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Catalogue of Latvian (Insecta: Coleoptera)

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Zoology and Ecology Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/tzec20 Catalogue of Latvian Dermestidae (Insecta: Coleoptera) a b c Andris Bukejs , Dmitry Telnov & Jiří Háva a Vienības iela 42-29, LV-5401 , Daugavpils , Latvia b Stopiņu novads , Dārza iela 10, LV-2130 , Dzidriņas , Latvia c Faculty of Forestry and Wood Sciences, Department of Forest Protection and Entomology , Czech University of Life Sciences , Kamýcká 1176, CZ-165 21 , Prague 6 – Suchdol , Published online: 05 Sep 2013.

To cite this article: Andris Bukejs , Dmitry Telnov & Jiří Háva (2013) Catalogue of Latvian Dermestidae (Insecta: Coleoptera), Zoology and Ecology, 23:3, 224-232, DOI: 10.1080/21658005.2013.830408 To link to this article: http://dx.doi.org/10.1080/21658005.2013.830408

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Catalogue of Latvian Dermestidae (Insecta: Coleoptera) Andris Bukejsa*, Dmitry Telnovb and Jiří Hávac aVienības iela 42-29, LV-5401, Daugavpils, Latvia; bStopiņu novads, Dārza iela 10, LV-2130, Dzidriņas, Latvia; cFaculty of Forestry and Wood Sciences, Department of Forest Protection and Entomology, Czech University of Life Sciences, Kamýcká 1176, CZ-165 21, Prague 6 – Suchdol, Czech Republic (Received 3 June 2013; accepted 21 July 2013)

A critically annotated catalogue of Dermestidae species recorded in Latvia is presented. In total, 33 species belonging to nine genera and four subfamilies are confirmed for Latvia. However, some of them, i.e. ater DeGeer, D. erichsoni Ganglbauer, D. undulatus (Brahm), Trinodes hirtus (Fabricius), and Globicornis emarginata (Gyllenhal), are known only from old records, and new material is needed to confirm their occurrence in the local fauna. One species, Megatoma graeseri (Reitter, 1887) is recorded in Latvia for the first time. A complete bibliography of Latvian skin is reviewed and analysed. The research history of Latvian Dermestidae is discussed. The Latvian fauna of skin beetles is represented by nine chorotypes. Cosmopolitan (13 or 39.4%) species are predominant in dermestid fauna of Latvia.

Išanalizavus bibliografiją ir tyrimų istoriją sudarytas naujas anotuotas Latvijos kailiavabalių rūšių katalogas. Patvirtinta, kad Latvijos faunoje iš viso yra 33 šių vabalų rūšys, priklausančios 9 gentims ir 4 pošeimiams. Apie Dermestes ater DeGeer, D. erichsoni Ganglbauer, D. undulatus (Brahm), Trinodes hirtus (Fabricius) ir Globicornis emarginata (Gyllenhal) priklausymą Latvijos faunai žinoma tik iš senųšaltinių.Būtina nauja medžiaga šiai informacijai patvirtinti. Megatoma graeseri (Reitter, 1887) Latvijoje užregistruotas pirmą kartą. Latvijos faunoje yra skiriami 9 kailiavabalių chorotipai; dominuoja kosmopolitinės (13, t.y. 39,4%) rūšys. Keywords: Coleoptera; Dermestidae; Latvia; fauna; catalogue; biogeography

Introduction The first information on skin beetles from the present The family Dermestidae is divided into six subfamilies territory of Latvia was published in the second half of the (Lawrence and Slipinski 2005; Kiselyova and McHugh eighteenth century by Fischer (1778, 1784, 1791). In the fi 2006; Bouchard et al. 2011) with 58 genera and approxi- rst edition of his monograph, which introduced the natu- – mately 1460 species and subspecies worldwide (Háva ral history of Livland (Fischer 1778), two species Der- – 2003). About 590 species and subspecies are known in mestes lardarius L. and D. murinus L. were mentioned. the Palaearctic (Háva 2007). The family, like most of its After some additions (Fischer 1784) in the second edition subfamilies, has a worldwide distribution. One fossil of his monograph, Fischer (1791) already mentioned four , five genera, and more than 30 species are presently species, namely , D. murinus L., known (Háva and Prokop 2004; Háva and Bukejs 2012; Attagenus pellio (L.) [as Dermestes], Anthrenus scrophu-

