ADDITIONS TO THE HETEROPTERA-FAUNA OF THE 1

By E. HEISS & J. RIBES **

ABSTRACT: The present paper is a first addition to tbe preliminary cataloguo Jf the Heteropteru-fauna of the Canary Islands (HEISS & BAEZ.1990) and includes corrections, new taxonomic assignmcnts and further citations from literature and several new records. 29 species are reported as new to the Canarian fauna, but one species is a synonym (Geocoris pvbescens Jak. = G.timidus Put.) and 8 spccies are regarded as doubtíidspccies inquirenda which should be deleted from the list. This brings the total number of species known to date from Canary Islands to 333. Key words: Heteropteru, catalogue, new additions, Canary Islands RESUMEN: La presente aportación es una primera addenda al catálogo preliminar de los Heterópferos des las islas Canarias (HEISS & BAEZ, 1990). Incluye enmiendas, nuevos cambios taxonómicos, datos complementarios y numerosas citas inéditas. Todo ello se desglosa en cuatro secciones distintas dentro del segundo capítulo: referencias adicionales y cambios en la literatura, citas canarias nuevas, citas isleñas nuevas y specics inquirenda. De este acervo hay que entresacar 29 especies para incorporar a la fauna canaria, con relación a lar 3 13 inventariadas, pero una es una sinonimia (Geocoris pvbescem lak. = G.rimidus Put.) y 8 especies más se consideran dudosas, por lo que se eliminan de la lista, quedando pues un total de 333 especies que se estiman válidas. Toda la bibliografía es nueva.

1.1 N TRO D UC TI ON The publication of “A preliminary catalogue of the Heíeropíera of the Canary Islands” by HEISS & BAEZ (1990) as a compilation of the then available data from literature was an attempt to provide a kind of practica1 “check-list” without pretension of completeness. Since its preparation (1987) and during publication time additional informa- tion concerning omissions. corrections, new taxonomic assignments and new records were gathered by both present authors and are summarized in this paper.

~~ Joacf Schraffl-Strasse 2A, A-6020 Innsbruck. Austria ** Valencia 123-125, ent. 3a. E-0801 1 Barcelona, Catalonia,

Bol. Mur. Mun. Fuucbil, 44 (238): 77- 102 , 112 ISSN 0870 - 3876 78’ Boletim do Museu Municipal do Fundia No. XLN, Art. 238

The recent publications by HOBERLANDT( 1987); PUTSHKOV (1987) and Sigara (Halicorixa) selecta (FIEBER,1848) BAENA & BAEZ (1990) also comprise references of Canarian Heteroprera which are Sigara lugubris (FIEBER, 1848) = S.stagnalis (LEAcH,~~17) of NOUALHIER included here. CARVALHO (1957-i960), STICHEL (1955-1962), SLATER (1964) and (1893) sec. BAENA & BAEZ (1990); S.slagnalis NIESER & MONTES (1984) VAZQUEZ (1 987) repeat some very doubtful ancient records which should be C removed from the Canarian list, unless their presence there is proved. For practica1 reasons we use the same sequence of categories and abbrevia- Family No t o n e c t i da e tions as used in the “preliminary catalog”. It differs from the recently published “Verzeichnis der Wanzen Mitteleuropas” (G~NTHER & SCHUSTER, 1990) by Anisops sardeus HERRICH-SCHAEFFER,1853 another collocation of some suprageneric categories within the basically identical BAENA & BAEZ (1990) system e.g. now Leptopodomorpha come after Gerromorpha and Tingoidea are at the Alegranza beginning of . Family H e b r i da e The indication 1/1 refers to the number of malelfemale spechens found. Hebrus pusillus (FALLEN, 1807) 2. TAXONOMIC RESULTS The taxonornic status of the reported ssp. canariensis PO1SS0Ny1954 is questioned by NIESER & MONTES (1 984) and BAENA & BAEZ(1990) 2.1 Additional references and taxonomic‘changes from literature Merragata hebroides BUCHANAN WHITE, 1877 Family Corixidae BAENA & BAEZ (1990) Subfam. C o r i x i n aie H,F

Corixa alfinis LEACH, 1817 Family Hydro m e f r ida e New to G, Alegranza, BAENA & BAEZ (1990) Hydrometra stagnorum (LINNAEUS, 1758) Sigara (Vermicorixa) lateralis (LEACH, 1817) New for F, L, cited in a tabie withouth giving data, BAENA & BAEZ (1990) The ssp.nakurui POISSON, 1958, which has beenreported from CanqIslands by LMDBERG (1961) and HOBERLANDT (1967) is a mere synonym of lateralis sec. FamilyMiridae JANSSON (1986), BAENA & BAEZ (1990) Subfam. D i cyph ina e Alegranza Cyrtopeltis (Canariesia) salviae CARAPEZZA, 1988 Sigara (Tropocorixa) hoggarica POISSON, 1929 CARAPEZZA (1988) BAENA & BAEZ (1990) .. New subgenus and species described on material from Tenerife. C T

Sigara (Vermicorixa) scripta (RAMBUR, 1842) Dicyphus baezi RIBES, 1983 It has been verified that the reported specimens belong to S.lateralis The unknown female has been discovered and a brief description is given: (LEACH, 1817), for which reason this species has been excluded from the Canarian Macmpterous. Hemelytra slightly surpassing the apex of abdomen (considerably fauna by BAENA & BAEZ (1990) in). Antennal segment 11 yellowish with dark basd 1/3 and black apical 1/3. in 111 only the basa1 1/4 is yellowish (11 with a wide dark subapical ring and iight coloured

. I 81 80 Bolethn do Museu Municipal do Funchal No. XLiV, Ah 238 Heiss & Ribes, Additions to he Heteroptera-hu~of the Canary 1s. 1

111 in ). Posterior lobe of pronotum with transverse rugosities (not distinct in ). Dark Cardiasleihus, Scoioposceiis, Xylocoris) are therefore placed here in Xylocorinae. coloration on corium less extended than in male. Morphology and pilosity as in male. Length 4.5 mm. Xylocoris obliquus COSTA, 1852 T - Aguamansa, 011 1.X.56. MORALES leg.er coll., RIBES der. GOMEZ MENOR (1956); STICHEL(1955 -62); not mentioned from the Canary lslands by PBRICART(1972) Subfam. M i r i n a e T

