27 June 1995 PROC. ENTOMOL. SOC. WASH. 97(2), 1995, pp. 379-395

EUROPIELLA REUTER (: ): RECOGNITION AS A HOLARCTIC GROUP, NOTES ON SYNONYMY, AND DESCRIPTION OF A NEW SPECIES, CARVALHOI, FROM NORTH AMERICA

RANDALL T. SCHUH, PER LINDSKOG, AND I. M. KERZHNER (RTS) Department of Entomology, American Museum ofNatural History, New York, New York 10024, U.S.A.; (PL) Department ofEntomology, Swedish Museum ofNatural History, Stockholm S- 104-05 Sweden; (IMK) Department ofInsect , Zoological Institute, Academy of Sciences, St. Petersburg, Russia 199034.

Abstract. -The subgenus Poliopterus Wagner of Fieber is placed in syn- onymy with Europiella Reuter. The identity of Holarctic species is clarified. New com- binations are created because of generic synonymy, species are transferred from other genera, and many names are placed in synonymy. Most species for which hosts are known feed on Artemisia, a few feeding on other Asteraceae such as Chrysothamnus, Helichrysum, and Tanacetum, with two species being recorded from the Lamiaceae. Key Words: Heteroptera, Miridae, Europiella, new species, new synonymy, Holarctic

Reuter (1909) used the name Europiella the history of synonymy for the Nearactic for two species of phyline Miridae from fauna, as well as more detailed distributions North America. Since that time many ad- of the species, can be found in Henry and ditional American species have been placed Wheeler (1988). in the genus. Wagner (1949) described the This paper is presented in honor of our subgenus Plagiognathus (Poliopterus) to long-time colleague and friend Jose Can- contain several species of Palearctic Phyli- dido de Melo Carvalho. His influence on nae related to P. albipennis (Fallen). Our the study ofthe Miridae has been profound, individual research efforts have shown that because of his monumental world catalog many species placed in Europiella and Pla- and his unparalleled descriptive efforts, par- giognathus are in fact congeneric, or in a ticularly on the Neotropical fauna. few cases conspecific, that many species from We thank Thomas J. Henry and Michael North America that have previously been D. Schwartz for reviewing the manuscript. placed in Europiella do not belong there, and some Palearctic species placed in Pla- Europiella Reuter giognathus sensu stricto and Chlorillus ac- tually belong to Europiella. There is much Type species. -Agalliastes stigmosus confusion regarding the correct application Uhler 1893. of species-group names, including substan- Europiella Reuter 1909: 83 (n. gen., desc.); tial synonymy. In the following pages we Knight 1968: 37 (key to spp.). provide solutions to these problems and de- Poliopterus Wagner 1949: 53 (n. subgen. of scribe a new species. Additional details on Plagiognathus, desc.). NEW SYNONYMY. 380 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON

Diagnosis.-Relatively small, length 2-5 stricto, but some species such as Plagiog- mm; some spp. sexually dimorphic, body nathus alpinus, have been placed in Chlor- form varying from elongate and nearly par- illus or by some authors. The struc- allel-sided to broadly ovoid, especially in ture of the genitalia indicates that they females. Coloration varying from almost should be placed in Europiella, as does com- completely black to almost completely green parison with some North American species, or white, usually with a few dark markings which are similar in coloration. All but two at the bases of spines on the legs. Vestiture of them feed on Artemisia. of dorsum with reclining simple setae and In the New World there are several spe- recumbent, woolly, sericeous setae; head cies that are very closely related to albipen- weakly produced and not overlapping an- nis, and a number ofothers that we treat as terior margin of pronotum. Male genitalia congeneric. Among these is stigmosa (Uhl- varying greatly in size, but pygophore al- er), the type species of Europiella Reuter ways large relative to total size ofabdomen; 1909. We are therefore treating Poliopterus right paramere truncate apically, with a more Wagner as the junior synonym of Euro- or less well-developed protrusion on either piella. The male genitalia of stigmosa are side, never lanceolate in form; vesica always shown in Figs. 12-14 and those of albipen- with two apical spines, these sometimes nis in Fig. 16. elongate and broadened basally (Figs. 12, The generic limits ofPlagiognathus are a 13), or much shorter and variously acumi- subject that can be properly addressed only nate and/or strongly curving (Figs. 6, 7, 9, with a much more wide-ranging analysis of 10, 15-17); secondary gonopore placed to the Holarctic , something that we do the side ofthe chitinous bands ofthe vesica not attempt in this paper. Because subgen- rather than between them and distinctly era have been used primarily in the Pale- proximad of attachment of spines. Often arctic, a consistent treatment of the world breeds exclusively on Artemisia, occasion- fauna including the use of Palearctic sub- ally on other Asteraceae, and rarely on other generic concepts would leave many species plant families. unplaced as to subgenus. It is for these rea- Discussion. -Wagner (1949) described sons that we treat Europiella as a distinct the subgenus Poliopterus, with albipennis genus. (Fallen) as the type. He consistently placed Knight(1968, 1969, 1970)diagnosedEu- it in the genus Plagiognathus Fieber by vir- ropiella without examining the structure of tue ofits members having two elongate chi- the male genitalia. He included some spe- tinous spines apically on the vesica and with cies which are clearly congeneric with stig- the secondary gonopore rather distant from mosa, the type. He also included many oth- them. Within Plagiognathus, Wagner rec- er species (as enumerated by Henry and ognized Poliopterus by the presence ofonly Wheeler 1988) which bear a superficial re- light colored setae on the dorsum. Wagner semblance to stigmosa and its congeners. (1975) placed 13 species from the Mediter- These species are found in desert areas of ranean Region in Poliopterus; the majority western North America feeding primarily of those for which hosts are known feed on on Atriplex, Sarcobatus (Chenopodiaceae), species ofArtemisia. and Lycium (Solanaceae), but nearly all of Members of another assemblage of Pa- them belong to a separate lineage. Com- learctic species, richly represented in the Far parison ofpreparations ofthe male genitalia East (Kerzhner 1 988a) and including one of paratypes of most of these species and European species, are green and dissimilar the genitalia of Megalopsallus atriplicis to albipennis in appearance. They were tra- Knight, the type species of Megalopsallus ditionally placed in Plagiognathus sensu Knight, indicates that most, if not all, of VOLUME 97, NUMBER 2 381