Downloaded by [Andris Bukejs] at 01:33 15 September 2013 Háva 2013; Ponomarenko and Kirejtshuk 2013). lariae (L.) [as Byrrchus]. Skin beetles are most commonly found on flowering The professor of Jelgava (former Mitau) gymnasium plants and dried carcasses and are also nidicolous Johan Groschke published a list of household pests. In the “ ” in nests of vertebrates and social Hymenoptera. monograph Beschreibungen der Provinz Kurland Dermestids are also found in human dwellings. Most dedicated to Kurzeme [Curland], he reported three species species feed almost exclusively on materials of animal of Dermestidae (Groschke 1805): Dermestes lardarius L., origin, such as bones, skin, fur, wool, silk, etc. However, D. murinus L., and Attagenus pellio (L.) [as Dermestes]. fi some species feed partly or entirely on vegetable matter. At the beginning of the nineteenth century, the rst Some species specialize in plant material (Zhantiev 1974, checklist of Coleoptera of the Latvian fauna was 1976; Bousquet 1990; Toskina 2007). Many species are published without indication of authorship. It included serious pests and can damage stored food and items of 725 species of Coleoptera from a large territory includ- organic origin, including museum collections (especially ing most of the present-day Latvia. According to Kawall ī zoological). A few species are predatory, Thorictinae (1868), the author was the resident of R ga Precht species are myrmecophilous. (1818). In this catalogue, 11 species are listed as skin To date, 33 species of Dermestidae are confirmed for beetles. According to contemporary systematics, only the Latvian fauna. Until now, 51 works containing nine species belonged to Dermestidae. Two taxa (listed information on skin beetles in Latvia have been published. as Dermestes pedicularis and Dermestes fumatus) are

*Corresponding author. Email: [email protected]

Ó 2013 Nature Research Centre Zoology and Ecology 225

currently representatives of Kateretidae and Byturidae or Fragmentary faunal data on Latvian skin beetles were Mycetophagidae, respectively. also published by Barševskis (1988, 1989, 1991, 1996, In 1829, the resident of Jelgava J. G. Fleischer 1997a, 2001), Barševskis et al. (2004); Barševskis, published another revision of beetles for the Baltic area. Bukejs, and Anichtchenko (2008), Bukejs (2006), Danka In this revision, 640 species of Coleoptera are and Stiprais (1972), Leiskina (2000), Telnov (1996, mentioned, which, according to the author, were 2001), Telnov et al. (2005, 2006, 2007, 2008, 2010, recorded for the fauna of “Ostseeprovinzen” (the Baltic 2011). Sea provinces) for the first time (Fleischer 1829). In 1997, the first catalogue of Latvian Coleoptera Among them, there were five skin species, was published (Telnov et al. 1997), containing 27 species namely Dermestes ater Deg [as Dermestes domesticus of the family Dermestidae. In the second edition of this Gebl.], D. laniarius Ill., D. affinis Gyll. [according to catalogue (Telnov 2004), this number had slightly the modern the last two taxa are conspecific], increased (28 species); however, Dermestes mustelinus Attagenus schaefferi (Hrbst.) and A. unicolor (Brahm) Erichson was removed from the list. [as Attagenus megatoma F.]. In the publication specifically dedicated to Latvian Kawall (1865, 1866) mentioned three species and Dermestidae (Bukejs and Barševskis 2007), faunal data one species of Dermestidae from Kurzeme [Curland]. on 21 species and a checklist including 32 species were At the end of the nineteenth century, Seidlitz’s mono- presented. graph “Fauna Baltica” was published (Seidlitz 1872–75, Ulanowski (1883) published faunal data on 12 1887–91). In the monograph, keys to determination of species of Dermestidae, namely Dermestes lardarius L., families, genera and species of the Baltic Coleoptera D. frischii Kugelann, D. laniarius Ill., D. murinus L., (including potential ones) were given. Unfortunately, D. undulatus (Brahm), Attagenus pellio (L.), A. unicolor species sampling localities are rarely mentioned. In both (Brahm) [as A. piceus Ol.], Anthrenus scrophulariae (L.), editions of this monograph, 16 species of Dermestidae A. museorum (L.), serra (F.) [as Tiresias], were mentioned. Only in the second edition, one Megatoma undata (L.) and Trogoderma versicolor species was mentioned twice under different names, (Creutzer). All records are from the eastern part of Ill. and D. affinis Gyll. (according to Latvia. However, many of these records are considered to modern taxonomy, D. affinis is a junior synonym of be doubtful (see also Barševskis 1993). Consequently, in D. laniarius). the current catalogue, data from Ulanowski (1883) are At the beginning of the twentieth century, another not used as well as in other accounts of Latvian beetles checklist of Baltic beetles was published (Rathlef 1905). (Spuris 1981, 1983, 1984, 1990, 1991a, 1991b; In the list and its supplement (Rathlef 1921), 17 species Barševskis 2003; Barševskis, Kalniņš, and Cibuļskis of Dermestidae were included. 2005; Bukejs 2013). A few other papers containing faunal data on Latvian The aim of the current work is to critically Dermestidae were published in the first half of the summarize previous bibliographical data to compile the twentieth century (Heyden 1903; Mikutowicz 1905; annotated catalogue of Latvian Dermestidae according to Lackschewitz and Mikutowicz 1939). modern systematics and to provide biogeographical Nine skin beetle species, namely Dermestes analysis of the Latvian fauna of skin beetles. lardarius L., D. lardarius L., D. frischii Kugelann, Attagenus pellio (L.), A. unicolor (Brahm) [as A. pieceus Downloaded by [Andris Bukejs] at 01:33 15 September 2013 Ol.], Anthrenus scrophulariae (L.), A. caucasicus Rtt., Methods and conventions A. museorum (L.) and Megatoma undata (L.), were Taxa listed in this catalogue are arranged in subfamilies recorded as pests from Latvian storehouses (Vītols and tribes. The order of higher taxa follows the 1941). Later, some skin beetle species were also Catalogue of Palaearctic Coleoptera, volume 4, chapter mentioned as pests by Danka (1950). Dermestidae (Háva 2007) and Bouchard et al. (2011), Additional information on Latvian Dermestidae can synonymy and nomenclature for the species level were be found in Šmits and Spuris (1966) and Spuris (1974) used after Háva (2007, 2011). For the same reason, as well as in Barševskis and Savenkovs (1991), Priedītis species and genera are arranged alphabetically. (1979), Rūtenberga (1980), Spuņģis (2008), and Species recorded from Latvia for certain are marked Trauberga (1957). with an index number, species erroneously mentioned for Eighteen skin beetle species are mentioned and the Latvian fauna are marked with (–) as excluded. faunal data from the eastern part of Latvia are reported Synonyms and references to bibliographic sources follow by Barševskis (1993). Later, Barševskis (1997b) reported scientific names of species. 16 skin beetle species, including the first Latvian record The general distribution of species was analysed of Megatoma pubescens (Zetterstedt) from the according to Háva (2003, 2007, 2013), Lafer (1992) and Augšdaugava region (Eastern Latvia). Zhantiev (1976). The classification of chorotypes follows Ten species of Dermestidae are reported from Silene the one suggested by Taglianti et al. (1999). Nature Park (Barševskis 2002) and 15 species from Abbreviations of chorotypes are as follows: COS – Cos- Gauja National Park (Kalniņš et al. 2007). mopolitan, OLA – Holarctic, SCO – Subcosmopolitan, 226 A. Bukejs et al.

