Wild Orchids

Total Page:16

File Type:pdf, Size:1020Kb

Wild Orchids Wild Orchids by Bob Armstrong and Marge Hermans from Southeast Alaska's Natural World 161 161 A dance fly that probes People the world over have into a green bog orchid long been fascinated with orchids. Wealthy blossom will likely collectors have paid fortunes to possess emerge with clumps exotic varieties, and adventurers have risked of pollen stuck to tiny their lives to extract rare varieties from disks on its head. The jungles and remote locations. insect’s subsequent In 2001, U.S. sales of potted orchids visits to other flowers (artificially raised or hybrids) approached of the same species $100 million; and according to an item in help promote the U.S. News and World Report, a single potted exchange of pollen white stem of a phalaenopsis orchid at the between plants (cross- elegant Takashimaya Floral Boutique in pollination). New York was priced at $175. Nurseries closely or using a magnifying glass or hand and other outlets currently ship millions lens. of orchids all over the United States and Fairy slipper and lady’s slipper blos- Canada, and their spectacular varieties and soms display the quintessential orchid-type colors are often the highlight of garden blossom. Each flower has several upright shows and greenhouse displays. parts (petals and sepals), and a lower petal (Preceding page) Most people are attracted by the com- modified into a showy pouch that appar- Calypso bulbosa, plex and elegant blossoms of orchids, and ently serves as a landing pad for pollinating “fairy slipper” tropical species especially show a stagger- insects. Insects are attracted to the blossoms ing variety of colors, petal shapes, and intricate mark- ings. A few wild orchids in Southeast Alaska have blossoms large and color- ful enough to remind us (Right) White bog of their more spectacular orchids may grow three tropical relatives. Those are feet high in Southeast the beautiful Calypso bulbosa, Alaska, often rising or “fairy slipper,” found above surrounding mostly on small islands of plants in a muskeg Southeast Alaska, often on and emitting a strong, the outer coast; and three pleasant aroma similar species of “lady’s slip- to that of vanilla. pers” occasionally found in the northern parts of our region. But a number of other wild orchids are quite com- mon in Southeast forests and bogs, and it’s easy to discover and appreciate their lovely, delicate blos- soms by just looking more 162 by their color and fragrance, then when they shoot out a drop of viscous fluid filled with probe the blossoms, they emerge carrying pollen grains. The fluid glues the pollen mass (Bottom) Rattlesnake pollen that they subsequently deposit on to the insect’s body so it, too, can be carried plantain is named for other plants. to subsequent blossoms. the variable striations on its leaves, which Some orchids go to great lengths to If pollination is successful, an orchid often resmemble trick insects into helping with the process may produce for each blossom a single oval the markings of of pollination. They have developed blos- or oblong capsule filled with tiny seeds. A rattlesnake skin. Its soms that mimic both the appearance and capsule may contain an enormous num- blossom, shown above the chemical sex signals, or pheromones, ber of seeds—as many as 3,770,000 were the photo of the leaves, of female insects. The blossoms of Chilo- counted in a single capsule of a tropical displays delicate petals glottis tropeziformis, an orchid not found in American orchid, Cynoches chlorochilon, for on a central spike. Southeast, looks and “smells” so much like example. And it’s a good thing, too. As A.F. a particular species of female wasp that Szczawinski writes in The Orchids of British males of the species are attracted to the Columbia, “Unless the conditions of tem- orchid flowers from long distances. And the perature, moisture, soil and shelter are ideal, orchid Coryanthes speciosa, one of the species failure for the seedlings is certain. There known as bucket orchids found in lowland is one chance in thousands that such ideal forests in Central and South America, lures conditions will occur.” male euglossine bees into its heart, coating Once orchid seeds land on the ground, them with both pollen and a waxy perfume they do not germinate as easily as the seeds that appears to be useful in attracting female of most