Pinalirus conspurcatus (REUTER, 1873) = Orrhops conspurcarus Family N a b i d a e KERZHNER(1988a) Subfam. Nabinae New generic assignment 1 l \ Halonabis sarepíanus occidentalis KERZHNER, 1963 Pinalitus insularis (REUTER, 1895) = Orrhops insularis KERZHNER (1988b) KERZHNER(1988a) The author has revised the material cited by LiNDBERG & WAGNER( 1965) and New generic assignment confirms that it belongs to H.occidentalis. KERZHNER (1963) has originally de- scribed occideníalis as a ssp. ofsareptanus but gave it specific rank later. PBRICART Phylocoris (Eckerleinius) meinanderi WAGMR, 1968 (1987), whom we follow here has treated it again as a subspecies. WAGNER 1968,1970-75,1978; LINNAVUOU (1972); RIBES (1978) F, L,Graciosa Family Reduviidae Subfam. E m e s i n a e Subfam. P h y 1 i n ae

Compsidoion (Chamaeliops) coiiare WAGNE~,1976 I Empicoris salinus (LINDBERG, 1932) WAGNER (1 978), add.ref.for T l VIDAL 1938; later repeated by DISPONS 1955; STICHEL (1955-62); ! WYGODZINSKY 1966; WBES (1973)

Maurodactylus gomercnsis WAGNER, 1954 I “Canary lslands” ASWOLE & ASHMOLE (1988a) T - 1/1,15. and 22.11.84, Teide on snow, RIBES der.er coll. The record given by VIDAL refers to a specimen, which has been found in Morocco on bananas, which were imported from Canary Islands. Now a male . specimen from Tenerife, HElSS leg., which belongs to the salinus-group proves the Tuponia (Chlorotuponia) verricaíu WAGNER, 1975 previous record. A revision of the western europeaa species of Empicoris is in WAGNER 1978, add.ref.for T preparation by PUTSHKOV and RIBES.

Ploiaria chilensis (PHILIPPI, 1862) Family Anthoc0rida.e WYGODZINSKY 1966, ASHMOLE & ASHMOLE (1988b), add.ref.for T Subfam. X y 1 o c o r i n a e Subfam. P e ira t i n a e The former subfamily Lyclocorinue has been given family rank (with the only Lycrocoris) by SCHUH & STYS (1991). Folloping their well founded assign- ’ The name Peiruíinae AMYOT el SERVILLE,( 1843) and PEIRATES SERVILLE, ment, the remaining genera reported from Canary Islands (Bruchysieles, (183 1) hasg been introduced by PUTSHKOV & PUTSHKOV (1985). Subsequently a2 Boletim do Museu Municipal do Funchal No. XLIV, Art. 238 1 1992 Heiss & Ribes, Additioos to the Heteroptera-fauna of the Canary 1s. I 83 i followed by KANYUKOVA (1988) but not by PUTSHKOV (1987) and GÜNTHER& Neides aduncus FIEBER, 1859 SCHUSTER (1990). 1 PBRICART(1984), additional record for l C i Ectomocoris chiragra (FABRICIUS, 1803) c Peirates chiragra l LINNAVUORI (1972a) 1 Family L y g a e i d a e New generic assignment Subfam. O r s i 1 i n a e

Subfam. H a r p a c t o r i n a e Nysius (Macropari.us) cymoides (SPRJOLA, 1840) Coranus kerzhneri PUTSHKOV, 1982 BL~TE(1929), LINDBERG (1936,1953) PUTSHKOV (1987) As part o€ the material cited by the latter author was used for the description “Canary Islands” t of N.latus WAGNER, 1958, the distribution of N-cymoides on the islands is not clear, but seems to be common. Family Aradigae PGTC F(?) Subfam. A r a d i n a e Nysius (sstr.) thymi latus WAGNER, 1958 = N’sius latus WAGNER, 1958 Aradus canariensis KORMILEV, 1954 LINNAVUORI (1986). ASHMOLE& ASHMOLE (1988a, 1989) According to PÉRlCART in-lirt. N.latus might be a synonym of Nericae T - add. reference from Teide on snow, 110 15.1134, RIBES der. (SCHILLiNG,1829). ASHMOLE & ASHMOLE (1988a) report some specimens from T, Teide on snow, 30.III.and 1.V.84, RIBES det. Family B e r y t i d a e New specific assignment. Subfam. Berytinae Subfam. I s c h n o r h y n c h i n a e Beryrinus (Lizinus) geniculatus HORVATH, 1885 PÉRKART(1984) Kleidocerys ericae (HORVATH, 1909) = K. lruncatulus truncatulus auct. T CARAYON, 1989 The author has investigated the European species and considers K.truncatulus Berytinus (s.str.) hirticornis pilipes PUTON, 1875 (WALKER, 1872) as an endemic species of Madeira. = hirticornis nigrolineatus in HEISS &¿ BAEZ (1990) Canaria specimens belong to the widespread Euro-Mediterranean K.ericae. PERICART(1984) Subspecific reassignment as ssp. pilipes represents the west mediterrgnean Subfamily G e o c o r i n a e subspecies. Recorded from W,T Geocoris (Eilatus) chloroticus PUTON, 1888 = G.nigriceps ,189 1 LINNAVUORI (1972b) Berytinus (Lizinus) montivagus (MEYER-DÜR,184 1) New subgeneric assignment. PBRICART(1 984), additional record for PÉRICART & RIBES (1992) C New synonymy.

Geocoris (s.str.) pubescens (JAKOVLEV,1871) = G.timidus (PUTON, 1873)

, 84 Fkdetim do Museu Municipal do Funchal No. XLIV, Art 238 1992 Heiss & Ribes, Additions to thc Heteropkra-fauna of the Canary 1s. 1 85

PÉRICART & RIBES, 1992 Corizus hyoscyami nigridorsum (PUTON, 1874) New synonymy. GOLLNER-SCHEIDMG (198O);VAZQUEZ (1987) New subspecifk assignment. Al1 records reported as C. nigridorsum belong to Subfam. O x y c a r e’n n a e this subspecies.