PAL – Palearctic, ASE – Asiatic-European, WPA – Note: Actual faunal data on this species in Latvia are W-Palearctic, SIE – Sibero-European, TUE – absent; therefore, its occurrence in local fauna needs Turano-European, EUR – European. further confirmation.

Results: annotated checklist of Latvian Dermestidae D. (Dermestinus) frischii Kugelann, 1792 = heyrovskyi Obenberger, 1917 Family Dermestidae Latreille, 1804 = pollinctus Hope, 1834 Subfamily Latreille, 1804 = rufimembris Pic, 1951 Tribe Dermestini Latreille, 1804 = sternimaculatiis Marcu, 1957 Dermestes Linnaeus, 1758 = vulpinus Herbst, 1792 Subgenus Dermestes s.str. Vītols 1941; Danka, Stiprais 1972; Barševskis 1988, 1993, 2001, 2002; Telnov et al. 1997, 2011; Telnov D. (Dermestes) ater DeGeer, 1774 2004; Kalniņš et al. 2007; Bukejs and Barševskis 2007. = cadaverinus Fabricius, 1775 = chinensis Motschulsky, 1866 D. (Dermestinus) gyllenhalii gyllenhalii Laporte, 1840 = domesticus Germar, 1824 = atomarius Erichson, 1846 = favarcqui Godard, 1883 Lackschewitz, Mikutowicz 1939 (atomarius Er.); š = felinus Fabricius, 1787 Bar evskis 1996, 2001; Telnov et al. 1997, 2006; Telnov š = hispidulus Montrouzíer, 1860 2004; Bukejs and Bar evskis 2007. = noxius Mulsant & Rey, 1868 = piceus Thunberg, 1781 D. (Dermestinus) laniarius Illiger, 1801 fi = rufoapicalis Pic, 1951 = af nis Gyllenhal, 1808 = subcostatus Murray, 1867 = catta Duftschmid, 1825 = subsulcatus Ballion, 1878 = macellarius Herbst, 1792 Fleischer 1829 (domesticus Gebl.); Seidlitz 1872–1875 = murinus Herbst, 1792 – – (domesticus Germ.), 1887–1891 (domesticus Germ.); Fleischer 1829; Seidlitz 1872 1875, 1887 1891 š Rathlef 1905 (cadaverinus F.); Trauberga 1957 (macellarius Hrbst.); Rathlef 1905; Bar evskis 1993; ņš (cadaverinus F.); Telnov et al. 1997; Telnov 2004; Telnov et al. 1997, 2007; Telnov 2004; Kalni et al. š Bukejs and Barševskis 2007. 2007; Bukejs and Bar evskis 2007. Note: Introduced species. There are no actual faunal data D. (Dermestinus) maculatus De Geer, 1774 on this species in Latvia; therefore, its occurrence in = cinereus L. Redtenbacher, 1868 local fauna needs further confirmation. = cyprius Pic, 1951 D. (Dermestes) bicolor bicolor Fabricius, 1781 = elongatus Hope, 1834 Telnov et al. 2005; Bukejs and Barševskis 2007. = kurseongensis Lepesme, 1939 Note: Introduced species. The species is known only = lupinus Mannerheim, 1843 from a single locality in Jelgava (Central Latvia). = marginatus Thunberg, 1781 = pakístanicus Havelka, 1951 D. (Dermestes) lardarius Linnaeus, 1758 = rattulus Mulsant & Rey, 1868