92 ---7

1~~~~~~~~~~~~1

15 16 17

13 14

Figs. 1-17. Male genitalia of Europiella spp. 1-5. E. carvalhoi. 1. Vesica. 2. Detail of apex of vesica. 3. Obverse view of distal portion of vesica. 4. Right paramere. 5. Left paramere. 6-8. E. decolor (western North America). 6. Vesica. 7. Apex of vesica, rotated 90 degrees. 8. Right paramere. 9-11. E. artemisiae (western North America). 9. Vesica. 10. Apex of vesica, rotated 90 degrees. I 1. Right paramere. 12-14. E. stigmosa. 12. Vesica. 13. Obverse view ofdistal portion of vesica. 14. Right paramere. 15-17. Comparative views of apex of vesica of Holarctic species. 15. E. artemisiae (Europe). 16. E. albipennis (Europe). 17. E. decolor (Europe). 382 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON them are congeneric. We provide below a Agalliastes tibialis Fieber 1864: 228 (n. sp., list of all names for which new combina- desc.). tions are formed in Megalopsallus. Agalliastes lanuginosus Jakovlev 1875: 172 Some species which are clearly congeneric (n. sp., desc.). with stigmosa have often been placed in Plagiognathus albipennis var. tibialis: Ja- othergenera. We have transferred all ofthose kovlev 1877: 279 (syn.). of which we are aware. Plagiognathus albipennis: Reuter 1878: 175 In the Holartic there are Artemisia-feed- (part; descr., variability, syn.). ing species placed in other genera of Phy- Plagiognathus collinus Wagner 1941: 249 linae (e.g. species placed in Phyllopidea (n. sp., desc., figs.). NEW SYNONYMY. Knight) as well as in other subfamilies of Plagiognathus arenicola Wagner 1941: 252 Miridae. Critical examination of these spe- (n. sp., desc., figs.). NEW SYrNONYMY. cies indicates that although the host pref- Plagiognathus (Poliopterus) collinus: Wag- erence of Europiella spp. is helpful in be- ner 1952a: 197 (key, desc., figs.). ginning to establish a basis for their generic Plagiognathus (Poliopterus) arenicola: identity, it is often only with examination Wagner 1952a: 197 (key, desc., figs.); ofthe male genitalia that generic placement Wagner and Weber 1964: 413 (key, desc., and specific identity can be determined with figs.); Wagner 1975: 35 (key, desc., figs.). certainty. Plagiognathus (Poliopterus) lanuginosus: We have organized most ofour treatment Josifov 1974: 14, 20 (restored from syn- on geographical grounds, because the per- onymy, list, host); Wagner 1975: 34 (key, tinent literature is organized on that basis. desc., figs.). Distribution.-From southernmost Swe- IDENTITY AND SYNONYMY OF den, southeastern Finland, and Leningrad EUROPIELLA ALBIPENNIS AND Two Province, Russia, at least to Bulgaria and RELATED SPECIES from France east at least as far as East Ka- Three closely related species ofEuropiella zakhstan. are widely distributed in the Palearctic, two Hosts.-Living exclusively on Artemisia of these also occurring in the Nearctic. No campestris in Northern Europe; records of other species ofthe genus are Holarctic. All Plagiognathus lanuginosus from A. mariti- three of these species have been identified ma in southern Europe probably refer to E. by various authors as Plagiognathus albi- decolor. pennis. We found that these taxa are reliably Notes. -In this species the dark spots on distinguished from one another by the form the hind tibia are usually larger than in re- of the vesical appendages as seen in dorsal lated species and the venter is black in both view (e.g. Figs. 15-17). The size and color sexes, even though the dorsum may be pale. are highly variable, but in some regions the The apex of the vesica is shown in Fig. 16. species can be recognized by external ap- Fallen (1829) described Phytocoris albi- pearance. pennis from four specimens, including male(s) and female(s), all collected by Zet- Europiella albipennis (Fallen), terstedt in July. Type locality: Sweden, SkAne NEW COMBINATION Prov. (Scania), Esper6d. Extant material Fig. 16 qualifying as syntypes is present only in the Phytocoris albipennis Fallen 1829: 107 (n. collection of Fallen, not in the collection of sp., desc.). Zetterstedt, both collections deposited in the Agalliastes albipennis: Fieber 1861: 311 Zoological Museum, Lund University. It (key); Puton 1873: 25 (syn.). consists ofone male labelled "P. albipennis VOLUME 97, NUMBER 2 383