plants. Their outer seed-coat must bees. be penetrated by the microscopic threads Here in Southeast Alaska, white bog of certain fungi species in the soil before orchids, which often grow in large patches they can germinate. Apparently the fungi in marshes and muskegs, emit a strong fra- are needed to convert starches in the seed grance that some describe as a combination into simpler sugars that the embryo orchid of vanilla, cloves, and other spices. They can use. seem to attract certain insects, such as mos- Whether or not pollination is successful, quitoes, dance flies, and moths, which help it may be years before a particular orchid transport their pollen from one flower to will bloom again. For this reason, as well another. The nectar in bog orchids is housed in a narrow pouch or “spur” that projects backward from the lip of the blossom, and it can only be reached by an insect with a long proboscis, or tubular mouth part. When the insect sticks its head into the blossom it bumps against and tears off tiny adhesive discs that stick to its forehead and proboscis, and pick up clusters of pollen that can then be deposited in another orchid blossom. Heart-leaved twayblades—tiny or- chids that are fairly common in moist or wet forests in our region—use a different approach. They give off a strong odor that attracts flies and fungus gnats. When they are touched by one of these insects, they 163 as their fragility and their In the case of the bog adder’s tongue dependence on associating (Malaxis paludosa), found in bogs in South- with fungi growing in the east Alaska, small projections known as soil around them, wild or- “bulbils” grow at the tips of the leaves and chids in many parts of the break off to form new plants. U.S. and the world have Two species of orchids seen fairly often become rare, and many in the deep shade of Southeast coniferous states have laws prohibit- forests may not at first be recognized as ing picking them. orchids. Completely pink or yellowish Some orchids also and semi-transparent, the coralroots are reproduce by vegetative saprophytic plants—they live off decaying multiplication. This is true organic matter and contain no chlorophyll of the rattlesnake plan- for making their own food. Coralroot flow- tain, found in primary ers nonetheless show orchid-like character- glacial forests and dense istics—their many-petaled blossoms have a forests along the beach characteristic protruding lip, though they are fringe in Southeast. This small and clustered along an upright spike. Spotted coralroot is a intriguing plant, with rosettes of mottled or Authoritative field guides list 21 spe- a saprophytic orchid striped evergreen leaves and delicate flowers cies of wild orchids found in Southeast. that lives off decaying on a slender spike, can spread very rapidly, To learn more about them we recommend organic matter in the “creeping” along under the ground by ex- Native Plants of Southeast Alaska by Judy soil. Small clusters are tending its underground stem, or rhizome. Hall, and Plants of the Pacific Northwest Coast often seen in the deep edited by Pojar and MacKinnon. shade of coniferous forests in Southeast Alaska. Elusive Promises Many people’s inor- advertised to “stimulate the sexual dinate fascination with propensities.” Calypso orchids, or “orchidelerium,” Women in Victorian England were bulbosa, “fairy over the centuries prob- forbidden to own orchids, whose name ably stems from myths and slipper” comes from the Latin orchis, meaning traditions in a number of testicle and suggesting sexual associations. cultures. The Aztecs and Even today the scent and flavor of vanilla Mayas of Middle America is widely marketed for its sensuous and used vanilla, made from the evocative qualities in soap and perfumes. seed capsules of orchids of the Vanilla The “fairy slipper” Calypso bulbosa genus, as an erotic drink; and once the is named for the beautiful goddess who Spanish carried vanilla flavoring to Europe ensnared Homer’s Ulysses for seven years in the 1700s, it was touted as a tincture on the island of Ogygia. The mountain or infusion to insure male potency. After lady-slipper, occasionally found in northern Thomas Jefferson brought vanilla to Southeast Alaska, is named Cypripedium, the United States from France, it was which means “the foot of Aphrodite,” the enthusiastically accepted, and occasionally ancient Greek goddess of love. 164.