Downloaded by [Andris Bukejs] at 01:33 15 September 2013 = atrobasalis Pic, 1951 = roeí Hope, 1834 = conjunetus Reitter, 1887 = semistriatus Boheman, 1851 = luganensis Stierlin, 1902 = senex Germar, 1824 = proximus Reitter, 1887 = sudanicus Gredler, 1878 Fischer 1778, 1791; Groschke 1805; Precht 1818; Kawall = truncatus T. L. Casey, 1916 1865; Seidlitz 1872–1875, 1887–1891; Rathlef 1905; = vulpinus Fabricius, 1781 Heyden 1903; Vītols 1941; Danka 1950; Trauberga Precht 1818 (vulpinus); Lackschewitz, Mikutowicz 1939 1957; Šmits, Spuris 1966; Spuris 1974; Barševskis 1988, (vulpinus Fbr.); Telnov et al. 1997; Telnov 2004; Bukejs 1993, 2002; Telnov et al. 1997; Telnov 2004; Bukejs and Barševskis 2007. 2006; Kalniņš et al. 2007; Bukejs and Barševskis 2007; Note: Introduced species. The species is known only Barševskis, Bukejs, and Anichtchenko 2008; Barševskis from a single locality in Liepāja (SW Latvia). et al. 2009. D. (Dermestinus) murinus murinus Linnaeus, 1758 Subgenus Dermestinus Zhantiev, 1967 = catta Panzer, 1789 D. (Dermestinus) erichsoni Ganglbauer, 1904 = marinus Fabricius, 1787 = tessellatus Erichson, 1846 nec Fabricius, 1775 = nebulosus DeGeer, 1774 Seidlitz 1872–1875 (tessellatus Er.), 1887–1891 = roseiventris Laporte, 1840 (tessellatus Er.); Rathlef 1905; Telnov et al. 1997; Telnov Fischer 1778, 1791; Groschke 1805; Precht 1818; 2004; Bukejs and Barševskis 2007. Kawall 1865; Seidlitz 1872–1875, 1887–1891; Rathlef Zoology and Ecology 227