[male symbol]" and a small blue square, and the male as lectotype of P. albipennis and one female labelled "P. albipennis [female have labelled it accordingly. symbol]" plus a few illegible characters, and Wagner (1941) stated that the type spec- "Mellby 12 jun." on the underside of the imens were presumably lost and designated label. The labels are in Zetterstedt's hand- from his collection "Neotypen," saying writing, and the small blue square is known nothing about their number, sex, and lo- to signify that Zetterstedt collected the spec- cality. Wagner's "neotypes" are not con- imen at Esper6d, Mellby Parish. Among specific with the true types (see below) and these specimens is pin without any speci- Wagner's nomenclatural act is invalid be- men with Fallen's label "P. albipennis [male cause more than one specimen was desig- symbol] [female symbol] Esperod." nated as the neotype. According to Wagner (1941: 256), F. Os- From the examination of Fallen's speci- siannilsson informed him that the type was mens we have determined that the name no longer present in the collection ofFallen, albipennis should be ascribed to a species but rather that only an empty pin remained. which is not the most common in the Pa- Ossiannilsson in a letter to Lindskog (Jan. learctic and which does not occur in North 1 5, 1982) explained that his reply to Wagner America, in spite ofmany indications in the was based on the opinion of museum au- literature to the contrary. thorities that in general only specimens car- Agalliastes tibialis was described from rying Fallen's labels could be considered as Sarepta (now Krasnoarmeysk, part of Vol- his types. It is known that after the death gograd) from Frey-Gessner's collection. The of Fallen in 1830 his collection passed to type specimens were the possession of Zetterstedt, who notori- apparently examined filled in with by Puton and Reuter, and Reuter (1878, pl. ously any missing species spec- 4, fig. 8) published Fieber's figures of a fe- imens from his own collection. Ossiannils- male. We cannot son wrote (in litt.) "In the present case this locate the type(s). Judging was apparently fully justified, as I should from the large, black, tibial spots noted in have realized iftime had been taken to look the original description and the figure, and more closely on Zetterstedt's label and Fal- the pale dorsal surface and black abdomen, len's text. Fallen described albipennis based this species in a almost certainly identical on four specimens. He probably retained with albipennis. two specimens for his own collection which Agalliastes lanuginosus was described later were lost somehow, and returned two from specimens collected at Sarepta by to the collector Zetterstedt, who in turn Christoph and at Akhtubinskaya steppe by placed these in Fallen's collection after he Becker. The only known syntype is a badly acquired it. These two specimens should ac- damaged specimen (probably female) from cordingly be considered as syntypes." Sarepta in the Zoological Institute, St. Pe- As Fallen indicated that the types were tersburg. Its remnants include the head with collected in July, and because Zetterstedt in antennae, prothorax, and all legs. Judging his handwritten notes in his copy ofFallen's from these parts and from the original de- (1829) work (now in the library ofthe Zoo- scription, the species is identical with al- logical Institute, St. Petersburg), indicated bipennis. that he collected the species at "l[ocis] aridis Plagiognathus arenicola was described Esp[eroed] 18 jul. freq[uens]," it seems from Germany and Poland and P. collinus doubtful that the female labelled 12 June from Poland. We examined the types and belongs to the syntype series although no paratypes of both species in the Zoological such doubt exists for the male. Museum Hamburg and paratypes in other We accept these arguments and designate collections. Some paratypes were dissected 384 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON for comparison of the male genitalia, and Plagiognathus extremus: Wagner 1954: 77 these are conspecific with albipennis. (n. status, descr.). Reuter (1878: 81, 82) described several Plagiognathus (Poliopterus) litoralis f. abro- varieties of P. albipennis, assigning them tani Wagner 1949: 53 (n. form, desc.). Greek letters. Reuter did not indicate spec- NEW SYNONYMY. imens for his varieties and did not label Europiella bakeri: Carvalho 1955: 227 (n. varieties in his collections. Stichel (1934: comb.); Knight 1969: 86 (disc., redesc., 282) named these varieties assmanni, al- host). bella, beckeri, and antennaria. The identity Plagiognathus (Poliopterus) abrotani: Wag- of these names cannot be clarified at pres- ner and Weber 1964: 413 (new status, ent, and it is clear that Stichel did not base key, desc., figs.). his names on specimens which he himself Psallus artemisicola Knight 1964: 149 (n. examined. sp., desc., host). NEW SYNONYMY. Europiella nigricornis Knight 1968: 40 (n. sp., desc., host). NEW SYNONYMY. Plagiognathus (Poliopterus) larae Kerzhner Europiella decolor (Uhler) 1978: 46 (n. sp., desc., figs.). NEW Figs. 6-8, 17 SYNONYMY. ? Phytocoris pallidulus Dahlbom 1851: 21 1 Psallus albipennis (not Fallen 1829): Whee- (n. sp., descr.). Questionable NEW ler and Hoebeke 1982: 696 (n. comb., syn., SYNONYMY. figs.). Agalliastes decolor Uhler 1893: 380 (n. sp., desc.). Distribution. - Broadly distributed Agalliastes apiatus Uhler 1895: 53 (n. sp., throughout the Holarctic. desc.). NEW SYNONYMY. Hosts. -Neararctic hosts include Arte- Agalliastes signatus Uhler 1895: 55 (n. sp., misia californica, A. campestris, A. dracun- desc.). NEW SYNONYMY. culus, A. filifolia, A. ludoviciana, A. nova, A. bakeri Bergroth 1898: 35 (n. tridentata, and Chrysothamnus sp. In the n. for Agalliastes signatus Uhler). western Palearctic known from Artemisia Plagiognathus albipennis var. extrema absinthium, A. maritima, and A. abrotani Reuter 1901: 187 (n. var., descr.). NEW (in botanical gardens). Kurile Islands spec- SYNONYMY. imens are from A. schmidtiana, belonging Chlamydatus uhlerianus Kirkaldy 1909: 390 to the "Maritimae" series. (unnecessary n. n. for Agalliastes signatus Notes.-The name decolor (Uhler) has Uhler). previously been applied only to the North Plagiognathus decolor: Reuter 1909: 81 American fauna. Our studies indicate, how- (disc.). ever, that the taxon to which this name ap- Europiella decolor: Van Duzee 1916: 47 plies also occurs in the Palearctic. (list). Uhler (1893) indicated in his discussion Chlamydatus apiatus: Van Duzee 1917: 417 ofdecolor that he examined four specimens (n. comb.). from American Fork, Utah, as well as larger Psallus waldeni Knight 1923: 468 (n. sp.). specimens collected near Los Angeles, Cal- NEW SYNONYMY. ifornia. We located in the National Museum Psallus bakeri: Knight 1941: 43 (key, desc.). of Natural History, Washington, D.C., a Plagiognathus (Poliopterus) litoralis Wag- single female specimen from American Fork ner 1949: 53 (n. sp., desc., figs.); Wagner that seems to pertain to material originally 1952a: 199 (key, desc., figs.). NEW examined by Uhler. In addition to the data SYNONYMY. noted above it bears the labels "COTYPE VOLUME 97, NUMBER 2 385 (by Uhler) Agalliastes decolor Uh 1." [ap- species. Additional single male and female parently affixed by H. H. Knight], and "Co- specimens identified as this species were type No. 52835 U.S.N.M." found in the collections in Helsinki together Uhler (1895) described Agalliastes apia- with the lectotype. All of them are labelled tus from specimens collected at Fort Col- "Tokmak," but the male has Sahlberg's code lins, Manitou, and Steamboat Springs, Col- label indicating that it was collected at Con- orado. He indicated that it also occurred in stantinovskaya. Other cases ofconfusion in Kansas. The collections ofthe USNM con- labelling specimens from these two locali- tain 10 specimens associated with the apia- ties were also encountered. All specimens tus identification. One ofthese was labelled belong to E. decolor. as a cotype by H. G. Barber. It is clearly The identity of Phytocoris pallidulus congeneric with decolor. Dahlbom, 1851 is unclear. Reuter (1878) Uhler (1895) described Agalliastes sig- placed this name in synonymy with P. al- natus from a single male specimen from bipennis. Wagner (1941) treated pallidulus Manitou, Colorado. We were not able to as a valid species, but later (Wagner 1952a) find any specimens labelled as signatus after considered it as a pale variety of P. albi- careful checking of the USNM collections. pennis sensu Wagner. Phytocoris pallidulus Bergroth (1898) proposed the replacement was described from two specimens (appar- name bakeri for the preoccupied signatus. ently a male and a female) collected by Our use ofthe name bakeri is based on spec- Dahlbom at Stensuga, Gotland Island, Swe- imens identified by H. H. Knight, which are den, from Umbelliferae. The type speci- clearly conspecific with decolor. We there- mens are apparently lost; according to R. fore treat bakeri as a junior synonym. Danielsson (in litt.), Zoological Museum, Wheeler and Hoebeke (1982) correctly Lund University, they cannot be located in synonymized Psallus waldeni Knight with the collections of Dahlbom, Thomson, or what they called albipennis (Fallen), but our other pertinent collections. Dahlbom's onrg- examination ofspecimens and literature in- inal description is inadequate for identifi- dicates that the senior synonym is in fact cation and was apparently based on teneral decolor (Uhler), not albipennis (Fallen). specimens. At least the description can We examined type specimens of Plagiog- hardly be referred to E. albipennis or E. nathus litoralis (described from Germany artemisiae (albipennis sensu Wagner). and the Netherlands), P. abrotani (from the Lindskog was unable to find either of these Hamburg Botanical Gardens), and P. /arae species from Gotland Island either in Swed- (from Kurile Islands), as well as many ad- ish collections or through intensive personal ditional specimens, and found these nom- collecting. The only species of Europiella inal taxa to be conspecific with decolor from found on Gotland Island is E. decolor which North America. is abundant there on Artemisia absinthium. Plagiognathus albipennis var. extrema But, as Dahlbom's description may refer to Reuter 1901 (p. 187) was described from some other small Miridae, especially to ten- several specimens (length 1.75-2.00 mm) eral specimens, we prefer to consider Phy- collected by J. Sahlberg at "Constantinov- tocoris pallidulus as a possible synonym of skaya" (Konstantinovskaya on River Chu, E. decolor rather than treat it as a valid about 20 km N ofBeshkek, Kirgizia). Wag- species. ner (1954) examined a female from the Uni- Europiella artemisiae (Becker), versity Zoological Museum, Helsinki la- NEW COMBINAION belled "Tokmak" (about 60 km E of Besh- Figs. 9-1 1, 1 5 kek). He designated it as lectotype and con- Capsus artemisiae Becker 1864: 487 (n. sp., sidered Plagiognathus extremus as a valid desc.). 386 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON

Plagiognathus solani Matsumura 1917:432 er: three females (received in 1866), two (n. sp., desc.). NEW SYNONYMY. males (1872), and one female (1874); one Plagiognathus albipennis var. obscura Sahl- additional male collected by Becker was re- berg 1920: 167 (n. var., desc.). NEW ceived with the Jakovlev collection. All SYNONYMY. males belong to artemisiae (Becker) (albi- Plagiognathus albipennis (not Fallen 1829): pennis sensu Wagner). The male bearing the Wagner 1941: 248 (desc., figs.). number 13737 is designated here as lecto- Plagiognathus (Poliopterus) albipennis (not type. In the Naturhistorisches Museum Fallen 1829): Wagner 1952a: 198 (key, Wien, there are two males received from desc., figs.); Kerzhner 1964: 761 (key, Becker in 1870 and one female received in figs.); Kerzhner 1988a: 853 (key, figs.); Li 1869. They were examined by Reuter. One and Zheng 1991: 90 (key, dist., figs.). male belongs to artemisiae, the other to al- Plagiognathus (Poliopterus) gracilis Wagner bipennis. 1956b: 74 (n. sp., desc., figs.). NEW Plagiognathus solani Matsumura was de- SYNONYMY. scribed from Sakhalin and Hokkaido. The name was synonymized with albipennis Distribution. -This species is widely dis- sensu Wagner by Miyamoto (1977) and tributed in the Palearctic, in the north to should put in synonymy with artemisiae Scandinavia and Chukotka. Our examina- (Becker). Also the varietal name obscura tion of collections from Canada and the (Sahlberg) given to dark specimens from Western United States indicates that al- Finland should be placed here in synonymy. though this species has not previously been Plagiognathus gracilis was described from recorded from North America, it ranges Austria and Croatia. The type series was from Alaska south to montane areas of the examined by us and paratypes dissected for northwestern United States (detailed local- study of the male genitalia, indicating that ity records to be published separately). gracilis is a junior synonym. Host. -Palearctic hosts include Artemisia vulgaris, the related A. montana, and A. rub- NEARc-Ic SPECIES ripes, as well as other species. The only Europiella angulata (Uhler) known Nearctic host is Artemisia ludovi- angulatus Uhler 1895: 53 (n. ciana. sp., desc.). Notes. -This is the most common species Europiella angulata: Knight 1968: 41 (n. of Europiella in the Palearctic. Wagner comb., disc., dist., host). (1941) designated a neotype for what he Europiella yampae Knight 1968: 43 (n. sp., considered to be Plagiognathus albipennis. desc., host). NEW SYNONYMY. But because Wagner misidentified the spe- cies under discussion as Phytocoris albipen- Distribution.-Montane western United nis Fallen, his neotype(s) belongs to another States. species, the oldest available name for which Hosts. -Artemisia arbustorum, A. dra- appears to Capsus artemisiae Becker. cunculus, A. ludoviciana, A. tridentata (As- Capsus artemisiae was described from teraceae). specimens collected by Becker in 1862 at Notes. -Uhler (1895) mentioned having Sarepta in southern Russia, from a plant he examined "Only one specimen ... a male identified as Artemisiafragrans. This plant . . ." [ofangulatus]. The specimen here rec- species does not grow in the Lower Volga ognized as the holotype of angulata, in the region, so probably Becker collected from USNM, fits Uhler's original description, and another species of Artemisia. In the Zoo- bears the additional label "Maurodactylus logical Institute, St. Petersburg, there are six angulatus Uhl." specimens which were received from Beck- Examination of the male genitalia of the VOLUME 97, NUMBER 2 387 holotype of angulatus and paratype males Measurements: of E. yampae Knight from the type locality Apex indicates that the two are synonymous. tylus Inter- Width Length -cuneal Width ocular pro- antennal Europiella artemisiae (Becker), see above. fiacture head space notum segment 2 M 8 1.87-2.11 0.67-0.73 0.41-0.47 0.86-0.96 0.60-0.70 Europiella carvalhoi Schuh, F 8 1.65-2.07 0.66-0.74 0.43-0.49 0.82-1.00 0.50-0.68 NEW SPECIES Holotype.--male, USA: Wyoming: Big Figs. 1-5 Horn Co.: 27 mi. W Burgess Jct. on Rt. 14, Diagnosis. -Recognized by the light col- 7700 ft., Aug. 12, 1986, Schuh, Schwartz, oration ofthe dorsum with pale simple setae and Stonedahl; Artemisia tridentata Nutt. and weakly-flattened, recumbent, sericeous (Asteraceae); deposited in the AMNH. setae, the elongate oval body shape, anten- Paratypes. -3 males, 9 females, same data nal segment one black, segment two black as holotype (AMNH, OSU, USNM). USA: basally and progressively lighter distally, and Nevada: Washoe Co.: 7 mi. W of Vya to- the form of the male genitalia, the vesica ward Decarville, 1800 m, July 2, 1979, R. with two relatively short apical spines of T. Schuh and B. M. Massie, ex Artemisia nearly equal length, the more ventral spine sp. (not tridentata) (Asteraceae) (AMNH), with a small notch at the apex. Europiella 5 males, 9 females. Oregon: Jackson Co.: carvalhoi is most similar among described Ashland, 6500 ft., September 24, 1968, species in body form and coloration to E. Oman (OSU), 4 males, 13 females; 1 mi. unipuncta Knight which has more differ- below summit of Mt. Ashland, 6500 ft., ently formed vesical spines and E. signi- September 24, 1968, J. D. Lattin (OSU), 7 cornis Knight which has totally pale anten- males, 7 females. Umatilla Co.: 2 mi. W nae. Tollgate, 4600 ft., August 4, 1986, Schuh, Description. -Male: Dorsum, including Schwartz, and Stonedahl, Artemisia triden- membrane pale, white to tan, tinged with tata Nutt. (Asteraceae) (AMNH, USNM), green; neck (when visible) castaneous; tho- 32 males, 59 females. Wheeler Co.: 2 mi. racic pleuron and venter and pregenital ab- W Mitchell on Rt. 26, June 22, 1979, M. dominal segments weakly infuscate, pygo- D. Schwartz, G. M. Stonedahl, ex Artemisia phore somewhat darker; antennal segment tridentata (AMNH), 7 males, 20 females; one black, segment two black proximally, Mitchell, June 22, 1979, R. T. Schuh, ex progressively lighter distally, segments three Artemisia tridentata wyomingensis and four weakly infuscate; legs weakly in- (AMNH), 2 males, 8 females. fuscate, tibiae narrowly black at femoral ar- Hosts.-Artemisia tridentata, Artemisia ticulation, tibial spines black with black tridentata wyomingensis (Asteraceae). bases; femora with a few diffuse scattered infuscate areas. Dorsum densely clothed with reclining Europiella consors (Uhler), pale simple setae and weakly flattened, re- NEW COMBINATION cumbent, sericeous setae. Maurodactylus consors Uhler 1895: 53 (n. Body elongate ovoid; frons bulging; eyes sp., desc.). globular, touching anterior margin of pro- Europiella fuscicornis Knight 1969: 82 (n. notum; posterior margin of vertex sinuous. sp., desc., host). NEW SYNONYMY. Pygophore very large, occupying well Europiella basicornis Knight 1970: 230 (n. more than halfofabdomen; vesica with two sp., desc.). NEW SYNONYMY. relatively short apical spines ofnearly equal length, the more ventral with a small notch Distribution.-Western North America. at the apex (Figs. 1-3); left paramere as in Hosts.-Artemisia dracunculus, A. filifol- Fig. 5; right paramere as in Fig. 4. ia, A. ludoviciana ludoviciana, A. tridentata, 388 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON

Chrysothamnus nauseosus, C. parryi, C. vis- imens ... collected on Bigelovia near Amer- cidiflorus, C. sp. (Asteraceae). ican Fork, June 22." We have located in the Notes.-Uhler (1895) described consors USNM 2 specimens, a male and a female, on the basis of "one specimen, a male ... that bear the correct data. A third specimen, from Colorado." We have examined a spec- a female, was actually a misidentified ex- imen in the USNM which fits Uhler's orig- ample of unipuncta Knight. inal description, and bears the label "Mau- Comparison of the male genitalia of pi- rodactylus consors col. Uhler." We therefore losula with those of the holotype ofMicro- have assumed that it is the specimen orig- phylidea pallens Knight indicates that the inally examined by Uhler. Dissections of two are synonymous. Comparison of the specimens which are almost certainly con- male genitalia ofpilosula with those of to- specific indicate that this is actually a spe- potypic males collected on the same date as cies of Europiella rather than belonging to the holotype ofEuropiella albata Knight in- the European Maurodactylus Reuter, and it dicates that the two are synonymous. Com- is therefore transferred. parison ofpilosula with E.flavicornis Knight Examination of the holotypes of Euro- indicates that the two are synonymous. piella basicornis Knight and E. fuscicornis Uhler (1895) described Tuponia subniti- Knight, a large amount of additional ma- da from Steamboat Springs, Colorado, on terial, and many dissections ofthe male gen- the basis oftwo specimens. We searched the italia, indicate that these two nominal spe- collections of the USNM, but were unable cies represent the same species and are both to find any specimens which appeared to junior synonyms of consors. have been examined by Uhler. We did find, Europiella decolor (Uhler), see above. however, specimens from Ft. Garland, Col- orado, collected by H. H. Knight and la- Europiella pilosula (Uhler) belled by him as "Tuponia subnitida Uhler, Atomoscelis pilosulus Uhler 1893: 377 (n. compared with type." These specimens agree closely with the Uhler's description of sub- sp., desc.). fact that Tuponia subnitida Uhler 1895: 45 (n. sp., nitida with the exception of the desc.). NEW SYNONYMY. antennal segment one is not "blackish" as Psalluspilosulus: Van Duzee 1915: 118 (list, indicated by Uhler, but rather only weakly All other n. comb.). brown. attributes, including size, pallens Knight 1968: 29 (n. shape, and distribution agree, however, and Microphylidea we are therefore treating subnitida as a syn- sp., desc.). NEW SYNONYMY. Europiella pilosula: Knight 1968: 44 (n. onym ofpilosula. comb., note). Europiella flavicornis Knight 1969: 82 (n. Europiella signicornis Knight sp., desc., host). NEW SYNONYMY. Europiella signicornis Knight 1969: 84 (n. Europiella pallida Knight 1969: 83 (n. sp., sp., desc., host). desc., host). (Syn. by Stonedahl 1990:79). Distribution. -Northern Arizona. Europiella albata Knight 1969: 85 (n. sp., desc.). NEW SYNONYMY. Host. -Artemisia sp. (Asteraceae). Distribution. -Interior western United Europiella stigmosa (Uhler) States. Figs. 12-14 Hosts. -Artemisia flifolia, A. tridentata Agalliastes stigmosus Uhler 1893: 379 (n. (Asteraceae). sp., desc.). Notes. -Uhler (1893) described Atomos- Europiella stigmosa: Reuter 1909: 84 (disc.); celispilosulus on the basis of "Several spec- Knight 1968: 43 (in part, dist., host). VOLUME 97, NUMBER 2 389

Distribution. Western United States. Species transferred from Europiella to Hosts. -Artemisia californica; Artemisia Megalopsallus Knight 1927 tridentata (Asteraceae). albipubescens Knight 1968 (Europiella), Notes. -In his original description, Uhler NEW COMBINATION said that "Specimens [ofstigmosa] were col- arizonae Knight 1968 (Europiella), NEW lected at American Fork, June 22." We lo- COMBINATION cated three specimens in the USNM which balli Knight 1968 (Europiella), NEW bearthese data. One ofthese, a female, bears COMBINATION an identification label which appears to have brevicornis Knight 1968 (Europiella), NEW been written by Uhler as well as a "lecto- COMBINATION type" label affixed by H. G. Barber, al- franseriae Knight 1969 (Europiella), NEW though we can find no evidence that Barber COMBINATION ever published this designation. The other grayiae Knight 1968 (Europiella), NEW two, both males, were in the collection of COMBINATION H. H. Knight and bear his identification la- humeralis Van Duzee 1923 (Sthenarus), bels, but there is no direct evidence that they NEW COMBINATION are part ofthe material examined by Uhler. knowltoni Knight 1970 (Europiella), NEW We have not designated a lectotype, but nei- COMBINATION ther do we consider the identity ofstigmosa lycii Knight 1968 (Europiella), NEW to be in question. COMBINATION montanae Knight 1968 (Europiella), NEW Europiella umbrina Reuter COMBINATION monticola Knight 1968 (Europiella), NEW Europiella umbrina Reuter 1909: 85 (n. sp., COMBINATION desc.). multipunctipes Knight 1970 (Europiella), Europiella similis Knight 1969: 81 (n. sp., NEW COMBINATION desc., host). NEW SYNONYMY. nicholi Knight 1968 (Europiella), NEW Europiella stigmosa: Knight 1968: 43 (mis- COMBINATION identification). nigrofemoratus Knight 1968 (Europiella), Distribution. -Western United States. NEW COMBINATION Hosts. -Artemisia californica, A. nova, A. punctipes Knight 1968 (Europiella), NEW tridentata (Asteraceae). COMBINATION Notes. -Henry and Wheeler (1988) listed rubicornis Knight 1968 (Europiella, NEW umbrina Reuter as a synonym ofstigmosa. COMBINATION Comparison of the types of stigmosa, um- rufiventris Knight 1968 (Europiella), NEW brina, and similis Knight, and the male gen- COMBINATION italia ofthese nominal species, indicates that sarcobati Knight 1969 (Europiella), NEW is stigmosa, but that COMBINATION umbrina distinct from sparsus Knight 1969 (Europiella), NEW similis is synonymous with umbrina. COMBINATION stitti Knight 1968 (Europiella), NEW Europiella unipuncta Knight COMBINATION Europiella unipuncta Knight 1968: 44 (n. viridiventris Knight 1968 (Europiella), NEW sp., desc., host). COMBINATION Distribution. -Interior western United States. PALEARcrc SPECIES Host. -Artemisia tridentata (Asteraceae). Europiella albipennis (Fallen), see above. 390 PROCEEDINGS OF THE ENTOMOLOGICAL SOCIETY OF WASHINGTON

