BARRY BOLTON’S CATALOGUE, 2020

CATAULACUS adlerzi Mayr, 1887; see under . adpressus. adpressus Bolton, 1974a: 30, figs. 15, 18 (w.) GHANA. Type-material: holotype worker. Type-locality: Ghana: Bunso, 8.vii.1969, pyrethrum knockdown B.6/4 (D. Leston). Type-depository: BMNH. Status as : Bolton, 1982: 355 (in key); Bolton, 1995b: 137. Distribution: Ghana. aethiops. Cataulacus jeanneli var. aethiops Santschi, 1924b: 220 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: syntype workers (number not stated). Type-localities: Democratic Republic of Congo (“Congo belge”): Kidada-Kitobola, 14/25.ii.1922 (H. Schouteden); Barumbu (Bequaert). Type-depository: MRAC. Junior synonym of pygmaeus: Bolton, 1974a: 48. Junior synonym of weissi: Bolton, 1982: 358; Bolton, 1995b: 137. alenensis. Cataulacus sulcatus var. alenensis Stitz, 1910: 137 (w.) EQUATORIAL GUINEA. Type-material: 2 syntype workers. Type-locality: Equatorial Guinea (“Spanisch Guinea”): Alen (Tessmann) . Type-depository: MNHU. [Misspelled as alensis by Santschi, 1937b: 102.] Subspecies of guineensis: Forel, 1913e: 668; Wheeler, W.M. 1922a: 913; Santschi, 1924b: 217; Emery, 1924d: 296; Santschi, 1937b: 102; Bernard, 1953b: 252. Junior synonym of guineensis: Bolton, 1974a: 55; Bolton, 1995b: 137. andamanensis. r. andamanensis Forel, 1903d: 406 (w.) (Andaman Is). Type-material: syntype workers (number not stated). Type-locality: India: Andaman Is (no collector’s name). Type-depository: MHNG. Subspecies of granulatus: Emery, 1924d: 298; Chapman & Capco, 1951: 85. Junior synonym of simoni: Bolton, 1974a: 72; Bolton, 1995b: 137. bakusuensis. Cataulacus pygmaeus var. bakusuensis Forel, 1913h: 350 (w.m.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: 1 syntype queen, 1 syntype male. Type-locality: Democratic Republic of Congo: Bakusu, dans un rameau d’Hevea (no collector’s name). Type-depository: MRAC. Subspecies of pygmaeus: Wheeler, W.M. 1922a: 915; Emery, 1924d: 297. Junior synonym of pygmaeus: Bolton, 1974a: 48; Bolton, 1995b: 137. batonga. Cataulacus baumi r. batonga Forel, 1913a: 114 (w.) ZIMBABWE. Type-material: syntype workers (number not stated). Type-locality: Zimbabwe (“Rhodesia”): Khami River, 22.x.1911 (G. Arnold). Type-depositories: MCZC, MHNG. Subspecies of baumi: Arnold, 1917: 389; Wheeler, W.M. 1922a: 912; Emery, 1924d: 296. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 137. baumi. Cataulacus baumi Forel, 1901d: 304 (w.q.m.) ANGOLA. Type-material: syntype workers (number not stated), 1 syntype queen, 1 syntype male. Type-locality: Angola: Mossamedes, Cubango-Cuito (no collector’s name). Type-depository: MHNG. [Also described as new by Forel, 1903e: 560.] Status as species: Forel, 1914d: 218; Arnold, 1917: 388; Wheeler, W.M. 1922a: 912; Emery, 1924d: 296; Santschi, 1930b: 72; Menozzi, 1930b: 111. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 137. bequaerti. Cataulacus bequaerti Forel, 1913b: 316 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: syntype workers (number not stated). Type-locality: Democratic Republic of Congo (“Congo belge”): Katanga, Kabanza (Kikondja), Riv. Lovoi (J. Bequaert)). Type-depositories: MHNG, MNHU, MRAC. Status as species: Wheeler, W.M. 1922a: 910; Emery, 1924d: 296; Bolton, 1974a: 40 (redescription); Bolton, 1982: 358 (in key); Bolton, 1995b: 137. Distribution: Democratic Republic of Congo. boltoni. Cataulacus boltoni Snelling, R.R. 1979a: 3, figs. 1-8 (w.) NIGERIA. Type-material: holotype worker. Type-locality: Nigeria: Gambari (= Cocoa Research Institute of Nigeria), 24.v.1975 (B. Taylor). Type-depository: BMNH. Status as species: Bolton, 1982: 356 (in key); Bolton, 1995b: 137. Distribution: Nigeria. brazzavillensis. Cataulacus kohli st. brazzavillensis Santschi, 1910c: 389, fig. 15 (w.m.) CONGO. Type-material: 3 syntype workers, 1 syntype male. Type-locality: Congo (“Congo français”): Brazzaville (A. Weiss). Type-depository: NHMB. Subspecies of kohli: Wheeler, W.M. 1922a: 914; Emery, 1924d: 297. Junior synonym of kohli: Bolton, 1974a: 22; Bolton, 1995b: 137. brevisetosus. Cataulacus brevisetosus Forel, 1901d: 305 (w.) ANGOLA. Type-material: holotype worker. Type-locality: Angola: Mossamedes, Cubango-Cuito (no collector’s name). Type-depository: MHNG. [Also described as new by Forel, 1903e: 560.] Subspecies of pygmaeus: Santschi, 1916b: 508; Wheeler, W.M. 1922a: 298, 916; Emery, 1924d: 297. Status as species: Arnold, 1917: 397; Santschi, 1930b: 56; Santschi, 1939b: 246; Bolton, 1974a: 31 (redescription); Bolton, 1982: 356 (in key); Bolton, 1995b: 137; Hita Garcia, et al. 2013: 209. Senior synonym of meduseus: Bolton, 1974a: 31; Bolton, 1995b: 137. Material of the unavailable name plebeja referred here by Bolton, 1974a: 31; Bolton, 1995b: 137. Distribution: Angola, South Africa, Tanzania, Zimbabwe. brookei. Cataulacus brookei Forel, 1901e: 378 (w.q.m.) BORNEO (East : Sarawak). Type-material: syntype workers (number not stated), 1 syntype queen, 1 syntype male. Type-locality: Malaysia (“Bornéo”): Sarawak (Haviland). Type-depository: MHNG. Status as species: Wheeler, W.M. 1919e: 92; Emery, 1924d: 298; Chapman & Capco, 1951: 84. Junior synonym of hispidulus: Bolton, 1974a: 66; Bolton, 1995b: 137. bulawayensis. Cataulacus baumi var. bulawayensis Arnold, 1917: 391. Type-material: syntype workers (number not stated), 1 syntype queen. Type-locality: Zimbabwe (“Rhodesia”): Bulawayo, 31.v.1914 (G. Arnold). Type-depository: MHNG. [First available use of Cataulacus baumi r. batonga var. bulawayensis Forel, 1914d: 218 (w.q.) ZIMBABWE; unavailable (infrasubspecific) name (Bolton, 1974a: 43).] As unavailable (infrasubspecific) name: Arnold, 1920a: 404 (in key); Wheeler, W.M. 1922a: 912. Subspecies of baumi: Emery, 1924d: 296. Junior synonym of intrudens: Bolton, 1995b: 137. carbonarius Mayr, 1870; see under PROCRYPTOCERUS. catuvolcus. Cataulacus catuvolcus Bolton, 1974a: 74, figs. 4, 37 (w.q.m.) PHILIPPINES (Romblon I.). Type-material: holotype worker, 8 paratype workers, 3 paratype queens, 2 paratype males. Type-locality: holotype Philippines: Romblon I., 2.v.1924 (J.W. Chapman); paratypes with same data. Type-depositories: MCZC (holotype); BMNH, MCZC (paratypes). Status as species: Bolton, 1995b: 137. Distribution: Philippines (Romblon). centrurus. Cataulacus centrurus Bolton, 1982: 359, fig. 33 (w.) CAMEROON. Type-material: holotype worker. Type-locality: Cameroon: Nkoemvon, 1979 (D. Jackson). Type-depository: BMNH. Status as species: Bolton, 1995b: 137. Distribution: Cameroon. cestus. Cataulacus cestus Bolton, 1982: 360 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: holotype worker, 4 paratype workers. Type-locality: holotype Democratic Republic of Congo (“B. Congo”): Ituri Forest, Beni-Irumu, ii.1948, no. 2122 (N.A. Weber); paratypes: 1 worker with same data, 1 worker with same data but no, 2120, 2 workers with same data but no. 2119. Type-depositories: MCZC (holotype); BMNH, MCZC (paratypes). Status as species: Bolton, 1995b: 137. Distribution: Democratic Republic of Congo. chapmani. Cataulacus chapmani Bolton, 1974a: 75, fig. 30 (w.m.) PHILIPPINES (Negros I., Luzon I.). Type-material: holotype worker, 17 paratype workers, 3 paratype males. Type-locality: holotype Philippines: Negros I., Dumaguete, 30.iv.1924 (J.W. Chapman); paratypes: 14 workers with same data, 2 males with same data but without date and 500 ft, 1 worker with same data but 28.iv.1932, 1 worker, 1 male with same data but undated and numbered 405, 1 worker Philippines: Luzon I., Lamao. Type-depositories: MCZC (holotype); BMNH, MCZC, USNM (paratypes). Status as species: Bolton, 1995b: 138. Distribution: Philippines (Luzon, Negros). chariensis. Cataulacus pygmaeus var. chariensis Santschi, 1911c: 358 (w.) CHAD. Type-material: holotype worker. Type-locality: Chad: Moyen Chari, Fort Archambault, 1904 (J. Decorse). Type-depository: MNHN. Subspecies of pygmaeus: Wheeler, W.M. 1922a: 915; Emery, 1924d: 297. Junior synonym of pygmaeus: Bolton, 1974a: 48; Bolton, 1995b: 138. convergens Mayr, 1887; see under PROCRYPTOCERUS. coriaceus. Cataulacus coriaceus Stitz, 1910: 138, fig. 7 (w.) CAMEROON. Type-material: holotype worker. Type-locality: Cameroon: Mundame (Conradt). Type-depository: MNHU. Status as species: Wheeler, W.M. 1922a: 912; Emery, 1924d: 296. Junior synonym of pullus: Bolton, 1974a: 26; Bolton, 1995b: 138. coriarius Mayr, 1870; see under PROCRYPTOCERUS. crassispina. Cataulacus erinaceus var. crassispina Santschi, 1917b: 287 (w.) CONGO. Type-material: holotype worker. Type-locality: Congo (“Congo français”): Goda, P. Charleuf (H. de Buysson). Type-depository: NHMB. Subspecies of erinaceus: Wheeler, W.M. 1922a: 913; Emery, 1924d: 296. Junior synonym of erinaceus: Bolton, 1974a: 52; Bolton, 1995b: 138. deformis Roger, 1863; see under CYPHOMYRMEX. degener. Cataulacus pygmaeus st. degener Santschi, 1916b: 507 (w.) (no state data). Type-material: holotype worker. Type-locality: Angola: Loanda, 14.i.1913 (no collector’s name). [Note: no data given in description; type-locality data present on holotype label (Bolton, 1974a: 31).]