1905; Trauberga 1957; Danka, Stiprais 1972; Barševskis = cylindricornis (Schrank, 1785) and Savenkovs 1991; Barševskis 1993, 2002; Telnov = fruteti Grimmer, 1841 et al. 1997; Telnov 2004; Kalniņš et al. 2007; Bukejs = macellarius (Fabricius, 1781) and Barševskis 2007; Spuņģis 2008; Barševskis et al. = pilosissimus Roubal, 1932 2004, 2009; Barševskis, Bukejs, and Anichtchenko 2008. = quisquiliorum Gistel, 1857 = schranki (Kugelann, 1792) – ( ) D. (Dermestinus) mustelinus Erichson, 1846 Fischer 1784 (Dermestes), 1791 (Dermestes); Groschke = diadema Motschulsky, 1849 1805 (Dermestes); Precht 1818 (Dermestes); Kawall = tessellatus A. G. Olivier, 1790 1866; Seidlitz 1872–1875, 1887–1891; Rathlef 1905; – – Seidlitz 1872 1875, 1887 1891; Rathlef 1905; Telnov Heyden 1903; Vītols 1941; Danka 1950; Trauberga et al. 1997; Telnov 2004 (removed from the list); Bukejs 1957; Šmits, Spuris 1966; Spuris 1974; Priedītis 1979; š and Bar evskis 2007 (as excluded species). Barševskis 1988, 1993, 2002; Telnov et al. 1997; D. (Dermestinus) undulatus (Brahm, 1790) Telnov 2004; Kalniņš et al. 2007; Bukejs and Barševskis = caninus Gravenhorst, 1807 2007; Barševskis, Bukejs, and Anichtchenko 2008. = cylindricus T. L. Casey, 1916 A. schaefferi schaefferi (Herbst, 1792) = tessellatus Illiger, 1801 = bicolor Harold, 1868 = variegatus Bridle, 1832 = dichrous LeConte, 1854 = vulpecula Herbst, 1792 = sparsus T. L. Casey, 1916 Telnov 2004; Bukejs and Barševskis 2007. Fleischer 1829 (schaefferi Gyl.); Barševskis 1988, 1991, Note: Faunal data are available only in Ulanowski 1993; Telnov et al. 1997; Telnov 2004; Bukejs (1883). Actual faunal data on this species in Latvia are and Barševskis 2007; Barševskis, Bukejs, and absent; therefore, its occurrence in local fauna needs Anichtchenko 2008. further confirmation. Subfamily T. L. Casey, 1900 A. smirnovi Zhantiev, 1973 Tribe Trinodini T. L. Casey, 1900 Barševskis 1988, 1989, 1993, 2001; Telnov et al. 1997, Genus Trinodes Dejean, 1821 2006, 2007; Telnov 2004; Barševskis et al. 2004; T. hirtus (Fabricius, 1781) Barševskis, Bukejs, and Anichtchenko 2008; Bukejs = ater Dalla Torre, 1879 2006; Kalniņš et al. 2007; Bukejs and Barševskis 2007. = castanea Dalla Torre, 1879 Note: Introduced species. In Latvia, it is found only in = testacea Dalla Torre, 1879 human habitations. Precht 1818 (Anthrenus); Telnov 2004; Bukejs and A. unicolor unicolor (Brahm, 1791) Barševskis 2007. = brevicornis (Herbst, 1792) Note: For Latvian fauna, this species is mentioned only = cylindricornis (Schrank, 1798) in old bibliographic sources (Precht 1818). Actual faunal = cylindricornis Say, 1825 data on this species in Latvia are absent; therefore, its = deficiens T. L. Casey, 1900 occurrence in local fauna needs further confirmation. = floricola Say, 1825 Subfamily Attageninae Laporte, 1840 = fulvipes Mulsant & Rey, 1868

Downloaded by [Andris Bukejs] at 01:33 15 September 2013 Tribe Attagenini Laporte, 1840 = macellarius (Duftschmid, 1825) Genus Attagenus Latreille, 1802 = megatoma (Fabricius, 1798) A. brunneus Faldermann, 1835 = megadoma Sturm, 1826 = bicolor T. L. Casey, 1900 = obscurus Say, 1825 = dalmatinus Küster, 1847 = piceus (A. G. Olivier, 1790 nec Thunberg, 1781) = elongatulus T. L. Casey, 1900 = schaefferi (C. Sahlberg, 1819) = elongatus T. L. Casey, 1916 = stygialis Mulsant & Rey, 1868 = extricatus T. L. Casey, 1900 = urbicola Gistel, 1857 = longicornis Pic, 1894 Fleischer 1829 (megatoma F.); Seidlitz 1872–1875 = marginicollis Handschuch in Küster, 1847 (megatoma Fbr.), 1887–1891 (piceus Ol.); Rathlef 1905 = picipennis Pic, 1894 (piceus Ol.); Vītols 1941 (piceus Ol.); Barševskis 1993; = sordidus Heer, 1841 Telnov et al. 1997, 2007; Telnov 2004; Kalniņš et al. Barševskis 1991, 1993; Telnov et al. 1997; Telnov 2004; 2007; Bukejs and Barševskis 2007; Barševskis, Bukejs, Kalniņš et al. 2007; Bukejs and Barševskis 2007. and Anichtchenko 2008. Note: Introduced species. In Latvia, this species is associated with human habitation only. Subfamily Leach, 1815 Tribe T. L. Casey 1900 A. pellio (Linnaeus, 1758) Genus Anthrenus Geoffroy, 1762 = atra (Herbst, 1792) Subgenus Anthrenus s.str. = bipunctatus (DeGeer, 1774) = Byrrhus Sulzer, 1776 228 A. Bukejs et al.