Neocampfofeiusaeonii (LINDBERG, 1953) = Camptofelus aeonii Sfictopleurus abuiilon (ROSSI, 1790) HOBERLANDT (1987) GOLLNER-SCHEIDING (1 975);VAZQUEZ (1987) New genus erected for this species Was unclear to LINDBERG (1953),who assigned al1 records to S.picrus. As GOLLNER foc.cif.indicates. that she has seen material from the Canary Islands, no Befhylimorphus leucophaes LINDBERG, 1953 details of distribution on the islands are known to date. SEIDENST~CKER(1964) New assignment to subfamily Oxycareninae not Bledionotinae Siictopleurus ribesi GOLLNER-SCHEIDING, 1975 GOLLNER-SCHEIDING (1975),vAZQUEZ (1987) Subfam. R h yp aroc h r o m i n a e T

Remaudiereana annulipes (BARENSPRUNG,1859) = Pachybrachius Sficfopleurussubfomentosus REY, 1888 annulipes SCUDDER (1962) GOLLNER-SCHEIDING (1975),VAZQUEZ (1987), MOULET (1991) New generic assignment Sticfopleurus subtomentosus REY,1888 = S.rivefi RoYER,~~~~(MOULET , 1991) T Eremocoris wagneri LINDBERG, 1961 = ? E.maderensis HEISS & BAEZ, 1990 LATER S (1 964); TAMANINI(1974) Family Sc u f e 11 e r i dae The latter author considers this species synonymous with E. maderensis WOLLASTON, withOut formally synonymizing them. Odontotarsus iniermedius HORVATH, 1923 HPGT GOLLNER-SCHEIDING(1990) Canary Islands. No data of dispersion on the islands are given. Family C o r e i d a e Subfam. C o r e i n a e Odonfoscelis dorsalis (FABRICIUS,1803) = 0.lineola (RAMBUR, 1842) in HEISS & BAU 1990 Haploprocta sulcicornis insularis LINDBERG, 1953 GYLLENSVARD1968 (0.dubiu.rJdorsalis!); GOLLNER-SCHEIDING1986 (1953); HOBERLANDT (1967); VAZQUEZ (1987) LINDBERG LiNDBERG 1961, reassigned previous records of O.fuliginosa to 0.dubia Al1 Canarian records belong to this ssp. r+ WAGNEK1957. As the latter species has later been synonymized with 0.lineola by KERZHNER 1976, HEISS & BAEZ reported LINDBERG’s and GYLLENSVARD’Sdubia Family Rhopalidae as lineola. According to GOLLNER-SCHEIDING “1ineola”-species from the Canary Subfam. R h o p a 1 i n a e Islands belong certainly to dorsalis.

1 Lepfoglqssus australis (FABRICIUS, 1775)=L..membranaceus (FAB- FamilyP ent atom i d a e RICIUS, 178 1) Subfam. A s o p i n a e ALLEN (1969) Macrorhaphis acufa DALLAS, 185 1

, 85 87 Boletim do Museu Municipal do Funchal No. XLN, Art. 238 1992 Heiss & Ribes, Additions to the Heteroptem- of the Canary 1s. 1

SCHMITZ (1 976:3 73) Immature species. The author reports that he has seen a specimen from the Canary Islands, but T - Tejeína,O/l 22.V.80, MORALES leg.et coll., RIBES dei.This is similar to gives no further details. Species witb African dishibution, also known from the Cape specimens collected in the desert of Tafilalt, Morocco in 1990. Verde Islands. SE Family A n t h o c o r i d a e Subfam. P e n t a t o m i n a e Subfam. D u f o ur i e 11 i n a e

Dolycoris baccarum (LINNAEUS,1758) Dufourieflus aler (DUFOUR1833) TAMANINi (1959); LINDBERG (196 1) F - Vega de Río Paimas, 10.V.88, a series on trunks of fig tree, MORALES leg., C IUBES der. etcoll. LMDBERG (1953) reports only D.numidicus from the Canary Islands, but Family Ly c t o c o r id a e TAMANMi (1959) excludes it from there and gives the only reference for baccarum. Contrariiy REMANE & KAMMERSCHEN (1991b) retain TAMANMI’s citation as very Lyctocoris dimidiatus (SPMOLA,1837) doubtful as al1 specimens they collected in Madeira and the Canary Islands belong (s.str.) T - Bajamar, 0/1 11.75, BACALLADO leg., RIBES der.et coll. to numidicus. They also state that only the males c& be distinguished. Whatever the case may be, fiuther material is necessary to prove if both species occur on the Family R e d u vi i d a e Canary Islands like Coranus aegyptius and Cgriseus (see section 2.2). Subfam. Ha r p a c t o r i n a e

Family C y d n i d a e Coranus griseus (ROSSI, 1790) Subfam. S e h i r i n ae T - Las Cañadas,I/O 26.11.64 RIBES leg.,Candelaria,25.11.91 YkLAMOS leg., EUBES der.sensu PUTSHKOV.The species is recorded as “C.aegyp2ius” auct. from Ochetosiethus iarsaiis (MULSANT, 1852) = 7 0,opacus of HEISS & BAEZ Europe. ( 1990) KERZHNER (1976) did not report opacus from Western Europe or Maghreb, Corunus aegyprius (FABRICIUS,1775) for which reason the Canaran specimens had betteE be assigned to tarsalis. T - Santa Cruz de Tenerife,I/O 26.111.82 CAMPOS leg., same locality,costa sur,2/2 7.and 22.XII.58, MORALES leg. er coll., RIBES der. el coll. Subfam. C y d n i n a e . G - San Sebastián,l/O 23.111.91, YÉLAMOSleg. Recorded also from Madeira; Ponta de SHo Lourenqo .2.V1.89 de FERRER leg. RlBES det.sensu PUTSHKOV. Aethus pallidus (PUTON, 1887) ,= Byrsinocoris paliidus WAGNER, 1964 The previous citations of this species are to be verified. New generic assignment due to synonymy Family T i n g i d a e 2.2 Additional records new to the Canary Islands Subfam. T i n g i n a e

