The Early Origin of Feathers
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
The Wingtips of the Pterosaurs: Anatomy, Aeronautical Function and Palaeogeography, Palaeoclimatology, Palaeoecology Xxx (2015) Xxx Xxx 3 Ecological Implications
Our reference: PALAEO 7445 P-authorquery-v11 AUTHOR QUERY FORM Journal: PALAEO Please e-mail your responses and any corrections to: Article Number: 7445 E-mail: [email protected] Dear Author, Please check your proof carefully and mark all corrections at the appropriate place in the proof (e.g., by using on-screen annotation in the PDF file) or compile them in a separate list. Note: if you opt to annotate the file with software other than Adobe Reader then please also highlight the appropriate place in the PDF file. To ensure fast publication of your paper please return your corrections within 48 hours. For correction or revision of any artwork, please consult http://www.elsevier.com/artworkinstructions. We were unable to process your file(s) fully electronically and have proceeded by Scanning (parts of) your Rekeying (parts of) your article Scanning the article artwork Any queries or remarks that have arisen during the processing of your manuscript are listed below and highlighted by flags in the proof. Click on the ‘Q’ link to go to the location in the proof. Location in article Query / Remark: click on the Q link to go Please insert your reply or correction at the corresponding line in the proof Q1 Your article is registered as a regular item and is being processed for inclusion in a regular issue of the journal. If this is NOT correct and your article belongs to a Special Issue/Collection please contact [email protected] immediately prior to returning your corrections. Q2 Please confirm that given names and surnames have been identified correctly. -
Final Summary Report of the Research Project (ERB, 624732)
Final Summary Report of the Research Project (ERB, 624732): Introduction: Birds represent one of the most remarkable radiations in the history of life. The origin of flight and the diversification of birds have become classic ‘textbook’ examples for discussions about evolutionary models, biomechanics, the history of life, and phylogenetics. Much attention has focused on the discovery and description of new fossils, both derived theropods and basal birds: in particular, sediments of the Jehol Group in Liaoning and other provinces in China have doubled the number of Mesozoic bird taxa and have considerably expanded our knowledge of derived theropod dinosaurs. All this wealth of fossil data and new phylogenetic studies have established the relationships of basal bird groups, as well as their phylogenetic nesting among theropod dinosaurs, with dromaeosaurid and troodontid theropods as their closest sister groups. The mode and tempo of avian evolution have received much attention in recent years, thanks to the assembly of more complete datasets, the ongoing flow of new specimens and taxa, and the development of new techniques of macroevolutionary analysis. Recent studies revealed an avian radiation characterized by the gradual acquisition of apomorphies followed by accelerated (yet clade heterogeneous) rates of evolution. However, in the context of Coelurosauria (a diverse clade including T. rex and its kin, the ostrich-mimic ornithomimosaurs, and the sickle-clawed dromaeosaurids, among others), previous analyses have considered the skeleton as a whole, apart from studies focusing on single continuous traits. Here, I document the evolutionary dynamics of each anatomical region of the coelurosaurian skeleton. I aim at disentangling the relative role of each body part in the evolutionary dynamics of Coelurosauria. -
From the Late Cretaceous of Patagonia
AMEGHINIANA (Rev. Asoc. Paleontol. Argent.) - 40 (1): 119-122. Buenos Aires, 30-03-2003 ISSN0002-7014 NOTA PALEONTOLÓGICA A new specimen of Patagonykus puertai (Theropoda: Alvarezsauridae) from the Late Cretaceous of Patagonia Luis M. CHIAPPE1and Rodolfo A. CORIA2 Introduction Systematic paleontology With their short and robust forelimbs, highly ki- THEROPODA netic skulls, and a host of unique skeletal features, COELUROSAURIA alvarezsaurids are highly specialized Mesozoic ALVAREZSAURIDAEBonaparte, 1991 coelurosaurian theropods known from the Late Cretaceous of Asia, North, and South America Genus PatagonykusNovas, 1997a (Chiappe et al., 2002). Despite abundant and well- preserved material from the Gobi Desert of Patagonykussp. cf. P. puertaiNovas, 1997a Mongolia, the phylogenetic relationships of al- Figures 1 and 2 varezsaurids