A. (Anthrenus) pimpinellae pimpinellae (Fabricius, Mroczkowski); Bukejs and Barševskis 2007 (fuscus 1775) A. G. Olivier and polonicus Mroczkowski); Barševskis, = dimidiatus Hauser, 1894 Bukejs, and Anichtchenko 2008 (fuscus A. G. Olivier = scrophulariae Geoffroy, 1785 and polonicus Mroczkowski). Seidlitz 1872–1875; 1887–1891; Rathlef 1905; Telnov et al. 1997; Telnov 2004; Barševskis et al. 2004; Bukejs Subgenus Nathrenus T. L. Casey, 1900 and Barševskis 2007. A. (Nathrenus) verbasci (Linnaeus, 1767) = adspersus Herbst, 1797 A. (Anthrenus) scrophulariae scrophulariae (Linnaeus, = caseyi Hinton, 1945 1758) = confusus Reitter, 1886 = histrio Fabricius, 1792 = destructor Melsheimer, 1844 = insulicola Obenberger, 1917 = florilegus Geoffřoy, 1785 = rubricollis J. R. Sahlberg, 1913 = fiinebris Reitter, 1889 = suecicus Palm, 1940 = nebulosus Reitter, 1886 = variegatus Scopoli, 1763 = nitidulus Küster, 1847 = verbasci Herbst, 1797 = pictus Germar, 1813 Fischer 1791 (Byrrchus scrophulariae L.); Precht 1818; = pistor T. L. Casey, 1900 Seidlitz 1872–1875, 1887–1891; Rathlef 1905; Vītols = substriatus T. L. Casey, 1900 1941; Priedītis 1979; Rūtenberga 1980; Barševskis 1988, = tricolor Herbst, 1797 1993, 2002; Telnov et al. 1997; Telnov 2004; Bukejs = varius (Fabricius, 1775) 2006; Kalniņš et al. 2007; Bukejs, Barševskis 2007; = versicolor Reitter, 1887 Barševskis, Bukejs, and Anichtchenko 2008. = vorax T. L. Casey, 1900 Precht 1818; Barševskis 1993, 2002; Bukejs and Subgenus Florilinus Mulsant & Rey, 1868 Barševskis 2007. A. (Florilinus) caucasicus Reitter, 1881 Note: This species is known from a single locality = armeniacus Zaitzev, 1919 (Ilgas) in the extreme SE of Latvia located at the border Vītols 1941. with and . Note: Introduced species. Tribe Megatomini Leach, 1815 A. (Florilinus) museorum (Linnaeus, 1761) Genus Ctesias Stephens, 1830 = museorum Fabricius, 1787 Subgenus Ctesias Stephens, 1830 = pellio Thunberg, 1815 = Tiresias Stephens, 1835 = vagus Gmelin, 1790 = varius Stephens, 1830 C. serra (Fabricius, 1792) = verbasci Fabricius, 1775 = viennensis (Herbst, 1797) Precht 1818; Seidlitz 1872–1875, 1887–1891; Rathlef Seidlitz 1872–1875 (Tiresias), 1887–1891 (Tiresias); 1905; Vītols 1941; Danka 1950; Trauberga 1957; Šmits Rathlef 1905; Telnov et al. 1997; Telnov 2004; Bukejs and Spuris 1966; Rūtenberga 1980; Barševskis, 1988, and Barševskis 2007; Barševskis, Bukejs, and 1993, 2002; Telnov et al. 1997; Telnov 2004; Bukejs Anichtchenko 2008.

Downloaded by [Andris Bukejs] at 01:33 15 September 2013 2006; Kalniņš et al. 2007; Bukejs and Barševskis 2007; Barševskis, Bukejs, and Anichtchenko 2008. Genus Globicornis Latreille, 1829 Subgenus Hadrotoma Erichson, 1846 A. (Florilinus) olgae Kalik, 1946 G. (Hadrotoma) emarginata (Gyllenhal, 1808) Barševskis 1997a; Telnov et al. 1997, 2006, 2008; Tel- = marginata (Paykull, 1798 nec Thunberg, 1781) nov 2004; Kalniņš et al. 2007; Bukejs and Barševskis Seidlitz 1872–1875 (marginata Payk.), 1887–1891 2007. (Hadrotoma marginata Payk.); Rathlef 1905; Telnov et al. 1997; Telnov 2004; Bukejs and Barševskis 2007. Subgenus Helocerus Mulsant & Rey, 1868 Note: For the Latvian fauna the species is mentioned A. (Helocerus) fuscus A. G. Olivier, 1789 only in old bibliographic sources. Actual faunal data on = claviger Erichson, 1846 this species in Latvia are absent, therefore its occurrence = polonicus Mroczkowski, 1951 in local fauna needs further confirmation. Mikutowicz 1905; Rathlef 1921; Barševskis 1993 (fuscus A. G. Olivier and polonicus Mroczkowski), 2002 (fuscus Genus Megatoma Herbst, 1792 A. G. Olivier and polonicus Mroczkowski); Telnov 1996 Subgenus Megatoma Herbst, 1792 (polonicus Mroczkowski), 2004 (fuscus A. G. Olivier M. pubescens (Zetterstedt, 1828) and polonicus Mroczkowski); Telnov et al. 1997 (fuscus = fuscus (Gebler, 1841) A. G. Olivier and polonicus Mroczkowski); Kalniņš = innotata Pic, 1902 et al. 2007 (fuscus A. G. Olivier and polonicus Barševskis 1997b; Bukejs and Barševskis 2007. Zoology and Ecology 229