Recommended publications
  • Generic and Subtribal Relationships in Neotropical Cymbidieae (Orchidaceae) Based on Matk/Ycf1 Plastid Data
    LANKESTERIANA 13(3): 375—392. 2014. I N V I T E D P A P E R* GENERIC AND SUBTRIBAL RELATIONSHIPS IN NEOTROPICAL CYMBIDIEAE (ORCHIDACEAE) BASED ON MATK/YCF1 PLASTID DATA W. MARK WHITTEN1,2, KURT M. NEUBIG1 & N. H. WILLIAMS1 1Florida Museum of Natural History, University of Florida Gainesville, FL 32611-7800 USA 2Corresponding author: [email protected] ABSTRACT. Relationships among all subtribes of Neotropical Cymbidieae (Orchidaceae) were estimated using combined matK/ycf1 plastid sequence data for 289 taxa. The matrix was analyzed using RAxML. Bootstrap (BS) analyses yield 100% BS support for all subtribes except Stanhopeinae (87%). Generic relationships within subtribes are highly resolved and are generally congruent with those presented in previous studies and as summarized in Genera Orchidacearum. Relationships among subtribes are largely unresolved. The Szlachetko generic classification of Maxillariinae is not supported. A new combination is made for Maxillaria cacaoensis J.T.Atwood in Camaridium. KEY WORDS: Orchidaceae, Cymbidieae, Maxillariinae, matK, ycf1, phylogenetics, Camaridium, Maxillaria cacaoensis, Vargasiella Cymbidieae include many of the showiest align nrITS sequences across the entire tribe was Neotropical epiphytic orchids and an unparalleled unrealistic due to high levels of sequence divergence, diversity in floral rewards and pollination systems. and instead to concentrate our efforts on assembling Many researchers have posed questions such as a larger plastid data set based on two regions (matK “How many times and when has male euglossine and ycf1) that are among the most variable plastid bee pollination evolved?”(Ramírez et al. 2011), or exon regions and can be aligned with minimal “How many times have oil-reward flowers evolved?” ambiguity across broad taxonomic spans.
    [Show full text]
  • E29695d2fc942b3642b5dc68ca
    ISSN 1409-3871 VOL. 9, No. 1—2 AUGUST 2009 Orchids and orchidology in Central America: 500 years of history CARLOS OSSENBACH INTERNATIONAL JOURNAL ON ORCHIDOLOGY LANKESTERIANA INTERNATIONAL JOURNAL ON ORCHIDOLOGY Copyright © 2009 Lankester Botanical Garden, University of Costa Rica Effective publication date: August 30, 2009 Layout: Jardín Botánico Lankester. Cover: Chichiltic tepetlauxochitl (Laelia speciosa), from Francisco Hernández, Rerum Medicarum Novae Hispaniae Thesaurus, Rome, Jacobus Mascardus, 1628. Printer: Litografía Ediciones Sanabria S.A. Printed copies: 500 Printed in Costa Rica / Impreso en Costa Rica R Lankesteriana / International Journal on Orchidology No. 1 (2001)-- . -- San José, Costa Rica: Editorial Universidad de Costa Rica, 2001-- v. ISSN-1409-3871 1. Botánica - Publicaciones periódicas, 2. Publicaciones periódicas costarricenses LANKESTERIANA i TABLE OF CONTENTS Introduction 1 Geographical and historical scope of this study 1 Political history of Central America 3 Central America: biodiversity and phytogeography 7 Orchids in the prehispanic period 10 The area of influence of the Chibcha culture 10 The northern region of Central America before the Spanish conquest 11 Orchids in the cultures of Mayas and Aztecs 15 The history of Vanilla 16 From the Codex Badianus to Carl von Linné 26 The Codex Badianus 26 The expedition of Francisco Hernández to New Spain (1570-1577) 26 A new dark age 28 The “English American” — the journey through Mexico and Central America of Thomas Gage (1625-1637) 31 The renaissance of science