. Type-depository: NHMB. Subspecies of pygmaeus: Wheeler, W.M. 1922a: 916; Santschi, 1924b: 221; Emery, 1924d: 297. Junior synonym of brevisetosus: Bolton, 1974a: 31. Junior synonym of jeanneli: Bolton, 1982: 358; Bolton, 1995b: 138. densipunctatus. Cataulacus johannae r. densipunctatus Stitz, 1923: 163 (w.) NAMIBIA. Type-material: 2 syntype workers. Type-locality: Namibia (“Deutsch Südwestafrika”): Tsumeb, 13-19.vi.1911 (W. Michaelsen). Type-depository: unknown, probably in ZMUH. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 138. difficilis. Cataulacus pygmaeus st. difficilis Santschi, 1916b: 506 (w.) BENIN. Type-material: holotype worker. Type-locality: Benin (“Dahomey”): (no further data) (Desanti). Type-depository: NHMB. Subspecies of pygmaeus: Wheeler, W.M. 1922a: 916; Emery, 1924d: 297. Status as species: Bolton, 1974a: 33 (redescription); Bolton, 1982: 357 (in key); Bolton, 1995b: 138. Distribution: Benin. donisthorpei. Cataulacus donisthorpei Santschi, 1937a: 61 (w.) KENYA. Type-material: 2 syntype workers. Type-locality: Kenya (“Kenya colony”): (no further data), nos 17 and 42 (no collector’s name, in Donisthorpe collection). Type-depository: BMNH. Junior synonym of striativentris: Bolton, 1974a: 36; Bolton, 1995b: 138. durbanensis. Cataulacus micans r. durbanensis Forel, 1914d: 219 (w.) SOUTH AFRICA. Type-material: holotype worker. Type-locality: South Africa: Natal, Durban, 15.i.1914 (G. Arnold). Type-depository: MHNG. Arnold, 1917: 395 (q.m.). Subspecies of micans: Arnold, 1917: 394; Wheeler, W.M. 1922a: 915; Emery, 1924d: 297. Junior synonym of wissmannii: Bolton, 1974a: 51; Bolton, 1995b: 138. ebrardi. Cataulacus ebrardi Forel, 1886c: cv (w.) MADAGASCAR. Type-material: syntype workers (number not stated). Type-locality: Madagascar: Morondava (Grandidier). Type-depository: MHNG. Forel, 1910a: 20 (q.). Status as species: Forel, 1891b: 145 (redescription); Dalla Torre, 1893: 138; Mayr, 1895: 129; Emery, 1895f: 343; Forel, 1910a: 20; Wheeler, W.M. 1922a: 1033; Emery, 1924d: 298; Bolton, 1974a: 41 (redescription); Bolton, 1982: 355 (in key); Bolton, 1995b: 138. Distribution: Madagascar. egenus. Cataulacus egenus Santschi, 1911c: 359, fig (w.) CONGO. Type-material: holotype worker. Type-locality: Congo (“Congo français”): Madingou (R.P. Zimmermann). Type-depository: NHMB. Wheeler, G.C. & Wheeler, J. 1954b: 151 (l.); Bolton, 1974a: 18 (q.m.). Status as species: Wheeler, W.M. 1922a: 199, 912; Santschi, 1924b: 217; Emery, 1924d: 296; Bolton, 1974a: 18 (redescription); Bolton, 1995b: 138; Hita Garcia, et al. 2013: 209. Senior synonym of simplex: Bolton, 1974a: 18; Bolton, 1995b: 138. Distribution: Cameroon, Congo, Democratic Republic of Congo, Ghana, Kenya, Nigeria, Uganda. elongatus. Cataulacus elongatus Santschi, 1924b: 221 (w.) ANGOLA. Type-material: holotype worker. Type-locality: Angola: Loanda (Le Moult). Type-depository: NHMB. Bolton, 1974a: 34 (q.). Status as species: Santschi, 1930b: 56; Bolton, 1974a: 34 (redescription); Bolton, 1982: 356 (in key); Bolton, 1995b: 138. Distribution: Angola, Cameroon, Ghana. erinaceus. Cataulacus erinaceus Stitz, 1910: 134, fig. 3 (w.) EQUATORIAL GUINEA, CAMEROON. Type-material: syntype workers (total number unknown). Type-localities: 3 syntypes Equatorial Guinea (“Spanisch Guinea”): Alen (Tessmann); unknown number of syntypes Cameroon: Mundame (Conradt). Type-depositories: MNHU, NHMB. Forel, 1913h: 350 (q.). Status as species: Forel, 1913b: 315; Forel, 1913e: 668; Forel, 1913h: 350; Forel, 1915c: 345; Forel, 1916: 427; Wheeler, W.M. 1922a: 198, 913; Emery, 1924d: 296; Bernard, 1953b: 252; Bolton, 1974a: 52 (redescription); Bolton, 1982: 356 (in key); Bolton, 1995b: 138. Senior synonym of crassispina: Bolton, 1974a: 52; Bolton, 1995b: 138. Material of the nomen nudum princeps referred here by Wheeler, W.M. 1922a: 913; Emery, 1924d: 296; Bolton, 1974a: 52; Bolton, 1995b: 138. Distribution: Cameroon, Congo, Democratic Republic of Congo, Equatorial Guinea, Gabon, Ghana, Liberia. fernandensis. Cataulacus sulcatus var. fernandensis Stitz, 1910: 137 (w.) EQUATORIAL GUINEA (Bioko I.). Type-material: holotype worker. Type-locality: Equatorial Guinea: Bioko I. (“Fernando Po”) (Zenker). Type-depository: MNHU. Subspecies of guineensis: Wheeler, W.M. 1922a: 913; Santschi, 1924b: 217; Emery, 1924d: 296; Menozzi, 1942: 177; Eidmann, 1944: 456. Junior synonym of guineensis: Bolton, 1974a: 55; Bolton, 1995b: 138. flagitiosus. Cataulacus flagitiosus Smith, F. 1862a: 49 (w.) (Sulawesi). Type-material: holotype worker. Type-locality: Indonesia: Sulawesi (“Celebes”), Tondano (A.R. Wallace). [Note: holotype is assured but is mislabelled SAR5 (where SAR = Sarawak). This species has only been collected in Sulawesi.] Type-depository: OXUM. Status as species: Smith, F. 1862d: 415; Mayr, 1863: 402; Roger, 1863b: 39; Smith, F. 1871a: 334; Emery, 1887b: 470; Dalla Torre, 1893: 138; Emery, 1901g: 567; Emery, 1924d: 298; Donisthorpe, 1932c: 469; Chapman & Capco, 1951: 84; Bolton, 1974a: 76 (redescription); Bolton, 1995b: 138. Distribution: Indonesia (Sulawesi). foveolatus. Cataulacus foveolatus Stitz, 1910: 140 (w.) EQUATORIAL GUINEA. Type-material: holotype worker. Type-locality: Equatorial Guinea (“Spanisch Guinea”): Uelleburg (Tessmann). Type-depository: MNHU. Status as species: Wheeler, W.M. 1922a: 913; Emery, 1924d: 296. Junior synonym of kohli: Bolton, 1974a: 22; Bolton, 1995b: 138. foveosquamosus. Cataulacus foveosquamosus Santschi, 1937a: 58, figs. 8, 9 (q.) SOUTH AFRICA. Type-material: holotype queen. Type-locality: South Africa: Kwazulu-Natal (“Zululand”) (I. Trägårdh). Type-depository: NHMB. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 138. fricatidorsus. Cataulacus otii st. fricatidorsus Santschi, 1914e: 26 (w.) SOUTH AFRICA. Type-material: 4 syntype workers. Type-locality: South Africa: Natal, Zululand, Dukudu, 27.vii.1905 (I. Trägårdh). Type-depository: NHMB. Subspecies of otii: Arnold, 1917: 401; Wheeler, W.M. 1922a: 915; Emery, 1924d: 297. Status as species: Bolton, 1974a: 42 (redescription); Bolton, 1982: 358 (in key); Bolton, 1995b: 138. Distribution: South Africa. gazanus. Cataulacus baumi var. gazanus Santschi, 1928f: 208 (w.) MOZAMBIQUE. Type-material: syntype workers (number not stated). Type-locality: Mozambique (“Est Africain Portugais”): Beira, 6.vi.1920 (G. Arnold). Type-depository: NHMB. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 138. gilviventris. Cataulacus lujae var. gilviventris Forel, 1913b: 316 (q.