Family M i r i d a e Tingis (Tmpidocheila) ajugarum (FREY - GESSNER, 1872) Subfam. P h y 1 i n a e L - Playa Blanca, 8/4 1.-6.111.90 on a small Lumiaceae, HEISS leg. Chamaepsalius signaiicornis (REUTER, 1904) H - Boca Cueva de Don Justo,O/l 2.VIII.87, ASHMOLE leg., RIBES dei.et coll. 88 89 1 Boletim do Museu Municipal do Funcha No. XLN,Art. 238 1992 Heiss & Ribes, Additioas to the Hderoptera-Fauna of the Canary 1s. 1

Family Ar ad i d a e T - Santa Cruz de Tende, 25.UI.82 OROMt leg., RIBES del .el coll. one de. Subfam. A r a d i n a e Subfam. P e n t a t o m i n a e Aradus reurerianus PUTON, 1875 P - Cumbre 2000m, Roque de los Muchachos,l/l lL,spring 1983 on rotten Sciocoris (Neosciocoris) conspurcarus majusculus LiNNAVUORi, 1964 Iogs of Pinus canariensis, FRANZ leg. HEISS der.,coll. FRANZ et HEISS (= S.carayoni WAGNER1965 ) T - ElGaitero, 110 17.VII1.79, AGUILARleg., RIBES det.,coll. Univ. La Laguna F - Corraiejo,l2.V.88 de FERRER leg. NBES det.This species and C.flssus MLS. & REY are difficult to distinguish from each other but the single female Family Lyg a e i d a e specimen shares the characters given for the West mediterraneanssp.of conspurcarus. Subfam. L y g a e i n a e Family C y d u i d a e Spilosiethus saxarilis (SCOPOLI, 1763) Subfam. C y d n i n a e L - Playa Grisineta,17.VIII.72 COMAS leg, RIBES der., Famara, XII. 91, a series on Androcymbiumgramineum ssp.psammophilum (Svent.) Kunkel (Liliaceae), Georomus punclulatus (COSTA, 1847) leg. HANDKE vid. HEISS. F - La Oliva,1/0 6.XII.79, OROMf leg., RiBES der el colf.

Subfam. G e o c o r i n a e Georomus longicornis WAGNER,1953 F - Presa de las Molinos,O/l 1.111.90, OROMí leg.,RIBES det.er coll. Geocoris (s-str.) collaris PUTON, 1878 VERISIMILITER F - , 1/0 13.11.77, OROMI leg., RIBES der.el coll."", 1/ O, FRANZ leg., RIBES det. 2.3 Additional records new to individual islands L - Playa Blanca,O/l and Jaiza,O/l 1.-6.III.90, HEISS leg.er coll., RIBESdet. Cyrtopelris (Nesidiocoris) lenuis (REUTER, 1895) Geocoris (s.str.) eryfrops (DUFOUR, 1857) = G. scutellaris PUTON, 1886 L - Famara,XII.91, leg. HANDKE. F - Corralejo, 12.V.88 de FERRER leg. RIBES der. a series of specimens. They are identical to specimens which were collected in the desert of Tafilalt, SE Morocco Eurystylus occidenialis WAGNER, 1954 in 1990. L - Playa Blanca,l.-6.111.91 a series on Suaeda sp., HEISS leg. L - Playa Blanca,0/2 1.-6.III.90, HEISS leg.er coll., RIBES det.The above synonymy is discussed by PÉRICART& RIBES (1992): Aeíorrhinella parviceps NOUALHIER, 1893 H - Mancafite,l/O 10.V.84, OROMí leg., NBES der.el coll. Subfam. R h yp aro c h r o m i n a e Hererocordylus (ssb.) tibialis (HAHN, 188 1) = H.tibialis mediferraneus Srygnocoris fuligineus (GEOFFROY, 1785) WAGNER,1962 T - La Esperanza, 22.X.86 ESCOLAleg., RIBES der., Coll. Mus. Zool. Barc. G - "La Gomera",l/O 4.V.86, FERRER ieg.,RIBES deí.er colf. The above synonymy is discussed in a paper (in print) by RIBES. Family P en t a t o m i d a e Subfam. P o d o p i n a e Lindbergopsallus hyperici (LINDBERG, 1953) H - supra Frontera,19.IV.91 a series on Hypericum canariense, HEISSleg. Venfocoris modesfus (JAKOVLEV, 1880) ‘90 Boietim do Museu Municipal do Funcbal No. XLíV, Art. 238 1992 Heiss & Ribes, Additions to the Heteroptem-fauna of the Canary 1s. 1 91

Lindbergopsallus instabilis (REUTER, 1904) P - Breña Baja,IV.88 and Sta.Cruz,VI1.89.,HEISS leg. ’ H - Valverde,0/2 19.1V.91 on Lavandula canariensis, together with the T - El Médano,FRANZ leg.,Puerto de la Cruz and Los Cristianos, HEISS leg.,all endemic Heterogaster canariensis Lbg., HEISS leg. PBRICARTdel.

