Europe (southern part). Asia Minor. ZOOSYST. ROSSICA Vol. 10• V.V.Anikin & V.J.Shchurov:Coleophoridae of Caucasus 177 C. majuscula Falkovitsh, 1991. Az: Lenkoran (Falk K. implicitella (Fuchs, 1903). Ar: Erevan (Fuchs, ovitsh, 1991). 1903). Az: Gosmaljan, old garden, I .VII. 1964, 1 o'(Z.). C. galbulipennella. (Zeller, 1838) (= otitae Zeller, RuKr:- vie. Gelendzhik, Mt. Markotkh, 28.VI.2000, 2 1838). Ar (Baldizzone, 1994). RuKr: Gelendzhik o'.RuS (Filipjev, 1926; examined). Distr., Betta, 13.I:X.1996, 2 o'.- Russia (Ulyanovsk and !Kaluga Prov., Ural); Europe, Asia Minor, Iran. Description of hitherto unknown females *C. millefolii(Zeller, 1849). RuKB: Kabardino-8al kar Nature Reserve, 8ezengi River, 9.VIl.1998, 1 o' (B.). - Russia(Ural); C and S Europe, North Africa. Amselghia azishtella Anikin, 1998 *C. striatipennella·(Tengstrtim,(1848)). RuKB: Ur (Figs 2, 3) van River, 11-12.V.1997, 5 o', 5 9 (B.). -Russia(Sara tov, Samara, Ulyanovsk, !Kaluga, !Tula, !Ore!, Novosi Description. Wing expanse 12-13 mm. Head, birsk, and Magadan Prov., Altai). Transpalaearctic. thorax and tegula light ochreous. Labial palpus Ionescumia clypeiferella (0. Hofmann, 1871). whitish ochreous; 2nd segment 1.3 times as RuKB: Teberda (Patzak, 1977). RuS (Filipjev, 1926; examined). - Russia (Astrakhan, Volgograd, Saratov, long as diameter of eye; 3rd segment OA times as long as 2nd. Basal antennal segment with Samara, and Ulyanovsk' Prov., Ural); Europe, Kazakhstan. .. scale tuft; flagellum whitish ochreous. Fore *I. isomoera (Falkovitsh, 1972). Az: Yardumly, wings ochreous brown, apex with brownish 27.VIII.1962, 2 o'(Z.). - N Africa, Spain, Asia Minor, yellow scales. Hind wings grey; fringesas col Uzbekistan, Mongolia. oured as wing. *Carpochena trientclla (Christoph, 1872) (= pili Female genitalia (Fig. 2). Papillae anales cornis Rebel, 1914; = ussuriella Caradja, 1920). RuKB: short, wide, with lower long bristles. Apo Planovskoe, 20.V.1997, 1 o'(B.). - Russia (Volgograd, Saratov, and Ulyanovsk Prov., Far East). S Palaearctic, physes posteriores 2.5 times as long as apo fromHungary to the Far East. physes anteriores. Subgenital plate trapezoid, C. unipunctella (Zeller, 1849). RuKB: Sovetskoe, narrower than in A. balkara Flkv. & Jalv., 19.Vl.1997, I o'(8.). RuS: Essentuki (Falkovitsh & along posterior margin short bristles. Antrum Jalava, 1997). - Russia (Saratov, Ulyanovsk, and Sam cup-like, moderately wide, with poorly ara Prov.); Europe(except northernpart). sclerotized, roundly-angulate protuberance. C. acqualclla (Christoph, 1872) (� heratella Toll & Upper part of ductus 0.8 times as long as spiny Amsel, 1967). RuS: Pyatigorsk (Patzak, 1977). - Russia portion; spines small and dense; lateral a�d (Volgograd and Saratov Prov., Ural); Kazakhstan, Mid central rods well-defined. Next portion of duc dle Asia, ln!,n, Afghanistan. tus only with central rod and equal in length to C. binotapennclla (Duponchel, 1843). RuKr: No vorossiisk, IO.Vlll.1992, 1 9 (T.); Novorossiisk Distr., previous. Signum large, basal part 30 mm, api- Shirokaya Balka, 24.VIII.1999, 2 o'. RuKB: Teberda cal part pointed. · · (Patzak, 1977). - Russia (Astrakhan, Volgograd, Sara Abdominal tergites (Fig. 3). 1st abdominal tov, Samara, Ulyanovsk, and Novosibirsk Prov., Ural); tergite without spinelets; patches of spinelets Middle and S Europe. on 2nd tergite scattered, about 1.2 times as *C. squalorclla (Zeller, 1849) (= delibutella Chris wide as long. toph, 1872). RuS (Filipjev, 1926; examined). - Russia Comparison. The female of this species is (Astrakhan, Volgograd, Saratov, and Ulyanovsk Prov., closely related to that of A. balkara Flkv. & !8ashkiria, Ural); Europe, Kazakhstan. Jalv. The main differences are as follows: fore C. armeniae (8aldizzone & Patzak, 1991). Ar: Ere wings darker; subgenital plate trapezoid; sec van (Baldizzone & Patzak, 1991). - Russia (Astrakhan Prov.); southern part of E Europe, Middle Asia, Asia tion of ductus bursae considerably longer; sig Minor. num large.