have remained unclear (Chiappe et Material. MCF-PVPH-102 (Museo Carmen Funes, al., 2002; Novas and Pol, 2002; Suzuki et al., 2002). Paleontología de Vertebrados; Plaza Huincul, The most basal alvarezsaurids are from South Neuquén, Argentina), an articulated phalanx 1 and 2 America and are limited to just two taxa from the (ungual) of the left manual digit I. Patagonian province of Neuquén (Argentina), Geographical provenance and geological horizon. Alvarezsaurus calvoi (Bonaparte, 1991) and Northern shore of Los Barriales lake, approximately Patagokykus puertai (Novas, 1997a), which although 80 km north Plaza Huincul. Because the specimen poorly represented are important because they was not collected by professionals, no precise strati- have been consistently interpreted as the most ple- graphic information is available; nonetheless, only siomorphic members of the group (Novas, 1996; the Late Cretaceous Neuquén Group outcrops in this Chiappe et al., 1998). area. Also, considering that the holotype specimen of Here we report on a second specimen of Patagonykus puertaiwas found in beds attributed to Patagonykus, which we assign to the species P. -
On Two Pterosaur Humeri from the Tendaguru Beds (Upper Jurassic, Tanzania)
“main” — 2009/10/20 — 22:40 — page 813 — #1 Anais da Academia Brasileira de Ciências (2009) 81(4): 813-818 (Annals of the Brazilian Academy of Sciences) ISSN 0001-3765 www.scielo.br/aabc On two pterosaur humeri from the Tendaguru beds (Upper Jurassic, Tanzania) FABIANA R. COSTA and ALEXANDER W.A. KELLNER Museu Nacional, Universidade Federal do Rio de Janeiro, Departamento de Geologia e Paleontologia Quinta da Boa Vista s/n, São Cristóvão, 20940-040 Rio de Janeiro, RJ, Brasil Manuscript received on August 17, 2009; accepted for publication on September 30, 2009; contributed by ALEXANDER W.A. KELLNER* ABSTRACT Jurassic African pterosaur remains are exceptionally rare and only known from the Tendaguru deposits, Upper Jurassic, Tanzania. Here we describe two right humeri of Tendaguru pterosaurs from the Humboldt University of Berlin: specimens MB.R. 2828 (cast MN 6661-V) and MB.R. 2833 (cast MN 6666-V). MB.R. 2828 consists of a three- dimensionally preserved proximal portion. The combination of the morphological features of the deltopectoral crest not observed in other pterosaurs suggests that this specimen belongs to a new dsungaripteroid taxon. MB.R. 2833 is nearly complete, and because of a long and round proximally placed deltopectoral crest it could be referred to the Archaeopterodactyloidea. It is the smallest pterosaur from Africa and one of the smallest flying reptiles ever recorded. These specimens confirm the importance of the Tendaguru deposits for the Jurassic pterosaur record. This potential, however, has to be fully explored with more field work. Key words: Tendaguru, Tanzania, Africa, Upper Jurassic, Pterosauria. INTRODUCTION in providing isolated remains up to now (Kellner and Mader 1997, Wellnhofer and Buffetaut 1999, Mader Africa shows a great potential for pterosaur material and Kellner 1999). -
Los Restos Directos De Dinosaurios Terópodos (Excluyendo Aves) En España
Canudo, J. I. y Ruiz-Omeñaca, J. I. 2003. Ciencias de la Tierra. Dinosaurios y otros reptiles mesozoicos de España, 26, 347-373. LOS RESTOS DIRECTOS DE DINOSAURIOS TERÓPODOS (EXCLUYENDO AVES) EN ESPAÑA CANUDO1, J. I. y RUIZ-OMEÑACA1,2 J. I. 1 Departamento de Ciencias de la Tierra (Área de Paleontología) y Museo Paleontológico. Universidad de Zaragoza. 50009 Zaragoza. [email protected] 2 Paleoymás, S. L. L. Nuestra Señora del Salz, 4, local, 50017 Zaragoza. [email protected] RESUMEN La mayoría de los restos fósiles de dinosaurios terópodos de España son dientes aislados y escasos restos postcraneales. La única excepción es el ornitomimosaurio Pelecanimimus polyodon, del Barremiense de Las Hoyas (Cuenca). Hay registro de terópodos en el Jurásico superior (Oxfordiense superior-Tithónico inferior), en el tránsito Jurásico-Cretácico (Tithónico superior- Berriasiense inferior) y en todos los pisos del Cretácico inferior, con excepción del Valanginiense. En el Cretácico superior únicamente hay restos en el Campaniense y Maastrichtiense. La mayor parte de las determinaciones son demasiado generales, lo que impide conocer algunas de las familias que posiblemente estén representadas. Se han reconocido: Neoceratosauria, Baryonychidae, Ornithomimosauria, Dromaeosauridae, además de terópodos indeterminados, y celurosaurios indeterminados (dientes pequeños sin dentículos). La mayoría de los restos son de Maniraptoriformes, siendo especialmente abundantes los dromeosáuridos. Las únicas excepciones son por el momento, el posible Ceratosauria del Jurásico superior de Asturias, los barionícidos del Hauteriviense-Barremiense de Burgos, Teruel y La Rioja, el posible carcharodontosáurido del Aptiense inferior de Morella y el posible abelisáurido del Campaniense de Laño. Además hay algunos terópodos incertae sedis, como los "paronicodóntidos" (entre los que se incluye Euronychodon), y Richardoestesia. -