Europiella alpina (Reuter 1875), Europiella gilva (Kulik), NEW COMBINATION NEW COMBINATION Plagiognathus alpinus Reuter 1875: 56 (n. Plagiognathus gilvus Kulik 1965: 155 (n. sp., descr.); Kerzhner 1964: 761 (key, sp., desc., figs.); Kerzhner 1979: 54 (key, figs.). syn., figs.); Kerzhner 1988a: 855 (key, Psallus pallidus Reuter 1880: 24 (n. sp., figs.). descr.) (syn. by Wagner 1958). Chlorilluspilosus Wagner 1969b: 34 (n. sp., Plagiognathus alpinus f. nigrescens Stichel desc., figs.). 1934: 279 (n. form, key). Plagionathus alpinus f. simplex Stichel 1956: Distribution.-Russia (Primorsk Tern- 332 (n. form, key). tory). Records from Korea (Josifov and Psallus (Psallus) alpinus: Wagner 1956a: 298 Kerzhner 1972) refer to E. livida. (disc., figs.); Wagner 1958: 325 (syn.). Host. -Rabdosia excisa (Lamiaceae). Plagiognathus (Chlorillus) alpinus: Ker- zhner 1962: 384 (disc., figs.). Europiella kiritschenkoi (Kulik), Chlorillus alpinus: Wagner 1963b: 155 NEW COMBINATION (disc.,); Wagner 1975: 13 (desc., key, figs.). Plagiognathus kiritschenkoi Kulik 1975: 587 (n. sp., desc., figs.); Kerzhner 1979: 55 Distribution. -Europe, western Palearc- (key, figs.); Kerzhner 1988a: 855 (key, tic Asia. Contrary to Wagner (1975), this Li and Zheng 1991: 90 (key, distr., species is not recorded from North Amer- figs.); ica. figs.). Host:-Mentha aquatica (Lamiaceae). Distribution.-Russia (Primorsk Tern- Europiella artemisiae (Becker), see above. tory); China (Heilongjiang). Host. -Artemisia sp. (Asteraceae). Europiella canoflavida (Qi and Nonnaizab), NEW COMBINATION Europiella leucopus (Kerzhner), Plagiognathus (Poliopterus) canoflavidus Qi NEW COMBINATION and Nonnaizab 1993: 31 (n. sp., desc., Plagiognathus leucopus Kerzhner 1979: 50 figs.). (n. sp., desc., figs.); Kerzhner 1988a: 854 Distribution. -China: Inner Mongolia. (key, figs.); Li and Zheng 1991: 90 (key, Note.-We have not examined speci- distr., figs.). mens of this species. It is probably synon- ymous with either E. decolor or E. arte- Distribution. -Russia (Primorsk Term- misiae, based on the published figures ofthe tory); China (Heilongjiang). male genitalia. Europiella livida (Reuter), Europiella decolor (Uhler), see above. NEW COMBINATION Europiella herbaalbae (Wagner), Plagiognathus lividus Reuter 1906: 73 (n. NEW COMBINATION sp., desc.); Kerzhner 1979: 51 (part; distr., Plagiognathus (Poliopterus) herbaalbae figs.); Kerzhner 1988a: 855 (key, figs.); Li Wagner 1969a: 13 (n. sp., desc., figs.); and Zheng 1991: 91 (key, distr., figs.). Wagner 1975: 29 (desc., key, figs.). Distribution.-Russia (Amur Province; Distribution.-Algeria, Libya, Tunisia. Primorsk Territory); Korea; eastern China Host. -Artemisia herba-alba (Astera- (to Sichuan). ceae). Host. -Artemisia gmelinii (Asteraceae). VOLUME 97, NUMBER 2 391