Lindbergopsallus laureti (LMDBERG, 1936) N’sius (s-str.) cymoides (SPINOLA, 1840) H - Ladera Mauricio,l/O 14.V.84, OROMi leg., RIBES det.et coll. H - supra Frontera,19.IV.91., HEISS leg. PÉRICART der. Records for the following islands are now confirmed (PÉRICART Brachysteles wollasroni BUCHANAN WHITE, 1879 det.,coll.HEISS) C - Barranco Oscuro,O/l 8.1.88., OROMf leg., RIBES det.ef coll. P - Breña Baja,IV.88,VI.89 HEISSleg. G - La Caieta,IV.72 KLIMESCH leg. Xylocoris contiguus WAGNER, 1954 T - Cañadas IV.88, La Orotava, Los Cristianos, HEISS leg. Las Cañadas, a P - Pico de la Cruz,l/O 1.111.87, OROMf leg., RIBES det.et coll.; Roque de los series 26.111.64, RIBES leg.,WAGNER der., coll. RiBES et Univ. La Laguna. Arenas Muchachos,O/l 1.III.87.,OROMf leg., RIBES det.,coll. Univ. La Laguna. Negras,28.V.83,MORALES leg., RIBES der. el coll. C - Sta.Lucía,Artenara,Tejeda, HEISS leg. Peiraies strepitans (RAMBUR, 1842) F - Morro Jable, 28.XI.87,de FERRER leg., PBRICARTder., RIBES coll. F - Presa de los Molinos,0/2 1.111.90, OROMt leg., RIBES der., coll. Univ. La L - Playa Blanca III.90., HEISS leg. Laguna et RIBES. Geocoris (Eilatus) chloroticus PUTON, 1888 = G. nigriceps REUTER, 1891 parvula (FALLÉN,1807) T - La Laguna,4.111.77., OROMf leg., RIBES der., coll. Univ. La Laguna. P - Cumbre Nueva,18.XII.82, OROMt leg., RIBES det.et coll. Geocoris (s.str.) pubescens (JAKOVLEV, 1871) Galeatus scrophicus SAUNDERS, 1874 H - Mirador de Jinama,1000m.18.IV.91., HEISS leg. L - Playa Blanca,O/l 1.-6.III.90, HEISS leg. Platyplax inermis (RAMBUR, 1842) Tingis (s.str.) maderensis (REUTER, 1890) . L - Playa Blanca,2/3 1.-6.111.90., HEISS leg. H - Mirador de Jinama lOOOm 18.1V.91 a series on Carduus sp., HEISS leg. Neocamprotelus aeonii (LINDBERG, 1953) Tingis (Tropidocheila) lituraia (FIEBER, 1844) ’‘ H -Mirador de Jinama, 1000 m 114 18.1V.91 under low vegetation ofMicromeria L - Playa Blanca,l/l 1.-6.III.90., HEISSleg. . sp. and Lotus sp., HEISS les, coll. HEISS and RIBES. G - Hermigua,O/l 28.XI.85, OROMf leg., RIBES det. et coll. Aneurus avenius tagasastei ENDERLEiN, 193 1 The specimens mentioned above differ from LMDBERG’Sdescription by H - supra Frontera,UI 2L 19.IV.91., under loose bark of Myncafqa, HEISS leg. antennal segment 11 shorter than 111 + IV (equal to 111 + IV) and by dark antennal segment 1 (yellowish-brown),sharing the other characters given.Comparison with Beryfinus (Lizinus) geniculaíus HORVATH,1895 iypes is necessary to clear the range of variability. L - La Geria,O/l 1.-6.III.90., HEISS leg. Oxycarenus lavalerae (FABRICIUS, 1787) Nysius (s.str.) ericae (SCHILLING, 1829) L-Playa Blanca,LosValles, 1.-6.111.90., HEiSS feg.Famara, XII.91.. HANDKE L - El Golfo, 1.-6.111.90 a small series, HEISS leg. PÉRICART det. leg. Confirmed material is further from:

, 1992 Heiss & Ribes, Additions to the Heteroptera-fauna of the CaMiy 1s. 1 93 92 Boletim do Museu Municipal do Funcha No. XLIV, Arí. 238

f% Acrosrernum rubescens (NOUALHIER,1893) Aphanus rolandri (LINNAEUS, 1758) El Golfo,2/0 1.-6.III.90 on Euphorbia regis-jubae, HEISS leg. Costa del F Jandia,O/l 19.11.71., SANTOS leg., RiBES det.,coll. Univ. La Laguna L - - Rubicón,O/l 25.11.91., GUSENLEITNER leg., HEISS coll. Beosus maritimus (SCOPOLI, 1763) Mecidea lindbergi WAGNER, 1954 H - Valverde, 19.IV.91., HEISSleg. L - Famara,Xl1.9l.,HANDKE leg. Emblerhis duplicarus SEIDENSTOCKER, 1963 Nezara viridula (LINNAEUS, 1758) P San Andrés,1/0 3.XI.77., FERNANDEZleg., IUBES der.et coll. - L - Playa Blanca,O/l.21.11.90., GUSENLEITNER leg., HEISS coll. Emblethis denticollis HORVATH,1878 Piezodorus puncíipes PUTON, 1889 G El Cedro, 1/0 12.VIII.77., OROM~leg., RIBESder. et coll. Specimen without - H Mirador de la Peña,l/O 19.1V.91 on Teline sp., HEISS leg. anterior lateroventral pronotal dilatations. - T Los Baldíos,O/l 20.X.8 1 .SANCHEZleg. Specimen with pronotal dilata- - Aethus pilosulus (KLUG. 1845) tions. Verified record. L - Playa de Famara,O/l6.11.89., OROMf leg., RIBES det.,coll. Univ. La Laguna Lamprodema maurum (FABRKIUS, 1803) 2.4 Species inquirenda 5 - MoJónBlanco,0/2 25.XI.88., OROMf leg.;Dunas de Orzola, 7.II.88., RIBES der., coll.Univ. La Laguna Here we list those species which have been cited in the literature from the “Canary Islands” and those most probably assigned to a wrong taxon. In both cases Peritrechus gracilicornis PUTON, 1877 neither the source material nor the presence on the islands could be verified, as no P - Los Tilos,24.IX.75., FERNANDEZleg., RiBES det.et coll. subsequent records are known.Their occurrence on the Canary Islands remains doubtful dessfume investigations show that these taxa really belong to the insular Proderus subetythropus (COSTA, 184 1) fauna. Meanwhile we propose,that these taxa should not be regarded as to belonging P Santa Cruz de la Palma,17.VIII.81., RAFAEL GP leg.,RiBES det.et coll. - to the fauna of the Canaqr Islands. Cercinthus lehmanni (KOLENATI,1856) Some of such species inquirenda were already listed in the preliminary catalogue and comprise mainly ancient citations e.g. by the first entomologist T Las Caletillas,l/O 11.11.82., AMAR leg., RIBES det.et coll. - reporting Heieroptera from the Canary Islands,such as M. BRULLÉ(1838) and other authors,who have later been repeated by recent authors without further investigation. Camptopus laierafis (GERMAR, 18 17) Their taxonomic assignment is very doubtful and no recent material has been found. G - El Cedro,O/l 5.V.86., FERRER leg., WBESdeí.et coll. The following species are concerned (page numbers refer to HEISS & BAEZ, 1990): Liorhyssus hyalinus (FABRICIUS,1794) Lygus pratensis (285) L Playa Blanca, El Golfo, 1.4.III.90 common underhalophytic plants, HEISS leg. - Aellopus syriacus (298) Lamproplax piceus (299) introduced species Srictopleurus ribesi G6LLNER-SCHEIDMG, 1975 Cenírocoris spiniger H El Golfo,Las Casitas,2/0 19.IV.91., HEISS lfg. (300) - Cercinthus eiegans (301) Carpocoris mediterraneus (303) Odontoscelis dorsalis (FABRICIUS, 1803) Eurydema venrrale (303) see REMANE & KAMMERSCHEN, 1991a L Playa Blanca,0/9 1.-6.III.90.in swimming pool, HEISS leg. - Holcostethus vernalis (303) .