Analyzing Pterosaur Ontogeny and Sexual Dimorphism with Multivariate Allometry Erick Charles Anderson [email protected]
Marshall University Marshall Digital Scholar Theses, Dissertations and Capstones 2016 Analyzing Pterosaur Ontogeny and Sexual Dimorphism with Multivariate Allometry Erick Charles Anderson [email protected] Follow this and additional works at: http://mds.marshall.edu/etd Part of the Animal Sciences Commons, Ecology and Evolutionary Biology Commons, and the Paleontology Commons Recommended Citation Anderson, Erick Charles, "Analyzing Pterosaur Ontogeny and Sexual Dimorphism with Multivariate Allometry" (2016). Theses, Dissertations and Capstones. 1031. http://mds.marshall.edu/etd/1031 This Thesis is brought to you for free and open access by Marshall Digital Scholar. It has been accepted for inclusion in Theses, Dissertations and Capstones by an authorized administrator of Marshall Digital Scholar. For more information, please contact [email protected], [email protected]. ANALYZING PTEROSAUR ONTOGENY AND SEXUAL DIMORPHISM WITH MULTIVARIATE ALLOMETRY A thesis submitted to the Graduate College of Marshall University In partial fulfillment of the requirements for the degree of Master of Science in Biological Sciences by Erick Charles Anderson Approved by Dr. Frank R. O’Keefe, Committee Chairperson Dr. Suzanne Strait Dr. Andy Grass Marshall University May 2016 i ii ii Erick Charles Anderson ALL RIGHTS RESERVED iii Acknowledgments I would like to thank Dr. F. Robin O’Keefe for his guidance and advice during my three years at Marshall University. His past research and experience with reptile evolution made this research possible. I would also like to thank Dr. Andy Grass for his advice during the course of the research. I would like to thank my fellow graduate students Donald Morgan and Tiffany Aeling for their support, encouragement, and advice in the lab and bar during our two years working together. -
Theory of the Growth and Evolution of Feather Shape
30JOURNAL R.O. PRUMOF EXPERIMENTAL AND S. WILLIAMSON ZOOLOGY (MOL DEV EVOL) 291:30–57 (2001) Theory of the Growth and Evolution of Feather Shape RICHARD O. PRUM* AND SCOTT WILLIAMSON Department of Ecology and Evolutionary Biology, and Natural History Museum, University of Kansas, Lawrence, Kansas 66045 ABSTRACT We present the first explicit theory of the growth of feather shape, defined as the outline of a pennaceous feather vane. Based on a reanalysis of data from the literature, we pro- pose that the absolute growth rate of the barbs and rachis ridges, not the vertical growth rate, is uniform throughout the follicle. The growth of feathers is simulated with a mathematical model based on six growth parameters: (1) absolute barb and rachis ridge growth rate, (2) angle of heli- cal growth of barb ridges, (3) initial barb ridge number, (4) new barb ridge addition rate, (5) barb ridge diameter, and (6) the angle of barb ramus expansion following emergence from the sheath. The model simulates growth by cell division in the follicle collar and, except for the sixth param- eter, does not account for growth by differentiation in cell size and shape during later keratiniza- tion. The model can simulate a diversity of feather shapes that correspond closely in shape to real feathers, including various contour feathers, asymmetrical feathers, and even emarginate prima- ries. Simulations of feather growth under different parameter values demonstrate that each pa- rameter can have substantial, independent effects on feather shape. Many parameters also have complex and redundant effects on feather shape through their influence on the diameter of the follicle, the barb ridge fusion rate, and the internodal distance. -
Onetouch 4.0 Scanned Documents