Europiella lividella (Kerzhner), Europiella ovatula (Wagner), NEW COMBINATION NEW COMBINATION Plagiognathus lividellus Kerzhner 1979: 51 Plagiognathus (Poliopterus) ovatulus Wag- (n. sp., desc., figs.); Kerzhner 1988a: 855 ner 1952b: 41 (n. sp., desc., figs.); Putsh- (key, figs.); Li and Zheng 1991: 90 (key, kov 1971: 35 (dist., host); Wagner 1975: distr., figs.). 31 (key, desc., figs.). Distribution.-Russia (Far East); China Distribution. -Croatia; Ukraine. (Heilongiang; Fujian, Ningxia). Hosts.-Helichrysum angustifolium, H. Hosts.-Artemisia montana, A. rubripes, arenarium (Asteraceae). and A. vulgaris (Asteraceae). Note. -This species does not differ from E. artemisiae in the structure of the aede- Europiella miyamotoi (Kerzhner), agus, but shows marked differences in size NEW COMBINATION and color, and because the host plants are Plagiognathus lividus (not Reuter 1906): also different, we regard it as a separate spe- Miyamoto 1969: 90 (desc., figs.); Ker- cies. It is closely related to the less well- zhner 1979: 54 (part). known E. tomentosa and possibly synony- Plagiognathus miyamotoi Kerzhner 1988a: mous with it. 955 (n. sp., key, figs.); Kerzhner 1988b: 64 (desc.). Europiella strawinskii Distribution.-Russia (Sakhalin and Ku- (Sienkiewicz), rile Islands); Japan. NEW COMBINATION Host. -Artemisia montana (= gigantea) Plagiognathus (Poliopterus) strawinskii (Asteraceae). Sienkiewicz 1986: 373 (n. sp., desc., figs.). Europiella moesta (Reuter), Distribution. -Romania. NEW COMBINATION Note. -Probably a synonym of E. deco- Plagiognathus albipennis var. moesta Reu- lor. ter 1906: 75 (n. var., desc.). Plagiognathus (Poliopterus) moestus: Wag- Europiella strigifemur ner 1954: 75 (n. stat., desc., figs.); Li and (Wagner), Zheng 1991: 90 (key, distr., figs.). NEW COMBINATION Distribution.-China: Sichuan. Collected Europiella (Poliopterus) strigifemur Wagner at high altitudes. 1964: 70 (n. sp., desc., figs.); Wagner 1975: Host.-Unknown. 28 (desc., key, figs.). Note. -This species is very close to E. Distribution. -Libya. artemisiae and does not differ from it in the Host. -Unknown. structure ofthe male genitalia, except in size. Note. -We examined only female speci- It is, however, markedly larger, very dark, mens ofthis species. They clearly belong to and strongly shining. the group of species closely related to Eu- ropiella albipennis. Europiella nigrocunealis (Putshkov), NEW COMBINATION Europiella tomentosa (Reuter), Plagiognathus (Poliopterus) nigrocunealis NEW COMBINATION Putshkov 1975: 283 (n. sp., desc.). Plagiognathus tomentosus Reuter 1888: 236 Distribution. -Armenia; Azerbaijan. (n. sp., desc.). Host. - Tanacetum vulgare (Asteraceae). Plagiognathus (Poliopterus) ovatulus (not 392 PROCEEDINGS OF THE ENTOMOL)GICAL SOCIETY OF WASHINGTON Wagner 1952b): Wagner and Weber 1964: position, and consequent possible second- 415 (desc.). ary homonymy ofP. decolor Lindberg with Plagiognathus (Poliopterus) tomentosus: Europiella decolor (Uhler) need re-exami- Wagner 1975: 31 (key, desc.). nation. See also notes under Psallus argy- rotrichus. Distribution.-Southern France; Sicily. Host.-Helichrysum spp. (Asteraceae). Plagiognathusflavipes Reuter 1875. This species is known from southwestern Europe, living on Lonicera. Wagner (1952a) COMMENTS ON ADDITONAL PALEARCrIC placed it in Plagiognathus (Poliopterus). We SPECIES examined about 10 specimens, including Psallus argyrotrichus Fieber 1861. some from Corsica, the type locality, iden- This taxon was described from Germany tified by Puton. The structure of the male and Spain. Wagner (1963a) examined four genitalia is typical of Plagiognathus sensu specimens from Spain belonging to the col- stricto. lections ofMeyer-Dur (now in the AMNH) Plagiognathus servadeii Wagner 1972. identified as P. argyrotrichus, designating one ofthem as the lectotype. He transferred Wagner (1972) described this species from argyrotrichus to Plagiognathus (Poliopterus) northern Italy and placed it in Plagiogna- and placed P. decolor Lindberg in synonymy thus (Poliopterus). Judging from the figure with it. We re-examined the material stud- of the right paramere accompanying the ied by Wagner and have determined that original description, it does not belong to the specimens are actually Europiella; the structure ofthe aedeagus will crotchi (Scott). Contrary to what Wagner not clearly place the species in either Eu- (1963a) indicated, the identification label ropiella or Plagiognathus, and we therefore attached to the specimens is written by provisionally leave the species in Plagiog- Meyer-Dur, not by Fieber. Although all of nathus sensu stricto. the specimens are badly rubbed, they have Plagiognathuspallescens Zheng and Li 1991. remnants of black setae on the hemelytra, which contradicts Fieber's original descrip- This species was described (Zheng and Li tion. It is clear that the specimens examined 1991) from China (Sichuan). The type series by Wagner do not belong to the type series (ofwhich we examined three specimens la- ofFieber and were misidentified. The iden- belled as paratypes), is mixed, as could be tity of P. argyrotrichus needs clarification. presumed from the great variability in the length, males 3.30-4.28, females 3.02-3.22. Plagiognathus bicolor (Jakovlev 1880). One male specimen labelled as a paratype This species, the only representative of fiom "Szechwan, 2000-2300 m, 1O.IX.1963," Plagiognathus (Zophocnemis) Kerzhner measures 4.5 mm in length and contradicts 1962, resembles some species ofEuropiella the original description regarding length and in coloration and size, but male genitalia collecting data; it bears a handwritten para- are typical of Plagiognathus sensu stricto, type label where the other two specimens and we therefore place it in that genus. available for our examination have printed labels. The genitalia ofthis specimen are of Plagiognathus decolor Lindberg 1934. the species figured with the original descrip- This species was described on the basis tion. A male from "Hsiao-chin" is 4.2 mm offour females from Spain (Lindberg 1934). long, has genitalia very different in structure Wagner (1963a) placed the taxon in syn- from the first discussed specimen, and is onymy with Psallus argyrotrichus Fieber and possibly not even congeneric with it. Nei- transferred argyrothichus to Plagiognathus ther ofthese specimens belong to Europiella (Poliopterus). The synonymy, taxonomic and could only be placed in Plagiognathus VOLUME 97, NUMBER 2 393 sensu stricto with difficulty. The third spec- Jakovlev, B. E. 1875. Bugs, Heteroptera, imen is a female measuring 3.2 mm in ofthe Astrakhan region. Bulletin de la Societe Im- periale des Naturalistes de Moscou 49(3): 145- length, from "Pao-shing, 950-1350 m, 174. [In Russian and German.] 1.VII. 1963," these data matching the orig- 1877. New bugs, Hemiptera Heteroptera, of inal description except for the date; it pos- the Astrakhan fauna. Second supplement. Bulletin sibly belongs to the group of species closely de la Societe Imperiale des Naturalistes de Moscou related to alpina. 52(2): 269-300. [In Russian and German.] . 1880. Bugs (Hemiptera, Heteroptera) ofRus- Salicarus bimaculatus Zheng and Li 1991. sia and neighboring countries. I. Trudy Russ. En- tomol. Obshch. 11: 200-220. This species was described from Sichuan, Josifov, M. 1974. Die Heteropteren der Bulgarischen China. Judging from the size variability in- Schwarzmeerkuiste. Izvestiya Instituta Zool. Muz., dicated in the original description (2.09-3.5 Sofia 39: 5-27. [In Bulgarian, German summary.] mm), it may be based on a mixed series. Josifov, M. and I. M. Kerzhner. 