    [Show full text]
  • List of Orchid Plantsfor Sale, April 2018
    List of Orchid Plants for Sale, April 2018 INSTRUCTIONS: 1. Enter the desired Quantity of Plants in the Column "Q". The "Total" column will update automatically. 2. Type your personal information in the cases below this list. Fill in the light green cases only. 3. Send your order to: [email protected] 1. SECTION: ORCHID SPECIES & HYBRIDS Climate Name Q US$ Total Cool Intermediate Acianthera casapensis 0 12 0 Cool Intermediate Acineta superba 0 22 0 Intermediate Ada brachypus 0 20 0 Intermediate Ada elegantula 0 20 0 Intermediate Ada euodes (Ada elegantula) 0 22 0 Intermediate Ada rolandoi 0 20 0 Intermediate Anguloa clowesii 0 28 0 Intermediate Anguloa eburnea 0 24 0 Intermediate Anguloa uniflora 0 28 0 Intermediate Anguloa virginalis 0 24 0 Cool Barbosella cucullata 0 16 0 Cool Barbosella prorepens 0 16 0 Warm Batemannia colleyi 0 20 0 Intermediate Bletia campanulata 0 16 0 Warm Intermediate Bletia catenulata 0 16 0 Warm Intermediate Bletia catenulata var. Coerulea 0 120 0 Warm Brassavola tuberculata (Brassavola ovaliformis) 0 20 0 Intermediate Brassia cauliformis 0 14 0 Intermediate Brassia cochleata (Brassia angusta) 0 14 0 Warm Intermediate Brassia longissima 0 16 0 Intermediate Brassia pascoensis 0 18 0 Intermediate Brassia villosa 0 14 0 Warm Intermediate Brassia wageneri 0 16 0 Cool Intermediate Bulbophyllum (equinolabium x carunculatum) 0 24 0 Warm Intermediate Bulbophyllum meridense 0 18 0 Cool Bulbophyllum steyermarkii 0 18 0 Intermediate Bulbophyllum weberbauerianum 0 18 0 Warm Catasetum discolor 0 24 0 Warm Catasetum expansum
    [Show full text]
  • A Phylogenetic Assessment of Lycaste and Anguloa (Orchidaceae)
    A PHYLOGENETIC ASSESSMENT OF LYCASTE AMD ANGULOA (ORCHIDACEAE) By ANGELA RYAN A thesis submitted for the degree of Doctor of Philosophy in the University of London DEPARTMENT OF CHEMISTRY UNIVERSITY COLLEGE LONDON 2001 ProQuest Number: U642610 All rights reserved INFORMATION TO ALL USERS The quality of this reproduction is dependent upon the quality of the copy submitted. In the unlikely event that the author did not send a complete manuscript and there are missing pages, these will be noted. Also, if material had to be removed, a note will indicate the deletion. uest. ProQuest U642610 Published by ProQuest LLC(2015). Copyright of the Dissertation is held by the Author. All rights reserved. This work is protected against unauthorized copying under Title 17, United States Code. Microform Edition © ProQuest LLC. ProQuest LLC 789 East Eisenhower Parkway P.O. Box 1346 Ann Arbor, Ml 48106-1346 ABSTRACT Parsimony analysis has been used to examine the phylogenetic relationships of two genera of Neotropical orchids, Lycaste and Angulos. Within these genera, difficulties occur when assigning names to plants using traditional morphological techniques. Many herbarium specimens are in bad condition and some descriptions are incomplete. To date, infrageneric classifications have been based on very few diagnostic characters. Here, three approaches have been evaluated: A systematic analysis of the morphology, an examination of two regions of DMA sequence and an analysis of the chemical composition of the floral fragrances. Morphological analysis showed that Lycaste is not monophyletic. There was a clear division between species currently ascribed to sect. Fimbriatae and the other species of the genus.