/ Chapter 2 THE FOSSIL RECORD OF BIRDS Storrs L. Olson Department of Vertebrate Zoology National Museum of Natural History Smithsonian Institution Washington, DC. I. Introduction 80 II. Archaeopteryx 85 III. Early Cretaceous Birds 87 IV. Hesperornithiformes 89 V. Ichthyornithiformes 91 VI. Other Mesozojc Birds 92 VII. Paleognathous Birds 96 A. The Problem of the Origins of Paleognathous Birds 96 B. The Fossil Record of Paleognathous Birds 104 VIII. The "Basal" Land Bird Assemblage 107 A. Opisthocomidae 109 B. Musophagidae 109 C. Cuculidae HO D. Falconidae HI E. Sagittariidae 112 F. Accipitridae 112 G. Pandionidae 114 H. Galliformes 114 1. Family Incertae Sedis Turnicidae 119 J. Columbiformes 119 K. Psittaciforines 120 L. Family Incertae Sedis Zygodactylidae 121 IX. The "Higher" Land Bird Assemblage 122 A. Coliiformes 124 B. Coraciiformes (Including Trogonidae and Galbulae) 124 C. Strigiformes 129 D. Caprimulgiformes 132 E. Apodiformes 134 F. Family Incertae Sedis Trochilidae 135 G. Order Incertae Sedis Bucerotiformes (Including Upupae) 136 H. Piciformes 138 I. Passeriformes 139 X. The Water Bird Assemblage 141 A. Gruiformes 142 B. Family Incertae Sedis Ardeidae 165 79 Avian Biology, Vol. Vlll ISBN 0-12-249408-3 80 STORES L. OLSON C. Family Incertae Sedis Podicipedidae 168 D. Charadriiformes 169 E. Anseriformes 186 F. Ciconiiformes 188 G. Pelecaniformes 192 H. Procellariiformes 208 I. Gaviiformes 212 J. Sphenisciformes 217 XI. Conclusion 217 References 218 I. Introduction Avian paleontology has long been a poor stepsister to its mammalian counterpart, a fact that may be attributed in some measure to an insufRcien- cy of qualified workers and to the absence in birds of heterodont teeth, on which the greater proportion of the fossil record of mammals is founded. -
Federal Register/Vol. 78, No. 156/Tuesday, August 13, 2013
49186 Federal Register / Vol. 78, No. 156 / Tuesday, August 13, 2013 / Rules and Regulations Black grouper, Mycteroperca bonaci individual fishing quota (IFQ) system. consistent with the terminology used in Yellowmouth grouper, Mycteroperca The new tilefish IFQ program became rest of that regulatory paragraph. The interstitialis effective November 1, 2009. After 3 introductory text on IFQ transfer Gag, Mycteroperca microlepis years of operation, it has become applications has been adjusted to more Scamp, Mycteroperca phenax apparent that some of the implementing Yellowfin grouper, Mycteroperca venenosa clearly explain why applications for Serranidae—Sea Basses regulations need to be clarified, permanent transfers are due before Black sea bass, Centropristis striata corrected, or modified to better reflect September 1, while applications for Sparidae—Porgies the intent of Amendment 1. temporary transfers are due before Jolthead porgy, Calamus bajonado On March 28, 2013, NMFS published October 10. Saucereye porgy, Calamus calamus a proposed rule in the Federal Register Whitebone porgy, Calamus leucosteus (78 FR 18947) proposing several minor Comments and Responses Knobbed porgy, Calamus nodosus corrections, clarifications, and Red porgy, Pagrus pagrus modifications to the regulations NMFS received no comments on the Scup, Stenotomus chrysops implementing Amendment 1. proposed rule. The following species are designated as Comments on the proposed rule were ecosystem component species: Classification Cottonwick, Haemulon melanurum accepted through April 29, 2013. No Bank sea bass, Centropristis ocyurus comments were received on the The Administrator, Northeast Region, Rock sea bass, Centropristis philadelphica measures, and the measures, as NMFS, determined that this final rule is Longspine porgy, Stenotomus caprinus proposed, are implemented by this final necessary for the management of the Ocean triggerfish, Canthidermis sufflamen rule. -
Magnificent Magpie Colours by Feathers with Layers of Hollow Melanosomes Doekele G
© 2018. Published by The Company of Biologists Ltd | Journal of Experimental Biology (2018) 221, jeb174656. doi:10.1242/jeb.174656 RESEARCH ARTICLE Magnificent magpie colours by feathers with layers of hollow melanosomes Doekele G. Stavenga1,*, Hein L. Leertouwer1 and Bodo D. Wilts2 ABSTRACT absorption coefficient throughout the visible wavelength range, The blue secondary and purple-to-green tail feathers of magpies are resulting in a higher refractive index (RI) than that of the structurally coloured owing to stacks of hollow, air-containing surrounding keratin. By arranging melanosomes in the feather melanosomes embedded in the keratin matrix of the barbules. barbules in more or less regular patterns with nanosized dimensions, We investigated the spectral and spatial reflection characteristics of vivid iridescent colours are created due to constructive interference the feathers by applying (micro)spectrophotometry and imaging in a restricted wavelength range (Durrer, 1977; Prum, 2006). scatterometry. To interpret the spectral data, we