94 Boietim do Museu Municipal do Funchai No. XLIV, Art. 238

Further doubtful literature citations: Tuponia (sstr.) tamaricis (PERRIS,1857) STlCHEL (Bd.11,1956); CARVALHO (1958); WAGNER (1970-75) Family C o r i x i d a e “CM~Islands” Subfam. C o r i x i n a e Distribution not clear sec.WAGNER (1975:432).The citation by STICHEL may refer to PüTON (1889) and LiNDBERG (1936),but this reference has later been Corixa punctata (ILLIGER, 1807) assigned to Tlefhierryi(REUTER, 1875) by LINDBERG (1953) BRULLÉ (18389; LINDBERG (1953); NIESER 8z MONTES (1984). Family R e d u vi i d a e C.geofioyiLEACH,LINDBERG 1936. Subfam. E m e s i n a e This species is not included in the Canarian fauna by BAENA &Bm(1990), but is reported fkom Madeh by HOBERLANDT (1967) Ploiaria domesrica SCOPOLI, 1786 DISPONS (1955,1960); STICHEL (1955-62); but not recorded from the Canary FamilyMiridae ’ Islands by WYGODZINSKY(1966) “Canary Islands” Miris fuscicornis BRULLÉ,1838 BRULLÉ(1838); LINDBERG (1936), OSHANIN (1906-1909,1912);CARVALHO Tinna grassalor (PUTON, 1874) (1959) WYGODZINSKY (1966) “Canary Islands” Although the first citation has been repeated several times, L~DBERG(1936) Cerascopus grassaforNOUALHIER, 1893 From C,,Arganiguin.This citation is has stated,that the identity of this and the following species could never have been mentioned with a 7 under Pioiaria canariensis NOUALHIER,1895 = I! chiiensis cleared.We propose that both names should be deleted from the list. PHILIPPI, 1862 by LINDBERG (1953). WYGODZINSKY loc.cit:211 states that the record given by NOUALHIER is Miris parvula BRULLÉ,1838 aimost certainly secondary also. See comments of M.fuscicornis Family Lygaeidae Subfam. P h y 1 i n a e Subfam. L y g a e i n a e

Conostethus roseus (FALLEN, 1829) Lygaeosoma sardeum eryrhropterum PUTON, 1876 CARVALHO (1 958) SLATER (1964),as ssp. “Canary Islands” L. relicuiatum var.e.NOUALHIER 1893; LINDBERG (1934) As several citations of the nominal form are known from P, G, T and C it is to Plagiognathus (s.str.) arbustorum (FABRICIUS, 1794) be verified,if ssp.erythroprerum does coexist in T. STICHEL (Bd.I1,1956); CARVALHO (1958) “Canary Islands” Subfam. O r s i 1 i n a e Psaiius (Iiops)puncticoiiis FIEBER, 1870 WAGNER (1970-75, Bd.11: 178) Nysius (Macroparius) graminicola (KOLENATI, 1846) The author indicates that this species is supposed to occur on the Canary SLATER (1964), following MANCINI (1956) Islands also,but there is no evidence so far. “Canary Islands”

e O l 1992 Heiss & Ribes, Additions to the Heteroptem-fauna of the Canary 1s. 1 97 96 Boietim do Museu Mdcipai do Funcbal No. XLIV. Art. 238

ancient citations of that region are very doubtful. Nysius (Tropinysius) configuus (WALKER, 1872) SLATER (1 964), following OSHANIN (19 12), who included Madeira, the type locaiity, into Canary 1slands.Very doubtfu1,as N-configuusseems to be endemic to Family P e n t a t o m i d ae Madeira proper. . Subfam. P e n t a t o m i n a e Sciocoris (Neosciocoris) conspurcafus Nysius (Anorfhuna)aflanfidum H ORVATH, 1.890 KLUG,1845 STICHEL SLATER (1964), following again OSHANIN (1912), who obviously included (1955-62) The citation Islands” is doubtful for this eremean species. the Azores,the type locality,in the Canary Islands. Most probably an erroneous “Canary citation. Most possibly confused with other species (S.fissur MULSANT et REY or S. conspurcatus majusculus LRWAVUORI,1964 = carayoni WAGNER,1965 sec. LINNAVUORI et RIBES SAULEDA Subfam. R h y p a r o c h r o m i n a e )1975) 62 (1979)