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: holotype queen. Type-locality: Democratic Republic of Congo (“Congo belge”): Katanga, Kabanza (Kikondja), Riv. Lovoi (J. Bequaert). Type-depository: MRAC. As unavailable (infrasubspecific) name: Wheeler, W.M. 1922a: 916; Emery, 1924d: 297. Junior synonym of brevisetosus: Bolton, 1974a: 31. Junior synonym of lujae: Bolton, 1982: 358; Bolton, 1995b: 138. granulatus. Formica granulata Latreille, 1802c: 275, pl. 12, fig. 75a-d (w.) (no state data, "Grand-Indes"). Type-material: holotype worker (in fragments). Type-locality: “Grand-Indes”. Type-depository: perhaps in MNHN (Latreille collection). Emery, 1893e: 215 (q.m.); Bingham, 1903: 123 (q.); Bolton, 1974a: 65 (m.). Combination in Cryptocerus: Lepeletier de Saint-Fargeau, 1835: 171; combination in Cataulacus: Smith, F. 1853: 226. Status as species: Lepeletier de Saint-Fargeau, 1835: 171; Smith, F. 1853: 226; Smith, F. 1858b: 196; Smith, F. 1862d: 414; Mayr, 1863: 402; Roger, 1863b: 39; Smith, F. 1871a: 335; Emery, 1889b: 507; Dalla Torre, 1893: 138; Emery, 1893e: 215; Emery, 1895k: 473; Emery, 1900d: 691; Forel, 1903a: 707; Bingham, 1903: 122; Forel, 1906b: 90; Forel, 1913e: 662; Wheeler, W.M. 1919e: 92; Wheeler, W.M. 1923b: 4; Emery, 1924d: 298; Wheeler, W.M. 1930h: 71; Chapman & Capco, 1951: 84; Bolton, 1974a: 64 (redescription); Bolton, 1995b: 138; Wu, J. & Wang, 1995: 61; Zhou, 2001b: 72; Jaitrong & Nabhitabhata, 2005: 17; Mohanraj, et al. 2010: 6; Pfeiffer, et al. 2011: 44; Guénard & Dunn, 2012: 41; Bharti, Guénard, et al. 2016: 35; Jaitrong, Guénard, et al. 2016: 34. Senior synonym of hispidus: Bolton, 1974a: 64; Bolton, 1995b: 138; Zhou, 2001b: 72. Distribution: (Guangxi, Hainan I.), India (India, Andaman Is), Indonesia (Java, Sumatra), Laos, Malaysia (Peninsula, Sarawak), , , , . . greggi. Cataulacus greggi Bolton, 1974a: 54, fig. 22 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: holotype worker, 2 paratype workers. Type-locality: holotype Democratic Republic of Congo (“Zaire”): Ituri Forest, Epulu, vii.1955, no. 10 (T. Gregg); paratypes: Democratic Republic of Congo (“Zaire”): Yangambi, x.1956 (N.L.H. Krauss). Type-depositories: MCZC (holotype); BMNH (paratypes). Status as species: Bolton, 1982: 357 (in key); Bolton, 1995b: 138. Distribution: Cameroon, Democratic Republic of Congo. guilelmi. Cataulacus huberi subsp. guilelmi Wheeler, W.M. 1925b: 1 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: holotype worker. Type-locality: Democratic Republic of Congo (“Central Africa”): Ituri (no further data). Type-depository: unknown, perhaps in NHRS. Junior synonym of huberi: Bolton, 1974a: 19; Bolton, 1995b: 138. guineensis. Cataulacus guineensis Smith, F. 1853: 225, pl. 20, fig. 5 (w.) (no state data, "Tropical West Africa"). Type-material: holotype worker. Type-locality: Tropical West Africa. Type-depository: OXUM. Santschi, 1910c: 390 (q.); Wheeler, W.M. 1922a: 199 (m.). Status as species: Smith, F. 1858b: 196; Smith, F. 1862d: 414; Mayr, 1863: 403; Roger, 1863b: 39; Dalla Torre, 1893: 138; Mayr, 1895: 129; André, 1895a: 4; Santschi, 1910c: 390; Forel, 1910e: 421; Forel, 1911e: 263; Santschi, 1914d: 373; Forel, 1916: 427; Wheeler, W.M. 1922a: 199, 913; Santschi, 1924b: 217; Emery, 1924d: 296; Santschi, 1939c: 10; Bolton, 1974a: 55 (redescription); Bolton, 1982: 356 (in key); Bolton, 1995b: 138; Hita Garcia, et al. 2013: 209. Senior synonym of alenensis: Bolton, 1974a: 55; Bolton, 1995b: 138. Senior synonym of fernandensis: Bolton, 1974a: 55; Bolton, 1995b: 138. Senior synonym of parallelus: Bolton, 1974a: 55; Bolton, 1995b: 138. Senior synonym of sulcatus: Forel, 1910e: 421; Bolton, 1974a: 55; Bolton, 1995b: 138. Senior synonym of sulcinodis: Bolton, 1974a: 55; Bolton, 1995b: 138. Distribution: Cameroon, Democratic Republic of Congo, Equatorial Guinea, Ghana, Ivory Coast, Kenya, Liberia, Nigeria, Togo, Uganda. hararicus. Cataulacus hararicus Forel, 1894b: 79 (w.) ETHIOPIA. Type-material: syntype workers (number not stated). Type-locality: Ethiopia (“Süd abessinien”): Harar (Ilg). Type-depository: MHNG. Status as species: Emery, 1895a: 180; Wheeler, W.M. 1922a: 914; Viehmeyer, 1923: 92; Emery, 1924d: 296. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 138. herteri. Cataulacus huberi r. herteri Forel, 1913b: 315 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: syntype workers (number not stated). Type-locality: Democratic Republic of Congo (“Congo belge”): Katanga, Welgelegen, 14.vi.1912 (J. Bequaert). Type-depositories: MHNG, MNHU, MRAC. Subspecies of huberi: Wheeler, W.M. 1922a: 914; Santschi, 1924b: 217; Emery, 1924d: 296; Santschi, 1937a: 57; Santschi, 1937d: 234; Menozzi, 1942: 177; Eidmann, 1944: 469. Junior synonym of huberi: Bolton, 1974a: 19; Bolton, 1995b: 138. hispidulus. Cataulacus hispidulus Smith, F. 1865: 76, pl. 4, fig. 7 (w.) INDONESIA (Sumatra). Type-material: holotype worker. Type-locality: Indonesia: Sumatra (“SUM”) (A.R. Wallace). Type-depository: OXUM. Bolton, 1974a: 66 (q.m.). Subspecies of granulatus: Dalla Torre, 1893: 138; Emery, 1924d: 298; Chapman & Capco, 1951: 84. Status as species: Smith, F. 1871a: 334; Mayr, 1872: 155; Emery, 1887b: 470; André, 1892b: 53; Wheeler, W.M. 1919e: 93; Donisthorpe, 1932c: 474; Bolton, 1974a: 66 (redescription); Bolton, 1995b: 138; Pfeiffer, et al. 2011: 44. Senior synonym of brookei: Bolton, 1974a: 66; Bolton, 1995b: 138. Distribution: Indonesia (Sumatra), Malaysia (Sarawak). hispidus. Cataulacus hispidus Smith, F. 1876d: 611, pl. 11, fig. 11 (w.) SINGAPORE. Type-material: holotype worker. Type-locality: Singapore (“Sing.”). Type-depository: OXUM. Status as species: Forel, 1903d: 406; Forel, 1909d: 224; Forel, 1911b: 205; Forel, 1911d: 388; Forel, 1912n: 60. Subspecies of granulatus: Dalla Torre, 1893: 138; Forel, 1911a: 24; Forel, 1911b: 205; Viehmeyer, 1916a: 140; Wheeler, W.M. 1919e: 93; Emery, 1924d: 298; Chapman & Capco, 1951: 85. Junior synonym of granulatus: Bolton, 1974a: 64; Bolton, 1995b: 138; Zhou, 2001b: 72. horridus. Cataulacus horridus Smith, F. 1857a: 81, pl. 2, fig. 3 (w.) BORNEO (East Malaysia: Sarawak), WEST MALAYSIA. Type-material: 4 syntype workers. Type-localities: 3 syntypes Malaysia (“Borneo”): Sarawak (“SAR”) (A.R. Wallace); 1 syntype Malaysia: Peninsula, Malacca (“Mal.23”) (A.R. Wallace). Type-depositories: BMNH, OXUM. Crawley, 1924: 401 (q.); Wheeler, G.C. & Wheeler, J. 1954b: 151 (l.). Combination in C. (Otomyrmex): Santschi, 1928h: 125. Status as species: Smith, F. 1858b: 196; Smith, F. 1862d: 415; Mayr, 1863: 403; Roger, 1863b: 39; Smith, F. 1871a: 335; Dalla Torre, 1893: 138; Emery, 1893e: 216; Forel, 1907a: 12; Forel, 1913k: 83; Viehmeyer, 1916a: 140; Wheeler, W.M. 1919e: 93; Crawley, 1924: 401; Emery, 1924d: 298; Santschi, 1928h: 125; Donisthorpe, 1932c: 451; Chapman & Capco, 1951: 85; Bolton, 1995b: 138; Jaitrong & Nabhitabhata, 2005: 17; Pfeiffer, et al. 2011: 44. Junior synonym of insularis: Bolton, 1974a: 84 (error). Senior synonym of insularis: Wheeler, W.M. 1919e: 93; Bolton, 1995b: 138. [Note: Bolton, 1974a: 84, gave insularis as senior synonym, but horridus has priority on first reviser principle, based on Wheeler's, 1919e: 93, attribution (Bolton, 1995b: 138).] Distribution: Indonesia (Java, Sumatra), Malaysia (Peninsula, Sabah, Sarawak), Thailand. huberi. Cataulacus huberi André, 1890: 326 (w.) SIERRA LEONE. Type-material: syntype workers (number not stated). Type-locality: Sierra Leone: (no further data) (A. Mocquerys). Type-depository: MNHN. André, 1895a: 4 (q.); Bolton, 1974a: 20 (m.). Status as species: Dalla Torre, 1893: 138; André, 1895a: 4; Mayr, 1895: 128; Santschi, 1910c: 390; Wheeler, W.M. 1922a: 914; Santschi, 1924b: 217; Emery, 1924d: 296; Wheeler, W.M. 1925b: 1; Bolton, 1974a: 19 (redescription); Bolton, 1982: 355 (in key); Bolton, 1995b: 138. Senior synonym of guilelmi: Bolton, 1974a: 19; Bolton, 1995b: 138. Senior synonym of herteri: Bolton, 1974a: 19; Bolton, 1995b: 138. Senior synonym of longispinus: Bolton, 1974a: 19; Bolton, 1995b: 138. Material of the unavailable name luebensis referred here by Bolton, 1974a: 19; Bolton, 1995b: 138. Distribution: Cameroon, Democratic Republic of Congo, Gabon, Ghana, Nigeria, Sierra Leone, Uganda, Zambia. impressus. Cataulacus impressus Bolton, 1974a: 35 (w.) UGANDA. Type-material: holotype worker. Type-locality: Uganda: (no further