performed optical The melanosomes come in many different shapes and forms, and modelling, applying the finite-difference time domain (FDTD) method their spatial arrangement is similarly diverse (Prum, 2006). This has as well as an effective media approach, treating the melanosome been shown in impressive detail by Durrer (1977), who performed stacks as multi-layers with effective refractive indices dependent on extensive transmission electron microscopy of the feather barbules the component media. The differently coloured magpie feathers are of numerous bird species. He interpreted the observed structural realised by adjusting the melanosome size, with the diameter of the colours to be created by regularly ordered melanosome stacks acting melanosomes as well as their hollowness being the most sensitive as optical multi-layers. -
The Molecular Evolution of Feathers with Direct Evidence from Fossils
The molecular evolution of feathers with direct evidence from fossils Yanhong Pana,1, Wenxia Zhengb, Roger H. Sawyerc, Michael W. Penningtond, Xiaoting Zhenge,f, Xiaoli Wange,f, Min Wangg,h, Liang Hua,i, Jingmai O’Connorg,h, Tao Zhaoa, Zhiheng Lig,h, Elena R. Schroeterb, Feixiang Wug,h, Xing Xug,h, Zhonghe Zhoug,h,i,1, and Mary H. Schweitzerb,j,1 aChinese Academy of Sciences Key Laboratory of Economic Stratigraphy and Palaeogeography, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, Nanjing 210008, China; bDepartment of Biological Sciences, North Carolina State University, Raleigh, NC 27695; cDepartment of Biological Sciences, University of South Carolina, Columbia, SC 29205; dAmbioPharm Incorporated, North Augusta, SC 29842; eInstitute of Geology and Paleontology, Lingyi University, Lingyi City, 27605 Shandong, China; fShandong Tianyu Museum of Nature, Pingyi, 273300 Shandong, China; gCAS Key Laboratory of Vertebrate Evolution and Human Origins of the Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 100044 Beijing, China; hCenter for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 100044 Beijing, China; iCollege of Earth and Planetary Sciences, University of Chinese Academy of Sciences, 100049 Beijing, China; and jNorth Carolina Museum of Natural Sciences, Raleigh, NC 27601 Contributed by Zhonghe Zhou, December 15, 2018 (sent for review September 12, 2018; reviewed by Dominique G. Homberger and Chenxi Jia) Dinosaur fossils possessing integumentary appendages of various feathers in Anchiornis, barbules that interlock to form feather morphologies, interpreted as feathers, have greatly enhanced our vanes critical for flight have not been identified yet (12). -
Running Birds from Quail to Ostrich Prioritise Leg Safety and Economy
© 2014. Published by The Company of Biologists Ltd | The Journal of Experimental Biology (2014) 217, 3786-3796 doi:10.1242/jeb.102640 RESEARCH ARTICLE Don’t break a leg: running birds from quail to ostrich prioritise leg safety and economy on uneven terrain Aleksandra V. Birn-Jeffery1,*,‡, Christian M. Hubicki2,‡, Yvonne Blum1, Daniel Renjewski2, Jonathan W. Hurst2 and Monica A. Daley1,§ ABSTRACT Daley, 2012; Dial, 2003; Jindrich et al., 2007; Rubenson et al., Cursorial ground birds are paragons of bipedal running that span a 2004). These athletes span the broadest body mass range among 500-fold mass range from quail to ostrich. Here we investigate the extant bipeds, over 500-fold from quail to ostrich. Birds thus provide task-level control priorities of cursorial birds by analysing how they a natural animal model for understanding the functional demands of negotiate single-step obstacles that create a conflict between body striding bipedalism and how these demands change with body size stability (attenuating deviations in body motion) and consistent leg (Gatesy and Biewener, 1991; Hutchinson and Garcia, 2002; Roberts force–length dynamics (for economy and leg safety). We also test the et al., 1998a). hypothesis that control priorities shift between body stability and leg Here, we ask two questions fundamental to locomotor behaviour: safety with increasing body size, reflecting use of active control to (1) what are the task-level leg control priorities of running animals; overcome size-related challenges. Weight-support demands lead to and (2) how do these priorities vary with terrain and body size? a shift towards straighter legs and stiffer steady gait with increasing Running animals must control their legs to balance numerous, body size, but it remains unknown whether non-steady locomotor sometimes conflicting, task-level demands including minimising priorities diverge with size.