Esuridea lathridioides PUTON, 1889 (maculafa,,1890) 3. E R RA TA SLATER following OSHANIN BLOETE and LINDBERG (1964), (1912), (1929) They refer to the published preliminary catalogue and should be changed as follows: (1936). A Madeiran species, see comments above. page 28 1 preliminar catalog - preliminary catalogue (2x) Family C o re i d a e 28 1 Josef-Schraffe-strasse - Josef-Schraffl-Strasse Subfam. P s e u d o p h 1 o e i n a e (Arenocorinae sec.VAZQUEZ,1987) 28 1 been recording - been recorded 286 Phyfodorus refamae - Phyfocoris refamae 28 8 Compsidolon (Coniorf odes) falleni (SCHILLING, 1829) VAZQUEZ 1987 hierroensis - hierroense This reference is obviously based on the record given by BRULLÉ(1838). 296 Ischnorrhynchinae - Ischnorhynchinae 296 Kleidoceris Kleidocerys Already LINDBERG (1953) had serious doubts about the correctness of the determi- - nation and regarded it as the common A. walfli. 298 veryfied -. verified 301 Psedophloeinae - 305 hirficornis (FABRICIUS, Actorrhinella - Aetorrhinella 1794) 305 (alluadi,Carpocoris) - alluaudi VAZQUEZ 1987 (alluadi,Dolycoris) alluaudi Also based on a reference by BRULLÉ1838 and is regarded as the common 305 - alluadi,Eudolycoris alluaudi CaSfinis by LINDBERG 1953. 305 - 3 12 strepifans,Pirates - Peirafes Family R ho p al i d ae Subfam. R h o p a 1 i n a e ACKNOWLEDGMENTS For presenting their interesting material we thank M. and P. ASHMOLE Sficfopleuruscrassicornis (LINNAEUS,1758) (Edinburgh), K. G. BEWHARDT (Osnabrück), J. COMAS (Barcelona), J. DE FERRER NOUALHIER (var. anlicus Rey); GOLLNER-SCHEIDING (1893) (1975); (Algeciras), H. FRANZ (Mbdling), J. GUSENLEITNER (Linz), M. MORALES (Santa VAZQUEZ (1 987) Cruz de Tenerife), T. YÉLAMOS (Barcelona) and are particularly grateful to P.OROMI According to GOLLNER this is an Euro-siberian species which does not occur (La Laguna). in northern Africa and the southern Mediterranean region. She states therefore,that 98 Boletim do Museu Municipal do Funchd No. ?UN,Art. 238 1992 Heiss & Ribes, Additions to the Heteroptera-fauna of the Canary 1s. I 99

REFERENCES GOLLNER-SCHEIDING,U.: 1975. Revision der Gattung Stictopleurus STAL, 1872 (Heteroptera,Rhopalidae). - Dtsch. Works which we refer to in the text and are not listed here, were already ent. Z..N.F., 22(1-3):1-60 included by HEISS & BAEZ (1990). 1980. Einige Bemerkungen zu den Gattungen Corizus Fallen,1814 und Xenogenus Berg, 1883 (Heteroptera, Rhopalidae).- Mitt.zoo1.M~~.Berlin,56( 1): 1 1 1-12 1 1986. Revision der Gattung Odontoscelis LAPORTE DE CASTELNAU,1832 (Heteroptera, ALLEN,R.C.: Scutelleridae). Dtsch. ent. 1-2):95-127 1969. A Revision ofthegenus Leptoglossus Guerin (:).- Ent.Amer.,45:35- - Z..N.F.,33( 140 1990. Revision der Gattung Odontotarsus LAPORTE DE CASTELNAU,1832 (Heteroptera: Scutelleridae). - Mitt. 2001.Mus. Berlin,66(2):333-370 ASHMOLE, N. P. & M. J. ASHMOLE: 1988a. hectdispersa1 on Tenerife,Canary 1slands:High altitude fallout and seaward drift.- GOMEZ-MEN0R.J.M.: Arclic and Alpine Research,20 (1): 1-12 1956. Antocóridos de España y Marruecos.- Inst.Est.Afr.,Madrid,122 pp.

1988b. communities supported by biological fallout on recent lava flows in the GÜNTHESH. & G. SCHUSTER Canary Islands. Ent. scand. Suppl. 3267- - 1990. Veneichnis der Wanzen Mitteleuropas (Heteroptera).-Dtsch.ent.Z.,N.F.,37(4-5):361- 1989. Natural History Excursions in Tenerife.A Guide to the Countryside,Plants and Ani- 396 mals.- Kidston Mil1 Press, Peebles, Scotland, 252 pp. HEISS,E. & M.BAEZ. 1990. A preliminary catalogue of the Heteroptera of the Canary Is1ands.- Vieraea 18:281- ASHMOLE,N.P.,ASHMOLE M.J. & P.OROMI: 3 15 1990. of recent lava flows on .- Vieraea 18: 17 1 - 187.

BAENA,M. & M.BAEZ: HOBERLANDT,L.: 1990. Los Heterópteros acuáticos de las Islas Canarias (Heteróptera; Nepomorpha, 1967. On some species of Heteroptera collected in the Madeiran IsIandsActa Faun. Ent. Gerromorpha).- Vieraea 19:233-244 Mus. Nat.Pragae, 12(117): 43-49 1987. Results of the Czechoslovak-Iranian entomological expeditions to Iran 1970,1973 CARAPEZZA,A. : 1988. Cyrtopeltis (Canariesia nmbgen.) salviae nsp. delle Isole Canarie (Heteroptera, and 1977. Heteroptera, Lygaeidae,Oxycareninae.-Acta Ent. Mus. Nat. Pragae, , Dicyphinae). - Fragm.Ent.Roma, 20 (2):137 -142 42:1 1-29 KANYUKOVA,E.V.: CARAYON,J.: 1988.23.Fam. Reduviidae in: VINOKUROV el al.:Keys for the identifícation of the 1989. Systématique et biologie des Kleidocerys d'Europe (Hem.Lygaeidae).- Eull. Soc. ent. Fr., 94 (5-6):149-164 ofthe Far East of USSR.2 1 .Order Heteroptera (Hemiptera)(in Russian).- Akad. Nauk SSSR,Leningrad,II: 869-873 CARVALH0,J.C.M.: KERZKNES1.M.: 1957-1960. Catálogo dos Mirídeos do Mundo.- Arquivos do Museu Naciona1,Rio de 1963. Einige Nabinae (Heteroptera.Nabidae) aus der Sammlung des Ungarischen Janeiro, Vol. XLIV, 1957 (Part I);Vol.XLV, 1958 (Part- _. 1I);Vol.XLVII. 1958 (Pari Naturwissenschaftlichen Museums in Budapest.- Ann. hist. -nat. Mus. nai. hung., lIl);Vol.XLVlII, 1959 (Part IV); VoLLI, 1960 (Part V) 55:457-461 DISPONS,P.: 1976. New and little-known Heteroptera from Mongolia and from adjacent regions of the 1955. Les Réduviidés de I'Afrique nord - occidentale.Biologie et Biogéographie. - Mém. USSR.111 (in Russian).- Nasekom. Mong.,4:87- 1 14 Mus. nat. Hist.nat. ,Paris,sér.Zool.lO (2):93-240 1988a. New and little known Hemiptera (Insecta,Heteroptera) from the Far East of USSR 1960. Commentaires sur la famille des Reduviidae. - Rev. Fr. d'Ent., XXVII(4):291-305 (in Russian). - AcadSci. USSR, Vladivostok, 1987,pp 1-83