data). Type-depository: MCZC. Status as species: Bolton, 1982: 357 (in key); Bolton, 1995b: 138. Distribution: Uganda. inermis. Cataulacus inermis Santschi, 1924b: 218, fig. 10 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: holotype worker. Type-locality: Democratic Republic of Congo (“Congo belge”): Kasai, Ngombe, 5.xi.1921 (H. Schouteden). Type-depository: MRAC. Status as species: Bolton, 1974a: 21 (redescription); Bolton, 1982: 355 (in key); Bolton, 1995b: 138. Distribution: Democratic Republic of Congo. insularis. Cataulacus insularis Smith, F. 1857a: 80, pl. 2, fig. 4 (m.) BORNEO (East Malaysia: Sarawak). Type-material: holotype male. Type-locality: Malaysia (“Borneo”): Sarawak (“SAR”) (A.R. Wallace). Type-depository: OXUM. Status as species: Smith, F. 1858b: 197; Smith, F. 1862d: 415; Mayr, 1863: 403; Roger, 1863b: 39; Smith, F. 1871a: 335; Dalla Torre, 1893: 138; Emery, 1924d: 298; Donisthorpe, 1932c: 451; Chapman & Capco, 1951: 85. Junior synonym of horridus: Wheeler, W.M. 1919e: 93; Bolton, 1974a: 84 (redescription); Bolton, 1995b: 138. [Note: see comment under horridus.] intermedius. Cataulacus intrudens st. intermedius Santschi, 1917b: 287 (w.) ZIMBABWE. Type-material: syntype workers (number not stated). Type-locality: Zimbabwe (“Rhodesia”): Bambesi, 25.vi.1914 (G. Arnold). Type-depositories: BMNH, NHMB. Subspecies of intrudens: Arnold, 1920a: 403; Wheeler, W.M. 1922a: 914; Emery, 1924d: 296. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 138. intrudens. intrudens Smith, F. 1876d: 609, pl. 11, fig. 7 (w.q.m.) SOUTH AFRICA. Type-material: lectotype worker, 2 paralectotype workers, 2 paralectotype males (designated by Bolton, 1974a: 42). [Note: the queen described by Smith is absent from the BMNH collection, presumed lost or destroyed.] Type-locality: lectotype South Africa: Natal, Durban, Weenen District, in acacia thorns (“76,48.Natal”) (J.M. Hutchinson); paralectotypes with same data. Type-depositories: BMNH (lectotype); BMNH, MCZC (paralectotypes). Combination in Cataulacus: Mayr, 1886c: 364. Status as species: Mayr, 1886c: 364; Dalla Torre, 1893: 138; Mayr, 1895: 129; Emery, 1895h: 41; Mayr, 1907b: 14; Arnold, 1917: 391; Wheeler, W.M. 1922a: 914; Emery, 1924d: 296; Arnold, 1926: 286; Menozzi, 1930b: 112; Menozzi & Consani, 1952: 68; Bolton, 1974a: 43 (redescription); Bolton, 1982: 357, 358 (in key); Bolton, 1995b: 138; Hita Garcia, et al. 2013: 209. Senior synonym of batonga: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of baumi: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of bulawayensis: Bolton, 1995b: 138. Senior synonym of densipunctatus: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of foveosquamosus: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of gazanus: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of hararicus: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of intermedius: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of johannae: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of pseudotrema: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of rugosus: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of subrugosus: Bolton, 1974a: 43; Bolton, 1995b: 138. Senior synonym of umbilicatus: Bolton, 1974a: 43; Bolton, 1995b: 138. [Note: intrudens and its current synonyms are best regarded as an unresolved species-complex.] Material of the unavailable names krugeri, tangana referred here by Bolton, 1974a: 43. Distribution: Angola, Ethiopia, Kenya, Madagascar, Malawi, Mozambique, Namibia, Somalia, Tanzania, South Africa, Zambia, Zimbabwe. jacksoni. Cataulacus jacksoni Bolton, 1982: 360 (w.) CAMEROON. Type-material: holotype worker. Type-locality: Cameroon: Nkoemvon, 1980 (D. Jackson). Type-depository: BMNH. Status as species: Bolton, 1995b: 138. Distribution: Cameroon. jeanneli. Cataulacus jeanneli Santschi, 1914b: 108, fig. 16 (w.) KENYA. Type-material: holotype worker. Type-locality: Kenya (“Afrique orientale anglaise”): Gazi, 20 km. S of Mombasa (st.no.6), xi.1911 (Alluaud & Jeannel). Type-depository: NHMB. [Misspelled as janneli by Santschi, 1926b: 244.] Subspecies of pygmaeus: Santschi, 1916b: 506; Wheeler, W.M. 1922a: 916; Emery, 1924d: 297. Junior synonym of brevisetosus: Bolton, 1974a: 31. Status as species: Santschi, 1924b: 220; Santschi, 1926b: 244; Bolton, 1982: 356 (in key), 358; Bolton, 1995b: 138; Hita Garcia, et al. 2013: 209. Senior synonym of degener: Bolton, 1982: 358; Bolton, 1995b: 138. Senior synonym of loveridgei: Bolton, 1982: 358; Bolton, 1995b: 138. Distribution: Angola, Cameroon, Ghana, Ivory Coast, Kenya, Tanzania, Uganda. johannae. Cataulacus johannae Forel, 1895c: 250 (w.q.) MADAGASCAR. Type-material: syntype workers, syntype queens (numbers not stated). Type-locality: Madagascar: Imerina oriental (M. Sikora). Type-depository: MHNG. Status as species: Wheeler, W.M. 1922a: 1033; Emery, 1924d: 298. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 138. kenyensis. Cataulacus jeanneli st. kenyensis Santschi, 1935b: 272, fig. 6a-c (w.) KENYA. Type-material: 2 syntype workers. Type-locality: Kenya: Nairobi, st.2, 1660 m., 1932-33 (Arambourg, Chappuis & Jeannel). Type-depository: NHMB. Junior synonym of weissi: Bolton, 1974a: 39. Status as species: Bolton, 1982: 357 (in key), 358; Bolton, 1995b: 138; Hita Garcia, et al. 2013: 209. Distribution: Kenya, Uganda. kohli. Cataulacus kohli Mayr, 1895: 127, fig. 2 (w.) SIERRA LEONE. Type-material: holotype worker. Type-locality: Sierra Leone: N’Gamie River, Samlia Falls. Type-depository: unknown; Mayr has “k. belgisches Museum”, therefore perhaps ISNB or MRAC. Status as species: Santschi, 1910c: 389; Wheeler, W.M. 1922a: 914; Santschi, 1924b: 217; Emery, 1924d: 297; Santschi, 1930a: 69; Bolton, 1974a: 22 (redescription); Bolton, 1982: 355 (in key); Bolton, 1995b: 138. Senior synonym of brazzavillensis: Bolton, 1974a: 22; Bolton, 1995b: 138. Senior synonym of foveolatus: Bolton, 1974a: 22; Bolton, 1995b: 138. Senior synonym of latipes: Bolton, 1974a: 22; Bolton, 1995b: 138. Distribution: Cameroon, Congo, Democratic Republic of Congo, Equatorial Guinea, Sierra Leone, Uganda. krugeri. Cataulacus rugosus var. krugeri Prins, 1965d: 104, pl. 1, figs. 1-3, pl. 2, figs. 1-2 (w.q.m.) SOUTH AFRICA; unavailable name. [Note: material included: South Africa: Pumbe (AcAx 1800), 12.v.1962 (A.J. Prins); 537 workers, 71 queens, 2 males, deposited in RPPP (no type status).] [Unavailable name (variety proposed after 1960): Bolton, 1974a: 43.] Material referred to intrudens by Bolton, 1974a: 43; Bolton, 1995b: 138. latipes. Cataulacus latipes Menozzi, 1933a: 106, fig. 4 (w.) UGANDA. Type-material: 4 syntype workers. Type-localities: 1 worker Uganda: Entebbe (E. Bayon); 3 workers Uganda: Victoria Nyanza, Sesse I., Bugala (E. Bayon). Type-depository: IEUB. Status as species: Menozzi, 1942: 177; Eidmann, 1944: 469. Junior synonym of kohli: Bolton, 1974a: 22; Bolton, 1995b: 138. latissimus. Cataulacus latissimus Emery, 1893e: 215, pl. 8, fig. 10 (w.) WEST MALAYSIA. Type-material: 2 syntype workers. Type-locality: Malaysia: Perak, near Malacca I. (Bedot & Pictet). Type-depository: MSNG. Viehmeyer, 1922: 212 (q.); Bolton, 1974a: 78 (m.). Status as species: Forel, 1912n: 60; Wheeler, W.M. 1919e: 94; Viehmeyer, 1922: 212; Emery, 1924d: 298; Santschi, 1928h: 125; Chapman & Capco, 1951: 85; Bolton, 1974a: 77 (redescription); Bolton, 1995b: 138; Pfeiffer, et al. 2011: 44. Senior synonym of mimula: Bolton, 1974a: 77; Bolton, 1995b: 138. Distribution: Brunei, Malaysia (Peninsula, Sarawak), Singapore. latus. Cataulacus latus Forel, 1891b: 144 (m.) INDIA (no state data). Type-material: syntype males (number not stated). [Note: syntype male series also contains 3 queens, not mentioned by Forel, 1891b. This series, together with some workers (now lost), correspond with the entry in Wroughton, 1892: 178, “120. Cat. latus (Forel MS. Poona Dists. 