Note: This species is known only from a single locality = parabile Beal, 1954 in Naujene (extreme SE of Latvia). Barševskis 1991, 1993; Telnov et al. 1997; Telnov 2004; Kalniņš et al. 2007; Bukejs and Barševskis 2007. M. undata (Linnaeus, 1758) Note: Introduced species. The species is known only = glabera (C. R. Sahlberg, 1819) from a single locality in Kuprava (E Latvia). = undulatus (Linnaeus, 1761) = unifasciata Ganglbauer, 1904 T. versicolor (Creutzer, 1799) Precht 1818 (Dermestes undatus); Kawall 1865 = costae Mulsant & Rey, 1868 (Attagenus undatus); Seidlitz 1872–1875, 1887–1891; = elongatulus (Duftschmid, 1825) Rathlef 1905; Lackschewitz, Mikutowicz 1939; Vītols = flexuosa C. G. Thomson, 1862 1941; Barševskis 1993, 2002; Telnov et al. 1997, 2005, = grandis (Thunberg, 1815) 2006, 2008, 2010, 2011; Leiskina 2000; Telnov 2001, = hieroglyphica Abeille de Perrin, 1872 2004; Bukejs and Barševskis 2007; Barševskis, Bukejs, = reductum Pic, 1919 and Anichtchenko 2008. = testaceicorne Perris, 1862 Seidlitz 1872–1875, 1887–1891; Rathlef 1905; Telnov Subgenus Pseudohadrotoma Kalík, 1951 1996, 2004; Telnov et al. 1997; Kalniņš et al. 2007; M. (Pseudohadrotoma) graeseri (Reitter, 1887) Bukejs and Barševskis 2007. = latenotata Pic, 1902 Note: Introduced species. Material examined:Cēsis district, Gauja National Park, Cēsis, Pārgaujas Str., 8 July 2004 (1, meadow, leg. I. Salmane). Material deposited in the private collection Taxonomical structure of Latvian Dermestidae fauna of D. Telnov. According to cladistic-based analysis of higher taxa within Note: New species for the Latvian fauna. the family Dermestidae (Lawrence and Slipinski 2005; Kiselyova and McHugh 2006), six subfamilies should be Genus Reesa Beal, 1967 recognized: Dermestinae (including Marioutinae), R. vespulae (Milliron, 1939) Thorictinae, Trinodinae (including Thylodriadinae), š Telnov et al. 2005, 2008; Bukejs and Bar evskis 2007. Orphilinae, Attageninae, and Megatominae. Note: Introduced species. In Latvian fauna, 33 species belonging to four subfamilies (Dermestinae, Trinodinae, Attageninae and Genus Trogoderma Dejean, 1821 Megatominae) are known. The number of known T. angustum (Solier, 1849) Dermestidae species in adjacent territories varies slightly: = bifasciata Redtenbacher, 1867 21 species in (Silfverberg 2010), 27 species in = boliviensis Pic, 1915 Lithuania (Tamutis, Tamutė, and Ferenca 2011). The Barševskis 1993; Telnov et al. 1997, 2007, 2008; Telnov taxonomical structure of dermestids of the Latvian fauna 2004; Barševskis et al. 2004; Barševskis, Bukejs, and and its comparison with those of the adjacent countries Anichtchenko 2008; Bukejs and Barševskis 2007. (Estonia and Lithuania) is shown in Table 1. Note: Introduced species. In Latvia, it is associated only with human habitation. Table 1. Taxonomical structure of Dermestidae of Latvia T. glabrum (Herbst, 1783) (LV), Lithuania (LT) and Estonia (EE). Downloaded by [Andris Bukejs] at 01:33 15 September 2013 = boron Beal, 1954 = elongatulum (Fabricius, 1801) Number of species = fuscicornis Mulsant & Rey, 1868 Taxa LV LT EE = limbatum Pic, 1934 Dermestinae = niger (Herbst, 1797) Dermestes 10 8 8 = ruficornis (Latreille, 1807) = subfasciatus (Gyllenhal, 1808) Trinodinae = versicolor (Illiger, 1801) Trinodes 110 Seidlitz 1872–1875 (elongatum Fbr.), 1887–1891 Attageninae (nigrum Herbst); Rathlef 1905 (nigrum Hrbst.); Lack- Attagenus 544 schewitz, Mikutowicz 1939 (nigrum Hrbst.); Trauberga 1957 (nigrum Hbst.); Barševskis 1993, 1997a; Telnov Megatominae et al. 1997, 2008; Telnov 2004; Bukejs and Barševskis Anthrenus 754 2007; Barševskis, Bukejs, and Anichtchenko 2008. Ctesias 111 Globicornis 121 T. variabile Ballion, 1878 Megatoma 321 Reesa 101 = persica Pic, 1914 Trogoderma 441 = versicolor var. turkestanicum Pic, 1914 = scabripennis Casey, 1916 Total 33 27 21 230 A. Bukejs et al.