    [Show full text]
  • Sede Amministrativa: Università Degli Studi Di Padova Dipartimento Di Filosofia
    Sede Amministrativa: Università degli Studi di Padova Dipartimento di filosofia ___________________________________________________________________ CORSO DI DOTTORATO DI RICERCA IN FILOSOFIA CICLO XXXI TITOLO TESI DARWIN’S CONTRIVANCES: ORCHIDS, EVOLUTION AND SCIENTIFIC ETHICS IN THE PHILOSOPHY OF SCIENCE OF XIX CENTURY Coordinatore: Ch.mo Prof. Francesca Menegoni Supervisore: Ch.mo Prof. Fabio Grigenti Dottorando : Antonio Danese INDEX 0 INTRODUCTION 5 0.1 Topic 5 0.2 Why Orchids? 9 0.3 Structure of the work 12 1 THE DARWINIAN THEORY 14 1.1 The origin of the origin of species 14 1.2 The first concept of species 16 1.3 The renouncement of immutability and the problem of classification 18 1.4 Variation 23 1.5 The ecological context and the tree of life 26 1.6 New way of observing nature 28 2 FORERUNNERS OF DARWIN IN THE BOTANICAL FIELD 29 2.1 Cell and transmutation 31 2.2 The molecular basis for the development of life sciences 31 2.3 The starting points for Darwinian botany 33 3 THE WORK ON PLANT SCIENCES 35 3.1 The botanist Darwin 36 3.2 The movements of climbing plants 40 3.3 The insectivorous plants 43 3.4 Cross-self fertilisation 45 3.5 The different forms of flowers on plants of the same species 46 3.6 The power of movement in plants 49 4 ORCHIDS IN VICTORIAN AGE 52 4.1 Classification of new species of orchids 53 4.2 Linnaeus 54 4.3 Orchids, society, and literature 56 4.4 Darwin and orchids 59 5 THE ORIGIN OF On the Various Contrivances by Which British and Foreign Orchids Are Fertilised by Insects and on the Good Effects of Intercrossing
    [Show full text]
  • A Critical Examination of Some of the Principal Arguments for and Against
    118 CHAPTER IV . Darwin on the fertilization of orchids — Some orchids fertilized by particular insects only – Case of the Angræcum sesquipe dale and of the Coryanthes Macrantha - Abundance of or chid seed and scarcity of seedling plants — Duke of Argyle ' s Reign of Law - Admits evolution of species under a guid ing will — The organs of reproduction of orchids a proof of this - Mr Wallace ' s observation upon this point - Hum ming - birds - Mr Wallace ' s observations upon the Reign of of Law , The Duke of Argyle ' s reply - The peculiarity of organs of reproduction of orchids not the result of natural selection - Owen ' s comparative anatomy - Fossil ancestors of the horse - Bucephalus - Extinct corals , Owen ' s views as to derivation - Mr Darwin ' s animals and plants under domestication - As to the stability of variations - Cross bred rats — Different degrees of variability - In what sense variations are accidental - Reference to fragments fallen from a cliff - Asa Gray ' s point that variation has been divinely directed . We come now to Mr Darwin ' s work on the fertilization of Orchids , his object evidently being to show that the extraordinary struc ture of the flowers of these plants was bene ficial to them , and thus to meet to some extent the objections made to the theory of natural selection , from the fact that there CHAP . IV . MODE OF FERTILIZATION . 149 are in nature many structures which have been created for the sake of beauty and variety . The pollen masses of almost all orchideous plants are enclosed in sacks in such a manner that the pollen cannot be shaken on to the stigma , nor the flowers fertilized by insects in the usual manner .
    [Show full text]
  • Stanhopeinae Mesoamericanae I
    LANKESTERIANA 8: 23-30. 2003. STANHOPEINAE MESOAMERICANAE I GÜNTER GERLACH1 y ROBERT L. DRESSLER2 1Botanischer Garten München-Nymphenburg Menzinger Str. 65, 80638 München, Germany. [email protected] 2Missouri Botanical Garden; Florida Museum of Natural History; Marie Selby Botanical Gardens Direccion para correo: 21305 NW 86th Ave., Micanopy, Florida 32667, U.S.A. [email protected] AB S T R A C T. Two new species of the subtribe Stanhopeinae tribe are described, illustrated and discussed: Coryanthes kaiseriana, from Costa Rica and Panama, and Paphinia vermiculifera from Panama. RESUMEN. Se describen, ilustran y comentan dos nuevas especies de la subtribu Stanhopeinae: Coryanthes kaiseriana, de Costa Rica y Panamá, y Paphinia vermiculifera de Panamá. PALABRAS CLAVE / KEY WORDS: Orchidaceae, Stanhopeinae, Coryanthes kaiseriana, Paphinia vermiculifera, Mesoamérica La investigación intensa y metódica de la subtribu por el espécimen tipo. De este modo, fue posible Stanhopeinae para el tratamiento de la F l o r a observar y entender el espectro de la variabilidad de M e s o a m e r i c a n a hace necesario aclarar algunos con- los táxones mesoamericanos de Stanhopeinae. A con- ceptos para entender diferentes especies. El estudio tinuación se describen, ilustran y comentan dos de tipos en el marco de este trabajo culminará en especies nuevas de esta subtribu: Coryanthes kaiseri - descripciones, combinaciones nuevas y nombres ana, de Costa Rica y Panamá, y Paphinia vermiculif - nuevos de plantas ya conocidas. Con este artículo era, de Panamá. empieza una serie de publicaciones con la meta de establecer una base firme para investigar la biología Coryanthes kaiseriana G.