14.vi.90. Specimens in USNM from Kanara, LVI/6 (Wroughton), which bear “cotype” labels, have no type-status (Bolton, 1974a: 78).] Type-locality: India: Poona, 16.vi.1890 (R.C. Wroughton). Type-depository: MHNG. Forel, 1903a: 706 (w., in key); Bingham, 1903: 122 (q.). Status as species: Forel, 1903a: 707; Bingham, 1903: 121; Emery, 1924d: 298; Donisthorpe, 1942d: 457; Chapman & Capco, 1951: 85; Bolton, 1974a: 78 (redescription); Bolton, 1995b: 138; Tiwari, 1999: 65; Bharti, Guénard, et al. 2016: 35. Distribution: India, Myanmar. linearis. Cataulacus wissmanni st. linearis Santschi, 1914b: 109, fig. 17 (w.) KENYA. Type-material: 2 syntype workers. Type-localities: 1 worker Kenya (“Afrique orientale anglaise”): Voi, Wa-Taita, st.no.60, iii.1912 (Alluaud & Jeannel); 1 worker Kenya: Mbuyuni, Pori, st.no.63 (Alluaud & Jeannel). Type-depository: NHMB. Subspecies of wissmannii: Wheeler, W.M. 1922a: 917; Emery, 1924d: 297. Junior synonym of wissmannii: Bolton, 1974a: 51; Bolton, 1995b: 138. lobatus. Cataulacus lobatus Mayr, 1895: 126, fig. 1 (w.) CAMEROON. Type-material: holotype worker. Type-locality: Cameroon: Kriegsschiffhafen, 15.iii.1892. Type-depository: NHMW. Status as species: Wheeler, W.M. 1922a: 914; Emery, 1924d: 297; Bolton, 1974a: 23 (redescription); Bolton, 1982: 355 (in key); Bolton, 1995b: 138. Distribution: Cameroon, Democratic Republic of Congo. longinodus. Cataulacus granulatus var. longinoda Forel, 1912n: 60 (q.) INDONESIA (Sumatra). Type-material: holotype queen. Type-locality: Indonesia: Sumatra, Indrapura (Tritschler). Type-depository: MHNG. Subspecies of granulatus: Emery, 1924d: 298; Chapman & Capco, 1951: 85. Status as species: Bolton, 1974a: 67 (redescription); Bolton, 1995b: 138. Distribution: Indonesia (Sumatra). longispinus. Cataulacus huberi var. longispinus Stitz, 1910: 139, fig. 8 (w.) CAMEROON. Type-material: holotype worker. Type-locality: Cameroon: Mundame (Conradt). Type-depository: MNHU. [Misspelled as longispinosus by Emery, 1924d: 296.] Subspecies of huberi: Wheeler, W.M. 1922a: 914; Emery, 1924d: 296; Wheeler, W.M. 1925b: 2. Junior synonym of huberi: Bolton, 1974a: 19; Bolton, 1995b: 138. loveridgei. Cataulacus jeanneli var. loveridgei Santschi, 1926b: 244 (w.) TANZANIA. Type-material: holotype worker. Type-locality: Tanzania (“Tanganika”): Morogoro, 21.xi.1917 (A. Loveridge). Type-depository: unknown. Junior synonym of brevisetosus: Bolton, 1974a: 31. Junior synonym of jeanneli: Bolton, 1982: 358; Bolton, 1995b: 138. luebensis. Cataulacus huberi st. herteri var. luebensis Santschi, 1937a: 57 (w.) DEMOCRATIC REPUBLIC OF CONGO; unavailable (infrasubspecific) name. [Note: material included: Democratic Republic of Congo (“Congo belge”): Luebo, Kamajembi, 22.ix.1921 (H. Schouteden); 2 workers, deposited in NHMB (no type status).] Declared as unavailable (infrasubspecific) name: Bolton, 1974a: 19. Material referred to huberi by Bolton, 1974a: 19; Bolton, 1995b: 138. lujae. Cataulacus lujae Forel, 1911c: 311 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: syntype workers (number not stated). Type-locality: Democratic Republic of Congo (“Belg. Kongo”): Kasai, Kondue (Luja). Type-depository: MRAC. Forel, 1914d: 220 (q.m.); Forel, 1916: 427 (q.m.). Subspecies of brevisetosus: Forel, 1914d: 220; Forel, 1916: 427; Arnold, 1917: 397. Subspecies of pygmaeus: Santschi, 1916b: 506 (footnote); Wheeler, W.M. 1922a: 199, 916; Emery, 1924d: 297. Junior synonym of brevisetosus: Bolton, 1974a: 31. Status as species: Forel, 1913b: 316; Santschi, 1924b: 219; Bolton, 1982: 356 (in key), 358; Bolton, 1995b: 138; Hita Garcia, et al. 2013: 209. Senior synonym of gilviventris: Bolton, 1982: 358; Bolton, 1995b: 138. Distribution: Cameroon, Democratic Republic of Congo, Ghana, Kenya, Nigeria. marginatus. Cataulacus marginatus Bolton, 1974a: 68, fig. 38 (w.) CHINA (Hainan I.). Type-material: holotype worker, 9 paratype workers. Type-locality: holotype China: Hainan I., grove near Hoi Man Chuen, SW of Nodoa, 4.vii.1929, Lingnan University 5th Hainan Is expedition, 1929 (no collector’s name); paratypes: 4 workers Hainan Is, Ta Hau, 7.vii.1935 (J.L. Gressitt), 1 worker with same data but 4-5.vii.1935, 4 workers Hainan Is, Nodoa, 15- 17.iii.1935 ((J.L. Gressitt). Type-depository: MCZC (holotype); BMNH, MCZC (paratypes). Status as species: Bolton, 1995b: 138; Guénard & Dunn, 2012: 41. Distribution: China (Hainan I.). marleyi. Cataulacus marleyi Forel, 1914d: 219 (w.) SOUTH AFRICA. Type-material: syntype workers (number not stated). Type-locality: South Africa: Natal, Krantz Kloof (H.B. Marley). Type-depository: MHNG. Arnold, 1917: 399 (q.). Status as species: Arnold, 1917: 398; Wheeler, W.M. 1922a: 914; Emery, 1924d: 297. Junior synonym of pygmaeus: Bolton, 1974a: 48. Junior synonym of traegaordhi: Bolton, 1982: 359; Bolton, 1995b: 139. mckeyi. Cataulacus mckeyi Snelling, R.R. 1979a: 5, figs. 9-18 (w.q.m.) CAMEROON. Type-material: holotype worker, 135 paratype workers, 20 paratype queens, 7 paratype males. Type-locality: holotype Cameroon: near Lac Tissongo, Douala-Edea Reserve, 3°29’N, 9°50’E, 5 km. S Sanaga River, 15 km. E Mouanko, 18.vii.1976 (D. McKey); paratypes with same data. Type-depositories: LACM (holotype); BMNH, DMPC, LACM, MCZC. Status as species: Bolton, 1982: 358 (in key); McKey, 1984: 81; Bolton, 1995b: 139. Distribution: Cameroon. meduseus. Cataulacus meduseus Santschi, 1939b: 245 (w.) SOUTH AFRICA. Type-material: holotype worker. Type-locality: South Africa: Natal, Durban, 24.i.1917 (C.P. Merve). Type-depository: SAMC. Junior synonym of brevisetosus: Bolton, 1974a: 31; Bolton, 1995b: 139. micans. Cataulacus rugosus subsp. micans Mayr, 1901b: 27 (w.q.m.) SOUTH AFRICA. Type-material: syntype workers, syntype queens, syntype males (numbers not stated). Type-locality: South Africa: Port Elizabeth, 1890 (H. Brauna). Type-depository: NHMW. Status as species: Forel, 1914d: 219; Arnold, 1917: 393; Wheeler, W.M. 1922a: 915; Emery, 1924d: 297; Bolton, 1974a: 46 (redescription); Bolton, 1982: 358 (in key); Bolton, 1995b: 139. Senior synonym of tristiculus: Bolton, 1974a: 46; Bolton, 1995b: 139. Distribution: South Africa. mimula. Cataulacus latissimus var. mimula Menozzi, 1923b: 210 (w.) BORNEO (Brunei). Type-material: syntype workers (number not stated, “plusiers ouvrières”). Type-locality: Brunei: Borneo septentrional (Staudinger & Bang-Haas). Type-depository: IEUB. Subspecies of latissimus: Chapman & Capco, 1951: 85. Junior synonym of latissimus: Bolton, 1974a: 77; Bolton, 1995b: 139. minor. Cataulacus reticulatus var. minor Smith, F. 1857a: 81 (w.) BORNEO (East Malaysia: Sarawak). Type-material: holotype worker. Type-locality: Malaysia (“Borneo”): Sarawak (A.R. Wallace). Type-depository: OXUM (absent from collection, presumed lost). Junior synonym of reticulatus: Bolton, 1974a: 82; Bolton, 1995b: 139. mocquerysi. Cataulacus mocquerysi André, 1889: 229 (w.) SIERRA LEONE. Type-material: holotype worker. Type-locality: Sierra Leone: (no further data). Type-depository: MNHN. [Misspelled as moquerisi by Emery, 1924d: 297.] Bolton, 1974a: 48 (q.). Status as species: Dalla Torre, 1893: 138; Wheeler, W.M. 1922a: 915; Emery, 1924d: 297; Bolton, 1974a: 47 (redescription); Bolton, 1982: 355 (in key); Bolton, 1995b: 139. Senior synonym of nainei: Bolton, 1974a: 47; Bolton, 1995b: 139. Distribution: Cameroon, Democratic Republic of Congo, Ghana, Liberia, Nigeria, Sierra Leone. moloch. Cataulacus moloch Bolton, 1982: 361, fig. 34 (w.) GHANA, NIGERIA. Type-material: holotype worker, 3 paratype workers. Type-locality: holotype Ghana: Pankese, 24.ix.1968 (C.A. Collingwood); paratypes: 1 worker with same data, 2 workers Nigeria: Onipe, CRIN, 11.vi.1975, tree 47-169 A63.1, black pod project (B. Taylor). Type-depository: BMNH. Status as species: Bolton, 1995b: 139; Hita Garcia, et al. 2013: 209. Distribution: Ghana, Kenya, Nigeria. muticus. Cataulacus muticus Emery, 1889b: 507, pl. 10, fig. 17 (w.) MYANMAR. Type-material: holotype worker. Type-locality: Myanmar (“Birmania”): Tenasserim, Thagata, Mt Mooleyit (L. Fea). Type-depository: MSNG. Status as species: Dalla Torre, 1893: 138; Emery, 1895k: 473; Forel, 1903a: 706; Bingham, 1903: 124; Emery, 1924d: 298; Chapman & Capco, 1951: 85; Bolton, 1974a: 69; Bolton, 1995b: 139; Mohanraj, et al. 2010: 6; Bharti, Guénard, et al. 2016: 35. Distribution: India (Andaman Is), Myanmar. nainei. Cataulacus mocquerysi var. nainei Forel, 1918a: 724 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: holotype worker. Type-locality: Democratic Republic of Congo (“Congo”): (no further data) (H. Kohl). Type-depository: MHNG. Subspecies of mocquerysi: Wheeler, W.M. 1922a: 915; Emery, 1924d: 297. Junior synonym of mocquerysi: Bolton, 1974a: 47; Bolton, 1995b: 139. nenassus. Cataulacus nenassus Bolton, 1974a: 70, fig. 41 (w.q.m.) INDONESIA (Java). Type-material: holotype worker, 9 paratype workers, 7 paratype queens, 2 paratype males. Type-locality: holotype Indonesia: Java, Semarang, no. g45, 7.ii.1928 (L.G.E. Kalshoven); paratypes: 9 workers, 6 queens, 2 males with same data as holotype, 1 queen ‘Java zee’, 11.ix.1920 (Kl. Kombuis). Type-depositories: MCZC (holotype); BMNH, MCZC, USNM (paratypes). Status as species: Bolton, 1995b: 139. Distribution: Indonesia (Java). oberthueri. Cataulacus (Otomyrmex) oberthueri Emery, in Forel, 1891b: 146, pl. 4, fig. 9 (w.) MADAGASCAR. Type-material: syntype workers (number not stated). Type-locality: Madagascar: Tamatave and Alaha Kato (E. Perrot). Type-depository: MRAC, MSNG. Status as species: Dalla Torre, 1893: 138; Emery, 1899f: 286; Wheeler, W.M. 1922a: 1033; Emery, 1924d: 299; Bolton, 1974a: 24 (redescription); Bolton, 1995b: 139. Distribution: Madagascar. orientalis. Cataulacus pullus var. orientalis Santschi, 1914b: 108 (w.) KENYA. Type-material: holotype worker. Type-locality: Kenya (“Afrique orientale anglaise”): Voi, 600 m., st.no.60, iii.1912 (Alluaud & Jeannel). Type-depository: NHMB. Subspecies of pullus: Wheeler, W.M. 1922a: 915; Emery, 1924d: 297. Junior synonym of pullus: Bolton, 1974a: 26; Bolton, 1995b: 139. otii. Cataulacus wissmanni r. otii Forel, 1901d: 304 (w.q.) SOUTH AFRICA. Type-material: syntype workers (number not stated), 1 syntype queen. [Note: syntype queen is not mentioned in the description.] Type-locality: South Africa: Natal, Durban (Haviland). Type-depository: MHNG. Status as species: Santschi, 1914e: 26; Arnold, 1917: 401; Wheeler, W.M. 1922a: 915; Emery, 1924d: 297. Subspecies of wissmanni: Santschi, 1930b: 56. Junior synonym of wissmannii: Bolton, 1974a: 51; Bolton, 1995b: 139. parallelus. Cataulacus parallelus Smith, F. 1853: 226, pl. 19, fig. 6 (q.) South Africa. Type-material: holotype queen. Type-locality: South Africa: Cape of Good Hope. [Note: type-locality incorrect or specimen incorrectly labelled; species distribution is forested West and Central Africa (Bolton, 1974a: 55).] Type-depository: OXUM. Status as species: Smith, F. 1858b: 196; Smith, F. 1862d: 414; Mayr, 1863: 403; Roger, 1863b: 39; Dalla Torre, 1893: 138; Arnold, 1917: 402; Wheeler, W.M. 1922a: 915; Emery, 1924d: 297. Junior synonym of guineensis: Bolton, 1974a: 55; Bolton, 1995b: 139. pilosus. Cataulacus pilosus Santschi, 1920c: 118 (w.q.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: 1 syntype worker, 1 syntype queen. Type-locality: Democratic Republic of Congo (“Congo belge”): Avakubi, 6.i.1914 (Bequaert). Type-depository: NHMB. Status as species: Wheeler, W.M. 1922a: 915; Bolton, 1974a: 35 (redescription); Bolton, 1982: 356 (in key); Bolton, 1995b: 139. Distribution: Democratic Republic of Congo. *planiceps. *Cataulacus planiceps Emery, 1891a: 148, pl. 1, figs. 8, 9 (w.) SICILIAN AMBER (Italy: Sicily, Oligocene). Type-material: 2 syntype workers. Type-locality: Italy: Sicily, in amber (Bombicci). Type-depository: MMUB. Status as species: Dalla Torre, 1893: 139; Handlirsch, 1907: 877; Bolton, 1974a: 87; Keilbach, 1982: 276; Bolton, 1995b: 139. *plebeius. *Cataulacus plebeius De Andrade & Baroni Urbani, 2004: 248, pl. 1, figs. B, C, E. BORNEO AMBER (Borneo: no state data, Miocene). Type-material: holotype worker, 1 paratype worker. Type-locality: holotype Borneo: fragment A (no further data); paratype: fragment B (no further data). Type-depository: apparently in JWPC. plebeja. Cataulacus pygmaeus st. lujae var. plebeja Santschi, 1916b: 508 (w.) ZIMBABWE; unavailable (infrasubspecific) name. [Note: material included: Zimbabwe (“Rhodesia”): Bulawayo, ix.1914 (G. Arnold), deposited in NHMB (no type status).] As unavailable (infrasubspecific) name: Wheeler, W.M. 1922a: 916. Declared as unavailable (infrasubspecific) name: Bolton, 1974a: 31. Material referred to brevisetosus by Bolton, 1974a: 31; Bolton, 1995b: 139. plectroniae. Cataulacus traegaordhi var. plectroniae Wheeler, W.M. 1922a: 199 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: 24 syntype workers. Type-locality: Democratic Republic of Congo (“Belgian Congo”): Stanleyville, from cavities of Plectronia (Lang & Chapin). Type-depository: MCZM. Junior synonym of weissi: Bolton, 1974a: 39; Bolton, 1995b: 139. pompom. Cataulacus pompom Snelling, R.R. 1979a: 8, figs. 19-26 (w.) WEST MALAYSIA. Type-material: holotype worker, 9 paratype workers. Type-locality: holotype Malaysia (“Malaya”): Selangor, Gombak, 2.x.1973 (B. Bolton); paratypes with same data. Type-depositories: BMNH (holotype); BMNH, LACM (paratypes). Status as species: Bolton, 1995b: 139. Distribution: Malaysia (Peninsula). porcatus. Cataulacus porcatus Emery, 1899f: 286 (w.q.) MADAGASCAR. Type-material: 2 syntype workers, 1 syntype queen. Type-locality: Madagascar: Antongil Bay, 1897-98 (A. Mocquerys). Type-depository: MSNG. Status as species: Wheeler, W.M. 1922a: 1033; Emery, 1924d: 298; Bolton, 1974a: 25 (redescription); Bolton, 1995b: 139. Distribution: Madagascar. praetextus. Cataulacus praetextus Smith, F. 1867: 528, fig. 5 (w.) BORNEO (no state data). Type-material: holotype worker. Type-locality: Borneo: (no further data). Type-depository: OXUM. Bolton, 1974a: 81 (q.m.). Status as species: Dalla Torre, 1893: 139; Wheeler, W.M. 1919e: 93; Emery, 1924d: 298; Chapman & Capco, 1951: 85; Bolton, 1974a: 80 (redescription); Bolton, 1995b: 139; Pfeiffer, et al. 2011: 44. Senior synonym of sumatrensis: Bolton, 1974a: 80; Bolton, 1995b: 139. Distribution: Indonesia (Sumatra), Malaysia (Peninsula, Sarawak), Thailand. princeps. Cataulacus princeps Forel, 1909b: 71. Nomen nudum (attributed to Emery). Material referred to erinaceus by Wheeler, W.M. 1922a: 913; Emery, 1924d: 296; Bolton, 1995b: 139. pseudotrema. Cataulacus baumi st. pseudotrema Santschi, 1926b: 244 (w.) TANZANIA. Type-material: 2 syntype workers. Type-locality: Tanzania (“Tanganika”): Duthumi, 18.ix.1919 (A. Loveridge). Type-depository: NHMB. Subspecies of baumi: Santschi, 1928f: 209. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 139. pullus. Cataulacus pullus Santschi, 1910c: 387, fig. 13 (w.) CONGO. Type-material: holotype worker. Type-locality: Congo (“Congo français”): Brazzaville (A. Weiss). Type-depository: NHMB. Status as species: Wheeler, W.M. 1922a: 915; Santschi, 1924b: 217; Emery, 1924d: 297; Bernard, 1953b: 252; Bolton, 1974a: 26 (redescription); Bolton, 1982: 355 (in key); Bolton, 1995b: 139; Hita Garcia, et al. 2013: 209. Senior synonym of coriaceus: Bolton, 1974a: 26; Bolton, 1995b: 139. Senior synonym of orientalis: Bolton, 1974a: 26; Bolton, 1995b: 139. Distribution: Cameroon, Congo, Democratic Republic of Congo, Gabon, Kenya. pygmaeus. Cataulacus pygmaeus André, 1890: 325 (w.) SIERRA LEONE. Type-material: holotype worker. Type-locality: Sierra Leone: (no further data) (A. Mocquerys). Type-depository: MNHN. Status as species: Dalla Torre, 1893: 139; Santschi, 1910c: 390; Wheeler, W.M. 1922a: 199, 915; Emery, 1924d: 297; Weber, 1943c: 378; Bolton, 1974a: 48 (redescription); Bolton, 1982: 357 (in key); Bolton, 1995b: 139; Hita Garcia, et al. 2013: 209. Senior synonym of bakusuensis: Bolton, 1974a: 48; Bolton, 1995b: 139. Senior synonym of chariensis: Bolton, 1974a: 48; Bolton, 1995b: 139. Distribution: Cameroon, Chad, Democratic Republic of Congo, Ghana, Ivory Coast, Kenya, Liberia, Sierra Leone, Tanzania. regularis. Cataulacus regularis Forel, 1892l: 252 (w.) MADAGASCAR. Type-material: syntype workers (number not stated). Type-locality: Madagascar: Bezanozano, Anosibé, ESE Antananarivo (Sikora). Type-depository: MHNG. Status as species: Dalla Torre, 1893: 139; Wheeler, W.M. 1922a: 1033; Emery, 1924d: 298; Bolton, 1974a: 27 (redescription); Bolton, 1995b: 139. Distribution: Madagascar. resinosus. Cataulacus taprobanae var. resinosa Viehmeyer, 1913: 145 (w.) INDONESIA (Sulawesi) (in copal). Type-material: 2 syntype workers. Type-locality: Indonesia: Sulawesi (“Celebes”) (no frther data), in copal. Type-depositories: MNHU; second specimen possibly in SMTD. [Misspelled as resinoda by Chapman & Capco, 1951: 86.] Subspecies of taprobanae: Emery, 1924d: 299. Status as species: Chapman & Capco, 1951: 86; Bolton, 1974a: 86; Bolton, 1995b: 139. Distribution: Indonesia (Sulawesi). reticulatus. Cataulacus reticulatus Smith, F. 1857a: 81, pl. 2, fig. 8 (w.) BORNEO (East Malaysia: Sarawak). Type-material: holotype worker. Type-locality: Malaysia (“Borneo”): Sarawak (“SAR.5”) (A.R. Wallace). Type-depository: OXUM. Junior synonym of granulatus: Dalla Torre, 1893: 138; Wheeler, W.M. 1919e: 92; Emery, 1924d: 298. Status as species: Smith, F. 1858b: 196; Smith, F. 1862d: 415; Mayr, 1863: 403; Roger, 1863b: 39; Smith, F. 1871a: 335; Donisthorpe, 1932c: 451; Bolton, 1974a: 82 (redescription); Bolton, 1995b: 139; Pfeiffer, et al. 2011: 44. Senior synonym of minor: Bolton, 1974a: 82; Bolton, 1995b: 139. Distribution: Malaysia (Sarawak). rudis Mayr, 1870; see under PROCRYPTOCERUS. rugosus. Cataulacus intrudens var. rugosus Forel, 1894b: 78 (w.) MOZAMBIQUE. Type-material: syntype workers (number not stated). Type-locality: Mozambique: Delagoa, in gall, 2.xi.1890 (Müller, Liengme). Type-depository: MHNG. Mayr, 1895: 129 (q.). Status as species: Mayr, 1895: 129; Mayr, 1901b: 27; Arnold, 1917: 392; Wheeler, W.M. 1922a: 916; Emery, 1924d: 297; Santschi, 1937d: 234; Prins, 1963: 104. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 139. satrap. Cataulacus satrap Bolton, 1982: 363 (w.) CAMEROON. Type-material: holotype worker, 1 paratype worker. Type-locality: holotype Cameroon: Nkoemvon, 1970, M12 (D. Jackson); paratype with same data. Type-depository: BMNH. Status as species: Bolton, 1982: 356 (in key); Bolton, 1995b: 139. Distribution: Cameroon. schoutedeni. Cataulacus schoutedeni Santschi, 1919c: 248 (w.q.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: 4 syntype workers, 1 syntype queen. Type-locality: Democratic Republic of Congo: Congo da Lemba, i-ii.1913 (R. Mayné). Type-depositories: MRAC, NHMB. Status as species: Wheeler, W.M. 1922a: 917; Santschi, 1924b: 217. Junior synonym of tardus: Bolton, 1974a: 27; Bolton, 1995b: 139. setosus. Cataulacus setosus Smith, F. 1860b: 114, pl. 1, fig. 7 (w.) INDONESIA (Bacan I.). Type-material: holotype worker. Type-locality: Indonesia: Moluccas, Bacan I. (“Bachian”), (“Bac.31.”) (A.R. Wallace). Type-depository: OXUM. Smith, F. 1863: 24 (q.). Status as species: Smith, F. 1862d: 415; Smith, F. 1863: 24; Mayr, 1863: 403; Roger, 1863b: 39; Smith, F. 1865: 76; Smith, F. 1871a: 334; Dalla Torre, 1893: 139; Emery, 1924d: 298; Donisthorpe, 1932c: 464; Donisthorpe, 1943d: 453; Chapman & Capco, 1951: 86; Bolton, 1974a: 71 (redescription); Taylor, 1991b: 601; Bolton, 1995b: 139. Distribution: Indonesia (Bacan, Waigeu), Papua New Guinea, Philippines (Mindanao). *silvestrii. *Cataulacus silvestrii Emery, 1891a: 147, pl. 1, figs. 5-7 (w.) SICILIAN AMBER (Italy: Sicily, Oligocene). Type-material: holotype worker. Type-locality: Italy: Sicily, in amber (Bombicci). Type-depository: MMUB. Status as species: Dalla Torre, 1893: 139; Handlirsch, 1907: 876; Bolton, 1974a: 86; Keilbach, 1982: 276; Bolton, 1995b: 139. simoni. Cataulacus simoni Emery, 1893f: 248 (w.) SRI LANKA. Type-material: 2 syntype workers . Type-localities: Sri Lanka (“Ceylan”): 1 syntype Kandy, i.-ii.1892 (E. Simon), 1 syntype Colombo, i.-ii.1892 (E. Simon). Type-depository: MSNG. Forel, 1909e: 393 (q.). Status as species: Forel, 1903a: 706; Bingham, 1903: 123; Forel, 1908a: 2; Forel, 1909e: 393; Emery, 1924d: 299; Chapman & Capco, 1951: 86; Bolton, 1974a: 72 (redescription); Bolton, 1995b: 139; Mathew & Tiwari, 2000: 322; Mohanraj, et al. 2010: 6; Guénard & Dunn, 2012: 41; Bharti, Guénard, et al. 2016: 35. Senior synonym of andamanensis: Bolton, 1974a: 72; Bolton, 1995b: 139. Distribution: China, India (India, Andaman Is), Sri Lanka. simplex. Cataulacus egenus st. simplex Santschi, 1914b: 111, fig 18 (w.) UGANDA. Type-material: holotype worker. Type-locality: Uganda: Central Region, i.1909 (Ch. Alluaud). Type-depository: NHMB, not in collection, presumed lost. Subspecies of egenus: Wheeler, W.M. 1922a: 913; Santschi, 1924b: 218; Emery, 1924d: 296; Santschi, 1930a: 69. Junior synonym of egenus: Bolton, 1974a: 18; Bolton, 1995b: 139. striativentris. Cataulacus wissmanni var. striativentris Santschi, 1924b: 219 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: syntype workers (number not stated).. Type-localities: Democratic Republic of Congo (“Congo belge”): Ubanghi, Banzyville (R.P. Augustin); Haut Uele, Moto, 1920 (L. Burgeon). Type-depositories: MRAC, NHMB. Status as species: Bolton, 1974a: 36 (redescription); Bolton, 1982: 356 (in key); Bolton, 1995b: 139; Hita Garcia, et al. 2013: 209. Senior synonym of donisthorpei: Bolton, 1974a: 36; Bolton, 1995b: 139. Distribution: Democratic Republic of Congo, Kenya. subrugosus. Cataulacus rugosus var. subrugosus Santschi, 1914e: 26 (w.) SOUTH AFRICA. Type-material: 4 syntype workers. Type-locality: South Africa: Natal, Zululand, Junction of Umfolozis, 29.vi.1905 (I. Trägårdh). Type-depository: NHMB. Santschi, 1937d: 236 (q.). Subspecies of rugosus: Arnold, 1917: 393; Wheeler, W.M. 1922a: 917; Emery, 1924d: 297; Santschi, 1937d: 234. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 139. suddensis. Cataulacus pygmaeus subsp. suddensis Weber, 1943c: 378 (w.m.) SOUTH SUDAN. Type-material: syntype workers (number not stated, “several workers”), 1 syntype male. Type-locality: South Sudan (“Anglo-Egyptian Sudan”): Upper White Nile, Adok, in the Sudd, 10.vii.1939, No. 1244 (N.A. Weber). Type-depositories: AMNH, MCZC. Junior synonym of pygmaeus: Bolton, 1974a: 49. Junior synonym of traegaordhi: Bolton, 1982: 359; Bolton, 1995b: 139. sulcatus. Cataulacus sulcatus Stitz, 1910: 136, figs. 4-6 (w.q.m.) CAMEROON. Type-material: syntype workers, syntype queens, syntype males (numbers not stated). Type-locality: Cameroon: Jaundestation (Zenker). Type-depository: MNHU. Subspecies of guineensis: Forel, 1911e: 263; Wheeler, W.M. 1922a: 913; Emery, 1924d: 296. Junior synonym of guineensis: Forel, 1910e: 421; Bolton, 1974a: 55; Bolton, 1995b: 139. sulcinodis. Cataulacus guineensis r. sulcinodis Emery, 1892d: 563, pl. 15, fig. 8 (w.) IVORY COAST. Type-material: holotype worker. Type-locality: Ivory Coast: Assinie, vii-viii.1886 (Ch. Alluaud). Type-depository: MSNG. Status as species: Dalla Torre, 1893: 139. Subspecies of guineensis: Emery, in Dalla Torre, 1893: 138 (footnote); Wheeler, W.M. 1922a: 913; Emery, 1924d: 296. Junior synonym of guineensis: Bolton, 1974a: 55; Bolton, 1995b: 139. sumatrensis. Cataulacus praetextus var. sumatrensis Forel, 1912n: 60 (w.) INDONESIA (Sumatra). Type-material: holotype worker. Type-locality: Indonesia: Sumatra, Indrapura (Tritschler). Type-depository: MHNG. Viehmeyer, 1916a: 140 (q.). Subspecies of praetextus: Viehmeyer, 1916a: 140; Emery, 1924d: 298; Chapman & Capco, 1951: 85. Junior synonym of praetextus: Bolton, 1974a: 80; Bolton, 1995b: 139. tangana. Cataulacus baumi st. pseudotrema var. tangana Santschi, 1928f: 209 (w.q.) TANZANIA; unavailable (infrasubspecific) name. [Note: material included: Tanzania (“Afrique orientale”): Tanga, 