· *C. salicorniae (Heinemann & Wocke, 1877). Ecebalia monoceros (Falkovitsh, 1975) RuKr: Krasnodar, Lenin, 3 I.VIII.1996, I <;?;Novorossi isk Distr., Shirokaya 8alka, 26.VIIl.1999; 2 o', I 9- (Figs 4, 5) RuKB: Planovskoe, 26.VlII.1997, 1 9 (8.). - Russia _ (Astrakhan, Volgograd, Saratov, Samara, Ulyanovsk; Description. Wing expanse 15 mm. Head, and Novosibirsk Prov.). W Palaearctic. thorax and tegula brownish. Labial palps on in Klinzigcdia phlomidis (Stainton, 1867). Ar (8aldiz ner side whitish grey, on outer side with dark zone, 1994). - Russia (Volgograd Prov.); Asia Minor, brown scales; 2nd segment 1.4-1.5 times as Turkmenistan, Iran, Syria. long as diameter of eye and 1.5-1.6 times as *K. wockcclla (Zeller, 1849). Ar: Erevan, the city long as 3rd segment. Basal segment of anten zqo, 20.V.1955, 1 o'(Z.). - Russia(Samara, Ulyanovsk, nae simple, brownish; flagellum whitish with and Tula Prov.); Europe, Asia Minor. dark brown rings. Fore wings brownish grey K. onopordiclla (Zeller, I 849). Ar (8aldizzone, with brownish yellow streaks along veins and 1994). Az: Gosmaljan, old garden, 1.VIl.1964, I o'(Z.). whitish scales between them. Surface of wing RuKB: Teberda (Patzak, 1977). RuS: 100 km NW of · with not compacted dark brownish scales. Hind Derbent, 26-28.V.1994, 2 9 (T.). Europe, Asia Minor. wing and cilia brownish grey. 178 V. V.Anikin & V.I.Shchurov:Coleophoridae o/Caucasus•ZOOSYST. ROSSICA Vol. 10
Figs 2-5. 2, 3, Amselghia. azishtellaAniki n, female: 2, genitalia; 3, anterior abdominal tergites; 4, 5, Ecebalia monoceros (Falkovitsh), female: 4, genitalia; 5, anterior abdominal tergites. ZOOSYST. ROSSICA Vol. 10• V.V.Anikin & V.I.Shchurov:Coleophoridae a/Caucasus 179 Female genitalia (Fig. 4). Apophyses posteri Anikin, V.V. & Falkovitsh, M.I. 1997. On the case ores 2.2 times as long as apophyses anteriores; bearer fauna of the lower Volga region (Lepidop all apophyses twisted towards their ends. Sub , tera: Coleophoridae). Zoosyst. ross., 5(2), 1996: genital plate suboblong, along posterior margin 303-308. Baldizzone, G. 1994. Coleopboridae dell' Area Irano short bristles; ostium bursae elongate ovate, Anatolica e regioni limitrofe (Lepidoptera). Assoc. ending in medial part of subgenital plate; later Natur. Piemont. Mem., 3: 1-423. ally on anterior margin two small triangular Baldizzone, G. & Patzak, H. 1991. Sur quelques projections. Tubular antrum widened to end, Coleophoridae de la region de Berdjansk (Ukraine). with a small gradual swelling on right side an Beilr. Entomol., 41(2): 351-369'. · teriorly. Spiny portion of ductus bursae 2.1 Falkovitsh, M.I. 1991. New species of casebearers times as long as subgenital plate, with dark in (Lepidoptera, Coleophoridae) of the fauna of the USSR. Entomo/. Obozr., 70 (3): 586-599. (In Rus ternal spinelets. Lateral and central rods well sian). defined. Next portion of ductus with central Falkovitsh, M.1. & Jalava, J. 1997. Coleophoridae rod in 1/3 length, ending with a spiral; bursa collected in the North Caucasus in 1990, with de copulatrix oval, with a small signum and nar scription of a new species (Lepidoptera). Zoosyst. row sclerotized plate. ross., 5(2), 1996: 309-311 .. Abdominal tergites (Fig. 5). 1st abdominal Filipjev,.N.N. 1926. Mikrolepidopterenfaunades Gou tergite with 20-23 spinelets; on the following vern·ements von Stavropol (Russland, Ciscaucasien). tergites, patches of spinelets compact, elon- · Arch. Naturgesch., 91 (I), 1925: 94-118. Fuchs, A; 1903. Neue Kleinfalter des Mittelmeerge gate, twice as long as wide. biets. Stell. entomol. Ztg., 64: 1-16. Comparison. The femaleof this species is re Patzak, H. 1977. Coleophoriden vom Kaukasus und latedto that of E. nomgona Flkv. The main dif aus Transkaukasien (Lepidoptera, Coleophoridae). ferences in the genitalia are as follows: sub Deutsche entomol. Z., 24(4/5): 277-281. genital plate longer; ostium bursae narrow and Reznik, S. Ya. 1977. A brief review of the genus Mul oval; spiny portion of ductus longer; the medial tico/oria Cap: (Lepidoptera, Coleophoridae ). In: part of rod not sclerotized and more spiralized. O.L. Kryzhanovskij & A.K. Galkin (eds). Sistema tika i faunistika · nasekomykh (Systematics and Acknowledgements faunistics of insects): 78-88. Leningrad: Zool. Inst. Akad. Nauk SSSR. (in Russian). Reznik,S.Ya. 1989. New and little known species of We are thankful to Dr. M.I. Falkovitsh (St.Peters casebearers of the genus Multicoloria Cap. (Lepi burg, Russia) for his valuable advice concerning the doptera, Coleophoridae) from arid zones of the taxonomy and nomenclature of the Coleophoridae. USSR. Trudy zoo!. Inst. Akad. Nauk SSSR, 200: 33- 39. (In Russian). References Stainton, H.T. 1867. The Tineina of Syria and Asia Minor. V1+84 p. London. Anikin, V.V. 1998. A new species of the genus Amsel ghia from the North Caucasus (Lepidoptera: Coleo phoridae). Zoosyst. ross., 7(2): 326. Received 7 April 2000