    [Show full text]
  • 2009 Orchid List
    2009 ORCHID LIST IMPORTANT ANNOUNCEMENT! Dear Friends, Here is our new 2009 List of new hard-to-find, interesting, unusual, and proven species. Following through with our promise to gradually move from print to the Internet, we no longer will print and mail color photos. Producing those beautiful color pages has been a lot of fun, but it is time to move on. There are only a few us working and we need to put all of our time and energy towards what we do best; producing more new items and introducing them to the market faster. We are putting a lot of energy into new photos for our website, though, and 2009 will have the most new photos posted for any year since we developed the website. If you must have some color printed material, for a limited time we can mail you a collection of some of our best color pages for $5.00. These color pages are already “collector’s items”. Also, some of you may not be aware of Jay Pfahl’s excellent website; the Internet Orchid Species Photo Encyclopedia – a priceless resource, you can see photos of thousands of orchid species there. This List is only a partial listing of all the species that will become available this year. Please check our website frequently to see all the latest species as they are added. The website also has “special offers”. Our lab operation is looking great - and there are hundreds of exciting species yet to be offered. You can help us save time by ordering on-line.
    [Show full text]
  • Lankesteriana International Journal on Orchidology ISSN: 1409-3871 [email protected] Universidad De Costa Rica Costa Rica
    Lankesteriana International Journal on Orchidology ISSN: 1409-3871 [email protected] Universidad de Costa Rica Costa Rica Cribb, Phillip FOREWORD Lankesteriana International Journal on Orchidology, vol. 10, núm. 2-3, diciembre, 2010, pp. 1-215 Universidad de Costa Rica Cartago, Costa Rica Available in: http://www.redalyc.org/articulo.oa?id=44340990001 How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative LANKESTERIANA 10(2-3): i. 2010. FOREWORD Paradise for an orchid collector is a trail that runs through rich orchid habitat. Preferably the trail should decrease in elevation from 3000 to 500 meters over a protracted distance, it should be in a high annual rainfall area with the rain distributed evenly throughout the year, it also should be in a region of extremely high biodiversity and very pronounced local endemism. The adjoining forests, cliffs and embankments would be festooned with the natural epiphytes and terrestrials of the zone. In the Western Hemisphere, prior to the development of roads and highways, such trails from the lowlands to the Andean highlands existed from northwestern Colombia to southern Ecuador and northern Peru and provided the means of communication for people traveling by foot or mule back. Each of those trails might have had more than 1,000 orchid species distributed along their length. Curiously, each trail may have had a very different species composition from the next closest trail.
    [Show full text]
  • A Comparison and Combination of Plastid Atpb and Rbcl Gene Sequences for Inferring Phylogenetic Relationships Within Orchidaceae Kenneth M
    Aliso: A Journal of Systematic and Evolutionary Botany Volume 22 | Issue 1 Article 36 2006 A Comparison and Combination of Plastid atpB and rbcL Gene Sequences for Inferring Phylogenetic Relationships within Orchidaceae Kenneth M. Cameron New York Botanical Garden Follow this and additional works at: http://scholarship.claremont.edu/aliso Part of the Botany Commons Recommended Citation Cameron, Kenneth M. (2006) "A Comparison and Combination of Plastid atpB and rbcL Gene Sequences for Inferring Phylogenetic Relationships within Orchidaceae," Aliso: A Journal of Systematic and Evolutionary Botany: Vol. 22: Iss. 1, Article 36. Available at: http://scholarship.claremont.edu/aliso/vol22/iss1/36 MONOCOTS Comparative Biology and Evolution Excluding Poales Aliso 22, pp. 447-464 © 2006, Rancho Santa Ana Botanic Garden A COMPARISON AND COMBINATION OF PLASTID atpB AND rbcL GENE SEQUENCES FOR INFERRING PHYLOGENETIC RELATIONSHIPS WITHIN ORCHIDACEAE KENNETH M. CAMERON The Lewis B. and Dorothy Cullman Program for Molecular Systematics Studies, The New York Botanical Garden, Bronx, New York 10458-5126, USA ([email protected]) ABSTRACT Parsimony analyses of DNA sequences from the plastid genes atpB and rbcL were completed for 173 species of Orchidaceae (representing 150 different genera) and nine genera from outgroup families in Asparagales. The atpB tree topology is similar to the rbcL tree, although the atpB data contain less homoplasy and provide greater jackknife support than rbcL alone. In combination, the two-gene tree recovers five monophyletic clades corresponding to subfamilies within Orchidaceae, and fully resolves them with moderate to high jackknife support as follows: Epidendroideae are sister to Orchidoideae, followed by Cypripedioideae, then Vanilloideae, and with Apostasioideae sister to the entire family.