6.v.1925 (G. Arnold), 4 workers, 1 queen, deposited in NHMB (no type status.] Declared as unavailable (infrasubspecific) name: Bolton, 1974a: 43. Material referred to intrudens by Bolton, 1974a: 43; Bolton, 1995b: 139. taprobanae. Cataulacus taprobanae Smith, F. 1853: 225, pl. 20, fig. 10 (w.) SRI LANKA. Type-material: holotype worker. Type-locality: Sri Lanka (“Ceylon”): (no further data) (G.H.K. Thwaites). Type-depository: BMNH. Forel, 1909e: 393 (q.); Wheeler, G.C. & Wheeler, J. 1954b: 150 (l.); Bolton, 1974a: 83 (m.). Status as species: Smith, F. 1858b: 195; Smith, F. 1862d: 414; Mayr, 1863: 403; Roger, 1863b: 39; Motschoulsky, 1863: 21; Smith, F. 1871a: 335; Dalla Torre, 1893: 139; Emery, 1893f: 248; Emery, 1901f: 121; Forel, 1903a: 706; Rothney, 1903: 97; Bingham, 1903: 123; Forel, 1908a: 2; Forel, 1909e: 393; Forel, 1911i: 226; Forel, 1913k: 83; Emery, 1924d: 299; Mukerjee, 1930: 155; Mukerjee, 1934: 4; Chapman & Capco, 1951: 86; Bolton, 1974a: 83 (redescription); Bolton, 1995b: 139; Mathew & Tiwari, 2000: 321; Mohanraj, et al. 2010: 6; Guénard & Dunn, 2012: 41; Bharti, Guénard, et al. 2016: 35. Distribution: China, India (India, Andaman Is), Sri Lanka. tardus. Cataulacus tardus Santschi, 1914d: 372, fig. 33 (w.) GUINEA. Type-material: holotype worker. Type-locality: Guinea (“Guinée française”): Mamou, 24.viii.1912 (F. Silvestri). Type-depository: NHMB. Status as species: Wheeler, W.M. 1922a: 917; Emery, 1924d: 297; Bolton, 1974a: 27 (redescription); Bolton, 1982: 355 (in key); Bolton, 1995b: 139; Hita Garcia, et al. 2013: 209. Senior synonym of schoutedeni: Bolton, 1974a: 27; Bolton, 1995b: 139. Distribution: Cameroon, Democratic Republic of Congo, Ghana, Guinea, Kenya. taylori. Cataulacus taylori Bolton, 1982: 364 (w.) NIGERIA. Type-material: holotype worker, 1 paratype worker. Type-locality: holotype Nigeria: Gambari, CRIN, 24.v.1976, black pod project (B. Taylor); paratype: Nigeria: Onipe, CRIN, 25.vii.1975, black pod project (B. Taylor). Type-depository: BMNH. Status as species: Bolton, 1995b: 139. Distribution: Nigeria. tenuis. Cataulacus tenuis Emery, 1899f: 288 (q.) MADAGASCAR. Type-material: holotype queen. Type-locality: Madagascar: Antongil Bay, 1897-98 (A. Mocquerys). Type-depository: MSNG. Santschi, 1913c: 310 (w.). Status as species: Santschi, 1913c: 310; Wheeler, W.M. 1922a: 1033; Emery, 1924d: 298; Bolton, 1974a: 37; Bolton, 1995b: 139. Distribution: Madagascar. theobromicola. Cataulacus theobromicola Santschi, 1939c: 8, fig. 2 (w.) DEMOCRATIC REPUBLIC OF CONGO. Type-material: holotype worker. Type-locality: Democratic Republic of Congo (“Congo belge”): Gazi, xii.1937 (Baenaert). Type-depository: MRHN. [Misspelled as theobromicolus by Bolton, 1974a: 28.] Status as species: Bolton, 1974a: 28 (redescription); Bolton, 1982: 355 (in key); Bolton, 1995b: 139. Distribution: Democratic Republic of Congo. traegaordhi. Cataulacus traegaordhi Santschi, 1914e: 24, fig. 3 (w.q.m.) SOUTH AFRICA. Type-material: syntype workers, syntype queen(s), syntype male(s) (numbers not stated). Type-localities: workers, queen, male South Africa: Natal, Zululand, Dukudu, 27.vii.1905 (I. Trägårdh); workers Natal, Umfolosi, 6.vi.1905 (I. Trägårdh). Type-depositories: MRAC, NHMB. Junior synonym of pygmaeus: Bolton, 1974a: 48. Status as species: Arnold, 1917: 399; Wheeler, W.M. 1922a: 199, 917; Santschi, 1924b: 220; Emery, 1924d: 297; Santschi, 1935b: 273; Weber, 1943c: 326; Bernard, 1953b: 252; Bolton, 1982: 358 (in key), 359; Bolton, 1995b: 139; Hita Garcia, et al. 2013: 209. Senior synonym of marleyi: Bolton, 1982: 359; Bolton, 1995b: 139. Senior synonym of suddensis: Bolton, 1982: 359; Bolton, 1995b: 139. Senior synonym of ugandensis: Bolton, 1982: 359; Bolton, 1995b: 139. Distribution: Cameroon, Democratic Republic of Congo, Ghana, Kenya, Nigeria, South Africa, South Sudan, Tanzania, Uganda. tristiculus. Cataulacus intrudens st. tristiculus Santschi, 1919b: 237 (w.q.m.) SOUTH AFRICA. Type-material: 2 syntype workers, 1 syntype queen, 1 syntype male. Type-locality: South Africa: Cape Province, Port Elizabeth, 1917 (T. Reese). Type-depository: NHMB. Subspecies of intrudens: Wheeler, W.M. 1922a: 914; Arnold, 1926: 286. Junior synonym of micans: Bolton, 1974a: 46; Bolton, 1995b: 139. ugandensis. Cataulacus traegaordhi var. ugandensis Santschi, 1914b: 110 (w.) UGANDA. Type-material: 5 syntype workers. Type-locality: Uganda: Unyoro Prov., near Hoima, i.1909 (Ch. Alluaud). Type-depository: NHMB. [Misspelled in original reference as trägardhi.] Subspecies of traegaordhi: Wheeler, W.M. 1922a: 917; Santschi, 1924b: 220; Emery, 1924d: 297; Menozzi, 1933a: 109. Junior synonym of pygmaeus: Bolton, 1974a: 48. Junior synonym of traegaordhi: Bolton, 1982: 359; Bolton, 1995b: 139. umbilicatus. Cataulacus umbilicatus Santschi, 1937a: 59, figs. 10-12 (q.) MOZAMBIQUE. Type-material: holotype queen. Type-locality: Mozambique (“Est Africain Portugais”): Beira, vii-viii.1936 (M. Grobham). Type-depository: BMNH. Junior synonym of intrudens: Bolton, 1974a: 43; Bolton, 1995b: 139. voeltzkowi. Cataulacus voeltzkowi Forel, 1907g: 84 (w.) COMORO IS. Type-material: syntype workers. Type-locality: Comoro Is (“Comoren”): Moheli (Voeltzkow). Type-depositories: MHNG, MNHU. Status as species: Wheeler, W.M. 1922a: 1033; Emery, 1924d: 297; Bolton, 1974a: 50 (redescription); Bolton, 1995b: 139. Distribution: Madagascar. vorticus. Cataulacus vorticus Bolton, 1974a: 38, figs. 16, 19 (w.q.m.) NIGERIA, DEMOCRATIC REPUBLIC OF CONGO. Type-material: holotype worker, 3 paratype workers, 4 paratype queens, 4 paratype males. Type-locality: holotype Nigeria: Gambari (= Cocoa Research Institute of Nigeria), 30.viii.1969, under bark of cocoa tree (B. Bolton); paratypes: 2 workers, 4 queens, 3 males with same data; 1 worker, 1 male Democratic Republic of Congo (“Zaire”): 91 mi. W of Popokabaka2.viii. 1957 (E.S. Ross & R.E. Leech). Type-depositories: BMNH (holotype); BMNH, MCZC (paratypes). Status as species: Bolton, 1982: 356 (in key); Bolton, 1995b: 139; Hita Garcia, et al. 2013: 209. Distribution: Cameroon, Democratic Republic of Congo, Kenya, Nigeria. wasmanni. Cataulacus (Otomyrmex) wasmanni Forel, 1897c: 193 (w.) MADAGASCAR. Type-material: holotype worker. Type-locality: Madagascar: Isle Ste Marie, Kalolo (Perrot). Type-depository: unknown, perhaps in SMFM . [Otomyrmex wasmanni Wasmann, 1897: 250. Nomen nudum (attributed to Forel).] Status as species: Emery, 1899f: 286; Wheeler, W.M. 1922a: 1033; Emery, 1924d: 299; Bolton, 1974a: 29 (redescription); Bolton, 1995b: 139. Distribution: Madagascar. weissi. Cataulacus weissi Santschi, 1913c: 310 (w.) CONGO. Type-material: holotype worker. Type-locality: Congo (“Congo français”): Brazzaville, 1907 (A. Weiss). Type-depository: NHMB. Forel, 1916: 427 (q.m.). As unavailable (infrasubspecific) name: Wheeler, W.M. 1922a: 916; Emery, 1924d: 297. Subspecies of pygmaeus: Santschi, 1916b: 506 (footnote). Subspecies of lujae: Santschi, 1924b: 219. Status as species: Forel, 1916: 427; Bolton, 1974a: 39 (redescription); Bolton, 1982: 357 (in key), 358; Bolton, 1995b: 139. Senior synonym of aethiops: Bolton, 1982: 358; Bolton, 1995b: 139. Senior synonym of plectroniae: Bolton, 1974a: 39; Bolton, 1995b: 139. Distribution: Congo, Democratic Republic of Congo, Ghana, Ivory Coast. wissmannii. Cataulacus wissmannii Forel, 1894b: 78 (w.) MOZAMBIQUE. Type-material: holotype worker. Type-locality: Mozambique: (no further data), 9.xi.1890 (A. Müller). Type-depository: MHNG. Santschi, 1937a: 61 (q.). Status as species: Arnold, 1917: 401; Wheeler, W.M. 1922a: 917; Emery, 1924d: 297; Santschi, 1937a: 61; Bolton, 1974a: 51 (redescription); Bolton, 1982: 357 (in key); Bolton, 1995b: 139; Hita Garcia, et al. 2013: 209. Senior synonym of durbanensis: Bolton, 1974a: 51; Bolton, 1995b: 139. Senior synonym of linearis: Bolton, 1974a: 51; Bolton, 1995b: 139. Senior synonym of otii: Bolton, 1974a: 51; Bolton, 1995b: 139. Distribution: Kenya, Mozambique, South Africa.