References Part A. Bibliography on Latvian Dermestidae Barševskis, A. 1988. “Фаунистические исследования жуков юго-восточной части Латвии в раpличных биотопах [Faunal Investigations on Beetles in Various Habitats of the South-eastern Part of Latvia].” В Экологическое воспита- ние в средней и высшей школе: теpисы республиканской научно-методической конференции.(Проблемы краеве- дения). Часть 3 [Ecological Education in Secondary and Higher School, Part 3], под ред. Э. А. Вайводе,63–75. Даугавпилс: ДПИ [DPI]. Barševskis, A. 1989. “Par dažām jaunām un mazpazīstamām vaboļu sugām Latvijas faunā [On New and Little-known ” Figure 1. Biogeographical structure of Latvian Species of Beetles in the Fauna of Latvia]. Latvijas – Dermestidae: COS – Cosmopolitan, OLA – Holarctic, SCO – Entomologs 32: 52 53. š “ ž ļ Subcosmopolitan, PAL – Palearctic, ASE – Asiatic-European, Bar evskis, A. 1991. Da as jaunas vabo u sugas Latvijas ā ” WPA – W-Palearctic, SIE – Sibero-European, faun [Some New Beetles Species in the Fauna of Latvia]. – TUE – Turano-European, EUR – European. Latvijas Entomologs 34: 28 32. Barševskis, A. 1993. Austrumlatvijas vaboles [The Beetles of Eastern Latvia]. Daugavpils: Saule. Barševskis, A. 1996. “Latvijas faunai jaunas un retas vaboļu (Coleoptera) sugas [New and Rare Species of Coleoptera in Biogeography of Latvian Dermestidae the Fauna of Latvia].” Daba un muzejs 6: 16–18. š “ ā ē Nine chorotypes represent the Latvian fauna of skin Bar evskis, A. 1997a. Materi li par Latvijas vabol m (Coleoptera) [Materials about Latvian Beetles beetles: ” – – (Coleoptera)]. Acta coleopterologica latvica 1 (2): 63 71. Cosmopolitan 13 species (39.40%): Dermestes ater, Barševskis, A. 1997b. “Augšdaugavas vaboļu (Insecta: D. lardarius, D. frischii, D. maculatus, Attagenus pellio, Coleoptera) faunistiskās īpatņības [Faunistical Peculiarities A. unicolor, Anthrenus pimpinellae pimpinellae, of Beetles (Insecta: Coleoptera) of Augšdaugva].” Daba un – A. scrophulariae scrophulariae, A. museorum, A. verba- muzejs 7: 8 9, (+ tab.). Barševskis, A. 2001. “New and Rare Species of Beetles sci, Reesa vespulae, Trogoderma variabile, T. versicolor; ” – (Insecta: Coleoptera) in the Baltic Countries and Byelorus. Subcosmopolitan Four species (12.12%): Attagenus Baltic Journal of Coleopterology 1(1–2): 3–18. smirnovi, Anthrenus fuscus, Trogoderma angustum, Barševskis, A. 2002. “Vaboļukārta (Coleoptera) [Beetles Trogoderma glabrum; (Coleoptera)].” В Silenes dabas parka fauna, flora un veģe- ā Holarctic – Four species (12.12%): Dermestes t cija [Fauna, Flora and Vegetation of Silene Nature Park], под ред. A. 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The northern border of the geographic range of Bukejs, A. 2006. “Materials about the Fauna of Beetles Dermestes gyllenhalii Laporte goes through Latvia. (Insecta: Coleoptera) of Naujene Rural Municipality (Daugavpils District, Latvia). Part 1.” Acta Biologica Universitatis Daugavpiliensis 6(1–2): 65–75. Acknowledgement Bukejs, A., and A. Barševskis. 2007. “Materials about Lat- vian Fauna of Dermestids (Coleoptera: Dermestidae).” Sincere thanks to Dr Sergejs Berdnikovs (Northwestern Acta Biologica Universitatis Daugavpiliensis 7 (1): fi University, Chicago, USA) for comments on the nal version 29–36. of the manuscript and linguistic corrections. Zoology and Ecology 231

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