1972. Heteroptera aus Korea. I. Teil (Ochtendae, Gerridae, Saldidae, Clearly, this is not a species of Salicarus, Nabidae, Anthocoridae, Miridae, Tingidae und butjudging from the description closely re- Reduviidae). Annales Zoologici (Warsaw) 29: 147- sembles species closely related to albipen- 180. nis, especially E. artemisiae. Kerzhner, I. M. 1962. Materials on the taxonomy of capsid bugs (Hemiptera, Miridae) in the fauna of the USSR. Entomologicheskoye Obozreniye 41: LITERATuRE CITED 372-387. [In Russian.) Becker, A. 1864. Naturhistorische Mitteilungen. Bul- 1964. Fam. Miridae. In Bei-Bienko, G. Ya., letin de la Societe Imperiale des Naturalistes de ed., Keys to the ofthe European USSR. 1: Moscou 37(2): 477-493. 700-765. Moscow and Leningrad, Nauka. [In Bergroth, E. 1898. Ueber einige amerikanische Cap- Russian.] [ English translation: 1967, Israel Pro- siden. Wiener Entomologische Zeitung 17: 33-35. gram for Scientific Translation, Jerusalem, pp.913- Carvalho, J. C. M. 1955. Keys to the genera of Mir- 1003.] idae ofthe world. Boletim Museu Paraense Emilio 1978. Heteroptera of Saghalien and Kurile Goeldi 11(2): 1-151. Islands. Trudy Biol.-Pochv. Inst. Dal'nevost. 1955. Analecta miridologica: Miscellaneous Nauch. Tsentra Akad. Nauk SSSR (n. s.) 50(153): observations in some American museums and 31-57. [In Russian.] bibliography (Hemiptera). Revista Chilena de En- 1979. New Heteroptera from the Far East of tomologia 4: 221-227. the USSR. Trudy Zoologischeskogo Instituta Aka- Dahlbom, A. G. 1851. Anteckningar Nfver Insekter, demii Nauk SSSR 81: 14-65. [In Russian.] som blifvit observerade pa Gottland och i en del 1988a. Fam. Miridae. In Ler, P. A., ed., Keys afCalmare Lin, under sommaren 1850. Kongliga to the Insects of the USSR Far East. Vol. 2: 788- Svenska Vetenskaps-Akademiens Handligar 1850: 857. Leningrad, Nauka. [In Russian.] [Issued March 155-229. 1988]. Falltn, C. F. 1829. Hemiptera Sveciae. Sectio prior, 1988b. New and Little Known Heteroptera Hemelytrata. Officina Berlingiana, Londini Goth- from the USSR Far East. Vladivostok, Akademia orum. 188 pp. Nauk SSSR. 83 pp. (1987). [In Russian.] [Issued Fieber, F. X. 1860-1861. Die europaischen Hemip- April 1988]. tera. Halbfluigler (Rhynchota Heteroptera). Carl Kirkaldy, G. W. 1909. Hemiptera: Old and new. No. Gerold's Sohn, Wien. 444 pp. (pp. 1-112, 1860; 2. Canadian Entomologist 41: 388-392. pp. i-iv, 113-444, 1861). Knight, H. H. 1923. Family Miridae (Capsidae), pp. 1864. Neuere Entdeckungen in europaischen 422-658. In Britton, W. E., ed., The Hemiptera Hemipteren. Wiener Entomologische Monat- or Sucking Insects of Connecticut. Connecticut schrift 8: 65-86, 205-234, 321-335. Geological and Natural History Survey Bulletin Gillette, G. P. and C. F. Baker. 1895. A preliminary 34: 1-807. list of the Hemiptera of Colorado. Colorado Ag- 1925. A list of Miridae and Anthocoridae ricultural Experiment Station Bulletin 31 (Tech. from Alberta, Canada (Hemiptera). Canadian En- Ser. 1), 137 pp. tomologist 57: 181-182. Henry, T. J. and A. G. Wheeler, Jr. 1988. Family . 1927. Notes on the distribution and host Miridae Hahn, 1833, pp. 251-507. 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1964. Phymatopsallus, new genus and new Bihang Kongliga Vetenskaps-Akademien Han- species of from North America (Hemip- dlingar 3(1): 1-66. tera, Miridae). Iowa State Journal of Science 39: . 1878. Hemiptera Gymnocerata Europae. 127-152. HWmipteres Gymnockrates d'Europe, du bassin de 1968. Taxonomic review: Miridae ofthe Ne- la M6diterranee et de l'Asie russe, 1. Acta Socie- vada Test Site and the western United States. Brig- tatis Scientiarum Fennicae 13: 1-188, 8 pls. ham Young University Science Bulletin 9(3): 1- * 1880. Diagnoses Hemipterorum novorum. 282. II. Ofversigt Finska Vetenskaps-Societatens For- 1969. New species of Pronotocrepis Kngt., handlingar 22: 9-24. Europiella Reut., and Hesperocapsus Kngt., from * 1888. Descriptio speciei nova sicilianae ge- western United States (Miridae, Hemiptera). Iowa neris Plagiognathus (Hemiptera, Capsidae). Na- State Journal of Science 44: 79-91. turalista Siciliana 7: 236. 1970. Tannerocoris new genus, and new spe- * 1901. Capsidae rossicae descriptae. Ofversigt cies of Miridae (Hemiptera) from the western Finska Vetenskaps-Societatens Forhandlingar B United States. Great Basin Naturalist 30: 227- 43: 161-194. 231. * 1906. Capsidae in prov. Sz'tschwan Chinae Kulik, S. A. 1965. New species of capsid bugs (Het- a DD. G. Potanin et M. Beresowski collectae. An- eroptera, Miridae) from East Siberia and the Far nuaire Musee Zoologique Academie Imperiale East. Izv. Irkutsk. Sel'skokhoz. Inst. 25: 147-156. Sciences St. Petersburg 10: 1-81. [In Russian.] 1909. Bemerkungen uber nearktischen Cap- 1975. A new species of the genus Plagiog- siden nebst Beschreibung neuer Arten. Acta So- nathus (Heteroptera, Miridae) from the vicinity of cietatis Scientiarum Fennicae 36(2): 1-86. Ussuriysk. Entomologischeskoye Obozreniye 54: Sahlberg, J. 1920. Enumeratio Hemipterorum Het- 587-588. [In Russian.] eropterorum faunae Fennicae. Editio secunda auc- Li, H.-y. and L.-y. Zheng. 1991. Genus Plagiognathus ta et emendata. Bidrag Kainnedom Finlands Natur Fieber (Hemiptera: Miridae) from China. Acta Folk 79(2): 1-227. Scientiarum Naturalium Univsitatis Nan Kaiensis Sienkiewicz, I. 1986. Description de Plagiognathus 3: 88-97. [In Chinese with English summary.] (Poliopterus) strawinskii n. sp. (Heteroptera, Mir- Lindberg. H. 1934. Inveta entomologica itineris His- idae). Nouvelle Revue Entomologique (n.s.) 3: 373- panici et Maroccani, quod a. 1926 fecerunt Harald 378. et Hakan Lindberg. Societas Scientiarum Fennica, Stichel, W. 1934. Illustrierte Bestimmungstabellen Commentationes Biologicae 4(12): 23 pp. der Deutschen Wanzen, Heft 10: 275-306. W. Sti- Matsumura, S. 1917. Oyo Konchu-gaku I [Applied chel, Berlin. Entomology 1]. Cited from Second Rev. Edition, _ 1956. Illustrierte Besstmmungstabellen der 1920, 731 + 12 pp., 50pls. Wanzen. II. Europa. Vol. 2, Heft 11, pp. 321-352. Miyamoto, S. 1969. Notes on the species ofthe genus Privately printed, Berlin-Hermsdorf. Plagiognathus Fieber in Japan and Saghaline. Sie- Stonedahl, G. M. 1990. Revision and cladistic anal- boldia 4: 85-94. ysis of the Holarctic genus Atractotomus Fieber . 1977. On the scientific names concerning (Heteroptera: Miridae: Phylinae). Bulletin of the Japanese Heteroptera (3). Rostria 28: 231-234. [In American Museum ofNatural History 198: 1-88. Japanese.] Uhler, P. R. 1893. Summary ofthe collection of He- Puton, A. 1873. Notes pour servir a l'etude des Hem- miptera secured by Mr. E. A. Schwarz in Utah. ipteres. Annals de la Societe Entomologique de Proceeding ofthe Entomological Society ofWash- France (5)3: 11-26. ington 2: 366-385. Putshkov, V. G. 1971. On the ecology of some little- _ 1894. Observations upon the heteropterous known species of Heteroptera of the European Hemiptera of Lower California, with descriptions USSR. Communication IV. Miridae. Vestnik ofnew species. Procceding ofthe California Acad- Zoologii 1971(5): 30-35. [In Russian.] emy of Sciences, ser. 2, 4: 223-295. 1975. New species of Miridae from the Cau- _ 1895. Descriptions ofnew genera and species casus (Hepteroptera). Doklady Akademii Nauk of Hemiptera, In Gillette, G. P. and C. F. Baker, Ukrainskoi SSR (B) 1975(3): 281-284. [In Rus- A Preliminary List ofthe Hemiptera ofColorado. sian.] Colorado Agricultural Experiment Station Bulle- Qi, B. and Nonnaizab. 1993. New species and a newly tin 31 (Tech. Ser. 1), 137 pp. recorded species ofPlagiognathus Fieber ofChina Van Duzee, E. P. 1915. New genera and species of (Hemiptera: Miridae). Entomotaxonomia 15: 29- North American Hemiptera. Pomona Journal of 34. [In Chinese with English summary.] Entomology and Zoology 7: 109-121. Reuter, 0. M. 1875. Genera Cimicidarum Europae. _ 1916. Checklist of the Hemiptera (excepting VOLUME 97, NUMBER 2 395

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