! Boleiim do Museu Municipal do Funcbal No. XLN, Art. 238 1O0 1992 Heiss & Ribes, Additions to the Heteroptera-fauna of the Canary 1s. 1 101

1988b. Revision of the Nabidae (Heteroptera) mentioned in LMDBERG's papers on the REMANE& & KAMMERSCHEN,D.: fauna of the Eastem and Southem Pa1aearctic.- Ann.Enf.Fenn.,54:135-137 199la. Zur Besicdlung der Mittelatlantischen Inseln (Kanaren,Madeira,Azoren) durch Taxa 1988c. 2 1.Fam.Miridae (Capsidae) in: VINOKUROV et al.:Keys for the identificationofthe der Gattung Eurydema LAP. (Heteroptera, Pentatomidae).- Marburger Eni. Publ., insccts of the Far East of USSR.2i.Order Heteroptcra (Hemiptcra)(in Russian).- 2 (5) :25-57 Akad. Nauk SSSR, 4,Leningrad. ik778-857 1991b. Zur Verbreitung und Artabgrenzung der Taxa der Gattung Dolycoris M. R. (Hctcroptcra, Pentatomidae, Pentatominae) in der Südwestpaliiarktis,insbesondere LINDBERG,H.: auf den Mittelatlantischen 1nseln.- Marburger Ent.PubI., t(5): 58-1 12 196 1. Hemiptera lnsularum Madeirensium.- Commentat.Bio., 24: 1-1 10 RIBESJ.: LINNAVUOR1,R.. 1973. Sobre Empicoris oalinus (LINDBERG) 1932 (Hem.Het. Reduviidae Emesinae).-Misc. 1964. Hemiptcra of Egypt,with remarks some species of the adjacent Eremian region.- on ZOO~.111(3):33-42 Ann. Zool. Fenn., i:306-356 1978. Un Mirido nuevo de las Islas Selvagens (Insecta,Heteroptera).- Conrrib. est. Hisr. 1972a. Studies on Palaearctic Hemiptera.- Ann.Ent.Fenn., 38 (1):40-50 Nat. Islas Salvajes/Mus.Cien.Nat.Tenerife, 1 I.pp 139-144 1972b. On the of the genus Geocoris Fn. (Het.,Lygaeidae).- Ann. Ent. Fenn., 38 (2):lOO-106 RIBESJ. & N.SAULEDA: 1979. Heterópteros de Alicante y zonas adyacentes.- Mediierrtinea, 3: 123-158 1975. Hemiptera of the Sudan,with remarks on some species of the adjaccnt countries 5. Pentatomidae. - Bo1.Soc.Port. Citnc. Nat.15:5- 128 SCHM1TZ.G.: 1976. La faune terrestre de I'Ile de Sainte-Htlbne,20.Heteroptera.-Ann.Mus. Roya1 Afr. 1986. Heteroptera of Saudi Arabia.- Fauna of Saudi Arabia, 8:31-197 Centr., Serie in 8°,Zool.Sci.,215:3 19-533

MANCIN1,C.: SCHüH,R.T. & STYS,P.: 1956. Emitteri Eterotteri del Gargano e delle Tremiti con osservazioni sulle specie a 1991. Phylogenetic analysis of Cimicomorphan family relationships (Heteroptera).- J. New distribuzione transadriatica.- Mem.Biogeografia Adriatka,IIi: 16 1-195 York Entomol. Soc. ,99(3):298-350

MOULET,P.: SCUDDER,G.G.E.: 1991. Notes,typifications et remarques sur quelques (Hemiptera,Heteroptera). - 1962. "he World Rhyparochrominae (Hemiptera:Lygaeidae).I.New Synonymy and Generic Rev. fr. Ent. (N.S.), 13(2):49-64 \ Changes.- Can.Eni. ,94(7):764-773

OSHANIN,B.: SEIDENSTOCKER,G.: 1906- 1909. Verzeichnis der Palaearktischen Hemipteren mit besonderer Beriicksichtigung 1964. Zur Systematik von Bledionotus, Bethylimorphus und Thaumastella HORVATH ihrer Verteilung im Russischen Reiche.- Ann.Mus.Zool.Ac.Imp.Sc. St.Petersburg, (Heteroptera, Lygaeidae). - Reichenbachia 3(25):2 69-279 Beilagen zu Bd.XI-XIV, 1087 pp. TAMANIN1,L.: PÉRICARTJ. & RIBESJ.: 1959. Caratteri generici di Dolycoris Muls. et Rey e Eudolycoris nov. gen. con tavola 1992. Quelques nouvelles synonymies et dtsignation de lectotypes dans le genre Geocoris dicotomica delle cntitá della sottoregione mediterranea (Heteroptera, Pentatomidae) (Heteroptera,Lygaeidae,Geocorinae).- Rev.fr. Ent. ,(N.S.),14(2):77-8 1 - Mem. Soc.Ent.ltal.,38: 73-83

PUTSHK0V.P.V.. VAZQUEZ,M.A.: 1987. Fauna Ukrainiy, 21.Heteroptera. 5.Reduviidae (¡u Russian).- Ahd. Nauk Ukrain 1987. Inventario de los Coreoidea, 1910 de la región paleártica occidental (Hemiptera, .SSR,Kiev,248 pp. Heteroptera). - Bol.R.Soc.Esp.Hist.Nat.(BioI.),83(1-4):229-247

PUTSHKOV,V.G & P.V.PUTSHKOV: V1DALJ.P.: 1985. A catalog of the ,Assasin-bugs of the World (Heteroptera,Reduviidae).- Akud. Nauk 1938. Contribution P I'étude des Hérnipteres du Maroc (2" note).- Bull. Soc. Hist. Nar. Afr. CIkrain.SSR,Kiev, 138 pp. Nord, XXIV