    [Show full text]
  • Classification, Evolution, and Phylogeny of the Families of Monocotyledons
    SMITHSONIAN CONTRIBUTIONS TO BOTANY NUMBER 71 Classification, Evolution, and Phylogeny of the Families of Monocotyledons Aaron Goldberg SMITHSONUN INSTITUTION PRESS Washington, D.C. 1989 ABSTRACT Goldberg, Aaron. Classification, Evolution, and Phylogeny of the Families of Monocotyle- dons. Smithsonian Contributions to Botany, number 71, 74 pages, 41 figures, 2 tables, 1 diagram, 1989.-To some extent classification is subjective. Taxonomists differ in the relative importance they ascribe to particular characters and in the degree of difference between related taxa they deem sufficient to constitute family or ordinal rank. About 250 monocot family names have been published. Those who have attempted an overview of the system at the family level and above in the last quarter century recognize between 45 and 103 monocot families in 14 to 38 orders. I accept 57 families in 18 orders. In Table 1 I give my ordinal allocation of the families and that of 11 recent authors to indicate where there is agreement and where there are differences to be resolved. I have constructed a dendrogram to suggest relationships and degree of advancement of the orders. I have written concise, uniform descriptions of all the families of monocots emphasizing those characters that show trends between families or occur in more than one family. Each family is illustrated by analytical drawings of the flower, fruit, seed, and usually inflorescence. Several species are usually used to show the range of major variation within families and trends toward related families. Monocots and dicots have existed concurrently for most of their history, have been subjected to many of the same ecological pressures, and consequently show similar evolutionary trends.
    [Show full text]
  • December 2017. Orchid Specialist Group Newsletter
    ORCHID CONSERVATION NEWS The Newsletter of the Orchid Specialist Group of the IUCN Species Survival Commission Issue 3 December 2017 CONSERVATION AND THE DYNAMICS OF CHANGE A tranquil scene before a storm 1 negatively impacted by loss of their particular Editorial pollinator(s). Change is inevitable, catastrophic weather-related events thankfully sporadic, but storm trajectories and outcomes are as unpredictable for human populations and their infrastructure as for orchids, their habitats, and for associated biodiversity. This year has seen exceptionally destructive hurricanes damaging Caribbean islands and Puerto Rico as well as parts of the south and southeastern US. Two super typhoons struck China. There have been devastating floods, fires, earthquakes and recent volcanic activity. Anthropogenic-driven disturbances may alter biodiversity dynamics and impact conservation of orchids but when change happens, whatever the cause or event, opportunities to analyse the impact and track recovery often emerge. Vale et al. (2013) have examined habitat disturbance including the impact of hurricanes and of overcollection as challenges to the conservation of the endangered endemic Broughtonia cubensis in Cuba. Following flooding after the construction of the Balbina dam in Coryanthes macrantha is the most widely distributed of the genus and is pollinated by Eulaema spp. Brazil, Storck-Tonon & Peres (2017) used the resulting archipelago of 3546 variously sized and separated Robert Pemberton has studied a naturalized orchid bee, islands to learn how an assemblage of euglossine orchid Euglossa delimma (formerly known as E. viridissimma) bees responded to habitat loss and varying degrees of and has shared some of his observations and related island isolation. We may not be able to study locations publications.
    [Show full text]